Citation
The caddis flies of Alachua County, with notes on those of Florida

Material Information

Title:
The caddis flies of Alachua County, with notes on those of Florida
Creator:
Nirenberg, Marshall Warren
Publisher:
Marshall Nirenberg
Publication Date:
Language:
English

Subjects

Subjects / Keywords:
Adult insects ( jstor )
Counties ( jstor )
Creeks ( jstor )
Female animals ( jstor )
Genitalia ( jstor )
Insects ( jstor )
Larvae ( jstor )
Pupae ( jstor )
Species ( jstor )
Streams ( jstor )
Poe Spring ( local )

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University of Florida
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University of Florida
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Copyright [name of dissertation author]. Permission granted to the University of Florida to digitize, archive and distribute this item for non-profit research and educational purposes. Any reuse of this item in excess of fair use or other copyright exemptions requires permission of the copyright holder.
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021604259 ( alephbibnum )
13213009 ( oclc )

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Full Text












THE CADDIS FLIES OF ALACHUA COUNTY,

WITH NOTES ON THOSE OF FLORIDA












By
MARSHALL NIRENBERG











A THESIS PRESENTED TO THE GRADUATE COUNCIL OF
THE UNIVERSITY OF FLORIDA
IN PARTIAL FULFILMENT OF THE REQUIREMENTS FOR THE
DEGREE OF MASTER OF SCIENCE










UNIVERSITY OF FLORIDA
February, 1952


















TABLE OF CONTENTS




X Inxtroducticn... R*.. ** e*,t*** @ *l41

/clmcTn~e^^fryentSe **************************9



4i Preservetico u es n ClrreFrgor,,e,;*..,, *...,...IS.1



64 Key to rF.uali L.* **.ii; .**00 ****. v *****.** 19

74 Armotated Li8..*...*......,4*.** *****29

84 unDsrVPd Prolftzs'**U *****we**>* f ** **i v n rl61



10 R eferences,.<........, ... ..... .. ............ .I1G

114 Chlok List of the Ceddis Flies of Florida

and IBdexi..,..,.,********. ,** .. ******.172

12* Biographiorl l .te....,. ,. .,.. ,,,......,,.,o..177





-14


IITRODUCTIO'T


Historically speaking, some of the caddie flies of the south-

east were described at a relatively early date, about one hundred

years ago. At that time Francis Walker (1852 and 1860) of the British

museumm described twenty species from Georgia which were sent to him

by Abbot, and.this constituted the earliest large collection made in

the southeast. The investigation of caddie flies in the south stag*

n.ted for the next eighty-five years, for esxamle, fifteen years ago,

two out of three species collected in the state would hrve been un-

described. Since then much incidental work has been done by visiting

trichopterists and by entomologists rorlrkin in allied fields,


Caddis flies are f-irly abundant, but as compared with other

states, Florida shows a general paucity in the number of species

present; a rnucity most probably due to lpck of lmowledge rather than

a lack of ceddia flies. One hundred and eighty-four species have

been found in Illinois, a thoroughly studied state. On the other hand,

only sixty-three species have been found in Florida, thirty-eight of

which occur in Alachua County. Some day when the state has been in-

vestit-ted a ruood deal more thoroughly, this number will undoubtedly

be pre.tly increased.


In the Trichoptera, as in other orders of insects, our know-

lpdae of the immiture stages has leted far behind that of the adults.

The larvae of many species of caddir flies amoim to occur in the state

still are unknown or unesaocinted.






-2-


Betten (1934),. alduf (1939), and Ross (1944), have all care-

fully and fully summarized the extensive literature deocribinc and

deteilin- the life histories and habits of caddis fliee. For this

reason, no further discussion of this subject will be included here.


The current status of the knowledge of cr.dis flies in Florida

can be stated concisely. The species are very incompletely known;

cornm-r-tively few larvae have been associated with the adults. The

larvae of some genera have never been found and. much more collecting

of adults is needed. Georr"r.hic variation of the Trichopterap of the

1Thited States has never been investie ted thoroughly. Little col-

lecting has been done in northwestern Florida and many new records

plus undescribed species should be found there. Obviously, there is

amle room for far more study.


This researrch problem was specifically limited to the caddie

flies of Alachua County; however, as I had access to material collec-

ted in northwestern Florida I decided to extend the scope of the

.ntida; to include all th.t was known end all that I could find out

Pbout the ceddie flies of Florida. This additional material consisted

of caddie flies collected by Dr. Lewis Berner in 1939 and 1940, and

also spPcinens which were in the collections of the University of

Florida.


7ttendin the study by the inclusion of this supplementary

nrterlpl -reatly broPdened the entire picture, but also, simultaneously,






-3-


complicated its for the oolleotions examined were sufficient to raise

a multitude of questions but were insufficient to answer may of them.

For eanry problem solved, two were discovered which ccvld not be ao-

p-rorad.


This report is the result of an investigation earrr.c onm for

two yearc-----from Fcriary, 1950 to Februlry, 19r6-----in .- '..ch time

I colleotec appro!rnPtely 10,000 speciiens and examirne an ,'l1tieonal

eight lrTndred inrdivdirrle vhioh were collected by other.


The purpose of this investigation is to present (1) an

annotated liaf of the caddis flies found in Alachua County, (2) a

oheck list of the oaddia flioj of Florida, (3) keys for the identifi-

cation of the larvae and adults found in Florida, vnd (4) a biblio-

graphy which inolides all references to Florida ead'li fliec. In

Ohort, this investigetion has two Frir~ry rpirposes: to report Cn the

caddie flies of Alatohna County and to gather together all tha-i: is

Imown about the crodiv flies of Florida. As such this report mirj.t

well aorve as a foundation for a future study of the oanidii flios of

Florida,


On the basis of the information gathered in this study, which

of necessity is very incomplete as collections from parts of Florida

other than Alaohua County are very spotty, the state of Floricn oan be

subdivided Into two areas hwv nG d3itinct caddsi fly faunara (1)

Iorthweteorn Florida, and (2) Peninsular Florida. Northwest-rn Florida






-4-


certainly has a fairly distinct fauna for many species of caddie flies

hrvinf northern effinities Pa& are found here which are not collected

in the Gainesville region. "The Apalachicola River draineTe has with-

out doubt been the main hirh::.y of ingress to Florida for the great

majority of animals which require flowing water or h-rdwood forests.

Ro ers (1933) found that the ecological conditions existing in the

Apalachicola drainage woould admirably explain the distribution of

many northern craneflies in Florida, Carr (1940) reports that the

most extensive invasion of Florida by the northern element is encoun-

tered in that portion of the panh-ndle which is drained by the Apala-

chicola River. Hubbell (1936t354) states that in this peculiar en-

vironment (the deep ravines of the Apalachicola region) a gre-t many

northern plants occur, evidently the remnants of a northern flora left

as relicts from Pleistocene times in these deep, moist, cool revines.

"It has elso been noted that many northern plants re-.ch their southern-

most limits in these rrvines and many typically southern plants inter-

nin'-l here with the northern species. Iot only is this true of plants,

but such has proved to be the case in the craneflies, the Odonata, the

Opilionids, the Orthoptera, the crayfish, the PrThibi:,ns, and the rep-

tiles." (nerner, 1950). Peninmnil-r Plorida has a number of endemic

species, but the majority of the fauna is composed of species having

a far riider reolra.nhlc range.


Oaddil flies are fairly nbundrnt in Florida --- they can easily

be found -- but they do not appear to be present in numbers as larre






-5-


as those occurring in many northern streams. I hnve never collected

caddie flies in the north or the west, but I have reed many descriptions

of the immense numbers of these insects which swarm in northern streams.

The rock dam near Poe Springs is the only locality in Alachua County

which apnropches these descriptions of northern abundance.


In the north caddie flies are an important link in the aquatic

food chain, for they convert small units of food into larger units,

and since they ere present in tremendous numbers, they are a group to

be reckoned with, Caddie flies constitute an important factor in the

diet of many northern fish.


This does not seem to be true in Florida. I do not believe

that caddie flies are abundant enough (except in certain scattered

localities) to be a major link in the aquatic food chain. Reid (1950)

found the number of caddis flies eaten by the Black Crappie, Pomoxli

nIorm-clntus, in Orange Lake was insignificant. William McLane

(personal communication) found the same condition to be true while

he was workln- with large-mouth bass in the St. Johnts River. Very

few caddie flies were found in the stomachs of the bass.


Caddie flies are important in Florida, however, as indicators

of stream pollution. Many stream forms of caddie flies have an ex-

tremnly low ecological tolerance and cannot withstand any stream pol-

lution. According to William Beck, Jr., of the Florida State Borrd











of Health (personal communication), Cheumatopsyohid larvae will

tolerate pollution of a minor degree while most other caddis flies

will not. Personal observation bears this outt for example, in the

summer of 1950 the East Branch of Hogtown Creek had a caddis fly

fauna composed almost entirely of Cheumatopsychid larvae, while the

West Branch had a more varied fauna. Many old, empty Leptooella

and Oeoetis cases were found in the East Branch showing that other

species had lived there. Since it is known that the East Branch

occasionally becomes polluted, it would seen likely that the oreek

was slightly polluted in the summer of 1950, and that only the

Cheumatopsychid larvae could withstand the pollution. Ross (1944)

corroborates these observations* He states that members of the

genus Cheumatopsyche can frequently succeed to some extent in streams

too polluted for almost any other caddie flies. Therefore, it appears

that caddis flies can be used to show compnrative pollution (1) if

a stream is unpolluted, there will be a varied caddie fly fauna present,

(2) if a stream is slightly polluted, only Cheumatopsychid Iyrvae will

be found, and (4) if a stream is highly polluted, no oaddis fly larvae

will be present.

Although this is the first systematic study of this group of

insects in the state, the Trichoptera of Florida, and ccpecially

Alachua County, were previously fairly well investigated. Caddie

flies were collected by Drs. Lewis Berner, J. Speed RoEers, Frank

Young and others, end vwre sent to authorities for i~Sntification.











V.-rioui ruthorities on caddis flies made quic': collecting trips

thro-,th'out the stte. Almost all of this has been incldent.!'l col*

lectlnr in scattered locations rather tha. thorou.-'h, s.ytem7tic,

faunal studieS. Furtherr.re, information on the caddis flies of

Florida was citttrred -- indceed much of it was unpublished -- and

has been inpcceesible to all but a spFC"lirit. For these reasons

this problem has been Turnued long the aforementioned lines; a

faunal study and a consolidation of knowledge.


The collector's names are abbreviated throughout the thesis.

The abbrevi-.tions refer to the following ment D. A. S., 3. A. Bc-r-

rington; W. M. o ., V. M. Becek, Jr.; L. 3., Lewis Derner; A. J. H.,

A. J. Henry; A. M. L., A. '. Lpessle; A. F. C., Archie F. Carr;

E. L., Ellis Lanquist; H. ,, TN. Marshall M. W. B., 'p:rahanll Niren-

bert J. S. R,, J. Speed RoCers; R. P. T. R. P. Trondon; F. N. Y.,

Frr nk TYoun; J. S. I., J. S. :cCulluph.


:'oEt of the drawings in this report which illustrates the

keys were copied, and come from many different sources. Although

some of the illustrations were taken from the illustrations accom-

,r.nyAng oriin.Il descriptions, it has ceened Fdvis-ble to use the

best ir.idnrs available, and thus some newer, redescriptions were

nsed in -'ince of the older, original descriptions.


All of the original LrPwinrt were made with a camera lucida

fron uncle.red specimens preserved in 85;' ethyl alcohol.


-7.










lTh source of each lllutrrtlon is a s followbr 'ics. 1-36,
Ross, 1941P; Iri. 37, "irenbTrc:; -i. 38, Ross, 1944: Pir-. 39, -aos,

1948 : 1'1?. 40-64, ross, 194; rFi. 65, Vorhies, 1909; Figs. 66-73,
Rnos, I2!'8!; Pig. 74, Theur; .i:'s. 75-79, Ross, 1944; Fig. 80, Ross,
19!P7-; Frgs. 81-99, Ross, 19'4; 1igs. 100-100A, Theur; PiVs3. 101-
106, Bose, 1944; Fir. 107, Ross, 19-41; Fi-s. 108-117, Ross, 1944;
'igf. 118, DPnninn, 1947; rlig. 119, Poss, 1941}; Vig. 120, Dennln;,
19'8: T'irf.121, Ross, 1948; irc. 122, Ross, 1944; Fig, 123, Denning,
19L7; igs. 124-125, Ross, 1944; rig. 126, Denning, 1948; Figs, 127-
128, Ross, 19W!; rlg, 129, Ross, 19-1; Firs. 130-131, ~enninr, 1947;
Yirs. 132-256, Ross, 1944; Fic, 157, Theur; ri,,c. 158-1767, oBe,
1944; fiG. 168, ?:irrnberg; nit. 169, Ross, 1947; Ti r. 170-171,
];irenbcr,-i: Fis. 172-173, Ross, 1944; Figs. 174-175, Teur; Figs.

176-177, Ross, 1944; Frc. 178, Bkans, 193S; rics. 179-180, Ross,
1944; PIGS. 181-183, ITrenberg; Figs, 184-185, Ross, 19441 Fics. 186-
187, .'It r; 'ic. 188, Ross, 19'4; rig. 189-190, Toss, 1947; ~ iez. 191-
192, Ror, 1944; 7.i. 193, Ross, 1941; 71-~. 194, Ross, 19'4; Fig. 195,
Boss, 19380; j'ir. 196-200, Rose, 1944; fi-c. 201-202, Theur; Fig, 203,
Ross, 1944: 'ir. 204, Ross, 1941; Tic. 205, Firenbers; Pig. 206,
Theur; Fie. 207, Tirenberr: rir. 208, Theur; ri'-. 209, ,lTtten Cn
:!ocel:, 1940; Firs. 210-211, loss, 1944.










ACRNME1DnaEIMED TS


This study could not have been completed without the help

of many individuals. I take great pleasure in acknowledging the

time and thought spent by these men and was greatly impressed by the

willingness with which this help was extended.


I especially wish to thank Dr, Lewis Perner for his guidance,

constructive criticism, and for the many specimens collected by him

in other parts of Florida, Dr. M{inter Testfall spent many hours

helping with the photography, Dr. Cornelius Betten sent a list of

all the caddie flies he had collected in Florida end a list of all

the oaddie flies recorded in the literature which were collected

in Florida. Dr. Herbert Rose of the Illinois Natural History Survey

sent me many of his reprints, gave advice and identified some

specimens. Dr. D. G1 Denning of the University of Wisconsin sent

a number of his reprints. Mr. Ellis Lanquist, Dr. J. Speed Rogers,

Dr. Frank Young, Mr* R. P. Trogdon, Dr. B A. Barrington, Mr. Jon

Herring and others too numerous to mention, collected specimens used

in this investigation. Mr. Bert Theuer helped greatly in the

preparation of the keys by drawing the heads of some larvae.


William Book, Jr., George N. 7Teld and William MoLane aided

by oantributing bits of essential information used in this study,


Last, but by no means least, I would like to thank all

those who contributed materially by offering criticism, advice

and suggestions.













Three different methods were used in collecting cpddin flies:

(1) the larvae and papae were collected from water, (2) the adults

vere collected at light, and (3) adults were collected during the

day by sweeePin vegetation overh-nr~ning in-ter or by ne-rchin- crevices

underneath bribes. Since no one method will yield a representr.tlve

collection of the area, a conrbin-tion of all three was used,


All ITearctic caddie fly lprvee rnd, .puae p rrea qu.tie, and

almost any type of ?qu-tie hl.bitat is a -otenltinl collecting site.

Stonor and pieces of :ootd were brought to the surface for examine-

tlon. Leaf-drlft, rouvtic vepet-tion, end debris vere du-ped in a

pile on shore, After drying for a few minutes, the lprv. e begin to

rovr and can easily be .rottec.. Some pupal and larval cases show a

high derTer of protective colorFtion tnd blend in well with their

surroundinns, but with practice these cases can be readily disttin*




Fron a quantitative point of view I found the best place to

collect larvae and nnrpe in Alachua County was in rpTida end riffles;

for example, the rock dam at Poe Sprin'!. The larvae are present in

this snot in tremen9ovs numbers. I also found nui.ntities of larvae

by exratinlnrg the lenf-,rift in some of the numerous, small, swiftly-

flowin,, urnpolluted streams scr.ttered thror-hort' the county; for

e2rample, ptrts of "o~to:n Creek. Larvae nr' be found in all conceiv-

able tyrnes of aquatic habitats, for there are records of immatures






-11.


being found in pitcher plants and the hollowed-out boles of tree

stumps that had collected a little pool of water,


Adults of most species of caddis flies will come to light on

warm, still nights, and I used this technique extensively. Specimens are

to be found on store vindons; pink neon lights especially attract these

inseots. TMhile some specimens can be found in town on store windows,

lighting on the shores of streams, lakes, oto* is essential. I have

had good results using a Coleman, single mantle, gasoline inltrn.

Caddis flies clinging to the lighting sheet can be collected by

holding a small, wide-mouth jar half full of 85" alcohol directly

beneath the insect and tapping directly above the insect w.ith the

forefinger. The startled caddie fly will leap backnanrds into the

open jar* Dipping an index finger in alcohol, scooping up the insect

rapidly but gently on the wet surface and then dipping it in the bottle

is a rapid method which I used for collecting iany small caddis flies

I have used an automatic light trap with success. The trap

was set in various places overlooking water and was periodically

checked.

In daylight collecting of adults, sweeping vegetation with

an insect net is a neo&ssity for some species are negatively photo-

tropic and cannot be caught at light. Resting places differ with

the species, but most eaddia flies prefer shaded, humid places. Vege-

tation overhanging water and crevices of tree trunks are favorite

resting places. These insects are very fast and agile, and swift






-12-





movemont i~ required to oaptura then,


Frequently the adults may be captured on sticks end vegei-

tion floating on the water. When floating vegetation is submerged,

trapped adults will quickly swim to the surface,










PRPS9TRVATIOT MD CLTA'RIIC-G


All cddfis flies, both immature and adult, were preserved in

85,' ethyl alcohol. One genus, ,Letocella, wao collected dry since

identific-tion in this genus depends lrrely on the p-ttern of hairs

on .the wrinrB. I eost specimencof this genus were killed in a cyanide

jar and were carefully pinned.


The genitalia were used to identify almost all species of

caddie flies. It was usually necessary to clear the genitalia to

see the dipgnontic characters. I have used the following clearing

procedureI The apical third of half of the abdomen was removed and

placed for five minutes in a hot five percent potassium hydroxide

solution heated in a water bath. It was then transferred to alcohol

and the viscera were gently squeeze out. The cleared gFnital cap-

sule and the cnecimen to.rhich it belongs were put in a small vial

rpnroximntely 74 by 4 mm. This vial was filled, stoppered with a

cotton plug and inverted in a ring neck 4 dram vial.


It was desirable to clear the entire body of the micro caddie

flies belonrnin: to the family Hydroptilidae. The Mbdomen was not

detached. The procedure used is the same as above except in this

case it was necessary to teat a slit in the base of the Pbdomen through

rhich the dissolved viscera was expelled.,






.14.


Two major methods were used to associate larval caddie flies

with their adult forms. The larvae or pupse were either reared in

cmes until the adult emerged, or the pup-e were dissected.


Laboratory Rearing


Reprinn caddis flies in the laboratory is a difficult pro-

cedure, for many species have a low ecoloLc-l tolerance, and n-tural

conditions mnist be clmulnted. Therefore, in trying to increase the

yield of reared adults, I developed a number of different rearing

methods: (1) a series of eight screen wire cages, each enclosing

a balanced, aereQted aquarium were used Both larvae and pupse were

kept in the nquaris. All the larvae died occoslon.lly some adults

emerged from the pupal case. Vhen caged together, the larvae are

cannibllttic, and the larvae that survive are usually killed by

the chaneI in their environment. Pupae, however, require nothing

to eat, and if the putp is mature, the pdult will emer~e within a

few days. Pupae of certain species are often rather scrarce, which,

since it is difficult to find enough individuals to nmrai certain that

re.nlts will be obtained, constitutes a. major icrAvwnt ce. (2) SCob-

terred rocks and vegetation covered with r.upne end Irrvr.e were taken

from their natural habitat and kept emoitened, but not submerged, in

large screen-wire cages. The enddis fly inn.atures were able to get







-1&5


all the o-yp.en they needed by 5'eepinr them in contact with air r-ther

than water, The larvae died, but many pupae survived and .,ere re-red

to the Piut -tage. Micfel and Milliron (1939) described a Esilrr

method of rearing caddis flies. They placed -i.p"e in shallow pie

p-ns containing dnamn srhpmnhain m oss. The moss kept the ..nae roist,

and the pupae received more oxygen than they would have received in

water. TIy method was quantitative rather than qualitative, for if

two or more species of a genus were present, a specific association

could not be n'de. (3) The pupae were separated under a binocular

microscope and npcked, in groups of five, in dG~pn cotton. ach group

was then put in a shell vial covered with a wire screen top. This

senTrpted the species qualitltively, however, only a small percentaPe,

approximately two percent, could be reared since r-!ny pupe were Un-

doubtedly injured in hnndlin. (a balanced

cqFrium- with a strong jet of air eere-ting the water and Incident.lly

stirrin-t the water swiftly and constantly. A slow flow of tap water

fell into the aquarium, and a'slnhonin' arrrncEment took care of the

overflow. The larvae were fed with plnlkton. Time and lack of

eqruiment prevented me from making full use of this method, but I

believe this method was the best and can be used to cdvanate with

both larvae and pupre.



Field Rerinw


On three different occasions I set series of modified, copper-

wire pillow cages filled with larvae and pupae in the west branch of






-16-


Iogtown Creek near Gainesville. TheoreticIlly, this should be an

excellent method for associating stges in the life history of cAldis

flies, but in practice it was unproductive. Uore extensive, con-

trolled field reorinr should yield good results.



1n- al Dissection

Certain pupae form the connecting link between larvae and

adults and can be used to associate the adults with their specific

larval forms. In most caddis fly groups the larval scleritts, which

are readily diagnostic to species, are packed into the posterior end

of the pupal chamber after the pupa is formed. Just before emergence

of the adult, the genitalia of both sexes becomes completely formed,

hardened.and colored, which therefore specifically identifies the

slpeimen.


If a cocoon or case is collected which has a matire pup? in

it, the larval sclerites and fully formed renitalir can be associated,

and it is thus possible to linkfthe a:1ult and larval forms of the

species. VTowever, it is ,unll:y necessary to collect n.any pupae bs-

fore one is fomud with mature genitalia.


The family Leptoceridae constitutes a major exception to this

type of linkage. In this family the pupal chamber has a slit and not

a mesh in the closing cap at both ends. Through these slits the pupa

pushes out all the larval sclerites, so that it is irmossible to get












associations of adult structures and Inrrvl sclri1tes in the asme

case, For this reason it is neeessary to rear these larvae.


Care was taken in trPnsportlng the larvae and pupae. I hlve

had bent results by packing both larvae and pupme in drmp moss. The

caddis flies were transferred to their permanent aquarium or cage as

soon as possible.


















ane
pr n t rn- -- -









ii































metanoum --



later I hu p-






-19-


KEY TO FAMILIES



Bays to genera and species are incorporated within the

annotated list Refer to the Cheek List And Index at the end of

this volume for page numbers.


Larvae

1. Larvae (not pupae) living in and carrying around a
definiete C&8**********************.****,****, *********.** 6

Larvae without cases&*.......**** .....*.......*. ...*****.....lO


2. Case like a snail's shell, fig. 210.
Belicopoyahidaa (Ielioopgyohe borealis)

Case not lie a snailr shell,*.....*..*...*...........****..5.


3S Larvae smaller than 4 mas pro-, meso- and metanotum
each with a single, solerotized shield embracing
the entire notum.
Hydroptilidae

Larvae usually larger; either mesa- or metanotum or
both without solerites or with solerotized shield
subdivided into separtto plates, fig* 11....................4


4. Clnws of hind legs very small, those of middle end
front legs large, fig, 2.
Molannidae (Molanna tryphena)

Claws of hind legs as long as those of middle legs,
fig. 8o*..**.*.***.****.>f..*.....**.*.........*..*.....**

5. Antennae long, at least eight times as long as wide,
and arising at base of mandibles, fig. 4...*.
Leptoceridae

Antennae much shooter, not more than three or four
times as long as wide, often very inconspicuous,
and arising at various points, figs. S and 6*....+..,.......6



























aal















A B















I S Ul0@TOK




















40



U *S






-21-


6. T.esonotum subnembranous, except for a- pair of
parenthesis-like, sclerotized bare, fig. 7.
Leptoceride.e

?Mesonotum 'dthout such barae.. .0 or.....* *&**.. *.. ..o... 0


7. Meson end metanotum entirely. membranous.
Phryganeidae (Bankuiola conoatenats)

Mesonotum with some conspiouous sclerotized
platess *.**. *.................... .******. ............8


8. Anal hooks formed of 2 or 3 long teeth ,ituatd one
over another, fig. 8 A.
Serioostomatidae (Sericatosma orsssioornis)

Anal hooks formed of a single large, tooth with
1 or more mail teeth on its dorsal edge, fig. 8 B........*.9


9. Pronotun with a deep furrow running almost the ..
full width of the soleritae head as in fig. 10.
Braohyeentridae

Larvae large, living in a case made of sticks,
fig. 135.
LimnopAhlidae (Pyonopsyohe ep. A)


10. Pro-* meso- end metanotum each with a single,
sclerotised shield embracing the entire notuim,
fig. 12i abdomen with many oonspieuous branched
gillls fig. 12.
HIydropsyohide.

Either mesoo or metanotum or both without solerites
or with salerotised shield subdivided into separated
plates, fig II******,** ********************.***.,******.II


11. Labrum with anterior end lateral portions expanded
into a ilde, mnembranous area, fig. 13.
Philopotamidne (rhiiarra)

Labrum shorter, entirely selerotised, fig. 14.
Psyohortriidae





















;ell
k 4"

FIG. 13 --- U TITZT-1



'Y'

4


FIG. 14-11-,ABR7


FIG. 12 ----- 1A7VA OF


SC2 Ft P2 Ft R4
antenna --------------------------- R5
median ocellus -------------
anterior wart ------------- ---
compound eye ----------
lateral ocellus ----------- M2
postero-lateral wart M3
pronoturn ------------
CU
a., t M4
tegul c u uM----
mesonotum -C"
scut C.-d
(posielol tuumm----- '4 b
metanotum ------- --------
...... Sc
............



2 R4
iA
r F1,
abdomen ............. 2
cvb cu. M, M M,



--1o
claspers genitalia
-.:aedeagu'---





1 2 3 4,
maxillary palpus,, 7
io claspers

labial paipus.----- me'so-
pleurae
metapleurae
c 0)(aj
trochanter, 'apical spurs
femur,
tib a' tarsus
claws_._. preapical spurs


FIG. 15 ADULT, ILLUSTRATIEG TEER11INOLOGY OF PARTS.











Adults

I. Small, hairy individuals not over 6 mm long; meao-
scutellum with posterior portion forming a tri-
angulpr, flat area with steep sides, fig* 161
mseoscutum without warts.
Hydroptilidae

Size ~mng of 5 to 40 mm; either mesoeoutellum evenly
oonvex, without a triangular posterior portion set
off by sharp sides, firs. 17 and 18; mesoaoutum
with warrts* ....**..***************. ... ******* *******


2. Ocelli present, fi. 15*........*........**,*,,*..*..*..........

Ocolli absent ese***** ***


*aegamented, fig* 19.
Limnonrilidne (Pyonopeyohe spi A)


r'axillnry palpi 4. or 5-segmented............................


4. Msxillary palpi


o. Maxillary palpi
times as long


4-segmented, fige 20.
Phryganeidae (Bankaiolsa conatenata)


with fifth segment two or three
as fourth, fig. 22.
Philopotamidae (Chimarra)


axillary palpi with fifth eeg~int not nore than
one and one-third times as long as fourth, figM 2$3......*....


6. Haxillary palpi with 5 or more segments, fig. 24*..e...,,*....*7

1laxillary palpi with less then 5 segents.............*......10


7. Terminal segment of maxillary palpi much longer than
prooeding and with close, suture-like, cross strike,
which are not possessed by the other r~gments, figs.
25 end 2a*n**********n************************


*230


8. Maxillary palpti


Maxillary palpi 5-segmented,fig 21s,*.... .....********.******















































In














rI.






-25.


Terminal sognont cf maxillory pwlpi without such
strike and similar in general structure to fourth
segment, usually of same length***........*......**,.....10


8. Anterior tibiae with a proapionrl pur*.
Psayoho' iidao

Anterior tibiae without n preapical Bpur*........* ....** ....*9


9. !!s3ooutum without warts, fig. 17. ...
Hydropsychidne

Tepsoutu:n with a pair of smail warts, f 27*.
Psyohotrildae

10. Ifiddle'tibiae without preapi cl spurs Pnd with a
row of black spines, fig*, 28.,*..., ...........,........,1

middlee tibiae with preapionl spurs, with or with*
out a row of sGinee..**..,..... ....,....,,..,,*****.,,,14


11. l esonotum with soutal warts represented by a long,
irregular line of estate spots, fig* 291 antennae
always very long and slender.
SLeptoeoridne ... .. .

Mesonotum with soutal warts either small, fIg, 30,
Or nbsent, fig, 188 antennae fairly short.....**********,,* 12


12. Male genitalia as in fig. 31S ventral aspect of
abdonon of femle as in fig. 32i
Ilelicopsychidtoe (Hfelioopsyche 'boroalia)

Genitalia of male and female not as in figf. 31
and 322.********t******w******. ***********e**e******e..... .13


13, I esonotum with a deep, antero-mesal fissure with
soutal warts as in fig, 33.
Seri noFstcomatide (Sericostoma craesioornis)

Mosonotum with only a shallow antero-mosal crenas,
with scutal-warts as in ffiCS. ..
Brachyeentrid roc

















































(lift"















.5s 10) as m OT.S,-






-27.




14. Middle femora eaoh with a row of -.10 blaek apltes
on nntoro.-vetrnl face, fir. 55.
Pol1anidne (Molanna tryhrena)

?fiddle femora each with none to 2 black spines on
antero-ventr.'l face,
Er.-cycontrldie


































AA








VIE; B, EEAG
F S F 4
MAF











U S 0M






029*


Key To The Speoies Of Chimarra



Larvae

1. Apex of frons as in fig. 36,
Chiirarrft aterrima

2. Apax of frons as in fi,. 37.
ChLmarra florida

3. Apex of frans ae in fig. 38.



Adults

1. Apex of abdomen with a pair of cleaners (maleas)*.*,,.****2

Apox of abdomen without claspers (females)... ***....*........*4


28 Color brown, genitalia as in fig* 39.
Chimarra perigua

Color blackish....,.......,.,, ... ...*******.****************.


S. Genitalia as in fig. 40.
ChiLmrra aterrima

dGnitalin as in fi g 41.
Chimarra florida


4. Color bro=n,
Chimarra periguaa

Color blaocish ..** **iii*es.*t**.*r.*******>** *5

S6. inth tergita produced into well-defined, lateral,
solorotised ears, fig, 42.
Chimarra florida

Ninth tergite without oars*
Chimarra aterrima











SChimarre aterrima Hagen


Range Eastem and southern states*

Desriptiont Ross, 1944.

Remarks: The larvae of this species live in small, cool, fast-

flowing, spring-fed creeks. Although I found a stream in Golden

Head State Park where 0. .torrima was the dominant caddis fly, it is

not usual for the species to be so abundant, The fact that I have

never collected adults at light suggests that they may be negatively

phototropic.

Previous Florida Reoordst Rose (1944) records this species from

Florida but does not give the locality. Totton personall commimi-.

oation) oolleoted C. aterriia in the vicinity of Lake Placid.

Specsa ens Examined Five larvae, 2i- miles W. of Gainesville, L. B.,

1/28/391 3 adults, 2-i miles W. of Gainesville, L. B., 1/28/39 1

larva, Alachua Co., 11/9/371 40 larvae, T1o,,tn Creek, In leaf detritus,

F, N. Y., 12/31/471 1 Inrva, Devil's Uillhopper, 10/25/37; 1 larva,

Devil's Millhopper, 10/2/51; 400 larvae man upae, Golden Head

State Park, M1 W. N, 8/12/601 40 larvae, Hogtown Creek, M. WV. N,

8/8/501 38 larvae, Hogtona Crook, M. N., 6/4/51.






-31-


Chimrrtr per~ga Ross



Range Florida, Georgia and Illinois*


Description& Rose,,1948, ,

Renarkt: Ross (1948) states that C. perigua was recorded erron-

eously as sooia and the latter species was thus listed as being

present in Florida. Chimarra sooia does not occur in the state.

According to Ross (1948), "Re-examination of a Irrge series of

Booia---.---( Indxiates) a wide eastern and northeastern range for

scoia and a small range of perig a which is peripheral on the southern

and western portions of the range of soola."

The larvae of C, periua live in clear, cold, rushing water

such as that pouring over the rock dam at Poe Springs. The adults

appear to be negatively phototropic, for though I "lighted" for

caddis flies many times in this locality, I did not collect any

adults.

The larva was Pseociatod with the adult by a process of

elimination. I collected larvae of three distinct species of

Chimarra and adults of three species C. aterrima Co perigua and

C. florida. The larva of C. nterrima was previously described and

was easily identified. I associated the larva of C* florida by

pupal dissection, which left one unidentified species, The adults

of C. perigua are very similar to C. sooa, and the third larva

was also extremely similar to that of Co socia. It seems obvious,






-S2-


therefore, that those larvae must be those of C, porigua,

The larva of C. periiua cannot be distinguished from that

of Ce sooia using the description given by Ross in 1944, I do not

have comparative material of C sooia and, therefore, cannot look

for differences in morpholoSgicn characters between the two species,

but a critical examination of both larvae is definitely in order,

Previous Florida Reeordat "Holotype, male---wSanta Fe River,

Alachua Co., Florida, April 6, 1940, L. Berner. Allotype, female---

Santa Fe River at Poe Springs, Alaohua Co., Florida, Maroh 12, 1938,

L. Berner. Paratypea-----Floridat same data as for allotype, 1

male." (Ross, 1948)

Spea 2ines fxaminea Two adults, Poe Springs, L. B., 3/12/481 3

larvae, 6 miles W. of Gainesville, B, A. B., 11/6/S39 1 larva, 5

miles W. of Gainesville, B. A. B., 12/?/39i 1 larva, Alaohua Co.,

L. B., 2/7/39 4 larvae, Poe Springs, 3/2/s38 10 pupae, Santa

Fe River, J. S. RI, 8A2/30; 8 larvae, Poe Springs, !I. We N.,

3/24/501 2 larvae, Poe Springs, U. WV, ., 8/8/650 14 larvae, .I

mile south of Gulf HaI.ock, Me W. N. 7/30/50S






-53 A.


C* thlrra florid Roes .


Ranget Florida and Georgia.


Desorintiont Rossa 1944.

Remarks The undeecribed 1erva of C. florid was nasooiatedl Vwith

its adult forn by nmean of r~~rl dissocotl. .. .


The larval stage of this species was only found in the most

rapid Torticns of very s ift caloareous streamse

Previous lrorid P.ecordet Oae male, Freeport, Walton County,' collected

by Dr, Lowis Berner, Aprr!l 3, 192?. .(from Rose, 1944)


Specimens aEamined: Three males, fast stream, .1 mille S of Gulf

Hammock, tLvy Co., sweeping, M, W, Ne,, 7/30/501 1 pupa, .1 mile S.

of Gulf Ham-ock,. Levy Go., i.1. Vi. No, 7/30/0sO 9 larvae, Poe Springs,

10/14/47; 1 female, Poe Springs, at light, M.. W. 2, 9/9/601 1 pupa,

Pow Springs, M1 W. N., 8/24/51; 1 pupa, no dnta; 1 male, sweeping, Poe

Springs, M. W. N., 3/24/50; 2 males, 6 females, Poe Springs, at light,

M, W. V., 7/2/s0.






















10






-- 45--,APAX
A:-.AL ITCOX AYAT- PITA:




Z


10 10







FIG. 47 ---- APEX OP FIG. 43 ---- AYWX OF
ABD( MUM











FIG. 49 --- X DIr L E, S FIG, 50-OAMUTS







...............






F IG. 51 --- T.4kNDIBLi7,S FIG. 52--wTUDIBLES






-34-


oey to the Psycharnyidae


Larvae

1, Anal hooks with a row of 4 or 6' lng teeth along
S inner ventral mn.rri.n, fit 44s tenth segwent'short*
with seareely any ventral margin fig. 45.
Psyohoaprla flavida

Anal hooked with at most very short, inner toeth, fig.
486 tenth segment longer and tubular, figs. 47 end


2S Mandibles short and triangular, eaoh with a large,
thiok brush on the mosal side, fig. 49.
Phylooentropus placidus

tMndibles longer, fig. 50, with only a thin brush
on loft mandible, none on ,rightf......u***...,*i* ,..* 3


3. Right mandible with two large dorsal teeth whioh
completely overhang and hide the ventral rows on
the left mandible the dorsal row of teeth over.
hangs and hides the ventral row, fig* 51.
Cyrrellue marginalia
Right mandible with a single dorsal tooth whioh only
partially hides the ventral row of teeth on the
/ left mandible the dorsal row of teeth does not hide
the ventrol row$ figs. 50 and 653....*.....,*.*.*r,,.*,***4


4. Basal segment of anal appendages (tenth segment)
without hair, fig, 47; left mandible wtth basal
tooth small and with a linear brush cn msanl
face near bnse, fig. 47.
Neureclinsis crepusoularis

Basal segment of anal appendnCos (tenth segment)
with long hair, fig. 481 left mandible with basal



































rr




?l". 7 ( TAlD TG




FI.5--wi&ILR

PAMPf PA3"






-36-


tooth large, subequal to one .above nod with
brush small, fig. 52.
Polyoentropus

Adulta

1. Front tibiae with a preapioal spur.........**...*....,......,.2

Front tibiae without a preanpial spur......*...** .,.......*. .6


2. Both pairs of wings with R2 present end branohing
from:R3 at radial crossvein, fig 56oS
Phyloeentropu placdus

Both pairs of wins with R2 either absent or
branching from RH near margin of wing, fig. 54.............


53 Hind wings with M.S-brancihd, fig. 55.
eureelipsia orepusoularie

Hind wings with M 2-branohed, fig. 6.......,,.,...,........4


4. Front or hind wings, or both, with 'R present,
fig. 55.
Polycentropus

Both wings with R2 absent, fig, 56...........................5


5. Maxillary palpi with second segment long, third
only slightly longer than second, fifth short,
fig. 57.
Cyrnellus marginalis

:axillary palpl with sooond segment short, third
three times as long as second, fifth long, fig.
68.
Nyotiophylax vestitus

6. laxillary palpi with second segment only one-*
half to one-third as long as third segment
and with the apex enlarged into a small cushion,
fig. 69,
Carnotina






437.


Uexillefr palpi with second cement as long as
third end uniformly cylindrical fig. 60*,...****......*..,7


7o Hind wings with apex evenly rounded1 figa 61,
LySpU diverea

Hind wings with apex tapering and somewhat
pointed. fig* 62*
S, Payohosyia flavida



T':1T1CTLIPSIS ToTachlrn


l7eureolinsis arepu eulrria ('alkeer)


This species has not been taken in Alachua County, but

Better (personal communication) collected it in Florida (location

not given).










PHYLOCENTROPUS Banks


Phylooentropus plaoiddus (Banks)


(Holooentropua laciduB Banks, 1905b
f.iocOentropua maximus Vorhies, 1909)

Rnnoet Eastern, central and saithern states










Description: Ross, 1944.

Remarks: This species appears to be rather rare in Alachua County,

for I was able to obtain only a few speoimans.* One larva was found

in a lake, the other in a slow stream with a soft, flooculent bottom.

Vorhies (1909, p. 712) gives an excellent description of the unusual

larval ease.


"The larva burron~ into the sand of the bottom of the

stream and forms a long, often complexly branched case of sand,

scarcely firm enough to keep its shape when it is eXtracted from

its position. This oann, about 5 rmo. in diameter, excepting some

smaller branches, is ofton as muoh as 65 mia in length, the rretter

portion buried, only 10-20 am. projecting from the stream bed."

Previous Florida Recordst Dead Lake, VWewahitohka, oollected by

J. G. Needhan (Betten, 1934, p. 213); speoamens collected by Betten

(personal comnunioation), loortion not given.

Specimens Examinedt One female, Waashington Co., L. B., 6/SO/40;

1 larva, Lake Santa Fe, 12/8/47; 1 larva, .low stream, 5 miles N.

W. of Gainesville on U. S. 441, i. W. N., 6/21/50; 1 male, Worth-

ington Springs, at light, M. W. N., 7/3/50.








-40-






-41-


rOLY Ti. "OPUS Curtis



-Ke yTo The Spocies Of Polyoentropus


Larvao

1. Spots on larval head as in fig. 68.
Polyoentropue sp A

Larval stage unknown.
Polyoentropve orascioornis


Adult

1, Genitalia of iamlen as in fig* 70o genitalia of females
as in fig. 71.
Polycontronpu on assicorni




Polyoentroeas orassioornia W'alker



The larva of this spsoieo, although it has a wideepread

range throughout eastern United Stotes tnd Canada, ia unknown. Banks

(1907) records P. orassicornie from Jaoksonvillel however, I have no

other record of its occurrence in Florida. Possibly Folyoentropu

sO. A Is the undescribed larva of P. crnsioornis.



Polycentropua sp. A



Although I collected the larvae of this species in a num.

ber of locations, a constant search for the adults failed to produce





-423


a singlo *speoimen The adults nay be negatively phototropso, for

night lighting and daylight sweeping were both ineffective in col-

looting them. I attempted to rear some larvae which I collected

in Big Hatchet Creek, but the larvae died before pupation.

These larvae appear to be very similar to the larvae of P.

remotus (fig. 68). Since P. remotus has been collected in British

Columbia, Illinole, Minnesota, New Hampshire and New York, a wide-

spread geographic distribution coupled with a looal occurrence ia

indicated. It is possible that the larvae of Polycentropus ap. A

are the undesoribcd larvae of P. ,rassioornia, which has previously

been collected in Florida, but it is also possible that they are

the larvae of P. remotus. Further study will undoubtedly permit

association of larva and adult*


Sieoimene Examineds One larva, Poe Sprines, J. S. M., 5/14/384 1

larva, Santa Fe River, 3/24/357 1 larva, Big Hatchlt Creek, M. Wo

N., 4/24/60; 2 larvae, stream, .1 mile S. of Gulf H1m-~nock, Levy

Co., Ie 1. 1 1., 7/50/6SO 3 larvae, Big Tatchot Croos, WI N,,

8/l0/60s 1 pupa, ravine stream, Golden Head State Park, Clay Co.,

M. V. li., 8A2/60.






4-42-


IYOTITOPHYLrA Brauer


Only two Nearctic species are Inown for this genus, one

of tlie from Florida. The Itrvrne have not yet been associated with

the adults


Kyotiophylatx Vstitus (Hagen)


(Polyoentronus vestitus Hagen# 1881
Polyoentropus affini Bannks 1897
1r lYla ax moeestue Danks, 1911
vOT estItu (Haren), Sibley, 1926)


Ranget Eastern, central end southern states.


Dereriptiont Ross, 1944.






-43-


Romarkce The larva of *. vestitus is unmownar however, it found,

the larva might very well turn out to be one of those which are now

classified under Psychomyild Genus A. Further study should clorifyj

this interesting enigma.


Nyctiophylax vestitus is rare in Alsahua County, having

been collected only at Poe Springs.


Previous Florida Reoords: This species has not previously been re-

ported frhn Florida. Betten (personal -comnunictticn), however, aol-

looted it in the state (locality not oited).


Specimens Examinedt One male, Blue Springs Creek, Jackson Oo., L.

B., 6/5/40)1 1 female, Poe Springs, at light, M9. V, N., 8,42/650 1

male, Poe Springs, at light, M. W. XN, 9/9/60,




CEtNOTINA Ross


No larvae of this genus have been discovered, and no morph-

ologiarl differences lhve been found to distinguish between the fe-

males of the various Epooles, the genitnl segments of vhloh form a

conical structure with only simple parts as shown in fig. 77T I

have examined specimens of the following ferTles, ~oich cannot be

identified specifically since they were not collected together with

males 1 female, Gainesville, at light, M. W. No, 5/3/60; 1 fensle,

Big Hatchet Creek, at light, M. W. N., 6/12/60 2 females, Vorth-



























( A L -j 7 T



IFRCU5
S Ak











ington Springs, at lirht, M* Vr NX, 7AS/560 2 females, St. John's

Rivers Putnnan Co~ at light, 7/66/60 60 females, Poe Springa, at

light, M. W. N,, 9A//60S 25 females, Poe Springs, at light, M. '.

, 8s/8/so5.



o:y To Tlhe Speoias Of Cornotina

Larva--**- Unknown


Adults



Females, fig. 77----Not keyed.


2. Corci vwith 3 or 4 long, black teeth on the wmeal
side near the base, fig. 78.
Cernotina oncaea

Ceroa without long, black meosa teeth..*..*........*,*....S3


, (Genitalia as in fig. 79.
Cernotinta spioaa

beenitalia as in fig. 80.
Cornotina truno.ma






-406


Cernotins oenloes os



r.n: The rnnae of this species ir *Aide, but C caloee aper-rs

to be locally distributed. It has been taken in Illinois* Missouri,

Oklahoma, Texas and Flor ida


Desoripticon Ross, 1944.


P.emgrhsa I collected, adults at Poe Sprntgs and Worthington Springs,

but continued inveetigatico within wneing; distance of the shore

y iel cd no larvaei. The larvae may rossihly be found in the deeper

parts of the Santa Fe River, beyond wading depth*


Emergence, met likely, takes place over the entire stummer,


Previous Florida Recordat Ross (1944) lists this species as occur-

ring in Florida but does not speiofy the locality* aetten (personal

coniuniocticn) reports tainag C. ealaea in the vicinity of Lake

Placid.


Speoisena Examined; Twelve males and feomles, Poe Springs, at

light, U t W. *N. 7/12/50s 100 malos and females, Worthingtcn Springs,

at light, M# 7, N,, 7L3/50; 20 males end forales, Poe SprinCs, at

ligit, gU W* N*, 9/t/50; 30 males and females, Poe Springs, at light,

M. W#. N, 7/3/51.






.47.


rn2 T' Carnl;' Cr-f. n- riCCrS.




qoaniwintliont anosS 1047,


Tr-Phs: I nollootoe this cr-oleo on oanl" tiwo o- 'oloai, t.toth t!nos

on ithe o cr o cCf it-o P1i-r '^ty To Yr tynociIro t'-o typo mntoriol

rIpls r ario eO

;:nro a n'Go. r of i:'Iv'durlu wro i*ourAd9 it n-'.crrv

ll!al t':-t the larvae emrn be found in the ryivr; hBoavorc, at corah

lia. t.r. rtdien-; el ctarco of t'c C:(oro .lor to roeval t':n. I 1.0.

limev the rro.lcn leto o te^.r to tbr.t oCf fer-m-**'*:c lrt.rroe

are to be Ccunti In the flconor nntera, boeycri vOtir depth.


T'e;'re s"s ctr *n'rr'o.--.crl datrovtrrca .1:-l car. tIo usvt to

it'':ero:'t':A-!.b'? tt .c!-o a rlcrlr o01 tao egernu. It'tv-;;- I eol-

loctod toa fe-nloc of Co trImco-n Wlich habd not bon collected

prov.cuaely, no rorcela-tlcl r't-r*tro -:ror founr Id thit nc'ld

be us1Te9 to Ci IneV!tui-clv t:-.on fro other-. fnr.cror in t':. gPanu8e


rlVe.o--0'o Prt-k r 0 't IIo9, :c;:tc:u: :o.vt?1, VoGlrt'ii- Co.9, fod

lo::ota ;lo.ol, Tir. :'p1 r o r -.- ':, 0/27/4s. (ross, 147a)


Aaolmrnas ; n. l Two rArlos, 383 tfenarlo, rivr Styx, at ig lt,
. ?. I., 7/A2/a/ 5 Inlec, g2 Cor lnr, river !tPx, M lr-t, V. C.

:;., n/bA/.











Cernctine rpicate Rose



aCrnet s Maine, Tcichi-.n, Ci1hne., end Florid-., OCernotina pcaeta

has not previously been reported from Floridn, It is obvious

though, from its spotty distribution, that S spioa.ta is far more

widely dietriluted than has 'baem thought and most probably occurs

In certain areas of the intervening territory,


Deaoriptiont Ross# 1944*

Re~mr-ks Ceornotia spio.at_ is umoonon in Alaohu.. CoWmty, for I

collected only four spoclimns at t o widely separated points. Most

likely these individuals had floan some distanaoe Two of the adults

were taken near Gainesville, the others on the banks of the Santa

Fe River.


P.evlous loridn Recordst 4one.

Specimens Examinedt Onemalet Blue Sprinj~ Crcee, Jaokl:sa Co., L.

B., 6/5/40r 1 male, one female Pond B, at light, M. W. N., 7AO/O60

2 mrles, Poe Springsa at light, UM, N,, 9/9/50






-049.


Psyho'yll.a Genus A


Pan: t Illinois, Florida, Michigan da r.d "isscnGin.

DescriptiLnt nossg 1944.


Remarket The adult of this curious larva is unklowno but it is

probably one of those now, known only. from the adult stage, auoch.,

as Tyctidopyl.x or Cornotina. In Alachun County the larva (fig,

81) ias only been colleste'1 in the Santa Fo River and 31ig atchet

reec.


Previous Florida Reoordea Rose (1944) records this larva from

Florida but does not givo tke locality-


Speolmens Examinedt Two larvae, Santa Pe Rivers 3/24/37; 2 larvae,

Alaohua Co., 1/b1/51; 3 larvae, Big Hatchet Creek, l. v,, N., 4/29/50s

1 larva, Big Hatohet Crook, !L. 1,. N*, 8/40/51i 2 larvae, Big I atahet






-SO.


Creek, MN,% No 4/29/50.



LPE McLaohlan


_es diversa (Banks)


Ross (1944) records this spooles as occurring in Florida

(looality not given), but I have not collected it in Alaohua County.

Perhaps it ooours in northwestern Florida.












PSYCEiOAY IA Piotalt



Of the throee described North Amer5oari speoiesg only P,

flavida has boon taken in Florida.






-52..


Psyohomia fla vida Hagen


(Pryachopa flavida Eagen, 1861
Suoiiia pulchella Banls, 1899
PyMhangi&a moeata Bank& 1907a)


Rane BEastorn central and southern states, Since ,flavida

has not previously been take in in Porida and Kentuokyp Tennessee

and Vorth Carolina were the nearest reported collecting siteae

finding this species in Florida indicates that it occurs in the in-

tervening territory.

Desoriptiont Ross, 1944.

Remarks Perhaps northwestern Florida is the only area in the state

this species can be found, for, although I collected no specimens in

Alaohua County, I examined one specimen which was collected in the

vicinity of Iarianna.

According to Rose (1944), the larvae are restricted to

swift, cold streams.


Previous Florida Recordst one,


Specimen Examinedt One female, Blue Springs Creek, Jackson Co., L,

B., E8//40*





















iloi





-Uynnej.jus -10.r7inalis
FIG. 74--- C
MEAD 07, TAIR-A.













A
FIG. 75 ---- Cyrnell-Qs mar,7inalis, 'AL1 &1, 7 ITALIA.
V61-TRUL ASPC'T 01?
A, EAT UL ASP,-rr
'1211'










FIG. 76--anneilus
;G7r inalis, FE7ALE
UM-TTALIA -






*54-


COYZSNLUS Banks



Cyrnellue marginalia (Banim)

(Qlyotiophylax marGinalis Banks2 1903b
Cyrnellus zernyii osely, 154)

iange~ Central end southeastern states also known from near the

mouth of the Amasn River in South America. In the Southeast, C.

marginalis has been reported from Tennessee, Kentuolj and Alabana

but not from Florida. Although I have not verified the correct-

ness of the identification of the specimens from South Amerioa,

finding this caddie fly in Florida is an indication that it must

ooour in at least limited portions of the intervening territory,

namely Mexico and Central America.


Descriptian: Ross, 1944*

Remarkst Only one North American species, marginalis, is known for

this genus. The immature stages have never been discovered, but

evidence presented here, I believe, will clarify the association of

the various stages. I found two species of larvae which were keyed

(Rose, 1944) to Psychohniid Genus At one of which is to be found

under Psayeholild Genus A in this thesis, and one of which was un-

described. I collected an innature pupa, however, in Lake Vlauberg

which contained the larval solerites of the undescribed species of
Psychomyid Genus A, The fact that these sclerites could be identi-

fied as Psychomyiid Genus A would eliminate Phylocentropus, Neurei*





.*i5


elip is and Polycentropu. The pupa had a preapical spur on the
front tibia whilh eliminated Cernotinaj, Igpe end Psayohonmia leav-

ing only Cyrnellus and yTtioetphylax The maxillary pelpi, the key

character for distinguishing between the two genera, are identical

with those of Cynellus, so I feel the assooi.tian is a valid one.

It appears, therefore, that tho Lake 1nauberg specimen is the pupa

of C, tmaginalis while the second species still romains iitho:t a

name In PsychoTlld Genue A*

This species is not very abundant in Alaohua County. Ify

records indicate that emergence occurs In the Spring*

Ross (1944) states that the adults are mainly found on

the shoroa of lIrge rivers but also on tse shores of small streams

and lakes. Although I collected adults from rivers and streams,
I found larvae only in lakes.

Previous Florida Recordst None.

Spooin ons _xaminedi Three larvae, Kingsley Lake, Clay Co,, bottom

sample 10 feet deep, silt over sand, Je So R. and As F.'C,, 5/18/33

1 larva, i ak.-anta PF, bottom example 4-5 feet deep, 12/2/477 2

larvae, Johnson Lake, Golden Iead State Park Clay Co*, Me W. N.,

8sA2/60s 1 larva, LerJ Co., E. L., G/A8/G50s 7 females Prairie

Crook, at liCht. # M Wt 1., 8/29/61; 1 male, 1 female, Viorthington
Springs, at light, U We, 1,# 7/13/801 2 females St. John's River,

Putnam Co., at light, 7/A6/50 19 males and females, River Styx, at





















































A,








STIUAO ST S:T,






-S7-


light, HI I .N, 7/17/50$ 1 pupa,, Lakei VTauberg,iM..W. TT.. 6/18/504




IE'Dr.o'PSYCnIDAE



Ross (1944, p* 76) characterizes the family IIydropyohidae

very rvll h en he says, "The larvae of all genera are remarkably

uniform in habits and appearance. They are wormlike, active and

yugnaoious, and possess rows of bushy abdominal gills. They pre-

fel the more rapid locations in stress, ,usually being conoentratod

around riffles, spilltays and rspi.De,. although they my also be.

found wherevertheee. is an appreciable current They make a re-

treat under and ab)t trac.h, logs, stonel and any other haVen.

In front-of this. ryc-reat they build a aet whioh is reputed to

strain food frm ro thei flwingwter. or pupation they spin an

ovoid coooon near the retreat, generally. ueing sand, atones and

bits of trash..


ICKy o The &ydropayohi!da



l1 Head with a d6nrp, U-shnapod ridge, fi(. 871
Maoronenmm carolina

Head without a sharp, U-shaped ridge, fig. 103OS......*****,*,2

2. Stridulator of fore Aoo~ not forked, fig* 808 '
W Dipecatrona mnodesta

Stridulator of fore coxa forked, fig. 89....,......*..***.*.S













































FI. 96SEAEFG, 7--liLL


ASEC. iSE"T












S3 Prosternal plate tith a pair of posterior soleritos,
fig. 90.
rydropsyche

Prosternal plate without a pair ef posterior solerites,
fig. 91.
Cheumatopsgohe

Adult

1. LarCe, Etrkidng caddle fly with piot.,red wings end
very long antennae, fig. Cr.
SaorW~Pmun- oarolina
Caddie fly not as in fig* 02............ ..* a,*4,*,,...* 2


2 Hind wings with apex round and with So and R bowed
deeply at apex, fig. 93.
Diplectrona modest

Hind wings either with So and Ri not markedly bowed,
or both wrngs with apical mrgln ino eed...,,****c**,****,.


e 41 jK0B8************************************************* ***

2021Pei *** ************************************************ 6


4. Base of aedeagus cylindrical, fig* 94.
Uydropqyohe in commodn
Base of aodeagus bulbous, fig. 95.
Cheumatonsycoie

6, Sternal plate of eighth cegpent separated to base
of soiment, fig. 96.
Chlouiir~topsyohCo

Stornal plates of eighth segment seopcrated only
tno-thirds distance to base of segment, fig. 97.
IHdropeycho incomrioda






-GO-


DIPTLT:C;A .c st-wood


Diploctrona nodostn Banks



Rnn.a: ,astern and southern states.


Descriptions Rosa, 1944.


Renmrkat Dipleotrona modest does not occur in Alaohua County and

most probably is limited to t'he northwestern part of Florida.


FrevlcTs FloridA. recordoat 2oc (194) .11st this apecies as occur-

ring in Floridr. but dose not cite the loc1Aity.


SpeciRmes ExE= ned: Six larvno, Torreya Stato Park, Liberty Co.,

5/7/331 1 larva, Crestview, Okaloosa Co.. L. B., 12/L2/87.










































in c ori:-,, o A7 OF 17
CR I 'i,,E 1,(71 A


FI G. 99 --- I-Tvdro Svche
inco'lmoda.











A B
FIG. 101---L.lyd
= ILche FIG.102 --- ad v he
Ln-_22-rao(Lp_, APEX OF inc6.-Tcla, M-1,11ALF,
AEDEAGUS. A, LATEPAj, GENT TALIA.
ASPECT; B, VENTRAL ASPECT.










HYDROPSYCIIE Piotet



The genuea fdropsyohe contained over fifty peCieso of onddis

flies, but only one, Hydropsyche inco-tnoda, is IknoMI to oaeur in

Alachua County. The larvae prefer clean, fast-flowlng, cool water


Key To The Spe.os Of Hlydropgsy(ie ,


Larvae,

1. Head as in fig. 100.
E dropsyo he incormoda

Head as in fig, 100A.
Itydropyoche ep. C


Adults

1. Genitalia of males as in fig. 101; genitalia of
females as in fig, .102.
fydropsyocho ineooimoda





lydropsyche incomrB~oda HEgen

Rancea Recorded from Florida, GeorEgi, ITorth Carolina and Illinois.


Desariptlont Roess 1944.


Remarks The immense numbers of enddis fly Inrivae swarming over

the rocks and loaves of eel grace (Vallieneria sp.) at the rook dam

never Poe Springs is certainly strilking. For shoor numbers, there

is no other place in Alaohua County which even approximates it.






-653


Cool, '.crric m ttaOr ryuCde r-: : -":. Ith cnsicdoranblo

epcad, sM t'o lari:, reOC" krve.o of V. Snooodc 2crm a8 conoploorsa

olo-.entA of t'o 1:r.3tt,;


?.o0 linrcr b'lyild rotreate in l pro.d 'iA trach rid bito of

dobric l t' on roolks &! vol rct. Xn frott of tetA rOtrcIt they

uilk' s rot to etrr" fccfZ front t':o flo.e- wsatr, lot for :rnpatiC

t".:", crn n oi!d c

Z necoasltaed thi rprovl oucldy nteon it-nt':ro ctnaoo wu th

t%'or rEult forw ty rctrir;In howveorl, C, tVh larvte t iro very wmn-

rn.'oliatio Mssd ia i loa e inO..ol.i'."er tolornnaeo, I o'tn-arnod tost

ror.'tto by tC':!-' r-ifl :;rctca ocf ptinO 3d pcH-;)% t': o in Er-f

eotto"ns 'Ocvatoo C*:lt:' c-zrod Cr4 <- ovrr 57y '.lre, 7: f C1i-

Ito acOroultini tn raadoe t'.r-a2. ct':; cf U.o thtval aoliurttroc,

tc;t:i wVero ncv'Vd :rto Vte' ;'oetor':c -rortlron of t'io Vncrtfod puncli




A Inr rx nrlr or of lro tni' ;.:;rco dloi gl tio 1i.l transg-

rpTrteO In ',ter*i "ar ticien io.r. t nveo rm d Svootr-timi for trs-ne-

poertatict n mau W fr r othe vat '"-roeroiro :roed,


'"oe tnd'to foI.) crrs! !7r"AO1*fln:ro to Ity tioir o:.so .o

tt:, t'o or'a* t '" "of r- '. m rI -lt' I toCd livO, wn'ult

eicrilro of j. aio;! n T'r.cod than w-:*mr.?at-or. Thteo hoir: eat

tie boiTy nrd ifnre rproveort9 O t'on frtom rottir-n rots ,'d ithe oc rFeic

flioa, mn rel3oncro 'boo'ce to -'-n rirre, c't.1: 1roTo t'rour'> t'e






-64-


surface film and flow awoay Complete eubmergenoo did not seem to
affect them in the least,

Previous Florida 'Peorclst Ross (1944) lists this pooies as occur-
ring in Florida but does not specify the locality.

Speoimns Examinedt Two males, 6 females. Tavarea, Lake Co., P.
E11 Y., 3/23/36 7 larvae, Sweetwmter Creek, Liberty Co., L. B,#
12/40/S7t 2 larvae, Shoal River, Okaloosa Co ,, L. B, 12I/11S/
16 larvae, Santa Fe River, J. S. R., 8/12/381 24 larvao, Hillsboro
River, L. J. M., 2/18/ 9~ 1 female, Santa Fe River, L. B., 3//40O
I male, Santo Fe River, 4/6/40; 3 females, Blue Springs Creok, Jaolk
son Co., L. B., 6/E/401 11 Irrvne, Poe Springs, 10A4/47; 54 larvae,
6 pupae, Poe Springs, M W. N, 3/24/501 3 female., Big nlathot
Crook a li, t li M, W, No 6 2/60t 1 male, 4 females, Hogtom

Creek, at light, MI W, T., 6/49/60s 104 larvae, Poe Springs, M, W.

iN, 6/21/60; 1 male, Poe Sprinnre, smeepinc, M. WVT, r*, 6/21/60; 275
males and females, Poe Springs, at light, '!. N, 7/2/50s 125
lervee iend pupae, etre~em .+ mile S. of Gulf Tammonok, Levy Co.,
M*, WT, ., 7/30/60; 9 males, 23 females, Poe Springe, at light, TM.
IT, ,# 9/9/501 approxz 2000 larvao and puppo, Poe Sprir-s, .T W,
IN, 6/6/50; 175 males and females, Poe SprinCe, at lirht, TI. W. i#,

7/s/51.






-656


Hydropeyhe spe A


FIG. 100A---mydrop2ydhe sp. Aj
H1iAD OF LARVA.


The larva (fig. 100?') of this specion is undsoribod, and I

have not-yet been able to ascoo~Ito it with an adult forms indeed no

adults of the genue Rydroppyohe, other than 1. incoioda, have been

collected in the state. I have not found those larvae in Alachua

County.


Specinen Examinedt Two larvae, Jasper, Eoamilton Co,, L, Be

2/4/383 2 larvae, Lenont, Iedison Co., L. B,, 2/6/38: 1 larva,

Drifton, Jeffereon Cow, L. B., 2/6/388 4 larvae, Golden ITeaa State

Park, Clay Co,, I. W, 1I., 8/12/50.











CT.!isATOPSYCHGIT allengren


ostt of the small, broan Trichoptera larvae with bushy ab-

dominal Gills belong to this gonue. Both larvae and adults are

very similar in appearance and behavior to the larvae and adults of

!Sydropsryhe.


According t~o illiam Beok, Jr. of the Florida State Board

of Health (personal co~unmioatioin) Cheumatopsychid larvae will

tolerate pollution of a minor degree while most other caddis flies

will not. Personal observation bears this out; for example, in the

summer of 1950 the Eest Brnnoh of Hogtown Creek had a caddie fly

fatma composed almost entirely of Cheumatopsychid larvae, while the

West Branch had a more varied fauna* Many old, empty Leptooelia

and Oeoetis oases s were found in the East Eranoh showing that other

species had lived there. Since it is known that the East Branch

occasionally becomes polluted, it would seem likely that the areek

was slightly polluted in the su.mor of 190Q, and that only the

Cheumatopspyhid larvae could withstand the pollution.


I collected many larvao in Alaohua County, for this genus

is abundantly represented. To structural characters haie yet been

found to separate the larvae to species; therefore, these records

will not be given. Liarrv of Cheumatopsyche are very coacan and

occur in most of the streams in the county*



































FG 03--











cheynaonynh 0












UF 7, .
710 10 -- Chmaryh buki






-G*S


Key to The Speates Of Chenatqovyche

Laryva-----atot xeod. See fig. 10.S


Adults

1, Genitilial of mnles as in fig. 104 egenitilia of re-.
males as in fig* 103.
CheumatoMgoh burkai


2, Genitaltf of males as in fig. 105) genitalia of efo.
males as in fig, 109.
C 10hou-.rnMAtofsy Cho afalis


3 G Oenitalia of sales as in fig. 1061 genitalia of fe-
males as in fig. 110.
Cheumatopsyohe 9ahanta


' 4. Genitalia of males as in fig. 107.
Cheunatopsyche pitaca





Choumatopgyohe buyrlsi Roes


Ranget Florida and Illinoisa, Undoubtedly further collcctir-. will

fill in the gaps in our knoiledGo reGcrdinG the distribution of this

species,


DosoriptiDon RosS, 1944*


RemArkas I aseoclated the previously und@ecribed larva with its,

adult form by rearing pupae in damp cotton. No morphological char-

aoters were found, however, to distinguish thia apooloe from the

others In the genus.
























A

c
FIG. 107-------,heuTMto,)s li. 2Lnaa, UALE GENITALIA.
A, LAfk]'-lZAL ASPECT; B, CAUDAL ASPECT CIF TE1171 T,
C, CLASHR, CATMAL ASPECT.




IV





A
FIG. 108----Cheumatopsyche burksi, ms"ilp'LE
GENITALIA. A, IXTEAL WSPEECT OF ITINTIT Al-.,D
TEENTH TERGITES; B, DORSAL ASPECT.











FIG. log --- C. FIG. 110 --- C
FEMALE
-Ea -li S aphanta, FE."LuE
GENITALIA. GEITITALIA.






-70-


Those lrtvae ooeur in immense numbers on leaves of eel grasn

(Vallisneria sap) and rooks at the dam in the Santa Fe River near

Poe Springs. The larvae are very similar to Hydropsychid larvae

and are oanTonly found assooint-., ":ith them. The 1..rval nets of

both Cheunato syhe burkri and Hydropsyahe .inCooroda are easily and

ofto oseen the submerodc rooks and vegetation.


Previous Flori'da Reeordat 1Ross (1944) rooorde'tnis t Os pocl from
Tava-aes.


Speoimoue Examineds Sinoo extraordinary numbers of lar vae can be

found at Poe Springs any tim of the 'year, I will omit ey records as

they are not indicative of the numbers present. One male, 7rdla!ia,

Putnn~ Co., R. B., 3/6/46i 250 males and fewless, Poe 3prings, at

light W. IT ., 7A12/50 1 mible, 5 fennalse, 'Torthington Springs, at

light, m. 7. ,, 7/3/50; 27 females, River Styx, at light, s1. 7. 0TX,

7/27/50 j. males 23 females, Poe Springs, nt lijht, T:. W, I,*

9/b/0so 280 males and females, Poe Springs, at light, Mt. WT. .,
7/3/51.




Choumra&topElohe analis (Banko)

(Hydropsyoho analis Banks, 1903b
Ilydronss'ch_ ^ it^i Bar'rs, 1903b)

Rane. Widespread, ocourring over most of the United States with

the exception of the Southwest.






-71-


Deaoriptiunt Ross, 1944.

Remarks Cheumatoppycae analis is a rare oaddis fly in Alachua

County. According to Rose (1944) the larvae inhabit snall streams

but also occur in larger rivers. They have a wide coological toler-

anoe and are often found in streams carrying considerable pollution.

Previous Florida Recordst this species has not previously been

taken in Florida.


Speoina Examineds One male* Ebro, ahington Co., L. S., 5/30/401

1 male, Gainesville, no date; 1 male, 1 female, Hogtown Creek, at

light, M. W. ., 6/19/601 4 males, 3 females, Hogtown Creek, at light,
M. W. X0 7?/160.




Chaumatopsyohe aphenta Ross


Ran!gy The range of this species is poorly delineated, It ocurs

in Illinois, Indiana, Arkansas and possibly Florida.

Desoriptiont Rose, 1944.

Remarks The identification of this species is based up n two spea-

imens, both of thich were females, These specimens did not quite

agree with the description of the species given by Ross (1944), so

I sent them to Dr. Ross for confirmation of identification. He said

they were certainly very close to Cheumatopsyche nphanta, but also






-72-


suggested that they might represent a different spooios present in
this locality* It therefore seems advisable to record this species

as questionable until males are secured which would then corroborate

the record.


Previous Florida Recordsa, !tone*


Speimaens Exanedt Two females, Poe Springs, at light, M, WVt N.,

9/9/60.




Choumatopsyehe pinaoa, Ross



Riget -Southeastern states.


Desoriotions Rose, 1941.


Remarks Cheumatoprohe pifnaooa as collected on two oocaiaons, both

times at light, at widely separated pointso-*-* Poe Springs end Hog-

town Creek.

The larva has not been found, ooasequently nothing is known

regarding the biology of thia species.


Provioue Florida Recordse This spooies has not previously been

recorded from Florida.


Specimens Examinedt One males Poe Springs, at light, M, W. i0., 7A2/601

1 male, Fogtown Creek, at light, M. 'W. ., 7A/6/0,






-753


IACRCTE'ULI Burmeister


Macronemum arolina (Banks)


Ranse Southern states.

Pesoriptoan Ross, 1944.

Remarkas The adults are among the largest aeddis flies found in

Florida and are very striking, with their extremely long antennae

and polished, brightly-colored wings (fig. 92).

This species is only occasionally found in Alaohua County.

1o larvae were collected; however, the larvae probably inhabit the

deeper portions of the Santa Fe River, for Ross (1944) says that the

species frequents large, rapid rivers.

Previous Florida Recordst Rose (1944) lists this species as ooour-

ring in Florida but does not specify the loonlity. Betten (personal

oonmmnlcation) has taken specimens at Lake ITewahitchka,

Sneoimons Examineds One male, Ebro, 7ashington Co., L. B., 5/30/40;

1 nmle. 6.4 miles W. of 7ashington Co. line at Hwy. 10, nalton Co.,

L.o B., 6/31/40s 2 males, Bayou "Marquis, Esonmbia Co., Lo BD, 6/1/40;

Smale, Poe Sprinegs at light, MS. V. N., 7A2/50; 1 male, Worthington

Springs8 at light# M, WO No,, 7/1/S50 1 role, Prarie Crook, at light,

MH O !., 8/29/60.


















"'TZ















T 7











FI G 113 APEX ,Jr, 11
Al T)
CF ARD U L 77
A] AL












FIG. FIG. 117---
-H(-)R A X TT-TORAX.











T-o1 1TTLDiAS


This family comprises most of the "micro" onddis flies.

The adults are extremely minutes some having a total length of only

one end one-half millemoters.


A good deal of work has recently been done on the HydroptilidRe

of Florida. Denning (1947 and 1948) studied the miero oaddis fly

fauna of south and central Floridn in some detail and deoribed a

number of now species from there. Rose (1948) ias recently described

some new species on the basis of materiel 'sent to him by a number of

entomologists collecting in the state.


Many fydroptilidno have a very wide geographic distribution

but seen to occur only in certain limited localities within its

ranges for instance, lMaatrichia yama ocoure from the extreme

southern portion of Mexico end Florida to New York and Montana*

but withn that broad area its distribution is spotty. It is very

evident that muah more work needs to be done on the geographic

distribution of the whole group of micro caddis flies




Key to The Genera Of Hydroptilidae

Larvae

X1* middle end hind legs almost three times as long as
front legs, fig, 1l11
QOyethira

Middle end hind legs not more than one and one-half
times as long as front legs, fig. 112*.............*...*.....











2. Anal legs distinctly projecting from body mass,
fig 115.......****************************f****************

Anal logs apparently combined with body mass and
only the claws projeotingr *******........... ...............


3. Thorncie termites clothod with long, slender, ereot
inconspicuous setae, fig. 114; case of sand grains,
evenly tapered and without posterior slit.
Neotrichia ranen

Thoracic tergites clothed with shorter, stout, black
setae which are conspicuous, fig. 1165 ease trcns-
luocnt, evenly tapered and with dorsal side either
ringed or fluted with raised ridges, fig. 132.
itypatri Cain iE~DURG.

4. Case long, smooth and round in cross section,
tapered at each end and with an indented slit
at both ends.
Orthotriohia

Case purse*lise (wcaants change purse),
Hydroptila


Adults

Ooelli prbsent*..*... ...es e** *********A******o



2. "Motasoutollum almost rectangular, fig. 1160
Orthotrichia

Metasoutellui pentagonal to triangular, fig. 117.
Hydroptila

S, Hind tibiae with only 1 preapical epur.
Neotriohia ransa

Tind tibino with 2 preapical spurs...........................4




















A0



.3'~
&I.foia ,7









9EE'TS C, FZ-AF GE 4 4 LI




A B
























































A. B


























7 A, 8 9






, ..-


4. m idlle tibiae without a preapioal spur.
1 aystriohi a '. '

HIiddle tibine with a preapioa aspur.
Oyeothira





OXY=TiTIRA Eaton



Six apeioes of this genus are reported from Florida; how-

ever, only one,.O, 0 anella was found in AlaohuaaCounty* No ohar-

aoters have been found which oan be used to determine the species

Of the larvae.




Key To The Species Of Oayethira


Larvae-----Not keyed.


Adults

1 .M ale genitali as in fig. 118t female genitalia as
in fig. 118C0
Oyothira floridn
2. TNale goritalia as in fig. 119.
C 3la'ira sa


8. Male genitalia






64 Male genitalia
5. Ma'le genitalia


as in fig.-120.
Oyote'irn gne1llo


as in fig. 121.
Oerethira lumosa


as in fig. 122.
Oxyethira verna











GC Male gonitalia as in fig. 123.
Ovyqthira walteri





(Oxethirajnenella Denning



Raols Florida and Lomiainna.

Desoriptiant Denning, 1948,


Romanrks Oxoethira eanxella was described by Dennimg in 1948 on the

basis of two male speimens; one from Winter Park, Florida and the

other from rew Orleane, Louisiana. This species is the only rep-

resentative of the genus Oxyethira that I found in Alaohua County*

Since a lea of janellZ were collected along with females

at both Poe Springs and Vorthingtan Springe, it oan be assumed

that the fonnele are the undoscribed forms of this spooieo. Teny

other females of Ox_0 thirn, collected at the River Styx and Dig

Hatohet Creel., could not be speoificnlly identified due to the fact

that no males were project. In all probability tV'se females be-

long to the apeclea 0. janellea howeverE bor.use of a lack of oomw

parative material, I do not feel that I can place a nnmo oa them

at this time.

Although almost all caddi fliees that are attracted to li)ht

come directly to the source of the light, I noticed that 0* janella

came to the nore dimly illuninatod portions of the hBeet, such as










the back of the sheet, indicating that they are less strongly photo-

tropio than most other species.

Previo's Florida Rocordst "Holotyne, nalo--*-Winter Trk, Florid..,

May 16, 1940, (H. T. Fernald), Par~lt --*-ew Orlems, Loulsiana,

October 10, 1945, (D, G. Donnin), I male." (Denning, 1948)

Speoimane Ixamineda One r lo, Hogtom Creek, at light, 1 k:,. i.,

/1/650 7 males, 02 females, Pee Springs, at light, M1. N.,

/7A/50t I mle, 11 females, WTorthington Springs, at light, I. V7. N.,

7/13/50s 2 nales, 16 females, Poe Springs, at light, I!. I ,I JI.,

7/5/51,




Oxethira florida Donning


Ranat South Florida.

Descritiont Denning, 1947.

lomarka: Oxyethira florida was described by Denning (1947) on the

basis of specimens collooted in "ianmi. I have not found this species

in Alachua County, and therefore, I assuam that it does not oceur as

far north as Oainesville.

Previous Florida Reoordset "Iolotype, ale-masMiafZS, Plorida#, Feb. 1,

1945, D. 0, Denning. Allotype, female*-wHaian, Oot. 1, 1944, D. G.

Denning. Paratypos, four males---' tLanI, Florida, nTv. 15-30, 1944*






.32-


Lliht trnp, D G. Denning. Peratypes, thee males, eleven foe-alos

---..--Tiai, Florida, Oct. 12-30. 1944o Light trap, D. G. Denning.

Paratypcss fiv-o nalos, six females, .1.i-ni, Florid Oct. 1, 1944.

Light trap, D, G* Dennange Farntyhtcs, five =aleo, nine fnelos,

Liami, Floridn, March 1-15, 1945. D. G. Denning Panrat'pos, coven-

teen nales, twcnty-nine females, IUiani, Florida, PFb. 2-8, 1945.

Light trapD.o G.: Denning." (Deaning, 1947)


Specimens Examinedt N neo




Oyothira gLtar Bosa



Range Oklahoma, Florida, Georgia and Louisiana.


Desoriptient Rose, 1941.


P.marke: Donning (1947) found 0. lasa to be a Lbndont in southern

Florida, bat it has not as yet bean collected in Alaohua County,

Since this species does occur in Georgia and Louisiana, it mnfht

be ass~jud that it does inhabit some part of northern Floridn.


Previous Florida ecords: "Floridat Miamit February 1"8, 1945,

light trap, D. G0 De inning, males. February 16 1944, Do G. Denning,

19 males. February 2n-28, 1944, D. G. Donning, 16 Pales*. !arch

1-16, 1945, li.gt trap, D. G. Denni-C, 2 TnMale3, ovorlhr 15-SO4

1944, li!ht trap, D. G..DonnrinC, 16 males. Deooober 4, 1944, D. G.

Donning, 14 males. December 14-15, 1944, ligCit tr#p, D, G. Denning,











28 males*


Sgomlnena Examinad c None.




O2rethira 1umos~as Ross



Rnan Florian.


Desoriptiont Rose, 1948.

Ronarkcas 9oyethira lumosa waas described by Rosa n the bnais of

one male collected at Daytina Beach. It has note to my ckowledee,

boon oolleoted slnor


Previous Plorida Records: -olotjgY <..k-"<---Tcaytoca Po oh, Fla.,

AuCusn 27, 1945, G. T. PieOcl." (Rose, 1948)

Spe'~naons Exmnined. ITone.




e athira vorarn Ros



RBanE* Nw Brunmwick, Illinois, Florida. rnd Loutisient

Desofnriti
nenar~et Aaoordinc to DonninG (1947) Cyohliirr. vrri.a is the nost

abundarit spooioa of iydroptil-dae in sout:crna 1lordrita; honovcr, I







084.


have not collected It In Alachua County. It ap -ers to have a very

rddo goosraphio distrirtion but only occur in cortAin aroas Within

its overall rsniee


Previous Florida Records: "Ploridat 'uian February 1-8, 1945,

LfghCt trap, D, G. Denuing, 168 males, 92 females. February 15,

1944, D. G6 Denning, 11 males. February 23, 1044, D. G. DonninG, S

na.les. March 15S0s, 1945, D. G. Denning, 12 males. April 10, 1944,

D. G. Denning, 56 nmloo. October 1-3, 1945, Licht Trap, D. GO Denning,

5 males 28 fenn les. October 12-30, 1944, Light Trap, D. G, Denri0ng,

25 males, Sl females. I:oveo:ber 15-10, 1945, Li(ht Trap, D. Ga Denning,

34 raloe. December 4, 1944, Light Trap, D. G. DerninG, 18 males.

December 14415, 1944s Light Trap, Da G. DonninG, 23 males. December

20, 1944, Light Trap, D G. Donnings, 32 males." (Denning, 1947)


Snecirnens Examinodt Cone,





OxyethIra alteri Denning


Ranger Florida, GeorCia and Louisiana*


Description Denning, 1947.


P.en-rrl:sz Denming (1947) collected 0. walteri in !iaamir however, I

have not found it in Alachuaa Oounty Since this cpeaecs ocura in

Goorgia, one might surrniz that it can be found somewhere in northern

Florida*
























































ameica "ALE GEMTAIA







*38.


Previous Florida Reoords: "'olotjiypro mrle*m-c-Plorinz Ujiarit

tovombor 50, 1944, Light trap, Di Go Denning. Paratypeos: ;Linr.it

Ilarch 15, 1945, D. 'G Denning, 6 mnrle. 'ny 10, 1^.4, D. G. DSrj.1inC,

12 males. :'wovembr 15, 1944, D. G. Denning, 4 -nles. Dooembor 20,

1944, D. G. Denning, I ale." (Donning, 1947)





Cr.THOTnECI If. EatCn



SoRX femalos of this gonus, found at several localitios

(Rivor Syxr iorthingrta Sprinco, Devil's -Tllhoppor and Poe Springs),

could not be identified speoofloally duo to lack of oomparative

material. A larva found in Lake Alice al2o could not be identified

to pooeies.



Koy To The Stocies Of Ortheotrie:ina


Larvae*-- -i-lot keyed


Adult

1. iIalo genitalia as in fig. 1241 female genitalia as
in fig, 127.
Orthotriohia amerioana


2. Unle qo nital'a as in fig. 125; fnrale cenal1-ia na
in fi. 128.
Ort'atrialain critata


38 Malo oenitalia as in fig* 126t foenles not k~yed.
Orthotrichia instabiliS



























Orthotriohia americana Banks


Reaggt This speolis has a wide range enoompassing an area from

Texas and Florida in the South to Nw York and Minnesota in the

North,

Description Ross, 1944.

Remnarkst Denning (1947) found 0 ameriant to be present throughout

the year in Miami howovor, I have not tabn any males in Alachua

County This species does occur in this section of Florida, though,

for peo lmons from the St. John's River in Putnam County were ex*

amined by me. Perhaps the unidontifiable fo-ales and tho larva

found in Lake Alice also belong to this species.


Previous Florida Recordes "Poridat Mianit DOtOber 1-S, 1944,
Light trap, D. G. Denning, 40 males, 7 females. October 12.4O, 1944,

D. G, Denning, 7 males, 2 fomales. November 15, 1944, D, 0G Donning,






488.


1 male, 1 female. February I-8, 1945, Light trap, D. Go Denninge
21 nalos, 15 fomalea. 7Tarch 15, 1945, Light trap, D. Go Denning,

1 male. ITay 10, 1944, D. Go Denning, 14 males, 12 females."

(Donning, 1947)

Specimens Examinedt Four males, 10 females, shore of the St. John's

River, at light, J. H., 7/5A/60.




Ofthotriohia oristata Morton


Range ost of eastern end part of western United States.

Description: Ross, 1944.

Remarks Denning (1947) collected one male and one female in niami.

I havo not found this spocios in Alachua County.

Previous Florida Recordes "Florida T HiaTmit .April 10, 1944, Do G.

Denning, 1 male, 1 female." (Denning, 1947) .

Speolmens Egminedt None

































A B c
FIG. 129 ---- Hydroptila berneri, IIiALE GENITALIA. A,
LATERAL ASPECT; B, CIA.SPERS; C, FE71ALE GENITALIA.













B

FIG. 130----Idr2p-bila wakulla. A, ITALE
GEIJITALIA; B. FEMALE GEITITALIA.






-90-


Orthotrichia instabiliae Denning


Rlen t Contra.l Florida,


Deaoriptionz Denning, 1948a.


nemarku: Denning (1948a) described 0. instabilia on the basis of

Nine males colleeted in Winter Park. I have not found this species

in Alachua County and can give no further data about it.


Previ ou3 florida Recordst "Holotype, nrmle--Z--.&inter Park,

Florida, .lay 16, 1940, (H. T. Fernald). Paratypee ---- Florida,

sem data as for holotype, 8 nales." (Donning, 1948a)


Snooimons fBlandnodt Yone.




HYDROPTIIA Dalman



This Conus embraoes abcut forty Fearotio apocloes comprising

cne-thilr of the Hydroptilidae. Only two species, H. berne. end

t. wakulla have been taken in Florida,


Key To The Speloes of Hydroptila


Larae.--Not keyed.

Adults

1. Hale conitalia as in AfiG 129 A and BS female genitalia
as in fic. 129 C0
fYdroptia berneri






-91-


2, Male genitalia as in efie 1SO A fA;o.lo genitalia As
in fig, 130 B.
Hydroptila watl1la




&ydroptila borneri Ross



Rangt Floridan.

Descriptiint Ross, 1941.


Remarks This species was described by Ross (1941) on the basis of

two males and one female collected at Poe Springs by Dr. Lewis

BDrner. I was able to collect topotypical material and was aloo able

to collect the uidosoribod larva. This larva, which is fairly

oonrno at the rook dam near Poe Springs makes a purseeshaped onae

of sand grains. The eases are attached to submerged vegetation and

rooks in the slower moving portions of the river around the dam,

The adults are attracted to light, but seem to come to the

light a little later in the evening than most of the other caddis

flies.

Previous Florida Reoordst Two males, 1 fealdo, Poe Springs, at

lidht, collooted by Lowis Bernor, 3/4/39. frontn Rose, 1941)


Speoimonsm Examineod Four males, 10 females, Poe Snpring, at light,

U* A7. 7/12/560 1 males Big Hatohet Creek-, at light W5 is,

4/i2/60 1 male. Poe Springs, at light, 1U. vT. It, 9/b/s0o 3 males,











5 female, Poe Springs, at light, Io VT. T,,* 7/3/561 6 larvae Poe

Springs, Ml' V 1. Nl., 3/24/60.




hydioWil.a xkulla Donning


Rgn2ea Florida.


Des oipticai Denning, 1947.


Remarket ydropftila waklla was described ly Denning (1947) an the

basis of 11 specimena collected at Wakulla Springs. I havo not fo~nd

this spoolo in Alaohua County end have found no othor data oonoern.

ing it in the lteoramture.

Previous Florida Recordst "nolo%'po, male-----Florid1t Tfakulla

Springs Ootober 23, 1945, D. G. Dening. Allotype, feomle*-m-*-

Same data as for holotype. araty -S data as for holotypoe

I male 8 fomales.s (Denninc. 1947)


Efpeoimiens Exawnined Ncmne.


-32-











NEOTRICIHIA Morton


1eotriohia ranea Denning


gafmLt Georgia end Florida,

Deooriptiont Dennlng, 1947.


Remarkast Notrishia reanea is undoubtedly the smallost caddis fly

that occurs in Alaohua CounCty a full-grown male is omly one and one*

half nillimeters long,


This speooie was described by Denning (1947) am the basis of

two males, one from Florida, the other froa Georgia. I have collected

the undesoribed female of the speoles.

Previoue Florida Peoordez "One naelo Ilionl, May 10, 1944, D. Go

Denning." (Denning, 1947)















































FIG 13 -- FG 14--M











Speoimlena IxaAmnedt Two males 2 famaleos, 7ort'iingc'o .prinsg, at

light T, V. II, 7A/SOij 1 female Poo Springs, at light M -T. T.,

7/i2/50t 1 malo, 1 fomalo, Poo Springs, at light, I o. I* E#, 9/b/50.




MlYATRICUIA Moeoly


This genus contain three North American apeoies, of which

omly one, M ay aa has been taean tn Florida,


layatrichia Mlosely


r nOt Very iido-spread over most of the eastern states. Ranges

front extreme southern t.oxloo to oew York end Iontana. Although the

records are scattered over a wide area, the spooios is quite rare

in collections*

Desoription s Rossa 1944.

ra, arkst P.oac (1944) lists this species as occurring in Floridn, I

exarinod sixz females whioh, due to the lack of comparative material,

cannot be identified to speoious however, these specimens (one from

the St. Jolm's River in Putnam County and the other five from Poe

Springs) most probably belong to I. ayara

Previous I'lorida Rooords: Rooc (1944) record this spoo5io from

Florida but does not qpecifj the locality.






*96.


Spooirnons Examinedt Ictae.





PHRYG.A!EID.AE



BA:K3IOcIA Martynov



Bankaiola conontenata (Walikr)



Waller (1852) described this species oa the basis of speci-

mans caught at St. John's Bluff, East Florida, which is near the

mouth of the St. John's River. I have not collected this species

but en including it for the sake of ocmpletoneass





LI!TCD'ILIDAE



PYCIPZYCTTi E Banks



The family Linmephilidao is a very largo one containinG

numerous species scattered throughout the United States, and thus

it is rather uprising to find caly one representative ooaurring in

Florida.




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xml record header identifier oai:www.uflib.ufl.edu.ufdc:UF0008998200001datestamp 2009-02-24setSpec [UFDC_OAI_SET]metadata oai_dc:dc xmlns:oai_dc http:www.openarchives.orgOAI2.0oai_dc xmlns:dc http:purl.orgdcelements1.1 xmlns:xsi http:www.w3.org2001XMLSchema-instance xsi:schemaLocation http:www.openarchives.orgOAI2.0oai_dc.xsd dc:title The caddis flies of Alachua County, with notes on those of Floridadc:creator Nirenberg, Marshall Warrendc:publisher Marshall Nirenbergdc:date 1952dc:type Bookdc:identifier http://www.uflib.ufl.edu/ufdc/?b=UF00089982&v=00001000546271 (alephbibnum)13213009 (oclc)dc:source University of Floridadc:language English


THE CADDIS FLIES OF ALACHUA COUNTY,
WITH NOTES ON THOSE OF FLORIDA
B7^v,e,
M ARSH ALL * NIRENBERG
A THESIS PRESENTED TO THE GRADUATE COUNCIL OF
THE UNIVERSITY OF FLORIDA
IN PARTIAL FULFILMENT OF THE REQUIREMENTS FOR THE
DEGREE OF MASTER OF SCIENCE
UNIVERSITY OF FLORIDA
February, 1952

TABLE OF COHTERTS
1» Introdxjotion.......
2.. Acímowled£#»ents«i,,*........ .......................*9
5. Collecting. * »»*10
Preservation esd Clearing..IS
5, Kearirtg##••#••«• ••••••••!•••....;••..*..•;••••• •#•••14
6* Kay tó FeuailioE****.*.*«¿«.•*••••*.••-••*♦•*•••*••*29
?• Annotated List**.......29
84 . Unsolved Prohlens *..**..***.161
9« Summary and Conoluslcns «.164
10. ..References...«*166
11* Check List of the Gaddis Flies of Florida
and Index*..*172
12» .Biographical Bote**».......*..**.***..177
i

IlJÜ&QDtrCTIOH
Historically speaking, some of the caddis flies of the south¬
east were described at a relatively early date, about one hundred
years ago. At that tine Francis Walker (1852 and i860) of the British
Museum described twenty species from Georgia which were sent to him
by Abbot, and this constituted the earliest large collection made in
the southeast. The investigation of caddis flies in the south stag¬
nated for the next eighty-five years, for example, fifteen years ago,
two out of three species collected in the state would have been un-
deseribed. Since then much incidental work has been done by visiting
trlChopterists and by entomologists working in allied fields.
Gaddis flies are fairly abundant, but as compared with other
states, Florida shows a general paucity in the number of species
present; a paucity most probably due to lack of knowledge rather than
a lack of caddis flies. One hundred and eighty-four species have
been found in Illinois, a thoroughly studied state. On the other hand,
only sixty-three species have been found in Florida, thirty-eight of
which occur in Alachua County. Some day when the state has been in¬
vestigated a good deal more thoroughly, this number will undoubtedly
be greatly increased.
In the Trichoptera, as in other orders of insects, our know¬
ledge of the immature stages has lagged far behind that of the adults.
The larvae of many species of caddis flies known to occur in the state
still are unknown or unassociated.

Betten (193*0* Balduf (1939)» end Eoss (1944), have ell care¬
fully and fully summarized the extensive literature describing and
detailing the life histories and habits of caddie files, for this
reason, no further discussion of this subject will be included here.
The current status of the knowledge of caddie flies in Florida
can be stated concisely. The species are very incompletely known?
comparatively few larvae have been associated with the adults. The
larvae of some genera have never been found and much more collecting
of adults is needed. Geographic variation of the Trichoptera of the
TMted States has never been investigated thoroughly, little col¬
lecting has been done in northwestern Florida and many new records
plus undescribed species should be found there. Obviously, there is
ample room for far more study.
This research problem was specifically United to the caddis
flies of Alachua County? however, as I had access to material collec¬
ted in northwestern Florida. 1 decided to extend the scope of the
study to include all that was known and all that I could find out
about the caddis flies of Florida. This additional material consisted
of caddis flies collected by Dr. lewis Berner in 1939 and 1940, and
also specimens which were in the collections of the University of
Florida.
Extending the study by the inclusion of this supplementary
materiel greatly broadened the entire picture, but also, simultaneously,

complicated it, for the collections examinad -were sufficient to false
a multitude of questions hut were insufficient to answer «easy of them*
For orrery problem solved, two were discovered which oculd not he an¬
swered*
this report is the result of an Investigation carried on for
two years——from February, 1950 to February, 1968——«in rMch time
1 collected approximately 10,000 specimens and examined an additional
eight hundred individuals which were collected by others*
The purpose of this investigation is to present* (1) an
annotated list of the «saddle flics found in Alachua County* (2) a
cheek list of the caddis flies of Florida, (S) keys for the identlfi*
cation of the larvae end adults found in Florida, and (4) a biblio¬
graphy which includes all references to Florida .caddis flies* In
short, this investigation has two primary purposes* to report m the
caddis flies of Alachua County and to gather together all that is
tocen about the caddis flies of Florida* As such this report might
well serve as a foundation for a future study of the onddie flies of
Florida*
On the basis of the Information gathered in this study, which
of necessity is very incomplete as collections from parts of Florida
Other than Alachua County are very spotty, the state of Florida om he
subdivided into two areas having distinct caddis fly faunas* (1)
Northwestern Florida, and (2) Peninsular Florida* Northwestern Florida

certainly has a fairly distinct fama for many species of caddis flies
having northern affinities gad are found here which are not collected
in the Gainesville region, "fhe Apalachicola River drainage has with¬
out douht been the main highway of ingress to Florida for the great
majority of animals which require flowing water or hardwood forests.
Rogers (1933) found that the ecological conditions existing in the
Apalachicola drainage would admirably explain the distribution of
many northern craneflies in Florida. Carr (1940) reports that the
most extensive invasion of Florida by the northern element is encoun¬
tered in that portion of the panhandle which is drained by the Apala¬
chicola River. Hubbell (1936*35^) states that la this peculiar en¬
vironment (the deep ravines of the Apalachicola region) a great many
northern plants occur, evidently the remnants of a northern flora left
as relicts from Pleistocene times in these deep, moist, eool ravines.
«It has also been noted that many northern plants reach their southern¬
most limits in these ravines and many typically southern plants inter¬
mingle here with the northern species, hot only is this true of plants,
but such has proved to be the ease in the craneflies, the Odonata, the
Opilionids, the Orthoptera, the crayfish, the amphibians, and the rep¬
tiles, " (Berner, 1950). Peninsular Florida has a number of endemic
species, but the majority of the fauna is composed of species having
a far wider geographic range.
Caddis flies are fairly abundant in Florida — they ecu easily
but they do not appear to be present in numbers as large
be found

5-
as those occurring in many northern streams* I have never collected
caddis flies In the north or the west, hut I have read many descriptions
of the immense numbers of these insects which swarm in northern streams.
The rock dam near Poe Springs is the only locality in Alachua County
which approaches these descriptions of northern abundance.
In the north caddie flies ere an important link in the aquatic
food chain, for they convert small units of food into larger units,
and since they are present in tremendous numbers, they are a group to
be reckoned with. Caddis flies constitute an important factor in the
diet of many northern fish.
This does not seem to be true in Florida. I to not believe
that caddis flies are abundant enough (except in certain scattered
localities) to be a ma^or link in the aquatic food chain. Reid (1950)
found the number of Caddis flies eaten by the Black Crappie, Fomoads
nlero-raacnlatus. in Orange Rake was Insignificant. William McLane
(personal communication) found the same condition to be true while
he was working with large-mouth bass in the St. John*s River. Very
few caddis flies were found in the stomachs of the bass.
Caddis flies ere important in Florida, however, as Indicators
of stream pollution. Many stream forms of caddis flies have an ex¬
tremely low ecological tolerance and cannot withstand any stream pol¬
lution. According to William Beck, Jr., of the Florida State Board

of Health (personal communication)* Cheumatopsychld larvae will
tolerate pollution of a minor degree -while most other caddis flies
•will not* Personal observation bears this out* for example* in the
summer of 1950 the Bast Branch of Hogtown Creek had a caddis fly
fauna composed almost entirely of Cheumatopsychid larvae* while the
West Branch had a more varied fauna* Many old* empty Leptocella
and Oeoetia cases -were found in the Bast Branch showing that other
species had lived there* Since it is known that the East Branch
occasionally becomes polluted* it -would seat likely that the oreek
was slightly polluted in the summer of 1950* and that only the
Cheumatopsychid larvae could withstand the pollution* Ross (1944)
corroborates these observations* lie states that members of the
genus Cheumatopsyche can frequently succeed to some extent in streams
too polluted for almost any other caddis flies* Therefore, it appears
that eaddis flies oan be used to show comparative pollution* (l) if
a stream is unpolluted* there will be a varied caddis fly fauna present*
(g) if a stream is slightly polluted* only Cheumatopsy chid larvae will
be found* and (£) if a stream is highly polluted, no caddis fly larvae
trill be present*
Although this is the first systematic study of this group of
insects in the state* the Triehoptera of florida, and ©specially
Alachua County* were previously fairly well investigated* Gaddis
files were collected by Drs* Lewis Berner* J* Speed Rogers, Frenk
Young and others* and were sent to authorities for iisntlfiqaticm*

Various authorities On caddis flies nade quick collecting trips
throughout the state. Almost 'all- of this has been incidental col** "
lectlhg in scattered locations rather than thorough,' systematic,
fatuxal studies. Furthermore, information on the caddis flies of
Florida was scattered —— indeed much of it was unpublished — and
has "been inaccessible to all but a specialist. For these reasons
this problem has been pursued along the aforementioned lines? a
faunal study and a consolidation of knowledge.
The collector's names are abbreviated throughout the thesis.
The abbreviations refer to the following men* 3* A, B.t 3, A, Bar¬
rington; W, I!, 3,, .Vi, M. Beck, Jr.; L, B.f Lewis Berner; A. J. H»,
A. J, Henry; A, M. L,, A. M, Laessle; A. F, C., Archie F. Carr;
1, L., Ellis Lanqulst; II. IU, IT. Marshall; M, W. II., Marshall Miren*
berg; J. S. B., J« Speed Bogers; B. 3, 5?*, B. P. !Erogdoa; F. H. I.,
Frank Young; J, S. H», J. S. McCulliigh,
Most of the drawings in this report which illustrates the
keys were copied, and come from many different sources. Although
some of the illustrations were taken from the illustrations accom¬
panying original descriptions, it has seemed advisable to use the
best drawings available, and thus some newer, redescriptions were
used in place of the older, original descriptions.
All of the original drawings were made with a camera lucida
from uncleared specimens preserved in 85$ ethyl alcohol.

â–  Big source of each illustration is a s follows?â–  Figs*.1*36,
Ross, 1944; ■ Fig. 37» Flrehberg; Fig. 33, Rocs, 1944; Fig, 39, Ross,
1948;'Pigs,-40-64, Ross, 1944; Fig, 65, Forhies, 19091 Figs. 66-73,
Ross, 1944; Fig. ?4, Theur; Figs, 75-79* Ross, 1944; Fig, 80, Rosa,
1947a; Figs. 81-99, Foss, 1944; Figs. 100-100A, Bieur; Figs, 101-
106, Ross, 1944; Fig. 107, Ross, 1941; Figs, 108-117, Ross* 1944;
Fig, 118, Denning, 1947; Fig, 119* Ross, 1941; Fig, 120, Denning,
1948; Fig. 121, Ross, 1948; Fig, 122, Ross, 1944; Fig, 123, Denning,
1947? Figs, 124-125, Ross, 1944; Fig. 126, Denning, 1948; Figs, 12?-
128, Ross, 1944; Fig, 129, Ross, 1941; Figs. I3O-I3I, Denning, 194?;
Figs, 132-256, Ross, 1944; Fig, 157, Theur; Figs. 158-16?, Ross,
1944; Fig, 168, Rirenherg; Fig. 169, Ross, 1947; Figs, 1?0-1?1,
ITirenberg; Figs. 172-173, Ross, 1944; Figs, 174-175, Theur; Figs.
176-177, Ross, 1944; Fig, 178, Banks, 1938; Figs, 179-180, Ross,
1944; Figs. ISI-I83* ITirenberg; Figs, 184-185, Roes, 1944; Figs, 186-
18?, Theur; Fig. 188, Ross, 1944; Fig. 189-190, Ross, 194?; Figs, 191-
192, Ross, 1944; Fig, 193, Ross, 1941; Fig. 194, Ross, 1944; Fig. 195,
Ross, 1938c; Fig, I96-2OO, Ross, 1944? Figs, 201-202, Theur; Fig, 203,
Ross, 1944; Fig. 204, Ross, 1941; Fig. 205, Rirenherg; Fig. 206,
Theur; Fig, 20?, ITirenherg; Fig. 208, Theur; Fig, -209, Betten and
Mosely, 1940; Figs, 210-211, Ross, 1944.

*9.
ACM0WLEDGEMEKTS
Ihis study eould not have been completed without the help
of many Individuals* I take great pleasure In acknowledging the
time and thought spent by these raen and was greatly impressed "by the
willingness with which this help was extended*
I especially wish to thank Dr* Lewis Berner for his guidance#
constructive criticism, and for the many specimens collected by him
In other parts of Florida* Dr. Minter Westfall spent many hours
helping with the photography* Dr* Cornelius Betten sent a list of
all the caddie flies he had collected in Florida and a list of all
the caddis flies recorded in the literature which were collected
in Florida. Dr* Herbert Ross of the Illinois Natural History Survey
sent me many of his reprints, gave advice and identified some
specimens* Dr* D* 0* Denning of the University of Wisconsin sent
a number of his reprints* Mr* Ellis Lanquist, Dr. J. Speed Rogers,
Dr* Frank Young, Mr* R* P* Trogdon, Dr. B* A* Barrington, Mr. Jon
Herring and others too numerous to mention, col looted specimens used
in this investigation. Mr* Bert Theuer helped greatly in the
preparation of the keys by drawing the heads of some larvae*
William Beck, Jr., George X* Reid and William McLane. aided
by contributing bits of essential information used in this study*
Last, but by no means least, I would like to -thank all
those who contributed materially by offering criticism, advice
and suggestions*

40-
coi.Lsc'iro
Tnree different methods were used in collecting caddis flies?
(1) the larvae and pupae were collected from water* (2) the adults
were collected at li#it, and (3) adtrlts were collected during the
day hy sweeping vegetation overhanging water or hy searching crevices
underneath bridges* Since no one method will yield a representative
collection of the area, a combination of all three was used*
•
All Hesrctie caddis fly larvae and pupae are aquatic, and
almost any type of aquatic habitat is a potential collecting site.
Stones and pieces of wood were brought to the surface for examine-
tlon. Leaf-drift, aquatic vegetation, and debris were dumped in a
pile on shore* After drying for a few dinutes, the larvae begin to
move end can easily be spotted. Some pupal and larval cases show a
high degree of protective coloration and blend in well with their
surroundings, but tdth practice these cases can be readily distin¬
guished, ...
From a quantitative point of view X found the best place to
collect larvae and pupae in Alachua County was in rapids and riffless
for example, the rock dam at Poe Springs. The larvae are present in
this spot in tremendous numbers, X also found quantities of larvae
by examining the leaf-drift in some of the numerous, small, swiftly-
flowing, unpolluted streams scattered throughout the county; for
example, parts of Hogtown Creek. Larvae my be found in all conceiv¬
able types of aquatic habitats, for there are record® of immature®

-11-
being found In pitcher plants and the hollewed-out boles of tree
stumps that had collected a little pool of water.
Adults of most species of caddie flies will come to light on
warm# still nights# and I used this technique extensively* Specimens are
to be found on store windows* pink neon lights especially attract these
Inseote* VJhile some specimens cen be found in town on store windows#
lighting on the shores of streams# lakes# eto* is essential* I have
had good results using a Coleman# single mantle# gasoline lantern*
Caddis flies clinging to the lighting sheet eon be collected by
holding a small# wide-mouth jar half full of 86% alcohol directly
beneath the insect and tapping directly above the insect with this
forefinger# the startled caddis fly will leap backwards into the
'â–  V. '
open jar* Dipping an index finger in alcohol# scooping up the insect
rapidly but gently on the wet surface and then dipping it in the bottle
is a rapid method which 1 used for collecting many small caddis flies*
I have used an automatic light trap with success# The trap
was set in various places overlooking water and was periodically
checked*
In daylight collecting of adults# sweeping vegetation with
an Insect net Is a necessity for some species are negatively photo-
tropic end cannot be caught at light* Resting places differ tilth
the species# but most caddis flies prefer shaded# humid places* Vege¬
tation overhanging water end crevices of tree trunks are favorite
resting places* These insects are very fast and agile# and swift

raovemnt is required to capturo the»»
Frequently the adults may he captured on sticks and vegeta¬
tion floating m the water* When floating vegetation is submerged,
trapped adults will quickly swim to the surface*

>13“
SBB8SH7AH0H ARD CXSAEIIBO
AH caddis flies, both immature and adult, were preserved la
855® ethyl alcohol. One genus, Lento cel la. was collected dry since
identificatipa la this genus depends largely on the pattern of hairs
on the wings. Most specimens.of this genus were hilled in a cyanide
jar and were carefully pinned.
The genitalia were used to identify almost ell species of
caddis flies. It was usually necessary to clear the genitalia to
see the diagnostic characters. I have used the following clearing
procedure* The apical third of half of the abdomen was removed and
placed for five minutes in a hot five percent potassium hydroxide
solution heated in a water bath. It was then transferred to alcohol
and the viscera were gently squeezed out. The cleared genital cap¬
sule and the specimen to.which it belongs were put in a small vial
approximately ?lJ- by 4 ram. This vial was filled, stoppered with a
cotton plug end inverted in a ring neck 4 dram vial.
It was desirable to clear the entire body of the micro caddis
flies belonging to the family Hydroptilidae, The abdomen was not
detached. The procedure used is the same as above except in this
case it was necessary to teat a slit in the base of the abdomen through
which the dissolved viscera was expelled..

«•14"»
HEARING
Two major methods were used to associate larval caddis flies
with their adult forms. The larvae or pupae were either reared in
cages until the adult emerged, or the pupae were dissected.
Laboratory Rearing
Rearing caddis files in the laboratory is a difficult pro¬
cedure, for many species have a low ecological tolerance* and natural
conditions must be simulated. Therefore, In trying to increase the
yield of reared adults, I developed a number of different rearing
methods! (1) a series of ei$it screen wire cages, each enclosing
a balanced, aereated aquarium were used* Both larvae and pupae were
kept In the aquaria. All the larvae died? occasionally some adults
emerged from the papal case. Shen caged together, the larvae are
cannibalistic, and the larvae that survive are usually killed by
the change in their environment. Pupae, however, require nothing
to eat, ató. if the pupa Is mature, the adult will emerge within a
few days. Pupae of certain species are often rather scarce, which*
since it Is difficult to find enough individuals to raaki certain that
adults will be obtained, constitutes a major disadvantage* (2) Sub¬
merged rocks end vegetation covered with pupae and larvae were taken
from their natural habitat and kept moistened, but not submerged, in
large screen-wire- cages. The caddis fly immature® were able to get

16-
all the oxygen they needed by keeping them in contact with air rather
than water» The larvae died, hat many pupae survived and were reared
to the adult stage* Miefeel and Mllliron (1939) described a similar
method of rearing caddis flies. They placed pupae In shallow pie
pans containing damn sphagnum toss, The toss kept the pupae moist*
and the pupa© received more oxygen than they would have received in
water. 15y method was quantitative rather than qualitative, for if
two or more species of a genus were present, a specific association
could not he made, (3) The pupae were separated under a binocular
microscope and packed, In groups of five, in damp cotton. S&eh group
was then put in a shell vial covered with a wire screen top, This
separated the species qualitatively, however, only a small percentage,
approximately two percent, could he reared since many pupae were un¬
doubtedly injured in handling, (4) Larvae were reared in a balanced
aquarium with a strong jet of air aerea.ting the water and incidentally
stirring the water swiftly and constantly. A slow flow of tap water
fell into the aquarium, and a'siphoning arrangement took care of the
overflow. The larvae were fed with plankton. Time and lack of
equipment prevented me from making full use of this method,- but I
believe this method was the best and can be used to advantage with
both larva® and pupae.
Field Rearing
On three different occasions I set series of modified, copper-
wire pillow cages filled with larvae and pupae in the west branch of

-16-
Hogtown Creek near Gainesville, Theoretically, this should he an
excellent method for associating stages in,the life history of caddis
flies, hut in practice it was unproductive. More extensive, con¬
trolled field rearing should yield good results. â– 
Pupal Dissection
Certain pupae fora the connecting link "between larvae end
adults and can "be used to associate the adults with their specific
larval forms. In most caddis fly groups the larval seleritfca, which
are readily diagnostic to species, are packed into the posterior end
of the pupal chamber after the pupa is formed, dust before emergence
of the adult, the genitalia of both sexes becomes completely formed,
hardened.and colored, which therefore specifically identifies the
specimen.
If a cocoon or ease is collected which has a. mature pupa in
it, the larval sclerltes end fully formed genitalia can be associated*
and it is thus possible to linktthe adult and larval forms of the
species. However, it Is usually necessary to collect many pupae be¬
fore on® is found with mature genitalia.
The family leptoeerlda# constitutes a major exception to this
type of linkage. In this family the pupal chamber has a slit and not
a mesh in the closing cap at both ends. Through these slits the pupa
pushes out all the larval sclerltes, bo that it is impossible,to get

17-
associations of adult structures and larval sclerites in the sane
case. For this reason it is necessary to rear these larvae.
Care was taken in transporting the larvae and pupae. I have
had heat results by packing "both larvae and pupae in clamp mss. The
caddis flies were transferred to their permanent aquarium or cage as
soon as possible.

FIG« 2 Mol arm a LARVA, LEGS. A, FLU'T LEG; 3, MIDDLE LEG
C, HIND LEG.
FIG« 3 Trian odes LARVA, LEGS. A, PROMT LEG; B, MIDDLE
LEG; C, HIID LEG.

>19'
S:
KST TO FAMILIES
Kays to genera and apeóles are incorporated within the
annotated Hat* Refer to the Cheok List And Bade* at the end of
this volume for page numbers*
Larvae
1* Larvae (not pupae) living in and aarrying around a
definite case*
Larvae without oases*#............................
,.2
10
2* Case like a snail's shell* fig* 210*
Helioopsychidae (Selloopgyche borealis)
Case not like a snail's Shell*.. 3
5* Larvae smaller then 4 ml pro»* meso- and raetanotum
eaoh with a single* sclerotised shield embracing
the entire noturn*
Hydroptilidae
. . . • 'V •
* .
Larvae usually larger» either meso- or raetanotum or
both without solerites or with ecleretlzed shield
subdivided into seporfcto plates* fig* 11*«*••••••*•••••••••*4
4* Claws of hind legs very small* those of middle end
front legs large, fig* 2*
Molannidae (Molanna tryphena)
Claws of hind legs as long as those of middle legs*
fig* 3* »*.*«•8'
8. Antennae long* at least eight times as long as wide*
and arising at base of mandibles, fig* 4««*««
Leptoceridae
Antennae much shorter* not more than three or four
times as long as wide, often very inconspicuous*
and arising at various points, figs. 5 and 6**........6


-21-
6» Mesonotum eubmembrcmoue, except for a pair of
parenthesis-like# sclerotiaed bare# fig* 7*
Leptocerid&e
Itesonotum without such bars**
7* Meso- and metanotum entirely, membranous.
Phryganeidae (Banks!ola oonoatenata)
Mesonotum with some conspiouous sclerotised
plates*..••*......* •••••••3
8# Anal hooks formed* of 2 or 3 long teeth situated one
over another# fig* 8 A*
Sérloostematidae (Serloostoma orassleornia)
Anal hooka formed of a single large# tooth Kith
1 or more «malt teeth «a its dorsal edge, fig. 8 3******«*#*9
9. Pronotun with a deep furrow running almost the
full width of the solerite} head as in fig* 10*
Braohyeentridae
Larvae large# living, in a ease made of sticks#
' fig* 186. -â– 
Limnophilidae (Pyonopayohe sp* A)
10* Pro»# meso- end metanotum each with a single#
solerotizod shield embracing the entire aotum#
fig* 12i abdomen with many conspicuous branched
gills# fig* 12*
Hydropsyohidae
Either meso- or metanotum or both without spieritea
or with solerotised shield subdivided into separated
plates# fig* 11#*#*****##*#***♦***#*##*##•*#**##*♦********* 11
11* Labrum with anterior and lateral portions expanded
into a wide# membranous area# fig* IS*
Philopotaaidoe (Ohiaarra) â– - -
Labrum shorter# entirely sclerotised# fig* 14#
Psyohorayiidae

.ñ
1'*' . ‘ ■ /
FIG. 13 LAURUM
FIG. 14 LA3RUM
FIG. 12 LARVA OF
Hydropsyohe.
u - - - -»«■■ y-- - ■■»■--*
median ocellus.. , //
anterior wart , \ //
compound eye ■■jfr-tósHk
lateral ocellus R
postero-lateral wart 3*V.
pronotu m
legula.-'" G'scutum.''A
mesonotum| scutellum
l.posfnotum..^.:;-*-''^'>y\^^^¿„„„„OTm
metanotum
\\\ s 'â– tV ^
jVXJOA\ \'rf .,
s¡.
J*l*11Ttp‘‘‘~Tn iaimlt f* 2
s\ \ \ v>& \
N> \ ’ \X. ' . F
Cu,b Cu,-» m3.„ m2
&Lclaspers—Seni,alia
W'-aedeagusJ
maxillary palpus3
labial palpus.
] meso^xxIOs/F
^pleurae
metapleurae11 //
coxaj 1/
trochanter>',
Fol .daspers
tibia'' tarsus"*^’
“'apical spurs
1 I '"preapical spurs
FIG. 15 ADULT, ILLUSTRATING TERMINOLOGY OF PARTS.

â– 23-
Adults
1. Símil# hairy individuals not over 6 im long? meso»
scutellum with posterior portion forming a tri¬
angular, flat area with steep sides# fig* 16}
maeoscutum without warts*
Hydroptilidae
Size rango of 5 to 40 ami either mesesoutellum evenly
convex# without a triangular posterior portion set
off fcy sharp sides# figs* 17 and 18} mesosoutua
â–  with warts* * ..... ......... *2
2# Ocelli present* fig* 15.......... *«•.•*••.••••••.*.....•.•3
Ocelli absent# ........ * ....... *6
3* Maxillary palpi 3-segmented, fig* 19*
liiranophilldae (Fyonopsyohe sp* A)
Maxillary palpi 4» or ©«segmented****•»•♦•♦•.<••••«•*••♦♦••*••••4
4» Maxillary palpi 4-segraented, fig* 20*
Fhryganeidae (Banks!ola eoncatenate)
Maxillary palpi 5—segmented* fig* 21»*••*••.*••••••*•♦••••*«•*•6
5* Maxillary palpi with fifth segment two or three
tines as long as fourth, fig* 22*
Phllopotamldae (Chimarra)
Maxillary palpi with fifth segment not acre than
one end one*»third times as long as fourth# fig* 23,.«.......*6
6. Maxillary palpi with 6 or more segments, fig, 24...............7
Maxillary palpi with less than S segments***#***...........***10
7* Terminal segment of maxillary palpi mu oh longer than
preceding and with close* suture-like# cross striae#
which are not possessed by the other segments* figs*
28 and 26.......*#.#.***««.«..A#»*..*.*...*....8


Terminal segment ©f maxillary palpi without such
stria© and similar in general structure to fourth
segment* usually of same length»»10
3* Anterior tibiae with a proapieal spur»
Fayohomylidae
Anterior tibiae without a preapieal spur#.....*9
9* Meaooutum without warts* > fig# 17»
Hydropsychidae
Hesooutua with a pair of - small -warts* fig# 27»
Psychoayildae
10* Middle; tibiae without preapieal spurs and with a
row of black spines* fig* 28*,*••*»««,*.*•*««.#.#*•***«»**«11
Middle tibiae with preapieal spurs* with or with¬
out a row of -spines*»*..*.**#*»...♦.**.**».,,»»,..*•*******14
11* Mesonotum with sou tal warts represented by a long*
-irregular line of setate spots* fig* 291 antennae
always very long and slender*
â–  Leptoceridae / ' : -
Mesonotum «4th scutal warts either small* fig# SO#
or absent* fig# 18| antennae fairly short***##*..**.*#*«***12
12* Male genitalia as in fig* Sit ventral aspect of
abdomen of female as in fig# St#
• Helicopsyohidae (Helloopsyche borealis)■■
Oenltalia of male and female not as in fig»# 31
and 32..........................................,,#.,*,.*.*13
IS. Mesonotum with a deep* antero-mesal fissure with
scutal warts as in fig* S3.
Seri costomatidae (Serlcostoma cragsicomis)
Mesonotura with only a shallow antero-mesal crease*
with scutal-warts as in fig*-54#- - - - - - ~- - - - â– 
Brachyoentridoe


-27-
14# Middle femora each with a row of 6-10 black apiñes
or entero-central face, fig* 35*
Molannidne (Molanna tryohena)
Middle femora each with none to 2 black spines on
antero-centrsl face*
Eraehycontridn*

>
FIG» 56—C.
atarrima, APEX
OF FRONS.
FIG. 37—0.
florida, APEX
OF FRONS.
FIG. 38 C.
perigua, AFEX
OF FRONS.
A B
FIG. 39- 0.
perigua, MaITe
GENITALIA. A,
CLASFER, CAUDAL
VIEW; B, AEDEAGUS.
FIG. 40---Chimarra aterrima, MALE
GENITALIA.' A, LAlEMli" VI EVf; B,
GLASPERS.
FIG. 41—Chimarra florida, FIG. 42——Chimarra
MALE GENITALIA. aterrima, FEMAIS
GENITALIA.

-29-
OHBtARRA St options
Key To The Species Of Chlnarra
Larvae
1. Apex of frons as in fig* 36.
â– â–  ChlBarra aterrima
'i V
2» Apex of irons ns in fig* 37.
■ • . ' Chliaarra florida
3* Apex of fresas as in fig* 30*
â–  Chirarra perilla
Adults
1* Apex of abdomen with a pair of claapers (ttalas}**..*****.#*«:.*2
Apex of abdomen without claapers (females) .***4
3* Color brown, genitalia as in fig* 39*
Chlmrra perlera
Color blaokish* 3
3* Genitalia as in fig* 40.
" Chimarra:aterrimaâ– 
Genitalia as in fig* 41*
Chlnarra florida
4* Coico* brean
Chliaarra perlgua
Color blaokish** •••••5
5* Ninth tergite produced into well-defined* lateral,
solorotised ears, fig* 42*
Chlmarra florida
Ninth tergite without oars*
Chliaarra aterrima

Chistar re. aterrlma Hagen
Range» Eastern end southern states*
Description» Ross, 1944.
Reatarles» The larras of this species lire in small, cool, fast-
flowing, spring-fed creeks* Although 1 found a stream in Golden
Head State Park where C* sterriroa was the dominant caddis fly, it is
not usual for the apeóles to be so abundant. The fact that 1 hare
nerer collected adults at light suggests that they may be negatirely
phototropio.
Prerious Florida Records» loss (1944) records this species from
Florida but does not gire the locality. Rotten (personal ccrawuni-.
cation) collected 0. aterrima in the riolnity of Lake Placid.
Specimens Examined» Fire larrae, 2^ miles W* of Gainesrille, L. B.,
l/28/39j 3 adults, 2! miles W. of Gainesrille, L» B., 1/28/391 1
larra, Alachua Co*, ll/d/z7i 40 larrae, .Bogtown Greek, in leaf detritus,
F, S, Y., 12/31/47» 1 larra, Deril's Millhopper, 10/25/3?» 1 larra,
Peril's Millhopper, IO/2/SI1 400 larrae and pupae, Golden Head
State Park, !4. W# H., 8/L2/50» 40 larrae, Hogtoen Creek, H. 17. 1.,
8/8/SO1 38 larrae, Hogtown Creek, M. W. H., 6/4/51.

31-
Chimarra -porlgua Ross
Range» Florida, Georgia and Illinois*
Description* Roas#, 3,948* ,
Remarks» Ross (1948) states that C* perlgua was recorded erron-
> • , ' i
cously as C. socla# and the latter species was thus listed as being
4 .• !
present in Florida* Chlmarra soola does not occur in the state*
í . ‘
According to Ross (1948)# "Re-examination of a large series of
soola—*——(indicates) a wide eastern and northeastern range for
socla end a small range of perlgua which is peripheral on the southern
and western portions of the range of soola**
The larvae of C# perlgua live in olear# oold# rushing water
such as that pouring over the rook dam at Poe Springs* The adults
appear to he negatively phototroplo, for though I "lighted* for
caddis flies many times in this locality# 1 did not collect any
adults*
The larva was associated with the adult hy a process of
elimination* I collected larvae of three distinct species of
Ghimarra and adults of three species* C* aterrima# C* perlgua and
C* florida* The larva Of C, aterrima was previously described and
was easily identified* I associated the larva of c* florida by
pupal dissection# Which left one unidentified species* The adults
of C* perlgua are very similar to C* soola* and the third larva
was also extremely similar to that of C. soola. It seems obvious#

•32*
therefore# that those larvec must bo those of C* porlgua*
The larva of C. perlgua cannot bo distinguished from that
of C, soola using the description given by Eoss in 1944. I do not
have comparative material of £» soola and# therefore# cannot look
for differences in morphological characters between the two species#
but a critical examination of both larva© is definitely in order*
Previous Florida Be cord at "llolotype# male—«—Santa Pe Elver#
Alachua Go.# Florida# April 6# 1940, 1». Berner. Allotype# female*-*
Santa Pe Elver at Poe Springs, Alaohua Co*# Florida# March 12# 1938#
L. Berner. Paratypos-——Florida* same data as for allotype# 1
male,” (Eoss# 1948)
Specimens Examined* Two adults# Poe Springs# I». B«* zflz/%%% 3
larvae# 6 miles W# of Gainesville# B# A. B*# ll/e/39; 1 larva# 5
miles W* of Gainesville, B. A, B„# 12A/59* 1 larva# Alaohua Co*#
I*. B#, 2A/S9* 4 larvae# Poe Springs# s/l2/ZBi 10 pupae# Santa
Pe Elver# J* S, R., sAs/SSj S larvae, Poe Springs#' 11* W# H*#
3/24/sOf 2 larvae# Poe Springe# M, W# l?*# b/Ib/S0i 14 larvae# .1
mil© south of Gulf Hammock, M# W, H## 7/so/so*

-S3 A.
â–  Ohimarra florida Rots. .. .
Range* Florida and Georgia.
Description* Ross# 1944#
Remarks* . She undescrihed larva of C, florida was associated with
its adult form hy means of pupal dissection.
She larval stage of this speoies «as only found in the most
rapid portions of vary swift# oaloareous streams#
Previous 'Florida. Records* • One male# Freeport# Walton Count;-, < collected
hy Dr* Lewis Berner, April 8# 1958* (from Ross, 1944)
Specimens Examined* Three males# fast stream# .1 mile S« of Gulf
Hammock, Levy Co*# sweeping# M* W, H,, ?/so/80f 1 pupa, *1 mile S*
of Gulf Ham-nock, Levy Co., II* W* I*# 7/8O/6O1 9 larvae# Poe Springs#
10/t4/47| 1 female# Poe Springs* at light# M, W, H*# 9/9/60; 1 pupa*
Pot Springs* M* w, H#, 3/24/51; 1 pupa* no data; 1 male, sweeping* Poe
Springs, M. W, H., 3/24/SO* 2 males* 8 females, Poe Springs* at light#
M. W* H„ 7A2/S0. ' " '


34-
PSYCHOMniDAE
Key to the PaychoEglldaa
Larvae r
1* Anal hooks tilth a row of 4 or 6 long teeth along
inner ventral margin, fig* 44s tenth segment short*
with scarcely May ventral margin» fig* 46.
Paychoapia fiavida
Anal hooks with at most very short* inner teeth, fig*
46s tenth aegra^t longer end tubular, figs* 47 and
43***•**••«••••#••••*»*•#•*••••••**•••*••*•••••••••*«*••••••2
2# Mandibles short end triangular* eaeh with a large#
thiok brush on the mesal side, fig* 49*
Phyloeentropas placidua
Mandibles longer, fig* 50» tilth only a thin brush
on left mandible# nono on right* ***** *••«*#.*#«*.* »***>*••*•• *3
3* Right mandible with two large dorsal teeth which
completely overhang and hid# the ventral row* on
the left mandible the dorsal row of teeth over¬
hangs end hides the ventral row# fig* 61*
Cymelius marginalia
* ! ! '
Right mandible with a single dorsal tooth which only
partially hides the ventral row of teethf on the >
/ left mandible the dorsal row of teeth does not hide
the ventral row, figs* 60 and 63..*.»,.......*..............4
4* Basal segment of anal appendages (tenth segment)
without hair, fig* 47* left mandible with basal
tooth small and with a linear brush on mesa!
face near base, fig# 47.
Keureclipais crepusoularls â– 
Basal segment of enal appendages (tenth segment)
with long hair, fig* 48» left mandible with basal

-FROTT GILO
UTIL OS
FIO. 53
FIG^ 55 FROFT A1*D
i tiro vraiGS
FIG. 5G FLOUT ATD
;rIl,D .L1.G3
FIG. 57 MAXILLARY
PALPOS
FIG. 58 I1AXILLARY
PALPUS
FIG. 59—MAXILLARY
PALPUS
FIG. 30—:'AXILLARY
PALPUS
FIG. 62—OILGS
FIG. 61 HIND WILG

•36-
tooth large $ subequal to one .above and with
brush small* fig* 62*
, Polyoentfopus
Adults
1* Front tibiae with a preapioal spur******•*•••«•*..••••..••*•••2
Front tibiae without a preapioal spur*•••••*•••••••• ••••••••«*6
2* Both pairs of wings with Eg present and branahing
fro®Rg at radial crossvein* fig* 63*
Phylocentropus plaoidus
Both pairs of wings with R2 either absent or
branching from Rg near margin of wing* fig* f>4............ .,3
3* Hind wings with IS 5-brsnched* fig* 56*
Keureolipsis orepusoularls
Hind wings with M 2-branehed* fig* 56****«»••••••»••••••••••••4
4.
Front or hind wings* or both* with Rg present*
fig. 66.
Pelyeentropue
Both wings with Rg absent* fig* ;06******»***#*»*.**«.*.**.*.**6
5* Maxillary palpi with second segment long* third
only slightly longer than second* fifth short*
fig. 67* •
Cyrnellus marginalia
Maxillary palpi with second se groen t short* third
three times as long as second* fifth long* fig*
68.
Kyotlophylax vestltus
6* Maxillary palpi with second segment only one*
half to one-third as long as third segment
and with the apex enlarged into a small cushion*
fig* 69* '
Carnot in a

-37.
Maxillary palpi with second segment as long as
third and uniformly cylindrical# fig* GO..*. ••••«•••«••?
7# Sind ninge with apex eirenly rounded# fig* 61*
Lypo dirersa
â– 4mm mmmmmmw
Sind â– wings with apex tapering and somewhat
pointed#â–  f i g* 62*
' PgyohOBgia flarlda
SBURBCfJPSIS Mohachlan
Heureolipsis crepusouloris (Walker)
This species has not bo go, taken in Alachua County# hut
Batten (personal communication) collected it in Florida (location
not giren)*

«•38*
PHYLOCENTROHJS Banka
Phylooentyopus placídus (Banka)
(Holooentropua placldua Banka* 1906b
PtyioQmtrowxa maxima 'fforhlea* 1909)
Ranget Eaatern* central «nd s outturn at ate a

-39
Description* Ross# 1944*
Remarks» This apeóles appears to fco rather rare in Alachua County#
for I was able to obtain only a few specimens* One larva was found
in a lake# the other in a ¿low stream with a soft# fleoculent bottom*
Torhlea (1909# p* 712) gires an excellent description of the unusual
larval ease*
"The larva burrows into the sand of the bottom of the
stream and forms a long# often complexly branched case of sand,
scarcely firm enough to beep its shape when it is extracted from
its position* This case, about 5 mm. in diameter# excepting some
smaller branches# is often as muoh as 65 ram* In length# the greater
portion burled# only 10-20 ram. projecting from the stream bed*"
Previous Florida Records» Dead Lake# Wewahitohba# collected by
J* G* Heedhan (Betten» 1934, p* 218)| specimens collected by Betten
(personal communication)# location not giren*
Specimens Examined* One female, Washington Co»# 1* B«, s/so/lOi
1 larva, Lake Santa Pe# 12/2/47» 1 larva, slow stream# 6 miles H*
W* of Gains swill e on U. S. 441, M* W. K.# 6/21/50» 1 male# Worth¬
ington Springs, at light# M, W. K„ t/iz/m.

*40*
â– U

*41*
POLYCEl'ITROPUS Curtis
. â–  Key To The Spoeies Of Polygontroous
Larvae
!• Spots on larval head as in fig* 68*
Polyoentropus sp* A
larval stage unknown*
' Polyoentropus orassioornls
Adults
1* Genitalia of wales as in fig* 70j genitalia of females
as in fig* 71.
Polycontropus orassioornls
V
Polyoentropus orassioornls Walter
Tli© larva of this species, although it has a widespread
â– *
range throughout eastern United States end Canada, is unknown* Banks
(1907) records P* crassicornlo from «Jacksonville! however, I have no
other record of its occurrence in Florida* Possibly Polyoentropus
sp* A is the ««described larva of P* orassioornls» ■
Polyoentropus sp* A
Although I collected the larvae of this speoies in a num¬
ber of locations, a constant search for the adults failed to produce

42-
a singlo specimen# The adults my bo negatively phototropio, for
night lighting and daylight sweeping were both Ineffective in col¬
lecting them# X attempted to rear some larva© which I collected
in Big Hatchet Creek, but the larvae died before pupation*
these larvae appear to be very similar to the larvae of P*
rento tus (fig* 68)* Sino© P* remotes has been collected to British
Columbia, Illinois, Minnesota, Hew Hampshire and Bew York, a wide¬
spread geographic distribution coupled with a looal occurrence i*
indicated# It is possible that the larvae of Polyeentropus sp# A
are the undeacrlbed larvae of P* crassloorala* which has previously
been collected to Florida, but it is also possible that they ar©
toe larvae of P# remotos* Further study will undoubtedly permit
association of larva and adult*
Specimens Sxamtoedt One larva, Poe Springs, larva, Santa Fo River, 5/24/37* 1 larva, Big Hatchet Creek, S* W*
H*, 4/24/SOf 2 larvae, stream, #1 mile S* of Gulf Hammock, Levy
Co., It. W. II*, 7/30/60i 3 larvae, Big Hatchet Creek, H* W* N,,
b/10/S0} 1 pupa, ravine stream, Golden Head State Park, Clay Co*,
M. W. H„ sAb/bO.

mrCTIOPHYLAT Brauer
Only two Hearctlc species are known for this genus, one
of then from Florida. The larvae have not yet bean associated with
the adults.
Kyotlophylax veatltua (Hagen)
(Polycentrorna vestítus Hagen, 1861
Polyoentropus afflnle Bonks# 189?
. iiW8t 1 onhslaaT inoestue Banks# 1911
hyotiopi^l'ajc ve'stl'tus (Hagen), Sibley, 1926)
Range» Eastern, central and southern states*
Description» Ross, 1944*

48-
Remafks? fhe larva of B* vestitus is unlmowi*! however, if found,
the larva might very well turn out to be one of those which are now
classified under Psyohomylid Genus A* Further study should clarify
this interesting enigma»
Wyctlophylax vestitus is rare in Alachua County, having
been collected only at Poe Springs.
Previous Florida Reoords* Shis species has not prevously been re¬
ported from Florida. Betten (personal communication), however, col¬
lected it in tho state (locality not cited)#
Specimens Examined* One male, Blue Springs Creek, Jackson Co», L.
B», 6/5/4D1 1 female, Poe Springs, at light, H* W» H», B/lz/SOi 1
male, Poe Springe, at light, M. ¥, B», 9/b/fiO»
CEKROTIM Ross
Bo larvae of this genus have been discovered, and no morph¬
ological differences have been found to distinguish between the fe¬
males of the various species, the genital segments of which fora a
conical structure with only simple parts as shown in fig» 77» I
have examined specimens of the following females, tdiich cannot be
identified specifically since they were not collected together with
males* 1 female, Gainesville, at light, M. W. !?», b/s/BOi 1 female,
Big Hatchet Creek, at light, M. ¥, K», 6/12/SOI 8 females, Worth-


•45*
Ingten Springs! at light# If* V/* IT,, 7As/S0i 2 females# St* John*a
River, Putnam Co*, at light# 7/l8/S0i 60 females, Poe Springs, at
light, M# W* H#, 9/d/S0t 25 fontales, Poe Springs, at light# M# W*
!U, 8A8/60*
‘ Koy go flho Species Of Cernotlna
Larvae———Unkn am
Adults
1* Males
Females, fig# 77———Hot keyed.
2* Corel with 3 or 4 long# black tooth on the raeeal
side near the base, fig* 78*
Cernotlna calcea
Corel without long, black mesal teeth*#**•***•»*••••***•*«•••*3
3* Genitalia as In fig* 79,
Cernotlna sploata
Genitalia as In fig* 80*
Cernotlna trunooaa

•46»
Cgrnotlna cnlcea 'loss
Ranee: The range of this species io 7rt.de» hut C* caloea appears
to be looally distributed* It has be® tafee» in Illinois» Missouri*
Oklahoma* Texas and Florida*
Description* Ross* 1944*
Remarks: I collected adults at Poe Springs and Worthington Springs*
but continued Investigation within wading distance of the shore
yielded no larvae« The larvae may possibly be found In the deeper
parts of the Santa Fe River, beyond wading depth*
Emergenoe, moat likely, tafees place over the entire sissuer*
Previous Florida Records: Ross (1944) lists this species as occur¬
ring in Florida but does not epeeify the locality* Betten (personal
ooramuntoaticn) reports taking C* oalcea in the vicinity of Lake
Placid*
Specimens Examined: Twelve males and females* Poe Springs, at
light, M* W* i?., 7/42/50* 100 males and females* Worthington Springs,
at light* 11# W* N«, 7/45/50* 20 mies and females, Poe Springs, at
light* H* W* I** 9^/sO* SO melee and femalee, Poe Springs, at light*
M* W. H„ 7/s/Sl.

•47»
CafWotSrm trtaieaaa Tioet
Ben ¡ge* tm®m cnly from florida*
Bjworlutlqp» Boos# 1047#
Rtmarbe* I eolloatod this epooioo m only two oreaelctie* both tires
m the shores of th® Hirer Sips* To ss®? 1»ostoigo# the typo «torio!
plot ts$ enoeirenc - or® th§ «sly eollootlcno of C* tronoooo esteasi#
Sino# a rndbor of iiKliridurle mr® found# It appears
that the larree am bo found in the rtrer i hcrerer# a ©ooroh
withlr, reding dl &%mw of the share foiled to roreol iheet# 1 1»»
liare the ©roblen Is siwilsr to that of £* M%mrs^******®'W lofreo
or© to bo found In the dcopor «atara# beyond reding depth#
There ere no rerphelosleal oharcotitr* tshiah e«n bo used t©
differentiate between the feast os of fhla genre# Altlioagh 1- eel*
looted. the feanloc of C* tyuneona which hod not' bren col looted
protricusly# no norpholonics.l chare, ©tere «oro found which eould
h© used to distinguish thorn from other «pioles In the ganre*
Pyqrfore. florid© Records* Hales* Daytona Bench# tolreia Co#* rel*
looted ^ BldSel* Beleeendre and Rett* C/27/4S* (Ross* 1047a)
Sreoiaan» .gmotoadt Tito solos* S3 femlos# Hirer Styx# at light*
It# W* »** 5 ítalos* i fósele©* Eire? Sty*# at ll^t* II* t.
!?•* o/sAU

•*48*.
Cernctlne gpioata Roes .
Rea ge* raicé* .Michigan, Oklahoma, end Florida* Ccraotlna epi cata
has not previously been reported from Florida, It is obvious
though, from its spotty distribution, that £, spioata is far more
widely distributed than has been thought* and most probably occurs
•'r' »
in certain areas of the intervening territory.
Description* Ross, 1944,
Remarks* Ceraotina spioata is uncommon in Alachua County, for I
collected only four specimens at two widely separated points* Most
likely these individuals had flown some distance, fwo of the adults
were taken near Gainesville, the others on the banka of the Santa
Fe River,
Previous Florida Records* Rone,
Specimens Examined* One male, Blue Springs Creek, Jackson Co*, Xi,
B., 6/5/40í 1 male, one female, Pona B, at light, M, W, H„ 7/LO/«>í
3 males, Poe Springe* at light, M, W, E,, 9/^/50,

•»49««
Payehogjylid Gsnn s A
Benge* Illinois* Florida, Michigan end ITieoensln*1,;. , .. .
■ Description! Ross* 1944»
Remarks* The adult of this curious larva is unknown, but it is
probably one of those new,known, only, fresa the ,adult stage#, such,,....
as Ifyotlophylax or Cenaotina* In Alachua County the larva (fig*
01) lms only been collected in the Santa Fa River and Big Hatchet
Creek*
Previous Florida Recordst Ross (1944) records this larva from
Florida but does not give the locality*
Specimens Bx&mined* Two larvae, Santa Pe River, 8/24/57? 2 larvae,
Alachua Co,, l/lb/51; 5 larvae, Big Hatchet Creek, M* W, N«, 4/29/60?
1 larva, Big Hatchet Greek, M, W. K*, o/io/si? % larva©, Big Hatohet

Creek# H* W. II.# 4/29/60.
LTPE MoLaohltm
Lype diversa (Banks)
Hoes (1944) records this species a® occurring in Florida
(locality not given)# bat I have not collected it in Alachua County.
Perhaps it occurs in northwestern Florida.

PSYCHOMYIA Piotet
Of the three described Horth American species, only P.
flavida has been taken in Florida.

«*53*
Peyohoada flavlda Hagen
(Peychomyta flavlda Hagen# 1861
. . Psyohomffia nuVoheTla Banka* 1899 â– '
Peyohonsrla moesta Bank^ 1907aj
Range* Eastern# central and southern states* Sine© P* flavlda
has not deviously been taken in Florida* and Kentucky ,> Tennessee
and Worth Carolina were the nearest reported collecting sites*
finding this species in Florida indicates that it occurs in the in¬
tervening territory*
Pesorintion* Ross* 1944*
Remarks* Perhaps northwestern Florida is the only area in the state
this species oaa be found* for* although I collected no specimens in
Alachua County* I examined one specimen which was collected in the
vicinity of Marianna#
According to Ross (1944)* the larvae are restricted to
swift* cold streams*
Previous Florida Reoerds* Rene*
Specimens Examined* One female# Blue Springs Creek* Jackson Co»# L*
B«* e/s/40*


-54-
CTIWBtUJS Besiks
Cymellua marginalia (Bonks)
(Hyotiophylax marginalia Bonks* 1905b
.. Cvrnellus sernylMoselv. 1954)
Rango» Control and southeastern states) also known froto near the
mouth of the Amasan Riser in South Araerioa* In the Southeast» C*
marginal 1 e has been reported frisa Tennessee» Kentuoky end Alabama
but not from Florida# Although I hare not verified the correct¬
ness of the identification of the specimens from South America»
finding this caddis fly in Florida is an indication that it must
occur in at least limited portions of the intervening territory,
namely Mexico and Central America*
Description* Ross, 1944*
Remarks* Only one Horth American species, marginalia. is known for
this genus. The Immature stages have never been discovered, but
evidence presented here, I believe, will clarify the association of
the various stages* I found two species of larvae Which were keyed
(Ross, 1944) to Paychetayiid Genus A} one of Which is to.be found
under Psyohonyild Genus A in this thesis, and one of Which was un-
desoribed* I collected an immature pupa, however, in hake Wauborg
which contained the larval solerites of the undescribed species of
Psyehomyiid Genus A# The fact that these solerites could be identi¬
fied as Psychomyiid Genus A would eliminate Fhyloeentropus, Reure-

elipsis and Polyeentropus* The pupa had a pre&plcal spur on the
front tibia which eliminated Gemot ina, Lype end Payohomyia; leav-
ing only Gyrnellus end Hyctiophylar* The maxillary palpi* the key
character for distinguishing between the two genera» are identical
with those of Cyme Hub» so 1 feel the association is a valid one»
It appears, therefore* that tho Lake Wauberg specimen is the pupa
of C* marginalia while the second species still remains without a
nan© in Psyohomylld Genus A*
This species is not very abundant in Alachua County, %
records indicate that emergence occurs in the Spring*
Ross (1944) states that the adults are mainly found cm
the shores of large rivers but also on the ehores of small streams
and lakes* Although I collected adults from rivers and streams»
I found larvae only in lakes*
Previous Florida Records* Hone*
Specimens Examined* Three larvae* Kingsley Lake, Clay Co*, bottom
sample 10 feet deep, silt over sand, J* S* R* and A* P* C*, 5/l8/sSt
1 larva. Lake Santa Re# bottom sample 4-5 feet deep, 12/2/47; 2
larvae, Johnson Lake, Golden Head State Park* Clay Co*» II* W* I*»
s/Ís/bOj 1 larva* Leyy Co*, £. L., 6/18/60; 7 females, Prairie
Creek, at light, fit# Tí* R*, 8/29/61; 1 male, 1 female* Worthington
Springs, at light* IÍ* W* R*, 7As/S0; 2 females, St* John’s River,
Putnam Co*, at light, 7/L5/&Ol 19 males and femles, River Styx, at


•67-
llght, M*1f. H.» 7/47/50í 1 pupa, .Lake Wauberg. M. W, ir, , 0,48/60*
HIBROPSIfOHIME
Ross (1944, p* 70) characterises the family Hydropsychidae
•very troll tfhen ho says, ’’His larvae of all genera are remarkably
uniform in habits end appearance* They are wormllke, active and
pugnacious, and poseessrows of bushy abdominal gills* fhey pre-
for the more rapid locations in streams, .usually being concentrated
around riffles, spllleays and rapids,, although they may also be .
found wherever there is an appreciable current* fhey make a re-
treat trader and ab^fc trash, logs, stones and any other haven.
In front of this, retreat they build-a net which is reputed to
strain food /roa the fleeing eater* For pupation they spin an
J£ ' r., ••
ovoid cocoon near the retreat, generally using sand, stones and
bits' of trash*. . •
Key fo fhe Hydropsyohldoa
â–  * .
. * ii '
Larvae
1* Head with a Sharp, U-shaped ridge, fig* 87*
Maoronenam Carolina
Head without a sharp, U-shaped ridge, fig* 10S****************2
2* Stridulator of fore coat not forked, fig* 88* '
*. Dlplectrona modesta
Stridulator of fore cora forked, fig* 89 **•* *•• •» ******* ***'**»S


3* Prosteraal plats with a pair of posterior ecleritos,
fig* 90.
Hydropsyohe
Prosteraal plate without a pair of posterior selerites*
. fig» 91.
Adulta
1# large, striking caddis fly with pictured «inga sad
•very long antennae, fig. 92. "
Haoreneaiun parolina
Caddis fly not as in fig* 92*.
2* Hind wings with apex round and with So end R^ bored
deeply at/.apex#-fig* 93*
Dlplectrona siesta
Hind wings either with So and R^ not markedly bowed,
or both wings with apioal margin ineised**....3
3* Hales*....4
Females*
4* Baso of aedeagus cylindrical, fig# 94*
Hydropsyohe incomoda
Base of aedeagus bulbous, fig. 95.
Cheamatopayche
6. Stoma! plates of" eighth segment separated to base
of segment, fig* 96*
Cheumatopsyche - *
Sternal plates of eighth segment separated only
two»thirds distance to baso of segment, fig. 97,
Hydropsyohe Incomoda.

•60»
DIPLSSTRGHA Westwood
Dlplectrona agosta Sanies
Bangsi Bastera and southern states#
Desorlptloni Boss* 1944*
Bemarka* Dlplectrona modesta does not occur to Alachua Count? and
most probably is limited to the northwestern part of Florida.
Freíd cus florida Records* Boss (1944) lists this species as occur*
ring.to Florida but does not cite the locality* .
Specimens BragAned* â–  Six larvae# forreya State Park*, .liberty Co.*
6/7/3S! 1 torva* Cresteiew, Okaloosa Co»* 1* B** 12/12/37*

\
FIG. 99 Hydropsyche
incomoda.
FIG. 101 Hydropsyche
incommode, APEX OF
AEDEAGUST A, LATERAL
ASPECT; 3, VENTRAL ASPECT
FIG. 1 (X—-Hydropsyche
in com ola, HEAD OF IÃœI-
DESCP.I32D LARVA.
FIG. 102 Hydropsyche
incomoda, FEMALE
GENITALÍA.

-62-
HEDROPSYCHE Pictet
She genua Ifodropsyche contains truer fifty speciee of caddie
flies, but only one, Hydropsyche incomoda, is known to occur in
Alachua County. The larvae prefer clean, fast-flowing, cool water •
Key So She Species Of Hydropsyoho â– 
Larvae •
1» Head as in fig* 100*.
’ ■ Bydropsyohe incomoda
Head as in fig, 100A*
Hydropsyche ep* C
Adults '
1. Genitalia of rales as in fig* 101s genitalia of
females as in fig, .102*
Hydropsy ohe incomoda
Hydropsyche lncomnoda Hagen
Range* Recorded from Florida, Georgia* North Carolina and Illinois,
Description* Ross, 1944,
Remarks* She immense numbers of caddis fly larvae swarming over
the rocks and leaves of eel grass (Valllsnerla sp.) at the rook dam
near Poe Springs is certainly striking; For sheer numbers, there
is no other place in Alaohua County which even approximates it.

&m%$ oMeeriest mt&r metes cmr the rest <3s» «ttb ©«¡slfcAer&lsílé
«peed* cn ©loatst of tTm fcoMhst*.
the terree build rotroats In end armad truth and Mts of
dehrte tooth on feotes «end ©el gross* la ¿ron* ef fht# tefenmt ftef
Mílñ © set te «trote feed fresa the Aeetns mt«f# tout f«r pi$wtt®
ttej1 spin t» we!4 eoeoo»*
I eesodtfttüd the preralouely «nteetes ian&ture stages oith
tteif adnlt fes» % mr&ngt fcmtver» es the terree wm> wr$ wm*
netoailstle m& hed o toa ©aeie§i©«'l 'teitaraiQo» I ehtaiited test
resolte taking «sell gtmpa et prpoe mñ f«etefeg fhea la. úmo
eottem. Smmfatm odtíltg «aorged frost «ear S90 pepee# «a.d o defln*
Ite ©ssoaiatle® «ne mñ® ihrau$i .«toady of the temí aoloritoe*
isfcleh mtw poteed inte ite posterior porti» of tías meeW popel
©ase*
A targe matter of líame «sd p»pb« died ohfte tetes trisas»
ported la mter# teetoteg them te «t «es» «#*4 ©egetetiim for trws»
perfeotton «« % far tt» test pr©©eá*ír© o sed#
B&m ©addle ftlos ontÉl «xtáeimtsr to tey their eggs* fe’
«teay fhe offset .of the «atoswaenee m fsáelts# I tóete Ileo* eitát
ejootewts of ||* teeaaaoflo ead pleoed the» itsdemter# She hairs-«a
the fcoty and *tnge proponte! thorn fría getting «ot# «5 the ©ftMio
file®# m relente# tetteé te tfce sorfeee# eesily teet© threogt the

•64-
surface film and flew away* Complete submergence did not seem to
affect them In the least»
Previous Florida Records* Ross (1944) lists this species as occur**
ring in Florida hut does not specify the locality»
Specimens Examined* Two males# 6 females# favores# Lake Co», F»
I» ¥», s/83/36* 7 larvae, Sweetwater Creek, Liberty Co», L» 6»,
ISAo/37* 2 larvae, Shoal River, Okaloosa Co»# L, B», 12Al/37*
16 larvae, Santa Fe River, J. S. R», 3A2/88* 24 larvae, Hillsboro
River, L. J. M», 2A8/S9* 1 female, Santa Fe River, L* B», 3A/*0l
1 male, Santa Fe River, 4/6/40* 3 females, Blue Springs Crook, Jacks*
son Co», L» B», 6/6/40* 11 larvae, Poe Springs, I0A4/4?; 54 larvae,
6 pupae, Poe Springs, M. If, H», 3/24/§0f 3 females, Big Hatchet
Crook, at light, M. W, H», gAs/bo * 1 male# 4 females# Hogtcwn
Creek, at light# M* W. IÍ», eA^/SOj 104 larvae, Poe Springs, M. W.
R», 6/21/60* 1 male, Poe Springs, sweeping, II* TT* R«, 6/21/50* 275
males and females, Poe Springs, at light# II» TT# R», 7/12/50* 12S
larvae end pupae, stream »1 mile S» of Gulf Hammock, Levy Co»,
II* Tí» K*# 7/30/50* 9 males, 23 females, Poe Springe* at light, M#
W# H», 9/9/sOj approx» 2000 larvae and pupae, Poe Springs, M* W»
N», 6/6/50* 176 males «aid females, Poe Springs# at light, K. T»\ H»,
7/3/51.

«6 5»
Hydropsyoho sp* A
FIS* 10QA«—»«Jlyaropayche sp* A*
H3AD OF LñWA.
The larva (fig* 100A) of this spoolos is undosaribod, sad I
have pot-yet been able to associate it •pith an adult forras indeed no
adulta of the genus Hydropeyohe, other than H* inoommoda, have been
collected in the state* I have not found those larvae in Alachua
County*
Specimens Examined* two larvae* Jasper, Hamilton Go*, L« B*,
g/é/sSj 2 larvae. Lament, Madison Co*, L* S*, 2/6^81 1 larva,
Drifton, Jefferson Co*, L* B*, 2/6/38s 4 larvae. Golden Head State
Park, Clay Co*, M* W. N*, 8As/®0*

-6G-
CH3Ü11&T0PSYCS5 Wallengren
Most of the small* brown Trichoptora larvae with bushy ab*
doralrtal gills belong to this genus* Both larvae and adults are
•very similar in appearance and behavior to the larvae and adults of
Hydropsyohe.
According to William Book* dr* of the florida State Board
of Health (personal communication)* Cheumatopsychid larvae will
tolerate pollution of a minor degree while most other caddis flies
will not. Personal observation hears this cut; for example, In the
summer of 1950 the East Branoh of Hogtown Creek had a caddis fly
fauna composed almost entirely of Cheumatopsychid larvae* while the
West Branch had a more varied fauna. Many old, empty Leptooella
and Oeoetis cases were found in the Bast Branch showing that other
species had lived there* Since it is known feat the Bast Branoh
occasionally becomes polluted, it would seem likely that the creek
was slightly polluted in the summer of 1960, and that only the
Cheumatopsychid larvae could withstand the pollution*
1 collected many larvae in Alachua County* for this genus
is abundantly represented. Bo structural characters have yet been
found to separate the larvae to species? therefore, these records
will not be given. Larvae of Cheumatopsyehe are vary common and
occur in most of the streams in the county*


-63-
Bey tú The Species Of Cheumatopayohe
Larvae-ot Keyed* See fig. IOS.
Adults '
1* (Senitalic, of moles as in fig. 104f genitalia of fe¬
males as in fig* 103. '*â– '
Cheumatopsyohe burles!
2* Genitalia of males as in fig* 105$ genitalia of fe¬
males as is fig* 108.
Chouaatopsyche analia
$* Genitalia of males as in fig* 106$ genitalia of fe¬
móles as in fig* 110.
C heurnat op ay ohe aphanta
4* Genitalia of males as in fig* 107.
• Cheumatopsyohe pinaea
- Ghourmtopsyohe burial Ross
* > .
Range» Florida and Illinois* Undoubtedly further collecting will
fill in the gaps In our knowledge regarding the distribution of this
species*
Deaorlptlon» loss» 1844*
Remarks» I associated the previously und®scribed larva with its
adult form % rearing pupae in damp ootton* Ko morphological char¬
acters were found, however* te distinguish this speolos from the
others in the genus*

A B
FIG. 108- Cheumatopsyche burksi, FEMALE
GENITALIA. A, LATERAL ASPECT OF NINTH AfiD
TENTH TERGITES; B, DORSAL ASPECT.
FIG. 109 C. FIG. 110 0.
analis, FEMALE aphanta, FEMALE
GENITALIA» 'GENITALIA.

-70-
íhoae larvae occur in immense numbers on leaves of eel grass
(Vallisneria sp*) and rooks at the dam la the Santa Re River near
Poe Springs* The larvae are very similar to Hydropsy ohid larvae
and are commonly found associated, with them* * the larval nets of
both Cheumtopsyehe burksl and Hydropsy che incommoda are;'., easily ¿and ■
often soon on the submerged rocks and vegetation*
Previous Florida Recordat Rose (1944) rooords this species from
favarae* .'’■■■
Speoimens Examined* ■ Since oxtrardinary numbers of larva© can be
found at Poe Springs any tino of the year, I ttí.11 omit sy records as
they are not indicative of the numbers present* One male# Wlaka,
Putnam Co*, R* B*» s/e/dGj 260 males and'females#.Foe.Springs# at
li$it# II* if* II,, 7/12/50j 1 ralo, S females, Worthington Springs# at
light, II* w. 11., 7/Í3/50i 27 females# River Styx, at light, II* W* I*,
7/27/501 3 males# 23 females, Poe Springs, at light, H* W, 17,#
9/9/5O1 230 males and females# Poe Springs, at light, H, W* I,#
7/s/si.
Choumatopsyohe snails (Banks)
(Hydropsyoho anells Banks, 1903b
. . liydropsycih'e Banks, 1909b)'
Range* Widespread, occurring over most of the United States with
the exception of the Southwest*

-71
Description* Ross, 1944,
Remarks* Cheumatopsyohe analla is a rare caddis fly In Alachua
County. According to Ross (1944) the larvae Inhabit small streams
but also occur in larger rivers* They have a wide ecological toler¬
ance and are often found in streams carrying considerable pollution*
Previous Florida Records* this spooles has not previously been
taken in Florida*
Specimens Examined* One male# Ebro, Washington Co*, t* B*, S/S0/40i
1 male, Gainesville, no date» 1 malo, 1 female, Hogtown Creek, at
light, M, W, H», 6/19/60» 4 males, S females, Hogtown Creek, at light,
M. W* H*, 7A/60*
Cheumtopsyehe aphsnta Ross
Range* the range of this species is poorly delineated* it occurs
in Illinois, Indiana, Arkansas and possibly Florida#
Description» Ross, 1944*
Remarks* the identification of this species is based upen two spec¬
imens, both of Which were females* These specimens did not quite
agree with the description of the species given by Ross (1944), so
I sent them to Dr* Ross for confirmation of identification* He said
they were certainly very close to Cheumatopsyohe aphanta, but also

.72**
suggested that they sight represent a different species present in
this locality* It therefore seems advisable to record this species
as questionable until males are scoured which would then corroborate
the record*
Previous florida Records» lone*
Specimens Examined* Two females, Poe Springs, at light, M, Y/* K*,
9/k/SO*
Chcumatopsyehe pinaoa Boss
Range* Southeastern states*
Description» Ross, 1941*
Remarks» Chousatopsyohe pinaoa was collected on two occasions, both
times at light, at widely separated points**—•—Poe Springs and Hog-
town Greek*
The larva has not been found, consequently nothing is known
regarding the biology of this species*
Previous Florida Records» This species has not previously been
recorded from Florida*
Specimens Examined» One male, Poe Springs, at light, M* W* 7/lz/60t
1 male, Hogtown Creek, at light, M, W. 1», l/l/so.

Memoran Burmeister
Maoroneraum parolina (Banks)
Range» Southern states.
Pesoriptlen* Roes» 1944#
Regarles* The adults are among the largest caddis flies found in
Florida «ad are very striking, with their extremely long antennae
and polished, brightly*colored wings (fig. 92).
This species is only occasionally found in Aleohua County.
Ho larvae were collected; however, the larvae probably inhabit the
deeper portions of the Santa fe River, for Ross (1944) says that the
species frequents large, rapid rivers.
Previous Florida Records* Ross (1944) lists this species as occur*
ring in Florida but does not specify the locality# Betten (personal
comamioation) has taken specimens at Lake Wewahitehka*
Speolmene Examined* One male, Ebro, Washington Co#, L« &#, 6/30/40;
1 male, 6#4 talles W* of Washington Co# line at Hey* 10, Walton Co#,
L» B., 6/31/40| 2 males, Bayou Marquis, Escambia Co*, L# B„ G/l/4®i
1 male, Poe Springs, at light, M# VI# R.# ?/i2/S0; 1 sale, Vlorthington
Springe, at light, M, W# H#, 7/lz/8Qt 1 male, Prarie Creek, at light,
M* W. H,#^8/29/60.


•75-
etdroptilidab
This family comprises most of the "micro* caddis flies.
The adults are extremely minutel some having a total length of only
one and one-half milléraeters.
A good deal of work has reoently been done an the Hydroptllidae
of Florida. Denning (1947 and 1948) studied the mioro caddie fly
fauna of south and central Florida in some detail and described a
number of new species from there. Ross (1948)' has recently described
some new species on the basis of material sent to him by a number of
entomologists collecting in the state.,
Many Hydroptllidae have a very wide geographic distribution
but seem to occur only in certain limited localities within its
ronget for instance* Mayatrlchia ay ama occurs from the extreme
southern portion of Mexico and Florida to Hew York and Montana*
but within that broad area its distribution is spotty. It is very
evident that muoh more work needs to be done or the geographic
distribution of the whole group of mioro oaddie flies.
Key To The genera Of Hydroptllidae
Larvae
1. Middle end hind legs almost three times as long as
front legs* fig. 111.
Oxyethira
Middle and hind legs not more than one «id one-half
times as long as front legs* fig. 112.....•••••••
2

-76-
2* Anal legs distinctly projecting from tody mass#
fig* 113..**•*....i**»****»*#*..****»*»*.. #•**,.»*****♦»•**••#8
Anal legs apparently combined with body mss and
only the clans projecting*.*.#**.#..•♦*...•*.**•..**••*•***•4
3* Thoracic iergites clothed with long# slender# erect#
inconspicuous setae# fig* 114} case of sand grains*
evenly tapered and without posterior slit*
Keotriohia ranea
Thoracic tergites clothed with shorter# stout# black
setae which are conspicuous* fig* 116* case trans¬
lucent, evenly tapered and with dorsal side either
ringed or fluted with raised ridges# fig* 132*
Uayatrlchia ayaraa
4# Case long# smooth and round in cross section#
tapered at each end and with an indented slit
at both ends.
Orthotrichia
Case purse-lifos (womans change purse)*
Hydroptlla
Adults
1. Ocelli absent
2
Ocelli present
..3
2* Metasoutellum almost rectangular# fig* 116.
Orthotrichia
Metasoutellum pentagonal to triangular# fig* 11?.
Hydroptlla
3# Hind tibiae with only 1 proapioal spur*
Heotriohla ranea
Hind tibiae with 2 preaploal spurs*...*
â– 4

FIG. 118——Cxyethira florida, A, HALE CENITi;.LI A; B,
AEDEAGTJS; C, FEMALE GENITALIA.
B
FIG. 120- Oxyethira .ianella, MALE GENITALIA. A, LATERAL
ASPECT j B, VENTRAL ASPECT OF NINTH STERNITE.

kr
y; •
FIG. 121- Cxyethira lumoaa, LIALE GENITALIA.
A",' LATERAL ASPECT;' B, CLASFSRS.
7
\ v
A B
FIG. 122--—Cxyethira verna, HALE GENITALIA.
ASPECT-;'"B, VENTRAL ASPECT; C, AEDEAGLS.
A, LATERAL
-\ \ i
\V,
' \V\
FIG. 123- Oxyethira waiteri, MALE GENITALIA. A,
GENITALIA SHOTTING CLASPERS; B, AEDEAGL’S.

>?0«*
•54», Middle tibiae without a proapical spur.
Mayatrlohla ayaraa
Middle tibiae with a preapioal spur.
Oxyethira
OXYBTHIM. Eaton
Six species of this genus áre reported from Florida? how¬
ever, only one, 0, .jane!la,was found in Alachua County* Ho ehar-
aotera hare been found which can be used to determine the species
of the larras*
Key To f he Speoles Of Oxyethira
larvae-—»—#ot teeyed.
Adults
1* Male genitalia as in fig* 118? female genitalia as
in fig* 1180.
Oxyethira florida
8* Male genitalia as in fig* 119*
■ $&*$&** é&sm.
, ' p:
S* Hale genitalia as in fig* 120*
Oxyethira .lanella
4* Male genitalia as in fig* 121#
. Oxyethira luaiosa
8* Male genitalia as in fig* 182*
Oxyethira rema

■*•80»
G. Mal© genitalia as in fig* 123*
Cbcyethlra «alter i
Oxyethira .janslla Denning
•ii
Range? Florida raid L«ats|.aiia*
Description? Denning, 1948*
Remarks? Oxethlra lanella •nag described by Denning in 1948 on the
basis of two male specimens; cate from Winter Park, Florida and the
other from Row Orleans, Louisiana* This species is the only rep»
resentatlve of the genus Oxethlra that I found in Alachua County*
Since nales of £* janolla were collected along with females
at both Poe Springs and Worthington Springs, It ean be assumed
that the females are the undcscribed forras of this species* Many
other females of Oxyethira* collected at the Hirer Styx and Big
Hatohet Creek, could not be specifically identified due to the fact
that no males were present, la all probability these females be*
long to the species £* .jaaellat however, because of a lack of com¬
parative material, I do not feel that I can place a name on them
at this time*
Although almost all caddis flies that are attracted to light
oorae direotly to the source of the light, 1 noticed that £• .janella
carao to the more dimly illuminated portions of the sheet, such as

-81-
th© back of the sheet, indicating that thoy are less strongly photo-
troplo than most other species*
Previous Florida Records* "Holotype, malo——«Winter Park, Florida,'
May 16, 1940, (H# f* Femnld) * Paratype-——Kg* Orlecns, Louisiana,
October 10, 1945, (D, <3* Denning), 1 male*" (Denning, 1948)
Specimens Bxaadnedt One.male, Hogtosm Creek,' at light, M# W, II,,
7/4/60; 7 males, 62 females, Pec Springs, at light, M* W, N.,
l/lz/tiGi 1 male, 11 females, Worthington Springs, at light, M# W* H#,
7/ls/sOi 2 males, 15 females, Poe Springs, at light, M. W* K#,
7/5/51,
Oxethíra florida Denning
%
Fanget South Florida,
Description* Denning, 1947»
Remarks; Oxyethlra florida ms described by Denning (1947) on the
basis of specimens collected in Miami# 1 have not found this species
la Alachua County, and therefore, 1 assume that it does not occur a®
far north as Gainesville#
Previous Florida Reoorfts* "Flolotype, male»»—Miami, Florida, Feb# 1,
1945, D# 0» Denning# Allotype, female-—«.Miami, Oct» 1, 1944, D» 0»
Denning* Paratypes, ■ four males»——Miami, Florida, Hoy# 16-30, 1944*

-32.
Light trap* D* 6. Denning. Peratypes, three males, eleven females
Miami, Florida, Oct* 12-30, 1044, Light trap, D* G* Denning*
Paratypes, five mies, sJbc females, Miami,. Florida,- Oct# 1, 1944,
Light trap, D, G, Denning# Datatypes, five sales, nine females,
Miami, Florida, March 1-15, 1945# D* G. Denning# Paratypes, seven¬
teen males, twenty-nine females, Miami, Florida, Feb# 2-8, 1945#
Light trap, D# G* Denning* * (Denning, 1947)
Specimens Examinedt Dene#
Oayethlra glasa Rose
flange* Oklahoma, Florida, Georgia and Louisiana#
Description* floss, 1941#
Remarks* Denning (1947) found £# glasa to bo abundant in southern
Florida, bat it has hot as yet been collected in Alaohua County*
Sine© this species does occur in Georgia end Louisiana, it might
be assumed that it does inhabit some part of northern Herida#
Previous Florida Records* “Florida* Miami* February 1-8, 1946,
light trap, D# G* Denning, 2 males# February 16, 1944, D* G# Denning,
19 males# February 23-28, 1944, D# G# Denning, 16 males* March
1-16, 1946, light trap, D# G, Denning, .2. rales# November 16-30, '
1944, light trap, B# G# Denning, 16 males* December 4, 1944, D* G.
Denning, 14 males* December 14*15, 1944, light trap, B* 0# Denning,

*»88»
28 mies* â– 
Specimens Examined* lime*
Oayethlra limosa Rose
Ban get Florida*
Description* Rose» 1948*
Remarks» Oayethira lumosa was described by Ross on Hie basis of
me male collected at Daytona Beach. It has not* to ay knowledge*
boon collected sine®.
Previous Florida Records* ^Bolotype. male .————Dayt oca Beach* Fla*»
August 27» 1945, <5. t. Riegel.” (Ross, 1948)
Specimens Examined» lone*
Oxyothira verna Ross
Range» low Brunswick* Illinois, Florida and Louisiana*
Dosorintlmt Ross* 1988a*
Remarks* According to Denning (1947) Oxyothira yerna is the most
abundant species of Kydi'optilidae in southern Florida? however, I

have not. collected it in Alachua County# It aproare to have a wry
wide geographic distribution but only occurs to certain areas within
its overall rango*
Previous Florida Records? ^Florida* Miami* February 1-8# 1945,
Light trap# B* G* Denning, 168 nales# 92 females* February 15#
1944# D# G* Denning# 11 males. February 23# 1944# D* G* Denning# 3
mies* March 15-30# 1945# D* G* Denning, 12 mies# April 10, 1944#
D* G* Denning, 56 males* October 1-3# 1945» Light Trap# D« G# Denning#
5 mies# 28 females* October 12-30# 1944# Light Trap# D* G# Denning#
25 males, 31 females* November 15-30, 1945#. Light Trap#. D. G* Denning,
34 males* December 4, 1944# Light Trap# D« G* Denning, 18 males* "
December 14-15, 1944, Light Trap, B* G# Denning, 23 males* December
20# 1944, Light Trap, D, G. Beaming, 32 males*w (Denning, 1947)
Specimens Examined* None*
Oxyethlra waiter! Denning
Range* Florida, Georgia and Louisiana*
Description* Denning, 1947*
Remarks* Denning (1947) collected 0# waiter! in Miamij however, I
have not found it to Alachua County* Sinee this species occurs to
Georgia, one might surmise that it con be found somewhere to northern
Florida*

i
A B
FIG. 126- Orthotrichia
instabilis, VALE GEN ITALIA.
A, VENTRAL ASPECT; B,
AEDEAGUS.

•86»
Previous Florida Reoerdst "Ilolotype, mole»»»—»»Floridat Miará t ■■•■--
lovember SO, 1944, Light trap, B* G, Denning* Paratypest Miami*.
March 15, 1945, D* 'G* Denning, 6 mies» May 10, 1944, 9* 0* Denning,
12 males* Movember 16, 1944, D* G. Denning, 4 moles* December 20,
1944, D* G# Denning, 1 male." (Donning, 1947)
(MH0TR1CHIA Baton ...
'i
Sob® females of this genus, found at several localities
(River Styx, Worthington Springs, Devil1 a Millhopper end Pm Springs),
could net be identified specifically due to lack of ooraparativo
material* A larva found in Lake Alice also could not be identified
to epodos.
Key To The Species Of Orthotrichia
Larvae»»»»»»—*»!! ot keyed
Adults
1* Male genitalia as in fig# 124} female genitalia as
in fig* 127*
Orthotrichia americana
2* Male genitalia as in fig* 125; female genitalia ae
in fig. 128*
.Orthotrichia or1stata
Si Male genitalia as in fig* 126; females not fcsyed*
Orthotrichia instabiliw

«S?«*
1
,f jj\o
w
1
FK
• >; ' ■ - a
V
A
A
( }. X
: i r.
,
VJ americana, FSTA
LE
PIC
cr:
x. 128-
L s+ipIr ,
*-"•* C 4
, AA'AL]
Orthotrlohla americana Banks
Mange* This species has a wide range encompassing an area fresa
Texas and Florida in the South to Hon York and Minnesota in the
Horth#
Pesorlptlon* Ross# 1944*
Remarks* Penning (194?) found £# americana to he present throughout
the year in Miami* however* 1 have not taken any males in Alachua
County* This species does occur in this section of Florida# though#
for specimens fresa the St* John*s River in Putnam County were ox*
«mined ty me* Perhaps the unidentifiable females and the torva
found in hake Alice also belong to this species*
Previous Florida Reoordst "Florida* Miami* October 1*3# 1944,
tight trap* P* C# Penning# 40 males# 7 females* October 12*30# 1944#
P* 0, Penning, 7 mies# t females* Hovember IS* 1944, P» 0# Penning*

—88—
1 male, 1 female* February 1-8, 1945, Light trap, D. G. Denning,
81 inales, 15 females* March 16, 1945* Light trap, D* G, Denning,
1 male# May 10, 1944, D, G, Denning, 14 males, 12 females.”
(Denning, 1947)
Specimene Examined» Four males, 10 females, shore of the St* John’s
River, at light, J. H., t/lh/m.
Orthotriohia oristata Morten
Range» Most of eastern and part of western United States*
Description? Ross* 1944*
Remarks» Denning (1947) collected one male and one female in Miami*
I have not found this species in Alachua County,
Previous Florida Records» "Florida» Miami»■,April 10* 1944, D, G.
Denning, 1 male, 1 female•" (Denning, 1947) .
Speelmans Examined? Hone

FIG. 130 Hydroptila wakulla♦ A, MALE
GENITALIA; B. FEMALE GENITALIA.

â– 90-
Orthotrlchla instabilis Doming
Range* Central florida*
Description» Penning» 1948a*
Remarks» Penning (1948a) described 0, tastabilla on the basis of
nine malea collected in Winter Park* I have not found this apeóles
in Alachua County and oan give no further data about it*
Previous Florida Records» - "Holotype* male-»-»»—Winter Park*
Florida* May 16* 1940, (H* T* Fernald)• Paratypes««~-~--Florida*
samo data as for holotype, 8 males*0 (Denning* 1948a)
â–  *
Specimens Examined» I fffPROPfm Palman
This genus embraces about forty Eearotio apéelos, comprising
cne-third of the Hydroptllldae* Only two species, H* berneri end
JU mkulla have been taken in Florida*
Key fo fhe Speeies of Hydroptlla
LajrmM—-Wot keyed.
Adults
1* Male genitalia as in fig* 129 A and Bf female genitalia
as in fig* 129 0*
Hydroptlla berneri

-91*
2* Mui© genitalia as in fig* ISO As fonal© genitalia as
in fig* ISO B,
Bydroptila wakulla
ffydroptila bernori Boss
Banget florida*
Deaorlptlcmt Boss* 1941*
Remarks* This species was described by Ross (1941) on the basis of
two mies and ene female collected at Foe Springs by Or* Lewis
Berner* i was able to collect topotypioel material and ms also able
to collect the undoaoribod larra* Shis larva, Which is fairly
oomon at the rook dam near Poo Springs, makes a purse-shaped case
of sand grains* The oases are attached to submerged vegetation and
rooks in the slower moving portions of the river around the dam*
The adults are attracted to light, lut seem to case to the
light a little later in the evening than most of the other caddis
flies*
Previous Florida Records* Two males, 1 female, Poo Springs, at
light, collected by Lewis Berner, s/4/S9. (from Boss, 1941)
Specimens Tixaalnod* Four males, 10 females, Poo Springs, at light,
II* Tf, II., 7A2/50J 1 male. Big Hatehet Greek, at light, H* W* H„
4A2/6O1 1 male, Poe Springs, at light, 1* W* K*, 9/9/50* 5 males.

5 females, Poe Springs, at light, !!• TT, IT*, 7/3/51! 6 larvae, Poe
Springs, M« W* H., s/24/50*
Rydroptlla «alalia Donning
Range» Florida*
Description* Denning, 1947*
Remarks» Hydroptlla makulla «as described by Denning (1947) on the
basis of 11 specimens collected at Wakulla Springs* I have not found
this spooles in Alachua County end have found no other data concern¬
ing it in the literature*
Previene florida Records* "Hole-type, male—--—Florida* Wakulla
Springs* October S3, 1945, D, 6, Denning* Allotype, female-——
Same data as for holotypo* Paratypes-——Sam® data as for holotypo*
1 male, 8 felBales*,, (Denning* 1947)
Specimens Examined* ffaae*

•»93~
HEOTRIGHIA Mortoa
Heotrlohia ranea Penning
Hanget Georgia and Florida»
Description! Denning, 1947•
Reaarkst Keotrlehla ranea le undoubtedly the smallest caddie fly
•feat ooours In Alachua County* a full-grown male is only one and one*
half millimeters long#
This speoles was described by Penning (1947) m the basis of
two males# one from Florida, fee other from Georgia# I hare o&leoted
the imdesoribed female of the species*
Previ cm g Florida P.eoorda* “One male# Miami* May 10, 1944, P. G#
Denning." (Denning, 1947)


Specimens Sxaalned* Two malea* 2 females* Worthingbcm Springs* at
light» M* W, »*» l/iz/mt 1 female* Poo Springs* at ll#t* H* W* H.,
7/12/sOi 1 malo* 1 female* Poo Springs* at light* 13* W# E*t 9^)/60«
M&YATRICHXA Moeely
This genua contains three Korth American apeóles» of which
only ene» M* ay ama» has been taken in Florida*
MayatrloMa ayaaa Hosely
Ranget Tory wide-spread over most of the eastern states* Ranges
from extreme southern Mexioo to Hew York end Montana* Although the
records are scattered over a wide area* the species is quite rare
in collections*
Description* Ross# 1944*
Remarks* . Ross (1944) lists this species as occurring in Florida# I
examined six •females which* due to the lack of comparative material*
cannot 1» identified to specios} however, these specimens (one from
the St* dohn*s River in Putnam County and the other five fresa Poe
Springs}' most probably belong to M* ay am#
Previous Florida Records* Ross (1944) records this species from
Florida bat does not specify tho locality*

-96.
Specimens Sxaainedt Ãœms.
PHRTGAKE3DAS
BAIIKBIOIA Martynov
Banksiola concatenate (Walker)
Walter (1862) described this species m the basis of speci¬
mens oaugbt at St» John's Bluff, Bast Florida, which is near the
mouth of the St* John's River* 2 have not collected this species
but am including it for the sake of completeness*
III.HEPHILIDAI
rXCKOPSTOHB Battles
The fatally Limnephilidae is a very large one containing
numerous species scattered throughout the United States, and thus
it Is rather suprising to find only one representative occurring in
Florida*

FIG. 155- Pyonopayohq sp. A,
Caa«.

Pycnopsycho sp* A
This larva is cue of the largest caddis flies found in
Alachua County* The ease (fig* ISS) is composed of a hbdge-podgc
of sticks# and is very well earacflemgcd#
1 did not collect this species# ¿id t do not know if it
occurs at present in Alachua County* Ho adults of this genus have
been collected in Florida* and, since no characters have been found
to separate the larvae of the various specie»,, this larva cannot
be identified further than genus.
The members of this family lay their eggs an vegetation
overhanging water rather then in the water and depend upe® rainfall
to wash the eggs into, the body of rater#
Specimens Examined* .One larva# 5 miles E* of Gainesville, B* A, B*
I2/8/59* 1 larva, Eogtovn Creek, A* J. H., g/lg/dS* d larvae. Hog*
tom Creek (in leaf drift), P* K, Y,, 12/si/4?s 1 larva, 2§ miles
W, of Gainesville, Z>* B«, 1/29/59* 1 larva, © miles W* of Gaines¬
ville* B. A, B„ 12AA9.

.99.
mumk Curtis
Molonna tryphena Batten
This is a Mew state record* X have not collected M*
tryphena In Alachua. County# tut Batten (personal eoneaunioatiao)
found It in Florida (locality not specified)*


«10l«*
tEPTOCBRIDAE
Proportionally, I believe there is a larger percentage of
species belonging to the fussily Leptoeoridao in Florida than there
is in northern states, The members- of this family constitute ap¬
proximately forty percent of tho total number of species found thue
far in Florida#
All the larvae in this family make oases of many shapes
(figs* 2.68, 169, 170, 171, 182, 185 and 200) whioh are constructed
of a variety of materials, such as sand, vegetation and bits of
debris* The adults are character!stieally slender and have long,
thin antennae*
Throughout the family the pupal chamber has a slit end not
a mesh in the closing cap at both ends* Through these slits the
pupa pushes oat all the larval solerites, so that it is impossible
to get associations of adult structures and larval sclerites in the
«
same case* For this reason it is necessary to rear larvae in order
tó associate immature and adult stages*
Key To Tho Genera Of Leptocerldae
larvae
1* Middle legs with clan stout and hook-shaped* tarsus
bent, .fig* 157*
Leptooerus amerlcanue
Middle legs with claw slender, slightly curved,
tarsus straight, fig* 130


-IGLU
g# Maxillary palpi nearly as Ions as stipes*- fig* 139;
rendibles long* sharp at apex* the teeth consid¬
erably below apex*
Oecotis
Maxillary palpi short, about half length of atipes*
fig* 140; mandibles shorter* blunt at apex, the
teeth near or at apex******»***»*****«##*#*****»***»»«**#«**3
S* Head with a suture-lit» line paralleling the
epicranial arma* fig* 141*
Athripeodos
Head -without a suturo-liko line paralleling the
epicranial arms**.***. 4
4* Meeonofun membranous with a pair of sclerotised*
narrow* curred or angled bare* fig* 14g*
Athriosodes
llosanotum without such a pair of eolerotised bars*#..«.*««•..«5
8* Anal segment developed into a pair of eolerotised,
concave plates* with apiñóse dorso-lateral and
mesal carinae, end an overhanging ventral flap*
fig*' 143*;
Setodes floridana
Anal segment convex and without carinae between anal
hooks, fig* 144. é **»***•*••** * »••••••* .•*•##«#**•*••*#•••#*6
6* Hind tibiao entirely eolerotised, Tdthout a fracture
in middle* fig* 186; abdomen without gills*
Leptooella
Hind tibiae with a fracture near middle which appears
to divide tibiao into two segments, fig* 145; ab¬
domen with at least a few gills*
friaonodes

americanus, MA.LE GENITALIA.
americanus, FEMALE GENITALIA

Multa
1. Front «inga with atoa of II atrophied» leaving only
two min veins â– 'between convex Rj_ 'and convex Cu^*
. triaenodes
Front wings with stem of II present,' so that three
main veins are present he tween convex Ej and
convex Cu¿, fig* 14G. •....>*••*...•«.••2
2# M apparently not branched, fig# 146*
Oecetia
1 obviously branched, fig# 147*#*** •*•*«•••*««*•*••»•••*•••*•#3
3# Epicranial stem distinct, lateral sutures absent
or indistinct, lateral sutures absent of in¬
distinct# fig* 148#
Setodos florldena
Epicranial stem absent or indistinct,' lateral sutures
well narked# fig*. 149»»*»#»»#*»*#*»*»»#*»»** *#*******.#*•**•4
4* ’lings white, usually spotted* •
Lcptooolla
Wings not white and spotted ##•••*•*••*#••«**#*••*••••••*«•**4*6
5* Front tibiae with 2 apical spurs# .
Athripsodes
Front tibiae without apical spurs*
- Lcptooerua araerloraua
LEFfOCEHUS Leach
Leptooorus amerloenua (Banks)
(Sestodos americana Sania, 1899
Seatodea granáis Banks,’ 1907a)

•10G»
Range* Recorded from Illinois, Re» York, Conneticut, Virginia,
Michigan, Indiana and TTisconsin* heptocorua araericanua 1ms not
previously been taken in Florida, therefore, finding this species
indicates that it occurs'in the area between Virginia and Florida,
Deaeriptimt Ross, 1944*
Remarks* . Emergence in this opooies talces place only; daring the
spring in Florida. RoSs (1944) collected â–  L. onoriocnita mainly
around glacial lolces end in the slow streams connecting them* In
these places the larvae were almost invariably found in water horse»
tail. I have not collected the larvae in Alachua County and have
no idea where1 they may-be found*. ': â– 
Previous Florida Records* Rone*
Specimens Examined* Yhirty-ona males end females, Gainesville, at
light, H# W* II*, 4/23/SOj 14 females, Gainesville, at light, II* W»
II*, s/s/80j 2 females, Gainesville, at light, II* Vf* U„ s/s/60s 8
females, Gainesville, at light, II# W, R*, g/s/so? 1 male, 1 female,
Gainesville, at light, W, !!„ gAo/so*


408*
1SP2CCSLLA Banks •
íhe adults of this genus are elongate, slender insects* ■'■
having long antennae cad enemy whito wings flecked with black or -
brown# Sosa® of the species aro very beautiful and showy*
It is rather difficult to identify the species of heptocolla
as our Imotiledge of the genus is incomplete and many oS the Identic
fying characters are dependent upon the easily destroyed pattern
of hairs on the wings* I collected many specimens which could not
be specifically identified because the hairs cn the wings liad rubbed
off*
It is essential to have both alcoholic specimens and dried*
pinned specimens for identification, 2hc specimens to be pinned
were killed a tm at a time in a jar containing strong cyanide*.
they were then pinned very carefully* as satisfactory results ware
obtained through gehtle handling* â– â–  haptocella is the only genus of
caddis flies In which pinned material is desirable#
Key fo The Species Of Loptooolla
Larvae
1# Head as in fig* 153*
Leptooella pavlda
2* Head as In fig# 154.
Leptooella albida


-no»
3* Head CG in-fig*.155*
Loptocolla candida
4* Hoad as in fig* ISO*
Lentocella exquisita
Adults
1* fíale genitalia as in fig# 163} wings as in fig# 163.
Leptooolla pavida
2. Wing.as in/fig* 164*
Leptocella alblda
3#. Malo genitalia as in fig* 169} 'tá.ng ns in fig* 16S#
Leptocelia candida
4# Male genitalia as in fig* 160} wing as in fig. 166#
Loptocolla exquisita
5. Male genitalia as in fig. 161*
leptocella tasara
6. Male genitalia as in fig* 162s wing as in fig* 16?.
Loptocolla gpiloaa.

•111-
FIG. 163——L. pav id a, WING
PATTERN. “
FIG. 164 L. albida, WING
PATTERN.
FIG. 165 L. o andida» FIG. 166—L. exquisita,
Wllíft' ‘PaTTERÍT. ViltlG PATTER^.
FIG. 167—L. spiloraa,
Wing patteríT.

Lentocella candida (Hagen;
(Setodes candida Hagen, 1861)
Range» Central and southern states*
Description* Ross, 1944*
Remarks* The description of this species needs revision for it is

aiiv
impossible to tell what form earlier authors had before them in using
the various names applied to L* candida»
I collected this species (me larva and me adult) on only
two oomelons» both times at Poe Springs* Ihe larva was found to a
shallow» sheltered spot near the rock dam where the water was only
about one foot deep and the current was considerably slowed* loss
(1944) found it In a wido variety of streams and rivers and also in
marshes»
Previous Florida Records» Hagen (1861) lists this species from Florida
but does not specify the locality*
Specimens Examined* One larva» Pm. Springs* M* W* IT*» s/sl/so*
1 male» Poe Springs, at light, M* W. IT., 7/b/fa*
- toptooolla exquisita (Walker)
(teptocerus exquisita Walker* 1852)
Range* Eastern, southern and central states*
Description* Ross* 1944*
Remarks» ■ I believe that this species must be rare since only two
specimens of h* exquisita were found in Alachua County* It may live
to the St* John’s %ver, as thirty-four males were collected while
lighting on the shore of that river one night* Ross (1944) found it

~U4~
only in laicas*
Previous Florida Records» Ross (1944) records L* ©sepilalta fresa
Florida tat does not speoify tho locality.
Specimens Examined* One male* Poe Springs, at light, M* W, IT*,
7/L2/6OS 84 malos, St* John*s River, Putnam Co*, at light, J* H*,
7Á2/60*
beptooella pavlda (Hagen)
Ranget Occurs mainly in tho eastern and southern states* ,
Description* Ross, 1944*
Remarks* X found L. pavida to bo the most abundant representative
of the genus Leptooella in Alaohua County* The somewhat smaller sise
of the insect (10 mm as compared with 12-17 mm of other members of
this genus) is a useful characteristic for identifying the species
in the field*
Ross (1944) associated the larva© and the adults on tho basis
of a grasp of nature pupae collected at Poo Springs by J* S* Rogers
in 1935* X found the species to be fairly common at the same place,
and X have also been able to rear the species*
Previous Florida Recordst Ross (1944) records a collection of larvae
and pupae made at Poo Springs, 4/15/55 by «J* 3* Rogers* Betten (1934)

-316-
records I»# pevida fresa the Chipóla River,
Spoolraons Examined» One female, Torreya State Park, Liberty Co#» L,
B# j 6Á0/5SI 1 female, Ebro, Washington «o., L, B„ s/so/m 10 fonales
Hogtotm Creek, at light, 12* W* I!#, B/lo/sOi 1 pupa, Po© Springs, II#
W, II,, s/ñl/SOi 1 fórnale» slow stream 10#5 miles li, W, of Gaines¬
ville on U« S# 441, i!# W, 13,, 6/21/6O1 1 female, Gainesville, at
light, IT, Vi, !i„ 6/g4/60} 1 malo, 2 females, Poe Springs, at light,
!1, Vi, l!,, l/lz/SOi 5 females, v/orthingtm Springs, at light, M, If,
I#* 7/45/60} 1 male, 16 females, St* John’s River, Putnam Co,, at
light, J, H„ 7/46/60} .2 males, 12 females, Poe Springs, at light,
M, V/, 13», 0/9/so} 2 sales, 2 females, Po© Springs, at light, M, W,
i,* 7^Ai# - â–  â–  â– 
Leptooella gplloma Ross
Range» foxas, Kansas, Missouri, Illinois and Florida,
Description* Ross, 1944,
Remarks» the larva of L# gplloma Is tmknomi and 1 collected only
one specimen of this species* a male, in Alachua County# Sine# this
male ms found at light in the city of Gainesville, no guess can bo
ventured as to its habitat,
the larvae of Leptooella sp, A aré probably either the tax-
described larvae of L# apilóme or L, tavara, for these two species

«fi¬
are the mlj1 species of this conus thus fair'found in Florida hawing
imdeacribed larra©#
Previous Florida Records* Hen©.
Sneeimens Bramlnedt One male* Gainesville# at light, II# ft# I?#,
e/gs/so#
Leptooella alblda (ftalfcer)
. >v.■ > •' * o
FIG. 168—*—*Loptooella albida»
LAIí/ii AOD OAStí.
Betten (personal oommioatien) found this speoies in the
vioinity of Lais© Plaold# I have not collected It in Alachua County#
end there are no previous Florida records of it in the literature*

<•217.
Leptocella talaya Ross
Ranges Central Florida*
Pesorlptlopt Boss, ,1944*
Remarkst there aro nine specimens in riy collection* which were
taken cm the shore of the St* John*3 Hirer, Putnam County* that may
he this species, hit since the hairs cm the wings are mostly gone,
and I hare no supplementary alcoholio material, the specific de»
termination is questionable* I bollero further collecting will show
that L* tarara does occur in Alachua County*
Leptooolla sp* A is probably the tmdoscribed larra of either
L, tarara or L* spiloma. Further collections, especially those from
oentml Florida, will make identification of this larra possible*
Frorious Florida Records» fh« following records are from toss (1944)*
Soren males*. 3 females, Chiefland, Pery Go#, collected by ¥* Stehr,
7/l7/S3s 1 sale, Winter Park, Grange Co#, collected by S* II# Paris,
s/25/?í S males, Winter Park* collected by E* II# Paris, s/ss/?* 1
male, Winter Park, collected by E* M# Paris, 4/c/?t 3 males, Tarares,
Lake Co*, oollectod by F# R, Young, 3/23/86*
Leptocella sp# A
This is a largo, striking, undescribed Leptocella larra


was collected % Mr* Bills Lanquist In Bevy County* It undoubtedly
is the larva of either L* gplloaa or L# tarara* for those two species
are the only species of this gents thus far found in Floil d® haring
undescribed larrae. Collections from other parte of Florida are
needed to determine the true status of these larrae*
I hare not been able to illustrate this larra because the
me specimen I had inadrortently disappeared# This larra» however,
was collected in a sand-hill lake, Cospel Pond, Lory County, in the
spring of 1950#
ATi-iRlPcaDSS BiUberg '
: The larvae of this genus are short end build some unusual
oases (figs* 170 and 171)* A large number of species hare been
desorlbed from North America, of which four hare been collected
in Florida#
Key To The Specioe Of Athripsodes
Larrae
1. Head as in fig* 172*
játolaaflfca fruakms&te
2* Head as:"ln fig* 173*
Athripsodes canocllatua
3* Head as in fig* 174*
Athripsodes sp. A


4* Head as in fig. 17 5 j ease as in fig# 171.
Athripsodes ep« B
Adults
1. Males#••.••••#*••••«•#2
Pernales»##..**.
2# Genitalia as in fig* 176#
Athripsodes cancellatus
S* Genitalia as in fig* 177*
Athripsodea tarsi-punetatus
4# Genitalia as in fig# 178#
Athripsodes solossonae
6# Description given under discussion of species.
Athripsodes floridanus
6# Genitalia as in fig* 179*
Athripsodes canooliatus
7* Genitalia as in fig* 180*
Athripsodes tarsi-punotatua
Athripsodea ¿loridanus (Banks)
(Leptooerus floridanus Banks# 1903
Athripsodes floridanus Milne# 1934)
Range* Florida*
Description* I have not seen this species* end there are no pub-

llshed illustrations of anatomical structures by naans of which en
identification can be nades se for the sake of conjletness, the
original description is included*
*Hettd'yellowish, clothed with long, tft&te hairi pronotum
Tilth long white hairs rest of thorax yellowish, with shorter and
nor© sparse White hairs antennae White, narrowly annulate tilth dark
browns legs.yellow, with short whit® hair, tarsi banded on tips with
browns wings pale brown, rather densely clothed with white hair,
mostly in email patches, giving the wing a ramoratc appearances
apical part slightly infuscateds( *) Length to tip of wings 10 m.”
(Banks, 1905b)
Milne (1934) presents a key by moans of which the females
can be identified* Athrlpsodas floridanus is identified by negative
characters (lack of anatomical features) rather than by positive
characters! therefore, the key is of little use in this instance*
Remarks; Athripsodes floridanus was described by Banks (1903b) on
the basis of case specimen, a female, collected by SIosnan near
Biscayne Bay* This species has not, to my knowledge, been taken
since*
Previous Florida Records* "One specimen from Biscayne Bay, Florida
(Slosson)*” (Banks, 1903b)
Specimens Examined* Bone*

<*¿25*
Athripsodes slossonae Banks
', fhis species uns described by Banka (1958) irosa specimens
colleotcd at Bellaire, Florida and Harrisburg, Pennsylvania* X
have not collected it in Alachua County, and there are no other
records published in the literature*
X believe that Athripeedes sp* A or sp* B is the undescribed
larva of A, slosoonae. Collections iron other parte of Florida are
needed to determine the affinity of these larvae.
Athripsodes caneellatua (Betten)
(heptooerus cancellatus Betten, 1954)
Range* Eastern and southern states, trestmrd through the Osarks to
Oldahom* â– 
Description* Boss, 1944#
Remarks* Athripsodes cancellatus Is rarely found in most of Alachua
County, but can be taken occasionally at Poe Springs* X collected a
male and a female there and found eleven empty larval cases in the
Santa Fe Biver uhlch I believe belong to this species* Repeated
collecting at this end other adjacent sites sere fruitless* Boss
(1944, p* 255) gives a good description of the case of A. cancellatus*

Ӓhe larval case is about 0 in# long* horn shaped* and the
main body is composed of fairly regular sand grains with a fe»
larger grains arranged along the side* giving very slightly the
appearance of a lateral extension.”
Previous Florida Reoordst This species has not previously been re»
ported from Florida# Batten (personal communication) has collected
it in the state but did not specify the locality#
Specimens Examined* One male* 1 female# Poe Springs* at light# II*
Yf# H*# 7/L2/5QJ 11 larval oases* Poe Springs* M. ¥* if,, 9/9/SQ#
Athrlpsodes tarsi*punetatua (Vorhiea)
(Leptooerus tarsi-punotatus Vorhies, 1909)
Batten (personal communication) collected this species in
Florida* 1 have not collected it in Alachua County, and there is
no published record of its occurring in the state#.

125-
Athripsodes an. A
»
FIG» 170—-—Athrlpsodes sp. A,
LARVA MD CASE.
These undescribed larvae, fig. 174, with their peculiar
flange type oases made of asnd grains, fig. 170, were collected in
the Santa Fe liver aome years ago. Repeated searches in 1960 and
1951, however, failed to bring to light a single specimen. The larvae,
most probably, are the undescribed iramatures of either A. floridanuB

126-
or iL* slossonae, for these two species are the only species thus fer
found in Florida haring undescribed larvae.
Specimens Examined» Two larvae» Santa Fe River, 3/24/37 j 1 larva,
Santa Fe River, J. S. R., 3A2/38.
Athripsodes sp. B
FIG. 171--»—Athripsodes sp. B,
CASE.


-129*
These undescribed larvae, fig. 178, have been collected
only in Swim Pond# in Marion County* The case# fig* 171# is made
of bits of leaves end vegetation rather than sand grains* The larvae
most probably# are the undesoribed immatures of either A* florldanua
or A, aloaaonae*
Specimens Examined» Two larvae# Stria Pond# Marlon Co## SAc/SSf
6 larvae# Stria Pend# Marita Co## 2/26/S7.
OECSTIS MoLaohlan
The genus Oecetls is# without doubt# the largest and probably
also the most abundant genus of oaddis flies in Florida* Of the
fourteen species present in the United States# eight have been col¬
lected in Alachua County# and of these# three# Oeoetls sp# A#
O# daytana and O# parva# oocur only in Florida*
The larvae an» predacious and have elongate# grasping
mouthparts* Some speed as have wide ecological tolerance and occupy
many habitats# Which include lakes# rivers# streams and pends*
Key To The Speeles Of Oeoetls
Larvae
1* Head as in fig. 184i log-cabin type case*
Qooetis ciñeraseens


2. Ilead as in fig* 186*
Oeoetis inocnsploua
3* Head as in fig. 186| ease as in Jig* 182.
Oooetia sp. B
4. Head as in fig. 187j ease as in fig. 183.
Oecetis sp* C
Adults
1. Mes... 2
Females**.* **..10
2. Genitalia as in fig. 188.
Oeoetis Inecpspleua
3* Genitalia as in fig* 189.
Oeoetis sp. A
4. Genitalia as in fig. 190*
Oeoetis dayteaaa
6* Genitalia as in fig. 191*
Oeoetis osteni
6. Genitalia as in fig. 192*
Oeoetis avara
7* Genitalia as in fig* 193*
Oeoetis georgla
8* Genitalia as in fig* 194#
Oeoetis eineraseens
9* Genitalia as in fig. 196.
Oeoetis parva


10* Genitalia, ge in fig* 196*
Oeoetlg lnconepicua
11* Genitalia as in fig* 197*
Oeeetís osteal
12* Genitalia as in fig* 198»
Oecetis avara
1S* Genitalia as la fig* 199*
Oeoetls einerasoena
Oeeetís inoonspioua (Walker) complex
Oeoetls inconsplona is me of the most common caddie flies
in ITorth America* At the present time* there are three closely
related species in Ml term the Oeeetis inoonspioua complex*
0. inoonspioua (Walter, 1852), £♦ portari Boas (194?) and 0* pratelia
Denning (1943b)* this complex deserves careful study for it poses
a number of highly interesting taxonomic problems*
Oeoetls porter! Boss and 0* pratelia Penning* both of Phioh
resemble 0. inoonepieua* were described within a few months of each
other from florida specimens and most probably are eynmomouB# How»
ever, they differ fresa JO* inoonspioua in a number of ways, Including
sise, wing maeulaticn and genitalia, so they might arbitrarily be
thought of as a species distinct from 0* inoonspioua, and, for the
purpose of this discussion may be called species A* Species A is
a highly variable conglomerate (both adults and larvae), and may

—iss*»
Itself be subdivided into other units upon further study#
Banks described two species on the same page of his 1899
paper——Oeootlaa flavlda from Kissimmee end Oeoetina floridana
tram Bisoayne Bay—-—¿which have both been eyncncraised to 0,
inconspioua but which really may be equivalent to species A. Hagan
(1861) described Sestodes sagltta from. Florida# and this was also
synonomised to £• Inconspioua» Furthermore# Inconspioua was
first described by Walker (1852) as Leptooerus lnccnsplouus on
the basis of three specimens collected in Georgia by Abbot# and
species A comes close to the description based on Walker*s types
given by Batten end Moscly (1940)* If this is true# species A
would be synonomlssed to 0, Inoonsplcua# and the northern species
would be synonosnised to the next species having precedence# 0#
flaveolata*
I have examined specimens of "Of* inconspioua11 from north¬
western Florida which are quite different (such larger# etc*) from
the material collected in Alachua County (species A) but are never¬
theless closely related to species A» I also examined one specimen
of species A, a femóle# from northwestern Florida (Santa Hosa County)#
Both seom to exist in Florida»
The solution of this problem requires an extensive col¬
lection of specimens from different parts of the United States
plus a careful perusal of the various types# and I am not in a
position to offer eny defínate conclusions at this time* I would#

heerwr * Wm to call attention to this confusing situation «id «g»
go it the following 1 úme «tilo h occur rea to »t (I) Clootie oerterl
*nd 2* nrctelle or© eynenenottSj (S) the Osaetie. locengolcm ©©Bp!®*
It really R ’?©ry widely distributed tptolet with highly, dtitlnet
eehepeelee, (3) tb© ©eaiplex It a grasp of distinct epea&ee, (4) a
eomfeifsatloa of points 2 ord 3 exists, and (8) if doflr.ito ©recios
art present* «pesies A -would be v^mones&ged to £• Iftooneeloua*
wMeh tms precedenoo, aid the northern sp-edea would be tmmmé*
A graph!© «ptepsl® of the syneooof that s»y exist, fell owe**
. hentooerua tr.eono^euua Walter (1882)
Go at ocio8 sagttta Hansn (1881)
üoeetina Habida Baefee (1803) a 0* Inoonanloua « ft
TBwefe Ma Bents (1800) * VtiS!$¡i~ 7)
^wotienorterl Rosa (1947b)
W^Ws TTrato'li’a Penning (1945b)
tP* A
2« Forthora fern, ©f £* • H* !?• Florida • £* flneeolat©
inemttntma (Walter) ferae (7) *“
for the paipai* of this paper 1 shell treat those epeelovoo
eolleoted in end around Aiaohue County «© Ooeetls op* ñ end' w»t of
those eollooted in northwestern Florida as £• taocoaolmm»

.158-
Oecetie sp* A
Rang»* Florida and Georgia»
Description» Roas» 1947b»
Remarks» For the purpose of this paper Q» porter! Ross (1947b)
8115 £* pratelia Denning (1943b) will arbitrarily be considered as
synonoraous end will be called Oecetis sp» A# The speoies were
described within a few months of each other on the basis of Florida
Specimens (O. porter! from Miami, Hew Smyrna and Daytona Beach and
0, pratelia from La Belle), and they appear to be morphologically
very similar»
The degree of variability was not recognized by Boss (1947b)
In hie description of 0. porter!» Because of additional information
before me it appears that the original description 1# not adequate
and will need revision.
I collected the previously undescribed larva, and it, also,
is highly variable, ranging from a light shad® with very indistinct
markings to a dark coloration with highly distinctive markings* The
larva shows a similarity to the drawing in Ross*s paper (1944, p, 245)
(and this thesis, fig. 186), but it is different nevertheless»
Oecetis sp. A Is the only representative of the Oecetis
in conspicua complex found in Alachua Counts'* I also examined me
speolmen, which was collected by Dr. Lewis Berner, from northwestern

-136-
Florida (Santa Rosa County). I do not believe that conclusions can
be dr aun, however, until further collections are nade*
In Alachua County the larvae occupy many habitats* and
emergence takes place throughout the year with the preponderant
portion of adults emerging during the warmer parts of the year*
Previcms Florida Records! Since the synonomy has not been definitely
proven* the rooords will net be included*
Specimens Examinedt Bight larvae* Kingsley Late* Clay Co*, Llenan
Dredge sample 10 feet deep between Maiden Cane, J* S* B* and A* F.
C,* 5/l9/sZt 1 female, Santa Rosa Co., L, B*, 6/l/40; 2 males, 2
females, Tavares, Late Co., F. H# Y«, 5/25/36* 6 larvae, 1 pupa,
Alachua Co*, no date! 1 larva, Late Santa Fe, bottom sample, l/d/48j
3 larvae, slow stream 10.5 miles H* W. of Gainesville on D* S* 441,
M* W, !!., 6/20/6O* 1 male, Biven*s Arm, at light, M. ’.T, H*, 8/23/60*
1 female, Gainesville, at light, «I* W* N,, 6/23/60* 1 male, 1 female,
Prarie Creek, at light, M. W. II», 8/29/50* 260 males end females, St.
John* 3 River, Putnam Co*, at light, J* H., 7 As/50 * 1 female, Pond
B, at light, M. If* H», 7Ao/SOj 1 male, Orange Point, Welaka, St*
John*s River, Putnam Co», sweeping, M, W* E,, 0/26/60* 1 female,
Gainesville, at light, II* Vi, E., 6/3/6O* 1 female, Gainesville, at
light, M* W, IT*, 5/9/50} 1 male, 2 females, Poe Springs, at light,
II* him E*, 9/9/6OÍ 1 male, 14 f ornate, Hewn an* a Late, at light, II* W,
E«, 6/23/50} 12 females, Worthington Springs, at light, M, VT* E«,
7/13/5O5 1 female, Devil’s MiUhopper, at light, M* Tí* IT,, o/s/so;

•287»
2 usías, roared from pupae found in stream 10*5 miles 14 ?'• of .
Gainesville on 14 S* 441, 14 IT, 14, S/Sl/sOf 8 males, 4 females. Big
Hatchet Creek, at light, 14 IT, 34, 6/12/60? 9 males, 17 females,
River Styx, at light, M* Yi« 14, l/ll/dOt 2 males, 3 females. Big
Hatohet Creek, at light, IS* W, 34, 8/2/61? 3 females, Gainesville,
at light, M* W* 14, 9/l9/ülí 1 female, Poe Springs, at light, M*
W, 14, 7/12/50? 3 males, 5 females, Poe Springs, at light, M* W.
34, 7/3/6I? 1 larva, Alachua Co., no date*
Oecetls in eon splcua (Walker)
(Lontoccrus inconsolouua "falker (1852)
'Matodes sagltta kagen (Í861)
Setoderar nicans 'Hagen (1861)
Set'odes flava blata, Hagen (1861)
'óecetina parvula^Bankg (1899)
Oeoetina' flavida Banks (1899)
Oeoetine* fiori'dana Banks (1899)
beoetina apioalle fcan ks (1907a)
Oeoetina inornate Banks (1907a))
Range» Very widely distributed throughout tbs Korth American
eontlnent* Appears to be fairly rare only in the northwest*
Description* Ross, 1944*
Remarks» Specimens of 0. Inconaplana were collected in northwestern
Florid® and differ from specimens collected in Alachua County in sise
(northwestern Florida specimens are fifty percent larger), color
(northwestern Florida specimens are darker), and genitalia (north-

•£S8*
west®*» Florida specimens hero shorter, stouter claspers and other
minor differences).
The larra of Oeoetis sp, A is very similar to, and may
easily be confused with, the larra of <3* inconspicua*
Previous Florida Reoords* Oaoetia inoansplcua is dted by Ross
(1944) as occurring in Florida, but I do not know whether Boas had
specimens of Oeoetis sp, A or 0. inconsptcua before him at the time#
Specimens Ssaminod* Twenty*foar males and females, Ebro, Washington
Co*, L. B„ 5/30/40.
Oeoetis detona Boss (1947a)
Range* Florida*
Description* Ross, 1947a*
Remarks* Oeoetis daytona appears to be rare in Alachua County, for
I hare only one specimen in ay collection* The larva of this species
is unknown and may be either Oeoetis sp, B or sp, C.
Previous Florida Records* Three males, 3 females, Daytona Beach, at
light, collected by 0. T. Riegel, ?/g7/45> 7 males, 12 females, Day*
tona Beaoh, at light, collected by 0, T, Riegel, 8/27/45? 1 male,
Daytona Beach, at light, collected by 0, T, Riegel, 7/31/45. These
records were taken from Ross (1947a),

Specimens Bxamlnedt One male» Devil's Millhopper, at light» M. W*
N*, 8/a/so.
Oeoetls ostenl Milne
Ran get The range of this species forms a band through the North*
east from Minnesota eastward to Rem Brunswick* also collected in
Florida* Oeoetls ostenl had not previously been reported from Florida,
therefore, establishing this locality record indicates that it can
be found in the intervening territory*
Description» Rose, 1944*
Remarks» The larva of 0* ostenl Is unknown* however, further study
may show a relationship between Oeoetls sp* B or sp* C and this
apeóles*
Ross (1944), in Illinois, collected the adults mainly around
glacial lakes but siso found them on the shores of rivers*
These caddis flies are common in Alaohua County and were
found associated with many habitats such as the River Styx, Eewnan’s
Lake, the city of Gainesville, Biven * a Arm, Devil’s Millhopper, Poe
Springs, Pond B, and Big Hatchet Creek* Ho clue as to the habitat
of the larva oen be deduced from these records.
Previous Florida Reoords» Betten (personal communication) collected

-140-
0, oatenl in Florida but did not specify the locality.
Specimens Examined* Two malea» 2 females* River Styx» at light* M*
W. 8„ 7/l?/601 1 male* Hewnen*s Lake, at light* M* W. N„ 6/23/60»
1 female* Gainesville* at light, M, W, H** s/o/sOj 1 female, Biven’s
Arm* at light, M, W. K,, 6/24/60» 6 females, Gainesville* at light,
M. W. I.» 6/3/501 1 female* Devil*s illllhopper, at light, U* W, II»,
8/a/S0i 1 female, Poe Springs* at light, M. W, IT,* 9/9/5O1 1 male*
1 female, Gainesville, at light, M, W. H„ 6/s/sii 1 male, 1 female,
Ebro, Washington Go,, L. B., 6/3/40» 1 female» Gainesville, at light,
M, W. N„ 5/9/SO1 1 female, Hewnan1* Lake, at light, M, W. 15., 10/2/SO*
7 females. Pond B, at light* M* W, H., 7A0/SO1 6 females, Big
Hatchet Creek, at light, M. W. II,, 6/12/SO 1 23 males and females*
Gainesville, at light, If, W, H„ 6/4/eO.
. ; .;1* j;
■ ■ J Tii;.* •• -
Oecetls avara (Banks)
(Setodes avara Banks, 1895)
r-<
Range» Most of the United States, southern Canada end Mexico,
Description* Ross, 1944,
Remarks* Oeoetie avara was collected only in the Senta Fe River at
Poe Springs in Alachua County where it thrives* Two distinct color
pattern phases were found at this looality* one, brown with many dark
spots on the wings, end the other, lighter, with no spots on the wings

•441.
whatsoever* Here spotted Individuals were found than unspotted ones*
Hobs (1944) suggests that an unspotted apeóles which occurs in Cali»
fornla* 0» dls.luneta» my simply be a color variant of O* avara*
The color variant at Poe Springs* however * did not agree with the
description of 0* disjunct»*
The min emergence of 0# avara probably occurs during the
early part of the summer*
Previous Florida Records» This species has not previously been re¬
ported from Florida*
Specimens Examined* Two-hundred males end females (spotted)* Poe
Springs* at light* M* W. H** 7/lZ/SOt 12 males and females (unspotted)*
Poe Springs* at light* M* W* IT.* V/lz/fjiOt S males* 3 females (spotted)*
Poe Springs* at light* H* W# IT** 9/9/SOi 2 wafer, 8 females, Poe
Springs* at light, H. W. IT., 7/s/si.
Oecetis georgia Boss
Ranger Georgia and Florida*
Description* Ross, 1941*
Remarks» Oecetis sp* B or sp* C may be the undeaoribed larva of
Oecetis goorgia* 1 have collected the undescribed females of this
spades in addition to the males

-142-
lothing is known regarding the haMta end habitats of 0*
gtorgia# .. . â–  â– 
Previous Florida Records* Hone#
Specimens Examinedt One male# Gainesville# at light# M# W# N##
6/23/SOI 1 male# 2 females, Poe Springs# at light# M# W# N#,
4 females# Hogtosn Creek, at light, W# W# ÍU# 6A9/S0j % female,
Hogtown Creek, at light, M# W. R#, 7/l/SO#
Oeoetis olnerascena (Hagen)
(Setodes ciñerascana Hagen, 1861
deeetliaW fumosa Banks, 1899)
Ron got Widely di stri bated throughout the eastern# ecu them and
central states.
Description* Ross# 1944.
Remarks» fhe log-cabin 111» case of Q# olnerasoens is often seen
in many types of habitats scattered throughout Alachua County1 for
example# it has been found in the Santa Fe River, Hogtown Creek, a
sink-hole pend, Rake «Tauberg and Rake Alice# fhe cases are hard to
distinguish, for they are often fastened to submerged logs, and they
blend in trail with their background#
fhe adult# can usually be separated from the other species
of Oeoetis at a glance, for 0# olnerasoens overages 11 to 13 milll-

.143.
netor« in length end is the largest species in this genus -which
occurs in Alachua County#
Previous Florlán Records* There is no published record of this
species being found in Florida* Bet ten (personal communication)
collected 0* cinerasoons in the state but did not specify the loca,
tioru
Specimens Examinedt One lares, Santa Fe River# 3A2/38* 4 pupae*
Johnson Lake* Golden Read State Park* Clay Co., M, W# it*# 9/lz/SOt
7 males# 4 females# River Styx* at light# M# W* It## 7A7/60* 1 larva*
Lake Alice* M# W* H„, Bfio/Sli 23 males end females* St. John’s River,
at light, J. B*, 7A6/50} 1 male, Blven’s Arm, at light# M, W. I*,
6/24/60* 1 male* Blue Springs Creek* Jackson Co,# L* B„, 6/6/40?
1'larva* Lake Alice* II* M*# 4/7/40*- 1 female# Devil’s Mlllhopper*
at light, M. ¥* !!., 8/h/80? 2 females, Pond B* at light* M. W, H«,
7Ao/601 Z larvae* Hogtown Creek, M. W. II,, 8/29/SO? 2 larvae,
sink»hole pond* Alachua Co*, A* H* L* s/26/S6$ 1 larva# Poe Springs#
6/a/47i 2 larvae* Santa Fe River# J# S, R., 8/21/34? 18 larvae,
Lake Wauberg, R* P* T*, no date? 1 female* Gainesville# at light*
M, W, II*, 6/25/60* 2 males#'1 female* Poe Springs* at light* It, W*
1.* 7/3/81? 1 female* Worthington Springs# at light, I.!* W, IT*, 7/lz/mt
1 nale* Gainesville# at light* I!. W. ÍÍ., 2/20/6O,

-144-
Oooetls parra (Banka)
(Setodina parra Sanies, 1907a)
Ranget Florida* '
Peserlptlent Roas, 1958a*
Remarkst This rara caddie fly was described on the basis of three
Specimens fresa Kissimmee in 1907* fo my knowledge, beside the two
specimens t collected, it has net been taken since*
nothing is known regarding the habits or larra of this species*
Previous Florida Records» Three specimens, Kissimmee, November. (from)
Bunks, 1907a*
Specimens Examined* One male, Poe Springs, at light, H* 1.7* R«,
9/VsOj 1 male, Gainesville, at light, H* W* !!., 0/4/6O*
Oecctle sp. B
This undescrlbed larva (figs* 182 and 186) 1ms not yet been
associated with the adult stage* It most probably will turn out to
bo the undesorited larva of either 0* deytona, £* oaten1, 0# goorgia*
or £♦ parva» Mr* Bills Xanquist collected two larvae in a sand-hill
lake in Bevy County, but aside from this, its habitat remains an .
enigma*

-145-
FIG. 162-m.——Oooetia ap, 3,
LARVAL case.
Spoolraena Examined* our larvae, Alaohua Co., T¡T. M. B., 1940 » 2
larvae, Gospel Pond, 4* ralles W. of Archer, Levy Co., E. L., 4/4/50.

-146-
Ocoetls so* C
FXO. 183—Oecetis gn* C.
vmr.—
This undescribed lan?a (figs. 183 and 187) also has not been
associated with the adult stage. The larval case sometimes has large
sand grains fastened to the sides giving it the appearance of having
lateral flanges. Oeoetia sp. C most probably will turn out to be the
undescribed larva of either 0. daytona, 0* oateni, £. gaorgia, or £.
parva.


443-
Speolmena Examined* Six lareae, no datas 1 larra* Gospel Pond,
4| alies W* of Archer, Lory Co.* B. L.» 4/4/SOi 8 larra®. Big Hatohet
Creek, bottom sample, M. W.-N., lo/s/so*
TRIA3N0D3S McLachlan
Key go The Species Of friaenodes
Larras
1, Seed as in fig. ease as in fig. 200.
friaenodes sp* A
Head as In fig* 20Zi ease es in fig* 200.
friaenodes sp* B
Adults
1* Male genitalia as in fig# 203#
friaenodes perna
Male genitalia as in fig* 204.
friaenodes florida â–  ;:r
i
friaenodes florida Ross '
Ranget Florida*
Description* Ross, 1941*
Remarks! friaenodes florida was described by Ross on the basis of
fire males sent to Mm by Dr* Lewis Berner. 1 haw collected the


/
•ISO'
ll
undoscribed female of this speciesi and I believe that the larva
will prove t^be either Triacnodcs ep# A or sp* D*
■.// : • .
Thp ad/ilts are rather striking in appearance, for they are
./ â–  / /; .
largo cacáis flies* golden tan In color, and they have extremely long
/ . • '/1.. .
antennae*
7 - ‘ : .
Previous Florida Records» Two malos* Gainesville* collected by Lewie
Berner, 4/2/3$? 3 males* Sbro, Vie shin gt cea Go., collected by Lewis
Borncr* s/gb/40* This record is from Sosa (1941)*
Speelmene^jsjwBdned* One female, Big Hatchet Creek, at light* M* if*
N.* 7/ís/ét 1 male* 1 female, Gainesville* at light* M* W# II#,
/ / ' "
5/3/CO5, 2 incles* 1 female* Gainesville* at light* H* W, K** 6/?/S0?
/'I ,,
19 malts 'and females* Gainesville, at light, M* W# IT», 4/3/50? 1 male#
Cainesviilo, at light, M* W* II#, 6/9/oli 17 females* Poo Springs* at
lig/t, i|* W* Tí,, 7/lg/sO? 3 males, 2 females* Gainesville, at light,
M*M* H** 6/2/50? 1 male» 2 females* Poe Springs* at light* K* W*
: //
//
Tif, 9/9/50*
//
/
i • i
7/
! 1
; i
; /
i
Trinen odes pema Ross
/ /
/ / Range? The only records of T, perna are 1 male and 2 females
/ f
I / from Illinois* 1 male from Ohio and 1 female from Oklahoma* This
/ !
I / spooies, as indicated by its occurrence in Florida* has a much more
Widespread distribution ever the eastern part of the United States*

-2 El-
Further collecting In the Intervening territory will undoubtedly
' f â– 
fill in the gaps In our knowledge regarding its distribution*
Description* Ross, 1944*
Remarks* ilj believe the larva of either Triaonodoe sp* A or op* B
is actually the undesoribed larva of f* porga»
; ’
7 Tfiaenodes perna appears to be rare in Alachua County,
/ . • . : •’ , •' * ■ ■ ■
for only two specimens were collected*
- .* • . • : '■ i
l
Previous Florida Records* Rone*'
Specimens Examlnedt Two males, Poe Springs, at light, M* W, I*,
7A2/6O.
Trinenodes sp* A and sp* B
7 ,' 1 • *
These larvae {figs# 201 end 202) arc undoubtedly the un-
/ / described larva© of T* perna and T* florido.i for I have records of
/ / ~ ~
only two speoies of this genus occurring in Florida, both species
/ have undosoribed larvae# end I collected these two undesoribed
?
species of larvae* Further collecting and rearing experiments are
necessary to show the true affinities of those larvae. Pupal dis¬
section cannot be used to associate the stages of Triaenodes, for the
larval sclerites of this genus, along with the other genera of
Leptoceridao, are ejected from the pupal case.

-asvo mflQ.
•y »ds W5I88WO-Í7?——902 *0M

453-
Soeeímcns Bxan&nedt
sp* A——One larva, Swim Pond* Marion Co*, 2/2C/37? 2
larva®, stream zk milco of Gainesville, L, B,,
S/lQ/mt 1 larva* Hogtown Greek, M, IT, H.f ó/w/sO*
sp, *-0ne larva* Santa Fe River* !!• T», !?,, s/gs/si*
SEtOPE8 Rambur
Setodes floridana Banks
This species was described by Banks (1905b) on the basis of
one specimen collected near Biscayno Bay* It has not, to my know-*
ledge,boon collected since.
BRACHYCEHTRIDAB
IHCRftSEam KcLaofalnn
Hloraeoaa sp, A
Many Ãœlcrnsom larvae were collected at the rook dam near
Poe Springs? however, as these larvae do not fit the description of
any known forms, I am designating By specimens as Ml or asoma op, A
(fige, 205 and 206),

•164-
PIG* 206—*—moraran* *p#
HEM) OFMEtfA* , - ' c ” ’
PIG* 208—Braohyoentrua sp* A,
HBAD OF“LARVA.

FIG. 207—.
Larval case
»Braohyo»ntrua ap. A,

-ise-
The adulta may he negatively phototropic, for a«e cane to
light, A single female specimen whioh could not he specifically
Identified was taken by sweeping, â– 
The larva builds a highly characteristic, cylindrical case
which is only found in the swiftest of currents, I tried to rear
this species in the laboratory but failed each time, for it was
rather difficult to simulate the natural conditions necessary for
the continued existence of this caddis fly#
Specimens Examined* Thirteen larvae, Pos Springs* A* M* I<«, z/isfahi
2 larvae, Poe Springe, 5/5/l?j 1 larva, Poe Springs, s/t2/S8j 1 pupa,
Poe Springs, M, W, H*, 2/24/SOj ,1 female, Poe Springs, sweeping, M,
V», II,, s/24/S0j 4 larvae, Santa Fe River, L, 3,, 2/ll/S9; 11 larvae
and pupae, Poe Springs, M, W, II,, 3/24/50; 2 larvae, Poe Springs,
li, Ví, 1,, ?/i9/61; S larvae, Poe Springs, L. B,, 3/L/4OJ 2 larvae,
ravine stream, Golden Head State Park, Clay Co*, M, Vi. n#, s/¡L2/S0*
BP-ACHYCgBTRISS Curtis
Srachycentres sp, A
A V â–  â–  . '
This undescribed larva was collected in northwest Florida
by Dr, Lewis Berner» however, no adult of this genus has as yet been
collected in the state* Further collecting in the northwestern
portion of the state should solve the problem of the relationships

of the larva*
Speoírapas Examinad* fno larvae* 5*1 ralloe W* of the Walton Co* line*
Okaloosa Co** 1* B*| 6^/51/40#
SmCGSTCHftSIDJJB
serioosycam Berthoia
Sari costorm crassicorals ha's cot boon collected in Alachua
County* but it does occur in tha northwestern part of the state* for
one apeoiraen was collected in forreya State Pari; by Dr* lewis Berner*
the larva of this species is unlmorai*
Previoua Florida Recordst Hone*
Specimens Examined* One male* lorraya State Parle* L* B** oAo/33*
HELICOPSfCEIDAS
HELICOPSYCHE Hagen

-ass-
Helloopsyche ’borealis (Hagen)
(Notidobla borealis Hagen» 1861
Hell copsyohe caYlFornioa Banks* 1899.
Hell oops:?che ^mtíiooKaiV ‘ Banks,; 19043)
Range» ’The continental range of this species is very wide, stretch¬
ing frost Mexico northeastward to Hove. Scotia and westward to Montana
and Oregon? the range embraces most of the forested areas of the
continent, forming a complete circle around the Great Plains*1* (Ross,
1944, p* 2(57)

'
Remarlest The curious snail-like cr,so of H * borealis (fig* 310) is
a unique feature of cool* fast-flowing, calcareous waters such as
is found at the rock dam near Poe Springs. Beth larca© and adults
(figs* SI end 32) are extremely common at this locality and 'can be
collected by thousands* The larva© ore encrusted m the mats of
Vegetation which cling to submerged rooks*
forhies (1909) reared and described all stages very com-
pletely* I was also able to roar this species.
The adults are attracted to light* and literally thousands
of specimens were collected by lighting near the rocíe dam for a
few hours. The species is so abundant at this locality that a
collector* working at a light, can continuously brush tho insects
Into an open jar of alcohol.
Emergence takes placo throughout the year as is indicated
by the collecting records*
Previoua Florida Records* Ross (1944) lists this species as oc¬
curring in florida hut does not specify the locality*
Specimens Exaninodt fwo-thousend mies «aid females* Poe Springs* at,
light, II. W. ?/lg/S0; S larvae* Poo Springs, J, S. R„ s/2l/34j "
6 mal.es, 9 fenoles» Poe Springs, sweeping, H* W* H,* s/m/so? 16
larvae, Poe Springs, I!. V/, II*, g/24/ñO; 1 larva, Santa Fo Elver, I».
B3 A/40; 1 male* Poe Springs* sweeping, M* W* IT., c/sl/SOf 1 female,
Hogtown Creek, at light, M* ¥* H», 7/l/sói 14 larvae, stream, *1

-160-
mile S, of Gulf Hammock, Lety Co,, H* W* B,, 7/so/sO; 1 pupa, Poe
Springs, M* W, H*, G/zi/SOf 400 males and females, Poe Springs,
at light, M, W, II», 9/9/sOi 5000 males end females, Poe Springs,
at light, M* Vi, 1 female, Worthington Springs, at
light, M. W, 11*, 7/45/60*

tSTSOCTSD PEOPLE'S
Extending this investigation to include the caddis flies of
Florida broadened the scope of the study considerably but also eon-
plicated the picture in many ways! for while enough specimens were
examined to visualize the problems entailed, not enough material was
available to solve these problems. In general, for every problem
solved, two problems were raised which are still unanswered. 1 be-
lleve that this study is as stimulatory in nature as it is definitive
because of these unsolved problems.
The problems are outlined here, for each one is more thoroughly
elaborated in the discussion of the species contained in the annotated
list.
1. The adults of Chi marra soda and Chi marra -perieua are extremely
similar, and furthermore, 1 collected the tmdescribed larva of
C. nerlgua and eould not distinguish it from 0. soda using the
description given by Boss in 1944. The two species had been
called soda up to 1948, when Ross first described nerlaua. Boss
(1948) states, BRe-examination of a large series of soda ———
(indicates) a wide eastern end northeastern ragge for soda and a
small range for nerieua which is peripheral on the southern and
western portions of the range of soda.”
A critical examination of both larvae and adults is definitely
in order

.162.
2. Polyttentrontts sp. A, which has a larra greatly.resembling the
larva of P. reap test. its unassociated axel may possibly he the un-
described larva-of P. cmssicornis. Adults of P. renotus have
never, been taken in Florida, but ray nevertheless occur in the
state.
3, flhe larva of Psyehosgriid Penns A Is unassociated with an adult
fora, fhe larvae of the entire genus Cernotina and the larva
of Uyctioxh.vlsn vest!tug are unknown and ray actually be the
adult form of the unassociated larva of Fsyehomyild Penns A»
h, fl%e larvae of the following species are still unknown. Before
our knowledge of the caddis fly fauna, of Florida Can be put on a
firm basis, it is important that they be discovered and described.
Ky&ropnyche sp. A, Oxyethira florida, glass. JfiBSÜB» lSS2i&* Z££M»
waiter!. Orthotrichla instabllis. Tlyuroutila weknlla. Pent rich! a
mea» heptocella, enUaaa. trvara, Athjijjpcj.es floridanus, gjLo8ggaa&,
Oecetir, daytonu, osteni, georgia. ram. griaenoAes perna. florida
and SeMcostoma crasslcorals.
5. ghe following are undescribed larvae} however, no adults which
could be associated with them have been collected in Florida.
Hvdrousvche sp. A, Pycnonsyche sp. A, Mierasema sp. A and BraChy»
gaateai »*>•' a.
6. hentocella, sp. A is an uadtscribed larva, and it is extremely
probable that this is the undescribed larva of either L, snlloaa

«3t62k
or I„ tavara because these are the only species of this genus which
occur in Florida .'and which have undescrihed larvae.:
7. Tvm species of the emus Athrinsodes.- 4. floriflanus ana A. slossonae.
occur in Florida* and both have undescrihed larvae* I have examined
two undescrihed species of larvae, Athrinsodes st>. A end Athrity
sodes sp. S, whiehalso occur in Florida. The problem is, which
-larva belongs to which species.
8. I have two undescrihed larvae, Qecetis sp. and Qecetis sp. 0,
which have not been associated with their adult stages, There
are four species of this genus which occur in Florida and 'which
have undescrihed larvae,
9. There are two undescrihed larvae of the genus Triaeno&«3. Triae-
nodes sp. A and Triaenodes sp, 3, which occur in Florida, and there
are two species of this genus* T. nema and T. florida, which occur
in Florida and have undescrihed larvae.
10. What are the physiological or genetic differences between the
spotted and unspotted forms of Qecetis avara?
11. The Qecetis inconsuictta-~»Oocctis sp, A complex is an intriguing
problem. In brief, two species* uorteri end nratelia may be sya-
onomised to inconsulcua. and the species which now bears the ñas»
ninconst?lcuan may be synonomized to the next name having precedence,
flaveolatfe. See the annotated list for a sacre extended discussion
of this problem.

>164»
StJKHAKT PXW COTCMJSIONS ,
1. Caddis flies, both adults and larvas, were collected throughout
Alachua County and were Identified.
2. Specimens collected by Drs, Berner, Young, Sogers and others fro®
various parts of Florida were examined and identified.
3. fhe literature was searched for all references dealing with caddis
files found in Florida, píese records were incorporated in the
discussion of the species.
4. Bearing techniques were experimented with,- end the new techniques
which were developed are described in detail.
5. Many state and regional geographic records ere recorded. These
records extend the range of some caddis flies by over one thousand
miles and add greatly to the knowledge of the range of many other
species. A number of very rare caddis flies were collected.
é, Ecological notes are included in the discussion of the individual
species.
?. Sixty-three species of caddis flies were found to occur in florida.
Sixteen undescribed larvae were found and these were associated,
with the adults whenever possible. She undescribed females of
five species were also found.

-3L6S-
8. Kony highly interesting taxonomic problems were brought to light,
9* A key with illustrations was compiled for the identification of
all the caddis fly larvae and adults known to occur in Florida,
10* On the basis of information gathered during the course of this
investigation, Florida can be divided into Wo soogeographic
«reas having fairly distinct caddis fly faunast (l) northwestern
Florida which contains caddis flies having northern affinities,
sad (2) peninsular Florida which contains twelve endemic species,
whose other caddis flies have southeastern affinities*
11, Evidence is presented which would indicate that caddis flies are
hardly as important in the aquatic food chain in Florida {except
in scattered localities) as they are in the 'North.
12, The fundamental importance of,caddis flies in Florida, has to a
large extent, been overlooked, They appear to' be highly sensitive
indicators of stream, pollution and can be used to show coscarative
degress of pollution.

«366*
lra?3CTJ3®3
Banka, laths» ■
1895» Hew neuropteroid insects. - Am,-Sat* Soc. 'Prang. 22!
313-6.
1897. Hew forth American neuopterold insects. Art, Sit. Soc.
Trans. 24»21-3l.
1899. Descriptions of new north American neuropteroid in-
<â– 
sects. An. Sit, Soc. Trans. 25»199-218.
1995b. Some new neuropteroid insects, 1?, T, Sit, Soc. Jour.
11! 236-2¿3,
190h&« A list of neuropteroid insects, exclusive of Odonata,
from the vicinity, of Washington, B. C. Wash. But. Soc.
Proc. 6(4) 1201-17.
1905a. Descriptions of new species of Hearctlc neuropteroid
insects from the Slack Mountains, IT. C, Am. Mas. fat.
Hist. Bul. 2U215-8.
1905b. Descriptions of new Hearctlc neuropteroid Insects.
Am. Bit* 3oc, Trans. 32:1-20, 2 pis,
1907a* Descriptions of new Trichoptera. Wash. Bit. Soc. Bros.
8(3-4)»II7-33, pis, 8-9.

«J6?<
Banks, ifethan.(continued)
19031), Heuropteroid insects - notes and descriptions. Act. lint,
Soc. ?rans. 34:.955-6?, pis. l?-9.
1909.
G\/o net/ caddiee flies. Eat. lews 20*342#
1911*
Descriptions of new species of ITorfch Anerican neurop*
teroid insects. Aa, Eat. Soe. Sirens* 37(4)*335*69»
pis. 11-3.
1914.
American Trichoptera - notes and descriptions. Can.
Ent. 46*149-56. 201-5, 252-8, 261-8, pis. 9» 10, 15, 20,
1938.
Hew native neuropteroid insects. Psyche 45(1)*72-9»
9 figs#
1943.
notes end descriptions of learctic frichoptera. Harv.
Univ, Hus. Coiap. 2ool. Dal. 92*341-69. 6 pis.
Berner, Lewis
1959.
fhe aayflies of Florida.. 1. of Florida Press 4(4)i
x-267. 88 figs., 19 naps, 24 plates.
Betten, Cornelius
1934.
Idle caddis flies of frichoptera of Hew York state.
1. Y. State Has. Bui. 292, 5?6 pp., ol text figs.,*
6? pis.

*46 8-
Batten, Cornelius and 'Martin 2. Mosely
1940. The Irancis Walker types of Trichoptera in the British
Museum. 248 pp., 122 figs, London.
Carpenter, V. tt.!
1933. Trichoptera from the mountains of llorth.Carolina and
Tennessee. Psyche 40:32-47, figs. I**!?*
Denning, Donald 0.
1937* The biology of son® Minnesota Trichoptera. Am. Sat.
Soc. Trans. 63:17-43. 45 figs.
1943. The Kydropsychi&ae of Minnesota (Trichoptera.). Ent,
Am. 23*101-71. 41 figs.
1947. Hydroptli&ae from southern United States. Can. Ent.
79:12-20. 10 figs.
1948a. Hew species of Trichoptera, Sot. Soc. Am. Ann* 4li
397-401. 6 figs.
19484. Descriptions of eight new species of Trichoptera.
Brooklyn Ent. Soc. Boll. 43(4):119-129* 11 figs.
Elkins, Winston A*
1936. The immature. stages of soma Minnesota Trichoptera.
Ent. Soc. Am. Ann. 29:656-81, 6 pis.

-ae9-
Frison, Theodore H.
1935. The stoneflies,.or Plecoptera, of Illinois, Ill, Mat.
Hist. Surv. Hal. 20:281-471. 343 figs.
Hagen, Herman A.
I86I. Synopsis of the Henroptera of Horth America, with a
list of the South American species* , Smithsn» Inst.
Mise, Collect. 347 pp. Trichoptera, pp, 249-98, 328-9*
Lloyd, John Tilomas
1915h. Wood-boring Trichoptera. Psyche 22:17-21, pi. 2, figs,
1-20.
1921. The biology of Horth American caddis fly larvae. Lloyd
Library of Botany, Pharmacy and Materia, Medica Bul.
21:1-124, figo. 1-197*
Mickel, Clarence I!., and Herbert S. Milliron
1939* Bearing the caddis fly, Tdrmcnhllus indivisas Walker,
and its hymenopterous parasite. Healteles biannalrtna
Grav. Snt. Soc. Am. Ann, 32(3)*575-80.
Milne, Loras J*,
1934-38* Studies in Horth American Trichoptera. Pt. 1,
1934: 1-19. Pt. 2, 19358 20-55* Pt. 3, 1936:58-128,
with 2 pis, Cambridge, Mass.

-170-
' Milne,. Margery J., and Loras J. Milas
1939* Evolutionary, trends la caddis worm case, construction.
Bat.'Soc. An. Ann. 32(3)*533-^2. 1 fig.
Reid, Ceorge IÍ.
1950» Food of the ‘black crappie, Pomoxjs nlgro-maculatus
(ReSeuer). in Orange Lake, Florida, Am. Fish. Soc.
' Trans. 79*145-154. 3 figs. ' ' '
Ross» Herbert,H,
1938a. Descriptions of Hearctic caddis flies (Trichoptera)
with special reference to the Illinois species. Ill.
Hat. Hist. Surv. Sul. 21(4)¡101-83. 123 figs.
1938h« Descriptions of new leptocerid Trichoptera* Eat, Soc.
Am, Ann. 31(l)J88-91. 8 figs*
1938c. Rectotypes of Horth American caddis flies in the Musetas
of Comparative Zoology. Psyche 45(1)*1-61. 10 pis.
1938d. Descriptions of new Forth American Trichoptera. Hash.
Ent. Soc. Proc. 40(5)*117-24. 2 pis.
1941b. Descriptions and records of Forth American Trichoptera.
Am. Bat. Soc. Trans. 67(1084)*35-126. 13 pis.
1944. The caddis flies, or Trichoptera, of Illinois. Bull.
111. Hat. Hist. Surv. 23(1)*1-326. 961 figs.

-171-
19‘?'7a. Descriptions ?n& records of Morth American Trlchop-
tera» 'With synoptic notes. Ara, SSit. Soc. Gratis.
73(2)s125-168,
19^7b. Motes and descriptions of líearctic Ky&roptilldee
(Trichoptera). Mash, Ent, Soc. dour, 38(6)J201-206.
6 figs.
X9**8. Bes* JTearotle.Rhyncophilláae and. Fhilopotarldae. Bit*
Soc, An. Ann. ^1:17-26. 8 figo.
Sibley, Charles K.
1926h. Trlchoptera, in A. preliminary biological survey of the
Lloyd-Oornell Reservation, lloyd library of Botany,
Pharnacy and Materia Heftlca Bal. 27(Ent. series $)i
102-8, 185-221, pis, 8-I3.
Vorhien, Charles T.
1909. Studies on the Trlchoptera of Wisconsin* Mis. Ac.?d
del. Trans. 16:6^7-738, pis. 52-51.

â– 172-
CHECK LIST OP THE CADDIS PLIES OP FLORIDA AND INDEX
1» Philopotamldae*,... •.•••••••*•••••••••••••••••••?• 29
Chlmarra»**********»*•*•**»•**••«*♦*»••»»«*•*#♦#******* 29
Chlmarra florida Roas»»«*,.«••••**.*..«.SS A
Chlmarra aterrlma Hagen*.•••» * 30
Chlmarra perlgua Rose........♦........•♦31
2* ' Payohofflyildaa . ••.•...•••.•••••••••34
87
Neuraellpela crepttaenlarla (Walloer) ***â– *#57
Phylocentropua. •.**38
Phylooentropua plaoidua (BanRe)**«*.«**»58
•1 ' • : • ; ^ . S ■■ , . s , t
Polyoentronua.......................... »....*•*41
Poiycentropua craasioornla Walker*...•♦•41
Polyoentropua ap« A*...*.............•••41
Kyctiophylas•»•••«»•«••...*.•••.«••.•**••••***••*••••••42
Kyotlophylax yeatltua (Hagan)*42
Cwnotina ••••••43
Carnotlaa oaloea Rosa**,••**•«»«••«•••,«46
Cernotlna tnmcona Rose.«*..♦♦.»♦•«...**47
Cernotlna spioata Ross *•••••••••••*••*•*43
Psyohoraylid Oentia A* •49
Lyp»*.... *50
Lypc diversa (Banka) ••••SO
Payohomyla*.••••... 51
Psyohomyia flavlda Hagen., ••••62

Cymollpa».
64
-173-
Cymellns marginalia (Banks) *##***#***#♦ 54
3» Hydropsychldao•.««••»*«•••••••••••«••«•>••••««..••«••««*57
Diplootrona..... ...60
Plpleotrona modesta Banks..............«60
Hydropsyohe............................................92
Hydropsy oho inoomoda Ha-on. •«.•••..«•••68
Hydropsy oho ap. A.. .....65
Choottt&t opsy oho»••••••••••«.•••••••«•••.•••*•>**»«•••«.•66
Choumatopsyohe harks! Hoss.68
Choumatopeyohe emails (Bonks)•«••••••••«70
Chsumatopaycho aphanta Ross..71
ChsMiaatopayoho plnaoa Boss.*»..... 78
Maoronemuia..... .....73
Maoronommn Carolina (Banks)........••»..73
4* Ilydroptllldao...............• •»*••••••».••***•«*•.«...*.75
Ogyethlra 78
OatyotHLra .janella Dennlnp;...............80
Qagathlra florida Panning...............81
' foyerthira glass Ross...........**.••••••88
Chtyethira Itnaosa loss.#*................83
Oxysthlra voma Ross*. *».».»».#...»*....83
Oaygthlra «alter! Panning............***84
Oythotrichla.».»»•»*.»»•»«•*»••••«•»•»*•»*»•••*»•*•*.•.86
' ¡ Ofthotrichla americana Banka.»*»*.»...«*87

-174-
• ''" OrfehOtflehta' orí g t ata Mor toa «♦. « 88
Orthotrlchla tastabille Penning»....«..*90
Hydroptlla» 90
. Hyd»Ofttíla bernorl Ross....««91
Hydroptila ttakolla Denning., #***;#*#** ...92
Heotriohla»9 S
ffeotriohla ranea Denning................95
Mayatrlohla ......95
Mayatrlohla ay ama loeely «.***».*»#.« »# •* .95
5. Fhryganeidae*...........................................96
Banksiola# ...**•♦*•***••*»«•»..*»»»•»•«».. *96
Banks!ola conoatanata (Walker)........«*96
6. Lliaaephilidap.«.»*** #96
Py on opsy ohe »...... .*.**.**96
Pyeoopsyohe sp* A*....................#*98
7* Molannldae*99
Hoianna*. .«...*..*99
Holanna tryphena Betten*#.**«.***«..#.#*99
8* Leptoeeridae* .•.•••.•••••.•••*•••101
Leptoeams.... ..IOS
Leptooerus asierloanns (Bank»}* #*» ♦ **■##* 108
Leptooella...... 108
Leptooella oandida (Hegen)#*...***..«**112
Leptooella exquisita (Walker) .118
Leptooella navida (Hagen) *#*#♦•***##♦# *114
Leptooella splloma Eosg..,............*115

-175-
loptocolla alMda (ffidice?)»«»■<«
tepfrooella tacara Boas*.*117
. . . ttptootllft sp. A*. *•*»*••»*.».»#»»*.* * «117
IthripaodM* »#**#**«#*»#4«* #**••»*«**« ******♦•««»#•**.* 119
Athripaodes florldanpa (Banka).....«..*121
Atkylpsedea alosgonae Bimka*******.....lgS
Athripaodes oanoollatua (Betten)«.....*123
Athyipaodes tsygi«punctatt»8 (Vorhlee) «*124
Athy i pa oda a sp# A»» ... ...125
Athyjpaodea sp* B,.126
Oeeetia*» ......... ..••«123
Oeeetia ap* A* .135
Oeoetia lnoon3pictm(Walker 13?
Ossetia daytona Ros8»»«*...»..m.»»»..*183
Oeoatis oaten! Milne..»».........*..*.*159
Oeoatis avara (Banka)......... *140
Oseetls geoygla Rose...................141
Oeeetia clnerascena (Hagen) *•••««»««**,142
Oeoatis parra (Banka)............ *144
Oeeetls sp* B*«•••••...•«••144
Oeoatis sp. C*.**#*««...»•••«...146
Tylaenodea .148
Tylaenodas florida Bogs* *•***.» **»• 148
frittenodes pema Ross. **#****.*»#•*•*»* *100
Trlaenodea ap* A..151
Trinenodes sp. B» ....................•«151

*176*
Sg|bod©8**»»« ••••»**•«« * *4** mimimi . »>15S
Set odes florlde.BR Beaks*.• .»•**•<«•* ***15§
9» Brachyoentridae...***.* ..........................«153
lücrai!C!iR>...t..»..« 153
Id orarea» ep* A*«*«*.**»*16S
Braehyoentrus* .«.........«**156
Braehy cen tras sp* A««< ....••156
10* Serioostoraatldae•**•*•• ..... * .,,.*157
Serlooetonia. >n»»« •**•«•*.*»***»• **•*•»,•*•«* *««»»•» *.*157
Serloostoma crassloomlB (Walker)....*.157
11* Hellcopsyehida© 1S7
Hellcopsyche... .157
Helleopayche boroalls (Hagen)*.............. ...*»108

477.
BIOGRAPHICAL HO®
Marshall Hirehberg vas horn In Haw York, Hew York, on
April 10, 1927. He moved to Orlando, Florida in 1939 snd has lived
there ever since. He attended Junior and senior high schools in
Orlando sad graduated from hi$3 school in 19*55* He matriculated at
the University of Florida in 19*55» and received the Bachelor of Science
degree with a major in Biolog? in 19*56. He worked in Orlando from the
time of graduation until January, 19*59, and then entered the Biology
Department of the University of Florida to do graduate work.
He held an undergraduate assistantship in the Biology Depart¬
ment from 19*'<5 to 19*58, and has been employed by the nutrition Labora¬
tory of the University of florida since 1950*
He is a member of the Florida Entomological Society, Phi Eta
Sigma and “hi Sigma.

This thesis was prepared under the direction of the Chairman
of the candidate's Supervisory Cosnittee and has been approved by
all members of the Committee* It was submitted to the Graduate
Council and was approved as partial fulfilment of the requirements
for the degree of Master of Science*
February 8, 1958
SBFBOTTSCBT COMMIffBH*
Chairman

6/12/08 10:59 AM
‘ . * -
UP Libraries: Digital Dissertation Project
Dear Dr. Marshall Nirenberg,
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