A revision of the Tyroglyphidæ of the United States

Material Information

A revision of the Tyroglyphidæ of the United States
Series Title:
Technical series / U.S. Dept. of Agriculture. Bureau of Entomology ;
Banks, Nathan, b. 1868
Place of Publication:
Washington, [D.C.]
Govt. Printing Office
Publication Date:
Physical Description:
34 p., 6 pl. : ill. ; 23 cm.


Subjects / Keywords:
Tyroglyphidae -- Classification
Mites -- Classification ( lcsh )
federal government publication ( marcgt )
non-fiction ( marcgt )


Includes bibliographical references (p. 26-27) and index.
Statement of Responsibility:
by Nathan Banks.

Record Information

Source Institution:
University of Florida
Rights Management:
This item is a work of the U.S. federal government and not subject to copyright pursuant to 17 U.S.C. §105.
Resource Identifier:
029623996 ( ALEPH )
62123778 ( OCLC )
QL461 .T39 no.13 1906 ( lcc )
632 ( ddc )

Full Text


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'18g!RD NovEA11! 14, 1906.
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L. 0. HOWARD, Entomologist and Chief of Buftii w,
C. L. MARLATT, Entomologid and Acting Chief in aWeti&
R. S._CLIvrou Odef Ch!rk.

F. H. CHITTENDEN', in charge pf bree4ing experiment.
A. D. HOPKINS, in charge of forest insed investigations.
W, D. HuAm, in charge bf cothm boll weevil in'Ve8h.qations.
F. M. WEBmm, in charge of cereal andforage-plant insect investigations.,
A. L. QVAINT ANCE, in charge, of deciduous-fruit insect investigations..,
FRANK BE-.vToN, in charge of, apimitural in cestigations.
D. M. RoGERs, in charge of gipsy and brom-tail moth work.
A. W. MORRILL, engaged in while jYyinvestigations.
E. S. G. TiTus, AUGutiT Brscx, 07To HEIDEMANNR. P. CuuRm, G..
CAUDELi, F. D. Cou E E. R. SA&14CFR, J.,1H. BEAmE, 1. Ji C. IT*
LILIAw L. HowENmuq FREDERt c Y KNABI A artist.
MABEL COLCORD, Librariaii 10
H. E. BtmKE, W. F. FisKE, J. L. WimB, J. F. Sv4uss,, engaged in fored.invOll
W. E. HINDS, J. C. C)RAwFo.Rr), W. A. HooKER, W. W. YOTHFM, Ar.,.C.
W. A PIERCE, F. C. BisHopp, C. R. Jomss, F. C., PRATr, C.' E. S" A*
MITCHELLP WILMO.-,,, NEWELL, J. B.. GARRL-rr, G,',W.'FvY-Nx, A, W. Bulail
CUSHMAN, W. H. GILSON, engagedin cotton boll wevilinvatigations.
6 1
G. I. REENTES) W. J. PHILLIPSj C. -N. AiNSLIE, engaged in'eereal and
inves igafions.
qed in decidu6w
E.T. RaILLIPS, J. M. RANKIN, engaged -in estilgaiq6ns.
0. J. GILLISS, T. A. KXLEHIER, W. A. KzLzH*' engagedin silk inv t* ati

Al 1,

Digitized by the Internet Archive
in 2013


L. 0. HOWARD, Entomologist and Chief of Bureau.





Assistand 11tomologist.




TEas] ngton, D. C., Se-tember 10, If
SIR: I have the honor to submit herewith a manuscript et
"A Revision of the Tyroglyphidiu of the United States," prepal
Mr. Nathan Banks, assistant entomologist. Mites belonging i
family Tyroglyphid- are destructive to a variety of stored pr(
and other commodities, certain species injure living plants,
others are useful because they destroy certain injurious insects.
group is therefore one of very considerable economic import
This paper 1)y Mr. Banks should prove a valuable contribution
study of these mites and greatly facilitate the determination
different species; I therefore recommend it for publication as Tec
Series, No. 13, of this Bureau.
Acting Ohief of Bur
'cr'ctar'y of Afltica lturc.

C 0 X T 1, X TrS

Habits and economic importance ----------------------------------------- 5
Structural characteristics ------------------------------------------------ 6
Transformations ---------------------------------------------------------- 6
Previous work on species occurring in the United States ------------------- 7
Occurrence of European species in the United States ------------------------ 8
Genera of the Tyroglyphidw --------------------------------------------- 9
Table of the genera -------------------------------------------------- 9
Genus Histiostoma --------------------------------------------- ---- 10
Table of the species --------------------------------------------- 10
Genus Glyciphagus -------------------------------------------------- 12
Genus Tyroglyphus ------------------------------------------------- 13
Table of the species --------------------------------------------- 13
Genus Rhizoglyphus ------------------------------------------------- 19
Table of the species --------------------------------------------- 19
Genus Monieziella --------------------------------------------------- 23
Table of the species --------------------------------------------- 23
Genus Carpoglyphus ------------------------------------------------ 24
Genus Trichotarsus -------------------------------------------------- 25
Bibliography ----------------------------------------------------------- 26
Index ------------------------------------------------------------------ 29



]I. Fig.


IVI. F ig

1.-Glyciphagus robustus -------------------------------
2.- Glyciphagus obesus: leg I ------------------------------
3.-Glyciphagus obesus ..................................
4.-istiostoma amcriCanvm: tarsus 1..................
5.-Glyciphugus obesus: genital plate...................
6. istiostomna amenia nm : tarsus IV ....................
7.-uist iostom a a merican urn: mouth parts .................
8.- H istiostoma bi'ecipes: palpus ----------------------------
9.-Ih stiostoma gracilipes ----------------------------------
1O.- Ilistiostomn breripes: tarsus IV .......................
I I.-H istiostoma breripes: tarsus I --------------------------
12.-Histiotoma gracilipes: tarsus I .......................
13.- 11M w ieziella ion gpes -----------------------------------
14.-T!i/oqlyph us furin and tarsus IV of male .............
l5.-Monieziella angust(( .. .. ... .... .. .. ...
16.- Tyroglphus f rimv: renter of male --------------------
17.-7:iroglyphu~s .ftrinw: leg I of male ----------------------
18. Monieziella brerit-rsis .................................
1 1. nieziella breita rsis: male venter and mandible .......
20.- TroglypIUs ameriC S -----us............................
21.- Tyroglyph: us a reric(wius: genitalia ----------------------
22.- Tqrogl qph us a ,mericanw.s: tarsi I and IV ................
23.-1 Trogph us linineri: vulva ..........................
24.-TqroglYphus liidneri: tarsus IV of male ................
25.- Tyro lyphus inneri: genitalia of male ..............
26.- Troglph us term in lis: leg I .........................
27.-7 Tqrogl yph us termbu ldis: tarsus IV ..............-- -.....
28.- 7:]roglyhs term inalis: vulva ........................
29.- T-r-glyphus -eri-...............................
30.- TvlroglIIph u.: bre rleps
31.- Tyroglyph s breuceps: tarsus IV ......................
32.- Tyrogq1yph us breviceps: leg I ----------------------------
33.- -Tyrogl yphus cocchilus: vulva and anal suckers ---------
34.-T!yrogI yph us cocciphiluis: male aperture -----------------
35.- Troglyphus cocciphwil: tarsus IV-male and fema!e ....
36.- Troglypus h eteromorphus: tarsus IV
37.-7yroglyphus heteromorphus: leg III of male ------------
38.-Tlroglyphus leteromorph us: tarsus I
39.- Tyroglyph s hetero moi'phus: male ....................
40.-C(arpoilyph us passularum .............................
41.-Troglyphus aruipes: tarsus IV ......................
42.-1Troglyphus tirmipes: tarsus I -----------------------
43.-ITyroglyphns a rmipes: vulva .........................
44.-Tyrogl'phws louegior: tarsi I and IV, and body hairs ....
45.-Rh izoglyphus tlrsilis: tarsus I .......................
46.-JRhizoqigyphus turuals.: tarsus I V -----------------------
47. -Rh izoglyphus hyacinthi: leg III of male -----------------
48.-MRhizoglyphus hyacinthi: tarsi I and IV -----------------
49.-Rhizogi yphus hyacinthi ................................
50.-Rh izoglyphus rhizophagus.............................
51.-Rhizoglyphus rhizophagus: tarsus I ...................
52.-Rhizoglyphus rhizophagus: tarsus IV ----------------
53.-Rhizoglyphus elongatus -------------------------------
54.-Rhizoglyphus elongatus: tarsus I .......................
55.-Rhizoglyphus longitarsis: tarsus IV ...................
56.-Rhizoglyphus rhizophagus: male venter ................
57.- Rhizoglyphus phylloxerw: tarsi IV-male and female ....
58.-Rhizoglyphus longitarsis: tarsus I ----------------------
59.-Rhizoglyphusphylloxer: tarsus I ---------------------
60. -Rhizoglyphus phylloxerx: leg III of male
61.-hizoglyphus phylloxer: half of front of body----



From an economic standpoint the Tyrois1yphida, are one of the most
important groups of mites. But owing to their small size and pale
color they have often been overlooked and the damage accredited to
some larger insect which hIappened to be present. 'By their rapidity
in breeding they make up for their minute size, so that articles, such
as flour and sugar, are often so badly infe4ed that the whole mass of
the substance appears to he in motion. It is chiefly thru their ravagwes
to stored foods that they are inmical to human elort. Dried fruits,
dried meats, and grain in mills are p I! aps most seiously affected by
them. Their frequency in cheese and sugar has won them the names
of "cheese mites" and 'sugar mites," while the disease known as
"grocer's itch" is due to their presence on the hands of persons
handling infested )roducts. A list of materials attacked by tyro-
glyphids would include cheese, flour, sugar, hanis, dried meats, hair
in furniture, mattresses, and pillows., grains in inills, cereal foods
many drugs. wine, dried fruits, seeds of all kinds, bulbs, roots of
plants, mushrooms, feathers, hay, scale-insects, pinned insects of the
entomologist's collection, and even the human corpse. Some species
are, however, of little economic interest and occur in the nests of mice,
moles, and ants, in decaying bark of trees, in sap from wounds in
trees, and a few are attached to certain insects. The species of the
genus Lonicz~dla do some good bv feeding oii scale-insects. The
"bulb mite" or "Eucharis mite, Rt oq!iypAv t ky/auc.t/ has long
been a prominent enemy to hot-house cultivation. It burrows into
the healthy tissue of bulbs and roots, thus giving entrance to destruc-
tive fungi and bacteria. This is the species infesting Bermuda lily
bulbs; and it has lately been shown that an allied species does great
damage to the roots of the vine in Europe. Another species has been
described that caused injury to the stems of carnations. Still another
R/izorlypus has been found to eat through the grafting wax on
grafted plants, bore beneath the bark, and so prevent the union of
graft and stock. The mushroom mites, both in this country and in
Europe, are prominent obstacles to successful mushroom culture.
Cellars apparently clean in the beginning of the season may be so
badly'infested by Christmas that crops are impossible.


The Tyroglyphidve are pale-colored, soft-bodied mites, devoi
trachea,; with small, appressed palpi; usually with prominent ch(
inandibles; with moderately long legs, ending in one claw, and o
a sucker or caroncle; and with a body about twice as long as br,
and broadest behind the middle. There is comnoly a disnct sux
between cephalothorax and abdomen. There are no eyes, unless
tain spots on the front margin of certain Hy popi and oil Cir pogh
represent these organs. The dorsum bears a few, usually long, h-
in size and arrangement constant for each species. The legs I
bristles. fine hairs, and sometimes distinct spines, the more promii
of which are constant in position for each species. One long br
near tip of the penultimate joint is especially prommint; an oth
a thickened or clavate hair iear base of tarsi I and II, and is prob-
a sense organ. The latter is always upon this joint, altho sev
authors have figure(l it on the penultimate joint in some species.
the venter are two apertures;: the genital is usually elongate
situate between hind coxav, and there are often U-shaped marks 4
side of it known as the genital suckers, which can be protruded.
anal opening is toward the ti) of th, l)odv. and is often but an elon
slit, with a sucking disk each side of it. In the genus Glycip/t
these openings are much larger, and the genital sometimes occu
the entire area l)etween the coxa'.
There are but slight differences in structure between the sexes;
in sonie ca. ses the nale has one pair of legs enlarged, or there ma
two forms of the male. In some pecies the male has two little suc
on the hind tarsi; in others there is a curved plumose bristle on
basal part of front legs. The males are smaller than the females,
have a less tumid abdoiiien.


The trnsformations of the Tyroglyphidw are among the most i
velous of the aninmil kingdom. All tyroglyphids lay eggs, ofte
large size, which are scattered haphazard over the infested mate
The young on hatching have six legs and at molting obtain two m
Thenceforward their life history may take the simple and direct ]
to the adult condition, but often passes through a stage called
Itpop its. This Ifypopwu is a very different creature than that f
which it developed-the octopod nymph. Its body is hard and c
nous, there is no mouth orifice, and no distinct mouth parts; the
are short and ill adapted to walking. On its ventral surface neal
tip is an area separated from the general surface and provided I


several circular marks or sucking disks. By means of these suckers
the Hypjopus attaches itself to an insect or other creature, and is trans-
ported to another locality, where it may find a suitable breeding place.
a ntelieo rog'lyphid fitted for
The Ifypopus is therefore a stage in the life of a t n
migration. The t!y pte,, upon reaching a suitable locality, molts into
an octopod nymph, which will feed and develop into an adult mite.
The causes that will induce a nymph to transform to a ffe/popw,. are
yet nanknown; Megnin supposed that dryness of the air or a scarcity
of food were necessary causes, but Michael has shown that JIqo)pZ
are developed in the absence of these conditions, and that this stage is
a natural and normal means of distributing the species. The structure
of the I[ypip)/us is characteristic for each species; but it has not vet
been found in all species, and in (il /;j)1wq/,8 the hypopial stage is
only partially. developed, so that specie should not be described from
this stage alone.
In the early days of acarology Ilfyjo),, stood for a separate 'enus,
allied more to Gaa, ... than to Tyeoflly/ii.'. Dujardin, in 1850, con-
cluded that Iyops was the pupal staoe of (-amasida,. As investi-
gation proceeded, Iypojnm.0 was so frequently found in association with
Tyroqlyp, auv that views were advanced as to their relationship. One
was that Ifyp)op. was a ferocious parasite, devouring the 7/y/to ,,y .
from within; another, that Jfyj/po)uN was the male of /ruqly/i '&,
and a third, that lpyl)opvus was the real adult of certain species of
Tyroljiyp1ots. The "Hypopus question" disturbed acarologists for a
long time, but was finally settled by the work of Megnin and Michael.


The Tyroglyphide of the United States have never been investigated
in a systematic manner. The principal economic species have been called
Tyror/lypl u8 6"iro or T. lon,,yo ', but without comparison with u ropean
specimens. Fitch, in his Third Report (1856), described a mite, Atca.
ribis, which may be a tyroglyphid. In 1868 Shimer described a mite
as Acarus nalus; this was interpreted by Riley and some Euro-
pean authors as a tyroglyphid, but from the description it is evidently
what Lignieres describes as Jfm si.arooptes cocci'(Npo. Riley, in 1874,
described a Tyroqiypk u" jityllox'erw as destroying the grape Phyllox-
era. I have identified as this a common species of RA ;zoiyt/yt,.s,.
Riley's opinion of the predaceous habit of this species was erroneous,
as it undoubtedly feeds on the roots of infested plants. In 1884 Haller
described Tyi gjyp]us crassipes and T. ertu.s from "Amerika." I
have not recognized them, and do not know whether they were from
the United States or not. In 1893 Osborn described a mite infesting
mushrooms as Tyroglyphus lintneri. I have seen numerous specimens


of this species, which is allied to what Canestrini has called Tyro6
lypkw1. 8iro. -In 1896 Felt described a mite injuring the roots of caj
nations as Tyroqlyphus I trohorpkWu. This species I have seen als
and retain it in the genus I/roqlyphus.


As regards the occurrence of European species, in this country,
have had great difficulties in arriving at definite conclusions, owing t
the fact that few European writers have appreciated the importance
of minute characters in classification. They have identified as on
species form, totally different and treated under different names forn
closely allied or identical. 'h( greatest trouble centers around Tyr(
qy1 A Its s/i'/" the type of the genus. Several European acarologisi
have figured this spc-cies. Michael is later thatin the others, and mei
tions the other figures in his references. (anestrini has figured nol
bristles than the others, and the' are longer. Tlie pair on dorsum (
ald)(nlien behind middle, according to Michael. are very short; wit
both Canestrin and Berlese thev are long. Canestrini and Michae
however, agree and differ frn(, Berlese in showing a pair of shoi
1)ritles near the larger huieral bristles. Michael figures the ma]
hind tarsus rather short and with the two suckers barely more tha
their diameter al)art, while, according to Berlese, this joint is ver
much longer and the two suckers are very remote from each othei
Canestrinis figure shows the l)eculiar thickened hair of the anteric
tarsi as arising from the preceding joint. Several European writer
(especially Robin, Pagenstecher, Karpelles, and Murray) have dent
tied specimens of Ab'UIov~f(Ir fl as T. ,#ro, and Doctor Oudemar
writes me that he does not know Ts jio; that the specimens formerl
placed by him in that species are Ah1urobbis" foinrbi. Berlese hE
suggested that these two species are one, the spurred male of Alez
rob b/s being a dimorphic forn. The habitat gives no clue to th
species. Specimens of a fTroglypl ?,8 are referred to elsewhere i
this paper which were taken from Limburger cheese, and doubtless (
European origin. These are not the T ;ro of any author, but moi
related to T. my1cop]4,t(,, M'gn. In a collection of mites obtained b
the Department of Agriculture from Doctor Berlese is a slide labele
by him as bl.qioj. The specimen seems to be T ,iro as figured b
Canestrini and himself; the hind tarsi are not nearly as long as the
figure for T. Ion for, and I can not see any hairs or pectinations 'o
the large bristles of the body. In the same collection is a sid
labeled T. kra-neri by Berlese. Michael considers this form to be who
he culls T. nycopagus Megn. Their figures show a species with vr
short abdominal bristles, and the inner cephalic ones much short
than the outer pair. The slide has three specimens, all females, wit


four subequal bristles on the cephalothorax, and the bristles at tip of
abdomen are as long as body; moreover, these bristles are provided
with fine, short hairs. In fact, it agrees very well with Michael's
figure of T. lonfior, except that the abdominal bristles are not quite
so long. Michael, however, figures the tarsus of this species as much
shorter than figured by Canestrini and in my specimens of T'. lon gior.
Michael, in detail figure of Alearohsas frrnw, male leg I, shows the
clavate hair arising from tip of tibia, whereas it should be from basal
part of tarsus; at least it is so in my specimens and so figured by
Berlese. Michael figures iierurobb.s farini with but two long poste-
rior bristles on the cephalothorax; (anestrini shows four of these,
but no long humeral bristle, as in Michael. My specimens have the
humeral bristle like Michael's figure and the cephalic as figured by
Canestrini. Berlese figures six cephalic 1)ristles, besides the frontal, on
this species. Berlese figures the hind tarsi of male Aleurubusfarsn w
with suckers far apart. My specimens agree with Michael in this
respect. Michael considers that his Olt Io(/lyj/jt, anovyimn 8 is the
same as Jltycoblnm aio//ymitl of Canestrini and T/ltodtctb/1u, alno i -
mu.Y of Berlese; yet Michael's figures show numerous differences from
their figures. Michael, for example, shows short spines on the body,
while they figure simple hairs.
In view of these and other discrepancies among European authors,
I have been sorely tempted to abandon all attempts at identification
and describe everything as new. However, I believe that we have
Tyroglypku loeflir, T. (Aleuiobuel.) farin, and Carapoqiqi) w)ji8-
8uiarumi in this country, and I should like to think that what I have
described below as T. (NiW'r/caims was the real -T. iro, but it certainly
is not the T. s;ro of Michael, the latest writer on the European fauna.


The genera known to me as occurring in the United States may be
separated by the following table. Various other genera are known
in Europe, and some of them, and possibly new genera, will be found
in our country when it is examined more thoroly for these mites.
The forms thus far collected are mostly of economic value, and have
been sent to this Department by various persons during the past
twenty-five years.
1. Dorsal tegument more or less granular; claws very weak, almost invisible;
some hairs of body plainly feathered; ventral apertures very large Glyciphagus.
Dorsal tegument not granular; claws distinct; no prominent feathered hairs;
ventral apertures small ----.----------------------------------------- 2
2. Mandibles not chelate, elongate and toothed below; body without long hairs;
palpi enlarged at tip and provided with two divergent bristles ----- Histiostoma.
Mandibles chelate; palpi not enlarged at tip, nor with the two bristles ------ 3


3. No clavate hair on base of tarsi I and II; no suture between ceploth
and abdomen; living on bees or in their nests ----------------Tricho
A clavate or thickened hair on base of tarsi I and II ---------------------
4. The bristle on penultimate joint of legs arises from near the middle; no sutur
between cephalothorax and abdomen ----------------------arpog,
The bristle on penultimate joint of legs arises from near tip; a suture between
cephalothorax and abdomen -------------------------------------------
5. Cephalothorax with four distinct and long bristles in a transverse row; tars
I and I about twice as long as preceding joint ----------------Tyrog
Cephalothorax with but two long, distinct bristles (beside the frontal pair)
but sometimes a very minute intermediate pair; tarsi I and II usually short
and not twice as long as preceding joint -------------------------------
6. Tarsi with some stout spines --------------------------------Rhizogl
Tarsi with only fine hairs --------------------------------------. Monie.
The hypopial stages are known for very few of our forms; it w
therefore be of little use to tabulate them.. These mites exceptp
possibly Tri cltasos) should be studied solely from the mature a
and not described from the hypopial stages.


Mandibles not chelate, but elongate, and toothed below; i
enlarged at tip and bearing two distinct divergent hairs; a disi
suture between cephalothorax and abdomen; male without
suckers; tarsi with distinct claws; cuticle not granulate; vel
apertures small; rarely with long bristles on body; no long pr
nent bristle at tip of penultimate joint of legs.
Tge.-IL rostro&ratws M(gn. (peetineunl Kramvr).
The peculiar mouth parts of this genus separate it rather sha
from all the other Tyroglyphide. It, however, has a well-deveh
hypopial stage, indicating affinity with Tyroglypku The specie,
variable in habits, but none, so far, has become of econ
The three species which have been found in this country ma
classified by the following table:
1. Tarsi I four or five times as long as broad; very slender ..................
Tarsi I scarcely three times as long as broad; body broad, especiall:
behind --------------------------------------------------------bre
2. Body about twice as long as broad, with several humps above on the abdc
men ---------------------------------------------------------- americo
Body scarcely one and one-half times as long as broad, without humps ol

Histiostoma gracilipes n. sp. (P1. I, fig. 9.)
Body hardly one and one-half times as long as broad, sides of a
men evenly rounded, and without humps above, emarginate bet
at posterior third of body there is a short curved hair,, a similar
at each posterior corner and on each humerus, and two or three



median pairs on the dorsum; all of these hairs are very short, curved,
and often invisible. Legs rather large and long; the tarsi very slen-
der, tarsus I (P1. I, fig. 12) nearly three times as long as penultimate
joint, above with two short spines near base, and before middle one
more, below with two rather beyond middle, and with several at tip,
and a long curved hair about three-fourths the length of tarsus; hind
tarsus fully three times as long as penultimate joint, with a short spine
above near base, a pair below beyond middle, and several near and at
tip, but no long hair. Hairs on tip of palpi much shorter than in other
Length, 0.30 to 0.35 mm.
Numerous specimens on decayed leaves, Washington. D. C.. August.
Histiostoma brevipes n. sp.
Body about one and one-fourth times as long as broad. subpyriform
in shape, broadly rounded behind, without very distinct humps, but
the posterior margin undulate; behind with four simple bristles, each
about as long as tarsus, and a pair of similar bristles on the posterior
sides; a similar humeral bristle each side. Legs rather short, first
pair heavy, provided with short spines: tarsus I (P1. I, fig. 11) about
two and one-half times as long as broad, with two spines below near
middle, one above before middle, and near it is the sense hair (instead
of at the base); at tip above is a spine, and near by is a fine apical
hair, two-thirds the length of the joint; tarsus IV (P. I, fig. 10) is
fully three times as long as the penultimate joint, with a spine above
near base, and one below near middle, and a few at tip. The palpus
(Pl. 1, fig. 8) has the usual two bristles, the apical one very long, longer
than tarsus I. and much longer than in allied species.
Length, 0.28 to 0.33 mm.
Several specimens from dead and diseased larvw of Cqlleiw roh;,li,,
Forst. in locust at Arlington, Va., July (Hopkins).
Very distinct by short tarsi, position of sense hair, and long hair to
palpus. In appearance it is much like Michael's I pyr'fori'nc, but
without the long apical hairs to posterior tarsi.
Histiostoma americanum n. sp.
Cephalothorax divided into two parts, a broad posterior part and
a narrow, elongate anterior portion, in some specimens almost broader
in front than behind, and on its anterior margin are two long bristles;
below are situated the mouth parts (Pl. I, fig. 7). Abdomen about
twice as long as broad, rather broader in front than behind, rather
rectangular in shape, almost straight across at base, weakly emargi-
nate behind; above with three large rounded humps each side, two
behind lower down on posterior surface, a median one at base
above, and less prominent ones on the lower sides; each of the larger
humps bears a short, stiff bristle. Legs quite short, with but few,


rather thickened, hairs; tarsus 1 (Pl. 1, fig. 4) nearly four
long as broad, with sense hair at extreme base, a spine slightly
and one near middle below, apical hair about one-half as long u
tarsus IV (P1. I, fig. 6) four times as long as penultimate joint.
spine above near base and one below near middle, apical h
Length, 0.20 mm.
Taken at Washington, D. C., in decaying matter, together
species of Rizogloylyps.

Cuticle of body more or less granulate; claws very small anc
spicuous; some of the hairs of body plumose, or formed in
aceous scales; the ventral apertures are yery large and occupy
space between the cox,; mandibles chelate, usually with a
between cephalothorax and abdomen; male without anal s
female with the bursa copulatrix projecting slightly at tip
TIpe. -G. dornesticus De Geer.
This genus is abundantly represented in Europe, but in this (
I have seen but few species or specimens. The genus should
tribe or subfamily in the Tyroglyphid-w on account of its widE
gence in structure from the typical Tyrog1ypoh1wo. These mit
been found feeding on all sorts of substances, but the name ir
that they are the true "sugar mites, and cause the disease kr
"grocer's itch." Two species are'Iknown to me.
Glyciphagus obesus n. sp. (P1. 1, fig. 3.)
Body about one and one-half times longer than broad, I
sided; pointed in front, broadly rounded behind. Dorsum wi
tered, irregular granules. Cephalothorax with four pairs ol
simple bristles; two pairs in front, a submedian pair behind, 9
in each posterior corner; and a short, broad hair or scale near
over coxe 11. Abdomen with a submedian pair of plumose
on basal third, a pair of simple bristles nearer to --margin I
terior third; three simple bristles each side, and five each side
near the tip, the outer one the longest. Legs rather short an'
I (Pl. I, fig. 2) and II with a curved, plumose bristle near 1
third joint and near middle of fourth joint, the latter joint
long, simple bristle at tip; tarsi I and II one and one-half times
than preceding joint, tarsi 111 and IV twice as long as preceding
Venter minutely granulate, genital aperture (P1. I, fig. 5) occ
all the space between coxte and mouth parts.
Length, 0.33 to 0.38 mm.
Taken from a necktie that had been for some time in a dra
Berkeley, Cal. (E. J. Wickson.)


Tech. Series 13 Bureau of Entomology. U. S, Dept. of Agriculture.








Fig. 1.-UYcquatrgiu robaxtit,. Fig. 2.-lYciphafju. obf.,, leg I. Fig. 3.-;lyciphagqi ib/.s/.s.
Fig. 4 -ttisti, twu, (mnricanll. tarsns I. Fig. 5 -b!/tCiph(UI. ,s is, geniital plate. Fig.
I.-tistiostoIn (I /tmi( u'uiu/1Wt, tarsus IV. Fig. 7.-lti.tistomau eim Wrican U/, inolith parts.
Fig.'stoma br rp ,., lallms. Fig. 9.-Ifistiostmna qracilipirs. Fi 1O.-listiost,,m
breipes, tarsus IV. Fig. l1.-Hitiorona brcripcs., tarsus I, Fig. 12.-HtiWi.toma r/ilipff+,
tarsus I.



Glyciphagus robustus n. sp. (Fl. i, fig. 1.)
Body short and broad, rather broader behind middle than else-
where, broadly rounded behind. Dorsum with many large, rounded
granules, irregularly arranged. A sul)median pair of long, sparsely
plumose bristles oii front margin; a still larger pair near the hind mar-
gin of the cephalothorax, as near to side as to middle; one bristle in
each posterior corner, and two shorter submarginal ones each side in
front of the last. Abdomen with about six pairs of long, dismal, plu-
mose bristles, the Subbasal pair not nearly as long as the others- and
six submarginal bristles each side, the two on the posterior margin not
nearly as long as the others. Legs rather short, but the tarsi are
slender; tarsi I and 1I twice as long as preceding joint- tarsus IV
more than three times as long as preceding" joint, all witl scattered,
simple hairs, a longer hair near tip of third and fourth joints of legs
I and I. Venter rather finely o-ianilate: the large, broad genital
aperturnelccupieulate ttheslarge,
aperture occupies aill the space between coxe II, III, and IV, but
does not extend forward between coxce I.
Length, 0.-24 mam.
Specimens from Leetonia, Ohio, in a lot of seeds, from Mr. H. E.
Genus TYROGLYPHUS Latreille.

A suture between cephalothorax and abdomen; mandibles chelate;
tarsi with distinct claws: cuticle without granulations: ventral aper-
tures small, four distinct posterior bristles on the cephalothorax; tarsi
rather slender, in some species with spines; male with anal suckers;
in some cases there is a dimorphic male, or the anterior legs of male
may be thickened.
T//k. 7. /iro L.
Oudemans uses the name Acarets for this genus, but I think the
application is strained and that Michael is right in this matter. I
include in Tyjrofllpl y the genus Alenro!tts, which is based on a male
character of not more than specific value.
There are doubtless a number of species in the linitA States. From
the materials at hand I separate. the following nine species:
1. Some bristles on tarsi I and 1I near middle are distinctly spine-like; the
sense-hair about its length from base of joint ----------------------------
No spine-like bristles near middle of tarsi; sense-hair not its length from base
of joint -------------------------------------------------------------- 4
2. Hind tarsi with two long hairs, one as long as the joint --------------- /erminalis.
Hind tarsi without such long hairs -------------------------------------- 3
3. Male with third legs enlarged ---------------------------------- heterouiorph ts.
Male without enlarged third legs ------------------------------------- arm ipes.



4. Of the terminal abdominal bristles at least six or more are very l
as long as the body ----------------------------------------------
Of the terminal abdominal bristles only two are about as long as th
men; leg I of male greatly thickened, and with a spine at apex of f
below . . . . . . .
5. Bristles of body distinctly pluinose or pectinate; tarsi very long----------
Bristles of body not pectinate -----------------------------------------
6. Cephalothorax very short; legs I and II of male stouter than usual .....
Cephalothorax longer, legs norlmial ...............................
7. On living trees, usually with scale insects; third and fourth joints of hi
legs miiore than twice as long as broad; abdominal bristles long ------ coc
On uinshroomis, or decaying matter; third and fourth joints of hind legs
twice as long as broad; abdominal bristles very long ...............
In mills, stored foods, grains, etc.; third and fourth joints of hind legs scarce
twice 0 long as broad; abdominal bristles shorter ---------------- ame
Tyroglyphus farinae De Geer. (Pl. II, fig. 14.)
Cephalothorax with four long subequal bristles above in a
verse row, a short pair in front over the mandibles; one oi
humerusn not as long as width of body. and a very short one ne
two pairs on the middle of dorsum, rather farther back than
the posterior pair the longer, but these not more than one-h
length of the abdomen: two hairs on each posterior side, not or
the length of abdomen, and near the tip are two more pairs of bi
one of which is nearly as long as the abdomen, the other pair
much shorter; a pair of short bristles near anus below (P1. II, fi
Legs rather short. and the front pair thickened; in the ma!
greatly so,. and the femur provided with a sharp, apical p
below at tip. and two small teeth on next joint. The long bri
penultimate joint is as long as the tarsus in all legs; tarsus I (
fig. 17) is scarcely twice as long as the preceding joint, tars
(P1. II. fig. 14) more than twice as long as penultimate join
third and fourth joints of the hind legs are about twice as 1,
broad; none of the hairs on the legs is spine-like.
Length, 0.45 to 0.60 ram.
Specimens have been examined from various localities, all, ho'
in the North, as Marblehead, Mass., Lancaster, Pa., Adrian,
and Minneapolis, Minn., in mills, granaries, and seeds.
I keep this species in the genus Tyroglyplue, since the genus
rohius is based on a secondary sexual character; in other resp&
a true Tyroqlyphu. If this genus is used, then other genera
be made for 7: letelrontorphus and T armipe8, which is, I
Tyroglyphus longior Gervais.
Cephalothorax with four long, subequal bristles in a vers
and two shorter bristles on front margin; two long hu ral hi
two pairs of submedian bristles on dorsum, the an pr
than one half the length of abdomen, the posterior pair as long


Tech. Series 13, Bureau of Entomology, U. S. Dept. of Agriculture.




Fig. 13.-Moviezielalongiprs. Fig. 14.-Tyroglyphus farina and tarsus IV of male. Fig. 15.-Mon-
iezi(.(a (n1aav.usta. Fig. W.- Tqforl qph is fitrimv, venter of male. Fig. 17.- Ty!y.lyp/h ts farbiw,
leg I of male. Fig. 18.- Mmoiziclla brerita'sis. Fig. 19.-Monieziella breiitarsis, imiale venter and



entire body, behind toward tip are six pairs of bristles, most of them
fully as long as entire body; all these bristles of body are seen to be
hairy (Pl. V, fig. 44): the cephalic bristles show these hairs more dis-
tinctly toward tip. The basal joints of the legs are of the usual
length, but the tarsi are extremely slender; tarsus I (P1. V, fig. 44)
i- about as long as the three preceding joints together, while tarsus IV
(P1. V, fig. 44) is about as long as the rest of the leg; the bristles oil
legs are about as usual, fine and slender, but that at the tip of the
penultimate joint of leg IV is scarcely half as long as the tarsus; the
hair near tip of tarsus is rather short on all legs.
Length, 0.40 to 0.60 mm.
This species is readily known by the hairy bristles of body and by
its extremely long tarsi. It has been received only a few times, but
usually in great abundance and usually attended with some predaceous
iites- CGeyletus or Gaas. Specimens have been examined from
grain in mill at Milwaukee, Wis.; from Winfield, Ontario, Canada, in
house and barn in great numbers, and Doctor Oudemans writes me
that he has seen it from California.
Tyroglyphus lintneri Osborn. (P1. III, fig. 29.)
Cephalothorax with four long, subequal bristles in a slightly curved
transverse row; a pair on front margin, longer than the mandibles;
two long humeral bristles and a short one close by; two pairs of sub-
median bristles on the dorsum of abdomen, the anterior pair about
one-half the length of the abdomen, the posterior pair as long as abdo-
men; and six each side near tip, all very long except the inner one,
which is scarcely as long as abdomen, and a short pair on venter near
the anus. The legs have the usual bristles, the long one at end of
penultimate joint is plainly longer than the tarsi in all the legs; the
hind tarsi (P1. III, fig. 24) are fully as long as the two preceding
joints together; there is a curved plumose bristle above on the third
joint of legs I and II. (Male genitalia, see P1. III, fig. 25.)
Length, 0.30 to 0.38 mm.
This mite is very similar in all respects to the T. americani ., but
the bristles of abdomen are longer and somewhat differently arranged.
In the male the tubercle-like suckers on hind tarsi (P1. III, fig. 24) are
farther apart than in T. aemercan ,s, in the female the vulva (P1. III,
fig. 23) shows a broader emargination behind than in that species.
These differences, associated with the different habitat, demand some
recognition in nomenclature; therefore I consider the mushroom mite
as a distinct species. It differs at once from T. longior in the simple
lbristles of the body.
This species has been received from Freehold, N. J.; Hazelton and
West Chester, Pa.; and from York Corner, Me. It does enormous
damage to mushrooms, but appears to be a native species. Lintner
records it from Jamesport, Suffolk County, N. Y.


Tyroglyphus americanus n. sp. (P1. 111, fig. 20).
Cephalothorax with four long, fine, equal bristles in a $
curved transverse row, a pair of shorter bristles on front in
two long funeral bristles, longer than width of body, a pair o
median bristles each side ol the dorsum, the anterior rather she:
posterior very long, as long as abdomen; on posterior margin
near the tip are six bristles each side, five of them about asi
abdomen, the other much shorter and near the median line. L
moderate length, like figures of 7. siro, but the tarsi (Pl. III, f
are more elongate and slender then in Michael's fig ure of that sj
the usual bristles are present, none spine-like; that at tip of pi
mate joints is very long; there is a curved plumose bristle
the third joint of legs I and II; the hair at tip of tarsus is no
half the length of the joint; in the male the sucker-like tuber(
tarsus IV are but little more than their diameter apart. (Ger
P1. Il1, fig. 21.)
Length 0.27 to 0.30 mm.
This species is close to Michael's identification of .T. 8,o, bi
much longer bristles and longer tarsi; indeed the hind tarsi are a
as the two preceding joints together; however, it can not be 16
since the bristles are not serrate. It' may be that some Eu
writers have mixt this species with longlior, but I regard Mi
identification of 7. lon Wor as correct, that is, a mite with long s
bristles, and the same as the form I regard as T. longior in this]
There are many specimens of this species in the collection
Department of Agriculture, as follows: Washington, D. C., on
plumis; Paola, Kans., in flaxseed; Minneapolis, Minn., in i
Savannah, Ga., in rice; College Station, Tex., in cotton seed; E
Wis., in flax mill, and on decaying orange at Washington, D. G
Many of the references to T. siro and 7. longior in our eco
literature doubtless refer to this species.
Tyroglyphus cocciphilus n. sp.
Cephalothorax with four long bristles in a transverse row, thi
dle pair plainly longer than the outer pair, yet the latter is as i
width of body; a short pair on the front margin, longer than there
bless; two pairs on dorsum, the basal pair about one-half the len
body, the posterior pair about as long as entire body; two long h
bristles each side and a short one near by; behind and near tip:
bristles, 3 on each posterior side and 3 each side near tip in a v4
or longitudinal line, all about as long as abdomen or a little ]Ion
short pair below near anus. Legs of moderate length, with the
bristles, none spine-like; the tarsus (P1. IV, fig. 35) about as h
two preceding joints together in all the legs, the bristle at

Tech. Series 13, Bureau of Entomology, U. S. Dept. of Agriculture.

0O o i

22 Co






Fig. 20.-T!Iroglqth 0 (IlCiUUI/1. Fig. 21.-TfrFfl hp/I U .'rs Ifl iicuuus, golitalia. I-ig. 22.-T/-
rofllylph,. us unricuu us. tarsi I and IV. Fig. 23 TOUq~ljh u lint u vitlva. Fig. 24.-T.fir-
gb!iph ,s liutniri, tarsus, IV ot male. Fig. 25.- T!1, (ll!yphs lid/i ', genitalia ()f male. Fig.
26.- T!iro!llyph is tt rio(nls, leg I. Fig. 27.-ITqmqlqphu. ,me nus. tarsus IV. Fig. 2,.-'-
rog!/1phus tcrmiualis, vulva. Fig. 29.-T!1r.qo1.qp us lidnl' II.

PLATE 111.


penultimate joint about as long as tarsus; third and fourth joints of
hind legs more than twice as long as broad; in the male the suckers on
hind tarsi (P1. IV, fig. 35) are more than twice their diameter apart
and nearly as close to each end of joint as to each other. (Vulva and
anal suckers, P1. IV, fig. 33; male aperture, Pl. IV. fig. 34.)
Length, 0.35 to 0.40 mm.
Specimens from Columbia, Pa., with Lecani tnt on plum; also with
oyster-shell scale oil osage orange (probably from Missouri), with mealy
bug on guava at Rock Ledge, Fla., and on orange leaves at Sanford, Fla.
This species is very close to I. lintneri Osb., and might have been
considered as only a variation of that species but for the totally differ-
ent habitat. The legs are rather more slender, but the hind tarsi are
no longer, and the hair at the tip of the penultimate joint is not as
long as in T. 1;nlbeo, nor as heavy.
Tyroglyphus breviceps n. sp. (Pl. IV, fig. 30).
Cephalothorax very short, in proportion to the length of the body;
four rather long, subequal bristles above in a transverse row, and a
short pair in front over mandibles; two moderately long humeral bris-
tles; two submedian pairs on the dorsum, the basal pair short, the
other scarcely as long as abdomen; six bristles each side and near tip,
rather widely separated at base, front two-thirds to fully the length
of the body; a short pair below near anus; tarsus about twice as long
as preceding joint, the bristle at tip of latter as long as tarsus on the
front legs (P1. IV, fig. 32), and three-fourths as long on the hind legrs;
each tarsus with a fine bristle beneath near middle; hind tarsus (P1.
IV) fig. 31), with apical hair not one-half the length of tarsus; the
third and fourth joints in hind leg not twice as long as broad.
Length, 0.35 to 0.50 mim.
The hairs are a little shorter than in E1 a. e 'ana and the legs
rather stouter, and especially so in the front legs of the male.
Specimens from Victoria, Tex., taken from dead larvwe of the cotton
boll weevil.
Tyroglyphus terminalis n. sp.
Cephalothorax with four bristles in a transverse row, the outer pair
nearly twice as long as the inner pair, a pair of short bristles on ante-
rior margin; two humeral bristles, about two-thirds the width of the
body, two pairs of bristles above on abdomen, and five pairs near tip,
all about one-half the length of the abdomen except one pair each side
near tip, which are about as long as the abdomen; all are simple.
Legs rather stout, especially the anterior pair (P1. III, fig. 26), the
bristles stout, and many of those on the tarsi are spine-like; tarsus I
is not twice the length of the penultimate joint, while the hind tarsi
(P1. III, fig. 27) are about as long as two preceding joints together;



the hair near tip of tarsus is very long in all legs, but the
have two long hairs near tip, one of them loge t
(Vulva, P1. 11., fig. 28.)
Length, 0.50 mm.
Specimens from Limburger cheese, Washing D. C
This species is near Michael's figure of Tyroglypk. any
the mite is not so slender. the bristles are longer, and
are longer. I am not certain that Michael has correct
Megnmins species.
Tyroglyphus heteromorphus Felt.
Male (P1. IV, fig. 39).-Cephalothorax with four long b
transverse row, but the inner pair is not one-half as long a.
pair; a pair of short bristles on anterior margin; two bristh
humerus, one short, the other as long as width of body; tw
submnedian bristles on dorsum, each about two-thirds the ler
abdomen; a rather long bristle on the middle each side, an
one behind this, then a short one, and three long ones each
tip, each but little shorter than abdomen. Legs with the
tles, that at apex of penultimate joint scarcely as long as t
sense hair on tarsi I (Pl. IV, fig. 38) and II is about its lei
the base; most of the tarsal bristles are distinctly spine-lik(
at tip of tarsus IV (Pl. IV, fig. 36) is not one-half the len
joint, the bristle at tip of penultimate joint of leg IV is not 4
long as tarsus, the latter joint as long as two preceding joint.
leg III (Ol. IV, fig. 37) enlarged. as in figure, ending in a'
and two very long bristles.
The female differs in having the bristles of body shorter,
the abdominal ones; there are four at tip which are not (
long as width oT body; the tarsi, especially the hind tarsi, U
than in the male. The abdomen is, of course, broader, anc
proportion to the cephalothorax.
Length 0.60 to 1 mm.
A peculiar species, which I have seen only from decaying
roots from near Washington, D. C. It was described as ir
roots of carnations at Berlin, Mass. The R/dzoglyphvu agil,
1903, may be a synonym ; however, there are several minor,
which may possibly be due to Michael's inaccurate figures.
Tyroglyphus armipes n. sp.
Cephalothorax with four long bristles in a transverse ro
inner pair is not half as long as the outer pair; a pair of s(
on front margin. Male with two bristles on each humerus,
the other nearly as long as width of body; three pairs of
dorsum, basal pair very short, the others long and reaching

Tech. Series 13, Bureau of Entomology, U. S. Dept. of Agriculture.

/- 33









1.1"I] ',

f f




Fig. 30.- T!rogljh us br diccps. Fig. 31.- T!rofl!! phwu, biwcic'eps, tarsus IV. Fig. 32.- Tyrorilph ts
breviceps, leg 1. Fig. 33.-Tyrorltyphits cocciphilts, vulva and anal suckers. Fig. 34.-T!ro.l !/-
phits coccipht tis, male aperture Fig. 35.-Tqrogl!!phs cocciph ilts, tarsus IV-male and female.
Fig. 36.- T/ruqltthus hrt Y)(cronmrphtis, tarsus IV. Fig. 37.-T!ro.ql1qphs /htc'rotiU)rphlus, leg III of
male. Fig. 38.-Tyroglyphits heteromorphus, tarsus I. Fig. 39.-Tyroglyph us l hdtromorphis,




of abdomen;. two on each posterior side, quite long, and behind are
four each side, three of which are about as long as the abdomen,
the inner pair much shorter. Legs rather slender, tarsi as long as
two preceding joints together; most of the tarsal hairs are spine-like;
the hair at tip of penultimate joint is rather shorter than tarsi, but in
leg IV (P1. V, fig. 41) it is about two-thirds as long as tarsus; the hair
at tip of tarsus not one-half length of tarsus. (Tarsus I, P1. V,
j Q.: 42.) In the female the bristles are much shorter, the humeral
-wistle not nearly as long as width of body, those on dorsum not reach-
ingc to tip, the posterior lateral ones very short, and those behind near
tip are scarcely one-half as long as width of body. (Vulva, Pl. V,
fig. 43.)
Length, 0.75 to 1 mm.
Specimens from Lincoln Nebr. and from Atlanta, Ga., on dead
larv, of corn pyralid.
Genus RHIZOGLYPHUS Claparede.
A suture between cephalothorax and abdomen; mandibles chelate;
tarsi with distinct claws; cuticle not granulate; ventral apertures
small; only two distinct posterior bristles on the cephalothorax, but in
some forms a minute intermediate pair is present; tarsi short and stout,
provided with some stout spines; male with anal suckers, and in some
cases there is a dimorphic male with the third pair of legs greatly
enlarged and ending in a large curved claw. The species are vege-
table feeders, and attack healthy living tissues, usually the part in the
Type. -P. hyacintdki Boisd. (echinopus Robin).
This genus was based on the absence of a caroncle to tarsus, a char-
acter of variable value in allied genera. Michael uses the dimorphic
male as a distinguishing character, but I would rather not use a sexual
character for a genus, so base the genus on the two posterior cephalic
bristles and spiny tarsi.
I distinguish six species in our fauna by the following table:
1. Tarsus I with a large spine close to the sense hair ------------------------ 3
Tarsus I without a spine near the sense hair ............................... 2
2. Abdominal bristles as long as width of body; tarsi longer --------------tarsalis
Abdominal bristles much shorter than width of body; tarsi shorter.. phylloxerw
3. Tarsus I fully two and one-half to three times as long as broad ------- longitarsis
Tarsus I not more than twice as long as broad --------------------------- 4
4. Body three times as long as broad; legs very short----------------- elongatus
Body less than three times as long as broad ----------------------------- 5
5. Hind tarsus two and one-half to three times as long as broad; abdominal bris-
tles longer; no dimorphic male seen --------------------------- rhizophagus
Hind tarsus about twice as long as broad; abdominal bristles very short; a
dimorphic male is common --------------------------------------- hyacinthi



Rhizoglyphus phylloxerse Riley. (Pl. VI, fig. 61.)
Cephalothorax with a pair of frontal bristles and a pair of
larger posterior bristles, also a pair of minute intermediate brisl
one bristle on each humerus; in the female there are six short bristles
near the tip of the abdomen, the longest pair not one-half te with
of the body; a submedian pair of short ones behind the middle of the
dorsum, and one short bristle on each posterior side. The le has
eight bristles at tip of abdomen, some about as long as width of ;
while the dimorphic male with the thickened leg III (P1. VI, ig. )
has these apical bristles about as long as the abdomen, and
pairs of long bristles on the dorsum. The legs are rather lo
tarsus I (P1. VI, fig. 59)has no spine above near the sense hair, but
toward the middle is a stifl bristle; the other spines are present
are long, the apical hairs are shorter than the joint; the bristle f
the penultimate joint is longer than the tarsus in all except the hind
legs: tarsus IV (P1. VI, fig. 57) is about as long as two preceding
joints together, and in the male is still more elongate; it has two spines
below near middle: the apical hairs are shorter than the joint. The
enlarged leg III of the dtimorphic male ends in a long claw; several
bristles are near by, but no tooth. In life these eites are rather
yellowish white, with chestnut-brown legs and a dark spot on each
posterior side of the abdomen.
Length 0.75 to 1 nm.
I identify this with Riley' s species, since his figures of the legs show
a rather slender tarsus. and the other characters shown by his figure
agree with this form.
Speciens have been examined from the roots of cowpeas from
Macon, (a.; from Auburn. Ky., on scabby potatoes; from Lawrence,
Mass., on young potato plants, and from Akron, Ohio, on rotten
potatoes; also from Illinois. infesting pine cones.
Michael has considered that Riley's species was a synonym of
R. eeinope.s M6gn. (I yac1l'tid Boisd.). This species is, however,
abundantly distinct from the "bulb mite," and perhaps does not now
occur in Europe, altho it was introduced into Prance. At that ti
it was supposed to feed on the Phylloxera.
Rhizoglyphus tarsalis n. sp.
This species is similar in nearly all respects to R. phylloxerxt
is, there is no spine on tarsus I (P1. V, fig. 45) near the sense hair,
and the tarsi are long. I have not seen, any males, but the fe
differs from R. pAylloleer a in having the bristles near tip of a
nearly as long as width of body, and in the plainly longer hind tarsi
(P1. V, fig. 46), which are longer than the preceding two joints
The specimens come from Spreckels, Cal., taken from sugar beets
by Mr. E. S. G.Titus.



Tech. Seres 13 Bureau of Entomo ogy U, S, Dept. of Agriculture

-~ 41


/ 7 -I \






< Vj


r (~





! 4


Fig. 40.- f rpob/phlts passdaritn. Fig. 41.- oTof!yli s a, iips, Itarsu,; IV. Fig. 42.-Ty',-
(Ifliphtil.' Uf/flj :. tarsus I. Fig. 43.- Tqt, 'qlqph,'. armit, .. vulva. Fig. 44-T *rglJ/ tt1il
lonw~'r, tarsi I and IV, and b(dv hairn. Fig. :.-Ihizq'2!tth ,s tor.o/i. tarsu-. I. Fig.
46.-Rhizoyl!ijhis trsqli. tarsts IN'. Fig. 47.-Rhiz,,!qlp t hmciuiti, l'g IlI of male. Fig.
4g.-Rii:aorlgph'. h!Jciathi. tarsi I and IV. Fig. 49.-fhiaip/llyphiDo hyacinthi.




Rhizoglyphus hyacinth Boisd. (P1. V, fig. 49.)
Cephalothorax with a pair of frontal bristles, and a pair of large
and long posterior bristles (intermediate bristles not visible); a rather
long humeral bristle; a submedian pair of short bristles on the dorsuin,
one on each posterior side and six near the tip, all short, the longest
scarcely one-half the width of body. Legs short and stout, the tarsus I
(P1. V, fig. 48, at right) but little longer than preceding joint, the
spine above is close to the sense hair, and the apical hairs are longer
than the joint; the bristle at tip of penultimate joint is longer than the
tarsus in all except the hind legs; the hind tarsus (P1. V, fig. 48, at
left) is about twice as long as broad, with two spines below near mid-
dle, the usual apical spines, and the apical hairs are longer than the
joint. In the male there are six hairs near tip of abdomen above, some
nearly as long as width of body, and below are four rather long, sub-
equal bristles in a straight transverse row; in the dimorphic male leg
11i (P1. V, fig. 47) is enormously thickened, and ends in a stout claw,
with a stout tooth at inner base. In color it is white with brownish
head and legs, and a dark spot on each posterior side of the abdomen.
Length, 0.55 to 0.75 mam.
Specimens have been taken from the bulbs of Bermuda lilies shipped
to this country.
Under the name of R. ec/K; pnw Megn. several European authors
have gathered various species; whether this form, which seems partial
to bulbs, is the same as Mtgnin's species 1 can not tell from descrip-
tions; however, I think it identical with the R. ec/#,nopus of Michael.
Its ravages in various bulbs and orchids have given it the name of
"bulb mite" and "Eucharis mite." The damage caused by it to
Bermuda lilies has been treated by Mr. A. F. Woods in 1897, in a
paper entitled "Bermuda lily disease." (See bibliography, p. 27.)
The R. menini of Haller appears to be a distinct species, with plainly
shorter bristles.
Rhizoglyphus rhizophagus n. sp. (P1. VI, fig. 50.)
Cephalothorax with a pair of frontal bristles, and a pair of long
posterior bristles (no intermediate bristles visible); a humeral bristle
fully one-half the width of body, two on each posterior side and six
at tip in the female, all short, the longest about one-half the width of
the body, and a submedian pair above on dorsum. Legs short, the
anterior pairs very heavy, the tarsi I (Pl. VI, fig. 51) and II with a
spine above near the sensory hair; and one below, rather before the
middle; apical hairs longer than the joint; the bristle from penulti-
mate joint longer than the tarsus in all legs, except the hind pair;
hind tarsus (P1. VI, fig. 52) two and one-half to three times as long as
broad, two spines below near middle, and the usual spines at apex;
the apical hairs as long as the joint. In the male the abdomen has
eight bristles near tip, one pair rather more than one-half the width

of the body; and on teventer (P. V1, fig..5)bhn
subequal bristles in a transvr rowe
of IV and not thickened in the many mal e
Length 0.65 to 0.80 mm.
This species differs from R. kyaiidti in
namely: the longer tarsi, especially hind tarsi; the
certain bristles, and the proportionately larger f
Specimens have been studied from Misso
apple trees; from Missouri, under a cottonwood st
from Glenellen, Cal.
Rhizoglyphus elongatus n. sp. (Pl. VI, fig. 53).
Cephalothorax with a pair of long frontal bristles, ar
posterior bristles, barely longer than the others, no intermei
visible; a huineral bristle each side no longer than cephalic
six short bristles near tip of abdomen, the superior pair
but no longer than frontal bristles. Body very elo
-three times as long as broad, mandibles large. Legs v
stout; tarsus I (Pl. VI, fig. 54), but little longer than penul
a spine above near the sense hair, one below beyond mid
near tip, apical bristles short; the bristle from tip of penu
very large and prominent, and longer than tarsus in a'
tarsus but little longer than front ones, and with short ap
Length 0.30 to 0.35 mam.
The only specimens seen were taken from the roots
October, 1879, probably in Missouri. Distinct by elongate
short legs, and short posterior bristles of cephalothorax.
Rhizoglyphus longitarsis n. sp.
Cephalothorax with a long frontal bristl a
long and large posterior bristles, pair of minute i
bristles; a long humeral bristle; the female with six bhi
of abdomen, none more than one-third the width of body;
the bristles are longer, some nearly as long as width of t
the venter, behind anus, are four bristles in a transverse
outer pair is very much longer than the inner pair. 'I
rather long; tarsus I (Pl. VI, fig. 58) about two and one-]
long as broad, and with a spine close to the sense bair, on(
middle and two others near tip; the apical hairs rly a
joint; the bristle at tip of penultimate joint is onger tba
in all legs, except the hind pair; the hind tarsus (Pl. V
very long and slender, with two spines beyond dl
hairs about two-thirds the length of the joint.
Length 0.75 mm.
I have seen specimens from Emporia, Kans, taken f
bulbs of Caladium esculenturn.

Tech. Series 13, Bureau of Entomology, U. S. Dept. of Agriculture.


Fig. 5.- Rhizofllh!.t rhizvqWphrtqi/. Fig. 51.-Rhi:.--)1/ph s ,'h rhi~otI)h b tarsus I. Fig. -2.-Ihi-
zfflh!)// I'hizsphuy/. taris IV. Fig. 5S.-R/,WitI/iphus(tfojifs.. Fig. 54.-Rhizo/I/phi s
/lft itis. tarsus 1. hg. 4).-Rhi!,l!ph.s lUl/it i.s, hiriv IV. Fii. iar6I .Jj,1;hi us N.hizo-
phaqu;ts, e'ale venter. Fig. 57.-Rhizoqiuph is jh! ii/hixr;. ar>1 IV--m lh ml, f'i milIc. Fig.
8.- i:i( /!/hi q i hs li,,,ijifusi, tarsis 1. Fig. 5.-R hizoy/1jhi/s j h uqb,xi --i, i ar i i' I. Fig.
60.-Rhiziiltphus ph!/llot.i r, leg III of male. Fig. j.-hZUt/hi/ph 1.1s p'l'ox(rt,, ha] 1 of front
of body


Genus MONIEZIELLA Berlese.

A suture between cephalothorax and abdomen; mandibles large,
chelate; tarsi with distinct claws; cuticle without granulations; ven-
tral apertures small; but two distinct posterior bristles on the cepha-
lothorax; tarsi without spines; male with ventral suckers; abdomen
usually more elongate than in TyroqlypIphts. All the species so far
known are predaceous or feed on recently killed animal matter.
Type.-Jl. entomophagus Lab.
The genus ilistiogaster is based on H. .arjyio Kramer, a species with
spiny tarsi, a peculiar male abdomen, and of very different habits; so
that I retain Jfonieziella as distinct from it.
There are probably from five to ten species in our fauna. I have
recognized the following three species:
1. Body about three or four times as long as broad; apparently no cephalic nor
humeral bristles --------------------------------------------------- angusta
Body scarcely twice as long as broad; cephalic and hurmeral bristles distinct. 2
2. Tarsi barely longer than preceding joint ---------------------------- brevitarsis
Tarsi nearly twice as long as preceding joint -------------------------- logipes

Monieziella angusta n. sp. (P1. II, fig. 15.)
Body three to four times as long as broad; apparently without any
cephalic or humeral bristles, and only four short ones near the tip of
abdomen. Legs very short; tarsi not longer than the penultimate
joint; the bristle from this joint longer than tarsi, especially in the
anterior legs, where it is very prominent.
Length, 0,30 to 0.35 mim.
Specimens taken from under scale insects (Aspidiotus) at Haywood,
Cal., and stated to be feeding on the coccids and their eggs.
This is probably what Doctor Riley figures in the Fifth Missouri
Report as Tyroglypus inalus; however, his figure shows some bristles
on the head which I can not see in the specimens before me. Michael
has identified Riley's species with the European X1. (I[istiogaster)
entomop]agus; but Michael's figures show a form very different;
more elongate and with shorter bristles. The Tyroglyp//.s mau.s of
Lignieres is probably a JMonieziella, but a different species, perhaps
f entomophagus, as asserted by Michael.
Monieziella longipes n. sp. (P1. 11, fig. 13.)
Cephalothorax with a pair of frontal bristles, and a pair of very
long posterior bristles; a long, fine humeral bristle each side; two
shorter bristles on the margin, and four rather long bristles near tip
of body, the-inner pair nearly as long as the abdomen. Legs rather
slender, and the tarsi nearly as long as the preceding two joints



together, and in the hind pair still longer; the hair at tip of the penul-
timate joint is longer than the tarsus in all except the hind legs.
Length, 0.33 mm.
Specimens from among .Mytilawj);, scales at Crescent City, Fla.
Monieziella brevitarsis n. sp. (Pl. II, fig. 18.)
Cephalothorax with a pair of short frontal bristles and a pair of
longer posterior bristles; a humeral bristle each side rather more than
one-half the width of body, and three bristles each side on posterior
margin toward tip, the outer one very short, the next longer, and the
inner pair about one-third the length of the abdomen. Body scarcely
twice as long as broad; mandibles (PI. II. tig. 119) large and prominent.
Legs short, with only a few short bristles, but that at tip of the penul-
timate joint is rather longer than the tarsus; the latter joint is only a
trifle longer than the preceding joint, but much more slender; the
sense hair on tarsi I and II is much curved. (Male venter, P1. II,
fig. 19.)
Length, 0.35 mm.
What is evidently the ITqpopils of this form has a projection on the
anterior part of the cephalothorax, with a black eye-spot each side;
the legs I and II are thick and heavy; leg III ends in a claw, as do
I and II; but leg IV terminates in two long bristles, the outer one
much the longer. hut both longer than the leg, and there is also a
short bristle near tip of the legs. The ventral sucking plate has six
suckers-two in front, smaller than others, and four in a curved row
,Specimens have been taken from Chi lo and Marshailville. Gia.I and it evidently feeds on the San Jose scale.


No suture between cephalothorax and abdomen; mandibles chelate;
tarsi with distinct claws: cuticle without granulations; ventral aper-
tures small; the epimera of the first two pairs of legs joined to each
other and to the sternum, thus forming a sort of skeleton; tarsal claw
arising from a clavate onvchium; the bristle on penultimate joint of
legs arises from near middle, not at tip of joint. The legs are rather
slender, not thickened in the male; the male has no anal suckers. On
the anterior margin of cephalothorax near base of mandibles there is
each side a rounded eye-like spot or projection, very doubtfully an
eye. No Ilypopuw is known.
Type.- C. .ssdarum Robin.
There are doubtless several species; I have described one from
Java; C. anonymous of Berlese and Michael has very short bristles on



cephalothorax, and Michael shows them spine-like, so that their
figures probably represent a species different front that of Canestrini.
and which I find in the United States. The species of this genus feed
on a great variety of substances, perhaps favoring dried fruits.
Carpoglyphus passularum Hering. (P1. V, fig. 40.)
Cephalothorax with a pair of short frontal bristles, apparently a
little back from the front margin: four posterior bristles in a curved,
transverse row, none very long: a humeral bristle each side, with a
short one near by; two submedian pairs of short bristles on the dorsuin;
two short ones on each posterior side margin; a short pair near tip
above, and three each side at tip, one pair very short, the others more
than one-half length of body. Legs moderately long. tarsi long and
tapering; the sense hair on tarsi I and II tapers to tip: the bristle
above on penultimate joint is situated near the middle and not at tip,
and is very large and prominent, and longer than the tarsus in all
except the hind pair of legs. There are a number of hairs on legs,
but no spines; the hair near apex of tarsus is long and rather farther
from the tip than usual in tyroglyphids.
Length, 0.40 to 0.50 mim.
Specimens have been seen from Fresno, Cal., on figs: from St.
Anthony Park, Minn., on dried figs and apples: from Albany. N. Y..
on smoked ham, and on polleiA of honey bees from Ohio.

Genus TRICHOTARSUS Canestrini.

No suture between cephalothorax and a)domen; mandibles chelate;
legs ending in distinct claws; male without suckers on hind tarsus; no
clavate or thickened hair on tarsi I and II; palpi not enlarged at tip;
cuticle not granulate; male without anal suckers; vulva of female
situated close to beak, between coxwe I; hypopial nymphs found upon
bees, adults in nests of bees.
Type.-. oS./ii Dufour.
The adult form is but little known; I have seen none from this
country; the hypopial nymphs are more common, and two forms
have been found in the United States. These agree well with the
European species, but as the adults are not known, their identification
is rather uncertain. They are of no economic value, and therefore I
shall simply list the two species as the European forms until such
time as mature specimens make possible a more exact identification
and satisfactory description.
Trichotarsus xylocope Donn.
Prof. H. Osborn has recorded this European species as taken from
a California Xylocopa, and sent to him. by Mr. D. W. Coquillett. I
have not seen it, and possibly some other species was concerned.



Trichotarsus osmim Duf.
I have taken specimens of a species of Trichotarsuts, which age
with the European form, from a species of Osm/ia at Sea Cliff, IN. Y.;
however, the adult females may present some differences. This
species differs from T. xylocopx in having two claws at the tips of
tarsi 1, 11, and III.

BANKS, N.-New genera and species of Acarians. 1902, pp. 171-176.
Records Triohotarsus osmiw and T. xylocopw from the United States.
DEARNESS, J.-A parasite of the San Jose scale. <31st Ann. Rept.
Entom. Soc. Ontario, 1901, pp. 87-88.
Form not named, probably a Monijeziella.
FELT, E. P.-Eleventh Report Inj. Insects of New York, 1896, pp.
Describes Tyroglyphus heteromorph us.
HALLER, G.-Beschreibung einiger neuen Milben. gesch.. Vol. L, Bd. 1, pp. 218-219, 1884.
Describes Tyroylyphus crassipes and T. curtus from "Amerika."
HOWARD. L. O.-The cheese, ham, and flour mites. series, Div. Ent., U. S. Dept. Agric., 1896, pp. 100-102, 2 figs.
Treats of Tyroglyphms stro and T. lor!ior.
LINTNER, J. A. -A mite infesting smoked meats. Insects N. Y., pp. 130-131. 1888.
Supposed to be Tlroglypiu s iro.
LINTNER, J. A. -The cheese mite infesting smoked meats. Rept. Inj. Insects N. Y., pp. 291-293, 1 fig., 1889.
Regarded as Tyroglyplhus siro.
LINTNER, J. A.-The cheese mite infesting flour. Insects N. Y., pp. 294-295, 1889.
Notes on Tyroglyphus siro.
LINTNER, J. A.-Tenth Rept. Inj. Insects N. Y., 1895, p. 452.
Notes on Tyroglyphus lintneri.
OSBORN, H.-A new mite infesting mushrooms, Tyroglyphuq Nntn.i.
OBORN, H.-Tricodactylus xylocopw in California. Vol. XXVII, pp. 1021-1022, 1893.
OSBORN, H., and F. A. SIRRINE. -Cheese mites. Agric. Exp. Sta., pp. 894-895, 1894.
Considered as Tyroglyphus siro.



RILEY, C. V.-Fifth Rept. Nox. Beneficial Insects of Missouri, 1873,
p. 82.
Description of a form near Acaris i ralus of Shimer, according to his opinion.
RILEY, C. V.-Sixth Rept. Nox. Beneficial Insects of Missouri, 1874,
pp. 52-55.
Describes Tyroglyphus phylloxerx.
RILEY., C. V., and L. 0. HOWARD.-Mites infesting an old grain ele-
vator. Identified as Tyrogylyphus longior.
RILEY, C. V., and L. 0. ttOwARD.-Mites in flaxseed. Vol. I, p. 285, 1889.
Identified as Tilroglyphus .b'o, but probably 7 .frie..
RILEY, C. V., and L. 0. HOWARI).--Mites in a warm house. Life, Vol. 111, pp. 162-163, 1890.
Questionably nained Tytroglyphaus siro.
RILEY, C. V., and L. 0. HOwARD.-Cheese mites. Vol. 1II, p. 165, 1890.
Considered as Tyroglyphus siro.
SAUNDERS, W.-On two mites. Treats of Tyroglyphus siro.
SHIMER, H. -Trans. Amer. Entom. Soc., Vol. I (1867-8), pp. 368-369.
Describes-Acarrus mwlus which has been wrongly considered as a tyroglyphid.
WVSASHBURN, F. L.-A troublesome mite. 1903 (1904), pp. 157-159, 1 fig.
Notes on Carpoglyphits passdairun.
WEBSTER, F. M. -The common cheese mite, Tyoglyp ts ,Zro, living
in Sporotriemn globuliferum. <32nd Ann. Rept. Entom. Soc.
Ontario, pp. 73-74, 1902.
WILSON, -A mite infesting a pork-packing house. Nat., 1882, p. 599.
Considered as Tyroglyphus longior.
WOODS, A. F.-Bermuda lily disease; a preliminary report of inves-
tigation. 1897.
Rhizoglyphus hyacinthi as an agent in distribution of disease.





Acarus mals, bibliographic reference ------------------------------------- 27
description of form supposedly allied ------------------------27
probablyf=Hemisarcoptes coccisugus, not a tyroglyphid -------- 7
ribis, possibly a tyroglyphid ---------------------------------------- 7
Tyroglyphus not a synonym --------------------------------------- 13
agilis. (See Rhizoglyphus agilis.)
Aleurobius, based on secondary sexual character ---------------------------- 14
Tyroglyphus ----------------------------------------------- 13, 14
faring= Tqroglyphs farin -------------------------------------- -9
americanum. (See ttistiostoina americaeorwn.)
americanas. (See Tyroglyphus americanus.)
angusta. (See Monieziella angusta.)
anonymous. (See (Carpoglyiphus an on!inaus, PhIcobius anonynrnw., and TVich odac-
tylus anonymus.)
Ants, Tyroglyphidc in nests --------------------------------------------- 5
Apple trees, infestation of roots by Rhizoglyphus rhizophags ----------------- 22
Apples, infestation by Carpoglyphits passularm .---------------------------- 25
arm ipes. (See Tyroglyphus arnuipes. )
Asparagus roots, decaying. infestation by Tyroglyphits heterom o phus ---------- 18
Aspidiotus, food of Voaleziella angtsa- ..-------------------------------------- 23
Bacteria, following work of Rhizogflyphits hyacti ..------------------------- 5
Barn, infestation by Tyroglyphus longior ------------------------------------ 15
Bees, hosts of Trichotarsus ..--------------------------------------------- 25-26
pollen, food of Carpoglyphuspassiarun -------------------------------)5
Beets, sugar, infestation by Rhizoglyph us tarsalis ---------------------------- 20
Boll weevil, infestation of dead larvT. by Tyroglyphus breticeps --------------- 17
breviceps. (See Tyroglyphus breiceps.)
brecipes. (See Histiostoina brevipes. )
brevitarsis. (See ilfonieziella brevitarsis.)
"Bulb mite." (See Rhizoglyphus hyacinthi.)
Bulbs, food of Tyroglyphidw --------------------------------------------- 5
Caladiun esculentuin, infestation of rotten bulbs by Rhizoglyphus longitarsis --- 22
Carnations, injury to roots by Tyroglyph1us heteromorphus ------------------- 8,18
stems by Rhizoglyphus ------------------------------- 5
carpio. (See Histiogaster carpio.)
Carpoglyphus anonymus, description of genus ------------------------------ 24
species ------------------------------ 25
passularum, bibliographic reference --------------------------- 27
description and food habits ----------------------- 25
type species --. -------------------------------- 24
Cereal foods, food of Tyroglyphida --------------------------------------- 5


Cheese, food of Tyroglyphid- ........................................... 5,2,27
Limburger, infestation by Tyroglyphus terminalis ------------------- is
mites ---------------------------------------------------------- 5, 26,27
Cheyletus, attending Tyroglyphus longior -------------------------..------ 15
Chilocorus, association of Monieziella brevitarsis ---------------------------- 24
Clover, infestation of roots by Rhizoglyphus elongatus ----------------------- 22
Coccids. (See Aspidiotus, Mytilaspis, and Scale-insects.)
cocciphilus. (See Tyroglyphus cocciphilus.)
Corn pyralid, infestation of dead larvw by Tyroglyphus aipes- .............. 19
Corpse, human, food of Tyroglyphide ------------------------------------ 5
Cotton seed, infestation by Tyroglyphus arnericanus ......................... 16
Cottonwood, Rhizoglyphus rhizophagws under stump -------------------2...... .2
Cowpeas, infestation of roots by Rhizoglyphws phylloxer .-------------------- 20
crassipes. (See Troglyphus crasst.pes.)
curtus. (See Tyroglyphus curtus.)
Cyllene robiniw, dead and diseased larv? infested by Ili.tiohnimma breripe ...... 11
domesticus. (See Gly'iphagus domesficus.)
Drugs, food of Tyroglyphida .............................................. 5
echinopus. (See Rhizoglyphus echinopus.)
elongalus. (See Rhizoglyphus elongatus. )
entoonophagpix. (See Monieziella entomnophagus.)
"Eucharis mite." (See Rh izoglyphus hyacinth i.)
farinw. (See Troglyphusfarinw.)
Feathers, food of Tyroglyphidw ------------------------------------------- 5
Figs, food of Carpogl'plis pasultrum ------------------------------------- 25
Flaxseed, infestation by Tyroglyplius u imericanus ---------------------------- 16
Flour, food of Tyroglyphid ---------------------------------- ----------- 5,26
Fruits, dried, food of Carpoglyphus ---------------------------------------- 25
Tyroglyphid -------------------------------------- 5
Fungi, following work of Rhizoglyphus hgacinthi ---------------------------- 5
Guwmisus, attending Tytroglyphus longior ---------------------------------- 15
Glyciphagus, decription of genus ---------------------------------------12
descriptions of species ---------------------------------------1 2-13
domesticius the type species ------------------------------------ 12
obesus n. Sp., description ------------------------------------- 12
partially developed hypopial stage ............................ 7
robustus n. sp., description and food habits --------------------- 13
gracilipes. (See Ilistiostoma gracilipes.)
Grafting wax, eaten thirn by Rbizoglyphus --------------------------------- 5
Grain, infestation by Tyroglyphus longior ----------------------------------- 15, 27
Grains in mills, food of Tyroglyphid .------------------------------------ 5
Granaries, infestation by Tgroglyphusfari ..------------------------------ 14
"Grocer's itch," caused by mites of genus Glyciphagus --------------------- 12
Tyroglyphida ---------------------------------.5
Hair in furniture, food of Tyroglyphid-e ----------------------------------- 5
Ham, smoked, food of Carpoglyphus passularun --------------------------- 25
Hams, food of Tyroglyphide --------------------------------------------- 5,26
Hay, food of Tyroglyphide ----------------------- ---------------------- 5
Hemisarcoptes coccisugus, Acarus malu. probably a synonym ------------------ 7
heteromorphus. (See Tyroglyphus heteronorphus.)
Histiogaster, carpio the type species --------------------------------------- 23
entomophagus. (See Monieziella entomophagus.)
americanum n. sp., description and food habits ----------------- 11-12


Histiostoma brevipes n. sp., description and food habits ---------------------- 11
description of genus ------------------------------------------ 10
descriptions of species ----------------------------------------1 0-12
gracilipes n. sp., description ----------------------------------- 10-11
pectineum =rostroserratus -------------------------------------- 10
pyriforme, comparison with tiosotoiua brecipes ------------------ 11
rostroserratus the type species ---------------------------------- 10
table of species ----------------------------------------------- 10
hyacinthi. (See Rhizoglyphus hyacinthi. )
House, infestation by Tyroglyphu.s longio, ---------------------------------- 15
Hypopial stages --------------------------------------------------------- 10
Hypopus stage, of Tyroglyphide ------------------------------------------ 6- 7
Insects, in collections, food of Tyroglvl)hid-e ------------------------------ 5
Tyroglyphide attached to Certain species ------------------------ 5
kraneri. (See Tyroglyphus kraimeri.)
Lecauiuni, association of TiroglyphAus cocciphilus ----------------------------- 17
Lilies, Bermuda, infestation of bulbs by Rhizoglyphits hyqacinthi .............5, 21
lintneri. (See Tyroglyphus lintneri. )
longior. (See Tyroglyphits longior. )
longipes. (See Mfonieziella longipes.)
longitarsis. (See Rhizoglyphius iongitarsis.)
malus. (See Acarus mals and Tyroglyphus muwis.)
Mattresses, infested by Tyroglyphide ------------------------------------ 5
Mealy bug, association of Tyroglqphi.s cocciphlis ---------------------------- 17
Meats, dried, food of Tyroglyphide. .. ..--------------------------------- 5
smoked, food of TyroglyphidT ------------------------------------ 26
Mice, Tyroglyphidce in nests -----.--------------------------------------- 5
mrgnini. (See Rhizoglyphus mvgnjui.)
Mills, infestation by Tyroglyphisfa'inx --------.-------------------------- 14
Monieziella (ingusta n. sp., description and food habits ----------------------- 23
brevitarsis n. sp., description and habits --.-_--. ----------------- 24
description of genus -----------------------------------------23
descriptions of species ------------------------------------23-24
enemies of scale-insects ---------------------------------------- 5
entomophagus the type species ---------------------------------- 23
longipes n. sp., description and habits ------------------------- 23-24
on San Jose scale, bibliographic reference ----------------------- 26
Moles, Tyroglyphidfe in nests ---------------.-------------------------- 5
Mushrooms, food of Tyroglyphidv --------------------------------------- 5
Tyroglyphus lintneri ----------------------------------. 7,15, 26
mycophagus. (See Tyroglyphus mycophagus.)
Mytilaspis, food of lionieziella longipes ------------------------------------- 24
obesus. (See Glyciphagus obesits.)
Onions, infestation by Rhizoglyphus rhizophagus ----------------------------- 22
Orange, decaying, infestation by Tyroglyphus aniericanus -------------------- 16
Orchids, infestation by Rhizoglyphus hyacinthi ------------------------------ 21
Osmia, host of Trichotarsus osmie ----------------------------------------- 26
osmix. (See Trichotarsus osmix.)
passularum. (See Carpoglyphus passula rum.)
pectineurn. (See Histiostoma pectineum.)
Phycobius anonymus, identity ---------------------------------------------- 9
Phylloxera, of grape, Tyroglyphus phylloxerx err-oneously considered enemy - 7


phylloxerx. (See Rhizoglyphius phylloxerx.)
Pillows, infested by Tyroglyphid- ........................................ 5
Pine cones, infestation by Rhizoglyphus phylloxer .------------------------- 20
Plums, rotten, infestation by Tyroglyphus arnericanus ----------------------- 16
Pollen of honey bees, food of Carpoglyphus passularum ---------------------- 25
Potato plants, young, infestation by Rhizoglyphus phylloxer ------------------ 20
Potatoes, rotten and scabby, infestation by Rhizoglyphus phylloxerwl ----------- 20
py/ riforme. (See llistiostom pyriforme.)
Rhizoglyphus gilis, possibly a synonym of Tyroglyphus hetercnnorphus --------- 18
(lescription of genus ----------------------------------------- 19
descriptions of species --------------------------------------- 20-22
erlhropus, identity ----------------------------------------- 21
R vhizogl?'phous phylloxera not a synonym -------------- 20
Robin, synonym of hyacthi .------------------------ 19
elongau n. sp., description and food habits -------------------- 22
hyacinth1), 1)ibliographic reference ----------------------------- 27
Boisd. --R. echinopus Mgn ------------------------- 20
description and food habits -------------------------- 21
enemy to bulbs and roots ---------------------------- 5
the type species ----------------------------------1
in decaying matter with tHistiostoma qmyertcanumi ---------------- 12
injury to roots of vine, stems of carnation, and wax on grafts.. 5
ion gitaris ii. sp., description an( food habits ------------------- 22
mg--i, identity..----------------------------------------- 21
phylloxer, (es(.ription and food habits ------------------------ 20
hot a sVnoIyiii of ik. echinopus ---------------------- 20
rIzoph(1gu n. sp., (lescription and food habits --------------- 21-22
table of species ---------------------------------------- 19
l(irsob's n. sp., description awd food habits ---------------------- 20
rhizophagus. (See Rhizogfl yph us rhizoplhugus. )
ribis. (See Acurus ib is. )
Rice, infestation by Tyroglypirus americanus -------------------------------- 16
robustus. (See (lyciphagus robustus. )
Roots, food of Tyroglyphid ---------------.---------------------------- 5
rostroserrt us. (See tlstiostoma rostroserralu..)
Sap, food of Tyroglyphid ------------------------------------------------ 5
Scale-insects. (See also Aspidiotus, Coccids, and Mytilaspis.)
food of Tyroglyphid -------------------------------------- 5
Mon ieziella enemies ---------------------------------------- 5
oyster-shell, association of Tyroglyphus cocciph ilus -------------------- 17
San Jose. (See also Aspidiotus. )
food of Monjeziella breritarsis----------------------------- 24
parasite ------------------------------------------------ 26
Seeds, food of Tyroglyphida --------------------------------------------- 5
infestation by Glyc iphagus robustus --------------------------------- 13
Tyroglyphus .farinx ----------------------------------- 14
siro. (See Tyroglyphus siro. )
Sporotrichum globuliferum, food of Tyroglyphus siro, bibliographic reference ... 27
Sugar beets. (See Beets, sugar.)
food of Tyroglyphidae ---------------------------------------------- 5
mites (see also Glyciphagus) --------------------------------------5
tarsalis. (See Rhizoglyphus tarsalis.)
terminals. (See Tyroglyphus terminalss)



Trichodactylus anonymous, identity ----------------------------------------- 9
xylocopw. (See also Trichotarsus xylocopw. )
in California, bibliographic reference -------------- 26
Trichotarsus, association with bees ---------------------------------------- 25
description of genus ------------------------------------------ 25
hypopial stages --------------------------------------------- 10
osmix, bibliographic reference -------------------------------- 26
on Osnia from New York ------------------------------ 26
the type species -------------------------------------- 25
records of species in United States .-------------------------- 25-26
xylocopx, bibliographic reference ------------------------------ 26
recorded on Xilocopa from California ---------------- 25
Tyroglyphidae, bibliography -------------------------------------- 2-27
economic importance -------------------------------------- 5
European species in United States -------------------------- -9
food habits ----------------------------------------------- 5
genera --------------------------------------------------- 9-26
Hypopzts stage -------------------------------------------- 6-7
previous work on species in United States ------------------- 7-8
structural characteristics ----------------------------------- 6
table of genera -------------------------------------------- 9-10
transformations ------------------------------------------- 6-7
Tyroglyphus, Aleurobius a synonym -----------------------------------1 14
acaericanus n. sp., description and food habits ------------------ 16
is it T. siro? ------------------------------------- 16
references in literature as T. siro and 1. longior. 16
armripes n. sp., description and food habits ------------------- 18-19'
breviceps n. sp., description and food habits -------------------- 17
cocciphtilus n. sp., description and habits --------------------1-17
crassipes, bibliographic reference ----------------------------- 26
uncertain origin and identity --------------------- 7
curtus, bibliographic reference -------------------------------- 26
uncertain origin and identity ------------------------
description of genus ----------------------------------------- 3
descriptions of species --------------------------------------- 14-19
farinw, bibliographic reference, as T. siro ---------------------- 27
description and habits -------------------------------- 14
heteromorph as, bibliographic reference ------------------------- 26
description and food habits --------------------- 18
injury to roots of carnations -------------------- 8
Rhizoglyphus agilis possibly a synonym -----------is
krameri, identity ---------------------------- 18--9
lintneri, allied to Canestrini's conception of Tyroglyphus siro ----- 7-8
bibliographic references ----------------------------- 26
description and food habits --------------------------- 15
infestation of mushrooms ---------------------------- 7-8
longior, bibliographic references ---------------------------- 26-27
description and habits ------------------------------- 14-15
identity -------------------------------------------- 7-9, 16
malus, identity ---------------------------------------------- 23
mycophagus, comparison of Tyroglyphus terminalis with Michael's
figure ---------------------------------------- 18
identity ---------------------------------------- 8-9


Tyroglyphus, not a synonym of Acarus -------------------------------------
phylloxerw, bibliographic reference ----------------------------
=Rhizoglyphus ...................................
not predaceous, but a root feeder ............
siro, bibliographic references under this name ------------------
the type species ........................................
uncertain identity ---------------------------------------
table of species .............................................
terminalis n. sp., description and food habits --------------------
Vine, injury to roots by Rhizoglyphus --------------------------------------
Wheat, infestation by Troglyphus americanux ...........................
W ine, food of Tyroglyphid e -----------------------------------------------
Xylocopa, host of Trichot(frsus xylocop;e------------------------------
xylocopx. (See Trichodactylws xylocopw and Thichotarsus xylocopae.)

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