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Molecular Phylogenetics of the Riodinidae (Lepidoptera)


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MOLECULAR PHYLOGENETICS OF THE RIODINIDAE (LEPIDOPTERA) By JONATHAN WALTER SAUNDERS A THESIS PRESENTED TO THE GRADUATE SCHOOL OF THE UNIVERSITY OF FLORID A IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE DEGREE OF MASTER OF SCIENCE UNIVERSITY OF FLORIDA 2010 1

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2010 Jonathan Walter Saunders 2

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To my Mom and Dad who fostered my love of the natural world and taught me the importance of education 3

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ACKNOWLEDGMENTS I thank the members of my committee for their encouragement and mentoring. And specifically I thank Dr. Tom Emmel for his multifaceted support and positive, uplifting spirit towards me, Dr. Charlie B aer for his generosity and forthrightness, and Dr. Mike Miyamoto for his patience, joke s, and ability to put my mind at ease about graduate school. I thank Mike Perry for all of his help getting me started on this project and Dr. J.D. Turner for shar ing with me both his butterflies and his love and knowledge of them. I also thank Dr. Rebecca Kimbal l for allowing me to use her facilities and answering my questions as well as Dr. Ed Braun, Dr. David Reed, and Julie Allen for being so willing to help me with my many questions. I thank my wife for her loving patience and for her support whic h allowed me to finish my study. And, I thank God for the ability to study his world full of wonders. 4

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TABLE OF CONTENTS page ACKNOWLEDG MENTS..................................................................................................4LIST OF TABLES............................................................................................................7LIST OF FI GURES..........................................................................................................8LIST OF OB JECTS.........................................................................................................9LIST OF ABBR EVIATIONS...........................................................................................10ABSTRACT ...................................................................................................................11CHAPTER 1 INTRODUC TION....................................................................................................122 MATERIALS A ND METHOD S................................................................................29Sequence Data Preparati on....................................................................................29Phylogenetic Analyse s............................................................................................303 RESULT S...............................................................................................................39Phylogenet ics.........................................................................................................39Myrmecoph ily.......................................................................................................... 404 DISCU SSION.........................................................................................................52Taxonom y...............................................................................................................52Phylogenetics and Gene Tr ees...............................................................................56Myrmecoph ily.......................................................................................................... 583G<2G<1G.......................................................................................................59The Hypotheses Ar e Not I ndependent .............................................................60The Indirect Method.........................................................................................60Conclusi on..............................................................................................................62APPENDIX A COMPARISON OF 3 PARTITION ING SCHEMES FOR THE TOTAL EVIDENCE AN ALYSES..........................................................................................69B BAYESIAN TREES WITH POST ERIOR PROBABI LITIES.....................................70C RANDOM TREES USED TO GENERA TE A NULL DISTRIBUTION FOR THE MYRMECOPHILY HYPOTHESE S.........................................................................74 5

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D PROCEDURE FOR SCORING RANDOM TREES WITH A WORKED EXAMPLE ...............................................................................................................75Procedur e...............................................................................................................75Worked Exam ple....................................................................................................78E SCORED RAND OM TREES...................................................................................89F RANDOM TREE NULL DIST RIBUTION RE SULTS...............................................90REFERENCE LIST........................................................................................................92BIOGRAPHICAL SKETCH ............................................................................................97 6

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LIST OF TABLES Table page 1-1 Taxonomy of riodinid specimens for this study with their respective identificatio n number s.........................................................................................22 2-2 Adapted PCR reaction components from (Nikla s Wahlbergs website: http://nymphalidae.utu.fi/N ymphalidae/Molecu lar.htm).......................................35 2-3 Primers of genes sequenced and their re spective s ources................................36 2-5 Sequences and accession numbers for three outgroups (denoted by *) and seven riodinids obtained from GenBank (Benson et al. 2006)............................38 3-1 Characteristics of the DNA sequen ce dat a.........................................................42 3-2 Likelihood values and AIC values from RAxML likel ihood analyses...................43 3-3 Parsimony score statistics fo r 1G, 2G, and 3G when mapped onto 100 random tree s......................................................................................................51 4-1 Parsimony scores from mapping gains and losses of myrmecophily onto the Myrmecophily Tree (F igure 4-2) .........................................................................65 4-2 Results from comparison of the hypotheses mapped on the Myrmecophily Tree next to the averages and 95% confidence intervals taken from random trees...................................................................................................................67 7

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LIST OF FIGURES Figure page 1-1 Phylogeny depicting evolutionary re lationships of butterfly families adapted from Wahlberg et al. 2 005..................................................................................21 1-2 Diagram of caterpillar bodies co mparing the auditory, feeding, and chemosensory emitting ant organs in three bu tterfly clades: Nymphidiini/Lemoniini, Eurybini, and Lycaeni dae...................................................................................25 1-3 Subfamily and tribal relationshi ps inferred from parsimony analysis on morphological data. This phylogeny is adapted from Harvey 1987....................26 1-4 A parsimony phylogeny from Campbell et al. (2000) based on the wingless gene excluding third-position transiti ons with bootstrap support values less than 50% collapsed into a polytomy...................................................................27 1-5 Two taxon sampling schema represented by A) and B) with the same number of taxa (and in this ca se, even the same topology)...............................28 3-1 RAxML most likely tree for t he 3 genes combined and partitioned by gene and by codon position, higher taxa i ndicated by colo red branc hes.....................44 3-2 RAxML most likely trees for indi vidual genes, higher taxa indicated by colored bran ches................................................................................................45 3-3 Consensuses of bootstrapped co mbined and individual RAxML trees (bootstraps indica ted at nodes) ..........................................................................46 3-4 50% majority rule consensus tr ee of the three most likely gene trees................50 4-1 Morphological phyloge netic hypothesis after Harvey 1987 including his subfamilies and tribes compared to my mo lecular phylogenetic hypothesis.......644-2 The Myrmecophily Tr ee.....................................................................................67 4-4 Adaptation of Cam pbells (1998) molecular phylo geny with myrmecophilous character state indicated by square next to genus.............................................68 D-1 Example of how to ma rk a phylogeny as in st ep.................................................75 D-3 Two interpretations of the evolutio n of myrmecophily based on the same tree. ...........................................................................................................................76 D-4 Figure D-4. Further ex planation for st ep 6) and 7)..............................................77 D-5 Step by step method for scoring a random tree (here it is random tree number 94 which is the most co mplex................................................................78 8

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LIST OF OBJECTS Object page C-1 Random Trees....................................................................................................74E-1 Scans of 100 sco red random trees.....................................................................89 9

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LIST OF ABBREVIATIONS Myrmecophily Tree The tree depicted in Figure 4-2, that is, the most likely tree based on the best partitioning scheme from AIC comparison mapped with the myrmecophilous/amyrmecoph ilous character states and gains/losses of said trait. 1G The evolutionary hypothesis st ating that myrmecophily evolved exactly one time in butterflies. 2G The evolutionary hypothesis st ating that myrmecophily evolved exactly two times in butterflies. 3G The evolutionary hypothesis st ating that myrmecophily evolved exactly three times in butterflies. 10

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Abstract of Thesis Pres ented to the Graduate School of the University of Florida in Partial Fulf illment of the Requirements for t he Degree of Master of Science MOLECULAR PHYLOGENETICS OF THE RIODINIDAE (LEPIDOPTERA) By Jonathan Walter Saunders May 2010 Chair: Charles Baer Cochair: Thomas Emmel Major: Zoology The butterfly family Riodinidae has many spec ies that exhibit diverse traits, but the taxonomy of the family is still unclear and a robust phylo geny derived from molecular data is lacking. In this study I generate a phy logeny of 68 novel riodinid taxa. I use two nuclear and one mitochondrial gene to investigat e the evolutionary relationships of the Riodinidae in likelihoo d and Bayesian analyses. The resulting phylogenies provide greater taxonomic resolution for the Riodi nidae, especially in the deeper level relationships. They support many but bri ng into question a few of the previously described taxonomic relationships. The phylogenies are also used to address hypotheses about the evolution of myrmecophily in butterflies. The hypothesis of three independent gains of myrmecophily in the bu tterflies is supported over the two gains hypothesis and the one gain hypothesis. The resu lts are also consistent with a lack of losses in the evolution of myrmecophily. 11

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12 CHAPTER 1 INTRODUCTION The metalmark butterflies of the family Riodinidae are known for their small size, their brilliant metallic colors, and the short ened forelegs of the ma les (Borror et al., 1989, DeVries, 1997). Riodinid species exhibit substantial phenotypic diversity. Various riodinid species resemble members of all ot her butterfly families. The metalmarks utilize a wide array of host plant families. The adults display complex mating behaviors. In some species the larvae engage in ant-caterpillar symbioses which are usually mutualistic, and are striking examples of my rmecophily (Devries 1997, Pierce et al. 2002). The position of the family Riodinidae in the larger phylogeny of Lepidopterans is well-resolved (see Figure 1-1), but the relationships of genera within the family are much more poorly understood (Ackery et al. 1999, de Jong et al. 1996, Kristensen 1976, Wahlberg et al. 2005). T herefore, a well-resolved phylogeny of the family is needed to contribute to our understanding of phenotypic evolution. Unfortunately, phylogenetic resolution has been impeded by the considerable phenotypic diversity exhibited by the riodinids, including the in tra-family and inter-fam ily mimicry mentioned earlier. There has been little progre ss in the understanding of phylogenetic relationships within the family over the 150 years of published major systematic treatments on the gr oup (Bates 1868, Stichel 1928, 1930-31, Harvey 1987, and Callaghan and Lamas 2001). The taxonomic rela tionships based on Harveys treatment of the Riodinidae are outlined in Table 1-1 as well as in the keys of the phylogenies in the results section. The subf amilies and tribes correspond to the phylogeny in Figure 13.

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Although a phylogeny of Riodinids is cha llenging to establish using morphological characters, its usefulness to the study of ev olution of metalmarks is obvious. The wing design diversity in the riodinids includes copy ing or mimicking species of at least eight other lepidopteran families. Riodinid genera also often contain aposomatic and cryptic species (see Dabrera 1994). Riodinid larvae also exhibit great divers ity in body morphologies and patterns. Their survival strategies in dealing with other insects vary from repelling insects with long bristles, to hiding inside or under leaves, to feeding ants in exchange for protection (DeVries 1997). The phenotypic diversity and plasticity found in this group are reasons why a molecular phylogeny is needed to under stand the evolution of the Riodinidae. One of the phenotypic traits of particu lar interest is myrmecophily, where caterpillars feed ants in exchange for protec tion by the ants from potential predators, including other ant species. Among butterflie s myrmecophily is unique to the Riodinidae and their sister family the Lycaenidae. As di scussed by DeVries (DeVries 1990, DeVries 1997), Non-myrmecophilous riodinids tend to hav e thicker cuticles and long setae that keep the ants at a distance, whereas myrme cophilous riodinids feed and communicate with ants in exchange for the ants active protection from predator s and parasitoids and do not have long, thick setae. Myrmecophily in these butterfly larvae is associated with three functional larval organs: an organ to feed ants, an organ to communicate acoustically with ants, and tentacle organs that appear to release chemicals which induce the ants to take a defensive posture and protect the caterpillar. Members of the Eurybiini tribe are an exception and lack tent acle organs. These organs are illustrated in Figure 1-2. 13

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The ant-feeding organ for all myrmecophilous riodinids, called the tentacle nectary organ, is found on abdominal segment 8 (A8) The ant-feeding organ in lycaenids, the dorsal nectary organ, is on abdominal segment 7 (A7) (DeVries 1991b), as denoted in Figure 1-2. When ants tap near the orifices of these or gans with their antennae, the caterpillars secrete a nutritious, sugaror amino-acid containing fluid through the nectary organs (DeVries 1990, 1997). Drops of this sweet fluid attract the ants and they widely drink it. Other studies have shown that acoustic co mmunication is important for regulating ant behavior towards caterpillars (Devries 19 90, Travassos and Pierce 2000). There are three ways in which the butterfly caterpillars produce stridulations to communicate with ants. These organs are shown in Figure 1-2: (1) Vibratory papillae (Ross 1966, DeVries 1997), (2) cuticle on a cuticle file system (Hill 1993), and (3) a cuticle/cranium spine system (DeVries and Penz 2002). These st ridulation mechanisms are found in the following clades respectively: (1) Riodinidae: tribes Lemoniini and Nymphidiini, (2) Lycaenidae, and (3) Riodinidae: tribe Eurybiini. As depicted in Figure 1-2, the tentacle organs, which appear to release signaling chemicals, are found on thoracic segment 3 in one riodinid clad e, absent in the Eurybiini (Riodinidae), and found on A8 in lycaenids (Kitching 1983, Devries 1997). DeVries (1991a, 1997) described three potential hypotheses regarding the evolution of myrmecophily in butterflies, but it is not yet know n which hypothesis is correct. 1G: Myrmecophily was gained once in the ancestor of the riodinid and lycaenid families and subsequently lost in the Lycaenidae and Riodinidae. 14

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2G: Myrmecophily was gained once in Riodinidae and once in Lycaenidae and also subsequently lost in the Riodinidae. 3G: Myrmecophily was gained on three independent occasions in the lycaenids and in the two riodinid clades (Eur ybiini and Lemoniin i/Nymphidiini). Please note that the phylogeny is curr ently unresolved to the degree that the frequency with which myrmecophily was lost in the different hypotheses is not entirely known for the multiple gains hypotheses. The weight of current evidence suggests that there were three independent gains of myrmecophily. Two differences are bes t explained by the 3G; the homoplasious nature of the three different acoustic or gans and the three different semiochemical organs, as indicated in Campbells thesis (1998). However, there are similarities between the ant organs which leave the possibilit y for either 1G or 2G to be correct, as supported by Harvey (1987). All three clade s have retractable, glandular, secretory organs and communicate with glandular ti ssue via ducts and setae (DeVries 1997, DeVries 1988). Additionally, all three cl ades have a paired ant organ on abdominal segment eight (A8) (see Figure 1-2). Also at first glance, there appears to be an underlying developmental mechanism responsib le for organs in A8 because they all arise from the same segment. Also, Stichel (1924) describes a Mesosemia moesia larvae as having a paired gland on A8 which corresponds to the location of the tentacle nectary organs of the other riodinids and t he tentacle organs of the lycaenids (as in Figure 1-2), but he does not refer to myrme cophily (Harvey 1987). Thus, it is still undetermined how myrmecophily developed in butterflies. In this study I will use a molecular phylogeny to test these hypotheses independently of the mo rphological characters being exam ined. This would allow for 15

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the presence or absence of myrmecophily to be mapped onto the phylogeny and thus allow for inference regarding evoluti onary gains and losses of this trait. 16

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17 The previously published phylogenies of the Riodinidae are useful but by no means complete. Previous phylogenetic anal yses were performed by Stichel, Harvey, Callaghan, Lamas, and Campbell (see Figures 1-3 and 1-4). The published molecular results on the Riodinidae are limited by taxon sampling, extent of molecular loci used, and are also limited to parsimony methods (Campbell et al. 2000). All other published phylogenies are based on mor phological characters. The taxon sampling scheme is import ant to break up long branches and to mitigate the effects of long branch attrac tion (Wheeler 1992, Gr aybeal 1998, Poe and Swofford 1999, Bergsten 2005). To better un derstand familial relationships through phylogenetics, sampling more genera from di fferent tribes would be better than having numerous species from just a fe w genera that do not cover all of the tribes in the family. This is because the evolutionary distances between the tribes in the first schema are greater than the evolutionary distances within the tribes in the first schema as compared to the second (see Figure 1-4). This is not onl y seen in theory but it is also exemplified in computer-simulated gene phyl ogenies and in molecular phylogenies from organisms. The second, less uniformly sampled schema de scribed above (corresponding to tree B) in Figure 1-5) has been shown to be worse in that it provides lower support for the correct trees (correct trees ar e assumed to be those used by the computer to make the simulated sequences in the computer-simul ated cases and the trees recovered using more genes, more taxa, and morphological data for the real-lif e data sets) and even higher support for incorrect trees in evolut ionary analyses (Pollock et al. 2002, Zwickl and Hillis 2002). More sequence data also prov ide greater phylogenetic signal in terms of accuracy and resolution due to more inform ation for the analyses (Zwickl and Hillis

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2002, Hillis et al. 2003). Not only do more se quence data provide the opportunity to find more informative substitutions to infer re lationships between genes, but the sequences themselves are used to determine the appr opriate models of evolution and their parameters for the analyses. Therefore more sequence dat a would allow for more informed parameters and, ther efore, a better analysis. Similarly, phylogenetic maximum likelihood as well as Bayesian methods have generally provided better molecular phylogenet ic trees than parsimony. They recover trees with topology that is closer to that of the true tree for both simulated and real-life data (under the assumption above that true tree is the one used by the computer to make the simulated sequence in the comput er-simulated cases and the trees recovered using more genes, more taxa, and morphology for the real-life data sets) as found by Huelsenbeck 1998, Alfaro et al. 2003, and Felsenstein 2004, and many more. This is because in true parsimony methodology the assu mption is that all substitutions are equally likely and the probability of back substitu tions or the possibility that different sites mutate at different rates are not ta ken into account while in maximum likelihood and Bayesian methods parameters are inferred to weight each substitution type, to model bias due to different base frequencies of each nucleotide, to account for rate heterogeneity between different sites, and to account for homoplasy. In maximum likelihood analyses these param eters are inferred from a pr eliminary tree while in Bayesian analyses they are calculated m any times as the analysis progresses. Rate heterogeneity is mainly due to differ ent selection pressures on different codon positions and differences in the coded am ino acids. Differing selection pressures on the different codon positions are seen bec ause, generally speaking, there are more 18

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third position substitutions that will not alter t he amino acid than there are in the third or second positions and similarly there are more first position substitutions that will not alter the amino acid than there are se cond position substitutions. This rate heterogeneity can be further accounted for in a phylogenetic analysis by partitioning the matrix by codon position. An unpublished phylogeny of the Riodini dae exists (Campbell 1998) but the sequences therein are inaccessible. To better understand evolution within the Riodinidae, a phylogeny with a larger taxon sa mpling and more loci is dearly needed. I undertook the present project in order to provide broader molecular coverage of this family and then create a sequence based phylogeny for the family. I then used the phylogeny to test the evolutionary theorie s of myrmecophily in the Riodinidae and its sister family, Lycaenidae. In this study I present a phylogenetic analysis based on the combined sequences of two nuclear genes and one mitochondr ial gene using maximum likelihood. Wingless ( wg ), Elongation factor 1-alpha ( Ef 1-a ), and Cytochrome oxidase subunit I ( CO1 ) were chosen for this study based on their phylogenet ic signal and the ability to combine these sequences with other lepidopt eran molecular data sets for a much larger analysis. Nuclear genes were chosen because of their efficacy in recovering deep phylogenetic relationships as seen in previous analyses (Hoelzer 1997, Springer et al. 2001, Lin and Danforth 2004). Wingless and Ef 1-a were previously shown to be useful for recovering deep lepidopteran evolutionary relationshi ps (Brower and DeSalle 1998, Campbell 1998, Wahlberg and Zimmerman 2000) and CO1 has been valuable in recovering shallower evolutionary relationships (Wahlberg and Zimmerman 2000). In addition to 19

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being housed in the mitochondria, which means it comes from a haploid genome and has a different cellular environm ent than the nuclear genes, Cytochrome oxidase subunit I is also likely to differ from the two nuc lear genes in evolutionary history due to its maternal inheritance and lack of recombination. The three genes are used here to strengthen congruence of i ndependent data sets. These three genes were also selected, in part, because they have been pr eviously used in lepidopteran molecular systematic work (Brower and DeSalle 1998, Wahlberg and Zimmerman 2000, Monteiro and Pierce 2001) and sequences her ein derived will allow for a broader scale molecular phylogenetic study of all lepidopterans when combined with the sequences from these other studies. 20

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Figure 1-1. Phylogeny depicting evolutionary relationships of butterfly families adapted from Wahlberg et al. 2005. 21

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22 Table 1-1. Taxonomy of riodinid specimens for this study with their respective identific ation numbers. These numbers were used to keep track of voucher specimens and corr esponding tissues (sample #) and to keep track of a specific isolation of DNA (Rio#). Note: accession numbers for 6 specimens taken from genbank can be found in Table 2-5 here they are marked with NA. Subfamily Tribe Subtribe Genus Species Sample # Rio # Riodininae Mesosemiini Mesosemiina Leucochimona aequitoralis 7617 87 Riodininae Mesosemiini Mesosemiina Leucochimona lagora 9302 33 Riodininae Mesosemiini Mesosemiina Leucochimona lagora 9306 27 Riodininae Mesosemiini Mesosemiina Mesophthalma idotea 6327 88 Riodininae Mesosemiini Mesosemiina Mesosemia latizonata 6516 85 Riodininae Mesosemiini Mesosemiina Mesosemia judicialis 8822 2 Riodininae Mesosemiini Mesosemiina Perophthalma lasus 9330 24 Riodininae Mesosemiini Mesosemiina Semomesia campanea 5223 52 Riodininae Mesosemiini Napaiena Cremna actoris 5260 53 Riodininae Mesosemiini Napaiena Hyphilaria nicia 5239 91 Riodininae Mesosemiini Napaiena Ithomiola callixena 5924 82 Riodininae Mesosemiini Napaiena Napaea eucharilla m17 89 Riodininae Helicopini Anteros allectus 7097 50 Riodininae Helicopini Helicopis endymaena 5318 54 Riodininae Helicopini Ourocnemis archytas 4621 90 Riodininae Helicopini Sarota acantus 9358 21 Riodininae Lemoniini Juditha caucana 9315 26 Riodininae Lemoniini Lemonias zygia 3950 55 Riodininae Lemoniini Synargis orestes 7605 29 Riodininae Lemoniini Thisbe fenestrella 6766 43 Riodininae Lemoniini Thysanota galena 5241 81 Riodininae Lemoniini Uraneis hyalina NA NA Riodininae Nymphidiini Nymphidiina Adelotypa balista 7807 37 Riodininae Nymphidiini Nymphidiina Calospila emylius 8532 96 Riodininae Nymphidiini Nymphidiina Menander coruscans 7339 49 Riodininae Nymphidiini Nymphidiina Pandemos pasiphae 8572 97 Riodininae Nymphidiini Nymphidiina Setabis gelasine 6330 38

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Table 1-1. Continued Riodininae Nymphidiini Nymphidiina Zelotaea phasma 4057 79 Riodininae Nym phidiini Theopina Protonymphidia senta 3948 59 Riodininae Nym phidiini Theopina Theope aureonitens 7789 65 Riodininae Nym phidiini Theopina Theope eudocia 9026 5 Riodininae Stalachtini Stalachtis calliope 6422 39 Riodininae Stalachtini Stalachtis phaedusa 7773 35 Riodininae Riodinini Caria castalia 7625 31 Riodininae Riodinini Cariomothis erythromelas 6518 41 Riodininae Riodinini Chamaelimnas iaeris 3942 57 Riodininae Riodinini Crocozona caecias 3995 62 Riodininae Riodinini Exoplisia cadmeis 6433 40 Riodininae Riodinini Lepricornis incerta 5041 80 Riodininae Riodinini Metacharis victrix 9347 23 Riodininae Riodinini Parcella amarynthina 4012 64 Riodininae Riodinini Riodina lysippus NA NA Riodininae Riodinini Ancyluris section Amarynthis meneria 5220 51 Riodininae Riodinini Ancyluris section Ancyluris eudaemon 6834 45 Riodininae Riodinini Ancyluris section Ithomeis satellites 3956 58 Riodininae Riodinini Ancyluris section Cyrenia martia 3992 63 Riodininae Riodinini Ancyluris section Isapis agyrtus 6815 44 Riodininae Riodinini Ancyluris section Melanis marathon 7612 30 Riodininae Riodinini Ancyluris section Monethe albertus 3977 61 Riodininae Riodinini Ancyluris section Necyria juturna 5966 93 Riodininae Riodinini Ancyluris section Panara pheraclus 3932 56 Riodininae Riodinini Ancyluris section Rhetus periander 3973 60 Riodininae Riodinini Ancyluris section Themone pais 7643 34 Riodininae Riodinini charis group Calephelis costaricicola 9356 22 Riodininae Riodinini charis group Charis anius 9365 19 Riodininae Riodinini charis group Detritivora barnesi 9327 25 Riodininae Eurybiini Alesa amesis 8380 84 Riodininae Eurybiini Eurybia unxia 9363 20 23

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Table 1-1. Continued Riodininae Symmachiini Esthemopsis lithosina 7813 32 Riodininae Symmachiini Mesene xanthe 7450 28 Riodininae Symmachiini Phaenochitonia pyrsodes 6926 47 Riodininae Symmachiini Symmachia falcistriga 8849 8 Riodininae Incertae Sedis Apodemia mormo NA NA Riodininae Incertae Sedis Argyrogrammana trochilia 6709 42 Riodininae Incertae Sedis Emesis temesa 6837 46 Riodininae Incertae Sedis Emesis sp. NA NA Riodininae Incertae Sedis Emesis mandana NA NA Riodininae Incertae Sedis Roeberella leneates 7457 36 Riodininae Incertae Sedis Calydna catana 6346 83 Riodininae Incertae Sedis Echenais thelephus 7168 48 Riodininae Incertae Sedis Pachythone lateritia 9050 18 Euselasiinae Euselasiini Euselasia serapis 8816 1 Euselasiinae Euselasiini Euselasia orfita NA NA Euselasiinae Euselasiini Euselasia sp. NA NA Euselasiinae Euselasiini Methone cecilia 8523 95 Styginae Stygini Styx infernalis NA NA Nemeobiinae Hamearis lucina NA NA 24

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25 Figure 1-2. Diagram of caterpillar bodies comparing the auditory, feeding, and chemosensory emitting ant organs in three bu tterfly clades: Nymphidiini/Lemoniini, Eurybini, and Lycaenidae. (Adapted from DeVries 1997)

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Figure 1-3. Subfamily and tribal relations hips inferred from parsimony analysis on morphological data. This phyloge ny is adapted from Harvey 1987. 26

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Figure 1-4. A parsimony phylogeny from Campbell et al. (2000) based on the wingless gene excluding third-position transiti ons with bootstrap support values less than 50% collapsed into a polytomy. 27

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Figure 1-5. Two taxon sampling schema r epresented by A) and B) with the same number of taxa (and in this case even the same topology). Tree A) represents a phylogeny with multiple genera per tribe whereas tree B) represents a phylogeny with multiple species per tribe but only one genus for each tribe. Longer clades between tribes are apparent in tree B) which is not ideal when analyzing inter-tribal relations hips in a family. Tree C) illustrates how adding a new genus, denoted by the asterisk, in Tribe 1 can break up the longer clade. 28

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CHAPTER 2 MATERIALS AND METHODS Sequence Data Preparation For my analysis I sampled 3 of the 4 subf amilies, all 11 tribes, and 69 of the 150 genera of Riodinidae described in Harvey (1987) along with 3 lycaenid outgroups, and I sampled six of the 69 genera twice (duplicat e genera) for a total of 78 specimens. Of these, I sequenced genes from a total of 68 new specimens (noted in Table 2-1). Samples of live specimens were stored in 100% EtOH. Using Qiagen DNeasy kits, the DNA was extracted at the Univ ersity of Florida in the Ba er genetics laboratory or the McGuire center genetics laboratory from abdominal tissue when possible, and from legs where abdomens were not available. I amplified three intr onless coding fragments: cytochrome oxidase subunit I (COI ), a mitochondrial gene, and wingless (Wg ) and Elongation Factor 1-alpha ( EF-1 ), two nuclear genes. This was done in 0.2ml PCR tube strips. The PCR mixture formula is found in Table 2-2. I amplified EF1a in 3 parts, CO1 in 2 parts, and wingless in one part (Table 2-3) on an Eppendorf Mastercycler thermocycler using 3 different protocols for the 6 fragments (see Table 2-4). I stored the amplified products in a 4 C refrigerator. I checked to see if amplification occu rred by running 5 l PCR product and 1 l 5x DNA loading dye in agarose gel at 100V for 20 minutes in a 0.5xTBE buffer with a 100bp ladder. I organized 96 PCR products to be sequenced at a time and submitted three 96well plates (of PCR product and correspondi ng forward and reverse primers ) to the Interdisciplinary Center for Biotechnology Research custom sequencing laboratory at 29

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the University of Florida where they purified and sequenced the PCR products using Sanger Sequencing and generated ABI 3130 sequences. Phylogenetic Analyses I assembled these generated sequences into contiguous sequences, edited the sequences for quality, deletin g ambiguous bases and a portion of the single stranded regions, and cut off the primers using the program SEQUENCHE R 4.8 (Gene Codes Corp., Inc., 1994). I exported the sequences in NEXUS format and aligned them and riodinid sequences taken from genbank (see Table 2-5) by eye into a matrix using MacClade 4.0 (Maddison and Maddison, 1992). I constructed and bootstrapped maximum likelihood trees usi ng RAxML 7.0.4 (Stamatakis 2006) for the three genes individually and also for the 3 genes comb ined using seven different matrices: an unpartitioned matrix, a matrix partitioned by gene, a matrix partitioned by codon position, a matrix partitioned by codon pos ition and further partitioned by gene, and three matrices where one of the three genes was partitioned from the other two and used PAUP 4.0 (Swofford 2003) to comput e consensus trees from the bootstrapped trees. More specifically I used the GTRMIX m odel of evolution in this analysis. GTRMIX incorporates a general time reversible (GTR) model that assumes a symmetric substitution matrix while allowing for differ ent base frequencies and di fferent substitution rates for each base. The MIX means that to account for rate heterogeneity among sites, it starts by categorizing each site into one of 25 rate categories, optimizes branch lengths and topology, and then replaces the categories with a gamma distribution and then further optimizes the tree using the gamma distribution for different rate categories. Thusly, I constructed seven different to tal evidence phylogenies (all three genes were used) based on seven different parti tioning schemes and compared them using 30

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AIC in order to determine the best scheme, that is, the most likely model and tree after accounting for over paramet erization. These schemes are as follows: an unpartitioned matrix, a matrix partitioned by gene, a matr ix partitioned by codon position, a matrix partitioned by gene and by codon position, and three matrices where one gene is partitioned from the other tw o genes and found that the matrix partitioned by gene and codon position is the best (see Table 3-2). Th is and the individual phylogenies are the trees presented here. For a co mparison of the most likely trees from the analyses based on three of the partitioning schemes see Appendix A. I also constructed multiple phylogenies using bayesian inference analysis (MrBayes, Huelsenbeck and Ronquist, 2001; Ronquist and Huelsenbeck 2003) using the GTR+I+G for the same partitioning schem es; I ran each for 10 million generations with a burnin of 1 milli on generations. The results of the Bayesian analysis did not differ significantly from those of the likeliho od analysis and are present ed in Appendix B. To assess the relative probabilities of di fferent numbers of independent origins of myrmecophily I scored the best maximum li kelihood tree for presence of absence of myrmecophily, leaving blank the genera of wh ich no data on myrmecophily was found. This best tree was the most likely tree t hat RAxML generated based on the best (from AIC comparison) partitioning sch eme for the 3 genes combined Hereafter this scored, most likely tree will be referred to as the Myrmecophily Tree. I also scored 100 random trees, generated using PAUP 4.0 (Swofford 2003) using the same taxa with a constrained, myrmecophilous outgroup as in the likelihood analysis, in the same manner to generate a null distri bution. Lack of myrmecophily, and with it possible loss of the trait, was either recorded based on an indication of amyrmecophilous larvae or 31

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inferred from notes on presence of long ant -repelling setae in riodinid larvae. These data were found in Harvey 1987, Horvitz et al. 1987, Devries 1997, and Kaminski 2008. 2G and 3G were tested by finding the most parsimonious evolutionary reconstruction for both 2G and 3G. I assigned gains according to 2G and 3G and counted the minimum number of losses needed per tree and then tallied the lo sses and gains for both hypotheses for each tree. I found the mini mum number of losses needed per tree in a similar fashion for the 1G. For more on how the phylogenies were scored, a worked example, and scanned copies of all 100 h and-scored random tree s see Appendix C, Appendix D, and Appendi x E respectively. 32

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Table 2-1. Novel specimen names, sample numbers assigned in the field and Rio numbers assigned after isolations. Genus Species Sample # Rio # Panara pheraclus 3932 56 Chamaelimnas iaeris 3942 57 Protonymphidia senta 3948 59 Lemonias zygia 3950 55 Ithomeis satellites 3956 58 Rhetus periander 3973 60 Monethe albertus 3977 61 Cyrenia martia 3992 63 Crocozona caecias 3995 62 Parcella amarynthina 4012 64 Zelotaea phasma 4057 79 Ourocnemis archytas 4621 90 Lepricornis incerta 5041 80 Amarynthis meneria 5220 51 Semomesia campanea 5223 52 Hyphilaria nicia 5239 91 Thysanota galena 5241 81 Cremna actoris 5260 53 Helicopis endymaena 5318 54 Ithomiola callixena 5924 82 Necyria juturna 5966 93 Mesophthalma idotea 6327 88 Setabis gelasine 6330 38 Calydna catana 6346 83 Stalachtis calliope 6422 39 Exoplisia cadmeis 6433 40 Mesosemia latizonata 6516 85 Cariomothis erythromelas 6518 41 Argyrogrammana trochilia 6709 42 Thisbe fenestrella 6766 43 Isapis agyrtus 6815 44 Ancyluris eudaemon 6834 45 Emesis temesa 6837 46 Phaenochitonia pyrsodes 6926 47 Anteros allectus 7097 50 Echenais thelephus 7168 48 Menander coruscans 7339 49 Mesene xanthe 7450 28 Roeberella leneates 7457 36 Synargis orestes 7605 29 Melanis marathon 7612 30 Leucochimona aequitoralis 7617 87 Caria castalia 7625 31 33

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Table 2-1. Continued Genus Species Sample # Rio # Themone pais 7643 34 Stalachtis phaedusa 7773 35 Theope aureonitens 7789 65 Adelotypa balista 7807 37 Esthemopsis lithosina 7813 32 Alesa amesis 8380 84 Methone cecilia 8523 95 Calospila emylius 8532 96 Pandemos pasiphae 8572 97 Euselasia serapis 8816 1 Mesosemia judicialis 8822 2 Symmachia falcistriga 8849 8 Theope eudocia 9026 5 Pachythone lateritia 9050 18 Leucochimona lagora 9302 33 Leucochimona lagora 9306 27 Juditha caucana 9315 26 Detritivora barnesi 9327 25 Perophthalma lasus 9330 24 Metacharis victrix 9347 23 Calephelis costaricicola 9356 22 Sarota acantus 9358 21 Eurybia unxia 9363 20 Charis anius 9365 19 Napaea eucharilla m17 89 34

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35 Table 2-2. Adapted PCR r eaction components from (http://nymphalidae.utu.fi/N ymphalidae/Molecular.htm). Reagent Volume deionized H2O 12.5 l 10x buffer 2.0 l MgCl2 2.0 l F Primer (1mM) 1.0 l R Primer (1mM) 1.0 l dNTP (1.25 mM) 0.4 l AmpliTaq polymerase 0.1 l specimen DNA 1.0 l

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36 Table 2-3. Primers of genes sequenced and their respective sources Gene Bp in Gene Forward primer Reverse primer Publication Bp in Fragment Sequence (5-3) EF-1 1240 bp Starsky Monteiro and Pierce (2001) 541 bp CAC ATY AAC ATT GTC GTS ATY GG Luke Monteiro and Pierce (2001) CAT RTT GTC KCC GTG CCA KCC Cho Monteiro and Pierce (2001) 517 bp GTC ACC ATC ATY GAC GC Verdi Monteiro and Pierce (2001) GAC ACC AGT TTC IAC TCT GCC EF51.9 Monteiro and Pierce (2001) 511 bp CAR GAC GTA TAC AAA ATC GG EFrcM4 Monteiro and Pierce (2001) ACA GCV ACK GTY TGY CTC ATR TC CO1 1498 bp LCO Wahlberg and Zimmerman (2000) 658 bp GGT CAA CAA TTC ATA AAG ATA TTG G HCO Wahlberg and Zimmerman (2000) TAA ACT TCA GGG TGA CCA AAA AAT CA Jerry Wahlberg and Zimmerman (2000) 829 bp CAA CAY TTA TTT TGA TTT TTT GG Pat Wahlberg and Zimmerman (2000) ATC CAT TAC ATA TAA TCT GCC ATA Wingless 403 bp LepWG1 Brower and DeSalle (1998) 403 bp ACT ICG CAR CAC GAR TGG AAT GTR CA LepWG2 Brower and DeSalle (1998) GAR TGY AAR TGY CAY GGY ATG TCT GG

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37 Table 2-4. Gene fragments and their respective thermocycler profiles adapted from Niklas Wahlbergs website (online Wahlberg accessed 1 March, 2008). CO1 95 C 5 min (both primer pairs) 94 C 30 sec 47 C 30 sec 72 C 1 min 30 sec repeat previous 3 steps 35 times 72 C 10 min 4 C hold wingless 95 C 7 min and ef51.9/EFrcM4 95 C 1 min 50 C 1 min 72 C 2 min repeat previous 3 steps 35 times 72 C 10 min 4 C hold Cho/Verdi 94 C 5 min and Starsky/Luke 94 C 30 sec 55 C 30 sec 72 C 1 min repeat previous 3 steps 35 times 72 C 5 min 4 C hold

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Table 2-5. Sequences and accession numbers fo r three outgroups (denoted by *) and seven riodinids obtained from GenBank (Benson et al. 2006) Genus Species (COI) (WG) (EF1a) Publication Euselasia orfita DQ018951 DQ018889 DQ018918Wahlberg et al. 2005 Styx infernalis DQ018949 DQ018887 DQ018916Wahlberg et al. 2005 Riodina lysippus DQ018952 DQ018919Wahlberg et al. (2005) Riodina lysippus AF233540 Campbell, Brower, Pierce (2000) Apodemia mormo AF170863 AF173403 Caterino, et al. (2001) Emesis mandana DQ018950 DQ018888 DQ018917Wahlberg et al. (2005) Uraneis hyalina AY954563 AY954593 AY954623 Braby, Vila, Pierce (2006) Hamearis lucina DQ018953 DQ018890 DQ018920Wahlberg et al. (2005) Miletus* ancon DQ018941 DQ018908Wahlberg et al. (2005) Miletus* ancon AF233550 Campbell, Brower, Pierce (2000) Poritia* erycinoides DQ018939 DQ018880 DQ018906Wahlberg et al. (2005) Curetis* bulis DQ018942 DQ018909Wahlberg et al. (2005) Curetis* bulis AF233549 Campbell, Brower, Pierce (2000) 38

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CHAPTER 3 RESULTS From the 68 novel organi sms, a total of 68 new cytochrome oxidase subunit 1 sequences, 66 new wingless sequences, and 66 new Elongation Factor 1sequences were generated. A total of 3,141 nucleotide sites were sampled in the phylogenetic analysis. Primers did not overlap for cytochrome oxidase subunit 1 and therefore this gene had a central gap of 11 bp for every sequence obtained. These were scored as missing data. No alignments required indels. Wingless (nuc) had a higher percentage of parsimony informative sites than did COI (mt) or EF 1(nuc), which had the lowest percentage of parsimony informative sites (see Table 3-1). Based on an AIC comparison, the matrix that is partitioned by gene and codon position is better than the others (see Table 3-2). Phylogenetics The results of phylogenetic analyses imply monophyly of the Lemoniini ( sensu Harvey 1987), Mesosemiini ( sensu Harvey 1987), Incertae Sedis ( sensu Harvey 1987 or Napeina sensu Hall 2003), Riodinini ( sensu Harvey 1987), all save one of the Helicopini ( sensu Reuter 1897), and all of the Symmachiini (sensu Harvey 1987) save one (see colored clades in Figure 1). T he Helicopini and Symmachiini both have one genus that is not in the tribe in the cu rrent analysis but previously described as contained in the tri be. Nymphidiini ( sensu Harvey 1987) and Emesini ( sensu Harvey 1987) are polyphyletic tribes (support indicated in Figure 3-3). Similar, poorly resolved topologies we re obtained for each gene when analyzed separately (see Figures 3-2 and 3-3, and for a consensus of the trees in Figure 3-2 see 39

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Figure 3-4). The Nymphidiin i/Nymphidiina subtribe ( sensu Lamas 2004) was supported by the analysis of all three genes w hen analyzed separately and together. The Riodina Ancyluris group ( sensu Harvey 1987), except for a slight aberration in the WG gene tree, and the Nymphidiini/Lemon iadina subtribe, except for a slight aberration in the COI gene tree, were also both recovered by analyzing each gene independently. The basal clade was recovered by COI and wingless but not by EF-1 Only part of the Mesosemiini/Mesosemiina subtribe was reco vered by analysis of the mitochondrial gene while the majority of the group was reco vered by each nuclear gene. Similarly, the topological location of Stalac htini (a mono-generic tribe in the Riodinidae) within the Nymphidiini was supported by the nuclear genes but not the mitochondrial gene ( COI ). The total evidence analysis supports the major clades (subfamilies and tribes) proposed by Harvey (1987) and Lamas et al (2004), with a few exceptions (see Fig 31).That is, all the tribes and subfamilies seen in Figure 1-3 except for the Emesini and Nymphidiini are either monophyletic or have just one mi splaced taxon in the best and bootstrapped total evidence trees. Specifica lly the exceptions to the conventional riodinid relationships are: Stalachtini and Lemoniini are both monophyle tic tribes but are also both within Nymphidiini, making Nymphidiini polyphyletic, Mesene, from Symmachiini, groups with a clade that is ba sal to the (Nymphiini, Stalachtini, and Lemoniini) clade, and Emesini, one of Harv eys incertae sedis group, splits and groups into five different clades (see Figure 3-1). Myrmecophily The parsimony analysis of the best like lihood tree and the random trees support the 3G. The difference in parsimony score s between 2G and 3G in the best tree was 5 while the average difference of the random trees was 0.98 with a standard deviation of 40

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0.876. Therefore, even though the 3G is expected to have a lower parsimony score than the 2G by random chance, the 3G, when m apped onto the Myrmecophily Tree, is better than the 2G by a significantly higher diffe rence than expected by random chance (Table 3-3). Also noteworthy, when scored with the 3G, the Myrm ecophily Tree (Figure 4-2) had no explicit losses whereas the average number of losses on the random trees was 12.16 (standard deviation: 2.13) and the 2G had 6 losses whereas the random trees had an average of 14.12 losses with a standard deviation of 2.13 losses, and the 1G had an average of 16.94 losses with standard devia tion of 2.32 losses (see Table 3-5). The fact that the Myrmecophily Tree require s no explicit losses means that a novel hypothesis with four or more gains would be less parsimonious than a hypothesis with three gains when mapped onto the Myrmecophily Tree so I did not test any hypothesis with more than 3 gains. 41

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Table 3-1. Characteristics of the DNA sequence data. Invariant Autapomorphic Cladistically Informative Total Sites wg 142 (35.2%) 29 (7.2%) 232 (57.6%) 403 COI 839 (56.0%) 103 (6.9%) 556 (37.1%) 1498 EF 1-a 810 (65.3%) 59 (4.8%) 371 (29.9%) 1240 42

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43 Table 3-2. Likelihood values and AIC va lues from RAxML likelihood analyses. The data partitions are indicated by parentheses and x CODON indicates that the matrix was partitioned by codon position. The AIC comparisons section portrays the difference of the AIC values, that is, it subtracts the AIC value for the partition scheme (1-7) depicted on the column from the AIC value of the scheme depict ed at the left of each row. E.G. 2111.4 comes from the the difference of the AIC values: 101410.3-99298.9. Partition scheme Log likeliRelative # of ParaRelative AIC=2kAIC pairwise comparisons hood (ln(L)) meters (k) 2ln(L) 1234567 1 (CO1 wg Ef 1-a ) -50705.15 0 101410.3 2111.4 2014.8 693.2 780.26 3348.3 6799.0 2 (CO1 ) (wg ) ( Ef 1-a) -49637.46 12 99298.9 96.6 1418.2 1331.1 1236.9 4687.6 3 (CO1 ) ( wg Ef 1-a) -49691.75 6 99395.5 1321.6 1234.5 1333.5 4784.2 4 (CO1 wg ) ( Ef 1-a) -50352.55 6 100717.1 87.1 2655.1 6105.8 5 (CO1 Ef 1-a ) (wg) -50309.02 6 100630.0 2568.0 6018.8 6 (CO1 wg Ef 1-a ) x CODON -49019.02 12 98062.0 3450.8 7 (CO1 ) (wg ) ( Ef 1-a) x CODON -47257.64 48 94611.3

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Figure 3-1. RAxML most likely tree for the 3 genes combined and partitioned by gene and by codon position, higher taxa indicated by colored branches. 44

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Figure 3-2. RAxML most likely trees for individual genes, higher taxa indicated by colored branches. 45

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46 A Figure 3-3. Consensuses of bootstrapped combined and individual RAxML trees (bootstraps indicated at nodes). Trees shown as follows: A) Three genes combined in a matrix partitioned by gene and by codon position 1000 bootstraps B) Bootstrap consensus COI gene tree of 1000 bootstraps C) Bootstrap consensus wingless gene tree of 1000 bootstr aps D) Bootstrap consensus Ef-1a gene tree of 1000 bootstraps.

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B Figure 3-3. Continued 47

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C Figure 3-3. Continued 48

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D Figure 3-3. Continued 49

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Figure 3-4. 50% majority rule consensus tree of the three mo st likely gene trees. 50

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Table 3-3. Parsimony score statisti cs for 1G, 2G, and 3G when mapped onto 100 random trees. 3G 2G 1G 2G-3G 1G-3G 1G-2G avg. 15.16 16.14 17.94 0.98 2.78 1.80 stdev. 2.13 2.15 2.32 0.88 1.80 1.26 range 10-21 12-22 12-24 0-4 0-9 0-6 51

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CHAPTER 4 DISCUSSION Taxonomy As seen in Figure 4-1 the results of the phylogenetic analyses corraborate the results of earlier morphologically-bas ed studies on the Riodi nidae and illuminate previously unresolved evolutionary relati onships within the Riodinidae (Hall 2002, Hall and Harvey 2002a, Hall and Harvey 2002b, Lamas 2004, Harvey 1987). There are seven major Riodinidae clades found in this study. These are: the Nymphidiini clade (which includes Nymphidiini sensu Harvey 1987, Lemoniini, Stalachtini, some Incertae Sedis Emesini, and the genus Mesene from Symachiini); the Riodinini clade (including the Riodinini sensu Harvey 1987); the Helicopini clade (including Charitini sensu Harvey 1987, Helicopini sensu Harvey 1987, and some Incertae Sedis Emesini sensu Harvey 1987); the Mesose miini clade (including Mesosemiini sensu Harvey 1987 and Napaeina sensu Hall 2003); the Eurybiini clade (including Eurybiini); the Euselas iinae clade (including Euselasiini sensu Lamas 2004); and the Nemeobiinae clade including Hamearis and Styx infernalis which makes up a monotypic tribe, Stygini, sensu Ehrlich 1958 (see Figure 3-1). I will address the taxonomic implications of my phylogenetic results starting with the basal clade of Riodinidae and working my way up the phylogeny shown in Figure 31. The basal clade is composed of Euselasi inae (Kirby 1871), Nemeobiinae (Bates 1868), and Stygini (Ehrlich 1958). Styginae, a monotypic subfamily comprised of the Andean Styx infernalis was not included in Riodinidae a ccording to Ehrlich (1958), but was considered a subfamily along with Lyc aeninae and Riodininae, based, in part, on 52

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the absence of a process of the trochanter. Stichel (1910-1911) also excluded Styx from Riodinidae due to the lack of the projecti on of the coxa on the male forelegs. However, according to the genet ic results reported herein, Styx and the Old World subfamily Nemeobiinae t ogether comprise the sister clade to Euselasinae, even though Lamas 2004 included Styx with Euselasinae separate from the Nemeobiinae. These results provide strong support for the placem ent of the controversial genus/subfamily Styx/Styginae within the Riodinidae most closel y related to the old world subfamily Nemeobiinae (Figures 31, 3-2, and 3-3). Mesosemiini (tribe) is the basal gr oup of the Riodinina e (subfamily). The Incertae Sedis group, sensu Harvey 1987 (or Napaeina) belong s with the Mesosemiini, as previously found by Hall (2003) who named it Napaeina, giving it subtribe status within the Mesosemiini and reducing the previous Mesosemiini ( sensu Harvey 1987) to a subtribe (Mesosemiina). Hereafter, Harveys Incertae Sedis is referred to as Napaeina. The other Incertae Sedis group sensu Harvey 1987 ( Incertae Sedis Emesini hereafter referred to as Emesini), however, is polyphyletic. Mesosemia (genus) is polyphyletic, as Harv ey predicted, with moderate bootstrap support (the highest bootstrap value indicating polyphyly is 64). And, as predicted by Hall (2003) Mesosemiina comes out as a si ster group of the Napaeina. Both Napaeina and Mesosemiina have 5 radial forewing ve ins, with the except ion of two genera, Mesophthalma and Perophthalma in the Mesosemiini (Harvey 1987). Along with Nymphidiini, Helicopini is also polyphyletic. Also, Helicopini sensu Lamas (2004) and Symmachiini sensu Harvey (1987) both contain one genus belonging to another tribe. These genera are Sarota and Mesene respectively (see Figure 3-1). 53

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Charitini is named after a misidentified type genus by Stichel. The existence of Helicopini makes Charitini par aphyletic, so Reuters Helicopini is used (Lamas 2004). Reuter (1897) also excluded Sarota (Sarotini) from his Helicopini tribe. This exclusion is supported by the mitochondrial gene tree but not by the nuclear gene trees or the total evidence tree (see Figures 3-1 and 3-2). Therefore, we need more data and analyses regarding Sarota to find its location in the Riodinidae. Based on the most likely tr ee, if Riodinini included Sarota then it would be monophyletic but, as the gene trees illuminate, Sarota probably belongs with the rest of the Helicopini. Harvey (1987) broke the Riodinini into two groups, the Ancyluris section and the Riodina section, based on the placement of a forewing vein (see Harvey 1987) and named after the genus Riodina The Ancyluris section is monophyletic, sharing an apparently derived placement of this vein (which Harvey hypothesized to be fused), while the Riodina section is paraphyletic (it is the re st of the Riodinini not sharing the synapomorphy). Another group in the Riodinini proposed by Hall and Harvey (2002b), the Charis group, also appears to be monophyletic with high support in my trees as well (bootstrap scores and corre sponding datasets: 93 in CO1 93 in wingless 98 in EF1a, and 100 in the combined analysis), though it also includes Crocozona which was not indicated by Hall and Harvey (see Figures 3-1, 3-2, and 3-3). Symmachiini does include Argyrogrammana as Stichel originally had it, even though Argyrogrammana lacks concealed abdominal scent organs, a trait shared by all other symmachiinids (Harvey 1987). This tr ait may not give good phylogenetic signal, because it was also found in Menander and Periplacis by Harvey. 54

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The Emesini, Harveys true Incertae Sedis clade, are all in cluded within the Riodininae, occurring mainly in sister clades of the Nymphidiin i/Lemoniini clade. However, a genus from Emesini also occurs in each of three tribes: Riodinini, Helicopini, and Symmachiini. Apodemia and Emesis group together as Harvey predicted, based on a characteristic of the pupal silk girdle (Harvey 1987). Lemoniini is monophyletic but Harveys Lemonias and Synargis sections are paraphyletic. Lemoniini splits the Nymphidiini tribe and the genus Calospila from Nymphidiini appears to belong withi n the Lemoniini. Stalachtin i, a mono-generic tribe comprised of Stalachtis also appears to be nested within the Nymphidiini. Based on these results, outgroup selecti on for higher level systematics of the Riodinidae is now possible. More intensive taxon sampling would better resolve many evolutionary relationships in the Riodinidae. The basal relationships in Riodinini and Helicopini are examples of said relations hips as well as the Mesosemiini, whose evolutionary relationships are also poorly resolved and would bene fit from a greater taxon sampling. When looking at t he phylograms (Figures 3-1 and 3-2) Stalachtis and Protonymphidium appear to have accelerated substitu tion rates relative to the taxa surrounding them: sampling additi onal taxa closely related to them would likely provide support or refute their grouping together. Though more research is still needed, this study provides a molecular phylogeny that was used to test the theories on the ev olution of myrmecophily and will serve as a foundation for future studies on the Riodinidae including insigh ts into the evolution of host plant usage, mimicry, and the co-evolu tion of myrmecophiles and their symbionts. 55

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It also provides novel gene sequences for 68 new taxa within the Riodinidae that can be used in a larger scale analysis of butterflies or insects. Phylogenetics and Gene Trees A noteworthy outcome of the phylogenetic analyses is the fact that the likelihood analysis of the matrix partitioned by codon position and gene is far better than any other likelihood analysis, including that of the matrix partiti oned by codon position alone and that of the matrix partiti oned by gene alone (see Table 3-2 for comparison). Greater congruence between this molecular tree and previous hypotheses based on morphology (e.g. Harvey 1987 and Lamas 2004) When looking at the topology of trees Sarota which was putatively misplaced in the trees from the analyses of both the codon position partitioned matrix and the gene par titioned matrix grouped with the other Helicopini in the tree from the analysis of the matrix partitioned by gene and codon position (see Appendix A). Although there are some minor diffe rences between genes, all three genes individually support most of the tribes described by Harv ey (1987), albeit with less support than in the total evidence tree (see Figures 3-2 and 3-3). Each of the three genes support the basal group (Euselasinae, Styginae, and Hamaerinae together), the Nymphidiini, most of the Mesosemiini, and the Riodinini (including the Charis group and Ancyluris group within it) although they do so with less support than when they are analyzed together. In other words, there are more polytomies when the genes are analyzed separately than in the combined analysis (see the consensus tree of all three individual gene trees in Figure 3-4). Although all three gene trees have similarities, there are differences as well. These differences pr ovide insight about some of the surprising results of this study. Wingless and elongation factor 1-alpha the two nuclear genes, 56

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provide better resolution (less polytomies) and higher support regarding the deeper relationships within the phylogeny of the Riodinidae than does the mitochondrial gene. Two of the surprising relationships uncovered in this analysis are supported by all three gene trees: Stalachtis is within the Nymphidiini and Mesene is closely related to two Incertae Sedis yet never near the other Symmachini. Also, the genus Mesosemia is polyphyletic in all three gene trees. As previously found (Hoelzer 1997, Springer et al 2001, Lin and Danforth 2004) there appear to be differences between the mitochondrial and nuclear genes. The reason that Sarota groups in Helicopini with such low support may be elucidated when looking at the different gene trees. Sarota is grouped near the ot her Helicopini genera but with low support in both the nuclear gene trees, however, it is grouped with Metacharis with high support (bootstrap value of 99) in the mitochondrial gene tree. Presumably, this is due to model based error, sampling error, or the different genes may truly have different coalescent histories; data from more genes are needed to resolve the position of Sarota within the Riodinidae. Additional notable differences between the mitochondrial and nuclear genes include a lycaenid (different family) outgroup, eurybinid ( Alesa ), and a member of the Mesosemiini ( Perophthalma ) grouping together, albeit with low support, in the mitochondrial gene tree but not in the nucle ar gene trees. This may be due to long branch attraction (note the relative long branches of Alesa and Curetis bulis which group as sister taxa in comparison to the surrounding branch lengths on the mitochondrial gene tree). The deeper relationships, between tribes, differ on each gene tree in the maximum likelihood tr ees, but these relationships break down 57

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into one polytomy in the 50% majo rity rule tree. The genera in the Incertae Sedis do not associate to the tribes with high support in many cases (see Figures 3-1 and 3-3). Myrmecophily As previously shown (DeVries 19 91b, Campbell 1998, DeVries 1997), myrmecophily appears to have been gained and/or lost multiple times in the Riodinidae + Lycaenidae clade. In the most likely to tal evidence tree the basal clades of the Riodinidae reportedly do not c ontain myrmecophily (DeVries 1997) and the two riodinid groups that do contain myrmecophily are topologically separated by multiple other clades (see Figure 4-2). With a robust mole cular phylogeny, the evolutionary dynamics of myrmecophily becomes testable. The inferr ed phylogeny allows us to test the three competing evolutionary hypothes es as to the origin of myrmecophily using parsimony following Campbell 1998, Nepokroeff et al. 2003, and Weiblen 2004. The three competing hypotheses (one, two, or thr ee gains) presented by Devries (1991a, 1997) were mapped onto the total evidence tree (Figure 4-2). The known and inferred character states were also mapped on the tree. These char acter states (presence or absence of myrmecophily) were taken or inferr ed from Harvey 1987, Horvitz et al. 1987, Devries 1997, DeVries et al. 2004, and Kaminski 2008, using presence of long, thick setae as an indication of amyrmecophily. In addition, one hundred random trees were generated in PAUP 4.0 (Swofford 2003) using the same taxa and outgroup as in the above likelihood analysis. The three hypotheses must be compared using an indirect method for two reasons. The first reason is, given our tree we would expect the number of evolutionary changes (= parsimony scores) per hypothes is to follow a trend from 3G<2G<1G by chance alone (here G<2G means the parsimony score for 3G is less than that for 2G, 58

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that is 3G is more parsimonious, when ma pped onto the tree) The second reason is, we cannot simply compare 3G mapped on t he Myrmecophily Tree to 3G mapped onto a random tree due to the fact that if any of our hypotheses (1G, 2G, or 3G) is correct for the Myrmecophily Tree, then we will see a reduction in tota l number of changes for each of the three hypotheses when mapped onto the Myrmecophily Tree (as compared to a random tree). 3G<2G<1G The first reason is due to the fact t hat in a relatively complex phylogeny (complexity defined here as the binomial variance of the evolutionary trait under consideration (myrmecophily or lack t hereof) and because there are16 taxa with myrmecophily present and 22 taxa with the tr ait absent the variance would be given by the equation: 38*16/38(116/38) = 9.26), a tree constrained to exactly two gains would expectedly be more parsimonious than a tree constrained to have exactly one gain due to the expected additional losses required fo r 1G. This is because each additional gain may be used to account for multiple losse s (compare B) and C) in Figure D-2) and a gain followed by a back-mutation loss is unlikely to occur, and likewise for 3G versus 2G. This is seen in the Myrmecophily Tr ee and in the random trees. When 2G and 3G are mapped onto the Myrmecophily Tree 3G is more parsimonious than 2G, but in this case, 2G is equally parsimonious to 1G which is quite possibly due to the fact that there are few outgroup taxa and only one character st ate is known in the outgroup (Table 4-1 and Figure 4-2). Furthermore, it is also conf irmed when looking at the results from the random trees (see Appendix F). 59

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The Hypotheses Are Not Independent We cannot simply compare 3G on the My rmecophily Tree to 3G on the random tree (and likewise for 1G and 2G) if any of the three hypotheses are true because we expect the 3G on the Myrmecophily Tree to be more parsimonious than 3G on a random tree (and likewise for 2G and 1G, that is 2G on the Myrmecophily Tree is expected to be more parsimonious than 2G on a random tree if any of the three hypotheses is true, etc.). This is because, given a phylogeny where 1G, 2G, or 3G is true, the complexity of t he tree will be reduced compared to a random tree. Take the case where 3G is known to be true and there are no losses. The comp lexity of the tree will be reduced because ther e are now only three clade s with myrmecophily, and effectively only three myrmecophilous taxa to take into account, so instead of a variance found in the following equation: 38*(16/38)(116/38) = 9.26, the variance will be 2.64 found in the in this way: 25*(3/25)(1-3/25) =2.64. So 1G (a si mpler case), when mapped on this tree, will also be more parsimonious than 1G mapped on a random tree (compare the three hypotheses mapped onto the Myrmecophily Tree against the three hypotheses mapped onto the random trees in Figure 4-2, Table 4-1, and Appendix F). The Indirect Method The difference between hypotheses must be used here. That is, we will test whether the difference in parsimony score s between the opposing hypotheses for the Myrmecophily Tree is larger than would be expec ted by chance. To exemplify this, take the above case (the tree where 3G is given to be true) and the expected result will be 3G is more parsimonious than 1G on this tree to a greater extent than on a random tree and likewise for 3G compared to 2G. However, 1G as compared to 2G will not show this trend, in fact we might expec t the differences to be similar between the true and random 60

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tree. In order to account for the expec ted difference between the hypotheses due to random chance I compared the parsimony sco res from the myrmecophily tree to the parsimony scores for 3G, 2G, and 1G when mapped onto 100 randomly generated trees. Given one of the hypotheses (1G, 2G, or 3G) is correct, the expected trends in the differences of parsimony scores when the hypotheses are mapped onto the trees are expressed in Table 4-2. Based on morphology (DeVries 1988, 1991 a, 1997, and Travassos et al 2008), myrmecophily appears to have evolved th ree times (3G). In support of the aforementioned previous work, the three apparently non-homolog ous ant-associated stridulation organs group with t he three myrmecophilous clades in this study. Vibratory papillae are found in clade 1, cuticular spines and cranial setae are found in clade 2, and a file and plate system is found in clade 3 (see Figures 1-2 and 4-2). Each of the three clades also has a distinct strategy for chemical communication (DeVries 1997). In support of this evidence for 3G, inference after mapping the different hypotheses onto the total evidence tree indica tes that the difference between 3G and 2G is greater than expected due to chance, the difference between 3G and 1G is greater than expected due to chance, and the difference between 2G and 1G is less than expected due to chance. This conclusion a ssumes that a gain is just as likely as a loss (see Table 4-1). When comparing the di fferences in parsimony scores for the Myrmecophily Tree versus the random trees we see the expected trends as predicted for 3G but the parsimony difference betw een 1G and 3G shows the expected trend, however it is not significant. C onversely, the difference between 2G and 3G is outside the 95% confidence interval as expected and the difference between the 1G and 2G is 61

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well within the 95% confidence interval as expected (see Tables 4-2 and 4-3 and Appendix F) The lower than ex pected difference between the 1G and 3G may be due to the fact that no ta xa with known character states we re sampled in the outgroup (see Figure 4-2). The results are also consistent with the interpretation that there have been no losses of myrmecophily within Riodinidae (the taxa within the nymphidine clade, which is clade 1 in Figure 4-2, have either been observed as myrmecophilous or there is no record as to their association with ants) which is different the findings of Campbell (1998). Note: in Figure 4-5 for a three gains hypothesis to occur there would have to be a loss of myrmecophily in the anc estor of the amyrmecophilous Helicopis (Campbell 1998). While the most likely total evidence tree presented here is in need of more data to improve resolution, there is however compelling evidence that myrmecophily evolved three times and has not been lost in the Ri odinidae. This analysis illustrates how a robust phylogeny enhances our understanding of interesting evolutionary phenomena. The three independent gains associate with t he three different types of sound producing apparatus, giving us further insight to their evolution. Further st udy regarding juvenile riodinids and additional sequence data are needed to better understand this interesting phenomenon. Conclusion This study generated many new butterf ly gene sequences, provides greater taxonomic resolution, supports the case for a three gain hypothesis regarding myrmecophily in the Lycaenidae + Riodinidae clade, and is a basis for future studies on the evolution found in the Rio dinidae and related taxa. Sequ ences were generated from 62

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68 novel riodinid specimens for a total of 200 new gene sequences from these taxa that will be made available for future studies, es pecially those that use the three genes employed in this study. The findings based on the analysis of these genes support and clarify relationships found in previous studies while raising qu estions to a few established taxonomic relationships. Based on the analysis of these gene sequences we now have a better understanding of the deeper level evolutionary relationships in the Riodinidae such as the placement of the subfam ilies and tribes as well as an affirmation of most of the shallow level relationships within these tribes There are some interesting phylogenetic results that raise questions to be analyzed in future studies. Finding more taxa and morphological or genetic markers will help clar ify such questions that arise from, for example, the extreme polyphyly found in the Emesini, the Mesosemia that looks like a super genus in need of splitting, and Stalachtin is position within the Nymphidiini. Also based on this analysis, there appears to have been three gains of myrmecophily in the riodinid and lycaenid clade and, more surprisingly, no losses. If this is true then individuals within Stalachtini as well as the other individuals within the nymphidine clade (clade 1 in Figure 4-2) will be myrmecophilous upon further examination. Greater taxon sampling as well as juvenile records are needed within the Riodinidae and Lycaenidae to more fully under stand the evolution of myrmecophily. This analysis is just one way in which this hypothesis can be used to more fully understand the mysteries found in this interesting family. The phylogeny herein provided can now be used to further explore host plant -butterfly co-evolution, ant-myrmecophile co-evolution, and the evolution of mimicry so vastly present in the Riodinidae. 63

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Figure 4-1. Morphological phy logenetic hypothesis after Harvey 1987 including his subfamilies and tribes compared to my molecular phylogenetic hypothesis. 64

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Table 4-1. Parsimony scores from mapping gains and losses of myrmecophily onto the Myrmecophily Tree (Figure 4-2). Hypothesis Gains Losses Total 1G 1 7 8 2G 2 6 8 3G 3 0 3 65

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Figure 4-2. The Myrmecophily Tree. Evol utionary gains and losses of myrmecophily mapped onto the best tree from the total evidenc e RAxML analysis with character states denoted. The hypothes is corresponding with each gain is indicated near the gain. Character states were inferred for Charis, Metacharis, Caria, Emesis, Anteros, Symmachia, Napaea, and Helicopis (Helicopis from information regarding absence of know n ant organs and the rest from presence of long, thick setae often used to repel insects). 66

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Table 4-2. Expected trends in differences of parsimony scores of the myrmecophily hypotheses when mapped onto the Myrmeco phily Tree (Figure 4-2) versus those mapped onto the random trees if the true myrmecophily hypotheses were known. The expected trend in par simony scores in a tree as complex as ours is 1G>2G>3G meaning that 3G is expected to be the most parsimonious (G>2G means that 1G is less parsimonious than 2G i.e. that the number of changes is greater for 1G compared to 2G). True hypothesis Expected trends Myrm. Tree Rand. Tree 1 gain 1G-3G < 1G-3G 1G-2G < 1G-2G 2G-3G ~ 2G-3G 2 gains 1G-3G ~ 1G-3G 1G-2G > 1G-2G 2G-3G < 2G-3G 3 gains 1G-3G > 1G-3G 1G-2G ~ 1G-2G 2G-3G > 2G-3G Table 4-3. Results from comparison of the hypotheses mapped on the Myrmecophily Tree next to the averages and 95% confidence intervals taken from random trees. Hypothesis indicates difference of hypotheses and notation follows that described in Table 4-2. Parsimony Scores Hypothesis 95% CI Avg. Myrm. T Tree 1G-3G 0-6 2.78 5 1G-2G 0-4 1.80 0 2G-3G 0-2 0.98 5 67

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Figure 4-3. Adaptation of Ca mpbells (1998) molecular phy logeny with myrmecophilous character state indicated by square next to genus. In this study if there was no myrmecophily indicated it was assumed to be amyrmecophilous. 68

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69 APPENDIX A COMPARISON OF 3 PARTITIONING SCHEMES FOR THE TOTAL EVIDENCE ANALYSES

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APPENDIX B BAYESIAN TREES WITH POST ERIOR PROBABILITIES 70

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71

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Above are three Bayesian gene trees with ta xa indicated by colored branches and posteriors indicated on branches. 72

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73 Appendix B Continued. Total evidence Bayesian phylogeny with ta xa indicated by colored branches and posteriors indicated on branches

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APPENDIX C RANDOM TREES USED TO GENERATE A NULL DISTRIBUTION FOR THE MYRMECOPHILY HYPOTHESES Object C-1. Random Trees 74

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75 APPENDIX D PROCEDURE FOR SCORING RANDOM TR EES WITH A WORKED EXAMPLE Procedure 1) Score character states on phylogeny (myrmecophilous or amyrmecophilous) and check to make sure you have correct number of each state. Note: in this case the outgroup is constrained to be myrmecophilous. And mark phylogeny branches as ancestral as possible for presence or absence of trait without requiring a gain (assuming parsimony is the standard) to simplify every step after step 1) (see Fig D-1) Figure D-1. Example of how to mark a phylogeny as in step 1). A) depicts a phylogeny whose taxa have known character stat es B) depicts the marking of the individual taxa and C) depicts how to simplify the tree based on parsimony. 2) Under the assumption of a myrmecophilous ancestral character state, find all clades where a character loss followed by a single more recent gain (one nearer the tips of the tree) will re sult in a more parsimonious character state evolution score than if the ancestral loss had not occurred (see Figure D-2 A and B). Make sure the loss is as ancestral as possible while only requiring one gain. (Compare A to B in Figure D-2: the loss depicted in B is more ancestral and more parsimonious than the loss in A so B is the chosen option.) Figure D-2. Three interpretations of the evolution of myrmecophily taken from the same phylogenetic tree. 3) Denote these changes and the total number of losses saved for each clade found in step 2) throughout the tree. E.G. in Figur e D-2 for B. I would write -> 3-1 losses implying that, respectively, the lo ss and gain seen on the tree, will save our score three losses (those depicted in C.) but cost one additional ancestral loss (at ) to the parsimony score.

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4) Check to see if any two of these clades can be combined to create a more parsimonious scenario than the two clades would create separately by placing an ancestral loss more basally than either of the two losses could have been placed separately (see Figure D-3). Figure D-3. Two interpretations of the ev olution of myrmecophily based on the same tree. In this figure, A. denotes a tree with two clades (described in step 2) and a total of four losses that can be replac ed with a single loss at in B. to reduce the number of changes from six to three. The numbers on the right follow the format introduced in step 3). 5) In order to compare schem es in the next step record all possible combinations of two clades from step 2) that are f ound in the previous step and any other combinations left (e.g. in Figure D-3 I listed the schem e from B. and the scheme from A. but in more co mplex trees there will be even more possibilities) along with their parsimony scores. Note: because there are at most three gains and the outgroup is constrained to be myrmecophilous there can be at the most two clades combined in this way. 6) Check the assumption in step 2), that is, check w hether a gain in the outgroup with a subsequent gain more recent on the tr ee (A-C in Figure D-4) or a gain in the ancestor of all taxa (D-F in Figur e D-4) is more parsimonious or equally parsimonious. Number the gain that is equally or more parsimonious to be included for comparison. 76

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Figure D-4. Further explanation for step 6) and 7). In this case for A) a gain in the outgroup followed by a subsequent gain is more parsimonious than a gain in the ancestor, for B) a gain in the out group followed by a subsequent gain is equally parsimonious to a gain in the ancestor followed by a loss and another gain and in C) a gain in the ancestor is more parsimonious than a gain in the outgroup followed by a loss and another gain. 7) If a gain in the outgroup followed by a subsequent gain (e .g., A-C in Figure D-4) is equally or more parsimonious than the gai n in the ancestor (e.g ., D-F in Figure D-4) then check to see if the gain in the outgroup can be combined with a clade from step 2) in a similar fashion to that described in step 4) to result in fewer changes (i.e. a more parsimonious result ) than if the two were not combined. 8) Record the gains (two or thr ee, depending on the hypothesis) and losses and total number of losses (e.g. as in step 3 )) saved for all possibl e combinations for both hypotheses. 9) Choose the most parsimonious option recorded for both 2G and 3G. (For 2G it will either be one clade from step 2 and a gain in the ancestor of the group or the gain in the outgroup and a subsequent gain. Fo r 3G it will either be two clades combined and a gain in the ancestor of t he group, the gain in the outgroup and a subsequent gain with a clade from step 2), or the gain in the outgroup combined with two subsequent gains). 10) Mark trees with gains and losses accordingly and record the gains and losses for each hypothesis. 11) To find the parsimony score for 1G take the clade that was used in step 9) and add the number saved on that clade to the parsimony score for the two steps hypothesis and subtract one (due to one less gai n). (E.g. in Figure D-3 the clade used in step 9) would be 2 -> 3-1 bec ause the parsimony score is bettered by a total of two (-1) changes due to the clade, clade 2, found in step 2), but this clade is not possible with only one hypothesis so we add the total changes saved and subtract one due to one less gain). 77

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Worked Example A Figure D-5. Step by step method for scoring a random tree (here it is random tree number 94 which is the most complex. A. is the random tree. B. shows step 1) from above the taxa with known evolutionary states are scored and then parsimony is invoked to group any of like states together and infer the characte r state of any branch unle ss there is a conflict with another taxon. C. shows step 2) where all gains followed by lo sses that result in a more parsimonious score are mapped. D. shows step 3) where the gains and losses are numbered. E. shows steps 4) and 5) where th e losses and gains followed by the total 78

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number of changes they save are recorded as well as combined and then recorded for 3G. Loss and gain save a total of th ree losses but cost one additional loss (the one labeled ) so ->3-1 is recorded. Loss and gai n save a total of four losses but cost one additional loss so ->4-1 is recorded. The rest save a total of 1 (2-1) changes and are not recorded. Both 34 and are equally parsimonious for 3G. F. shows step 6) and because 10 11 is worse than other hypotheses we predict that there must be a gain in the ancestor for 2G and 3G based on parsimony. Also there must be a gain in the ancestor of t he tree for 1G because the outgroup is myrmecophilous. Therefore step 7) is ski pped. G. shows step 9) where the most parsimonious scheme is chosen for 2G and 3G (chosen schemes are indicated by stars). H. shows step 10) where the tree is mapped with gains and losses accordingly and tallied for 2G and, because t he trees are already complicat ed, I deleted all the gains and losses from the previous steps except for the chosen options whereas on paper I would use a different color pen (see the scans in Appendix E). I. shows step 10) for 3G. And J. shows step 11) where t he parsimony score for 1G is calculated just off of the numbers used for 2G. In this case the extra gain (going from the one to 2G) saved the tree 4-1 changes but also cost an additional gai n so I added 2 (4-1-1) to the total score of 2G. (Similarly, if in another tree, the extra gain saved the tree 7-1 changes I would add 5 to the parsimony score of 2G). Ther efore the parsimony scores for 1G, 2G, and 3G are 24,22, and 21 respectively. 79

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B Figure D-5. Continued 80

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C Figure D-5. Continued 81

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D Figure D-5. Continued 82

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E Figure D-5. Continued 83

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F Figure D-5. Continued 84

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G Figure D-5. Continued 85

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H Figure D-5. Continued 86

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I Figure D-5. Continued 87

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J Figure D-5. Continued 88

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APPENDIX E SCORED RANDOM TREES Object E-1. Scans of 100 scored random trees 89

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90 APPENDIX F RANDOM TREE NULL DISTRIBUTION RESULTS Parsimony scores for 1G, 2G, and 3G and differences in parsimony scores for 100 random trees generated using PAUP 4.0 (Swofford 2003).The tree number corresponds to the random trees in Appendix C and Appendix E. random tree # 3G pscore 2G pscore 1G pscore p-score (2G-3G) p-score (1G-3G) p-score (1G-2G) 1 15 17 20 2 5 3 2 16 17 19 1 3 2 3 17 18 20 1 3 2 4 14 15 15 1 1 0 5 16 18 21 2 5 3 6 13 14 16 1 3 2 7 19 20 22 1 3 2 8 10 13 19 3 9 6 9 12 12 14 0 2 2 10 14 14 14 0 0 0 11 15 15 17 0 2 2 12 14 15 15 1 1 0 13 17 18 21 1 4 3 14 16 17 18 1 2 1 15 14 16 17 2 3 1 16 13 15 16 2 3 1 17 17 18 19 1 2 1 18 17 18 20 1 3 2 19 15 16 17 1 2 1 20 16 16 17 0 1 1 21 12 12 15 0 3 3 22 14 15 18 1 4 3 23 18 19 20 1 2 1 24 19 20 22 1 3 2 25 20 20 22 0 2 2 26 17 17 19 0 2 2 27 13 15 18 2 5 3 28 14 14 14 0 0 0 29 16 16 16 0 0 0 30 15 15 17 0 2 2 31 15 17 19 2 4 2 32 17 19 21 2 4 2 33 15 17 21 2 6 4 34 13 15 17 2 4 2 35 17 18 19 1 2 1 36 12 13 15 1 3 2 37 14 15 16 1 2 1 38 17 18 19 1 2 1 39 14 15 17 1 3 2 40 19 19 19 0 0 0 41 15 19 21 4 6 2 42 15 16 17 1 2 1 43 13 13 15 0 2 2 44 12 12 15 0 3 3 45 15 15 16 0 1 1 46 12 14 17 2 5 3 47 16 16 17 0 1 1 48 11 12 12 1 1 0 49 13 14 16 1 3 2 50 14 14 14 0 0 0

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Parsimony scores for 1G, 2G, and 3G and differences in parsimony scores for 100 random trees generated using PAUP 4.0 (Swofford 2003) continued. random tree # 3G pscore 2G pscore 1G pscore p-score (2G-3G) p-score (1G-3G) p-score (1G-2G) 51 15 16 18 1 3 2 52 16 17 18 1 2 1 53 13 14 16 1 3 2 54 15 15 17 0 2 2 55 14 15 17 1 3 2 56 14 14 14 0 0 0 57 10 13 16 3 6 3 58 17 18 20 1 3 2 59 17 17 18 0 1 1 60 16 16 18 0 2 2 61 16 17 19 1 3 2 62 18 19 21 1 3 2 63 15 16 17 1 2 1 64 13 13 13 0 0 0 65 19 19 20 0 1 1 66 16 16 17 0 1 1 67 16 16 16 0 0 0 68 15 15 15 0 0 0 69 13 14 19 1 6 5 70 14 14 17 0 3 3 71 18 18 18 0 0 0 72 14 16 21 2 7 5 73 13 14 18 1 5 4 74 16 16 16 0 0 0 75 19 21 21 2 2 0 76 16 18 20 2 4 2 77 17 18 19 1 2 1 78 15 17 19 2 4 2 79 14 16 21 2 7 5 80 18 18 19 0 1 1 81 17 18 19 1 2 1 82 16 18 20 2 4 2 83 14 16 19 2 5 3 84 14 15 17 1 3 2 85 16 18 21 2 5 3 86 17 17 19 0 2 2 87 16 17 20 1 4 3 88 14 14 19 0 5 5 89 15 16 18 1 3 2 90 13 14 16 1 3 2 91 16 18 19 2 3 1 92 13 14 16 1 3 2 93 15 15 17 0 2 2 94 21 22 24 1 3 2 95 16 17 19 1 3 2 96 15 18 20 3 5 2 97 18 19 19 1 1 0 98 11 12 15 1 4 3 99 14 15 16 1 2 1 100 16 19 22 3 6 3 Average: 15.16 16.14 17.94 0.98 2.78 1.80 91

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LIST OF REFERENCES Ackery, P. R., R. de Jong, and R. I. VaneWright. 1999. The butterflies: Hedyloidea, Hesperioidea, and Papilionoidea. Pages 264-300 in: Lepidoptera: Moths and Butterflies. 1. Evolutio n, Systematics, and Biogeography. Handbook of Zoology Vol. IV, Part 35. N. P. Kristensen, ed. De Gruyter, Berlin and New York. Alfaro ME, Zoller S, Lutzoni F. 2003. Bayes or bootstrap? A simulation study comparing the performance of Bayesian Markov chain Monte Carlo sampling and bootstrapping in assessing phylogenetic confidence. Mol. Biol. Evol 20:255 Bates, H.W. 1868. A catalogue of Erycini dae, a family of di urnal Lepidoptera. Journal of the Linnean Society London, 9: 373-459. Benson, D.A., Karsch-Mizrachi, I., Lipm an, D.J., Ostell, J., Wheeler, D.L. 2006. GenBank Nucleic Acids Res 34, D16D20 Bergsten, J. 2005. A review of long-clade attraction. Cladistics 21: 163-193 Borror, D.J., C. A. Tripplehorn, and N.F. Johnson. 1989. Introduction to the Study of Insects. Harcourt-Brace-Jovanovich Co llege Publishers, Or lando, Florida. Brower, A.V.Z. & R. DeSalle (1998). Patte rns of mitochondrial versus nuclear DNA sequence divergence among nymphalid butterflies: the utility of wingless as a source of characters for phylogenetic inference. Insect. Mol. Biol. 7:73-82. Braby, M. F., Vila, R., and Pierce, N.E. 2006. Molecular phylogeny and systematics of the Pieridae (Lepidoptera: Papilionoidea): higher cla ssification and biogeography. Zoological Journal of the Linnean Society 147: 239. Campbell D.L. 1998. Higher-level phylogeny and molecular evolution of the Riodinidae (Lepidoptera). PhD thesis. Harvard Univ. Campbell, D. L., Brower, A. V. Z. and Pierce, N. E. 2000. Molecular evolution of the wingless gene and its implications for the phylogenetic placement of the butterfly family Riodinidae (Lepi doptera: Papilionoidea). Mol. Biol. Evol 17: 684-696. Caterino,M.S., Reed,R.D., Kuo,M.M. and S perling,F.A. 2001. A partitioned likelihood analysis of swallowtail butterfly phylogeny (Lepidopter a:Papilionidae) Syst. Biol 50: 106-127. DAbrera B. 1994. Butterflies of the Neotropical Region, Pa rt VI Riodinidae. Victoria, Australia: Hill House. de Jong, R., R. I. Vane-Wrigh t, and P. R. Ackery. 1996. The higher classification of butterflies (Lepidoptera): problems and prospects. Entomologica Scandinavica 27:65-101. 92

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DeVries, P. J. 1997. The Butterflies of Costa Rica and their Natural History. II: Riodinidae. Princeton University Press, New Jersey. DeVries, P. J. 1988. The larval ant-or gans of Thisbe irenea (Riodinidae) and their effects upon attending ants. Zoological Journal of the Linnean Society 94: 379393. DeVries, P. J. 1990. Enhancement of symb ioses between butterfly caterpillars and ants by vibrational communication. Science 248: 1104 -1106. DeVries, P. J. 1991a. Call production by myrmecophilous riodinid and lycaenid butterfly caterpillars (Lepidoptera): morphological, ac oustical, functional, and evolutionary patterns. American Museum Novitates 3025: 1-23. DeVries, P. J. 1991b. Ecolog ical and evolutionary patterns in riodinid butterflies. IN: AntPlant Interactions. C. Huxley and D. F. Cutler (eds.) Oxford Univ Press. DeVries, P. J., B. Cabral and C. M. Penz. 2004. The early stages of Apodemia paucipuncta (Riodinidae): myrmecophily, a new caterpillar ant-organ and consequences for classification Milwaukee Public Museum Contributions to Biology and Geology 102: 1-13. DeVries, P. J. and C. M. Penz. 2002. Early stages of the entomophagous metalmark butterfly Alesa amesis (Riodinidae: Eurybiini). Journal of the Lepidopterists' Society 56: 265-271. Ehrlich, P.R. 1958. The com parative morphology, phylogeny and higher classification of the butterflies (Lepido ptera: Papilionoidea). Univ. Kans. Sci. Bull 39:305-370. Felsenstein, J. (2004): Inferring Phylogenies Sinauer Associates, Sunderland, MA. Graybeal, A. 1998. Is it better to add taxa or characters to a difficult phylogenetic problem? Syst. Biol 47:9. Hall, J. P. W., Harvey DJ. 2002a. Basal s ubtribes of the Nymphidiini (Lepidoptera: Riodinidae): phylogeny and myrmecophily. Cladistics 18: 539. Hall, J. P., and Harvey, D. J. 2002b. A phylogenetic review of Charis and Calephelis (Lepidoptera: Riodinidae). Ann. Ent. Soc. Amer 95: 407-421. Hall, J. P. W. 2003. Phylogenet ic reassessment of the five forewing radial-veined tribes of Riodininae (Lepidoptera: Riodininae. The Royal Entomological Society 28: 2337. Hall, J. P. W. 2002. Phylogeny of the riodi nid butterfly subtribe Theopeina (Lepidoptera: Riodinidae: Nymphidiini). Syst. Ent 27: 139-167. 93

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Harvey, D.J., 1987. The higher classification of the Riodinidae (Lepidoptera). Ph.D. Dissertation, University of Texas, Austin Hill C. J. 1993. The my rmecophilous organs of Arhopala madytus Fruhstorfer (Lepidoptera: Lycaenidae). J. Aust. Ent. Soc. 32: 283. Hillis, D. M., Pollock, D. D., McGuire, J. A. and Zwickl, D. J. 2003. Is sparse taxon sampling a problem for phylogenetic inference? Syst. Biol. 52: 124. Hoelzer GA. 1997. Inferring phylogenies fr om mtDNA variation: mitochondrial-gene trees versus nuclear-gene trees revisited. Evolution 51: 622 Horvitz, C., Turnbull, C.and Harvey, D.J. 1987. The biology of immature Eurybia elvina (Lepidoptera: Riodinidae), a myrm ecophilous metalmark butterfly. Ann. Ent. Soc. Amer. 80: 513-519. Huelsenbeck, J. P. 1998. Systematic bias in phylogenetic analysis: is the Strepsiptera problem solved? Syst. Biol 47: 519 537. Huelsenbeck, J. P., and F. Ronquist. 2001. MrBayes: Bayesian inference of phylogeny. Bioinformatics 17:754-755. Kaminski, L.A. 2008. Immature stages of Caria plutargus (Lepidoptera: Riodinidae), with discussion on the behavioral and morphological defensive traits in nonmyrmecophilous riodinid butterflies. Ann. Ent. Soc. Amer. 101: 906-914. Kirby, W.F. 1871. A Synonymic Catalogue of Diurnal Lepidoptera. John Van Voorst, London. Kitching, L. 1983. Myrmecophilous organs of the larvae and pupa of the lycaenid butterfly Jalmenus evagoras (Donovan). J. Nat. Hist 17: 471-481. Kristensen N. P. 1976. Remarks on the fam ily-level phylogeny of butterflies (Insecta, Lepidoptera, Rhopalocera). Zeit. Zool. Syst. Evol 14: 25-33. Lamas, G. 2004. Atlas of Neotropical Lepidoptera. Checklis t: Part 4A Hesperioidea Papiionoidea. Gainesville: Scientific Publishers/Association of Tropical Lepidoptera. Lin, C. P., and Danforth, B. N. 2004. How do insect nuclear and mitochondrial gene substitution patterns differ? Insight s from Bayesian analyses of combined datasets. Mol. Phylogenet. Evol 30:686 Maddison, W. P. and Maddison, D. R. 1992. MacClade: Analysis of phylogeny and character evolution, Sinauer Associates, Sunderland, MA. 94

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97 BIOGRAPHICAL SKETCH Jonathan was born in Miami, FL, U.S.A. in 1984, the son of James Saunders and Shirley Saunders. In 2003 he was admitted to the University of Florida, majored in Wildlife Ecology and Conservation and mi nored in mathematics and received a Bachelor of Science degree fr om the College of Agricultural and Life Sciences in 2007.




Permanent Link: http://ufdc.ufl.edu/UFE0041641/00001

Material Information

Title: Molecular Phylogenetics of the Riodinidae (Lepidoptera)
Physical Description: 1 online resource (97 p.)
Language: english
Creator: Saunders, Jonathan
Publisher: University of Florida
Place of Publication: Gainesville, Fla.
Publication Date: 2010

Subjects

Subjects / Keywords: character, cytochrome, elongation, evolution, factor, lepidoptera, myrmecophily, oxidase, phylogenetics, riodinidae, wingless
Zoology -- Dissertations, Academic -- UF
Genre: Zoology thesis, M.S.
bibliography   ( marcgt )
theses   ( marcgt )
government publication (state, provincial, terriorial, dependent)   ( marcgt )
born-digital   ( sobekcm )
Electronic Thesis or Dissertation

Notes

Abstract: The butterfly family Riodinidae has many species that exhibit diverse traits, but the taxonomy of the family is still unclear and a robust phylogeny derived from molecular data is lacking. In this study I generate a phylogeny of 68 novel riodinid taxa. I use two nuclear and one mitochondrial gene to investigate the evolutionary relationships of the Riodinidae in likelihood and Bayesian analyses. The resulting phylogenies provide greater taxonomic resolution for the Riodinidae, especially in the deeper level relationships. They support many but bring into question a few of the previously described taxonomic relationships. The phylogenies are also used to address hypotheses about the evolution of myrmecophily in butterflies. The hypothesis of three independent gains of myrmecophily in the butterflies is supported over the two gains hypothesis and the one gain hypothesis. The results are also consistent with a lack of losses in the evolution of myrmecophily.
General Note: In the series University of Florida Digital Collections.
General Note: Includes vita.
Bibliography: Includes bibliographical references.
Source of Description: Description based on online resource; title from PDF title page.
Source of Description: This bibliographic record is available under the Creative Commons CC0 public domain dedication. The University of Florida Libraries, as creator of this bibliographic record, has waived all rights to it worldwide under copyright law, including all related and neighboring rights, to the extent allowed by law.
Statement of Responsibility: by Jonathan Saunders.
Thesis: Thesis (M.S.)--University of Florida, 2010.
Local: Adviser: Baer, Charles F.
Local: Co-adviser: Emmel, Thomas C.
Electronic Access: RESTRICTED TO UF STUDENTS, STAFF, FACULTY, AND ON-CAMPUS USE UNTIL 2012-04-30

Record Information

Source Institution: UFRGP
Rights Management: Applicable rights reserved.
Classification: lcc - LD1780 2010
System ID: UFE0041641:00001

Permanent Link: http://ufdc.ufl.edu/UFE0041641/00001

Material Information

Title: Molecular Phylogenetics of the Riodinidae (Lepidoptera)
Physical Description: 1 online resource (97 p.)
Language: english
Creator: Saunders, Jonathan
Publisher: University of Florida
Place of Publication: Gainesville, Fla.
Publication Date: 2010

Subjects

Subjects / Keywords: character, cytochrome, elongation, evolution, factor, lepidoptera, myrmecophily, oxidase, phylogenetics, riodinidae, wingless
Zoology -- Dissertations, Academic -- UF
Genre: Zoology thesis, M.S.
bibliography   ( marcgt )
theses   ( marcgt )
government publication (state, provincial, terriorial, dependent)   ( marcgt )
born-digital   ( sobekcm )
Electronic Thesis or Dissertation

Notes

Abstract: The butterfly family Riodinidae has many species that exhibit diverse traits, but the taxonomy of the family is still unclear and a robust phylogeny derived from molecular data is lacking. In this study I generate a phylogeny of 68 novel riodinid taxa. I use two nuclear and one mitochondrial gene to investigate the evolutionary relationships of the Riodinidae in likelihood and Bayesian analyses. The resulting phylogenies provide greater taxonomic resolution for the Riodinidae, especially in the deeper level relationships. They support many but bring into question a few of the previously described taxonomic relationships. The phylogenies are also used to address hypotheses about the evolution of myrmecophily in butterflies. The hypothesis of three independent gains of myrmecophily in the butterflies is supported over the two gains hypothesis and the one gain hypothesis. The results are also consistent with a lack of losses in the evolution of myrmecophily.
General Note: In the series University of Florida Digital Collections.
General Note: Includes vita.
Bibliography: Includes bibliographical references.
Source of Description: Description based on online resource; title from PDF title page.
Source of Description: This bibliographic record is available under the Creative Commons CC0 public domain dedication. The University of Florida Libraries, as creator of this bibliographic record, has waived all rights to it worldwide under copyright law, including all related and neighboring rights, to the extent allowed by law.
Statement of Responsibility: by Jonathan Saunders.
Thesis: Thesis (M.S.)--University of Florida, 2010.
Local: Adviser: Baer, Charles F.
Local: Co-adviser: Emmel, Thomas C.
Electronic Access: RESTRICTED TO UF STUDENTS, STAFF, FACULTY, AND ON-CAMPUS USE UNTIL 2012-04-30

Record Information

Source Institution: UFRGP
Rights Management: Applicable rights reserved.
Classification: lcc - LD1780 2010
System ID: UFE0041641:00001


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'100 tree(s) from PAUP*'
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(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Semomesia_5223_52_________________________X) Argyrogrammana_6709_42) Juditha_9315_26__________________ANTS__O_o__o) ((Ancyluris_6834_45 Hamearis_lucina) Stalachtis_6422_39)) Detritivora_9327_25) (((Protonymphidia_3948_59_________ANTS__O_o__o Echenais_7168_48) T_Poritia_erycinoides) (Helicopis_5318_54 Styx_infernalis))) Ithomiola_5924_82) (((Chamaelimnas_3942_57 Melanis_7612_30_________________________X______) Mesophthalma_6327_88_____________________X) Methone_8523_95)) Hyphilaria_5239_91) ((((Theope_7789_65__________________ANTS__O_o__o Theope_9026_5____________________ANTS__O_o__o) T_Miletus_ancon) Euselasia_8816_01_________________________X) Necryia_5966_93)) (((((((((((((((Adelotypa_7807_37_________________ANTS__O_o__o (((Setabis_6330_38_________________ANTS__O_o__o Perophthalma_9330_24_______________________X) Eurybia_9363_20_________________ANTS__O_o__o) (Thisbe_6766_43___________________ANTS__O_o__o Euselasia_orfita____________________________X))) (Ithomeis_7773_35 Themone_7643_34)) Pandemos_8572_97) (Synargis_7605_29_________________ANTS__O_o__o Monethe_3977_61)) (Calydna_6346_83 Symmachia_8849_08___________________________)) Zelotea_4057_79) ((Menander_7339_49_________________ANTS__O_o__o Sarota_9358_21) ((Cariomothis_6518_41 Leucochimona_9302_33_______________________X) Ourocnemis_4621_90___________________________))) Mesene_7450_28_________________________X______) (((Brachyglemis_3956_58 Crocozona_3995_62) Phaenochitonia_6926_47) Charis_9365_19_______________________________)) (Thysanota_5241_81_________________ANTS__O_o__o (Metacharis_9347_23__________________________ Emesis_mandana___________________________X_____))) (((Lemonias_3950_55_________________ANTS__O_o__o Napaea_m17_89_______________________________) Exoplisia_6433_40) (Leucochimona_9306_27_______________________X Anteros_7097_50______________________________))) (((((Calephelis_9356_22____________________X______ Cremna_5260_53) Mesosemia_6516_85_________________________X) (Pachythone_9050_18 Parcella_4012_64)) ((Esthemopsis_7813_32 Mesosemia_8822_02_________________________X) Leucochimona_7617_87_______________________X)) Caria_7625_31_______________________________)) Rhetus_3973_60) Calospila_8532_96_________________ANTS__O_o__o) (Panara_3932_56 Apodemia_mormo))) (((Amarynthis_5220_51 Roeberella_7457_36) Lepricornis_5041_80) ((Cyrenia_3992_63 Isapis_6815_44) Emesis_6837_46_______________________________))) Riodina_lysippus) Alesa_8380_84____________________ANTS__O_o__o)*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Stalachtis_6422_39) (((((((((((((((((Adelotypa_7807_37_________________ANTS__O_o__o Cariomothis_6518_41) Roeberella_7457_36) Chamaelimnas_3942_57) (Protonymphidia_3948_59_________ANTS__O_o__o (Calydna_6346_83 Cyrenia_3992_63))) Eurybia_9363_20_________________ANTS__O_o__o) (Caria_7625_31_______________________________ Methone_8523_95)) ((Metacharis_9347_23__________________________ Parcella_4012_64) Panara_3932_56)) (Leucochimona_9302_33_______________________X Sarota_9358_21)) Mesosemia_8822_02_________________________X) Phaenochitonia_6926_47) Menander_7339_49_________________ANTS__O_o__o) Thysanota_5241_81_________________ANTS__O_o__o) ((Cremna_5260_53 (Esthemopsis_7813_32 Rhetus_3973_60)) (Euselasia_8816_01_________________________X T_Miletus_ancon))) ((Crocozona_3995_62 Melanis_7612_30_________________________X______) Ithomeis_7773_35)) Napaea_m17_89_______________________________) Charis_9365_19_______________________________) Mesophthalma_6327_88_____________________X)) (Mesene_7450_28_________________________X______ Euselasia_orfita____________________________X)) ((Amarynthis_5220_51 Calephelis_9356_22____________________X______) Helicopis_5318_54)) Ithomiola_5924_82) (((((Lemonias_3950_55_________________ANTS__O_o__o Brachyglemis_3956_58) Echenais_7168_48) Themone_7643_34) Detritivora_9327_25) ((Argyrogrammana_6709_42 (Mesosemia_6516_85_________________________X Hamearis_lucina)) (Exoplisia_6433_40 (((Isapis_6815_44 Ourocnemis_4621_90___________________________) Monethe_3977_61) ((Pandemos_8572_97 T_Poritia_erycinoides) Styx_infernalis)))))) Ancyluris_6834_45) Anteros_7097_50______________________________) Theope_7789_65__________________ANTS__O_o__o) Leucochimona_9306_27_______________________X) Emesis_mandana___________________________X_____) Perophthalma_9330_24_______________________X) Lepricornis_5041_80) (Hyphilaria_5239_91 Zelotea_4057_79)) ((((((Alesa_8380_84____________________ANTS__O_o__o Theope_9026_5____________________ANTS__O_o__o) Calospila_8532_96_________________ANTS__O_o__o) (Leucochimona_7617_87_______________________X Riodina_lysippus)) Symmachia_8849_08___________________________) Pachythone_9050_18) ((Setabis_6330_38_________________ANTS__O_o__o Juditha_9315_26__________________ANTS__O_o__o) (((((Thisbe_6766_43___________________ANTS__O_o__o Synargis_7605_29_________________ANTS__O_o__o) Apodemia_mormo) Semomesia_5223_52_________________________X) Emesis_6837_46_______________________________) Necryia_5966_93))))*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o (Methone_8523_95 Pachythone_9050_18)) Necryia_5966_93) Crocozona_3995_62) Synargis_7605_29_________________ANTS__O_o__o) Napaea_m17_89_______________________________) (((((Theope_7789_65__________________ANTS__O_o__o Rhetus_3973_60) Hamearis_lucina) Leucochimona_7617_87_______________________X) Thysanota_5241_81_________________ANTS__O_o__o) Metacharis_9347_23__________________________)) (Ithomiola_5924_82 Mesosemia_8822_02_________________________X)) Mesosemia_6516_85_________________________X) Mesene_7450_28_________________________X______) ((Setabis_6330_38_________________ANTS__O_o__o ((((((Brachyglemis_3956_58 ((((Charis_9365_19_______________________________ Echenais_7168_48) Zelotea_4057_79) Esthemopsis_7813_32) T_Miletus_ancon)) Hyphilaria_5239_91) (Cariomothis_6518_41 Ithomeis_7773_35)) Pandemos_8572_97) Calephelis_9356_22____________________X______) Phaenochitonia_6926_47)) Eurybia_9363_20_________________ANTS__O_o__o)) Stalachtis_6422_39) Menander_7339_49_________________ANTS__O_o__o) Amarynthis_5220_51) Mesophthalma_6327_88_____________________X) Lemonias_3950_55_________________ANTS__O_o__o) ((Thisbe_6766_43___________________ANTS__O_o__o Sarota_9358_21) Parcella_4012_64)) ((((Protonymphidia_3948_59_________ANTS__O_o__o ((Alesa_8380_84____________________ANTS__O_o__o T_Poritia_erycinoides) Argyrogrammana_6709_42)) Juditha_9315_26__________________ANTS__O_o__o) Panara_3932_56) (Euselasia_8816_01_________________________X (Isapis_6815_44 Melanis_7612_30_________________________X______)))) Themone_7643_34) Emesis_mandana___________________________X_____) (Leucochimona_9302_33_______________________X Monethe_3977_61)) ((Chamaelimnas_3942_57 (Exoplisia_6433_40 Euselasia_orfita____________________________X)) Cremna_5260_53)) (((((Calydna_6346_83 ((Caria_7625_31_______________________________ Semomesia_5223_52_________________________X) Cyrenia_3992_63)) Roeberella_7457_36) Riodina_lysippus) Apodemia_mormo) Symmachia_8849_08___________________________)) (Lepricornis_5041_80 (Leucochimona_9306_27_______________________X Ourocnemis_4621_90___________________________))) (Detritivora_9327_25 Anteros_7097_50______________________________)) Adelotypa_7807_37_________________ANTS__O_o__o) Calospila_8532_96_________________ANTS__O_o__o) Helicopis_5318_54) ((Theope_9026_5____________________ANTS__O_o__o (Ancyluris_6834_45 Emesis_6837_46_______________________________)) (Perophthalma_9330_24_______________________X Styx_infernalis)))*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Calospila_8532_96_________________ANTS__O_o__o) (((((Setabis_6330_38_________________ANTS__O_o__o (Lemonias_3950_55_________________ANTS__O_o__o ((((((Thisbe_6766_43___________________ANTS__O_o__o Apodemia_mormo) Calydna_6346_83) Cyrenia_3992_63) (Argyrogrammana_6709_42 Perophthalma_9330_24_______________________X)) Semomesia_5223_52_________________________X) (Synargis_7605_29_________________ANTS__O_o__o Methone_8523_95)))) Helicopis_5318_54) (Ourocnemis_4621_90___________________________ Pandemos_8572_97)) (Detritivora_9327_25 ((Euselasia_8816_01_________________________X Styx_infernalis) Rhetus_3973_60))) Eurybia_9363_20_________________ANTS__O_o__o)) ((Leucochimona_9306_27_______________________X T_Poritia_erycinoides) Stalachtis_6422_39)) Euselasia_orfita____________________________X) Phaenochitonia_6926_47) ((((Adelotypa_7807_37_________________ANTS__O_o__o ((((Thysanota_5241_81_________________ANTS__O_o__o Napaea_m17_89_______________________________) Emesis_6837_46_______________________________) Calephelis_9356_22____________________X______) Chamaelimnas_3942_57)) Crocozona_3995_62) ((Cremna_5260_53 Parcella_4012_64) Roeberella_7457_36)) Riodina_lysippus)) T_Miletus_ancon) Mesosemia_6516_85_________________________X) Isapis_6815_44) Amarynthis_5220_51) Ithomeis_7773_35) Mesene_7450_28_________________________X______) Melanis_7612_30_________________________X______) Theope_9026_5____________________ANTS__O_o__o) Esthemopsis_7813_32) ((((Alesa_8380_84____________________ANTS__O_o__o (((((((((((Juditha_9315_26__________________ANTS__O_o__o Hamearis_lucina) Panara_3932_56) (Lepricornis_5041_80 Monethe_3977_61)) Zelotea_4057_79) ((Ancyluris_6834_45 (Caria_7625_31_______________________________ Necryia_5966_93)) Cariomothis_6518_41)) Anteros_7097_50______________________________) Hyphilaria_5239_91) Sarota_9358_21) Exoplisia_6433_40) (Leucochimona_9302_33_______________________X Mesophthalma_6327_88_____________________X)) (Pachythone_9050_18 Emesis_mandana___________________________X_____))) Themone_7643_34) Mesosemia_8822_02_________________________X) Charis_9365_19_______________________________)) Ithomiola_5924_82) Menander_7339_49_________________ANTS__O_o__o) Protonymphidia_3948_59_________ANTS__O_o__o) ((Theope_7789_65__________________ANTS__O_o__o Echenais_7168_48) (Brachyglemis_3956_58 Leucochimona_7617_87_______________________X))) Metacharis_9347_23__________________________) Symmachia_8849_08___________________________)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((Uraneis_hyalina____________________ANTS__O_o__o ((Eurybia_9363_20_________________ANTS__O_o__o ((((((((Theope_7789_65__________________ANTS__O_o__o Synargis_7605_29_________________ANTS__O_o__o) Calydna_6346_83) Themone_7643_34) Calephelis_9356_22____________________X______) Esthemopsis_7813_32) Helicopis_5318_54) Necryia_5966_93) (Theope_9026_5____________________ANTS__O_o__o (Detritivora_9327_25 Hyphilaria_5239_91)))) Menander_7339_49_________________ANTS__O_o__o)) Isapis_6815_44) ((Adelotypa_7807_37_________________ANTS__O_o__o Mesosemia_6516_85_________________________X) Setabis_6330_38_________________ANTS__O_o__o)) Methone_8523_95) Hamearis_lucina) ((((((Alesa_8380_84____________________ANTS__O_o__o Metacharis_9347_23__________________________) Lemonias_3950_55_________________ANTS__O_o__o) (Sarota_9358_21 Emesis_mandana___________________________X_____)) Cyrenia_3992_63) Thisbe_6766_43___________________ANTS__O_o__o) (((Calospila_8532_96_________________ANTS__O_o__o Apodemia_mormo) (Cariomothis_6518_41 Charis_9365_19_______________________________)) Semomesia_5223_52_________________________X))) ((((((((((((((((Protonymphidia_3948_59_________ANTS__O_o__o Anteros_7097_50______________________________) Exoplisia_6433_40) Ithomeis_7773_35) (Mesene_7450_28_________________________X______ Pandemos_8572_97)) ((((Brachyglemis_3956_58 Lepricornis_5041_80) Melanis_7612_30_________________________X______) ((((Chamaelimnas_3942_57 Styx_infernalis) (Ithomiola_5924_82 (Leucochimona_9302_33_______________________X (Mesosemia_8822_02_________________________X Roeberella_7457_36)))) Mesophthalma_6327_88_____________________X) Monethe_3977_61)) Perophthalma_9330_24_______________________X)) T_Poritia_erycinoides) Stalachtis_6422_39) Napaea_m17_89_______________________________) Symmachia_8849_08___________________________) Riodina_lysippus) Ourocnemis_4621_90___________________________) Phaenochitonia_6926_47) Caria_7625_31_______________________________) Euselasia_orfita____________________________X) (((((Juditha_9315_26__________________ANTS__O_o__o Euselasia_8816_01_________________________X) T_Miletus_ancon) Emesis_6837_46_______________________________) ((Amarynthis_5220_51 Leucochimona_9306_27_______________________X) Rhetus_3973_60)) Pachythone_9050_18)) ((((((Argyrogrammana_6709_42 Cremna_5260_53) Crocozona_3995_62) Leucochimona_7617_87_______________________X) Echenais_7168_48) Parcella_4012_64) Zelotea_4057_79))) Panara_3932_56) Ancyluris_6834_45) Thysanota_5241_81_________________ANTS__O_o__o)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((Uraneis_hyalina____________________ANTS__O_o__o (Lepricornis_5041_80 (Zelotea_4057_79 T_Poritia_erycinoides))) (((((((((((Adelotypa_7807_37_________________ANTS__O_o__o (Juditha_9315_26__________________ANTS__O_o__o Melanis_7612_30_________________________X______)) ((Theope_7789_65__________________ANTS__O_o__o (((((Detritivora_9327_25 Mesosemia_8822_02_________________________X) Themone_7643_34) (Perophthalma_9330_24_______________________X Hamearis_lucina)) Monethe_3977_61) Leucochimona_9302_33_______________________X)) Theope_9026_5____________________ANTS__O_o__o)) (((((Alesa_8380_84____________________ANTS__O_o__o Eurybia_9363_20_________________ANTS__O_o__o) (Panara_3932_56 Phaenochitonia_6926_47)) Semomesia_5223_52_________________________X) Pandemos_8572_97) (Menander_7339_49_________________ANTS__O_o__o Ancyluris_6834_45))) (Calospila_8532_96_________________ANTS__O_o__o Emesis_mandana___________________________X_____)) ((((Thysanota_5241_81_________________ANTS__O_o__o (((Crocozona_3995_62 Euselasia_orfita____________________________X) ((Esthemopsis_7813_32 Ithomiola_5924_82) Isapis_6815_44)) Ithomeis_7773_35)) Argyrogrammana_6709_42) (Chamaelimnas_3942_57 T_Miletus_ancon)) Ourocnemis_4621_90___________________________)) Anteros_7097_50______________________________) (Hyphilaria_5239_91 Napaea_m17_89_______________________________)) (Thisbe_6766_43___________________ANTS__O_o__o Exoplisia_6433_40)) (Lemonias_3950_55_________________ANTS__O_o__o Cariomothis_6518_41)) Echenais_7168_48) (Amarynthis_5220_51 Styx_infernalis))) Mesophthalma_6327_88_____________________X) (Protonymphidia_3948_59_________ANTS__O_o__o ((((((((Setabis_6330_38_________________ANTS__O_o__o Stalachtis_6422_39) Synargis_7605_29_________________ANTS__O_o__o) ((((((Calephelis_9356_22____________________X______ (((Cremna_5260_53 (Mesosemia_6516_85_________________________X Metacharis_9347_23__________________________)) Apodemia_mormo) Riodina_lysippus)) (Caria_7625_31_______________________________ (Charis_9365_19_______________________________ Helicopis_5318_54))) (Calydna_6346_83 Pachythone_9050_18)) (Necryia_5966_93 Parcella_4012_64)) (((Euselasia_8816_01_________________________X Leucochimona_7617_87_______________________X) Sarota_9358_21) Methone_8523_95)) Cyrenia_3992_63)) Leucochimona_9306_27_______________________X) Roeberella_7457_36) ((Brachyglemis_3956_58 Symmachia_8849_08___________________________) Mesene_7450_28_________________________X______)) Rhetus_3973_60) Emesis_6837_46_______________________________)))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((Uraneis_hyalina____________________ANTS__O_o__o (Napaea_m17_89_______________________________ Anteros_7097_50______________________________)) ((((((Adelotypa_7807_37_________________ANTS__O_o__o Cariomothis_6518_41) Ithomeis_7773_35) Exoplisia_6433_40) Crocozona_3995_62) ((((((Theope_7789_65__________________ANTS__O_o__o Calospila_8532_96_________________ANTS__O_o__o) Leucochimona_7617_87_______________________X) Pachythone_9050_18) (Detritivora_9327_25 Rhetus_3973_60)) Mesosemia_6516_85_________________________X) (Thisbe_6766_43___________________ANTS__O_o__o (Calephelis_9356_22____________________X______ Esthemopsis_7813_32)))) ((Chamaelimnas_3942_57 Emesis_6837_46_______________________________) Mesosemia_8822_02_________________________X))) Emesis_mandana___________________________X_____) ((Alesa_8380_84____________________ANTS__O_o__o (((Thysanota_5241_81_________________ANTS__O_o__o (Argyrogrammana_6709_42 Monethe_3977_61)) ((Charis_9365_19_______________________________ Metacharis_9347_23__________________________) Isapis_6815_44)) Apodemia_mormo)) Themone_7643_34)) Roeberella_7457_36) (((((((((Theope_9026_5____________________ANTS__O_o__o Lepricornis_5041_80) Stalachtis_6422_39) (Menander_7339_49_________________ANTS__O_o__o Euselasia_8816_01_________________________X)) ((Amarynthis_5220_51 (((((Calydna_6346_83 (Cremna_5260_53 Hyphilaria_5239_91)) ((Caria_7625_31_______________________________ Methone_8523_95) Riodina_lysippus)) Phaenochitonia_6926_47) (Melanis_7612_30_________________________X______ Euselasia_orfita____________________________X)) Mesophthalma_6327_88_____________________X)) (Mesene_7450_28_________________________X______ Hamearis_lucina))) Brachyglemis_3956_58) Echenais_7168_48) Cyrenia_3992_63) T_Miletus_ancon) (Lemonias_3950_55_________________ANTS__O_o__o (Panara_3932_56 Styx_infernalis)))) ((((((((Protonymphidia_3948_59_________ANTS__O_o__o (Leucochimona_9302_33_______________________X Symmachia_8849_08___________________________)) Helicopis_5318_54) ((Setabis_6330_38_________________ANTS__O_o__o Zelotea_4057_79) T_Poritia_erycinoides)) Leucochimona_9306_27_______________________X) Synargis_7605_29_________________ANTS__O_o__o) Ithomiola_5924_82) (((Juditha_9315_26__________________ANTS__O_o__o (((Necryia_5966_93 (Perophthalma_9330_24_______________________X Sarota_9358_21)) (Pandemos_8572_97 Parcella_4012_64)) Semomesia_5223_52_________________________X)) Ourocnemis_4621_90___________________________) Ancyluris_6834_45)) Eurybia_9363_20_________________ANTS__O_o__o))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((Uraneis_hyalina____________________ANTS__O_o__o ((Menander_7339_49_________________ANTS__O_o__o (Leucochimona_7617_87_______________________X Leucochimona_9306_27_______________________X)) Exoplisia_6433_40)) ((Synargis_7605_29_________________ANTS__O_o__o Cariomothis_6518_41) ((Amarynthis_5220_51 Anteros_7097_50______________________________) ((((Brachyglemis_3956_58 (Crocozona_3995_62 (Euselasia_8816_01_________________________X Roeberella_7457_36))) Caria_7625_31_______________________________) Ithomeis_7773_35) Calephelis_9356_22____________________X______)))) (Panara_3932_56 Perophthalma_9330_24_______________________X)) ((Calospila_8532_96_________________ANTS__O_o__o (Cremna_5260_53 Hyphilaria_5239_91)) ((Thysanota_5241_81_________________ANTS__O_o__o Detritivora_9327_25) Pachythone_9050_18))) (((((((((((((((Adelotypa_7807_37_________________ANTS__O_o__o (Esthemopsis_7813_32 Leucochimona_9302_33_______________________X)) Lepricornis_5041_80) (Echenais_7168_48 (Metacharis_9347_23__________________________ Apodemia_mormo))) Emesis_mandana___________________________X_____) (Ancyluris_6834_45 Necryia_5966_93)) Melanis_7612_30_________________________X______) ((Alesa_8380_84____________________ANTS__O_o__o (Chamaelimnas_3942_57 Euselasia_orfita____________________________X)) Ithomiola_5924_82)) Riodina_lysippus) Themone_7643_34) (Protonymphidia_3948_59_________ANTS__O_o__o (((Calydna_6346_83 Mesophthalma_6327_88_____________________X) T_Poritia_erycinoides) Symmachia_8849_08___________________________))) (((((((((((((Setabis_6330_38_________________ANTS__O_o__o (Thisbe_6766_43___________________ANTS__O_o__o Stalachtis_6422_39)) Hamearis_lucina) Styx_infernalis) Lemonias_3950_55_________________ANTS__O_o__o) Cyrenia_3992_63) ((Juditha_9315_26__________________ANTS__O_o__o Argyrogrammana_6709_42) Napaea_m17_89_______________________________)) Pandemos_8572_97) ((Eurybia_9363_20_________________ANTS__O_o__o (Mesosemia_8822_02_________________________X Rhetus_3973_60)) Charis_9365_19_______________________________)) Parcella_4012_64) (Isapis_6815_44 Sarota_9358_21)) Mesene_7450_28_________________________X______) Theope_9026_5____________________ANTS__O_o__o) ((((Helicopis_5318_54 Ourocnemis_4621_90___________________________) Monethe_3977_61) Semomesia_5223_52_________________________X) Methone_8523_95))) Mesosemia_6516_85_________________________X) T_Miletus_ancon) Zelotea_4057_79) Phaenochitonia_6926_47)) Theope_7789_65__________________ANTS__O_o__o) Emesis_6837_46_______________________________)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((Uraneis_hyalina____________________ANTS__O_o__o Helicopis_5318_54) Apodemia_mormo) Leucochimona_9306_27_______________________X) ((((((((((((((((Protonymphidia_3948_59_________ANTS__O_o__o Symmachia_8849_08___________________________) Hamearis_lucina) (Cremna_5260_53 Zelotea_4057_79)) (Mesosemia_6516_85_________________________X Parcella_4012_64)) Juditha_9315_26__________________ANTS__O_o__o) Styx_infernalis) T_Miletus_ancon) Euselasia_orfita____________________________X) Pandemos_8572_97) ((((Euselasia_8816_01_________________________X Ithomiola_5924_82) Monethe_3977_61) Ourocnemis_4621_90___________________________) Leucochimona_7617_87_______________________X)) Isapis_6815_44) ((((((((((Setabis_6330_38_________________ANTS__O_o__o ((((((((((((Theope_7789_65__________________ANTS__O_o__o Amarynthis_5220_51) Rhetus_3973_60) (Thisbe_6766_43___________________ANTS__O_o__o Cyrenia_3992_63)) ((Synargis_7605_29_________________ANTS__O_o__o Themone_7643_34) Roeberella_7457_36)) ((Crocozona_3995_62 Stalachtis_6422_39) Necryia_5966_93)) ((Thysanota_5241_81_________________ANTS__O_o__o Detritivora_9327_25) T_Poritia_erycinoides)) Caria_7625_31_______________________________) Methone_8523_95) ((Echenais_7168_48 Ithomeis_7773_35) Metacharis_9347_23__________________________)) Cariomothis_6518_41) Menander_7339_49_________________ANTS__O_o__o) Riodina_lysippus)) Pachythone_9050_18) (Lepricornis_5041_80 Mesene_7450_28_________________________X______)) Phaenochitonia_6926_47) (Lemonias_3950_55_________________ANTS__O_o__o (Leucochimona_9302_33_______________________X Mesophthalma_6327_88_____________________X))) Theope_9026_5____________________ANTS__O_o__o) Eurybia_9363_20_________________ANTS__O_o__o) Argyrogrammana_6709_42) (Brachyglemis_3956_58 Exoplisia_6433_40)) Chamaelimnas_3942_57)) Alesa_8380_84____________________ANTS__O_o__o) (Ancyluris_6834_45 (Panara_3932_56 Emesis_mandana___________________________X_____))) Sarota_9358_21) Melanis_7612_30_________________________X______)) ((((Adelotypa_7807_37_________________ANTS__O_o__o ((Calephelis_9356_22____________________X______ (Charis_9365_19_______________________________ Anteros_7097_50______________________________)) Napaea_m17_89_______________________________)) (Hyphilaria_5239_91 Perophthalma_9330_24_______________________X)) Calospila_8532_96_________________ANTS__O_o__o) (((Calydna_6346_83 Mesosemia_8822_02_________________________X) Esthemopsis_7813_32) Semomesia_5223_52_________________________X))) Emesis_6837_46_______________________________)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o (Helicopis_5318_54 Pachythone_9050_18)) Leucochimona_7617_87_______________________X) ((((((((((((((((Protonymphidia_3948_59_________ANTS__O_o__o Riodina_lysippus) Calydna_6346_83) Pandemos_8572_97) ((Echenais_7168_48 Phaenochitonia_6926_47) Esthemopsis_7813_32)) Eurybia_9363_20_________________ANTS__O_o__o) Hamearis_lucina) Brachyglemis_3956_58) Mesosemia_8822_02_________________________X) ((((Alesa_8380_84____________________ANTS__O_o__o (Ourocnemis_4621_90___________________________ Roeberella_7457_36)) Styx_infernalis) ((Theope_7789_65__________________ANTS__O_o__o (Juditha_9315_26__________________ANTS__O_o__o Napaea_m17_89_______________________________)) (((Cariomothis_6518_41 Zelotea_4057_79) Sarota_9358_21) Mesene_7450_28_________________________X______))) Symmachia_8849_08___________________________)) Ithomeis_7773_35) Anteros_7097_50______________________________) ((Lemonias_3950_55_________________ANTS__O_o__o Metacharis_9347_23__________________________) (Chamaelimnas_3942_57 Euselasia_orfita____________________________X))) Stalachtis_6422_39) Theope_9026_5____________________ANTS__O_o__o) Setabis_6330_38_________________ANTS__O_o__o) Argyrogrammana_6709_42)) ((((((Synargis_7605_29_________________ANTS__O_o__o Calephelis_9356_22____________________X______) Amarynthis_5220_51) Lepricornis_5041_80) Ithomiola_5924_82) (((Crocozona_3995_62 T_Poritia_erycinoides) ((Isapis_6815_44 Semomesia_5223_52_________________________X) Mesosemia_6516_85_________________________X)) Perophthalma_9330_24_______________________X)) Mesophthalma_6327_88_____________________X)) (Menander_7339_49_________________ANTS__O_o__o Calospila_8532_96_________________ANTS__O_o__o)) ((Cyrenia_3992_63 Exoplisia_6433_40) Monethe_3977_61)) Methone_8523_95) Detritivora_9327_25) T_Miletus_ancon) Leucochimona_9306_27_______________________X) Necryia_5966_93) Charis_9365_19_______________________________) Parcella_4012_64) Emesis_6837_46_______________________________) Melanis_7612_30_________________________X______) (Leucochimona_9302_33_______________________X Themone_7643_34)) (((Thysanota_5241_81_________________ANTS__O_o__o Hyphilaria_5239_91) (Caria_7625_31_______________________________ Cremna_5260_53)) Apodemia_mormo)) Panara_3932_56) (Thisbe_6766_43___________________ANTS__O_o__o ((Ancyluris_6834_45 Euselasia_8816_01_________________________X) Rhetus_3973_60))) (Adelotypa_7807_37_________________ANTS__O_o__o Emesis_mandana___________________________X_____))*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((Uraneis_hyalina____________________ANTS__O_o__o (((((((((Protonymphidia_3948_59_________ANTS__O_o__o Isapis_6815_44) (Mesosemia_6516_85_________________________X Mesosemia_8822_02_________________________X)) Hyphilaria_5239_91) Cariomothis_6518_41) Parcella_4012_64) Leucochimona_9302_33_______________________X) Helicopis_5318_54) Anteros_7097_50______________________________) Hamearis_lucina)) Riodina_lysippus) Apodemia_mormo) Detritivora_9327_25) ((Adelotypa_7807_37_________________ANTS__O_o__o Ourocnemis_4621_90___________________________) Lepricornis_5041_80)) Themone_7643_34) (((((Menander_7339_49_________________ANTS__O_o__o (Exoplisia_6433_40 Ithomiola_5924_82)) Thysanota_5241_81_________________ANTS__O_o__o) Methone_8523_95) Necryia_5966_93) (Chamaelimnas_3942_57 ((Phaenochitonia_6926_47 Emesis_mandana___________________________X_____) Stalachtis_6422_39)))) ((Alesa_8380_84____________________ANTS__O_o__o (Euselasia_orfita____________________________X T_Miletus_ancon)) (((((((((((Eurybia_9363_20_________________ANTS__O_o__o (Brachyglemis_3956_58 Mesene_7450_28_________________________X______)) ((((Calospila_8532_96_________________ANTS__O_o__o (((Echenais_7168_48 Rhetus_3973_60) Perophthalma_9330_24_______________________X) (Ithomeis_7773_35 Metacharis_9347_23__________________________))) (Caria_7625_31_______________________________ Roeberella_7457_36)) Charis_9365_19_______________________________) Cremna_5260_53)) Crocozona_3995_62) Sarota_9358_21) Theope_7789_65__________________ANTS__O_o__o) Pachythone_9050_18) Theope_9026_5____________________ANTS__O_o__o) (Napaea_m17_89_______________________________ (Pandemos_8572_97 Semomesia_5223_52_________________________X))) Melanis_7612_30_________________________X______) T_Poritia_erycinoides) Leucochimona_7617_87_______________________X))) Synargis_7605_29_________________ANTS__O_o__o) Juditha_9315_26__________________ANTS__O_o__o) Monethe_3977_61) (Lemonias_3950_55_________________ANTS__O_o__o Euselasia_8816_01_________________________X)) Amarynthis_5220_51) (((((((Setabis_6330_38_________________ANTS__O_o__o Argyrogrammana_6709_42) (Cyrenia_3992_63 Esthemopsis_7813_32)) Styx_infernalis) Leucochimona_9306_27_______________________X) ((Thisbe_6766_43___________________ANTS__O_o__o Zelotea_4057_79) (Ancyluris_6834_45 (Calephelis_9356_22____________________X______ Emesis_6837_46_______________________________)))) ((Mesophthalma_6327_88_____________________X Panara_3932_56) Symmachia_8849_08___________________________)) Calydna_6346_83))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Methone_8523_95) Euselasia_orfita____________________________X) (Ancyluris_6834_45 Calydna_6346_83)) ((((((Adelotypa_7807_37_________________ANTS__O_o__o Sarota_9358_21) Charis_9365_19_______________________________) (Theope_7789_65__________________ANTS__O_o__o (Menander_7339_49_________________ANTS__O_o__o (Napaea_m17_89_______________________________ Zelotea_4057_79)))) (Argyrogrammana_6709_42 (Emesis_mandana___________________________X_____ Styx_infernalis))) Eurybia_9363_20_________________ANTS__O_o__o) (Emesis_6837_46_______________________________ Exoplisia_6433_40))) (((((Setabis_6330_38_________________ANTS__O_o__o Brachyglemis_3956_58) (Thisbe_6766_43___________________ANTS__O_o__o (((Cariomothis_6518_41 Mesophthalma_6327_88_____________________X) Ourocnemis_4621_90___________________________) (Leucochimona_7617_87_______________________X Symmachia_8849_08___________________________)))) Semomesia_5223_52_________________________X) Helicopis_5318_54) Calospila_8532_96_________________ANTS__O_o__o)) ((((Chamaelimnas_3942_57 Hamearis_lucina) ((((Hyphilaria_5239_91 Roeberella_7457_36) Panara_3932_56) (Pachythone_9050_18 T_Miletus_ancon)) T_Poritia_erycinoides)) Mesosemia_6516_85_________________________X) Mesosemia_8822_02_________________________X)) Necryia_5966_93) (Leucochimona_9302_33_______________________X Mesene_7450_28_________________________X______)) ((Caria_7625_31_______________________________ (Cyrenia_3992_63 Pandemos_8572_97)) Stalachtis_6422_39)) Protonymphidia_3948_59_________ANTS__O_o__o) ((((((((Cremna_5260_53 Rhetus_3973_60) ((((Ithomiola_5924_82 Leucochimona_9306_27_______________________X) Riodina_lysippus) Lepricornis_5041_80) Melanis_7612_30_________________________X______)) Apodemia_mormo) Phaenochitonia_6926_47) Esthemopsis_7813_32) Isapis_6815_44) Ithomeis_7773_35) Parcella_4012_64)) Metacharis_9347_23__________________________) ((Amarynthis_5220_51 Crocozona_3995_62) Detritivora_9327_25)) Monethe_3977_61) Euselasia_8816_01_________________________X) ((Lemonias_3950_55_________________ANTS__O_o__o Themone_7643_34) Thysanota_5241_81_________________ANTS__O_o__o)) Alesa_8380_84____________________ANTS__O_o__o) (Theope_9026_5____________________ANTS__O_o__o ((((Synargis_7605_29_________________ANTS__O_o__o Echenais_7168_48) Anteros_7097_50______________________________) Perophthalma_9330_24_______________________X) Calephelis_9356_22____________________X______))) Juditha_9315_26__________________ANTS__O_o__o)*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Menander_7339_49_________________ANTS__O_o__o) Amarynthis_5220_51) Ancyluris_6834_45) Ithomiola_5924_82) ((Euselasia_8816_01_________________________X Roeberella_7457_36) (Leucochimona_7617_87_______________________X Panara_3932_56))) ((((Alesa_8380_84____________________ANTS__O_o__o (Cariomothis_6518_41 (Esthemopsis_7813_32 Melanis_7612_30_________________________X______))) Chamaelimnas_3942_57) Mesene_7450_28_________________________X______) (Symmachia_8849_08___________________________ Apodemia_mormo))) Perophthalma_9330_24_______________________X) Euselasia_orfita____________________________X) (((((Thysanota_5241_81_________________ANTS__O_o__o (Leucochimona_9302_33_______________________X T_Miletus_ancon)) Methone_8523_95) Rhetus_3973_60) (Sarota_9358_21 Styx_infernalis)) Detritivora_9327_25)) (Hyphilaria_5239_91 Emesis_mandana___________________________X_____)) ((((((((Adelotypa_7807_37_________________ANTS__O_o__o (Juditha_9315_26__________________ANTS__O_o__o Brachyglemis_3956_58)) Exoplisia_6433_40) ((((Eurybia_9363_20_________________ANTS__O_o__o ((Theope_9026_5____________________ANTS__O_o__o Riodina_lysippus) Thisbe_6766_43___________________ANTS__O_o__o)) Pachythone_9050_18) Theope_7789_65__________________ANTS__O_o__o) ((Echenais_7168_48 Stalachtis_6422_39) Helicopis_5318_54))) (Calydna_6346_83 Napaea_m17_89_______________________________)) Mesophthalma_6327_88_____________________X) Calospila_8532_96_________________ANTS__O_o__o) (Leucochimona_9306_27_______________________X Zelotea_4057_79)) Crocozona_3995_62)) Cremna_5260_53) (((Protonymphidia_3948_59_________ANTS__O_o__o (((Calephelis_9356_22____________________X______ Lepricornis_5041_80) (Caria_7625_31_______________________________ Ourocnemis_4621_90___________________________)) ((Charis_9365_19_______________________________ (((((Isapis_6815_44 (Metacharis_9347_23__________________________ Hamearis_lucina)) Ithomeis_7773_35) Parcella_4012_64) Phaenochitonia_6926_47) Mesosemia_6516_85_________________________X)) Emesis_6837_46_______________________________))) Argyrogrammana_6709_42) Anteros_7097_50______________________________)) Lemonias_3950_55_________________ANTS__O_o__o) Monethe_3977_61) Mesosemia_8822_02_________________________X) Setabis_6330_38_________________ANTS__O_o__o) Necryia_5966_93) (((Synargis_7605_29_________________ANTS__O_o__o ((Pandemos_8572_97 T_Poritia_erycinoides) Themone_7643_34)) Semomesia_5223_52_________________________X) Cyrenia_3992_63))*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((Uraneis_hyalina____________________ANTS__O_o__o (((((((Alesa_8380_84____________________ANTS__O_o__o Hyphilaria_5239_91) (Theope_9026_5____________________ANTS__O_o__o Mesene_7450_28_________________________X______)) Stalachtis_6422_39) Ancyluris_6834_45) Calospila_8532_96_________________ANTS__O_o__o) ((Brachyglemis_3956_58 Perophthalma_9330_24_______________________X) Cariomothis_6518_41)) Monethe_3977_61)) (((((Theope_7789_65__________________ANTS__O_o__o Emesis_mandana___________________________X_____) (Synargis_7605_29_________________ANTS__O_o__o Ourocnemis_4621_90___________________________)) Necryia_5966_93) Themone_7643_34) Leucochimona_9302_33_______________________X)) Thysanota_5241_81_________________ANTS__O_o__o) ((((((((Setabis_6330_38_________________ANTS__O_o__o ((((Menander_7339_49_________________ANTS__O_o__o Mesophthalma_6327_88_____________________X) Juditha_9315_26__________________ANTS__O_o__o) T_Poritia_erycinoides) (((Esthemopsis_7813_32 Phaenochitonia_6926_47) (Sarota_9358_21 Zelotea_4057_79)) (((Exoplisia_6433_40 Symmachia_8849_08___________________________) (Ithomiola_5924_82 Melanis_7612_30_________________________X______)) ((Lepricornis_5041_80 Pachythone_9050_18) (Methone_8523_95 T_Miletus_ancon)))))) Isapis_6815_44) ((Emesis_6837_46_______________________________ Leucochimona_7617_87_______________________X) Anteros_7097_50______________________________)) Metacharis_9347_23__________________________) Parcella_4012_64) ((Euselasia_8816_01_________________________X Rhetus_3973_60) Riodina_lysippus)) (((Lemonias_3950_55_________________ANTS__O_o__o Ithomeis_7773_35) Charis_9365_19_______________________________) Cyrenia_3992_63)) Echenais_7168_48)) Apodemia_mormo) Panara_3932_56) ((Caria_7625_31_______________________________ Semomesia_5223_52_________________________X) Hamearis_lucina)) ((Calydna_6346_83 Crocozona_3995_62) (Cremna_5260_53 Euselasia_orfita____________________________X))) Calephelis_9356_22____________________X______) (Thisbe_6766_43___________________ANTS__O_o__o Mesosemia_8822_02_________________________X)) ((((Adelotypa_7807_37_________________ANTS__O_o__o ((Eurybia_9363_20_________________ANTS__O_o__o Helicopis_5318_54) (Detritivora_9327_25 Napaea_m17_89_______________________________))) Argyrogrammana_6709_42) Amarynthis_5220_51) (Pandemos_8572_97 Styx_infernalis))) (Leucochimona_9306_27_______________________X Roeberella_7457_36)) (Chamaelimnas_3942_57 Mesosemia_6516_85_________________________X)) Protonymphidia_3948_59_________ANTS__O_o__o)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((Uraneis_hyalina____________________ANTS__O_o__o Perophthalma_9330_24_______________________X) ((((((((((((((Protonymphidia_3948_59_________ANTS__O_o__o Esthemopsis_7813_32) (((((Synargis_7605_29_________________ANTS__O_o__o (Cyrenia_3992_63 Lepricornis_5041_80)) Euselasia_8816_01_________________________X) Leucochimona_9302_33_______________________X) Ithomeis_7773_35) Hyphilaria_5239_91)) Mesosemia_6516_85_________________________X) Pandemos_8572_97) (Lemonias_3950_55_________________ANTS__O_o__o ((Juditha_9315_26__________________ANTS__O_o__o Semomesia_5223_52_________________________X) (Argyrogrammana_6709_42 Zelotea_4057_79)))) Phaenochitonia_6926_47) Setabis_6330_38_________________ANTS__O_o__o) Methone_8523_95) Calospila_8532_96_________________ANTS__O_o__o) Thysanota_5241_81_________________ANTS__O_o__o) Alesa_8380_84____________________ANTS__O_o__o) (Chamaelimnas_3942_57 Pachythone_9050_18)) (Calephelis_9356_22____________________X______ Exoplisia_6433_40)) Isapis_6815_44)) Parcella_4012_64) ((((((((((((((Adelotypa_7807_37_________________ANTS__O_o__o Themone_7643_34) Mesophthalma_6327_88_____________________X) Napaea_m17_89_______________________________) Leucochimona_9306_27_______________________X) (Hamearis_lucina Styx_infernalis)) Euselasia_orfita____________________________X) Apodemia_mormo) Echenais_7168_48) (((((((((Eurybia_9363_20_________________ANTS__O_o__o Cremna_5260_53) ((Cariomothis_6518_41 (Rhetus_3973_60 Anteros_7097_50______________________________)) ((Necryia_5966_93 Panara_3932_56) Stalachtis_6422_39))) Theope_7789_65__________________ANTS__O_o__o) Charis_9365_19_______________________________) ((Brachyglemis_3956_58 Mesene_7450_28_________________________X______) Ithomiola_5924_82)) Riodina_lysippus) Leucochimona_7617_87_______________________X) ((Menander_7339_49_________________ANTS__O_o__o T_Poritia_erycinoides) (Ancyluris_6834_45 Emesis_6837_46_______________________________))) Metacharis_9347_23__________________________)) (((Caria_7625_31_______________________________ Ourocnemis_4621_90___________________________) Roeberella_7457_36) Helicopis_5318_54)) Theope_9026_5____________________ANTS__O_o__o) Thisbe_6766_43___________________ANTS__O_o__o) Melanis_7612_30_________________________X______) (((((Amarynthis_5220_51 (Crocozona_3995_62 (Mesosemia_8822_02_________________________X T_Miletus_ancon))) Emesis_mandana___________________________X_____) (Detritivora_9327_25 Monethe_3977_61)) Sarota_9358_21) (Calydna_6346_83 Symmachia_8849_08___________________________))))*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o ((Adelotypa_7807_37_________________ANTS__O_o__o ((Thisbe_6766_43___________________ANTS__O_o__o Apodemia_mormo) Cyrenia_3992_63)) Hyphilaria_5239_91)) ((((Cremna_5260_53 Rhetus_3973_60) Leucochimona_9302_33_______________________X) (Napaea_m17_89_______________________________ Ourocnemis_4621_90___________________________)) Monethe_3977_61)) Emesis_mandana___________________________X_____) ((Alesa_8380_84____________________ANTS__O_o__o (Perophthalma_9330_24_______________________X T_Miletus_ancon)) (((((Calephelis_9356_22____________________X______ (Ithomiola_5924_82 Riodina_lysippus)) Helicopis_5318_54) Semomesia_5223_52_________________________X) (Chamaelimnas_3942_57 Themone_7643_34)) Exoplisia_6433_40))) ((((((((Protonymphidia_3948_59_________ANTS__O_o__o Hamearis_lucina) (Eurybia_9363_20_________________ANTS__O_o__o ((Mesene_7450_28_________________________X______ (Parcella_4012_64 Zelotea_4057_79)) Necryia_5966_93))) ((Brachyglemis_3956_58 Leucochimona_7617_87_______________________X) (Metacharis_9347_23__________________________ Sarota_9358_21))) Pandemos_8572_97) Caria_7625_31_______________________________) ((((Theope_7789_65__________________ANTS__O_o__o (((((Lemonias_3950_55_________________ANTS__O_o__o Emesis_6837_46_______________________________) (Echenais_7168_48 Styx_infernalis)) Anteros_7097_50______________________________) Ithomeis_7773_35) (((Amarynthis_5220_51 Methone_8523_95) Symmachia_8849_08___________________________) Stalachtis_6422_39))) ((Synargis_7605_29_________________ANTS__O_o__o Panara_3932_56) Lepricornis_5041_80)) T_Poritia_erycinoides) Mesosemia_6516_85_________________________X)) (Theope_9026_5____________________ANTS__O_o__o Melanis_7612_30_________________________X______)) Phaenochitonia_6926_47)) Pachythone_9050_18) Leucochimona_9306_27_______________________X) Ancyluris_6834_45) Roeberella_7457_36) Mesophthalma_6327_88_____________________X) Isapis_6815_44) ((Argyrogrammana_6709_42 ((Crocozona_3995_62 Esthemopsis_7813_32) (Detritivora_9327_25 Euselasia_8816_01_________________________X))) (Charis_9365_19_______________________________ Mesosemia_8822_02_________________________X))) Juditha_9315_26__________________ANTS__O_o__o) ((Setabis_6330_38_________________ANTS__O_o__o Cariomothis_6518_41) Menander_7339_49_________________ANTS__O_o__o)) Thysanota_5241_81_________________ANTS__O_o__o) Euselasia_orfita____________________________X) Calospila_8532_96_________________ANTS__O_o__o) Calydna_6346_83)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((Uraneis_hyalina____________________ANTS__O_o__o ((Menander_7339_49_________________ANTS__O_o__o Isapis_6815_44) (Emesis_6837_46_______________________________ Metacharis_9347_23__________________________))) T_Poritia_erycinoides) Monethe_3977_61) Melanis_7612_30_________________________X______) ((((Calospila_8532_96_________________ANTS__O_o__o (((Argyrogrammana_6709_42 Necryia_5966_93) Brachyglemis_3956_58) Leucochimona_9302_33_______________________X)) T_Miletus_ancon) Calydna_6346_83) (Esthemopsis_7813_32 (Parcella_4012_64 Semomesia_5223_52_________________________X)))) Theope_9026_5____________________ANTS__O_o__o) (((((((((((Adelotypa_7807_37_________________ANTS__O_o__o Rhetus_3973_60) Eurybia_9363_20_________________ANTS__O_o__o) ((((((((((((((Alesa_8380_84____________________ANTS__O_o__o Mesophthalma_6327_88_____________________X) Anteros_7097_50______________________________) ((Crocozona_3995_62 (Mesosemia_6516_85_________________________X Styx_infernalis)) Lepricornis_5041_80)) Roeberella_7457_36) Echenais_7168_48) Caria_7625_31_______________________________) Synargis_7605_29_________________ANTS__O_o__o) (Juditha_9315_26__________________ANTS__O_o__o (Thisbe_6766_43___________________ANTS__O_o__o (Ourocnemis_4621_90___________________________ ((Zelotea_4057_79 Emesis_mandana___________________________X_____) Euselasia_orfita____________________________X))))) ((Leucochimona_7617_87_______________________X Sarota_9358_21) Themone_7643_34)) Cyrenia_3992_63) Setabis_6330_38_________________ANTS__O_o__o) (Mesene_7450_28_________________________X______ Phaenochitonia_6926_47)) (Amarynthis_5220_51 (Charis_9365_19_______________________________ (Exoplisia_6433_40 (Hyphilaria_5239_91 Symmachia_8849_08___________________________))))) Riodina_lysippus)) Methone_8523_95) Ancyluris_6834_45) Leucochimona_9306_27_______________________X) Mesosemia_8822_02_________________________X) ((((Protonymphidia_3948_59_________ANTS__O_o__o Ithomiola_5924_82) Pachythone_9050_18) Pandemos_8572_97) (Theope_7789_65__________________ANTS__O_o__o (Chamaelimnas_3942_57 Euselasia_8816_01_________________________X)))) ((Lemonias_3950_55_________________ANTS__O_o__o (Helicopis_5318_54 Apodemia_mormo)) Napaea_m17_89_______________________________)) Hamearis_lucina) Cariomothis_6518_41)) Thysanota_5241_81_________________ANTS__O_o__o) (Calephelis_9356_22____________________X______ ((Cremna_5260_53 Perophthalma_9330_24_______________________X) (Ithomeis_7773_35 Panara_3932_56)))) (Detritivora_9327_25 Stalachtis_6422_39))*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((Uraneis_hyalina____________________ANTS__O_o__o (((((Adelotypa_7807_37_________________ANTS__O_o__o Pachythone_9050_18) Thisbe_6766_43___________________ANTS__O_o__o) Cariomothis_6518_41) ((((Eurybia_9363_20_________________ANTS__O_o__o Ithomiola_5924_82) Roeberella_7457_36) (Charis_9365_19_______________________________ Hamearis_lucina)) ((((((((Calospila_8532_96_________________ANTS__O_o__o (Juditha_9315_26__________________ANTS__O_o__o Exoplisia_6433_40)) Styx_infernalis) (Hyphilaria_5239_91 Themone_7643_34)) Ancyluris_6834_45) (Lepricornis_5041_80 Napaea_m17_89_______________________________)) Sarota_9358_21) (((Detritivora_9327_25 Leucochimona_9302_33_______________________X) Esthemopsis_7813_32) (Parcella_4012_64 T_Miletus_ancon))) Phaenochitonia_6926_47))) Mesosemia_8822_02_________________________X)) Mesophthalma_6327_88_____________________X) ((((Protonymphidia_3948_59_________ANTS__O_o__o (Alesa_8380_84____________________ANTS__O_o__o (Menander_7339_49_________________ANTS__O_o__o Metacharis_9347_23__________________________))) Theope_9026_5____________________ANTS__O_o__o) Apodemia_mormo) (Chamaelimnas_3942_57 Leucochimona_7617_87_______________________X))) (((((((((((((Setabis_6330_38_________________ANTS__O_o__o Echenais_7168_48) (Calephelis_9356_22____________________X______ (((((Emesis_6837_46_______________________________ Mesene_7450_28_________________________X______) Anteros_7097_50______________________________) Leucochimona_9306_27_______________________X) Euselasia_8816_01_________________________X) Helicopis_5318_54))) Isapis_6815_44) Pandemos_8572_97) (Amarynthis_5220_51 Perophthalma_9330_24_______________________X)) ((Mesosemia_6516_85_________________________X Zelotea_4057_79) Symmachia_8849_08___________________________)) Ithomeis_7773_35) Semomesia_5223_52_________________________X) (Caria_7625_31_______________________________ Euselasia_orfita____________________________X)) (((Synargis_7605_29_________________ANTS__O_o__o Ourocnemis_4621_90___________________________) (Argyrogrammana_6709_42 Emesis_mandana___________________________X_____)) (((((Thysanota_5241_81_________________ANTS__O_o__o ((Necryia_5966_93 Rhetus_3973_60) Stalachtis_6422_39)) Crocozona_3995_62) Brachyglemis_3956_58) T_Poritia_erycinoides) Monethe_3977_61))) Lemonias_3950_55_________________ANTS__O_o__o) Riodina_lysippus) Melanis_7612_30_________________________X______)) Theope_7789_65__________________ANTS__O_o__o) (Methone_8523_95 Panara_3932_56)) (Cremna_5260_53 Cyrenia_3992_63)) Calydna_6346_83)*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Menander_7339_49_________________ANTS__O_o__o) (((Thisbe_6766_43___________________ANTS__O_o__o ((Argyrogrammana_6709_42 Euselasia_8816_01_________________________X) Cremna_5260_53)) Apodemia_mormo) Chamaelimnas_3942_57)) (Exoplisia_6433_40 (Rhetus_3973_60 Symmachia_8849_08___________________________))) Cariomothis_6518_41) ((((((((Alesa_8380_84____________________ANTS__O_o__o Esthemopsis_7813_32) ((Emesis_6837_46_______________________________ (Mesophthalma_6327_88_____________________X Metacharis_9347_23__________________________)) (Mesene_7450_28_________________________X______ Panara_3932_56))) Leucochimona_9306_27_______________________X) (Methone_8523_95 Sarota_9358_21)) (Calephelis_9356_22____________________X______ T_Poritia_erycinoides)) Caria_7625_31_______________________________) (((Calydna_6346_83 Zelotea_4057_79) (Monethe_3977_61 Riodina_lysippus)) ((Cyrenia_3992_63 Phaenochitonia_6926_47) Lepricornis_5041_80))) Mesosemia_8822_02_________________________X)) Theope_9026_5____________________ANTS__O_o__o) ((((Setabis_6330_38_________________ANTS__O_o__o Emesis_mandana___________________________X_____) Themone_7643_34) Detritivora_9327_25) Ithomeis_7773_35)) ((Calospila_8532_96_________________ANTS__O_o__o Mesosemia_6516_85_________________________X) Helicopis_5318_54)) Perophthalma_9330_24_______________________X) Synargis_7605_29_________________ANTS__O_o__o) (Protonymphidia_3948_59_________ANTS__O_o__o Charis_9365_19_______________________________)) (Amarynthis_5220_51 Pandemos_8572_97)) Leucochimona_9302_33_______________________X) ((((Adelotypa_7807_37_________________ANTS__O_o__o (Ourocnemis_4621_90___________________________ Parcella_4012_64)) Ancyluris_6834_45) ((((Theope_7789_65__________________ANTS__O_o__o Hamearis_lucina) ((Juditha_9315_26__________________ANTS__O_o__o (Thysanota_5241_81_________________ANTS__O_o__o Napaea_m17_89_______________________________)) Leucochimona_7617_87_______________________X)) Styx_infernalis) (Lemonias_3950_55_________________ANTS__O_o__o Crocozona_3995_62))) (Eurybia_9363_20_________________ANTS__O_o__o Pachythone_9050_18))) ((((((Brachyglemis_3956_58 ((Necryia_5966_93 Anteros_7097_50______________________________) Stalachtis_6422_39)) Euselasia_orfita____________________________X) T_Miletus_ancon) Isapis_6815_44) Echenais_7168_48) (Hyphilaria_5239_91 Melanis_7612_30_________________________X______))) (Ithomiola_5924_82 Roeberella_7457_36)) Semomesia_5223_52_________________________X)*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((Uraneis_hyalina____________________ANTS__O_o__o (Cremna_5260_53 (Necryia_5966_93 Euselasia_orfita____________________________X))) ((Calospila_8532_96_________________ANTS__O_o__o (Argyrogrammana_6709_42 ((Brachyglemis_3956_58 (Emesis_6837_46_______________________________ Leucochimona_9306_27_______________________X)) (((Isapis_6815_44 Monethe_3977_61) T_Poritia_erycinoides) Mesosemia_8822_02_________________________X)))) Rhetus_3973_60)) Semomesia_5223_52_________________________X) ((Calephelis_9356_22____________________X______ (Charis_9365_19_______________________________ Crocozona_3995_62)) Cyrenia_3992_63)) (((Theope_7789_65__________________ANTS__O_o__o Detritivora_9327_25) Emesis_mandana___________________________X_____) ((Thisbe_6766_43___________________ANTS__O_o__o (Pandemos_8572_97 Symmachia_8849_08___________________________)) T_Miletus_ancon))) (((((Alesa_8380_84____________________ANTS__O_o__o Panara_3932_56) Perophthalma_9330_24_______________________X) (Ancyluris_6834_45 Calydna_6346_83)) (Cariomothis_6518_41 Anteros_7097_50______________________________)) (Eurybia_9363_20_________________ANTS__O_o__o Themone_7643_34))) ((((((Adelotypa_7807_37_________________ANTS__O_o__o Leucochimona_9302_33_______________________X) (Setabis_6330_38_________________ANTS__O_o__o Ourocnemis_4621_90___________________________)) (Menander_7339_49_________________ANTS__O_o__o Synargis_7605_29_________________ANTS__O_o__o)) (((((((((((((((Protonymphidia_3948_59_________ANTS__O_o__o Echenais_7168_48) Chamaelimnas_3942_57) ((((((Lemonias_3950_55_________________ANTS__O_o__o Mesophthalma_6327_88_____________________X) Sarota_9358_21) Metacharis_9347_23__________________________) Mesene_7450_28_________________________X______) Thysanota_5241_81_________________ANTS__O_o__o) ((((Hyphilaria_5239_91 Roeberella_7457_36) Hamearis_lucina) Pachythone_9050_18) Riodina_lysippus))) Euselasia_8816_01_________________________X) Mesosemia_6516_85_________________________X) (Helicopis_5318_54 (Lepricornis_5041_80 Melanis_7612_30_________________________X______))) Exoplisia_6433_40) Methone_8523_95) Zelotea_4057_79) Ithomiola_5924_82) Styx_infernalis) Apodemia_mormo) (Juditha_9315_26__________________ANTS__O_o__o Amarynthis_5220_51)) Stalachtis_6422_39) Napaea_m17_89_______________________________)) Theope_9026_5____________________ANTS__O_o__o) (Leucochimona_7617_87_______________________X Parcella_4012_64))) ((Caria_7625_31_______________________________ Phaenochitonia_6926_47) Esthemopsis_7813_32)) Ithomeis_7773_35)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o ((((((((Adelotypa_7807_37_________________ANTS__O_o__o ((Caria_7625_31_______________________________ Pachythone_9050_18) (Ithomeis_7773_35 Leucochimona_7617_87_______________________X))) (Alesa_8380_84____________________ANTS__O_o__o Mesene_7450_28_________________________X______)) ((Theope_9026_5____________________ANTS__O_o__o T_Poritia_erycinoides) ((Synargis_7605_29_________________ANTS__O_o__o Necryia_5966_93) (Euselasia_8816_01_________________________X (Lepricornis_5041_80 Phaenochitonia_6926_47))))) (((((Protonymphidia_3948_59_________ANTS__O_o__o Melanis_7612_30_________________________X______) Perophthalma_9330_24_______________________X) Leucochimona_9302_33_______________________X) ((Isapis_6815_44 Parcella_4012_64) Napaea_m17_89_______________________________)) (Chamaelimnas_3942_57 Cremna_5260_53))) Sarota_9358_21) Cyrenia_3992_63) Semomesia_5223_52_________________________X) Emesis_mandana___________________________X_____)) Cariomothis_6518_41) Zelotea_4057_79) Stalachtis_6422_39) Calephelis_9356_22____________________X______) (Calospila_8532_96_________________ANTS__O_o__o Charis_9365_19_______________________________)) Ithomiola_5924_82) Calydna_6346_83) Ourocnemis_4621_90___________________________) Thisbe_6766_43___________________ANTS__O_o__o) Symmachia_8849_08___________________________) (((((((((Setabis_6330_38_________________ANTS__O_o__o (Amarynthis_5220_51 (Detritivora_9327_25 (((Echenais_7168_48 Themone_7643_34) Metacharis_9347_23__________________________) Hyphilaria_5239_91)))) ((Helicopis_5318_54 Methone_8523_95) Hamearis_lucina)) Mesophthalma_6327_88_____________________X) Rhetus_3973_60) Pandemos_8572_97) Apodemia_mormo) (Emesis_6837_46_______________________________ Esthemopsis_7813_32)) Theope_7789_65__________________ANTS__O_o__o) ((((((Eurybia_9363_20_________________ANTS__O_o__o (Juditha_9315_26__________________ANTS__O_o__o T_Miletus_ancon)) Argyrogrammana_6709_42) ((Lemonias_3950_55_________________ANTS__O_o__o Anteros_7097_50______________________________) (Crocozona_3995_62 Leucochimona_9306_27_______________________X))) Mesosemia_6516_85_________________________X) (((Menander_7339_49_________________ANTS__O_o__o Thysanota_5241_81_________________ANTS__O_o__o) ((Brachyglemis_3956_58 Monethe_3977_61) Exoplisia_6433_40)) Styx_infernalis)) Roeberella_7457_36))) Mesosemia_8822_02_________________________X) Panara_3932_56) (Ancyluris_6834_45 Euselasia_orfita____________________________X)) Riodina_lysippus)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((Uraneis_hyalina____________________ANTS__O_o__o Mesophthalma_6327_88_____________________X) Panara_3932_56) (((((Juditha_9315_26__________________ANTS__O_o__o Metacharis_9347_23__________________________) (((Thisbe_6766_43___________________ANTS__O_o__o ((Thysanota_5241_81_________________ANTS__O_o__o (Ithomiola_5924_82 Mesosemia_6516_85_________________________X)) (Methone_8523_95 (Napaea_m17_89_______________________________ Stalachtis_6422_39)))) Charis_9365_19_______________________________) Synargis_7605_29_________________ANTS__O_o__o)) Euselasia_8816_01_________________________X) Ancyluris_6834_45) Cremna_5260_53)) ((Chamaelimnas_3942_57 Melanis_7612_30_________________________X______) T_Miletus_ancon)) ((((Calospila_8532_96_________________ANTS__O_o__o Euselasia_orfita____________________________X) Lepricornis_5041_80) (Echenais_7168_48 Parcella_4012_64)) Leucochimona_9306_27_______________________X)) (((((((((((((((Adelotypa_7807_37_________________ANTS__O_o__o Riodina_lysippus) Eurybia_9363_20_________________ANTS__O_o__o) (((((((((Protonymphidia_3948_59_________ANTS__O_o__o (Calephelis_9356_22____________________X______ Roeberella_7457_36)) (Brachyglemis_3956_58 Esthemopsis_7813_32)) Leucochimona_7617_87_______________________X) ((((Alesa_8380_84____________________ANTS__O_o__o Exoplisia_6433_40) Necryia_5966_93) (Cyrenia_3992_63 Mesene_7450_28_________________________X______)) Themone_7643_34)) Menander_7339_49_________________ANTS__O_o__o) (((Calydna_6346_83 Ithomeis_7773_35) Perophthalma_9330_24_______________________X) Zelotea_4057_79)) Setabis_6330_38_________________ANTS__O_o__o) Ourocnemis_4621_90___________________________) (Theope_7789_65__________________ANTS__O_o__o Monethe_3977_61))) Cariomothis_6518_41) ((Theope_9026_5____________________ANTS__O_o__o Sarota_9358_21) (((Argyrogrammana_6709_42 Detritivora_9327_25) Rhetus_3973_60) Semomesia_5223_52_________________________X))) (Anteros_7097_50______________________________ T_Poritia_erycinoides)) Symmachia_8849_08___________________________) Phaenochitonia_6926_47) Mesosemia_8822_02_________________________X) Caria_7625_31_______________________________) Styx_infernalis) Hamearis_lucina) (Lemonias_3950_55_________________ANTS__O_o__o ((Amarynthis_5220_51 Emesis_mandana___________________________X_____) Helicopis_5318_54))) Apodemia_mormo) Hyphilaria_5239_91)) Leucochimona_9302_33_______________________X) Crocozona_3995_62) Pachythone_9050_18) Emesis_6837_46_______________________________) Isapis_6815_44) Pandemos_8572_97)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((Uraneis_hyalina____________________ANTS__O_o__o (Adelotypa_7807_37_________________ANTS__O_o__o Echenais_7168_48)) (((((Theope_7789_65__________________ANTS__O_o__o Emesis_mandana___________________________X_____) Perophthalma_9330_24_______________________X) Symmachia_8849_08___________________________) (Euselasia_8816_01_________________________X Anteros_7097_50______________________________)) (((((((((Calospila_8532_96_________________ANTS__O_o__o Calephelis_9356_22____________________X______) Charis_9365_19_______________________________) (Synargis_7605_29_________________ANTS__O_o__o Semomesia_5223_52_________________________X)) ((Ancyluris_6834_45 ((Argyrogrammana_6709_42 Sarota_9358_21) Necryia_5966_93)) Pandemos_8572_97)) Roeberella_7457_36) Leucochimona_9306_27_______________________X) T_Miletus_ancon) (((Thisbe_6766_43___________________ANTS__O_o__o ((Exoplisia_6433_40 Apodemia_mormo) Hamearis_lucina)) Leucochimona_7617_87_______________________X) Brachyglemis_3956_58)) Ithomeis_7773_35))) (Euselasia_orfita____________________________X Styx_infernalis)) Thysanota_5241_81_________________ANTS__O_o__o) Panara_3932_56) ((Calydna_6346_83 Emesis_6837_46_______________________________) Mesosemia_6516_85_________________________X)) (((((Protonymphidia_3948_59_________ANTS__O_o__o Mesophthalma_6327_88_____________________X) Stalachtis_6422_39) Theope_9026_5____________________ANTS__O_o__o) T_Poritia_erycinoides) Mesosemia_8822_02_________________________X)) Cyrenia_3992_63) (((Alesa_8380_84____________________ANTS__O_o__o Zelotea_4057_79) (Lemonias_3950_55_________________ANTS__O_o__o Melanis_7612_30_________________________X______)) ((((Eurybia_9363_20_________________ANTS__O_o__o (Mesene_7450_28_________________________X______ Rhetus_3973_60)) Riodina_lysippus) (Lepricornis_5041_80 Parcella_4012_64)) Cremna_5260_53))) Detritivora_9327_25) Setabis_6330_38_________________ANTS__O_o__o) (((Menander_7339_49_________________ANTS__O_o__o Themone_7643_34) Methone_8523_95) (((((Juditha_9315_26__________________ANTS__O_o__o ((((Caria_7625_31_______________________________ Pachythone_9050_18) Metacharis_9347_23__________________________) (Ithomiola_5924_82 Monethe_3977_61)) Ourocnemis_4621_90___________________________)) Leucochimona_9302_33_______________________X) (Chamaelimnas_3942_57 Phaenochitonia_6926_47)) Cariomothis_6518_41) Crocozona_3995_62))) ((((Esthemopsis_7813_32 Napaea_m17_89_______________________________) Hyphilaria_5239_91) Isapis_6815_44) Helicopis_5318_54)) Amarynthis_5220_51)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o (((((((((Menander_7339_49_________________ANTS__O_o__o Calospila_8532_96_________________ANTS__O_o__o) Lepricornis_5041_80) Echenais_7168_48) Esthemopsis_7813_32) Mesene_7450_28_________________________X______) Lemonias_3950_55_________________ANTS__O_o__o) Phaenochitonia_6926_47) Mesophthalma_6327_88_____________________X) Ithomeis_7773_35)) (Adelotypa_7807_37_________________ANTS__O_o__o Euselasia_orfita____________________________X)) Hamearis_lucina) ((Necryia_5966_93 Roeberella_7457_36) T_Miletus_ancon)) ((((Eurybia_9363_20_________________ANTS__O_o__o Cyrenia_3992_63) Mesosemia_8822_02_________________________X) Methone_8523_95) Amarynthis_5220_51)) Monethe_3977_61) Juditha_9315_26__________________ANTS__O_o__o) Theope_9026_5____________________ANTS__O_o__o) ((((Theope_7789_65__________________ANTS__O_o__o Themone_7643_34) (Synargis_7605_29_________________ANTS__O_o__o Cremna_5260_53)) T_Poritia_erycinoides) (Napaea_m17_89_______________________________ Emesis_mandana___________________________X_____))) (Alesa_8380_84____________________ANTS__O_o__o (((((((Setabis_6330_38_________________ANTS__O_o__o Caria_7625_31_______________________________) ((Argyrogrammana_6709_42 Ithomiola_5924_82) Ourocnemis_4621_90___________________________)) Euselasia_8816_01_________________________X) Emesis_6837_46_______________________________) Parcella_4012_64) Crocozona_3995_62) Leucochimona_9302_33_______________________X))) Pachythone_9050_18) (((Chamaelimnas_3942_57 Anteros_7097_50______________________________) Apodemia_mormo) Leucochimona_9306_27_______________________X)) Calephelis_9356_22____________________X______) ((Ancyluris_6834_45 Melanis_7612_30_________________________X______) Leucochimona_7617_87_______________________X)) ((Brachyglemis_3956_58 Symmachia_8849_08___________________________) (Helicopis_5318_54 Stalachtis_6422_39))) Protonymphidia_3948_59_________ANTS__O_o__o) Thisbe_6766_43___________________ANTS__O_o__o) (Detritivora_9327_25 Hyphilaria_5239_91)) Exoplisia_6433_40) (Isapis_6815_44 Metacharis_9347_23__________________________)) Panara_3932_56) Riodina_lysippus) Sarota_9358_21) Styx_infernalis) Rhetus_3973_60) Pandemos_8572_97) ((Calydna_6346_83 ((Cariomothis_6518_41 Semomesia_5223_52_________________________X) Zelotea_4057_79)) (Charis_9365_19_______________________________ Mesosemia_6516_85_________________________X))) (Thysanota_5241_81_________________ANTS__O_o__o Perophthalma_9330_24_______________________X))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((Uraneis_hyalina____________________ANTS__O_o__o Leucochimona_9302_33_______________________X) (((((((((((((((((Adelotypa_7807_37_________________ANTS__O_o__o Cyrenia_3992_63) (Eurybia_9363_20_________________ANTS__O_o__o Mesene_7450_28_________________________X______)) (Emesis_6837_46_______________________________ Mesosemia_8822_02_________________________X)) Detritivora_9327_25) Euselasia_orfita____________________________X) Brachyglemis_3956_58) ((Alesa_8380_84____________________ANTS__O_o__o (Exoplisia_6433_40 Mesophthalma_6327_88_____________________X)) Setabis_6330_38_________________ANTS__O_o__o)) ((Echenais_7168_48 Ithomiola_5924_82) Rhetus_3973_60)) (((((((Protonymphidia_3948_59_________ANTS__O_o__o (Melanis_7612_30_________________________X______ T_Poritia_erycinoides)) (Theope_9026_5____________________ANTS__O_o__o Phaenochitonia_6926_47)) Lemonias_3950_55_________________ANTS__O_o__o) Sarota_9358_21) ((Thysanota_5241_81_________________ANTS__O_o__o ((((((((((Calephelis_9356_22____________________X______ (Helicopis_5318_54 Perophthalma_9330_24_______________________X)) Styx_infernalis) Metacharis_9347_23__________________________) Apodemia_mormo) (Euselasia_8816_01_________________________X Semomesia_5223_52_________________________X)) (Chamaelimnas_3942_57 Cremna_5260_53)) Riodina_lysippus) Caria_7625_31_______________________________) (((Charis_9365_19_______________________________ Pachythone_9050_18) Ithomeis_7773_35) Monethe_3977_61)) (Leucochimona_9306_27_______________________X Symmachia_8849_08___________________________))) Stalachtis_6422_39)) (Thisbe_6766_43___________________ANTS__O_o__o ((Calydna_6346_83 Anteros_7097_50______________________________) (Zelotea_4057_79 T_Miletus_ancon)))) Napaea_m17_89_______________________________)) Methone_8523_95) Menander_7339_49_________________ANTS__O_o__o) Juditha_9315_26__________________ANTS__O_o__o) ((Ancyluris_6834_45 Emesis_mandana___________________________X_____) ((Cariomothis_6518_41 Esthemopsis_7813_32) Pandemos_8572_97))) Roeberella_7457_36) Amarynthis_5220_51) Ourocnemis_4621_90___________________________) Argyrogrammana_6709_42)) Hamearis_lucina) Theope_7789_65__________________ANTS__O_o__o) ((Isapis_6815_44 (Leucochimona_7617_87_______________________X (Necryia_5966_93 Panara_3932_56))) Themone_7643_34)) ((Calospila_8532_96_________________ANTS__O_o__o (Lepricornis_5041_80 Mesosemia_6516_85_________________________X)) ((Crocozona_3995_62 Hyphilaria_5239_91) Parcella_4012_64))) Synargis_7605_29_________________ANTS__O_o__o)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((Uraneis_hyalina____________________ANTS__O_o__o (((((((((((Adelotypa_7807_37_________________ANTS__O_o__o Leucochimona_9306_27_______________________X) ((((((Protonymphidia_3948_59_________ANTS__O_o__o ((((Theope_9026_5____________________ANTS__O_o__o ((Menander_7339_49_________________ANTS__O_o__o ((((Juditha_9315_26__________________ANTS__O_o__o (Ourocnemis_4621_90___________________________ Phaenochitonia_6926_47)) Ancyluris_6834_45) Synargis_7605_29_________________ANTS__O_o__o) (((Amarynthis_5220_51 Echenais_7168_48) Euselasia_orfita____________________________X) Symmachia_8849_08___________________________))) Charis_9365_19_______________________________)) ((Caria_7625_31_______________________________ ((Mesophthalma_6327_88_____________________X Styx_infernalis) Emesis_mandana___________________________X_____)) T_Poritia_erycinoides)) Isapis_6815_44) Hamearis_lucina)) Pandemos_8572_97) (((Eurybia_9363_20_________________ANTS__O_o__o ((Melanis_7612_30_________________________X______ Apodemia_mormo) Perophthalma_9330_24_______________________X)) Brachyglemis_3956_58) Necryia_5966_93)) Semomesia_5223_52_________________________X) ((Calospila_8532_96_________________ANTS__O_o__o Monethe_3977_61) Crocozona_3995_62)) (((((Thisbe_6766_43___________________ANTS__O_o__o Mesosemia_8822_02_________________________X) (Calephelis_9356_22____________________X______ T_Miletus_ancon)) Hyphilaria_5239_91) Helicopis_5318_54) Exoplisia_6433_40))) Mesene_7450_28_________________________X______) Ithomeis_7773_35) (Parcella_4012_64 Zelotea_4057_79)) ((Cyrenia_3992_63 (Metacharis_9347_23__________________________ Roeberella_7457_36)) (((Euselasia_8816_01_________________________X Leucochimona_7617_87_______________________X) Riodina_lysippus) Mesosemia_6516_85_________________________X))) (Theope_7789_65__________________ANTS__O_o__o Panara_3932_56)) Lepricornis_5041_80) Setabis_6330_38_________________ANTS__O_o__o) (Thysanota_5241_81_________________ANTS__O_o__o Leucochimona_9302_33_______________________X)) Rhetus_3973_60)) (Emesis_6837_46_______________________________ Pachythone_9050_18)) (Lemonias_3950_55_________________ANTS__O_o__o Sarota_9358_21)) (Cremna_5260_53 Napaea_m17_89_______________________________)) (Chamaelimnas_3942_57 Stalachtis_6422_39)) (Calydna_6346_83 Esthemopsis_7813_32)) (Argyrogrammana_6709_42 Themone_7643_34)) (Alesa_8380_84____________________ANTS__O_o__o ((Detritivora_9327_25 Methone_8523_95) Anteros_7097_50______________________________))) Ithomiola_5924_82) Cariomothis_6518_41)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((Uraneis_hyalina____________________ANTS__O_o__o Stalachtis_6422_39) ((((Alesa_8380_84____________________ANTS__O_o__o (((Thisbe_6766_43___________________ANTS__O_o__o Lepricornis_5041_80) Apodemia_mormo) Necryia_5966_93)) (Charis_9365_19_______________________________ (Metacharis_9347_23__________________________ Monethe_3977_61))) Roeberella_7457_36) Leucochimona_7617_87_______________________X)) ((Setabis_6330_38_________________ANTS__O_o__o (Cariomothis_6518_41 (((Isapis_6815_44 Zelotea_4057_79) Ourocnemis_4621_90___________________________) (Leucochimona_9302_33_______________________X Mesosemia_6516_85_________________________X)))) Mesosemia_8822_02_________________________X)) ((Detritivora_9327_25 Echenais_7168_48) (Sarota_9358_21 T_Poritia_erycinoides))) Amarynthis_5220_51) Caria_7625_31_______________________________) Hyphilaria_5239_91) Cyrenia_3992_63) (Theope_7789_65__________________ANTS__O_o__o Euselasia_orfita____________________________X)) (Adelotypa_7807_37_________________ANTS__O_o__o Methone_8523_95)) Melanis_7612_30_________________________X______) ((((((((((((Protonymphidia_3948_59_________ANTS__O_o__o ((((Lemonias_3950_55_________________ANTS__O_o__o (Cremna_5260_53 Hamearis_lucina)) Symmachia_8849_08___________________________) Pachythone_9050_18) Themone_7643_34)) ((Menander_7339_49_________________ANTS__O_o__o (Synargis_7605_29_________________ANTS__O_o__o (Argyrogrammana_6709_42 Chamaelimnas_3942_57))) (Mesene_7450_28_________________________X______ (Parcella_4012_64 Phaenochitonia_6926_47)))) Calephelis_9356_22____________________X______) (Napaea_m17_89_______________________________ ((Semomesia_5223_52_________________________X Riodina_lysippus) (Anteros_7097_50______________________________ Styx_infernalis)))) (Calospila_8532_96_________________ANTS__O_o__o Thysanota_5241_81_________________ANTS__O_o__o)) (Eurybia_9363_20_________________ANTS__O_o__o (((((Juditha_9315_26__________________ANTS__O_o__o ((Brachyglemis_3956_58 Emesis_6837_46_______________________________) Panara_3932_56)) Ithomeis_7773_35) Perophthalma_9330_24_______________________X) (Esthemopsis_7813_32 ((Euselasia_8816_01_________________________X T_Miletus_ancon) Emesis_mandana___________________________X_____))) (Ithomiola_5924_82 Mesophthalma_6327_88_____________________X)))) Helicopis_5318_54) Theope_9026_5____________________ANTS__O_o__o) (Ancyluris_6834_45 (Calydna_6346_83 Leucochimona_9306_27_______________________X))) (Exoplisia_6433_40 Pandemos_8572_97)) Rhetus_3973_60) Crocozona_3995_62))*
(T_Curetis_bulis_______________________ANTS__O_o__o Uraneis_hyalina____________________ANTS__O_o__o ((((((((((((((((((((((Adelotypa_7807_37_________________ANTS__O_o__o Thisbe_6766_43___________________ANTS__O_o__o) (((((Menander_7339_49_________________ANTS__O_o__o Calydna_6346_83) (Napaea_m17_89_______________________________ Panara_3932_56)) ((Helicopis_5318_54 Anteros_7097_50______________________________) (Ithomiola_5924_82 Melanis_7612_30_________________________X______))) Esthemopsis_7813_32) Monethe_3977_61)) Metacharis_9347_23__________________________) ((((Brachyglemis_3956_58 Leucochimona_9302_33_______________________X) Styx_infernalis) Crocozona_3995_62) ((Mesene_7450_28_________________________X______ (Necryia_5966_93 Emesis_mandana___________________________X_____)) Rhetus_3973_60))) ((((((Protonymphidia_3948_59_________ANTS__O_o__o (Alesa_8380_84____________________ANTS__O_o__o (Zelotea_4057_79 T_Poritia_erycinoides))) ((Caria_7625_31_______________________________ Methone_8523_95) Pandemos_8572_97)) Ourocnemis_4621_90___________________________) (Amarynthis_5220_51 Semomesia_5223_52_________________________X)) Mesophthalma_6327_88_____________________X) Setabis_6330_38_________________ANTS__O_o__o)) Hyphilaria_5239_91) Roeberella_7457_36) (Euselasia_8816_01_________________________X (Sarota_9358_21 Apodemia_mormo))) Leucochimona_9306_27_______________________X) (((((((Eurybia_9363_20_________________ANTS__O_o__o Argyrogrammana_6709_42) ((Lemonias_3950_55_________________ANTS__O_o__o Parcella_4012_64) Chamaelimnas_3942_57)) Cremna_5260_53) (Theope_7789_65__________________ANTS__O_o__o (Cyrenia_3992_63 Stalachtis_6422_39))) ((Detritivora_9327_25 (Lepricornis_5041_80 Phaenochitonia_6926_47)) Emesis_6837_46_______________________________)) Symmachia_8849_08___________________________) ((Theope_9026_5____________________ANTS__O_o__o (Leucochimona_7617_87_______________________X Perophthalma_9330_24_______________________X)) Synargis_7605_29_________________ANTS__O_o__o))) Exoplisia_6433_40) Juditha_9315_26__________________ANTS__O_o__o) Isapis_6815_44) ((Calospila_8532_96_________________ANTS__O_o__o Euselasia_orfita____________________________X) Riodina_lysippus)) Thysanota_5241_81_________________ANTS__O_o__o) Charis_9365_19_______________________________) Calephelis_9356_22____________________X______) T_Miletus_ancon) (((Ancyluris_6834_45 Hamearis_lucina) (Echenais_7168_48 Pachythone_9050_18)) Mesosemia_8822_02_________________________X)) (Cariomothis_6518_41 Ithomeis_7773_35)) Mesosemia_6516_85_________________________X) Themone_7643_34))*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Detritivora_9327_25) ((((((((Adelotypa_7807_37_________________ANTS__O_o__o Napaea_m17_89_______________________________) (Leucochimona_7617_87_______________________X Leucochimona_9302_33_______________________X)) (((Protonymphidia_3948_59_________ANTS__O_o__o Thisbe_6766_43___________________ANTS__O_o__o) (((((Setabis_6330_38_________________ANTS__O_o__o (Thysanota_5241_81_________________ANTS__O_o__o (Metacharis_9347_23__________________________ Rhetus_3973_60))) Mesene_7450_28_________________________X______) ((Eurybia_9363_20_________________ANTS__O_o__o Themone_7643_34) ((Crocozona_3995_62 (Lepricornis_5041_80 Roeberella_7457_36)) Hyphilaria_5239_91))) Methone_8523_95) Menander_7339_49_________________ANTS__O_o__o)) ((Ancyluris_6834_45 (Caria_7625_31_______________________________ (Helicopis_5318_54 Pachythone_9050_18))) Calydna_6346_83))) Echenais_7168_48) Monethe_3977_61) Pandemos_8572_97) Semomesia_5223_52_________________________X) Hamearis_lucina)) ((((Alesa_8380_84____________________ANTS__O_o__o ((Theope_7789_65__________________ANTS__O_o__o Chamaelimnas_3942_57) (Anteros_7097_50______________________________ Euselasia_orfita____________________________X))) Zelotea_4057_79) (Brachyglemis_3956_58 Isapis_6815_44)) Amarynthis_5220_51)) Ithomiola_5924_82) Stalachtis_6422_39) (((Juditha_9315_26__________________ANTS__O_o__o Mesosemia_6516_85_________________________X) Emesis_mandana___________________________X_____) (Cremna_5260_53 Esthemopsis_7813_32))) ((((((Calospila_8532_96_________________ANTS__O_o__o Calephelis_9356_22____________________X______) Exoplisia_6433_40) (Panara_3932_56 Perophthalma_9330_24_______________________X)) Ourocnemis_4621_90___________________________) Parcella_4012_64) T_Miletus_ancon)) Melanis_7612_30_________________________X______) Styx_infernalis) ((((Emesis_6837_46_______________________________ Phaenochitonia_6926_47) Sarota_9358_21) T_Poritia_erycinoides) Necryia_5966_93)) Charis_9365_19_______________________________) Symmachia_8849_08___________________________) Ithomeis_7773_35) Cariomothis_6518_41) ((Synargis_7605_29_________________ANTS__O_o__o Riodina_lysippus) Cyrenia_3992_63)) (((Theope_9026_5____________________ANTS__O_o__o Euselasia_8816_01_________________________X) Apodemia_mormo) Lemonias_3950_55_________________ANTS__O_o__o)) Leucochimona_9306_27_______________________X) Mesosemia_8822_02_________________________X) Mesophthalma_6327_88_____________________X) Argyrogrammana_6709_42)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((Uraneis_hyalina____________________ANTS__O_o__o (Synargis_7605_29_________________ANTS__O_o__o Pandemos_8572_97)) (Theope_7789_65__________________ANTS__O_o__o Symmachia_8849_08___________________________)) Leucochimona_9306_27_______________________X) ((((((((((((((((((((((((((((Protonymphidia_3948_59_________ANTS__O_o__o (Ithomiola_5924_82 Leucochimona_7617_87_______________________X)) (((Perophthalma_9330_24_______________________X T_Miletus_ancon) Riodina_lysippus) Hamearis_lucina)) Lepricornis_5041_80) Calydna_6346_83) (Cyrenia_3992_63 Isapis_6815_44)) Detritivora_9327_25) Echenais_7168_48) Lemonias_3950_55_________________ANTS__O_o__o) (((Menander_7339_49_________________ANTS__O_o__o Mesophthalma_6327_88_____________________X) Mesene_7450_28_________________________X______) Esthemopsis_7813_32)) Roeberella_7457_36) (Theope_9026_5____________________ANTS__O_o__o Helicopis_5318_54)) (Parcella_4012_64 Anteros_7097_50______________________________)) Euselasia_8816_01_________________________X) Ancyluris_6834_45) (Thysanota_5241_81_________________ANTS__O_o__o Calephelis_9356_22____________________X______)) Leucochimona_9302_33_______________________X) (Ithomeis_7773_35 Monethe_3977_61)) ((Mesosemia_6516_85_________________________X Methone_8523_95) Metacharis_9347_23__________________________)) Necryia_5966_93) Caria_7625_31_______________________________) Emesis_6837_46_______________________________) Semomesia_5223_52_________________________X) Chamaelimnas_3942_57) Euselasia_orfita____________________________X) ((Cariomothis_6518_41 T_Poritia_erycinoides) Themone_7643_34)) Ourocnemis_4621_90___________________________) Exoplisia_6433_40) Juditha_9315_26__________________ANTS__O_o__o)) (Argyrogrammana_6709_42 (Stalachtis_6422_39 Emesis_mandana___________________________X_____))) (((((((((((Adelotypa_7807_37_________________ANTS__O_o__o Melanis_7612_30_________________________X______) ((((Crocozona_3995_62 Styx_infernalis) Pachythone_9050_18) Panara_3932_56) (Phaenochitonia_6926_47 Apodemia_mormo))) (Amarynthis_5220_51 Hyphilaria_5239_91)) Rhetus_3973_60) Sarota_9358_21) Thisbe_6766_43___________________ANTS__O_o__o) Eurybia_9363_20_________________ANTS__O_o__o) Charis_9365_19_______________________________) Zelotea_4057_79) ((((Alesa_8380_84____________________ANTS__O_o__o Napaea_m17_89_______________________________) Mesosemia_8822_02_________________________X) (Brachyglemis_3956_58 Cremna_5260_53)) Calospila_8532_96_________________ANTS__O_o__o)) Setabis_6330_38_________________ANTS__O_o__o))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((Uraneis_hyalina____________________ANTS__O_o__o Lemonias_3950_55_________________ANTS__O_o__o) (Stalachtis_6422_39 Anteros_7097_50______________________________)) (((Eurybia_9363_20_________________ANTS__O_o__o Brachyglemis_3956_58) Mesosemia_6516_85_________________________X) (Calospila_8532_96_________________ANTS__O_o__o ((Chamaelimnas_3942_57 ((Emesis_6837_46_______________________________ ((Mesophthalma_6327_88_____________________X Panara_3932_56) Ourocnemis_4621_90___________________________)) Napaea_m17_89_______________________________)) (Isapis_6815_44 Rhetus_3973_60))))) (Theope_7789_65__________________ANTS__O_o__o (((Juditha_9315_26__________________ANTS__O_o__o Semomesia_5223_52_________________________X) Riodina_lysippus) Euselasia_orfita____________________________X))) (Setabis_6330_38_________________ANTS__O_o__o (Exoplisia_6433_40 Mesene_7450_28_________________________X______))) ((((((((((((((((((Adelotypa_7807_37_________________ANTS__O_o__o Theope_9026_5____________________ANTS__O_o__o) (Alesa_8380_84____________________ANTS__O_o__o Leucochimona_7617_87_______________________X)) (Argyrogrammana_6709_42 ((Crocozona_3995_62 (Mesosemia_8822_02_________________________X (Necryia_5966_93 Styx_infernalis))) Lepricornis_5041_80))) Calydna_6346_83) (((Caria_7625_31_______________________________ (Echenais_7168_48 (Hyphilaria_5239_91 (Zelotea_4057_79 T_Miletus_ancon)))) Charis_9365_19_______________________________) Pandemos_8572_97)) Detritivora_9327_25) Amarynthis_5220_51) Ithomeis_7773_35) Phaenochitonia_6926_47) (((((((((((Protonymphidia_3948_59_________ANTS__O_o__o Ithomiola_5924_82) Sarota_9358_21) Leucochimona_9302_33_______________________X) (Cariomothis_6518_41 Methone_8523_95)) Leucochimona_9306_27_______________________X) Apodemia_mormo) Parcella_4012_64) Esthemopsis_7813_32) (Helicopis_5318_54 Pachythone_9050_18)) ((Menander_7339_49_________________ANTS__O_o__o Thysanota_5241_81_________________ANTS__O_o__o) Thisbe_6766_43___________________ANTS__O_o__o)) Emesis_mandana___________________________X_____)) Ancyluris_6834_45) Roeberella_7457_36) Perophthalma_9330_24_______________________X) Cyrenia_3992_63) (((Calephelis_9356_22____________________X______ Cremna_5260_53) Themone_7643_34) Metacharis_9347_23__________________________)) (Euselasia_8816_01_________________________X Symmachia_8849_08___________________________)) Melanis_7612_30_________________________X______) ((Monethe_3977_61 Hamearis_lucina) T_Poritia_erycinoides))) Synargis_7605_29_________________ANTS__O_o__o)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((Uraneis_hyalina____________________ANTS__O_o__o Panara_3932_56) Cyrenia_3992_63) (Menander_7339_49_________________ANTS__O_o__o Isapis_6815_44)) (((Calospila_8532_96_________________ANTS__O_o__o ((Detritivora_9327_25 Monethe_3977_61) Echenais_7168_48)) Synargis_7605_29_________________ANTS__O_o__o) (Ancyluris_6834_45 Ithomeis_7773_35))) Mesosemia_8822_02_________________________X) (((((((((((Adelotypa_7807_37_________________ANTS__O_o__o (Thisbe_6766_43___________________ANTS__O_o__o Necryia_5966_93)) Protonymphidia_3948_59_________ANTS__O_o__o) Metacharis_9347_23__________________________) (Melanis_7612_30_________________________X______ (Sarota_9358_21 Zelotea_4057_79))) Calephelis_9356_22____________________X______) Eurybia_9363_20_________________ANTS__O_o__o) ((((((Alesa_8380_84____________________ANTS__O_o__o (Amarynthis_5220_51 Leucochimona_7617_87_______________________X)) (Emesis_6837_46_______________________________ Anteros_7097_50______________________________)) (((Lemonias_3950_55_________________ANTS__O_o__o Cremna_5260_53) T_Miletus_ancon) Argyrogrammana_6709_42)) ((Chamaelimnas_3942_57 (Ithomiola_5924_82 Mesophthalma_6327_88_____________________X)) Hyphilaria_5239_91)) (((((((Brachyglemis_3956_58 Lepricornis_5041_80) Phaenochitonia_6926_47) (((Caria_7625_31_______________________________ Parcella_4012_64) Rhetus_3973_60) Riodina_lysippus)) ((Cariomothis_6518_41 (Methone_8523_95 Pachythone_9050_18)) Stalachtis_6422_39)) Esthemopsis_7813_32) Apodemia_mormo) Symmachia_8849_08___________________________)) Styx_infernalis)) ((((((((Setabis_6330_38_________________ANTS__O_o__o (((Thysanota_5241_81_________________ANTS__O_o__o (Leucochimona_9302_33_______________________X Themone_7643_34)) Hamearis_lucina) Helicopis_5318_54)) Exoplisia_6433_40) Pandemos_8572_97) (Mesosemia_6516_85_________________________X Emesis_mandana___________________________X_____)) ((Theope_7789_65__________________ANTS__O_o__o (Charis_9365_19_______________________________ Mesene_7450_28_________________________X______)) (Theope_9026_5____________________ANTS__O_o__o Roeberella_7457_36))) Leucochimona_9306_27_______________________X) Euselasia_8816_01_________________________X) Euselasia_orfita____________________________X)) ((Crocozona_3995_62 T_Poritia_erycinoides) Napaea_m17_89_______________________________)) Perophthalma_9330_24_______________________X) (((Juditha_9315_26__________________ANTS__O_o__o Calydna_6346_83) Ourocnemis_4621_90___________________________) Semomesia_5223_52_________________________X)))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((Uraneis_hyalina____________________ANTS__O_o__o ((((((((((((((Protonymphidia_3948_59_________ANTS__O_o__o (Thisbe_6766_43___________________ANTS__O_o__o Thysanota_5241_81_________________ANTS__O_o__o)) ((Crocozona_3995_62 Rhetus_3973_60) (Isapis_6815_44 Apodemia_mormo))) ((Detritivora_9327_25 Perophthalma_9330_24_______________________X) Leucochimona_7617_87_______________________X)) Theope_9026_5____________________ANTS__O_o__o) ((((Brachyglemis_3956_58 Symmachia_8849_08___________________________) (Caria_7625_31_______________________________ Panara_3932_56)) (Calephelis_9356_22____________________X______ (Ithomeis_7773_35 T_Poritia_erycinoides))) (Ourocnemis_4621_90___________________________ Anteros_7097_50______________________________))) ((Chamaelimnas_3942_57 Lepricornis_5041_80) Mesophthalma_6327_88_____________________X)) Charis_9365_19_______________________________) Pachythone_9050_18) Mesosemia_8822_02_________________________X) ((Theope_7789_65__________________ANTS__O_o__o Emesis_6837_46_______________________________) Roeberella_7457_36)) (Echenais_7168_48 Parcella_4012_64)) ((((((((Lemonias_3950_55_________________ANTS__O_o__o Sarota_9358_21) ((Argyrogrammana_6709_42 Riodina_lysippus) Phaenochitonia_6926_47)) (Ancyluris_6834_45 (Cremna_5260_53 Stalachtis_6422_39))) Leucochimona_9306_27_______________________X) Esthemopsis_7813_32) (Leucochimona_9302_33_______________________X Styx_infernalis)) (Napaea_m17_89_______________________________ Emesis_mandana___________________________X_____)) Amarynthis_5220_51)) (((((Eurybia_9363_20_________________ANTS__O_o__o Pandemos_8572_97) Calospila_8532_96_________________ANTS__O_o__o) Monethe_3977_61) Cariomothis_6518_41) Synargis_7605_29_________________ANTS__O_o__o)) Hamearis_lucina)) Mesene_7450_28_________________________X______) (((((Adelotypa_7807_37_________________ANTS__O_o__o (((Alesa_8380_84____________________ANTS__O_o__o Themone_7643_34) Menander_7339_49_________________ANTS__O_o__o) Euselasia_orfita____________________________X)) Euselasia_8816_01_________________________X) Helicopis_5318_54) (Juditha_9315_26__________________ANTS__O_o__o (Melanis_7612_30_________________________X______ Semomesia_5223_52_________________________X))) (Calydna_6346_83 ((Exoplisia_6433_40 Methone_8523_95) Necryia_5966_93)))) Setabis_6330_38_________________ANTS__O_o__o) Metacharis_9347_23__________________________) ((((Cyrenia_3992_63 Ithomiola_5924_82) Mesosemia_6516_85_________________________X) Hyphilaria_5239_91) T_Miletus_ancon)) Zelotea_4057_79)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o ((((((((Ancyluris_6834_45 Echenais_7168_48) Calephelis_9356_22____________________X______) Caria_7625_31_______________________________) (Ithomiola_5924_82 T_Poritia_erycinoides)) ((Emesis_6837_46_______________________________ Esthemopsis_7813_32) Euselasia_orfita____________________________X)) Sarota_9358_21) Mesosemia_8822_02_________________________X) Calydna_6346_83)) Melanis_7612_30_________________________X______) (Ithomeis_7773_35 Mesophthalma_6327_88_____________________X)) ((Protonymphidia_3948_59_________ANTS__O_o__o Rhetus_3973_60) Thysanota_5241_81_________________ANTS__O_o__o)) Cariomothis_6518_41) Roeberella_7457_36) ((Alesa_8380_84____________________ANTS__O_o__o (Eurybia_9363_20_________________ANTS__O_o__o ((Detritivora_9327_25 Napaea_m17_89_______________________________) Lepricornis_5041_80))) Symmachia_8849_08___________________________)) ((((((Lemonias_3950_55_________________ANTS__O_o__o Leucochimona_9302_33_______________________X) Cremna_5260_53) Chamaelimnas_3942_57) Apodemia_mormo) (Mesene_7450_28_________________________X______ T_Miletus_ancon)) (Amarynthis_5220_51 Helicopis_5318_54))) ((Synargis_7605_29_________________ANTS__O_o__o (Isapis_6815_44 Methone_8523_95)) Exoplisia_6433_40)) Zelotea_4057_79) ((Setabis_6330_38_________________ANTS__O_o__o (Ourocnemis_4621_90___________________________ Panara_3932_56)) Crocozona_3995_62)) ((Theope_7789_65__________________ANTS__O_o__o (Theope_9026_5____________________ANTS__O_o__o Styx_infernalis)) Thisbe_6766_43___________________ANTS__O_o__o)) ((((((Adelotypa_7807_37_________________ANTS__O_o__o (Menander_7339_49_________________ANTS__O_o__o Semomesia_5223_52_________________________X)) Argyrogrammana_6709_42) Themone_7643_34) Necryia_5966_93) Monethe_3977_61) Metacharis_9347_23__________________________)) ((((Juditha_9315_26__________________ANTS__O_o__o Phaenochitonia_6926_47) (Brachyglemis_3956_58 ((Cyrenia_3992_63 (Mesosemia_6516_85_________________________X (Parcella_4012_64 Stalachtis_6422_39))) (Pachythone_9050_18 (Anteros_7097_50______________________________ Emesis_mandana___________________________X_____))))) ((Leucochimona_9306_27_______________________X Riodina_lysippus) Pandemos_8572_97)) Hamearis_lucina)) (Calospila_8532_96_________________ANTS__O_o__o (Charis_9365_19_______________________________ Leucochimona_7617_87_______________________X))) Hyphilaria_5239_91) Perophthalma_9330_24_______________________X) Euselasia_8816_01_________________________X)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((Uraneis_hyalina____________________ANTS__O_o__o ((((Theope_9026_5____________________ANTS__O_o__o Lemonias_3950_55_________________ANTS__O_o__o) ((Calydna_6346_83 (Helicopis_5318_54 Symmachia_8849_08___________________________)) (Necryia_5966_93 Anteros_7097_50______________________________))) Perophthalma_9330_24_______________________X) (((Juditha_9315_26__________________ANTS__O_o__o (Emesis_6837_46_______________________________ (Methone_8523_95 Hamearis_lucina))) Metacharis_9347_23__________________________) (Riodina_lysippus Styx_infernalis)))) Esthemopsis_7813_32) (Alesa_8380_84____________________ANTS__O_o__o Detritivora_9327_25)) Adelotypa_7807_37_________________ANTS__O_o__o) Theope_7789_65__________________ANTS__O_o__o) Pandemos_8572_97) Echenais_7168_48) Euselasia_8816_01_________________________X) Napaea_m17_89_______________________________) Euselasia_orfita____________________________X) (((Menander_7339_49_________________ANTS__O_o__o Melanis_7612_30_________________________X______) T_Miletus_ancon) ((Calephelis_9356_22____________________X______ Cyrenia_3992_63) Lepricornis_5041_80))) (((((((Protonymphidia_3948_59_________ANTS__O_o__o (((((((((Setabis_6330_38_________________ANTS__O_o__o Mesophthalma_6327_88_____________________X) ((Eurybia_9363_20_________________ANTS__O_o__o Emesis_mandana___________________________X_____) (Leucochimona_9302_33_______________________X (Mesosemia_6516_85_________________________X Phaenochitonia_6926_47)))) Chamaelimnas_3942_57) Leucochimona_7617_87_______________________X) (((Thysanota_5241_81_________________ANTS__O_o__o Zelotea_4057_79) Monethe_3977_61) ((Exoplisia_6433_40 Isapis_6815_44) Parcella_4012_64))) Mesene_7450_28_________________________X______) (Argyrogrammana_6709_42 ((Cariomothis_6518_41 Roeberella_7457_36) Leucochimona_9306_27_______________________X))) (((((Synargis_7605_29_________________ANTS__O_o__o (((((Ancyluris_6834_45 Ourocnemis_4621_90___________________________) Brachyglemis_3956_58) Cremna_5260_53) Apodemia_mormo) Mesosemia_8822_02_________________________X)) T_Poritia_erycinoides) Rhetus_3973_60) Stalachtis_6422_39) Ithomeis_7773_35)) (((Calospila_8532_96_________________ANTS__O_o__o Amarynthis_5220_51) ((Thisbe_6766_43___________________ANTS__O_o__o Hyphilaria_5239_91) Sarota_9358_21)) Panara_3932_56))) Charis_9365_19_______________________________) Ithomiola_5924_82) Semomesia_5223_52_________________________X) (Caria_7625_31_______________________________ Themone_7643_34)) Pachythone_9050_18) Crocozona_3995_62))*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Rhetus_3973_60) (((Protonymphidia_3948_59_________ANTS__O_o__o ((Theope_9026_5____________________ANTS__O_o__o T_Poritia_erycinoides) Cremna_5260_53)) Calydna_6346_83) Setabis_6330_38_________________ANTS__O_o__o)) (Argyrogrammana_6709_42 Isapis_6815_44)) Roeberella_7457_36) Phaenochitonia_6926_47) Synargis_7605_29_________________ANTS__O_o__o) (Metacharis_9347_23__________________________ Zelotea_4057_79)) Emesis_mandana___________________________X_____) ((Charis_9365_19_______________________________ Hamearis_lucina) Mesosemia_8822_02_________________________X)) Leucochimona_9302_33_______________________X) (Thysanota_5241_81_________________ANTS__O_o__o Necryia_5966_93)) Methone_8523_95) Apodemia_mormo) (Cariomothis_6518_41 (Crocozona_3995_62 (((Ithomiola_5924_82 Mesophthalma_6327_88_____________________X) Riodina_lysippus) ((Lepricornis_5041_80 Napaea_m17_89_______________________________) Ourocnemis_4621_90___________________________))))) Emesis_6837_46_______________________________) Alesa_8380_84____________________ANTS__O_o__o) Eurybia_9363_20_________________ANTS__O_o__o) ((((Adelotypa_7807_37_________________ANTS__O_o__o Themone_7643_34) Symmachia_8849_08___________________________) (Amarynthis_5220_51 ((Hyphilaria_5239_91 Monethe_3977_61) Melanis_7612_30_________________________X______))) (Detritivora_9327_25 Euselasia_orfita____________________________X))) ((Theope_7789_65__________________ANTS__O_o__o Styx_infernalis) (Euselasia_8816_01_________________________X Sarota_9358_21))) ((((Menander_7339_49_________________ANTS__O_o__o ((Ithomeis_7773_35 Anteros_7097_50______________________________) T_Miletus_ancon)) Calospila_8532_96_________________ANTS__O_o__o) Mesene_7450_28_________________________X______) ((Lemonias_3950_55_________________ANTS__O_o__o ((Ancyluris_6834_45 Semomesia_5223_52_________________________X) ((((Helicopis_5318_54 Mesosemia_6516_85_________________________X) Leucochimona_7617_87_______________________X) Stalachtis_6422_39) Leucochimona_9306_27_______________________X))) Echenais_7168_48))) Caria_7625_31_______________________________) Calephelis_9356_22____________________X______) (Juditha_9315_26__________________ANTS__O_o__o Chamaelimnas_3942_57)) Exoplisia_6433_40) Thisbe_6766_43___________________ANTS__O_o__o) Perophthalma_9330_24_______________________X) Esthemopsis_7813_32) Pandemos_8572_97) Cyrenia_3992_63) Pachythone_9050_18) (Panara_3932_56 Parcella_4012_64)) Brachyglemis_3956_58)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((Uraneis_hyalina____________________ANTS__O_o__o Hamearis_lucina) Symmachia_8849_08___________________________) (((((((Alesa_8380_84____________________ANTS__O_o__o Pandemos_8572_97) Charis_9365_19_______________________________) Theope_9026_5____________________ANTS__O_o__o) (((Ancyluris_6834_45 Cremna_5260_53) Roeberella_7457_36) Ourocnemis_4621_90___________________________)) (Leucochimona_9302_33_______________________X Phaenochitonia_6926_47)) Argyrogrammana_6709_42) ((Theope_7789_65__________________ANTS__O_o__o (Napaea_m17_89_______________________________ Semomesia_5223_52_________________________X)) Styx_infernalis))) ((((((Adelotypa_7807_37_________________ANTS__O_o__o Euselasia_orfita____________________________X) (((((((Protonymphidia_3948_59_________ANTS__O_o__o (Lemonias_3950_55_________________ANTS__O_o__o Cyrenia_3992_63)) ((Calospila_8532_96_________________ANTS__O_o__o Echenais_7168_48) Perophthalma_9330_24_______________________X)) Exoplisia_6433_40) (Hyphilaria_5239_91 Panara_3932_56)) Riodina_lysippus) Menander_7339_49_________________ANTS__O_o__o) (Leucochimona_9306_27_______________________X (Melanis_7612_30_________________________X______ Zelotea_4057_79)))) ((((Detritivora_9327_25 Helicopis_5318_54) T_Poritia_erycinoides) ((Esthemopsis_7813_32 Themone_7643_34) Mesophthalma_6327_88_____________________X)) Ithomiola_5924_82)) (((((Setabis_6330_38_________________ANTS__O_o__o (Methone_8523_95 Emesis_mandana___________________________X_____)) Caria_7625_31_______________________________) Parcella_4012_64) ((Eurybia_9363_20_________________ANTS__O_o__o Brachyglemis_3956_58) Emesis_6837_46_______________________________)) Lepricornis_5041_80)) (((Thisbe_6766_43___________________ANTS__O_o__o (((Leucochimona_7617_87_______________________X Mesosemia_6516_85_________________________X) Metacharis_9347_23__________________________) Monethe_3977_61)) Mesosemia_8822_02_________________________X) Crocozona_3995_62)) (Chamaelimnas_3942_57 (Mesene_7450_28_________________________X______ T_Miletus_ancon)))) (Synargis_7605_29_________________ANTS__O_o__o Anteros_7097_50______________________________)) ((((Juditha_9315_26__________________ANTS__O_o__o Stalachtis_6422_39) Apodemia_mormo) Ithomeis_7773_35) ((Euselasia_8816_01_________________________X Rhetus_3973_60) (Isapis_6815_44 (Pachythone_9050_18 Sarota_9358_21))))) (Calephelis_9356_22____________________X______ Necryia_5966_93)) (((Thysanota_5241_81_________________ANTS__O_o__o Calydna_6346_83) Amarynthis_5220_51) Cariomothis_6518_41))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o ((((((Theope_9026_5____________________ANTS__O_o__o Euselasia_8816_01_________________________X) (((Lepricornis_5041_80 (Riodina_lysippus T_Miletus_ancon)) Hamearis_lucina) Pachythone_9050_18)) (((Calospila_8532_96_________________ANTS__O_o__o Ourocnemis_4621_90___________________________) Thysanota_5241_81_________________ANTS__O_o__o) ((Synargis_7605_29_________________ANTS__O_o__o Metacharis_9347_23__________________________) Caria_7625_31_______________________________))) ((((Thisbe_6766_43___________________ANTS__O_o__o Ithomeis_7773_35) Pandemos_8572_97) Themone_7643_34) Parcella_4012_64)) (Amarynthis_5220_51 Monethe_3977_61)) Roeberella_7457_36)) Apodemia_mormo) (Eurybia_9363_20_________________ANTS__O_o__o Lemonias_3950_55_________________ANTS__O_o__o)) (Adelotypa_7807_37_________________ANTS__O_o__o Hyphilaria_5239_91)) (Calydna_6346_83 Mesosemia_6516_85_________________________X)) (Isapis_6815_44 Mesophthalma_6327_88_____________________X)) ((((((Protonymphidia_3948_59_________ANTS__O_o__o ((((((((Setabis_6330_38_________________ANTS__O_o__o T_Poritia_erycinoides) Necryia_5966_93) Calephelis_9356_22____________________X______) Phaenochitonia_6926_47) (Theope_7789_65__________________ANTS__O_o__o Brachyglemis_3956_58)) (Leucochimona_9306_27_______________________X Stalachtis_6422_39)) Styx_infernalis) Exoplisia_6433_40)) (Ancyluris_6834_45 ((((Cyrenia_3992_63 Sarota_9358_21) Panara_3932_56) Semomesia_5223_52_________________________X) Mesene_7450_28_________________________X______))) Cariomothis_6518_41) Euselasia_orfita____________________________X) Napaea_m17_89_______________________________) Emesis_mandana___________________________X_____)) (Cremna_5260_53 Rhetus_3973_60)) Leucochimona_9302_33_______________________X) (Detritivora_9327_25 Leucochimona_7617_87_______________________X)) (Mesosemia_8822_02_________________________X Anteros_7097_50______________________________)) ((Alesa_8380_84____________________ANTS__O_o__o Methone_8523_95) (((((((Menander_7339_49_________________ANTS__O_o__o Crocozona_3995_62) Charis_9365_19_______________________________) Chamaelimnas_3942_57) Melanis_7612_30_________________________X______) (Echenais_7168_48 Ithomiola_5924_82)) Juditha_9315_26__________________ANTS__O_o__o) Helicopis_5318_54))) Symmachia_8849_08___________________________) ((Emesis_6837_46_______________________________ Zelotea_4057_79) Esthemopsis_7813_32)) (Argyrogrammana_6709_42 Perophthalma_9330_24_______________________X))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((Uraneis_hyalina____________________ANTS__O_o__o ((((Thysanota_5241_81_________________ANTS__O_o__o Caria_7625_31_______________________________) Calephelis_9356_22____________________X______) (Echenais_7168_48 Leucochimona_7617_87_______________________X)) Styx_infernalis)) ((Mesosemia_6516_85_________________________X Necryia_5966_93) Symmachia_8849_08___________________________)) Ithomiola_5924_82) ((Synargis_7605_29_________________ANTS__O_o__o Amarynthis_5220_51) Cyrenia_3992_63)) (((((((((Adelotypa_7807_37_________________ANTS__O_o__o ((Brachyglemis_3956_58 Euselasia_8816_01_________________________X) ((Methone_8523_95 Napaea_m17_89_______________________________) Emesis_mandana___________________________X_____))) Phaenochitonia_6926_47) (Alesa_8380_84____________________ANTS__O_o__o (((Lemonias_3950_55_________________ANTS__O_o__o (Melanis_7612_30_________________________X______ Metacharis_9347_23__________________________)) Euselasia_orfita____________________________X) Calospila_8532_96_________________ANTS__O_o__o))) (Theope_7789_65__________________ANTS__O_o__o Themone_7643_34)) Setabis_6330_38_________________ANTS__O_o__o) (Cariomothis_6518_41 Ourocnemis_4621_90___________________________)) (Eurybia_9363_20_________________ANTS__O_o__o (((Theope_9026_5____________________ANTS__O_o__o Zelotea_4057_79) Leucochimona_9302_33_______________________X) ((((((((Menander_7339_49_________________ANTS__O_o__o Pandemos_8572_97) ((((Calydna_6346_83 (Stalachtis_6422_39 T_Poritia_erycinoides)) Rhetus_3973_60) Parcella_4012_64) ((Lepricornis_5041_80 Semomesia_5223_52_________________________X) Sarota_9358_21))) (Ancyluris_6834_45 Helicopis_5318_54)) Argyrogrammana_6709_42) Mesophthalma_6327_88_____________________X) (Thisbe_6766_43___________________ANTS__O_o__o Perophthalma_9330_24_______________________X)) Juditha_9315_26__________________ANTS__O_o__o) (((Detritivora_9327_25 Hamearis_lucina) Isapis_6815_44) Apodemia_mormo))))) (((Chamaelimnas_3942_57 Cremna_5260_53) Anteros_7097_50______________________________) Monethe_3977_61)) (((((Protonymphidia_3948_59_________ANTS__O_o__o Leucochimona_9306_27_______________________X) Esthemopsis_7813_32) ((Hyphilaria_5239_91 Ithomeis_7773_35) T_Miletus_ancon)) (((Emesis_6837_46_______________________________ Roeberella_7457_36) Exoplisia_6433_40) (Panara_3932_56 Riodina_lysippus))) (((Charis_9365_19_______________________________ (Crocozona_3995_62 Pachythone_9050_18)) Mesene_7450_28_________________________X______) Mesosemia_8822_02_________________________X))))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((Uraneis_hyalina____________________ANTS__O_o__o (Caria_7625_31_______________________________ (Detritivora_9327_25 Monethe_3977_61))) ((((((Alesa_8380_84____________________ANTS__O_o__o Styx_infernalis) Mesosemia_6516_85_________________________X) Symmachia_8849_08___________________________) Ithomiola_5924_82) (Amarynthis_5220_51 (Charis_9365_19_______________________________ (Melanis_7612_30_________________________X______ (Mesene_7450_28_________________________X______ Napaea_m17_89_______________________________))))) Semomesia_5223_52_________________________X)) ((((((((Setabis_6330_38_________________ANTS__O_o__o (Juditha_9315_26__________________ANTS__O_o__o ((Exoplisia_6433_40 (Leucochimona_9302_33_______________________X (Leucochimona_9306_27_______________________X T_Poritia_erycinoides))) Parcella_4012_64))) (((Thisbe_6766_43___________________ANTS__O_o__o ((Cariomothis_6518_41 Ithomeis_7773_35) Apodemia_mormo)) Pandemos_8572_97) ((((Synargis_7605_29_________________ANTS__O_o__o Ancyluris_6834_45) Cremna_5260_53) Calephelis_9356_22____________________X______) (((Emesis_6837_46_______________________________ ((Helicopis_5318_54 Necryia_5966_93) Stalachtis_6422_39)) (Mesosemia_8822_02_________________________X (Metacharis_9347_23__________________________ Anteros_7097_50______________________________))) Euselasia_8816_01_________________________X)))) (Brachyglemis_3956_58 Methone_8523_95)) ((Theope_7789_65__________________ANTS__O_o__o (Calydna_6346_83 (Crocozona_3995_62 Ourocnemis_4621_90___________________________))) Sarota_9358_21)) (Thysanota_5241_81_________________ANTS__O_o__o Zelotea_4057_79)) Roeberella_7457_36) T_Miletus_ancon) Echenais_7168_48)) ((Protonymphidia_3948_59_________ANTS__O_o__o Perophthalma_9330_24_______________________X) (Cyrenia_3992_63 (Mesophthalma_6327_88_____________________X Panara_3932_56)))) ((Chamaelimnas_3942_57 Rhetus_3973_60) Hamearis_lucina)) Lemonias_3950_55_________________ANTS__O_o__o) (((((Adelotypa_7807_37_________________ANTS__O_o__o Phaenochitonia_6926_47) ((Theope_9026_5____________________ANTS__O_o__o Leucochimona_7617_87_______________________X) ((Lepricornis_5041_80 Themone_7643_34) Euselasia_orfita____________________________X))) Calospila_8532_96_________________ANTS__O_o__o) Argyrogrammana_6709_42) Emesis_mandana___________________________X_____)) (((Eurybia_9363_20_________________ANTS__O_o__o Isapis_6815_44) Riodina_lysippus) ((Esthemopsis_7813_32 Pachythone_9050_18) Hyphilaria_5239_91))) Menander_7339_49_________________ANTS__O_o__o)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((Uraneis_hyalina____________________ANTS__O_o__o Theope_7789_65__________________ANTS__O_o__o) (Menander_7339_49_________________ANTS__O_o__o (((Thysanota_5241_81_________________ANTS__O_o__o Styx_infernalis) Chamaelimnas_3942_57) Brachyglemis_3956_58))) Synargis_7605_29_________________ANTS__O_o__o) (Crocozona_3995_62 Cyrenia_3992_63)) ((((((((((((((((((Adelotypa_7807_37_________________ANTS__O_o__o (Echenais_7168_48 Leucochimona_9306_27_______________________X)) (Eurybia_9363_20_________________ANTS__O_o__o Hyphilaria_5239_91)) (((((((((Alesa_8380_84____________________ANTS__O_o__o Euselasia_orfita____________________________X) ((Argyrogrammana_6709_42 Mesophthalma_6327_88_____________________X) ((Helicopis_5318_54 T_Miletus_ancon) (Symmachia_8849_08___________________________ (Anteros_7097_50______________________________ Emesis_mandana___________________________X_____))))) T_Poritia_erycinoides) Emesis_6837_46_______________________________) Cariomothis_6518_41) Cremna_5260_53) Leucochimona_7617_87_______________________X) Riodina_lysippus) (Ancyluris_6834_45 Mesene_7450_28_________________________X______))) (Thisbe_6766_43___________________ANTS__O_o__o Parcella_4012_64)) Zelotea_4057_79) (((Euselasia_8816_01_________________________X Mesosemia_8822_02_________________________X) (Ithomeis_7773_35 ((Methone_8523_95 Apodemia_mormo) Stalachtis_6422_39))) Exoplisia_6433_40)) Ourocnemis_4621_90___________________________) (Lemonias_3950_55_________________ANTS__O_o__o Hamearis_lucina)) Protonymphidia_3948_59_________ANTS__O_o__o) ((Theope_9026_5____________________ANTS__O_o__o ((((Isapis_6815_44 Ithomiola_5924_82) Metacharis_9347_23__________________________) Perophthalma_9330_24_______________________X) ((Leucochimona_9302_33_______________________X Phaenochitonia_6926_47) Napaea_m17_89_______________________________))) Themone_7643_34)) Calospila_8532_96_________________ANTS__O_o__o) (Monethe_3977_61 (Necryia_5966_93 Sarota_9358_21))) ((((Setabis_6330_38_________________ANTS__O_o__o Pachythone_9050_18) Semomesia_5223_52_________________________X) ((Esthemopsis_7813_32 Mesosemia_6516_85_________________________X) Rhetus_3973_60)) Roeberella_7457_36)) Charis_9365_19_______________________________) Panara_3932_56) Calydna_6346_83) Caria_7625_31_______________________________) Juditha_9315_26__________________ANTS__O_o__o)) ((Amarynthis_5220_51 (Lepricornis_5041_80 Pandemos_8572_97)) ((Calephelis_9356_22____________________X______ Melanis_7612_30_________________________X______) Detritivora_9327_25)))*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Parcella_4012_64) ((Setabis_6330_38_________________ANTS__O_o__o (((Synargis_7605_29_________________ANTS__O_o__o Calephelis_9356_22____________________X______) (Metacharis_9347_23__________________________ Sarota_9358_21)) Argyrogrammana_6709_42)) Theope_9026_5____________________ANTS__O_o__o)) (((((Ancyluris_6834_45 Perophthalma_9330_24_______________________X) Charis_9365_19_______________________________) Leucochimona_9302_33_______________________X) Esthemopsis_7813_32) T_Poritia_erycinoides)) (((Alesa_8380_84____________________ANTS__O_o__o Cremna_5260_53) Caria_7625_31_______________________________) T_Miletus_ancon)) ((Calydna_6346_83 Stalachtis_6422_39) Pandemos_8572_97)) Cariomothis_6518_41) Hyphilaria_5239_91) ((((((Adelotypa_7807_37_________________ANTS__O_o__o (Helicopis_5318_54 Styx_infernalis)) Detritivora_9327_25) Semomesia_5223_52_________________________X) (Lemonias_3950_55_________________ANTS__O_o__o ((Calospila_8532_96_________________ANTS__O_o__o Rhetus_3973_60) Thysanota_5241_81_________________ANTS__O_o__o))) Panara_3932_56) (((((Cyrenia_3992_63 (Ithomeis_7773_35 Mesene_7450_28_________________________X______)) ((Mesosemia_6516_85_________________________X Napaea_m17_89_______________________________) Necryia_5966_93)) Mesophthalma_6327_88_____________________X) Exoplisia_6433_40) (Roeberella_7457_36 Anteros_7097_50______________________________)))) Pachythone_9050_18) Theope_7789_65__________________ANTS__O_o__o) Monethe_3977_61) Zelotea_4057_79) ((((((((((Protonymphidia_3948_59_________ANTS__O_o__o (((Ithomiola_5924_82 Riodina_lysippus) Emesis_mandana___________________________X_____) Leucochimona_7617_87_______________________X)) Euselasia_orfita____________________________X) Lepricornis_5041_80) Ourocnemis_4621_90___________________________) (Menander_7339_49_________________ANTS__O_o__o ((((((Thisbe_6766_43___________________ANTS__O_o__o (Isapis_6815_44 Themone_7643_34)) Euselasia_8816_01_________________________X) Echenais_7168_48) Mesosemia_8822_02_________________________X) Brachyglemis_3956_58) Apodemia_mormo))) Juditha_9315_26__________________ANTS__O_o__o) (Amarynthis_5220_51 Symmachia_8849_08___________________________)) ((Crocozona_3995_62 Methone_8523_95) Phaenochitonia_6926_47)) ((Eurybia_9363_20_________________ANTS__O_o__o Emesis_6837_46_______________________________) Melanis_7612_30_________________________X______)) (Chamaelimnas_3942_57 Hamearis_lucina))) Leucochimona_9306_27_______________________X)*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Setabis_6330_38_________________ANTS__O_o__o) Hyphilaria_5239_91) Sarota_9358_21) (((Cremna_5260_53 Panara_3932_56) (Stalachtis_6422_39 Themone_7643_34)) Napaea_m17_89_______________________________)) (Chamaelimnas_3942_57 Necryia_5966_93)) Ithomeis_7773_35) (((((Protonymphidia_3948_59_________ANTS__O_o__o (Alesa_8380_84____________________ANTS__O_o__o ((Lemonias_3950_55_________________ANTS__O_o__o Monethe_3977_61) Emesis_mandana___________________________X_____))) Ancyluris_6834_45) Juditha_9315_26__________________ANTS__O_o__o) Exoplisia_6433_40) (Euselasia_8816_01_________________________X Euselasia_orfita____________________________X))) ((((((((((Eurybia_9363_20_________________ANTS__O_o__o ((Calospila_8532_96_________________ANTS__O_o__o Echenais_7168_48) Lepricornis_5041_80)) (((((Thysanota_5241_81_________________ANTS__O_o__o Methone_8523_95) (Calydna_6346_83 Crocozona_3995_62)) Caria_7625_31_______________________________) Helicopis_5318_54) Metacharis_9347_23__________________________)) (Menander_7339_49_________________ANTS__O_o__o Esthemopsis_7813_32)) (Symmachia_8849_08___________________________ Riodina_lysippus)) Perophthalma_9330_24_______________________X) ((Calephelis_9356_22____________________X______ (Mesosemia_8822_02_________________________X Parcella_4012_64)) Semomesia_5223_52_________________________X)) Emesis_6837_46_______________________________) Cyrenia_3992_63) T_Poritia_erycinoides) Mesosemia_6516_85_________________________X)) Mesene_7450_28_________________________X______) ((((Adelotypa_7807_37_________________ANTS__O_o__o Phaenochitonia_6926_47) (Pandemos_8572_97 Styx_infernalis)) Zelotea_4057_79) Leucochimona_9302_33_______________________X)) (Synargis_7605_29_________________ANTS__O_o__o Detritivora_9327_25)) Apodemia_mormo) Theope_7789_65__________________ANTS__O_o__o) (Leucochimona_7617_87_______________________X Melanis_7612_30_________________________X______)) ((Ourocnemis_4621_90___________________________ T_Miletus_ancon) (Anteros_7097_50______________________________ Hamearis_lucina))) (((Amarynthis_5220_51 Rhetus_3973_60) Mesophthalma_6327_88_____________________X) Leucochimona_9306_27_______________________X)) Theope_9026_5____________________ANTS__O_o__o) (Argyrogrammana_6709_42 Roeberella_7457_36)) ((Cariomothis_6518_41 Charis_9365_19_______________________________) (Ithomiola_5924_82 Pachythone_9050_18))) (Thisbe_6766_43___________________ANTS__O_o__o Brachyglemis_3956_58)) Isapis_6815_44)*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((Uraneis_hyalina____________________ANTS__O_o__o T_Poritia_erycinoides) ((((((Setabis_6330_38_________________ANTS__O_o__o (Thysanota_5241_81_________________ANTS__O_o__o (Amarynthis_5220_51 (Calydna_6346_83 Stalachtis_6422_39)))) Charis_9365_19_______________________________) (Calephelis_9356_22____________________X______ Exoplisia_6433_40)) (((Synargis_7605_29_________________ANTS__O_o__o (Ithomiola_5924_82 (Pandemos_8572_97 Anteros_7097_50______________________________))) (Phaenochitonia_6926_47 T_Miletus_ancon)) Parcella_4012_64)) ((Eurybia_9363_20_________________ANTS__O_o__o Apodemia_mormo) Mesophthalma_6327_88_____________________X)) (Thisbe_6766_43___________________ANTS__O_o__o Argyrogrammana_6709_42))) Euselasia_8816_01_________________________X) Hamearis_lucina) Roeberella_7457_36) Ancyluris_6834_45) ((Juditha_9315_26__________________ANTS__O_o__o ((Mesene_7450_28_________________________X______ ((Mesosemia_8822_02_________________________X Pachythone_9050_18) Metacharis_9347_23__________________________)) Rhetus_3973_60)) (((Napaea_m17_89_______________________________ Riodina_lysippus) Euselasia_orfita____________________________X) Semomesia_5223_52_________________________X))) ((Cyrenia_3992_63 Emesis_mandana___________________________X_____) Leucochimona_9302_33_______________________X)) ((((((((((Adelotypa_7807_37_________________ANTS__O_o__o Styx_infernalis) ((((((((Protonymphidia_3948_59_________ANTS__O_o__o (Emesis_6837_46_______________________________ Hyphilaria_5239_91)) (((Chamaelimnas_3942_57 Cremna_5260_53) Ithomeis_7773_35) Monethe_3977_61)) Symmachia_8849_08___________________________) (Leucochimona_7617_87_______________________X Melanis_7612_30_________________________X______)) Detritivora_9327_25) (Alesa_8380_84____________________ANTS__O_o__o Echenais_7168_48)) Esthemopsis_7813_32) (((Lemonias_3950_55_________________ANTS__O_o__o Leucochimona_9306_27_______________________X) Ourocnemis_4621_90___________________________) Methone_8523_95))) ((Helicopis_5318_54 Mesosemia_6516_85_________________________X) Themone_7643_34)) Calospila_8532_96_________________ANTS__O_o__o) Theope_7789_65__________________ANTS__O_o__o) Menander_7339_49_________________ANTS__O_o__o) Caria_7625_31_______________________________) Crocozona_3995_62) Panara_3932_56) (Cariomothis_6518_41 Necryia_5966_93))) (((Theope_9026_5____________________ANTS__O_o__o Brachyglemis_3956_58) Sarota_9358_21) ((Isapis_6815_44 Lepricornis_5041_80) Zelotea_4057_79))) Perophthalma_9330_24_______________________X)*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((Uraneis_hyalina____________________ANTS__O_o__o (((((((((Setabis_6330_38_________________ANTS__O_o__o Anteros_7097_50______________________________) Cremna_5260_53) T_Poritia_erycinoides) Euselasia_orfita____________________________X) ((Thisbe_6766_43___________________ANTS__O_o__o Argyrogrammana_6709_42) (Leucochimona_9306_27_______________________X Phaenochitonia_6926_47))) Echenais_7168_48) Stalachtis_6422_39) (Charis_9365_19_______________________________ Mesene_7450_28_________________________X______)) (((((Eurybia_9363_20_________________ANTS__O_o__o Rhetus_3973_60) (Leucochimona_9302_33_______________________X (Mesosemia_6516_85_________________________X Perophthalma_9330_24_______________________X))) (Calephelis_9356_22____________________X______ Metacharis_9347_23__________________________)) Pandemos_8572_97) Ithomeis_7773_35))) (Isapis_6815_44 Roeberella_7457_36)) ((Protonymphidia_3948_59_________ANTS__O_o__o Calydna_6346_83) Chamaelimnas_3942_57)) Adelotypa_7807_37_________________ANTS__O_o__o) Lepricornis_5041_80) (((((Alesa_8380_84____________________ANTS__O_o__o Sarota_9358_21) Hyphilaria_5239_91) ((Theope_9026_5____________________ANTS__O_o__o (Monethe_3977_61 Styx_infernalis)) Panara_3932_56)) Emesis_mandana___________________________X_____) Exoplisia_6433_40)) (((Lemonias_3950_55_________________ANTS__O_o__o Necryia_5966_93) Crocozona_3995_62) ((Ancyluris_6834_45 ((Brachyglemis_3956_58 (Caria_7625_31_______________________________ (Cyrenia_3992_63 ((Esthemopsis_7813_32 Ithomiola_5924_82) Ourocnemis_4621_90___________________________)))) ((Mesophthalma_6327_88_____________________X Riodina_lysippus) Themone_7643_34))) Symmachia_8849_08___________________________))) Calospila_8532_96_________________ANTS__O_o__o) Theope_7789_65__________________ANTS__O_o__o) (((Menander_7339_49_________________ANTS__O_o__o Synargis_7605_29_________________ANTS__O_o__o) (((((Cariomothis_6518_41 (Detritivora_9327_25 Mesosemia_8822_02_________________________X)) (Emesis_6837_46_______________________________ Melanis_7612_30_________________________X______)) T_Miletus_ancon) Methone_8523_95) ((Euselasia_8816_01_________________________X Apodemia_mormo) Parcella_4012_64))) Hamearis_lucina)) ((((Juditha_9315_26__________________ANTS__O_o__o Pachythone_9050_18) (Amarynthis_5220_51 (Semomesia_5223_52_________________________X Zelotea_4057_79))) (Thysanota_5241_81_________________ANTS__O_o__o (Leucochimona_7617_87_______________________X Napaea_m17_89_______________________________))) Helicopis_5318_54))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((Uraneis_hyalina____________________ANTS__O_o__o (Thysanota_5241_81_________________ANTS__O_o__o (Euselasia_8816_01_________________________X Helicopis_5318_54))) Setabis_6330_38_________________ANTS__O_o__o) (((((((((((((((Alesa_8380_84____________________ANTS__O_o__o (Lemonias_3950_55_________________ANTS__O_o__o Charis_9365_19_______________________________)) Monethe_3977_61) Semomesia_5223_52_________________________X) Necryia_5966_93) (((Thisbe_6766_43___________________ANTS__O_o__o Calydna_6346_83) Caria_7625_31_______________________________) ((((((((Ancyluris_6834_45 Themone_7643_34) Mesosemia_8822_02_________________________X) Parcella_4012_64) Calephelis_9356_22____________________X______) ((Brachyglemis_3956_58 Stalachtis_6422_39) Lepricornis_5041_80)) Emesis_6837_46_______________________________) Mesophthalma_6327_88_____________________X) Euselasia_orfita____________________________X))) Theope_9026_5____________________ANTS__O_o__o) (((((Cariomothis_6518_41 Leucochimona_9302_33_______________________X) Ithomiola_5924_82) Pandemos_8572_97) (Rhetus_3973_60 T_Poritia_erycinoides)) Esthemopsis_7813_32)) (Melanis_7612_30_________________________X______ Mesosemia_6516_85_________________________X)) Leucochimona_7617_87_______________________X) T_Miletus_ancon) Hamearis_lucina) Cyrenia_3992_63) Echenais_7168_48) (Detritivora_9327_25 Panara_3932_56)) Exoplisia_6433_40)) Argyrogrammana_6709_42) (((((Adelotypa_7807_37_________________ANTS__O_o__o Apodemia_mormo) (((((Eurybia_9363_20_________________ANTS__O_o__o Theope_7789_65__________________ANTS__O_o__o) Mesene_7450_28_________________________X______) Leucochimona_9306_27_______________________X) Methone_8523_95) Zelotea_4057_79)) ((Protonymphidia_3948_59_________ANTS__O_o__o Roeberella_7457_36) Anteros_7097_50______________________________)) (Menander_7339_49_________________ANTS__O_o__o Amarynthis_5220_51)) ((((((Juditha_9315_26__________________ANTS__O_o__o Ourocnemis_4621_90___________________________) Sarota_9358_21) Napaea_m17_89_______________________________) Perophthalma_9330_24_______________________X) Hyphilaria_5239_91) Emesis_mandana___________________________X_____))) Riodina_lysippus) Ithomeis_7773_35) (Chamaelimnas_3942_57 (Crocozona_3995_62 Phaenochitonia_6926_47))) Symmachia_8849_08___________________________) (((Synargis_7605_29_________________ANTS__O_o__o Pachythone_9050_18) Metacharis_9347_23__________________________) (Cremna_5260_53 (Isapis_6815_44 Styx_infernalis)))) Calospila_8532_96_________________ANTS__O_o__o)*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Synargis_7605_29_________________ANTS__O_o__o) (Hyphilaria_5239_91 (Leucochimona_7617_87_______________________X Leucochimona_9302_33_______________________X))) (Argyrogrammana_6709_42 (Cariomothis_6518_41 Isapis_6815_44))) ((Caria_7625_31_______________________________ (((Exoplisia_6433_40 Metacharis_9347_23__________________________) Leucochimona_9306_27_______________________X) Anteros_7097_50______________________________)) Symmachia_8849_08___________________________)) (Protonymphidia_3948_59_________ANTS__O_o__o T_Poritia_erycinoides)) Calydna_6346_83) Themone_7643_34) (Crocozona_3995_62 Esthemopsis_7813_32)) Brachyglemis_3956_58) Hamearis_lucina) (Ithomiola_5924_82 (Methone_8523_95 Riodina_lysippus))) ((Eurybia_9363_20_________________ANTS__O_o__o ((((Charis_9365_19_______________________________ Pandemos_8572_97) Stalachtis_6422_39) Detritivora_9327_25) Zelotea_4057_79)) Roeberella_7457_36)) Ancyluris_6834_45) Apodemia_mormo) Helicopis_5318_54) ((Theope_9026_5____________________ANTS__O_o__o Panara_3932_56) Phaenochitonia_6926_47)) Juditha_9315_26__________________ANTS__O_o__o) ((Thisbe_6766_43___________________ANTS__O_o__o (Calephelis_9356_22____________________X______ (Echenais_7168_48 Necryia_5966_93))) Mesene_7450_28_________________________X______)) Monethe_3977_61) (Menander_7339_49_________________ANTS__O_o__o Emesis_mandana___________________________X_____)) T_Miletus_ancon) Napaea_m17_89_______________________________) Mesophthalma_6327_88_____________________X) (((((((Adelotypa_7807_37_________________ANTS__O_o__o (((((Lemonias_3950_55_________________ANTS__O_o__o Amarynthis_5220_51) Rhetus_3973_60) (((Chamaelimnas_3942_57 ((Cremna_5260_53 Ourocnemis_4621_90___________________________) Melanis_7612_30_________________________X______)) Mesosemia_6516_85_________________________X) (Mesosemia_8822_02_________________________X Pachythone_9050_18))) Styx_infernalis) (Thysanota_5241_81_________________ANTS__O_o__o (Emesis_6837_46_______________________________ Euselasia_orfita____________________________X)))) Euselasia_8816_01_________________________X) ((Theope_7789_65__________________ANTS__O_o__o Lepricornis_5041_80) (Calospila_8532_96_________________ANTS__O_o__o Cyrenia_3992_63))) Alesa_8380_84____________________ANTS__O_o__o) Perophthalma_9330_24_______________________X) Sarota_9358_21) Parcella_4012_64)) Setabis_6330_38_________________ANTS__O_o__o) Ithomeis_7773_35) Semomesia_5223_52_________________________X)*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Amarynthis_5220_51) Theope_7789_65__________________ANTS__O_o__o) Thysanota_5241_81_________________ANTS__O_o__o) Mesophthalma_6327_88_____________________X) Euselasia_8816_01_________________________X) ((Protonymphidia_3948_59_________ANTS__O_o__o (Perophthalma_9330_24_______________________X Emesis_mandana___________________________X_____)) Cyrenia_3992_63)) Crocozona_3995_62) (((Adelotypa_7807_37_________________ANTS__O_o__o T_Miletus_ancon) (Ithomeis_7773_35 Metacharis_9347_23__________________________)) Menander_7339_49_________________ANTS__O_o__o)) Calephelis_9356_22____________________X______) Isapis_6815_44) Mesosemia_6516_85_________________________X) (Chamaelimnas_3942_57 Cremna_5260_53)) ((Eurybia_9363_20_________________ANTS__O_o__o Synargis_7605_29_________________ANTS__O_o__o) Pachythone_9050_18)) ((Alesa_8380_84____________________ANTS__O_o__o Theope_9026_5____________________ANTS__O_o__o) (Argyrogrammana_6709_42 Styx_infernalis))) (Rhetus_3973_60 Hamearis_lucina)) ((((Setabis_6330_38_________________ANTS__O_o__o Melanis_7612_30_________________________X______) Stalachtis_6422_39) (((((((Calospila_8532_96_________________ANTS__O_o__o (Calydna_6346_83 Phaenochitonia_6926_47)) ((Charis_9365_19_______________________________ Methone_8523_95) (Panara_3932_56 Apodemia_mormo))) (Caria_7625_31_______________________________ (((Lepricornis_5041_80 Necryia_5966_93) Napaea_m17_89_______________________________) Themone_7643_34))) Anteros_7097_50______________________________) Monethe_3977_61) Esthemopsis_7813_32) Sarota_9358_21)) Zelotea_4057_79)) Pandemos_8572_97) (((((Cariomothis_6518_41 Roeberella_7457_36) Leucochimona_7617_87_______________________X) (Emesis_6837_46_______________________________ Symmachia_8849_08___________________________)) Semomesia_5223_52_________________________X) T_Poritia_erycinoides)) Hyphilaria_5239_91) Riodina_lysippus) ((Ancyluris_6834_45 Detritivora_9327_25) Mesene_7450_28_________________________X______)) Leucochimona_9302_33_______________________X) Mesosemia_8822_02_________________________X) (Thisbe_6766_43___________________ANTS__O_o__o Euselasia_orfita____________________________X)) ((Lemonias_3950_55_________________ANTS__O_o__o Ithomiola_5924_82) Leucochimona_9306_27_______________________X)) ((Juditha_9315_26__________________ANTS__O_o__o (Helicopis_5318_54 Ourocnemis_4621_90___________________________)) ((Echenais_7168_48 Exoplisia_6433_40) Parcella_4012_64))) Brachyglemis_3956_58)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((Uraneis_hyalina____________________ANTS__O_o__o Leucochimona_9306_27_______________________X) ((((((((Adelotypa_7807_37_________________ANTS__O_o__o Anteros_7097_50______________________________) Setabis_6330_38_________________ANTS__O_o__o) Euselasia_orfita____________________________X) Alesa_8380_84____________________ANTS__O_o__o) (((Eurybia_9363_20_________________ANTS__O_o__o Perophthalma_9330_24_______________________X) (((((((((((((Theope_7789_65__________________ANTS__O_o__o (((Thisbe_6766_43___________________ANTS__O_o__o Echenais_7168_48) (Ithomiola_5924_82 Mesosemia_8822_02_________________________X)) Thysanota_5241_81_________________ANTS__O_o__o)) Ithomeis_7773_35) Styx_infernalis) (Brachyglemis_3956_58 Lepricornis_5041_80)) ((Theope_9026_5____________________ANTS__O_o__o Ourocnemis_4621_90___________________________) Cremna_5260_53)) Helicopis_5318_54) Charis_9365_19_______________________________) Emesis_mandana___________________________X_____) Apodemia_mormo) (Synargis_7605_29_________________ANTS__O_o__o (Cyrenia_3992_63 Semomesia_5223_52_________________________X))) T_Miletus_ancon) ((Calospila_8532_96_________________ANTS__O_o__o Caria_7625_31_______________________________) (Juditha_9315_26__________________ANTS__O_o__o Hamearis_lucina))) Leucochimona_7617_87_______________________X)) Exoplisia_6433_40)) (((((((Protonymphidia_3948_59_________ANTS__O_o__o (Menander_7339_49_________________ANTS__O_o__o Lemonias_3950_55_________________ANTS__O_o__o)) Themone_7643_34) ((Ancyluris_6834_45 (((((((Argyrogrammana_6709_42 Napaea_m17_89_______________________________) (Crocozona_3995_62 (Leucochimona_9302_33_______________________X T_Poritia_erycinoides))) Calephelis_9356_22____________________X______) (((Emesis_6837_46_______________________________ Stalachtis_6422_39) Hyphilaria_5239_91) Mesene_7450_28_________________________X______)) (Cariomothis_6518_41 Rhetus_3973_60)) Zelotea_4057_79) ((Mesophthalma_6327_88_____________________X (Methone_8523_95 Sarota_9358_21)) Riodina_lysippus))) Necryia_5966_93)) Mesosemia_6516_85_________________________X) Melanis_7612_30_________________________X______) Isapis_6815_44) Monethe_3977_61)) ((Chamaelimnas_3942_57 Pachythone_9050_18) Roeberella_7457_36)) Metacharis_9347_23__________________________)) (Amarynthis_5220_51 (Esthemopsis_7813_32 Symmachia_8849_08___________________________))) ((Calydna_6346_83 ((Euselasia_8816_01_________________________X ((Panara_3932_56 Phaenochitonia_6926_47) Pandemos_8572_97)) Parcella_4012_64)) Detritivora_9327_25))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Hyphilaria_5239_91) Pachythone_9050_18) Lemonias_3950_55_________________ANTS__O_o__o) (Adelotypa_7807_37_________________ANTS__O_o__o Helicopis_5318_54)) Synargis_7605_29_________________ANTS__O_o__o) ((((Setabis_6330_38_________________ANTS__O_o__o Exoplisia_6433_40) Leucochimona_7617_87_______________________X) T_Poritia_erycinoides) ((((((((Thisbe_6766_43___________________ANTS__O_o__o Anteros_7097_50______________________________) (Ancyluris_6834_45 (Necryia_5966_93 Emesis_mandana___________________________X_____))) (((((((((Argyrogrammana_6709_42 Mesosemia_6516_85_________________________X) Crocozona_3995_62) Parcella_4012_64) Ithomiola_5924_82) (Detritivora_9327_25 Mesene_7450_28_________________________X______)) Euselasia_8816_01_________________________X) Calephelis_9356_22____________________X______) Lepricornis_5041_80) Ourocnemis_4621_90___________________________)) Sarota_9358_21) Emesis_6837_46_______________________________) Thysanota_5241_81_________________ANTS__O_o__o) Echenais_7168_48) Cyrenia_3992_63))) ((Calospila_8532_96_________________ANTS__O_o__o Napaea_m17_89_______________________________) Mesosemia_8822_02_________________________X)) Amarynthis_5220_51) Leucochimona_9306_27_______________________X) Menander_7339_49_________________ANTS__O_o__o) Theope_7789_65__________________ANTS__O_o__o) ((Stalachtis_6422_39 Apodemia_mormo) Symmachia_8849_08___________________________)) Roeberella_7457_36) ((((((((Protonymphidia_3948_59_________ANTS__O_o__o Panara_3932_56) Caria_7625_31_______________________________) Cariomothis_6518_41) ((Cremna_5260_53 ((Leucochimona_9302_33_______________________X Monethe_3977_61) (Melanis_7612_30_________________________X______ Metacharis_9347_23__________________________))) T_Miletus_ancon)) (Brachyglemis_3956_58 ((Esthemopsis_7813_32 Pandemos_8572_97) Riodina_lysippus))) (((Chamaelimnas_3942_57 Methone_8523_95) Isapis_6815_44) (Charis_9365_19_______________________________ Mesophthalma_6327_88_____________________X))) Ithomeis_7773_35) (Alesa_8380_84____________________ANTS__O_o__o ((Theope_9026_5____________________ANTS__O_o__o Phaenochitonia_6926_47) (Semomesia_5223_52_________________________X Themone_7643_34))))) (Eurybia_9363_20_________________ANTS__O_o__o Perophthalma_9330_24_______________________X)) Rhetus_3973_60) Styx_infernalis) Zelotea_4057_79) (Calydna_6346_83 Euselasia_orfita____________________________X)) Juditha_9315_26__________________ANTS__O_o__o) Hamearis_lucina)*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((Uraneis_hyalina____________________ANTS__O_o__o ((((Lemonias_3950_55_________________ANTS__O_o__o (Brachyglemis_3956_58 Echenais_7168_48)) Calospila_8532_96_________________ANTS__O_o__o) ((((((((Calephelis_9356_22____________________X______ Cyrenia_3992_63) (Panara_3932_56 T_Poritia_erycinoides)) ((Phaenochitonia_6926_47 (Sarota_9358_21 Anteros_7097_50______________________________)) Zelotea_4057_79)) Charis_9365_19_______________________________) Methone_8523_95) Symmachia_8849_08___________________________) Ithomeis_7773_35) Pandemos_8572_97)) (((Thysanota_5241_81_________________ANTS__O_o__o Themone_7643_34) (Leucochimona_7617_87_______________________X Stalachtis_6422_39)) Chamaelimnas_3942_57))) (Juditha_9315_26__________________ANTS__O_o__o T_Miletus_ancon)) (((((((((Alesa_8380_84____________________ANTS__O_o__o Thisbe_6766_43___________________ANTS__O_o__o) Mesosemia_8822_02_________________________X) Mesene_7450_28_________________________X______) (((Synargis_7605_29_________________ANTS__O_o__o Calydna_6346_83) Cremna_5260_53) Monethe_3977_61)) Pachythone_9050_18) ((Amarynthis_5220_51 Metacharis_9347_23__________________________) Rhetus_3973_60)) Isapis_6815_44) Riodina_lysippus) Ancyluris_6834_45)) Detritivora_9327_25) Menander_7339_49_________________ANTS__O_o__o) Lepricornis_5041_80) (Mesophthalma_6327_88_____________________X Ourocnemis_4621_90___________________________)) Perophthalma_9330_24_______________________X) Necryia_5966_93) (((((((((Adelotypa_7807_37_________________ANTS__O_o__o ((((Ithomiola_5924_82 Roeberella_7457_36) Mesosemia_6516_85_________________________X) Styx_infernalis) Semomesia_5223_52_________________________X)) Hamearis_lucina) Cariomothis_6518_41) Leucochimona_9302_33_______________________X) Apodemia_mormo) Emesis_mandana___________________________X_____) Melanis_7612_30_________________________X______) ((((Setabis_6330_38_________________ANTS__O_o__o Parcella_4012_64) Napaea_m17_89_______________________________) (Theope_9026_5____________________ANTS__O_o__o (Emesis_6837_46_______________________________ Hyphilaria_5239_91))) Helicopis_5318_54)) Theope_7789_65__________________ANTS__O_o__o)) (((((Protonymphidia_3948_59_________ANTS__O_o__o (Eurybia_9363_20_________________ANTS__O_o__o Leucochimona_9306_27_______________________X)) (Argyrogrammana_6709_42 Caria_7625_31_______________________________)) Exoplisia_6433_40) Crocozona_3995_62) ((Esthemopsis_7813_32 Euselasia_8816_01_________________________X) Euselasia_orfita____________________________X)))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o (((Menander_7339_49_________________ANTS__O_o__o ((Cariomothis_6518_41 (Leucochimona_9302_33_______________________X Metacharis_9347_23__________________________)) (Chamaelimnas_3942_57 Perophthalma_9330_24_______________________X))) Cremna_5260_53) Exoplisia_6433_40)) ((((((((((Alesa_8380_84____________________ANTS__O_o__o Riodina_lysippus) Themone_7643_34) Crocozona_3995_62) (Methone_8523_95 Euselasia_orfita____________________________X)) Juditha_9315_26__________________ANTS__O_o__o) ((Theope_9026_5____________________ANTS__O_o__o (((Charis_9365_19_______________________________ Ithomeis_7773_35) Hamearis_lucina) ((((Cyrenia_3992_63 Styx_infernalis) Mesophthalma_6327_88_____________________X) Helicopis_5318_54) Mesene_7450_28_________________________X______))) ((Thisbe_6766_43___________________ANTS__O_o__o Synargis_7605_29_________________ANTS__O_o__o) Amarynthis_5220_51))) Anteros_7097_50______________________________) Ithomiola_5924_82) Mesosemia_8822_02_________________________X) Eurybia_9363_20_________________ANTS__O_o__o)) Lepricornis_5041_80) Argyrogrammana_6709_42) (Setabis_6330_38_________________ANTS__O_o__o Rhetus_3973_60)) Calydna_6346_83) Panara_3932_56) Thysanota_5241_81_________________ANTS__O_o__o) (((Lemonias_3950_55_________________ANTS__O_o__o Leucochimona_7617_87_______________________X) ((Calephelis_9356_22____________________X______ Pachythone_9050_18) T_Poritia_erycinoides)) Pandemos_8572_97)) ((Leucochimona_9306_27_______________________X Phaenochitonia_6926_47) (Necryia_5966_93 T_Miletus_ancon))) Zelotea_4057_79) Brachyglemis_3956_58) Caria_7625_31_______________________________) Detritivora_9327_25) (Isapis_6815_44 (Symmachia_8849_08___________________________ Emesis_mandana___________________________X_____))) Parcella_4012_64) Melanis_7612_30_________________________X______) (((((Adelotypa_7807_37_________________ANTS__O_o__o Theope_7789_65__________________ANTS__O_o__o) (Emesis_6837_46_______________________________ Apodemia_mormo)) Stalachtis_6422_39) (Esthemopsis_7813_32 ((Hyphilaria_5239_91 Napaea_m17_89_______________________________) Mesosemia_6516_85_________________________X))) ((((Calospila_8532_96_________________ANTS__O_o__o Euselasia_8816_01_________________________X) Semomesia_5223_52_________________________X) Echenais_7168_48) (Roeberella_7457_36 Sarota_9358_21)))) (Protonymphidia_3948_59_________ANTS__O_o__o (Ancyluris_6834_45 (Monethe_3977_61 Ourocnemis_4621_90___________________________))))*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Thisbe_6766_43___________________ANTS__O_o__o) Roeberella_7457_36) Emesis_mandana___________________________X_____) (((((((Theope_9026_5____________________ANTS__O_o__o (Cyrenia_3992_63 (((Helicopis_5318_54 (Ourocnemis_4621_90___________________________ Perophthalma_9330_24_______________________X)) Pandemos_8572_97) (Mesosemia_6516_85_________________________X Euselasia_orfita____________________________X)))) (Napaea_m17_89_______________________________ Parcella_4012_64)) Lemonias_3950_55_________________ANTS__O_o__o) Thysanota_5241_81_________________ANTS__O_o__o) (((Euselasia_8816_01_________________________X Riodina_lysippus) (Leucochimona_9306_27_______________________X Panara_3932_56)) Symmachia_8849_08___________________________)) Phaenochitonia_6926_47) Lepricornis_5041_80)) ((Theope_7789_65__________________ANTS__O_o__o (Brachyglemis_3956_58 Charis_9365_19_______________________________)) (Ancyluris_6834_45 Mesene_7450_28_________________________X______))) (Emesis_6837_46_______________________________ Styx_infernalis)) ((Alesa_8380_84____________________ANTS__O_o__o (Chamaelimnas_3942_57 Detritivora_9327_25)) Hamearis_lucina)) ((((Setabis_6330_38_________________ANTS__O_o__o Mesosemia_8822_02_________________________X) ((Synargis_7605_29_________________ANTS__O_o__o ((((Amarynthis_5220_51 Leucochimona_9302_33_______________________X) Rhetus_3973_60) Caria_7625_31_______________________________) Methone_8523_95)) (((((((((Argyrogrammana_6709_42 Sarota_9358_21) Isapis_6815_44) Calephelis_9356_22____________________X______) Themone_7643_34) Stalachtis_6422_39) Ithomiola_5924_82) (Exoplisia_6433_40 Necryia_5966_93)) Anteros_7097_50______________________________) Pachythone_9050_18))) Monethe_3977_61) Menander_7339_49_________________ANTS__O_o__o)) Echenais_7168_48) Apodemia_mormo) T_Miletus_ancon) ((Protonymphidia_3948_59_________ANTS__O_o__o Juditha_9315_26__________________ANTS__O_o__o) Leucochimona_7617_87_______________________X)) Semomesia_5223_52_________________________X) Eurybia_9363_20_________________ANTS__O_o__o) Cremna_5260_53) Calospila_8532_96_________________ANTS__O_o__o) Crocozona_3995_62) ((Adelotypa_7807_37_________________ANTS__O_o__o Esthemopsis_7813_32) ((((((Calydna_6346_83 Ithomeis_7773_35) Hyphilaria_5239_91) Metacharis_9347_23__________________________) T_Poritia_erycinoides) Zelotea_4057_79) ((Cariomothis_6518_41 Melanis_7612_30_________________________X______) Mesophthalma_6327_88_____________________X))))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o ((((((((Theope_7789_65__________________ANTS__O_o__o Ithomeis_7773_35) T_Miletus_ancon) Panara_3932_56) Helicopis_5318_54) Chamaelimnas_3942_57) (((Thysanota_5241_81_________________ANTS__O_o__o Argyrogrammana_6709_42) (Roeberella_7457_36 Stalachtis_6422_39)) Esthemopsis_7813_32)) Cyrenia_3992_63) Metacharis_9347_23__________________________)) (Crocozona_3995_62 Pandemos_8572_97)) (Lepricornis_5041_80 Leucochimona_7617_87_______________________X)) Detritivora_9327_25) Riodina_lysippus) (Caria_7625_31_______________________________ (Methone_8523_95 (Ourocnemis_4621_90___________________________ (Perophthalma_9330_24_______________________X Phaenochitonia_6926_47))))) ((((Euselasia_8816_01_________________________X Sarota_9358_21) Pachythone_9050_18) Styx_infernalis) Apodemia_mormo)) Emesis_6837_46_______________________________) Monethe_3977_61) Mesosemia_6516_85_________________________X) (Calephelis_9356_22____________________X______ Emesis_mandana___________________________X_____)) Menander_7339_49_________________ANTS__O_o__o) Calospila_8532_96_________________ANTS__O_o__o) (Eurybia_9363_20_________________ANTS__O_o__o (Brachyglemis_3956_58 Rhetus_3973_60))) ((Theope_9026_5____________________ANTS__O_o__o (((((((Amarynthis_5220_51 Themone_7643_34) Anteros_7097_50______________________________) Euselasia_orfita____________________________X) Hamearis_lucina) Mesosemia_8822_02_________________________X) Echenais_7168_48) Semomesia_5223_52_________________________X)) (Ithomiola_5924_82 Melanis_7612_30_________________________X______))) (((((((Protonymphidia_3948_59_________ANTS__O_o__o Zelotea_4057_79) (Thisbe_6766_43___________________ANTS__O_o__o (Hyphilaria_5239_91 Leucochimona_9306_27_______________________X))) Parcella_4012_64) Isapis_6815_44) Calydna_6346_83) Synargis_7605_29_________________ANTS__O_o__o) Alesa_8380_84____________________ANTS__O_o__o)) (Lemonias_3950_55_________________ANTS__O_o__o Cariomothis_6518_41)) Juditha_9315_26__________________ANTS__O_o__o) T_Poritia_erycinoides) Mesene_7450_28_________________________X______) (((((((Adelotypa_7807_37_________________ANTS__O_o__o ((Ancyluris_6834_45 Leucochimona_9302_33_______________________X) Cremna_5260_53)) Necryia_5966_93) Symmachia_8849_08___________________________) Mesophthalma_6327_88_____________________X) Napaea_m17_89_______________________________) Charis_9365_19_______________________________) (Setabis_6330_38_________________ANTS__O_o__o Exoplisia_6433_40)))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((Uraneis_hyalina____________________ANTS__O_o__o Sarota_9358_21) (Mesophthalma_6327_88_____________________X Riodina_lysippus)) Perophthalma_9330_24_______________________X) ((((((((((Adelotypa_7807_37_________________ANTS__O_o__o (Crocozona_3995_62 Napaea_m17_89_______________________________)) Thysanota_5241_81_________________ANTS__O_o__o) T_Poritia_erycinoides) Leucochimona_9306_27_______________________X) Ithomeis_7773_35) Pachythone_9050_18) (((Protonymphidia_3948_59_________ANTS__O_o__o Ourocnemis_4621_90___________________________) Parcella_4012_64) (Exoplisia_6433_40 (Panara_3932_56 Phaenochitonia_6926_47)))) ((Theope_7789_65__________________ANTS__O_o__o Themone_7643_34) (Calospila_8532_96_________________ANTS__O_o__o ((Emesis_6837_46_______________________________ Styx_infernalis) Helicopis_5318_54)))) Mesosemia_6516_85_________________________X) Emesis_mandana___________________________X_____)) (((((Alesa_8380_84____________________ANTS__O_o__o (((((((((((Setabis_6330_38_________________ANTS__O_o__o Calephelis_9356_22____________________X______) Cyrenia_3992_63) (((Eurybia_9363_20_________________ANTS__O_o__o Synargis_7605_29_________________ANTS__O_o__o) Leucochimona_7617_87_______________________X) Monethe_3977_61)) (Mesosemia_8822_02_________________________X (Methone_8523_95 Semomesia_5223_52_________________________X))) Metacharis_9347_23__________________________) Pandemos_8572_97) Zelotea_4057_79) (Detritivora_9327_25 Melanis_7612_30_________________________X______)) Charis_9365_19_______________________________) ((Theope_9026_5____________________ANTS__O_o__o Leucochimona_9302_33_______________________X) (((((Menander_7339_49_________________ANTS__O_o__o ((((((Lemonias_3950_55_________________ANTS__O_o__o ((Thisbe_6766_43___________________ANTS__O_o__o (Stalachtis_6422_39 T_Miletus_ancon)) Hamearis_lucina)) Caria_7625_31_______________________________) Isapis_6815_44) ((Calydna_6346_83 Mesene_7450_28_________________________X______) Euselasia_orfita____________________________X)) ((((Amarynthis_5220_51 Esthemopsis_7813_32) (Chamaelimnas_3942_57 Anteros_7097_50______________________________)) Cremna_5260_53) Hyphilaria_5239_91)) Argyrogrammana_6709_42)) Apodemia_mormo) (Euselasia_8816_01_________________________X Symmachia_8849_08___________________________)) Rhetus_3973_60) Brachyglemis_3956_58))) Ithomiola_5924_82)) Juditha_9315_26__________________ANTS__O_o__o) Lepricornis_5041_80) Necryia_5966_93) (((Ancyluris_6834_45 Cariomothis_6518_41) Roeberella_7457_36) Echenais_7168_48)))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((Uraneis_hyalina____________________ANTS__O_o__o Monethe_3977_61) ((Charis_9365_19_______________________________ Cremna_5260_53) Detritivora_9327_25)) (((((((((((((((((((((Adelotypa_7807_37_________________ANTS__O_o__o Sarota_9358_21) (Setabis_6330_38_________________ANTS__O_o__o ((Emesis_6837_46_______________________________ (Methone_8523_95 Anteros_7097_50______________________________)) Euselasia_orfita____________________________X))) Napaea_m17_89_______________________________) Eurybia_9363_20_________________ANTS__O_o__o) Cyrenia_3992_63) (Exoplisia_6433_40 Symmachia_8849_08___________________________)) Themone_7643_34) ((Ithomeis_7773_35 Riodina_lysippus) Panara_3932_56)) (((((((((((Alesa_8380_84____________________ANTS__O_o__o Mesosemia_8822_02_________________________X) Mesosemia_6516_85_________________________X) (Caria_7625_31_______________________________ Pandemos_8572_97)) (Lemonias_3950_55_________________ANTS__O_o__o Hamearis_lucina)) (Euselasia_8816_01_________________________X Ourocnemis_4621_90___________________________)) Zelotea_4057_79) Theope_7789_65__________________ANTS__O_o__o) (Synargis_7605_29_________________ANTS__O_o__o Brachyglemis_3956_58)) T_Poritia_erycinoides) ((Crocozona_3995_62 Necryia_5966_93) Pachythone_9050_18)) Thysanota_5241_81_________________ANTS__O_o__o)) ((Hyphilaria_5239_91 Apodemia_mormo) Melanis_7612_30_________________________X______)) Perophthalma_9330_24_______________________X) Protonymphidia_3948_59_________ANTS__O_o__o) Ancyluris_6834_45) Calephelis_9356_22____________________X______) ((Echenais_7168_48 Semomesia_5223_52_________________________X) T_Miletus_ancon)) Juditha_9315_26__________________ANTS__O_o__o) Calospila_8532_96_________________ANTS__O_o__o) ((((Theope_9026_5____________________ANTS__O_o__o (((Thisbe_6766_43___________________ANTS__O_o__o Cariomothis_6518_41) (Argyrogrammana_6709_42 Phaenochitonia_6926_47)) Stalachtis_6422_39)) (Chamaelimnas_3942_57 Lepricornis_5041_80)) Metacharis_9347_23__________________________) ((((Amarynthis_5220_51 (Ithomiola_5924_82 Styx_infernalis)) Leucochimona_7617_87_______________________X) Calydna_6346_83) Leucochimona_9302_33_______________________X))) Roeberella_7457_36) ((((Menander_7339_49_________________ANTS__O_o__o (Parcella_4012_64 Rhetus_3973_60)) Esthemopsis_7813_32) (((Helicopis_5318_54 Leucochimona_9306_27_______________________X) Mesophthalma_6327_88_____________________X) Isapis_6815_44)) Mesene_7450_28_________________________X______)) Emesis_mandana___________________________X_____))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((Uraneis_hyalina____________________ANTS__O_o__o Echenais_7168_48) Euselasia_orfita____________________________X) (((((((((((((((((Adelotypa_7807_37_________________ANTS__O_o__o Calephelis_9356_22____________________X______) Monethe_3977_61) ((Menander_7339_49_________________ANTS__O_o__o Cyrenia_3992_63) Thysanota_5241_81_________________ANTS__O_o__o)) (((Alesa_8380_84____________________ANTS__O_o__o Pachythone_9050_18) (Mesosemia_6516_85_________________________X Ourocnemis_4621_90___________________________)) Ithomiola_5924_82)) Anteros_7097_50______________________________) ((Esthemopsis_7813_32 Euselasia_8816_01_________________________X) Mesosemia_8822_02_________________________X)) Mesene_7450_28_________________________X______) Symmachia_8849_08___________________________) ((Lemonias_3950_55_________________ANTS__O_o__o Amarynthis_5220_51) T_Miletus_ancon)) Hamearis_lucina) (Leucochimona_7617_87_______________________X Napaea_m17_89_______________________________)) T_Poritia_erycinoides) Mesophthalma_6327_88_____________________X) Argyrogrammana_6709_42) (Setabis_6330_38_________________ANTS__O_o__o Themone_7643_34)) Emesis_mandana___________________________X_____) ((Calospila_8532_96_________________ANTS__O_o__o (Brachyglemis_3956_58 ((((Cremna_5260_53 Emesis_6837_46_______________________________) Rhetus_3973_60) (Crocozona_3995_62 Panara_3932_56)) Isapis_6815_44))) (((Cariomothis_6518_41 Charis_9365_19_______________________________) Parcella_4012_64) Leucochimona_9306_27_______________________X)))) Riodina_lysippus) Caria_7625_31_______________________________) Hyphilaria_5239_91) Helicopis_5318_54) Roeberella_7457_36) Exoplisia_6433_40) (((((((Protonymphidia_3948_59_________ANTS__O_o__o Eurybia_9363_20_________________ANTS__O_o__o) Sarota_9358_21) (Detritivora_9327_25 Methone_8523_95)) Theope_9026_5____________________ANTS__O_o__o) ((Theope_7789_65__________________ANTS__O_o__o (Calydna_6346_83 Necryia_5966_93)) Phaenochitonia_6926_47)) Melanis_7612_30_________________________X______) Thisbe_6766_43___________________ANTS__O_o__o)) (((((((((Juditha_9315_26__________________ANTS__O_o__o Chamaelimnas_3942_57) Lepricornis_5041_80) Leucochimona_9302_33_______________________X) Semomesia_5223_52_________________________X) Ancyluris_6834_45) (Synargis_7605_29_________________ANTS__O_o__o Styx_infernalis)) Zelotea_4057_79) (((Ithomeis_7773_35 Perophthalma_9330_24_______________________X) Apodemia_mormo) (Pandemos_8572_97 Stalachtis_6422_39))) Metacharis_9347_23__________________________))*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((Uraneis_hyalina____________________ANTS__O_o__o ((Alesa_8380_84____________________ANTS__O_o__o Brachyglemis_3956_58) (Lemonias_3950_55_________________ANTS__O_o__o ((((Cremna_5260_53 Apodemia_mormo) Roeberella_7457_36) Themone_7643_34) Lepricornis_5041_80)))) Ithomeis_7773_35) ((((((Adelotypa_7807_37_________________ANTS__O_o__o Cyrenia_3992_63) Leucochimona_9302_33_______________________X) Riodina_lysippus) (((((Juditha_9315_26__________________ANTS__O_o__o Necryia_5966_93) (((Chamaelimnas_3942_57 ((Echenais_7168_48 Stalachtis_6422_39) Ithomiola_5924_82)) (Melanis_7612_30_________________________X______ Sarota_9358_21)) Euselasia_orfita____________________________X)) Symmachia_8849_08___________________________) Exoplisia_6433_40) Euselasia_8816_01_________________________X)) Thysanota_5241_81_________________ANTS__O_o__o) T_Miletus_ancon)) Amarynthis_5220_51) (((((((((((Protonymphidia_3948_59_________ANTS__O_o__o Isapis_6815_44) Mesosemia_6516_85_________________________X) Ourocnemis_4621_90___________________________) ((Menander_7339_49_________________ANTS__O_o__o (Metacharis_9347_23__________________________ Panara_3932_56)) Synargis_7605_29_________________ANTS__O_o__o)) Leucochimona_9306_27_______________________X) Emesis_mandana___________________________X_____) (((Calydna_6346_83 Napaea_m17_89_______________________________) Mesosemia_8822_02_________________________X) Perophthalma_9330_24_______________________X)) Parcella_4012_64) Emesis_6837_46_______________________________) Argyrogrammana_6709_42) ((Theope_7789_65__________________ANTS__O_o__o (Caria_7625_31_______________________________ Styx_infernalis)) (((((((Theope_9026_5____________________ANTS__O_o__o Hamearis_lucina) (Thisbe_6766_43___________________ANTS__O_o__o T_Poritia_erycinoides)) (Monethe_3977_61 Rhetus_3973_60)) Calospila_8532_96_________________ANTS__O_o__o) Pachythone_9050_18) Crocozona_3995_62) Cariomothis_6518_41)))) Mesene_7450_28_________________________X______) Charis_9365_19_______________________________) ((((Setabis_6330_38_________________ANTS__O_o__o Zelotea_4057_79) Anteros_7097_50______________________________) (((Esthemopsis_7813_32 Hyphilaria_5239_91) Helicopis_5318_54) (Mesophthalma_6327_88_____________________X Semomesia_5223_52_________________________X))) ((Eurybia_9363_20_________________ANTS__O_o__o Ancyluris_6834_45) (Calephelis_9356_22____________________X______ Pandemos_8572_97)))) Methone_8523_95) Detritivora_9327_25) Phaenochitonia_6926_47) Leucochimona_7617_87_______________________X)*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Brachyglemis_3956_58) Detritivora_9327_25) Hyphilaria_5239_91) Parcella_4012_64) Sarota_9358_21) Cariomothis_6518_41) Zelotea_4057_79) ((Synargis_7605_29_________________ANTS__O_o__o Cyrenia_3992_63) Anteros_7097_50______________________________)) (((((((((Theope_7789_65__________________ANTS__O_o__o Mesophthalma_6327_88_____________________X) (Lemonias_3950_55_________________ANTS__O_o__o Esthemopsis_7813_32)) Menander_7339_49_________________ANTS__O_o__o) Exoplisia_6433_40) Stalachtis_6422_39) Semomesia_5223_52_________________________X) (Ancyluris_6834_45 (Isapis_6815_44 (Leucochimona_9302_33_______________________X Necryia_5966_93)))) (Crocozona_3995_62 Leucochimona_7617_87_______________________X)) (Helicopis_5318_54 Rhetus_3973_60))) Adelotypa_7807_37_________________ANTS__O_o__o) (((((Alesa_8380_84____________________ANTS__O_o__o Metacharis_9347_23__________________________) Caria_7625_31_______________________________) (((Emesis_6837_46_______________________________ Euselasia_orfita____________________________X) Lepricornis_5041_80) Ourocnemis_4621_90___________________________)) Symmachia_8849_08___________________________) Methone_8523_95)) (((((Charis_9365_19_______________________________ Phaenochitonia_6926_47) T_Poritia_erycinoides) Hamearis_lucina) Riodina_lysippus) T_Miletus_ancon)) ((Calydna_6346_83 Roeberella_7457_36) Apodemia_mormo)) ((((((((Theope_9026_5____________________ANTS__O_o__o ((((Argyrogrammana_6709_42 Ithomeis_7773_35) Styx_infernalis) Echenais_7168_48) Ithomiola_5924_82)) Leucochimona_9306_27_______________________X) Emesis_mandana___________________________X_____) Chamaelimnas_3942_57) Themone_7643_34) Melanis_7612_30_________________________X______) Pachythone_9050_18) Monethe_3977_61)) (((Setabis_6330_38_________________ANTS__O_o__o Cremna_5260_53) Mesene_7450_28_________________________X______) Thysanota_5241_81_________________ANTS__O_o__o)) Calephelis_9356_22____________________X______) Euselasia_8816_01_________________________X) (((Protonymphidia_3948_59_________ANTS__O_o__o ((((Calospila_8532_96_________________ANTS__O_o__o ((Amarynthis_5220_51 Mesosemia_8822_02_________________________X) Pandemos_8572_97)) Perophthalma_9330_24_______________________X) Thisbe_6766_43___________________ANTS__O_o__o) Panara_3932_56)) Napaea_m17_89_______________________________) (Eurybia_9363_20_________________ANTS__O_o__o Mesosemia_6516_85_________________________X))) Juditha_9315_26__________________ANTS__O_o__o)*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Hamearis_lucina) (Theope_7789_65__________________ANTS__O_o__o Panara_3932_56)) ((Argyrogrammana_6709_42 (Hyphilaria_5239_91 (Mesosemia_6516_85_________________________X Phaenochitonia_6926_47))) Ithomiola_5924_82)) (((Leucochimona_7617_87_______________________X (Methone_8523_95 Anteros_7097_50______________________________)) Stalachtis_6422_39) Emesis_mandana___________________________X_____)) ((((((Adelotypa_7807_37_________________ANTS__O_o__o Charis_9365_19_______________________________) Chamaelimnas_3942_57) Riodina_lysippus) Cyrenia_3992_63) Calephelis_9356_22____________________X______) Zelotea_4057_79)) Necryia_5966_93) (Juditha_9315_26__________________ANTS__O_o__o (Mesene_7450_28_________________________X______ Euselasia_orfita____________________________X))) ((Lemonias_3950_55_________________ANTS__O_o__o Thisbe_6766_43___________________ANTS__O_o__o) T_Miletus_ancon)) (((((Menander_7339_49_________________ANTS__O_o__o Pandemos_8572_97) (Brachyglemis_3956_58 (Calydna_6346_83 (Caria_7625_31_______________________________ (Cariomothis_6518_41 (Euselasia_8816_01_________________________X (Semomesia_5223_52_________________________X Styx_infernalis))))))) (Metacharis_9347_23__________________________ T_Poritia_erycinoides)) Isapis_6815_44) (Leucochimona_9306_27_______________________X Parcella_4012_64))) Crocozona_3995_62) Thysanota_5241_81_________________ANTS__O_o__o) Symmachia_8849_08___________________________) (((((((((Protonymphidia_3948_59_________ANTS__O_o__o (Alesa_8380_84____________________ANTS__O_o__o Apodemia_mormo)) Eurybia_9363_20_________________ANTS__O_o__o) Mesosemia_8822_02_________________________X) Napaea_m17_89_______________________________) Detritivora_9327_25) Mesophthalma_6327_88_____________________X) (((Setabis_6330_38_________________ANTS__O_o__o (((Theope_9026_5____________________ANTS__O_o__o ((((((((Synargis_7605_29_________________ANTS__O_o__o ((Amarynthis_5220_51 Monethe_3977_61) Echenais_7168_48)) Ithomeis_7773_35) Themone_7643_34) Melanis_7612_30_________________________X______) Rhetus_3973_60) (Emesis_6837_46_______________________________ Esthemopsis_7813_32)) (Helicopis_5318_54 Roeberella_7457_36)) ((Ancyluris_6834_45 Leucochimona_9302_33_______________________X) Perophthalma_9330_24_______________________X))) (Cremna_5260_53 Lepricornis_5041_80)) Exoplisia_6433_40)) Calospila_8532_96_________________ANTS__O_o__o) Pachythone_9050_18)) Ourocnemis_4621_90___________________________) Sarota_9358_21))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((Uraneis_hyalina____________________ANTS__O_o__o Juditha_9315_26__________________ANTS__O_o__o) (((((((((((((((((((((Protonymphidia_3948_59_________ANTS__O_o__o Emesis_mandana___________________________X_____) ((((Alesa_8380_84____________________ANTS__O_o__o (Cariomothis_6518_41 Zelotea_4057_79)) Emesis_6837_46_______________________________) Panara_3932_56) Calydna_6346_83)) (Theope_7789_65__________________ANTS__O_o__o Lemonias_3950_55_________________ANTS__O_o__o)) Argyrogrammana_6709_42) Perophthalma_9330_24_______________________X) Amarynthis_5220_51) Brachyglemis_3956_58) Crocozona_3995_62) (((((((((Calospila_8532_96_________________ANTS__O_o__o Cyrenia_3992_63) Ithomeis_7773_35) ((Leucochimona_7617_87_______________________X Phaenochitonia_6926_47) Parcella_4012_64)) Thisbe_6766_43___________________ANTS__O_o__o) (Napaea_m17_89_______________________________ Symmachia_8849_08___________________________)) Themone_7643_34) Synargis_7605_29_________________ANTS__O_o__o) Stalachtis_6422_39) Ancyluris_6834_45)) Charis_9365_19_______________________________) (Cremna_5260_53 T_Miletus_ancon)) Semomesia_5223_52_________________________X) Thysanota_5241_81_________________ANTS__O_o__o) Menander_7339_49_________________ANTS__O_o__o) Detritivora_9327_25) Leucochimona_9306_27_______________________X) Lepricornis_5041_80) Setabis_6330_38_________________ANTS__O_o__o) (((((((Calephelis_9356_22____________________X______ Mesene_7450_28_________________________X______) Chamaelimnas_3942_57) (Ourocnemis_4621_90___________________________ Pachythone_9050_18)) (Echenais_7168_48 Sarota_9358_21)) Apodemia_mormo) Pandemos_8572_97) Euselasia_8816_01_________________________X)) Isapis_6815_44) ((Esthemopsis_7813_32 Roeberella_7457_36) Styx_infernalis))) Theope_9026_5____________________ANTS__O_o__o) ((Adelotypa_7807_37_________________ANTS__O_o__o Riodina_lysippus) Melanis_7612_30_________________________X______)) Hyphilaria_5239_91) ((((Eurybia_9363_20_________________ANTS__O_o__o (Caria_7625_31_______________________________ Hamearis_lucina)) (Mesophthalma_6327_88_____________________X Methone_8523_95)) (Monethe_3977_61 T_Poritia_erycinoides)) Exoplisia_6433_40)) ((((((((Helicopis_5318_54 Ithomiola_5924_82) (Mesosemia_8822_02_________________________X Metacharis_9347_23__________________________)) Mesosemia_6516_85_________________________X) Euselasia_orfita____________________________X) Necryia_5966_93) Rhetus_3973_60) Leucochimona_9302_33_______________________X) Anteros_7097_50______________________________))*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Leucochimona_9306_27_______________________X) Mesophthalma_6327_88_____________________X) (Melanis_7612_30_________________________X______ (((Methone_8523_95 Perophthalma_9330_24_______________________X) Necryia_5966_93) Zelotea_4057_79))) Thysanota_5241_81_________________ANTS__O_o__o) ((((((((Adelotypa_7807_37_________________ANTS__O_o__o ((((((((((Theope_7789_65__________________ANTS__O_o__o Helicopis_5318_54) ((Thisbe_6766_43___________________ANTS__O_o__o Semomesia_5223_52_________________________X) Ancyluris_6834_45)) (((Calephelis_9356_22____________________X______ Anteros_7097_50______________________________) Apodemia_mormo) (Emesis_6837_46_______________________________ Riodina_lysippus))) Pachythone_9050_18) Isapis_6815_44) Mesosemia_6516_85_________________________X) (Synargis_7605_29_________________ANTS__O_o__o Styx_infernalis)) Hyphilaria_5239_91) ((Chamaelimnas_3942_57 Napaea_m17_89_______________________________) ((((Cremna_5260_53 Detritivora_9327_25) Esthemopsis_7813_32) Echenais_7168_48) Pandemos_8572_97))) Rhetus_3973_60)) (Calydna_6346_83 Euselasia_8816_01_________________________X)) (Protonymphidia_3948_59_________ANTS__O_o__o Lemonias_3950_55_________________ANTS__O_o__o)) (Leucochimona_9302_33_______________________X Roeberella_7457_36)) Argyrogrammana_6709_42) Ourocnemis_4621_90___________________________) ((((Setabis_6330_38_________________ANTS__O_o__o (((Juditha_9315_26__________________ANTS__O_o__o Mesosemia_8822_02_________________________X) Lepricornis_5041_80) (Exoplisia_6433_40 Themone_7643_34))) Metacharis_9347_23__________________________) Amarynthis_5220_51) (Monethe_3977_61 Phaenochitonia_6926_47))) Euselasia_orfita____________________________X)) Parcella_4012_64) (Symmachia_8849_08___________________________ T_Poritia_erycinoides)) Emesis_mandana___________________________X_____) (((((((Eurybia_9363_20_________________ANTS__O_o__o Leucochimona_7617_87_______________________X) T_Miletus_ancon) Theope_9026_5____________________ANTS__O_o__o) Stalachtis_6422_39) Sarota_9358_21) Menander_7339_49_________________ANTS__O_o__o) Charis_9365_19_______________________________)) Brachyglemis_3956_58) (Crocozona_3995_62 Ithomiola_5924_82)) (Cyrenia_3992_63 Mesene_7450_28_________________________X______)) Alesa_8380_84____________________ANTS__O_o__o) Hamearis_lucina) (((Calospila_8532_96_________________ANTS__O_o__o Cariomothis_6518_41) Caria_7625_31_______________________________) Ithomeis_7773_35)) Panara_3932_56)*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Isapis_6815_44) ((Calospila_8532_96_________________ANTS__O_o__o Stalachtis_6422_39) Ithomiola_5924_82)) (Caria_7625_31_______________________________ Hamearis_lucina)) ((Pandemos_8572_97 Perophthalma_9330_24_______________________X) Zelotea_4057_79)) Lemonias_3950_55_________________ANTS__O_o__o) (((((Protonymphidia_3948_59_________ANTS__O_o__o Styx_infernalis) (((((Alesa_8380_84____________________ANTS__O_o__o (Roeberella_7457_36 Euselasia_orfita____________________________X)) Cariomothis_6518_41) (Amarynthis_5220_51 Mesosemia_8822_02_________________________X)) (((Synargis_7605_29_________________ANTS__O_o__o (Echenais_7168_48 (Leucochimona_9302_33_______________________X Necryia_5966_93))) Methone_8523_95) Napaea_m17_89_______________________________)) Theope_7789_65__________________ANTS__O_o__o)) Theope_9026_5____________________ANTS__O_o__o) Melanis_7612_30_________________________X______) Ancyluris_6834_45)) (((((Adelotypa_7807_37_________________ANTS__O_o__o Cyrenia_3992_63) (((((Chamaelimnas_3942_57 Ourocnemis_4621_90___________________________) Lepricornis_5041_80) (Metacharis_9347_23__________________________ Phaenochitonia_6926_47)) (Esthemopsis_7813_32 Apodemia_mormo)) Ithomeis_7773_35)) (Eurybia_9363_20_________________ANTS__O_o__o Calephelis_9356_22____________________X______)) Parcella_4012_64) Detritivora_9327_25)) Semomesia_5223_52_________________________X) (((((((Setabis_6330_38_________________ANTS__O_o__o ((Helicopis_5318_54 Panara_3932_56) Leucochimona_9306_27_______________________X)) Riodina_lysippus) (Mesene_7450_28_________________________X______ Mesophthalma_6327_88_____________________X)) Leucochimona_7617_87_______________________X) Thisbe_6766_43___________________ANTS__O_o__o) T_Poritia_erycinoides) Rhetus_3973_60)) Cremna_5260_53) Calydna_6346_83) (Argyrogrammana_6709_42 Sarota_9358_21)) Thysanota_5241_81_________________ANTS__O_o__o) (((Charis_9365_19_______________________________ Euselasia_8816_01_________________________X) (Hyphilaria_5239_91 (Pachythone_9050_18 Symmachia_8849_08___________________________))) ((Emesis_6837_46_______________________________ (Monethe_3977_61 Themone_7643_34)) Anteros_7097_50______________________________))) Mesosemia_6516_85_________________________X) (Menander_7339_49_________________ANTS__O_o__o Emesis_mandana___________________________X_____)) ((Juditha_9315_26__________________ANTS__O_o__o Crocozona_3995_62) ((Brachyglemis_3956_58 T_Miletus_ancon) Exoplisia_6433_40)))*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o (Hyphilaria_5239_91 Styx_infernalis)) Ithomeis_7773_35) (Panara_3932_56 Euselasia_orfita____________________________X)) (Euselasia_8816_01_________________________X Melanis_7612_30_________________________X______)) (((((Eurybia_9363_20_________________ANTS__O_o__o Ancyluris_6834_45) (Calephelis_9356_22____________________X______ Hamearis_lucina)) ((Thysanota_5241_81_________________ANTS__O_o__o ((Calydna_6346_83 Mesosemia_8822_02_________________________X) Cyrenia_3992_63)) Metacharis_9347_23__________________________)) Argyrogrammana_6709_42) (Cariomothis_6518_41 Parcella_4012_64))) ((((((((((Protonymphidia_3948_59_________ANTS__O_o__o Ithomiola_5924_82) Helicopis_5318_54) Zelotea_4057_79) Alesa_8380_84____________________ANTS__O_o__o) Calospila_8532_96_________________ANTS__O_o__o) Riodina_lysippus) Mesosemia_6516_85_________________________X) ((((((Theope_7789_65__________________ANTS__O_o__o Brachyglemis_3956_58) Chamaelimnas_3942_57) Leucochimona_7617_87_______________________X) (Thisbe_6766_43___________________ANTS__O_o__o Ourocnemis_4621_90___________________________)) ((((Menander_7339_49_________________ANTS__O_o__o Echenais_7168_48) Exoplisia_6433_40) Detritivora_9327_25) Juditha_9315_26__________________ANTS__O_o__o)) Leucochimona_9306_27_______________________X)) Caria_7625_31_______________________________) Leucochimona_9302_33_______________________X)) (Lepricornis_5041_80 Apodemia_mormo)) (Charis_9365_19_______________________________ (((((((Cremna_5260_53 Stalachtis_6422_39) Anteros_7097_50______________________________) Semomesia_5223_52_________________________X) Mesene_7450_28_________________________X______) Monethe_3977_61) Sarota_9358_21) Themone_7643_34))) (Adelotypa_7807_37_________________ANTS__O_o__o Pandemos_8572_97)) (Theope_9026_5____________________ANTS__O_o__o T_Miletus_ancon)) (((((Lemonias_3950_55_________________ANTS__O_o__o Phaenochitonia_6926_47) Roeberella_7457_36) Perophthalma_9330_24_______________________X) Necryia_5966_93) Rhetus_3973_60)) (Synargis_7605_29_________________ANTS__O_o__o (Amarynthis_5220_51 Pachythone_9050_18))) Napaea_m17_89_______________________________) Setabis_6330_38_________________ANTS__O_o__o) (Crocozona_3995_62 Symmachia_8849_08___________________________)) (Emesis_6837_46_______________________________ (((Esthemopsis_7813_32 (Mesophthalma_6327_88_____________________X Emesis_mandana___________________________X_____)) (Methone_8523_95 T_Poritia_erycinoides)) Isapis_6815_44)))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Pandemos_8572_97) Roeberella_7457_36) Adelotypa_7807_37_________________ANTS__O_o__o) (Theope_7789_65__________________ANTS__O_o__o (Helicopis_5318_54 Ithomiola_5924_82))) (((((Cariomothis_6518_41 (Emesis_6837_46_______________________________ Zelotea_4057_79)) Chamaelimnas_3942_57) Hyphilaria_5239_91) Methone_8523_95) Pachythone_9050_18)) Detritivora_9327_25) Euselasia_8816_01_________________________X) (((((((((Protonymphidia_3948_59_________ANTS__O_o__o (Themone_7643_34 T_Miletus_ancon)) Caria_7625_31_______________________________) ((((Eurybia_9363_20_________________ANTS__O_o__o ((((((Charis_9365_19_______________________________ Anteros_7097_50______________________________) (Mesophthalma_6327_88_____________________X Symmachia_8849_08___________________________)) Apodemia_mormo) Parcella_4012_64) Exoplisia_6433_40) T_Poritia_erycinoides)) Sarota_9358_21) Mesosemia_8822_02_________________________X) Cyrenia_3992_63)) Necryia_5966_93) Cremna_5260_53) Panara_3932_56) (((((((Alesa_8380_84____________________ANTS__O_o__o ((Setabis_6330_38_________________ANTS__O_o__o Lepricornis_5041_80) Calydna_6346_83)) Leucochimona_9306_27_______________________X) Lemonias_3950_55_________________ANTS__O_o__o) Mesene_7450_28_________________________X______) Ancyluris_6834_45) Synargis_7605_29_________________ANTS__O_o__o) (Brachyglemis_3956_58 Echenais_7168_48))) Mesosemia_6516_85_________________________X) ((Juditha_9315_26__________________ANTS__O_o__o Metacharis_9347_23__________________________) (Leucochimona_7617_87_______________________X Phaenochitonia_6926_47)))) ((Theope_9026_5____________________ANTS__O_o__o (((((((Menander_7339_49_________________ANTS__O_o__o ((((Esthemopsis_7813_32 Rhetus_3973_60) (((Isapis_6815_44 (Perophthalma_9330_24_______________________X Styx_infernalis)) Leucochimona_9302_33_______________________X) Ourocnemis_4621_90___________________________)) (Ithomeis_7773_35 Emesis_mandana___________________________X_____)) Melanis_7612_30_________________________X______)) Amarynthis_5220_51) Monethe_3977_61) Crocozona_3995_62) Thysanota_5241_81_________________ANTS__O_o__o) Stalachtis_6422_39) Euselasia_orfita____________________________X)) (Calospila_8532_96_________________ANTS__O_o__o Napaea_m17_89_______________________________))) Thisbe_6766_43___________________ANTS__O_o__o) Semomesia_5223_52_________________________X) (Calephelis_9356_22____________________X______ Riodina_lysippus)) (Argyrogrammana_6709_42 Hamearis_lucina))*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((Uraneis_hyalina____________________ANTS__O_o__o Detritivora_9327_25) Leucochimona_9302_33_______________________X) Calospila_8532_96_________________ANTS__O_o__o) Alesa_8380_84____________________ANTS__O_o__o) Necryia_5966_93) ((((Lemonias_3950_55_________________ANTS__O_o__o Leucochimona_9306_27_______________________X) (Argyrogrammana_6709_42 Calydna_6346_83)) (((Juditha_9315_26__________________ANTS__O_o__o Monethe_3977_61) Phaenochitonia_6926_47) Sarota_9358_21)) Ourocnemis_4621_90___________________________)) Theope_9026_5____________________ANTS__O_o__o) ((Charis_9365_19_______________________________ (((Napaea_m17_89_______________________________ Emesis_mandana___________________________X_____) Styx_infernalis) Apodemia_mormo)) Mesophthalma_6327_88_____________________X)) (((((((Adelotypa_7807_37_________________ANTS__O_o__o (Hyphilaria_5239_91 Panara_3932_56)) (Protonymphidia_3948_59_________ANTS__O_o__o Thisbe_6766_43___________________ANTS__O_o__o)) Calephelis_9356_22____________________X______) (Emesis_6837_46_______________________________ T_Miletus_ancon)) ((((Setabis_6330_38_________________ANTS__O_o__o (((((Echenais_7168_48 Isapis_6815_44) Stalachtis_6422_39) T_Poritia_erycinoides) Exoplisia_6433_40) Ithomiola_5924_82)) Euselasia_orfita____________________________X) Caria_7625_31_______________________________) ((((((((((((Eurybia_9363_20_________________ANTS__O_o__o ((((Theope_7789_65__________________ANTS__O_o__o Metacharis_9347_23__________________________) ((Synargis_7605_29_________________ANTS__O_o__o Amarynthis_5220_51) (Lepricornis_5041_80 Symmachia_8849_08___________________________))) (Brachyglemis_3956_58 Cariomothis_6518_41)) Thysanota_5241_81_________________ANTS__O_o__o)) (Mesosemia_6516_85_________________________X Rhetus_3973_60)) Parcella_4012_64) ((Ithomeis_7773_35 Roeberella_7457_36) (Mesosemia_8822_02_________________________X Zelotea_4057_79))) (Cremna_5260_53 (Cyrenia_3992_63 (((Pachythone_9050_18 Semomesia_5223_52_________________________X) Hamearis_lucina) Themone_7643_34)))) Mesene_7450_28_________________________X______) (Pandemos_8572_97 Riodina_lysippus)) ((Chamaelimnas_3942_57 Anteros_7097_50______________________________) Leucochimona_7617_87_______________________X)) Euselasia_8816_01_________________________X) (Helicopis_5318_54 Methone_8523_95)) Melanis_7612_30_________________________X______) Esthemopsis_7813_32))) Menander_7339_49_________________ANTS__O_o__o) Crocozona_3995_62)) (Ancyluris_6834_45 Perophthalma_9330_24_______________________X))*
(T_Curetis_bulis_______________________ANTS__O_o__o (Uraneis_hyalina____________________ANTS__O_o__o (((Synargis_7605_29_________________ANTS__O_o__o Monethe_3977_61) Lepricornis_5041_80) Melanis_7612_30_________________________X______)) (((Adelotypa_7807_37_________________ANTS__O_o__o Styx_infernalis) Cariomothis_6518_41) (((((Protonymphidia_3948_59_________ANTS__O_o__o (((((((((((((((((((Theope_7789_65__________________ANTS__O_o__o Theope_9026_5____________________ANTS__O_o__o) (Cremna_5260_53 Isapis_6815_44)) Exoplisia_6433_40) Methone_8523_95) Necryia_5966_93) Stalachtis_6422_39) (((Menander_7339_49_________________ANTS__O_o__o T_Miletus_ancon) Metacharis_9347_23__________________________) Apodemia_mormo)) ((Cyrenia_3992_63 Esthemopsis_7813_32) Zelotea_4057_79)) Symmachia_8849_08___________________________) Pachythone_9050_18) Hamearis_lucina) Amarynthis_5220_51) Charis_9365_19_______________________________) Themone_7643_34) Euselasia_8816_01_________________________X) Helicopis_5318_54) (Leucochimona_7617_87_______________________X Mesosemia_6516_85_________________________X)) Ourocnemis_4621_90___________________________) Echenais_7168_48)) ((((Alesa_8380_84____________________ANTS__O_o__o Mesene_7450_28_________________________X______) (((((((((Setabis_6330_38_________________ANTS__O_o__o (((Argyrogrammana_6709_42 Riodina_lysippus) Hyphilaria_5239_91) Mesophthalma_6327_88_____________________X)) (Calephelis_9356_22____________________X______ ((((Caria_7625_31_______________________________ Emesis_6837_46_______________________________) Pandemos_8572_97) T_Poritia_erycinoides) Napaea_m17_89_______________________________))) Ithomiola_5924_82) ((Calospila_8532_96_________________ANTS__O_o__o Ithomeis_7773_35) Crocozona_3995_62)) (Thisbe_6766_43___________________ANTS__O_o__o Parcella_4012_64)) (Mesosemia_8822_02_________________________X Euselasia_orfita____________________________X)) (Detritivora_9327_25 (Leucochimona_9306_27_______________________X Panara_3932_56))) Calydna_6346_83) (Thysanota_5241_81_________________ANTS__O_o__o Sarota_9358_21))) (((Juditha_9315_26__________________ANTS__O_o__o ((Chamaelimnas_3942_57 Roeberella_7457_36) Emesis_mandana___________________________X_____)) Ancyluris_6834_45) Phaenochitonia_6926_47)) (((Eurybia_9363_20_________________ANTS__O_o__o Semomesia_5223_52_________________________X) Lemonias_3950_55_________________ANTS__O_o__o) Leucochimona_9302_33_______________________X))) Rhetus_3973_60) (Perophthalma_9330_24_______________________X Anteros_7097_50______________________________)) Brachyglemis_3956_58)))*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o (Juditha_9315_26__________________ANTS__O_o__o (Argyrogrammana_6709_42 Roeberella_7457_36))) (((Adelotypa_7807_37_________________ANTS__O_o__o (((((Setabis_6330_38_________________ANTS__O_o__o Semomesia_5223_52_________________________X) Emesis_6837_46_______________________________) (Brachyglemis_3956_58 Pachythone_9050_18)) ((Thisbe_6766_43___________________ANTS__O_o__o Styx_infernalis) (Cariomothis_6518_41 Emesis_mandana___________________________X_____))) Calydna_6346_83)) Calephelis_9356_22____________________X______) Isapis_6815_44)) (Cremna_5260_53 ((Mesene_7450_28_________________________X______ Ourocnemis_4621_90___________________________) Riodina_lysippus))) (Menander_7339_49_________________ANTS__O_o__o (Thysanota_5241_81_________________ANTS__O_o__o Ithomeis_7773_35))) Hyphilaria_5239_91) Eurybia_9363_20_________________ANTS__O_o__o) ((((Crocozona_3995_62 Perophthalma_9330_24_______________________X) T_Poritia_erycinoides) Mesosemia_6516_85_________________________X) Monethe_3977_61)) Parcella_4012_64) (((Theope_9026_5____________________ANTS__O_o__o Euselasia_8816_01_________________________X) (Ancyluris_6834_45 Panara_3932_56)) (((Chamaelimnas_3942_57 Necryia_5966_93) Leucochimona_7617_87_______________________X) Leucochimona_9306_27_______________________X))) ((((((Alesa_8380_84____________________ANTS__O_o__o Euselasia_orfita____________________________X) ((Theope_7789_65__________________ANTS__O_o__o (Detritivora_9327_25 (Methone_8523_95 Napaea_m17_89_______________________________))) Cyrenia_3992_63)) Synargis_7605_29_________________ANTS__O_o__o) ((Charis_9365_19_______________________________ Leucochimona_9302_33_______________________X) Mesosemia_8822_02_________________________X)) ((Phaenochitonia_6926_47 Rhetus_3973_60) Symmachia_8849_08___________________________)) (Echenais_7168_48 T_Miletus_ancon))) Pandemos_8572_97) ((Calospila_8532_96_________________ANTS__O_o__o Lepricornis_5041_80) Anteros_7097_50______________________________)) Metacharis_9347_23__________________________) Helicopis_5318_54) Sarota_9358_21) ((Protonymphidia_3948_59_________ANTS__O_o__o Caria_7625_31_______________________________) ((Esthemopsis_7813_32 Stalachtis_6422_39) ((Mesophthalma_6327_88_____________________X Hamearis_lucina) Apodemia_mormo)))) (Amarynthis_5220_51 Themone_7643_34)) ((Lemonias_3950_55_________________ANTS__O_o__o (Ithomiola_5924_82 (Melanis_7612_30_________________________X______ Zelotea_4057_79))) Exoplisia_6433_40))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o (((((((Adelotypa_7807_37_________________ANTS__O_o__o ((Cremna_5260_53 Apodemia_mormo) ((Leucochimona_9302_33_______________________X T_Miletus_ancon) Monethe_3977_61))) (Theope_9026_5____________________ANTS__O_o__o ((Calospila_8532_96_________________ANTS__O_o__o Methone_8523_95) Pachythone_9050_18))) (((((Setabis_6330_38_________________ANTS__O_o__o (Isapis_6815_44 Mesene_7450_28_________________________X______)) Zelotea_4057_79) ((Juditha_9315_26__________________ANTS__O_o__o Lepricornis_5041_80) Riodina_lysippus)) Echenais_7168_48) Lemonias_3950_55_________________ANTS__O_o__o)) Rhetus_3973_60) ((Alesa_8380_84____________________ANTS__O_o__o Styx_infernalis) Exoplisia_6433_40)) (Esthemopsis_7813_32 Semomesia_5223_52_________________________X)) ((Perophthalma_9330_24_______________________X Euselasia_orfita____________________________X) Themone_7643_34))) (Synargis_7605_29_________________ANTS__O_o__o (Cyrenia_3992_63 Leucochimona_9306_27_______________________X))) Emesis_mandana___________________________X_____) Eurybia_9363_20_________________ANTS__O_o__o) Crocozona_3995_62) Detritivora_9327_25) (Euselasia_8816_01_________________________X Pandemos_8572_97)) Ancyluris_6834_45) (Cariomothis_6518_41 (Emesis_6837_46_______________________________ Leucochimona_7617_87_______________________X))) Caria_7625_31_______________________________) ((((((((((Menander_7339_49_________________ANTS__O_o__o Phaenochitonia_6926_47) Mesophthalma_6327_88_____________________X) ((Ithomiola_5924_82 Roeberella_7457_36) Metacharis_9347_23__________________________)) Melanis_7612_30_________________________X______) (Argyrogrammana_6709_42 Mesosemia_8822_02_________________________X)) (Chamaelimnas_3942_57 Charis_9365_19_______________________________)) (Panara_3932_56 Symmachia_8849_08___________________________)) T_Poritia_erycinoides) (Calydna_6346_83 Necryia_5966_93)) Mesosemia_6516_85_________________________X)) Hyphilaria_5239_91) Ithomeis_7773_35) (((((((Protonymphidia_3948_59_________ANTS__O_o__o Stalachtis_6422_39) Calephelis_9356_22____________________X______) Parcella_4012_64) Amarynthis_5220_51) Brachyglemis_3956_58) Sarota_9358_21) (Thisbe_6766_43___________________ANTS__O_o__o Ourocnemis_4621_90___________________________))) (Theope_7789_65__________________ANTS__O_o__o Hamearis_lucina)) (Helicopis_5318_54 Napaea_m17_89_______________________________)) (Thysanota_5241_81_________________ANTS__O_o__o Anteros_7097_50______________________________))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((Uraneis_hyalina____________________ANTS__O_o__o Calydna_6346_83) (((Protonymphidia_3948_59_________ANTS__O_o__o Helicopis_5318_54) Theope_9026_5____________________ANTS__O_o__o) Symmachia_8849_08___________________________)) ((((((((Adelotypa_7807_37_________________ANTS__O_o__o (((((((((((Eurybia_9363_20_________________ANTS__O_o__o ((Melanis_7612_30_________________________X______ Pandemos_8572_97) (Semomesia_5223_52_________________________X Euselasia_orfita____________________________X))) (Lemonias_3950_55_________________ANTS__O_o__o Napaea_m17_89_______________________________)) Metacharis_9347_23__________________________) Mesosemia_8822_02_________________________X) Phaenochitonia_6926_47) (Roeberella_7457_36 Apodemia_mormo)) Sarota_9358_21) (Necryia_5966_93 T_Miletus_ancon)) Amarynthis_5220_51) Anteros_7097_50______________________________) (Ancyluris_6834_45 Chamaelimnas_3942_57))) Methone_8523_95) Perophthalma_9330_24_______________________X) ((((((((Synargis_7605_29_________________ANTS__O_o__o Hamearis_lucina) Mesosemia_6516_85_________________________X) Stalachtis_6422_39) (Argyrogrammana_6709_42 Mesophthalma_6327_88_____________________X)) ((Brachyglemis_3956_58 ((Detritivora_9327_25 Pachythone_9050_18) Leucochimona_9306_27_______________________X)) Styx_infernalis)) (Cyrenia_3992_63 Leucochimona_9302_33_______________________X)) (Euselasia_8816_01_________________________X Ithomeis_7773_35)) (Calephelis_9356_22____________________X______ Ourocnemis_4621_90___________________________))) Panara_3932_56) Calospila_8532_96_________________ANTS__O_o__o) ((Caria_7625_31_______________________________ Cremna_5260_53) ((Charis_9365_19_______________________________ Monethe_3977_61) Lepricornis_5041_80))) Exoplisia_6433_40)) ((Alesa_8380_84____________________ANTS__O_o__o ((((((((((Setabis_6330_38_________________ANTS__O_o__o Riodina_lysippus) Parcella_4012_64) Emesis_mandana___________________________X_____) Esthemopsis_7813_32) ((Cariomothis_6518_41 Ithomiola_5924_82) Crocozona_3995_62)) Isapis_6815_44) Leucochimona_7617_87_______________________X) Thysanota_5241_81_________________ANTS__O_o__o) Rhetus_3973_60) (Emesis_6837_46_______________________________ Mesene_7450_28_________________________X______))) Zelotea_4057_79)) Echenais_7168_48) ((Theope_7789_65__________________ANTS__O_o__o ((Juditha_9315_26__________________ANTS__O_o__o Hyphilaria_5239_91) T_Poritia_erycinoides)) (Thisbe_6766_43___________________ANTS__O_o__o Themone_7643_34))) Menander_7339_49_________________ANTS__O_o__o)*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o ((Isapis_6815_44 ((Ithomeis_7773_35 Symmachia_8849_08___________________________) Leucochimona_9306_27_______________________X)) Lepricornis_5041_80)) Brachyglemis_3956_58) (Amarynthis_5220_51 Necryia_5966_93)) Theope_7789_65__________________ANTS__O_o__o) Styx_infernalis) Mesosemia_8822_02_________________________X) Pandemos_8572_97) (Argyrogrammana_6709_42 ((Monethe_3977_61 Emesis_mandana___________________________X_____) Euselasia_orfita____________________________X))) Thisbe_6766_43___________________ANTS__O_o__o) Detritivora_9327_25) (Calydna_6346_83 Leucochimona_7617_87_______________________X)) Rhetus_3973_60) (((((Protonymphidia_3948_59_________ANTS__O_o__o (Juditha_9315_26__________________ANTS__O_o__o ((Synargis_7605_29_________________ANTS__O_o__o (Emesis_6837_46_______________________________ ((Perophthalma_9330_24_______________________X Themone_7643_34) Phaenochitonia_6926_47))) Parcella_4012_64))) ((((((Alesa_8380_84____________________ANTS__O_o__o ((Cremna_5260_53 (Hyphilaria_5239_91 T_Poritia_erycinoides)) ((Echenais_7168_48 Hamearis_lucina) Semomesia_5223_52_________________________X))) Helicopis_5318_54) Mesene_7450_28_________________________X______) Eurybia_9363_20_________________ANTS__O_o__o) (Cyrenia_3992_63 Pachythone_9050_18)) Ancyluris_6834_45)) (((((Crocozona_3995_62 Zelotea_4057_79) Panara_3932_56) Exoplisia_6433_40) Sarota_9358_21) Ithomiola_5924_82)) Chamaelimnas_3942_57) ((Caria_7625_31_______________________________ (Roeberella_7457_36 Stalachtis_6422_39)) Charis_9365_19_______________________________))) ((Esthemopsis_7813_32 Methone_8523_95) ((Melanis_7612_30_________________________X______ Anteros_7097_50______________________________) Mesophthalma_6327_88_____________________X))) Theope_9026_5____________________ANTS__O_o__o) (Menander_7339_49_________________ANTS__O_o__o Thysanota_5241_81_________________ANTS__O_o__o)) ((Adelotypa_7807_37_________________ANTS__O_o__o ((Setabis_6330_38_________________ANTS__O_o__o (Calephelis_9356_22____________________X______ Napaea_m17_89_______________________________)) Metacharis_9347_23__________________________)) Mesosemia_6516_85_________________________X)) ((Calospila_8532_96_________________ANTS__O_o__o Ourocnemis_4621_90___________________________) T_Miletus_ancon)) Leucochimona_9302_33_______________________X) Cariomothis_6518_41) ((Lemonias_3950_55_________________ANTS__O_o__o (Euselasia_8816_01_________________________X Riodina_lysippus)) Apodemia_mormo))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o (Setabis_6330_38_________________ANTS__O_o__o Echenais_7168_48)) Mesophthalma_6327_88_____________________X) (((((Charis_9365_19_______________________________ Mesene_7450_28_________________________X______) Styx_infernalis) Pachythone_9050_18) Helicopis_5318_54) Hyphilaria_5239_91)) Alesa_8380_84____________________ANTS__O_o__o) Brachyglemis_3956_58) (((((((((Protonymphidia_3948_59_________ANTS__O_o__o Perophthalma_9330_24_______________________X) Theope_9026_5____________________ANTS__O_o__o) (((((Calospila_8532_96_________________ANTS__O_o__o Ithomeis_7773_35) ((Synargis_7605_29_________________ANTS__O_o__o Mesosemia_6516_85_________________________X) Ancyluris_6834_45)) Panara_3932_56) Leucochimona_9306_27_______________________X) Euselasia_orfita____________________________X)) Metacharis_9347_23__________________________) Lemonias_3950_55_________________ANTS__O_o__o) (Exoplisia_6433_40 Methone_8523_95)) Calydna_6346_83) ((Emesis_6837_46_______________________________ Leucochimona_9302_33_______________________X) (Monethe_3977_61 Themone_7643_34))) Riodina_lysippus)) (Cremna_5260_53 (Leucochimona_7617_87_______________________X (Necryia_5966_93 Stalachtis_6422_39)))) ((((Theope_7789_65__________________ANTS__O_o__o Rhetus_3973_60) (((((Menander_7339_49_________________ANTS__O_o__o Euselasia_8816_01_________________________X) Isapis_6815_44) Mesosemia_8822_02_________________________X) Hamearis_lucina) Roeberella_7457_36)) Caria_7625_31_______________________________) Emesis_mandana___________________________X_____)) Ithomiola_5924_82) Symmachia_8849_08___________________________) Esthemopsis_7813_32) (Cariomothis_6518_41 (Chamaelimnas_3942_57 Ourocnemis_4621_90___________________________))) Eurybia_9363_20_________________ANTS__O_o__o) (((((Adelotypa_7807_37_________________ANTS__O_o__o Pandemos_8572_97) (((Juditha_9315_26__________________ANTS__O_o__o Argyrogrammana_6709_42) Melanis_7612_30_________________________X______) Phaenochitonia_6926_47)) (((Thysanota_5241_81_________________ANTS__O_o__o Calephelis_9356_22____________________X______) Cyrenia_3992_63) Anteros_7097_50______________________________)) ((Amarynthis_5220_51 (Zelotea_4057_79 T_Poritia_erycinoides)) Detritivora_9327_25)) ((((Lepricornis_5041_80 Parcella_4012_64) Semomesia_5223_52_________________________X) Sarota_9358_21) T_Miletus_ancon))) Crocozona_3995_62) (Thisbe_6766_43___________________ANTS__O_o__o Napaea_m17_89_______________________________)) Apodemia_mormo)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o T_Miletus_ancon) ((Amarynthis_5220_51 Monethe_3977_61) ((((Hyphilaria_5239_91 Mesosemia_6516_85_________________________X) Mesophthalma_6327_88_____________________X) Perophthalma_9330_24_______________________X) Necryia_5966_93))) Helicopis_5318_54) Styx_infernalis) Parcella_4012_64) Theope_7789_65__________________ANTS__O_o__o) Cyrenia_3992_63) Emesis_6837_46_______________________________) Panara_3932_56) Cremna_5260_53) (Calospila_8532_96_________________ANTS__O_o__o Leucochimona_7617_87_______________________X)) (((Alesa_8380_84____________________ANTS__O_o__o (Caria_7625_31_______________________________ Euselasia_8816_01_________________________X)) Symmachia_8849_08___________________________) Calephelis_9356_22____________________X______)) Pandemos_8572_97) (((((((((((((Protonymphidia_3948_59_________ANTS__O_o__o (Juditha_9315_26__________________ANTS__O_o__o (Thisbe_6766_43___________________ANTS__O_o__o Chamaelimnas_3942_57))) ((Phaenochitonia_6926_47 T_Poritia_erycinoides) Themone_7643_34)) Detritivora_9327_25) Charis_9365_19_______________________________) ((((Thysanota_5241_81_________________ANTS__O_o__o Leucochimona_9302_33_______________________X) (Stalachtis_6422_39 Riodina_lysippus)) (Lepricornis_5041_80 Mesosemia_8822_02_________________________X)) Cariomothis_6518_41)) Isapis_6815_44) Napaea_m17_89_______________________________) Ithomiola_5924_82) Methone_8523_95) Semomesia_5223_52_________________________X) Metacharis_9347_23__________________________) Euselasia_orfita____________________________X) Ancyluris_6834_45)) (((Theope_9026_5____________________ANTS__O_o__o Anteros_7097_50______________________________) Ourocnemis_4621_90___________________________) Crocozona_3995_62)) ((((((((Adelotypa_7807_37_________________ANTS__O_o__o Roeberella_7457_36) Setabis_6330_38_________________ANTS__O_o__o) ((Synargis_7605_29_________________ANTS__O_o__o (Echenais_7168_48 Esthemopsis_7813_32)) ((Calydna_6346_83 Leucochimona_9306_27_______________________X) Sarota_9358_21))) ((Argyrogrammana_6709_42 Emesis_mandana___________________________X_____) (Zelotea_4057_79 Apodemia_mormo))) Brachyglemis_3956_58) Melanis_7612_30_________________________X______) Mesene_7450_28_________________________X______) (Exoplisia_6433_40 Ithomeis_7773_35))) (Menander_7339_49_________________ANTS__O_o__o Pachythone_9050_18)) (Eurybia_9363_20_________________ANTS__O_o__o Lemonias_3950_55_________________ANTS__O_o__o)) Hamearis_lucina) Rhetus_3973_60)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o ((Adelotypa_7807_37_________________ANTS__O_o__o (((Theope_7789_65__________________ANTS__O_o__o Symmachia_8849_08___________________________) Calephelis_9356_22____________________X______) Ourocnemis_4621_90___________________________)) Styx_infernalis)) Ithomeis_7773_35) (Thysanota_5241_81_________________ANTS__O_o__o Perophthalma_9330_24_______________________X)) (Cyrenia_3992_63 Mesosemia_8822_02_________________________X)) Cremna_5260_53) ((Synargis_7605_29_________________ANTS__O_o__o Helicopis_5318_54) (Mesophthalma_6327_88_____________________X Euselasia_orfita____________________________X))) Leucochimona_9306_27_______________________X) (((((Alesa_8380_84____________________ANTS__O_o__o Pandemos_8572_97) Emesis_6837_46_______________________________) Leucochimona_7617_87_______________________X) (Necryia_5966_93 Anteros_7097_50______________________________)) Zelotea_4057_79)) Exoplisia_6433_40) Thisbe_6766_43___________________ANTS__O_o__o) Setabis_6330_38_________________ANTS__O_o__o) Parcella_4012_64) Theope_9026_5____________________ANTS__O_o__o) T_Miletus_ancon) (((Amarynthis_5220_51 Semomesia_5223_52_________________________X) ((Argyrogrammana_6709_42 Esthemopsis_7813_32) (Cariomothis_6518_41 Emesis_mandana___________________________X_____))) Hamearis_lucina)) (Protonymphidia_3948_59_________ANTS__O_o__o Methone_8523_95)) ((Juditha_9315_26__________________ANTS__O_o__o Themone_7643_34) Calydna_6346_83)) ((((Calospila_8532_96_________________ANTS__O_o__o Euselasia_8816_01_________________________X) Lepricornis_5041_80) Panara_3932_56) Metacharis_9347_23__________________________)) Echenais_7168_48) ((((Menander_7339_49_________________ANTS__O_o__o Ancyluris_6834_45) Chamaelimnas_3942_57) (Detritivora_9327_25 Napaea_m17_89_______________________________)) Roeberella_7457_36)) (Crocozona_3995_62 Rhetus_3973_60)) Caria_7625_31_______________________________) ((Leucochimona_9302_33_______________________X Riodina_lysippus) Mesosemia_6516_85_________________________X)) ((((Lemonias_3950_55_________________ANTS__O_o__o Brachyglemis_3956_58) Ithomiola_5924_82) (Monethe_3977_61 Phaenochitonia_6926_47)) Mesene_7450_28_________________________X______)) (Eurybia_9363_20_________________ANTS__O_o__o (Sarota_9358_21 Stalachtis_6422_39))) Apodemia_mormo) (((Charis_9365_19_______________________________ (Isapis_6815_44 T_Poritia_erycinoides)) Melanis_7612_30_________________________X______) Pachythone_9050_18)) Hyphilaria_5239_91)*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Hyphilaria_5239_91) Theope_9026_5____________________ANTS__O_o__o) (((((Juditha_9315_26__________________ANTS__O_o__o Emesis_6837_46_______________________________) Panara_3932_56) ((Ancyluris_6834_45 (Pachythone_9050_18 T_Poritia_erycinoides)) Detritivora_9327_25)) Melanis_7612_30_________________________X______) (Cremna_5260_53 (Roeberella_7457_36 Symmachia_8849_08___________________________)))) Isapis_6815_44) (Phaenochitonia_6926_47 Styx_infernalis)) Anteros_7097_50______________________________) Caria_7625_31_______________________________) Mesosemia_8822_02_________________________X) Apodemia_mormo) Chamaelimnas_3942_57) Euselasia_orfita____________________________X) Adelotypa_7807_37_________________ANTS__O_o__o) Leucochimona_9302_33_______________________X) (Theope_7789_65__________________ANTS__O_o__o Emesis_mandana___________________________X_____)) (((Amarynthis_5220_51 Ourocnemis_4621_90___________________________) Necryia_5966_93) Calephelis_9356_22____________________X______)) Pandemos_8572_97) Lepricornis_5041_80) (Hamearis_lucina T_Miletus_ancon)) (Brachyglemis_3956_58 Zelotea_4057_79)) (((Alesa_8380_84____________________ANTS__O_o__o (Argyrogrammana_6709_42 (((Calydna_6346_83 (Perophthalma_9330_24_______________________X Themone_7643_34)) Crocozona_3995_62) (Leucochimona_7617_87_______________________X Parcella_4012_64)))) (Esthemopsis_7813_32 Napaea_m17_89_______________________________)) Sarota_9358_21)) Semomesia_5223_52_________________________X) ((Setabis_6330_38_________________ANTS__O_o__o Monethe_3977_61) Menander_7339_49_________________ANTS__O_o__o)) (((((((Protonymphidia_3948_59_________ANTS__O_o__o Calospila_8532_96_________________ANTS__O_o__o) Riodina_lysippus) Mesene_7450_28_________________________X______) (Exoplisia_6433_40 Leucochimona_9306_27_______________________X)) Methone_8523_95) (Cyrenia_3992_63 Rhetus_3973_60)) (((((((Eurybia_9363_20_________________ANTS__O_o__o Charis_9365_19_______________________________) ((Lemonias_3950_55_________________ANTS__O_o__o Synargis_7605_29_________________ANTS__O_o__o) Echenais_7168_48)) (((Helicopis_5318_54 Mesophthalma_6327_88_____________________X) (Ithomiola_5924_82 Mesosemia_6516_85_________________________X)) Metacharis_9347_23__________________________)) (Ithomeis_7773_35 Stalachtis_6422_39)) Euselasia_8816_01_________________________X) Cariomothis_6518_41) (Thisbe_6766_43___________________ANTS__O_o__o Thysanota_5241_81_________________ANTS__O_o__o))))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o ((Theope_9026_5____________________ANTS__O_o__o (((Caria_7625_31_______________________________ Roeberella_7457_36) Symmachia_8849_08___________________________) Cariomothis_6518_41)) Cyrenia_3992_63)) Argyrogrammana_6709_42) Setabis_6330_38_________________ANTS__O_o__o) (Detritivora_9327_25 Melanis_7612_30_________________________X______)) T_Poritia_erycinoides) (Eurybia_9363_20_________________ANTS__O_o__o Crocozona_3995_62)) Pandemos_8572_97) (((((((((((Protonymphidia_3948_59_________ANTS__O_o__o Pachythone_9050_18) Leucochimona_9302_33_______________________X) ((Lemonias_3950_55_________________ANTS__O_o__o Sarota_9358_21) Zelotea_4057_79)) Leucochimona_9306_27_______________________X) Thisbe_6766_43___________________ANTS__O_o__o) Juditha_9315_26__________________ANTS__O_o__o) Brachyglemis_3956_58) Monethe_3977_61) Ancyluris_6834_45) (Mesene_7450_28_________________________X______ Necryia_5966_93)) ((((Alesa_8380_84____________________ANTS__O_o__o Rhetus_3973_60) (((Chamaelimnas_3942_57 Emesis_6837_46_______________________________) Styx_infernalis) ((((Helicopis_5318_54 Leucochimona_7617_87_______________________X) Lepricornis_5041_80) Hyphilaria_5239_91) Phaenochitonia_6926_47))) Methone_8523_95) Napaea_m17_89_______________________________))) ((Thysanota_5241_81_________________ANTS__O_o__o Semomesia_5223_52_________________________X) Anteros_7097_50______________________________)) (Synargis_7605_29_________________ANTS__O_o__o Exoplisia_6433_40)) (Adelotypa_7807_37_________________ANTS__O_o__o Panara_3932_56)) ((((Theope_7789_65__________________ANTS__O_o__o Apodemia_mormo) Hamearis_lucina) Ithomiola_5924_82) Cremna_5260_53)) (((Calospila_8532_96_________________ANTS__O_o__o Riodina_lysippus) Calephelis_9356_22____________________X______) Ourocnemis_4621_90___________________________)) Metacharis_9347_23__________________________) (Charis_9365_19_______________________________ T_Miletus_ancon)) Euselasia_orfita____________________________X) Perophthalma_9330_24_______________________X) Themone_7643_34) Mesosemia_6516_85_________________________X) (Ithomeis_7773_35 Mesophthalma_6327_88_____________________X)) (Parcella_4012_64 Stalachtis_6422_39)) Euselasia_8816_01_________________________X) Menander_7339_49_________________ANTS__O_o__o) Amarynthis_5220_51) ((Calydna_6346_83 (Esthemopsis_7813_32 Isapis_6815_44)) Echenais_7168_48)) Mesosemia_8822_02_________________________X) Emesis_mandana___________________________X_____)*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((Uraneis_hyalina____________________ANTS__O_o__o (Calospila_8532_96_________________ANTS__O_o__o ((Thisbe_6766_43___________________ANTS__O_o__o Melanis_7612_30_________________________X______) Mesene_7450_28_________________________X______))) (((((((((((((((((((Protonymphidia_3948_59_________ANTS__O_o__o (Cariomothis_6518_41 Mesosemia_8822_02_________________________X)) (Exoplisia_6433_40 Sarota_9358_21)) (Theope_7789_65__________________ANTS__O_o__o (Detritivora_9327_25 Pachythone_9050_18))) Hyphilaria_5239_91) ((Menander_7339_49_________________ANTS__O_o__o Pandemos_8572_97) ((Leucochimona_7617_87_______________________X Parcella_4012_64) Apodemia_mormo))) ((Echenais_7168_48 Emesis_mandana___________________________X_____) (((Helicopis_5318_54 Styx_infernalis) Anteros_7097_50______________________________) Symmachia_8849_08___________________________))) (Calephelis_9356_22____________________X______ Zelotea_4057_79)) ((Setabis_6330_38_________________ANTS__O_o__o Chamaelimnas_3942_57) Cremna_5260_53)) Charis_9365_19_______________________________) Synargis_7605_29_________________ANTS__O_o__o) Argyrogrammana_6709_42) Euselasia_8816_01_________________________X) Juditha_9315_26__________________ANTS__O_o__o) ((Eurybia_9363_20_________________ANTS__O_o__o Methone_8523_95) (Napaea_m17_89_______________________________ Themone_7643_34))) ((((((Calydna_6346_83 Roeberella_7457_36) Emesis_6837_46_______________________________) Riodina_lysippus) Semomesia_5223_52_________________________X) Perophthalma_9330_24_______________________X) Isapis_6815_44)) Leucochimona_9302_33_______________________X) (Ancyluris_6834_45 Crocozona_3995_62)) Caria_7625_31_______________________________) (Lepricornis_5041_80 Necryia_5966_93))) Thysanota_5241_81_________________ANTS__O_o__o) Rhetus_3973_60) Euselasia_orfita____________________________X) (Stalachtis_6422_39 Hamearis_lucina)) Cyrenia_3992_63) (Metacharis_9347_23__________________________ Panara_3932_56)) ((((((((Adelotypa_7807_37_________________ANTS__O_o__o Monethe_3977_61) (((Alesa_8380_84____________________ANTS__O_o__o Mesosemia_6516_85_________________________X) (Lemonias_3950_55_________________ANTS__O_o__o Ourocnemis_4621_90___________________________)) (Ithomiola_5924_82 T_Miletus_ancon))) Mesophthalma_6327_88_____________________X) T_Poritia_erycinoides) Theope_9026_5____________________ANTS__O_o__o) Amarynthis_5220_51) (((Brachyglemis_3956_58 Phaenochitonia_6926_47) Esthemopsis_7813_32) Ithomeis_7773_35)) Leucochimona_9306_27_______________________X))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((Uraneis_hyalina____________________ANTS__O_o__o (((((Amarynthis_5220_51 Monethe_3977_61) ((Leucochimona_7617_87_______________________X Themone_7643_34) Panara_3932_56)) Cyrenia_3992_63) Leucochimona_9302_33_______________________X) Argyrogrammana_6709_42)) Riodina_lysippus) ((((((Adelotypa_7807_37_________________ANTS__O_o__o ((((((((Protonymphidia_3948_59_________ANTS__O_o__o (Lepricornis_5041_80 Roeberella_7457_36)) ((((Ithomeis_7773_35 Semomesia_5223_52_________________________X) Necryia_5966_93) (Mesosemia_8822_02_________________________X Parcella_4012_64)) T_Poritia_erycinoides)) Cariomothis_6518_41) Chamaelimnas_3942_57) (Pandemos_8572_97 Anteros_7097_50______________________________)) T_Miletus_ancon) (((((((Theope_7789_65__________________ANTS__O_o__o Calospila_8532_96_________________ANTS__O_o__o) Synargis_7605_29_________________ANTS__O_o__o) (Euselasia_8816_01_________________________X Melanis_7612_30_________________________X______)) Pachythone_9050_18) (Theope_9026_5____________________ANTS__O_o__o Euselasia_orfita____________________________X)) Mesosemia_6516_85_________________________X) Hamearis_lucina)) Stalachtis_6422_39)) Emesis_6837_46_______________________________) Rhetus_3973_60) Ancyluris_6834_45) (Caria_7625_31_______________________________ Crocozona_3995_62)) Napaea_m17_89_______________________________)) Eurybia_9363_20_________________ANTS__O_o__o) Detritivora_9327_25) ((((Thysanota_5241_81_________________ANTS__O_o__o ((Esthemopsis_7813_32 Metacharis_9347_23__________________________) Emesis_mandana___________________________X_____)) (Calydna_6346_83 Symmachia_8849_08___________________________)) Mesophthalma_6327_88_____________________X) Brachyglemis_3956_58)) (Echenais_7168_48 Leucochimona_9306_27_______________________X)) Calephelis_9356_22____________________X______) Cremna_5260_53) (Alesa_8380_84____________________ANTS__O_o__o Apodemia_mormo)) (Ourocnemis_4621_90___________________________ Sarota_9358_21)) Perophthalma_9330_24_______________________X) ((((((Setabis_6330_38_________________ANTS__O_o__o Menander_7339_49_________________ANTS__O_o__o) ((Exoplisia_6433_40 Phaenochitonia_6926_47) (Mesene_7450_28_________________________X______ Methone_8523_95))) Isapis_6815_44) ((((Juditha_9315_26__________________ANTS__O_o__o Hyphilaria_5239_91) (Helicopis_5318_54 Ithomiola_5924_82)) Thisbe_6766_43___________________ANTS__O_o__o) Zelotea_4057_79)) (Charis_9365_19_______________________________ Styx_infernalis)) Lemonias_3950_55_________________ANTS__O_o__o))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o ((((Menander_7339_49_________________ANTS__O_o__o ((Crocozona_3995_62 Napaea_m17_89_______________________________) Mesosemia_6516_85_________________________X)) Mesophthalma_6327_88_____________________X) Mesene_7450_28_________________________X______) Calephelis_9356_22____________________X______)) (((Eurybia_9363_20_________________ANTS__O_o__o Argyrogrammana_6709_42) (Cremna_5260_53 Euselasia_orfita____________________________X)) Ourocnemis_4621_90___________________________)) (((Thisbe_6766_43___________________ANTS__O_o__o Emesis_mandana___________________________X_____) Mesosemia_8822_02_________________________X) (Brachyglemis_3956_58 Themone_7643_34))) Juditha_9315_26__________________ANTS__O_o__o) (((((((((Protonymphidia_3948_59_________ANTS__O_o__o Symmachia_8849_08___________________________) ((Thysanota_5241_81_________________ANTS__O_o__o Phaenochitonia_6926_47) Styx_infernalis)) Sarota_9358_21) Pachythone_9050_18) T_Miletus_ancon) (Ithomiola_5924_82 (Metacharis_9347_23__________________________ Pandemos_8572_97))) (((Monethe_3977_61 Hamearis_lucina) Stalachtis_6422_39) Zelotea_4057_79)) (Hyphilaria_5239_91 Apodemia_mormo)) (((Amarynthis_5220_51 (Esthemopsis_7813_32 Roeberella_7457_36)) Riodina_lysippus) Leucochimona_9306_27_______________________X))) Lepricornis_5041_80) (Calospila_8532_96_________________ANTS__O_o__o Chamaelimnas_3942_57)) ((((((((Setabis_6330_38_________________ANTS__O_o__o Isapis_6815_44) Calydna_6346_83) Caria_7625_31_______________________________) Parcella_4012_64) (Leucochimona_7617_87_______________________X Rhetus_3973_60)) (Cyrenia_3992_63 Emesis_6837_46_______________________________)) Echenais_7168_48) Euselasia_8816_01_________________________X)) (Charis_9365_19_______________________________ Anteros_7097_50______________________________)) ((((Theope_9026_5____________________ANTS__O_o__o Methone_8523_95) T_Poritia_erycinoides) (Synargis_7605_29_________________ANTS__O_o__o Exoplisia_6433_40)) (Lemonias_3950_55_________________ANTS__O_o__o Necryia_5966_93))) Ithomeis_7773_35) Helicopis_5318_54) Cariomothis_6518_41) Leucochimona_9302_33_______________________X) ((((Alesa_8380_84____________________ANTS__O_o__o Melanis_7612_30_________________________X______) Ancyluris_6834_45) (Panara_3932_56 (Perophthalma_9330_24_______________________X Semomesia_5223_52_________________________X))) Detritivora_9327_25)) Adelotypa_7807_37_________________ANTS__O_o__o) Theope_7789_65__________________ANTS__O_o__o)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((Uraneis_hyalina____________________ANTS__O_o__o Caria_7625_31_______________________________) Calospila_8532_96_________________ANTS__O_o__o) Isapis_6815_44) (Cyrenia_3992_63 Melanis_7612_30_________________________X______)) ((((((((((((((((((Adelotypa_7807_37_________________ANTS__O_o__o Emesis_mandana___________________________X_____) (((Ancyluris_6834_45 Hamearis_lucina) Leucochimona_9302_33_______________________X) Mesophthalma_6327_88_____________________X)) (Argyrogrammana_6709_42 Parcella_4012_64)) Semomesia_5223_52_________________________X) (Charis_9365_19_______________________________ Echenais_7168_48)) (Helicopis_5318_54 ((Mesosemia_6516_85_________________________X (Napaea_m17_89_______________________________ Riodina_lysippus)) Monethe_3977_61))) Lepricornis_5041_80) ((((((Eurybia_9363_20_________________ANTS__O_o__o Stalachtis_6422_39) Zelotea_4057_79) Chamaelimnas_3942_57) (Ourocnemis_4621_90___________________________ Roeberella_7457_36)) Cariomothis_6518_41) Leucochimona_7617_87_______________________X)) Perophthalma_9330_24_______________________X) ((((Lemonias_3950_55_________________ANTS__O_o__o Ithomeis_7773_35) (((Crocozona_3995_62 Styx_infernalis) Apodemia_mormo) Emesis_6837_46_______________________________)) ((Thysanota_5241_81_________________ANTS__O_o__o Rhetus_3973_60) Esthemopsis_7813_32)) (Metacharis_9347_23__________________________ Euselasia_orfita____________________________X))) ((Theope_7789_65__________________ANTS__O_o__o T_Miletus_ancon) ((((Theope_9026_5____________________ANTS__O_o__o Phaenochitonia_6926_47) (Euselasia_8816_01_________________________X Pandemos_8572_97)) Panara_3932_56) (Mesosemia_8822_02_________________________X Sarota_9358_21)))) Symmachia_8849_08___________________________) Brachyglemis_3956_58) (((((Thisbe_6766_43___________________ANTS__O_o__o Ithomiola_5924_82) Amarynthis_5220_51) Synargis_7605_29_________________ANTS__O_o__o) Themone_7643_34) Anteros_7097_50______________________________)) Protonymphidia_3948_59_________ANTS__O_o__o) (Menander_7339_49_________________ANTS__O_o__o (Hyphilaria_5239_91 Mesene_7450_28_________________________X______))) Necryia_5966_93) (((((Alesa_8380_84____________________ANTS__O_o__o T_Poritia_erycinoides) (Juditha_9315_26__________________ANTS__O_o__o Leucochimona_9306_27_______________________X)) Detritivora_9327_25) Pachythone_9050_18) Calydna_6346_83))) Cremna_5260_53) Exoplisia_6433_40) Calephelis_9356_22____________________X______) Setabis_6330_38_________________ANTS__O_o__o) Methone_8523_95)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Semomesia_5223_52_________________________X) Zelotea_4057_79) (Theope_7789_65__________________ANTS__O_o__o Riodina_lysippus)) Rhetus_3973_60) (Helicopis_5318_54 Themone_7643_34)) Esthemopsis_7813_32) Mesosemia_8822_02_________________________X) (Thisbe_6766_43___________________ANTS__O_o__o (Argyrogrammana_6709_42 Isapis_6815_44))) (((Brachyglemis_3956_58 Caria_7625_31_______________________________) ((Emesis_6837_46_______________________________ Leucochimona_9306_27_______________________X) Pachythone_9050_18)) Exoplisia_6433_40)) T_Miletus_ancon) Methone_8523_95) Roeberella_7457_36) (Emesis_mandana___________________________X_____ Hamearis_lucina)) Ithomiola_5924_82) (Euselasia_8816_01_________________________X ((Perophthalma_9330_24_______________________X Euselasia_orfita____________________________X) Sarota_9358_21))) Calospila_8532_96_________________ANTS__O_o__o) (Detritivora_9327_25 Leucochimona_9302_33_______________________X)) Crocozona_3995_62) Cremna_5260_53) Adelotypa_7807_37_________________ANTS__O_o__o) Setabis_6330_38_________________ANTS__O_o__o) ((Calydna_6346_83 (((Echenais_7168_48 (Pandemos_8572_97 T_Poritia_erycinoides)) Phaenochitonia_6926_47) Leucochimona_7617_87_______________________X)) Chamaelimnas_3942_57)) Mesophthalma_6327_88_____________________X) Ourocnemis_4621_90___________________________) (Cyrenia_3992_63 Mesene_7450_28_________________________X______)) Eurybia_9363_20_________________ANTS__O_o__o) ((Protonymphidia_3948_59_________ANTS__O_o__o Parcella_4012_64) ((((((((Alesa_8380_84____________________ANTS__O_o__o Juditha_9315_26__________________ANTS__O_o__o) (Calephelis_9356_22____________________X______ (Hyphilaria_5239_91 (Melanis_7612_30_________________________X______ Napaea_m17_89_______________________________)))) Panara_3932_56) Metacharis_9347_23__________________________) ((Theope_9026_5____________________ANTS__O_o__o Lepricornis_5041_80) (Monethe_3977_61 Styx_infernalis))) Apodemia_mormo) Lemonias_3950_55_________________ANTS__O_o__o) Stalachtis_6422_39))) ((((((Menander_7339_49_________________ANTS__O_o__o (Ancyluris_6834_45 Charis_9365_19_______________________________)) Amarynthis_5220_51) Ithomeis_7773_35) (Thysanota_5241_81_________________ANTS__O_o__o Necryia_5966_93)) Symmachia_8849_08___________________________) Anteros_7097_50______________________________)) (Synargis_7605_29_________________ANTS__O_o__o Cariomothis_6518_41)) Mesosemia_6516_85_________________________X)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Emesis_6837_46_______________________________) ((((((((((Adelotypa_7807_37_________________ANTS__O_o__o (Setabis_6330_38_________________ANTS__O_o__o (Ancyluris_6834_45 (Monethe_3977_61 Phaenochitonia_6926_47)))) Lepricornis_5041_80) Crocozona_3995_62) Thysanota_5241_81_________________ANTS__O_o__o) ((Calospila_8532_96_________________ANTS__O_o__o (Detritivora_9327_25 Mesosemia_8822_02_________________________X)) Hamearis_lucina)) (Helicopis_5318_54 T_Poritia_erycinoides)) ((Calydna_6346_83 Styx_infernalis) Hyphilaria_5239_91)) Roeberella_7457_36) Amarynthis_5220_51) Exoplisia_6433_40)) Ithomeis_7773_35) (Chamaelimnas_3942_57 Metacharis_9347_23__________________________)) (((Eurybia_9363_20_________________ANTS__O_o__o (Synargis_7605_29_________________ANTS__O_o__o (Anteros_7097_50______________________________ T_Miletus_ancon))) Brachyglemis_3956_58) Cariomothis_6518_41)) Calephelis_9356_22____________________X______) ((((((((((((((((((((Protonymphidia_3948_59_________ANTS__O_o__o ((Alesa_8380_84____________________ANTS__O_o__o (Mesophthalma_6327_88_____________________X Riodina_lysippus)) Sarota_9358_21)) (Theope_7789_65__________________ANTS__O_o__o Ithomiola_5924_82)) Mesosemia_6516_85_________________________X) Parcella_4012_64) ((Charis_9365_19_______________________________ Leucochimona_9306_27_______________________X) Cyrenia_3992_63)) Pachythone_9050_18) Theope_9026_5____________________ANTS__O_o__o) Menander_7339_49_________________ANTS__O_o__o) Semomesia_5223_52_________________________X) Methone_8523_95) Euselasia_orfita____________________________X) Perophthalma_9330_24_______________________X) Esthemopsis_7813_32) Emesis_mandana___________________________X_____) ((Argyrogrammana_6709_42 (Necryia_5966_93 Pandemos_8572_97)) Symmachia_8849_08___________________________)) Panara_3932_56) ((Juditha_9315_26__________________ANTS__O_o__o Zelotea_4057_79) Thisbe_6766_43___________________ANTS__O_o__o)) Melanis_7612_30_________________________X______) Leucochimona_7617_87_______________________X) Napaea_m17_89_______________________________)) Isapis_6815_44) Ourocnemis_4621_90___________________________) (Euselasia_8816_01_________________________X Stalachtis_6422_39)) Caria_7625_31_______________________________) Cremna_5260_53) Lemonias_3950_55_________________ANTS__O_o__o) Apodemia_mormo) Rhetus_3973_60) Echenais_7168_48) Themone_7643_34) (Leucochimona_9302_33_______________________X Mesene_7450_28_________________________X______))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Charis_9365_19_______________________________) Mesene_7450_28_________________________X______) Sarota_9358_21) Riodina_lysippus) Ancyluris_6834_45) (((Alesa_8380_84____________________ANTS__O_o__o Necryia_5966_93) ((Calydna_6346_83 Metacharis_9347_23__________________________) ((Cyrenia_3992_63 Leucochimona_9302_33_______________________X) Exoplisia_6433_40))) (Thysanota_5241_81_________________ANTS__O_o__o Calephelis_9356_22____________________X______))) (Cariomothis_6518_41 (Themone_7643_34 Styx_infernalis))) Lepricornis_5041_80) (Ithomiola_5924_82 (Perophthalma_9330_24_______________________X Semomesia_5223_52_________________________X))) (Eurybia_9363_20_________________ANTS__O_o__o Pandemos_8572_97)) Echenais_7168_48) (Lemonias_3950_55_________________ANTS__O_o__o (Argyrogrammana_6709_42 Melanis_7612_30_________________________X______))) ((Theope_9026_5____________________ANTS__O_o__o Symmachia_8849_08___________________________) Ithomeis_7773_35)) (Anteros_7097_50______________________________ T_Miletus_ancon)) (((((((((Adelotypa_7807_37_________________ANTS__O_o__o Cremna_5260_53) T_Poritia_erycinoides) (Protonymphidia_3948_59_________ANTS__O_o__o (((Brachyglemis_3956_58 ((Esthemopsis_7813_32 Pachythone_9050_18) Rhetus_3973_60)) Hyphilaria_5239_91) (Crocozona_3995_62 Euselasia_orfita____________________________X)))) Isapis_6815_44) Phaenochitonia_6926_47) (Thisbe_6766_43___________________ANTS__O_o__o Chamaelimnas_3942_57)) (((((((Calospila_8532_96_________________ANTS__O_o__o Apodemia_mormo) Stalachtis_6422_39) ((Amarynthis_5220_51 Mesophthalma_6327_88_____________________X) Helicopis_5318_54)) ((Monethe_3977_61 Roeberella_7457_36) Panara_3932_56)) Leucochimona_9306_27_______________________X) Ourocnemis_4621_90___________________________) (Leucochimona_7617_87_______________________X Mesosemia_6516_85_________________________X))) ((((Setabis_6330_38_________________ANTS__O_o__o Detritivora_9327_25) (Synargis_7605_29_________________ANTS__O_o__o Parcella_4012_64)) Mesosemia_8822_02_________________________X) Emesis_mandana___________________________X_____)) (Caria_7625_31_______________________________ Napaea_m17_89_______________________________))) (Juditha_9315_26__________________ANTS__O_o__o Euselasia_8816_01_________________________X)) Methone_8523_95) (((Theope_7789_65__________________ANTS__O_o__o Zelotea_4057_79) Emesis_6837_46_______________________________) Hamearis_lucina)) Menander_7339_49_________________ANTS__O_o__o)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o ((Ithomiola_5924_82 Pandemos_8572_97) Mesosemia_8822_02_________________________X)) (((((((((((((((((((Adelotypa_7807_37_________________ANTS__O_o__o Helicopis_5318_54) (Thisbe_6766_43___________________ANTS__O_o__o ((Caria_7625_31_______________________________ (Phaenochitonia_6926_47 Rhetus_3973_60)) (Parcella_4012_64 Stalachtis_6422_39)))) Roeberella_7457_36) Juditha_9315_26__________________ANTS__O_o__o) Anteros_7097_50______________________________) (Necryia_5966_93 T_Miletus_ancon)) Pachythone_9050_18) Theope_9026_5____________________ANTS__O_o__o) Euselasia_8816_01_________________________X) (((((((Menander_7339_49_________________ANTS__O_o__o (Esthemopsis_7813_32 Semomesia_5223_52_________________________X)) Lepricornis_5041_80) Emesis_6837_46_______________________________) Ourocnemis_4621_90___________________________) ((Chamaelimnas_3942_57 Symmachia_8849_08___________________________) (Detritivora_9327_25 Monethe_3977_61))) (Ancyluris_6834_45 Panara_3932_56)) Napaea_m17_89_______________________________)) Calephelis_9356_22____________________X______) Styx_infernalis) Cremna_5260_53) Setabis_6330_38_________________ANTS__O_o__o) Riodina_lysippus) Lemonias_3950_55_________________ANTS__O_o__o) Calydna_6346_83) ((Thysanota_5241_81_________________ANTS__O_o__o Exoplisia_6433_40) Mesosemia_6516_85_________________________X)) ((Brachyglemis_3956_58 Crocozona_3995_62) Sarota_9358_21))) Melanis_7612_30_________________________X______) ((Theope_7789_65__________________ANTS__O_o__o Apodemia_mormo) Themone_7643_34)) Mesene_7450_28_________________________X______) Hamearis_lucina) ((Ithomeis_7773_35 Perophthalma_9330_24_______________________X) Euselasia_orfita____________________________X)) Zelotea_4057_79) Methone_8523_95) Metacharis_9347_23__________________________) Eurybia_9363_20_________________ANTS__O_o__o) (Cariomothis_6518_41 Leucochimona_9302_33_______________________X)) (Calospila_8532_96_________________ANTS__O_o__o T_Poritia_erycinoides)) ((((Amarynthis_5220_51 Cyrenia_3992_63) Leucochimona_7617_87_______________________X) Leucochimona_9306_27_______________________X) (Hyphilaria_5239_91 Emesis_mandana___________________________X_____))) Isapis_6815_44) Mesophthalma_6327_88_____________________X) Charis_9365_19_______________________________) Argyrogrammana_6709_42) (Alesa_8380_84____________________ANTS__O_o__o Echenais_7168_48)) (Protonymphidia_3948_59_________ANTS__O_o__o Synargis_7605_29_________________ANTS__O_o__o))*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((Uraneis_hyalina____________________ANTS__O_o__o ((((Menander_7339_49_________________ANTS__O_o__o Lemonias_3950_55_________________ANTS__O_o__o) Riodina_lysippus) Crocozona_3995_62) Mesene_7450_28_________________________X______)) Semomesia_5223_52_________________________X) ((Calydna_6346_83 Parcella_4012_64) Styx_infernalis)) (((Setabis_6330_38_________________ANTS__O_o__o (Ancyluris_6834_45 Metacharis_9347_23__________________________)) Euselasia_orfita____________________________X) Cremna_5260_53)) Themone_7643_34) T_Poritia_erycinoides) (Calospila_8532_96_________________ANTS__O_o__o ((Thisbe_6766_43___________________ANTS__O_o__o Leucochimona_7617_87_______________________X) Cariomothis_6518_41))) (Theope_9026_5____________________ANTS__O_o__o Cyrenia_3992_63)) Emesis_6837_46_______________________________) (Esthemopsis_7813_32 Lepricornis_5041_80)) Phaenochitonia_6926_47) (((((((((Adelotypa_7807_37_________________ANTS__O_o__o Thysanota_5241_81_________________ANTS__O_o__o) Rhetus_3973_60) Pandemos_8572_97) ((((Amarynthis_5220_51 Hamearis_lucina) (Detritivora_9327_25 Hyphilaria_5239_91)) Leucochimona_9306_27_______________________X) (Isapis_6815_44 Mesosemia_6516_85_________________________X))) Leucochimona_9302_33_______________________X) Apodemia_mormo) Alesa_8380_84____________________ANTS__O_o__o) Protonymphidia_3948_59_________ANTS__O_o__o) (((((Theope_7789_65__________________ANTS__O_o__o Brachyglemis_3956_58) (Charis_9365_19_______________________________ Mesosemia_8822_02_________________________X)) Calephelis_9356_22____________________X______) (Argyrogrammana_6709_42 Ithomeis_7773_35)) (Synargis_7605_29_________________ANTS__O_o__o Necryia_5966_93)))) (Juditha_9315_26__________________ANTS__O_o__o (Melanis_7612_30_________________________X______ T_Miletus_ancon))) ((((((((((Eurybia_9363_20_________________ANTS__O_o__o (Mesophthalma_6327_88_____________________X (Pachythone_9050_18 Panara_3932_56))) Symmachia_8849_08___________________________) Emesis_mandana___________________________X_____) Euselasia_8816_01_________________________X) ((((Caria_7625_31_______________________________ Methone_8523_95) Ithomiola_5924_82) (((Chamaelimnas_3942_57 Stalachtis_6422_39) Anteros_7097_50______________________________) Ourocnemis_4621_90___________________________)) Zelotea_4057_79)) (Helicopis_5318_54 Napaea_m17_89_______________________________)) ((Echenais_7168_48 Exoplisia_6433_40) Perophthalma_9330_24_______________________X)) Monethe_3977_61) Roeberella_7457_36) Sarota_9358_21))*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Rhetus_3973_60) Menander_7339_49_________________ANTS__O_o__o) ((Charis_9365_19_______________________________ Lepricornis_5041_80) Leucochimona_9306_27_______________________X)) Cyrenia_3992_63) Phaenochitonia_6926_47) Ancyluris_6834_45) ((Protonymphidia_3948_59_________ANTS__O_o__o Mesosemia_8822_02_________________________X) Napaea_m17_89_______________________________)) (((((Amarynthis_5220_51 Esthemopsis_7813_32) Semomesia_5223_52_________________________X) Mesosemia_6516_85_________________________X) Chamaelimnas_3942_57) Ithomeis_7773_35)) Emesis_mandana___________________________X_____) Thisbe_6766_43___________________ANTS__O_o__o) (((((Alesa_8380_84____________________ANTS__O_o__o (Leucochimona_7617_87_______________________X Melanis_7612_30_________________________X______)) ((Theope_7789_65__________________ANTS__O_o__o (Calephelis_9356_22____________________X______ (Isapis_6815_44 Ourocnemis_4621_90___________________________))) Hyphilaria_5239_91)) (Theope_9026_5____________________ANTS__O_o__o Methone_8523_95)) (Thysanota_5241_81_________________ANTS__O_o__o Hamearis_lucina)) Panara_3932_56)) Exoplisia_6433_40) Perophthalma_9330_24_______________________X) Calydna_6346_83) (((Cariomothis_6518_41 Monethe_3977_61) (Detritivora_9327_25 Mesene_7450_28_________________________X______)) (Apodemia_mormo Styx_infernalis))) Anteros_7097_50______________________________) (Caria_7625_31_______________________________ Leucochimona_9302_33_______________________X)) Synargis_7605_29_________________ANTS__O_o__o) Eurybia_9363_20_________________ANTS__O_o__o) (Ithomiola_5924_82 Riodina_lysippus)) ((((((((Adelotypa_7807_37_________________ANTS__O_o__o (((Juditha_9315_26__________________ANTS__O_o__o Helicopis_5318_54) Euselasia_orfita____________________________X) T_Poritia_erycinoides)) Necryia_5966_93) (Argyrogrammana_6709_42 (Cremna_5260_53 Pachythone_9050_18))) (Crocozona_3995_62 (Parcella_4012_64 Sarota_9358_21))) T_Miletus_ancon) Pandemos_8572_97) Metacharis_9347_23__________________________) Calospila_8532_96_________________ANTS__O_o__o)) (Echenais_7168_48 Themone_7643_34)) (((Lemonias_3950_55_________________ANTS__O_o__o (Symmachia_8849_08___________________________ Zelotea_4057_79)) Emesis_6837_46_______________________________) (((Brachyglemis_3956_58 Euselasia_8816_01_________________________X) Stalachtis_6422_39) Roeberella_7457_36))) (Setabis_6330_38_________________ANTS__O_o__o Mesophthalma_6327_88_____________________X))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Riodina_lysippus) ((((Theope_9026_5____________________ANTS__O_o__o Mesosemia_8822_02_________________________X) Ithomeis_7773_35) Calephelis_9356_22____________________X______) ((((((Calydna_6346_83 Emesis_mandana___________________________X_____) (Detritivora_9327_25 Monethe_3977_61)) Symmachia_8849_08___________________________) (Charis_9365_19_______________________________ Sarota_9358_21)) ((((Hyphilaria_5239_91 Ourocnemis_4621_90___________________________) Hamearis_lucina) (Apodemia_mormo T_Miletus_ancon)) Parcella_4012_64)) Leucochimona_9302_33_______________________X))) Helicopis_5318_54) Protonymphidia_3948_59_________ANTS__O_o__o) Cyrenia_3992_63) (((((((Adelotypa_7807_37_________________ANTS__O_o__o Exoplisia_6433_40) Napaea_m17_89_______________________________) T_Poritia_erycinoides) Ithomiola_5924_82) ((Amarynthis_5220_51 Chamaelimnas_3942_57) Perophthalma_9330_24_______________________X)) Necryia_5966_93) (((((Theope_7789_65__________________ANTS__O_o__o Ancyluris_6834_45) Menander_7339_49_________________ANTS__O_o__o) Esthemopsis_7813_32) Cremna_5260_53) Lepricornis_5041_80))) Leucochimona_7617_87_______________________X) Alesa_8380_84____________________ANTS__O_o__o) Mesene_7450_28_________________________X______) (Juditha_9315_26__________________ANTS__O_o__o Thisbe_6766_43___________________ANTS__O_o__o)) (((Setabis_6330_38_________________ANTS__O_o__o Mesosemia_6516_85_________________________X) Methone_8523_95) ((Mesophthalma_6327_88_____________________X Pachythone_9050_18) Stalachtis_6422_39))) Roeberella_7457_36) (Eurybia_9363_20_________________ANTS__O_o__o Thysanota_5241_81_________________ANTS__O_o__o)) (Isapis_6815_44 Panara_3932_56)) (((Lemonias_3950_55_________________ANTS__O_o__o Anteros_7097_50______________________________) (Synargis_7605_29_________________ANTS__O_o__o Echenais_7168_48)) Melanis_7612_30_________________________X______)) (((((Calospila_8532_96_________________ANTS__O_o__o (((Argyrogrammana_6709_42 Euselasia_8816_01_________________________X) ((Caria_7625_31_______________________________ Phaenochitonia_6926_47) Metacharis_9347_23__________________________)) Pandemos_8572_97)) (Cariomothis_6518_41 Leucochimona_9306_27_______________________X)) Rhetus_3973_60) Euselasia_orfita____________________________X) ((Emesis_6837_46_______________________________ Styx_infernalis) Semomesia_5223_52_________________________X))) Brachyglemis_3956_58) (Crocozona_3995_62 Zelotea_4057_79)) Themone_7643_34)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Euselasia_orfita____________________________X) (Methone_8523_95 Phaenochitonia_6926_47)) Menander_7339_49_________________ANTS__O_o__o) (((Alesa_8380_84____________________ANTS__O_o__o (Ithomeis_7773_35 Leucochimona_9306_27_______________________X)) Ithomiola_5924_82) (Napaea_m17_89_______________________________ Zelotea_4057_79))) ((((Eurybia_9363_20_________________ANTS__O_o__o T_Miletus_ancon) Calephelis_9356_22____________________X______) (Caria_7625_31_______________________________ Sarota_9358_21)) Pachythone_9050_18)) ((Charis_9365_19_______________________________ (((Exoplisia_6433_40 Rhetus_3973_60) Panara_3932_56) Mesosemia_8822_02_________________________X)) ((Mesophthalma_6327_88_____________________X Styx_infernalis) Perophthalma_9330_24_______________________X))) Hamearis_lucina) Cremna_5260_53) Isapis_6815_44) Calydna_6346_83) ((Calospila_8532_96_________________ANTS__O_o__o (Chamaelimnas_3942_57 Cyrenia_3992_63)) Thysanota_5241_81_________________ANTS__O_o__o)) Symmachia_8849_08___________________________) Roeberella_7457_36) ((Brachyglemis_3956_58 Stalachtis_6422_39) Euselasia_8816_01_________________________X)) ((Protonymphidia_3948_59_________ANTS__O_o__o Pandemos_8572_97) Melanis_7612_30_________________________X______)) Necryia_5966_93) Echenais_7168_48) (Theope_7789_65__________________ANTS__O_o__o T_Poritia_erycinoides)) (((((((Setabis_6330_38_________________ANTS__O_o__o Crocozona_3995_62) ((Thisbe_6766_43___________________ANTS__O_o__o Parcella_4012_64) Lepricornis_5041_80)) (Helicopis_5318_54 Anteros_7097_50______________________________)) Detritivora_9327_25) Apodemia_mormo) Cariomothis_6518_41) Mesene_7450_28_________________________X______)) Monethe_3977_61) Synargis_7605_29_________________ANTS__O_o__o) Lemonias_3950_55_________________ANTS__O_o__o) (Amarynthis_5220_51 (Leucochimona_7617_87_______________________X Themone_7643_34))) Theope_9026_5____________________ANTS__O_o__o) Esthemopsis_7813_32) Mesosemia_6516_85_________________________X) Adelotypa_7807_37_________________ANTS__O_o__o) Ourocnemis_4621_90___________________________) Semomesia_5223_52_________________________X) (Leucochimona_9302_33_______________________X Emesis_mandana___________________________X_____)) (Juditha_9315_26__________________ANTS__O_o__o Emesis_6837_46_______________________________)) (Ancyluris_6834_45 (Hyphilaria_5239_91 Riodina_lysippus))) Metacharis_9347_23__________________________) Argyrogrammana_6709_42)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o (Protonymphidia_3948_59_________ANTS__O_o__o (Setabis_6330_38_________________ANTS__O_o__o Monethe_3977_61))) (Zelotea_4057_79 Emesis_mandana___________________________X_____)) (Napaea_m17_89_______________________________ T_Poritia_erycinoides)) Parcella_4012_64) Hyphilaria_5239_91) ((Argyrogrammana_6709_42 Calephelis_9356_22____________________X______) Pandemos_8572_97)) (Juditha_9315_26__________________ANTS__O_o__o (Calydna_6346_83 Panara_3932_56))) Styx_infernalis) Methone_8523_95) Apodemia_mormo) Detritivora_9327_25) (Lemonias_3950_55_________________ANTS__O_o__o Synargis_7605_29_________________ANTS__O_o__o)) ((Amarynthis_5220_51 (Charis_9365_19_______________________________ (Crocozona_3995_62 (Cyrenia_3992_63 (Esthemopsis_7813_32 Perophthalma_9330_24_______________________X))))) Stalachtis_6422_39)) ((Theope_7789_65__________________ANTS__O_o__o Roeberella_7457_36) Caria_7625_31_______________________________)) Brachyglemis_3956_58) ((Alesa_8380_84____________________ANTS__O_o__o (Menander_7339_49_________________ANTS__O_o__o Pachythone_9050_18)) ((Cariomothis_6518_41 Isapis_6815_44) (Leucochimona_9302_33_______________________X Mesosemia_8822_02_________________________X)))) (((((((((Adelotypa_7807_37_________________ANTS__O_o__o Thisbe_6766_43___________________ANTS__O_o__o) Semomesia_5223_52_________________________X) (Chamaelimnas_3942_57 Lepricornis_5041_80)) (Euselasia_8816_01_________________________X Leucochimona_7617_87_______________________X)) (((((Exoplisia_6433_40 Ithomeis_7773_35) Melanis_7612_30_________________________X______) Symmachia_8849_08___________________________) Hamearis_lucina) Leucochimona_9306_27_______________________X)) Helicopis_5318_54) Metacharis_9347_23__________________________) Echenais_7168_48) ((((Eurybia_9363_20_________________ANTS__O_o__o Sarota_9358_21) ((((Calospila_8532_96_________________ANTS__O_o__o (((Cremna_5260_53 Anteros_7097_50______________________________) Rhetus_3973_60) Necryia_5966_93)) Riodina_lysippus) Ancyluris_6834_45) Euselasia_orfita____________________________X)) ((((Thysanota_5241_81_________________ANTS__O_o__o Phaenochitonia_6926_47) (Mesene_7450_28_________________________X______ Ourocnemis_4621_90___________________________)) (Ithomiola_5924_82 T_Miletus_ancon)) (Mesophthalma_6327_88_____________________X Mesosemia_6516_85_________________________X))) Themone_7643_34))) (Theope_9026_5____________________ANTS__O_o__o Emesis_6837_46_______________________________))*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o ((Chamaelimnas_3942_57 Euselasia_orfita____________________________X) (Metacharis_9347_23__________________________ Methone_8523_95))) (Charis_9365_19_______________________________ Anteros_7097_50______________________________)) Apodemia_mormo) ((Menander_7339_49_________________ANTS__O_o__o Ourocnemis_4621_90___________________________) Leucochimona_7617_87_______________________X)) (Cariomothis_6518_41 Symmachia_8849_08___________________________)) ((Alesa_8380_84____________________ANTS__O_o__o (((((((((((((((Setabis_6330_38_________________ANTS__O_o__o Napaea_m17_89_______________________________) (Amarynthis_5220_51 Brachyglemis_3956_58)) (((Caria_7625_31_______________________________ ((Ithomiola_5924_82 Leucochimona_9302_33_______________________X) Mesene_7450_28_________________________X______)) Riodina_lysippus) Parcella_4012_64)) Argyrogrammana_6709_42) (Eurybia_9363_20_________________ANTS__O_o__o Mesosemia_6516_85_________________________X)) Echenais_7168_48) Isapis_6815_44) Juditha_9315_26__________________ANTS__O_o__o) Theope_9026_5____________________ANTS__O_o__o) (Calephelis_9356_22____________________X______ Melanis_7612_30_________________________X______)) Synargis_7605_29_________________ANTS__O_o__o) Theope_7789_65__________________ANTS__O_o__o) Rhetus_3973_60) (((Thisbe_6766_43___________________ANTS__O_o__o Euselasia_8816_01_________________________X) Monethe_3977_61) Necryia_5966_93)) Ithomeis_7773_35)) Phaenochitonia_6926_47)) Exoplisia_6433_40) Semomesia_5223_52_________________________X) Hamearis_lucina) (Esthemopsis_7813_32 Helicopis_5318_54)) Cyrenia_3992_63) Emesis_6837_46_______________________________) (Mesosemia_8822_02_________________________X T_Poritia_erycinoides)) (Panara_3932_56 Styx_infernalis)) Themone_7643_34) (Crocozona_3995_62 (Perophthalma_9330_24_______________________X (Sarota_9358_21 T_Miletus_ancon)))) ((Cremna_5260_53 Stalachtis_6422_39) Mesophthalma_6327_88_____________________X)) (((((Protonymphidia_3948_59_________ANTS__O_o__o Calydna_6346_83) Lemonias_3950_55_________________ANTS__O_o__o) Hyphilaria_5239_91) Leucochimona_9306_27_______________________X) (Lepricornis_5041_80 Roeberella_7457_36))) Thysanota_5241_81_________________ANTS__O_o__o) (Adelotypa_7807_37_________________ANTS__O_o__o Ancyluris_6834_45)) ((Detritivora_9327_25 Pachythone_9050_18) Pandemos_8572_97)) Calospila_8532_96_________________ANTS__O_o__o) Emesis_mandana___________________________X_____) Zelotea_4057_79)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((Uraneis_hyalina____________________ANTS__O_o__o (((((((((((((((Adelotypa_7807_37_________________ANTS__O_o__o Charis_9365_19_______________________________) Sarota_9358_21) ((Ancyluris_6834_45 (Calydna_6346_83 Mesosemia_8822_02_________________________X)) Roeberella_7457_36)) Ourocnemis_4621_90___________________________) Zelotea_4057_79) (((Alesa_8380_84____________________ANTS__O_o__o ((((((Setabis_6330_38_________________ANTS__O_o__o (((((((Theope_9026_5____________________ANTS__O_o__o Semomesia_5223_52_________________________X) ((Cariomothis_6518_41 Cremna_5260_53) Melanis_7612_30_________________________X______)) Echenais_7168_48) (Amarynthis_5220_51 Parcella_4012_64)) Pandemos_8572_97) Leucochimona_9302_33_______________________X) Necryia_5966_93)) Emesis_6837_46_______________________________) Helicopis_5318_54) Thisbe_6766_43___________________ANTS__O_o__o) Mesophthalma_6327_88_____________________X) Calephelis_9356_22____________________X______)) ((Menander_7339_49_________________ANTS__O_o__o Pachythone_9050_18) Ithomiola_5924_82)) Mesosemia_6516_85_________________________X)) Anteros_7097_50______________________________) Methone_8523_95) Perophthalma_9330_24_______________________X) Theope_7789_65__________________ANTS__O_o__o) Calospila_8532_96_________________ANTS__O_o__o) Rhetus_3973_60) (((((((((Protonymphidia_3948_59_________ANTS__O_o__o Metacharis_9347_23__________________________) Lepricornis_5041_80) Esthemopsis_7813_32) Chamaelimnas_3942_57) T_Poritia_erycinoides) Themone_7643_34) (((((((((((Eurybia_9363_20_________________ANTS__O_o__o (Brachyglemis_3956_58 Emesis_mandana___________________________X_____)) Caria_7625_31_______________________________) ((Juditha_9315_26__________________ANTS__O_o__o Mesene_7450_28_________________________X______) Monethe_3977_61)) Detritivora_9327_25) Napaea_m17_89_______________________________) Cyrenia_3992_63) Argyrogrammana_6709_42) ((Isapis_6815_44 Apodemia_mormo) Ithomeis_7773_35)) Phaenochitonia_6926_47) Panara_3932_56) (Exoplisia_6433_40 Hamearis_lucina))) Lemonias_3950_55_________________ANTS__O_o__o) (Euselasia_orfita____________________________X T_Miletus_ancon))) Stalachtis_6422_39) Euselasia_8816_01_________________________X)) (Leucochimona_7617_87_______________________X Riodina_lysippus)) Leucochimona_9306_27_______________________X) Thysanota_5241_81_________________ANTS__O_o__o) Symmachia_8849_08___________________________) (Synargis_7605_29_________________ANTS__O_o__o ((Crocozona_3995_62 Hyphilaria_5239_91) Styx_infernalis)))*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Phaenochitonia_6926_47) Crocozona_3995_62) (((((((((Theope_7789_65__________________ANTS__O_o__o ((((Euselasia_8816_01_________________________X Roeberella_7457_36) Stalachtis_6422_39) Leucochimona_9306_27_______________________X) Isapis_6815_44)) (Menander_7339_49_________________ANTS__O_o__o (Rhetus_3973_60 Semomesia_5223_52_________________________X))) (Mesophthalma_6327_88_____________________X Pandemos_8572_97)) (Leucochimona_7617_87_______________________X Monethe_3977_61)) (Esthemopsis_7813_32 Hamearis_lucina)) ((((Lemonias_3950_55_________________ANTS__O_o__o Anteros_7097_50______________________________) (Mesosemia_6516_85_________________________X Themone_7643_34)) Sarota_9358_21) Zelotea_4057_79)) (Metacharis_9347_23__________________________ Panara_3932_56)) (Juditha_9315_26__________________ANTS__O_o__o (Amarynthis_5220_51 T_Poritia_erycinoides))) ((Brachyglemis_3956_58 Helicopis_5318_54) Exoplisia_6433_40))) Napaea_m17_89_______________________________) ((Argyrogrammana_6709_42 ((Chamaelimnas_3942_57 Styx_infernalis) Riodina_lysippus)) Pachythone_9050_18)) Detritivora_9327_25) ((((((Setabis_6330_38_________________ANTS__O_o__o Charis_9365_19_______________________________) Symmachia_8849_08___________________________) Mesene_7450_28_________________________X______) (Perophthalma_9330_24_______________________X Euselasia_orfita____________________________X)) (Cremna_5260_53 Echenais_7168_48)) T_Miletus_ancon)) Ithomiola_5924_82) Theope_9026_5____________________ANTS__O_o__o) Methone_8523_95) Ancyluris_6834_45) Thysanota_5241_81_________________ANTS__O_o__o) Protonymphidia_3948_59_________ANTS__O_o__o) Leucochimona_9302_33_______________________X) (Mesosemia_8822_02_________________________X Necryia_5966_93)) Emesis_mandana___________________________X_____) ((Calospila_8532_96_________________ANTS__O_o__o Apodemia_mormo) (((((Synargis_7605_29_________________ANTS__O_o__o Cyrenia_3992_63) ((Calephelis_9356_22____________________X______ Ithomeis_7773_35) Emesis_6837_46_______________________________)) Ourocnemis_4621_90___________________________) Melanis_7612_30_________________________X______) (Calydna_6346_83 Cariomothis_6518_41)))) (Adelotypa_7807_37_________________ANTS__O_o__o Parcella_4012_64)) Hyphilaria_5239_91) Caria_7625_31_______________________________) (Alesa_8380_84____________________ANTS__O_o__o Eurybia_9363_20_________________ANTS__O_o__o)) Lepricornis_5041_80) Thisbe_6766_43___________________ANTS__O_o__o)*
(T_Curetis_bulis_______________________ANTS__O_o__o ((((((Uraneis_hyalina____________________ANTS__O_o__o Perophthalma_9330_24_______________________X) Theope_9026_5____________________ANTS__O_o__o) (Menander_7339_49_________________ANTS__O_o__o Apodemia_mormo)) ((((((((((((Adelotypa_7807_37_________________ANTS__O_o__o ((Lemonias_3950_55_________________ANTS__O_o__o Phaenochitonia_6926_47) (Stalachtis_6422_39 Symmachia_8849_08___________________________))) (Euselasia_8816_01_________________________X Helicopis_5318_54)) T_Poritia_erycinoides) Melanis_7612_30_________________________X______) Mesophthalma_6327_88_____________________X) ((Theope_7789_65__________________ANTS__O_o__o Calephelis_9356_22____________________X______) Metacharis_9347_23__________________________)) ((Leucochimona_9302_33_______________________X Monethe_3977_61) T_Miletus_ancon)) Echenais_7168_48) Emesis_mandana___________________________X_____) Riodina_lysippus) ((((Setabis_6330_38_________________ANTS__O_o__o (((Thisbe_6766_43___________________ANTS__O_o__o Panara_3932_56) Charis_9365_19_______________________________) Caria_7625_31_______________________________)) Styx_infernalis) Ithomeis_7773_35) ((Synargis_7605_29_________________ANTS__O_o__o Anteros_7097_50______________________________) Ithomiola_5924_82))) (Parcella_4012_64 Roeberella_7457_36))) (Mesosemia_8822_02_________________________X Ourocnemis_4621_90___________________________)) ((Protonymphidia_3948_59_________ANTS__O_o__o (((Eurybia_9363_20_________________ANTS__O_o__o ((Calospila_8532_96_________________ANTS__O_o__o Semomesia_5223_52_________________________X) Mesosemia_6516_85_________________________X)) Calydna_6346_83) (((((Juditha_9315_26__________________ANTS__O_o__o (((((Thysanota_5241_81_________________ANTS__O_o__o Hyphilaria_5239_91) ((((Amarynthis_5220_51 Leucochimona_7617_87_______________________X) Leucochimona_9306_27_______________________X) Hamearis_lucina) (((Chamaelimnas_3942_57 Euselasia_orfita____________________________X) Cyrenia_3992_63) Exoplisia_6433_40))) Argyrogrammana_6709_42) ((Detritivora_9327_25 Pachythone_9050_18) (Napaea_m17_89_______________________________ (Pandemos_8572_97 Zelotea_4057_79)))) Emesis_6837_46_______________________________)) (((Cariomothis_6518_41 Sarota_9358_21) Cremna_5260_53) Mesene_7450_28_________________________X______)) (Lepricornis_5041_80 (Necryia_5966_93 Themone_7643_34))) Rhetus_3973_60) Methone_8523_95))) ((Alesa_8380_84____________________ANTS__O_o__o (Ancyluris_6834_45 Crocozona_3995_62)) Esthemopsis_7813_32))) (Brachyglemis_3956_58 Isapis_6815_44))*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((((((((((((Uraneis_hyalina____________________ANTS__O_o__o (Exoplisia_6433_40 Leucochimona_9306_27_______________________X)) Calydna_6346_83) Setabis_6330_38_________________ANTS__O_o__o) Helicopis_5318_54) Mesophthalma_6327_88_____________________X) ((((((((((Eurybia_9363_20_________________ANTS__O_o__o (Menander_7339_49_________________ANTS__O_o__o ((((Thysanota_5241_81_________________ANTS__O_o__o (Leucochimona_7617_87_______________________X (Themone_7643_34 Riodina_lysippus))) Ithomiola_5924_82) Parcella_4012_64) Detritivora_9327_25))) Euselasia_8816_01_________________________X) Cariomothis_6518_41) Apodemia_mormo) Zelotea_4057_79) Stalachtis_6422_39) ((((Theope_7789_65__________________ANTS__O_o__o Leucochimona_9302_33_______________________X) Amarynthis_5220_51) (Esthemopsis_7813_32 Ourocnemis_4621_90___________________________)) ((Chamaelimnas_3942_57 T_Miletus_ancon) (Cremna_5260_53 Styx_infernalis)))) Rhetus_3973_60) Crocozona_3995_62) ((Cyrenia_3992_63 (Roeberella_7457_36 Emesis_mandana___________________________X_____)) Ithomeis_7773_35))) Thisbe_6766_43___________________ANTS__O_o__o) Alesa_8380_84____________________ANTS__O_o__o) Pandemos_8572_97) Mesosemia_6516_85_________________________X) (Theope_9026_5____________________ANTS__O_o__o Ancyluris_6834_45)) Isapis_6815_44) Hamearis_lucina) ((Lepricornis_5041_80 Euselasia_orfita____________________________X) (Necryia_5966_93 Symmachia_8849_08___________________________))) Caria_7625_31_______________________________) Adelotypa_7807_37_________________ANTS__O_o__o) ((Lemonias_3950_55_________________ANTS__O_o__o Perophthalma_9330_24_______________________X) Napaea_m17_89_______________________________)) ((Juditha_9315_26__________________ANTS__O_o__o ((Argyrogrammana_6709_42 Phaenochitonia_6926_47) Emesis_6837_46_______________________________)) Synargis_7605_29_________________ANTS__O_o__o)) (Brachyglemis_3956_58 Mesene_7450_28_________________________X______)) (((Calospila_8532_96_________________ANTS__O_o__o Echenais_7168_48) Monethe_3977_61) ((((Calephelis_9356_22____________________X______ T_Poritia_erycinoides) Panara_3932_56) Metacharis_9347_23__________________________) (Charis_9365_19_______________________________ (Methone_8523_95 Anteros_7097_50______________________________))))) Sarota_9358_21) Pachythone_9050_18) (((Protonymphidia_3948_59_________ANTS__O_o__o Hyphilaria_5239_91) Semomesia_5223_52_________________________X) Mesosemia_8822_02_________________________X)) Melanis_7612_30_________________________X______)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((Uraneis_hyalina____________________ANTS__O_o__o Lepricornis_5041_80) ((((((Eurybia_9363_20_________________ANTS__O_o__o Euselasia_8816_01_________________________X) Apodemia_mormo) ((((((Theope_7789_65__________________ANTS__O_o__o Lemonias_3950_55_________________ANTS__O_o__o) ((Amarynthis_5220_51 Chamaelimnas_3942_57) Perophthalma_9330_24_______________________X)) Calospila_8532_96_________________ANTS__O_o__o) Parcella_4012_64) Cariomothis_6518_41) Euselasia_orfita____________________________X)) Synargis_7605_29_________________ANTS__O_o__o) ((Mesosemia_8822_02_________________________X (Methone_8523_95 Emesis_mandana___________________________X_____)) Pandemos_8572_97)) Hamearis_lucina)) (((Adelotypa_7807_37_________________ANTS__O_o__o (((Protonymphidia_3948_59_________ANTS__O_o__o Helicopis_5318_54) ((((Theope_9026_5____________________ANTS__O_o__o (Brachyglemis_3956_58 Exoplisia_6433_40)) T_Miletus_ancon) Juditha_9315_26__________________ANTS__O_o__o) ((((Mesophthalma_6327_88_____________________X T_Poritia_erycinoides) Rhetus_3973_60) (Panara_3932_56 Sarota_9358_21)) Roeberella_7457_36))) (Argyrogrammana_6709_42 Zelotea_4057_79))) Ourocnemis_4621_90___________________________) ((Setabis_6330_38_________________ANTS__O_o__o Echenais_7168_48) Ithomeis_7773_35))) Cyrenia_3992_63) Hyphilaria_5239_91) ((((((((((((Menander_7339_49_________________ANTS__O_o__o Monethe_3977_61) Thisbe_6766_43___________________ANTS__O_o__o) ((Calydna_6346_83 (Detritivora_9327_25 (Isapis_6815_44 Themone_7643_34))) Anteros_7097_50______________________________)) Charis_9365_19_______________________________) Ithomiola_5924_82) Pachythone_9050_18) (Caria_7625_31_______________________________ Melanis_7612_30_________________________X______)) Necryia_5966_93) Mesene_7450_28_________________________X______) ((Leucochimona_9302_33_______________________X Stalachtis_6422_39) Semomesia_5223_52_________________________X)) (((Cremna_5260_53 Napaea_m17_89_______________________________) (Crocozona_3995_62 Emesis_6837_46_______________________________)) Riodina_lysippus)) (((Mesosemia_6516_85_________________________X Symmachia_8849_08___________________________) Metacharis_9347_23__________________________) Phaenochitonia_6926_47))) Styx_infernalis) ((Alesa_8380_84____________________ANTS__O_o__o (Thysanota_5241_81_________________ANTS__O_o__o Esthemopsis_7813_32)) (Ancyluris_6834_45 Leucochimona_9306_27_______________________X))) Calephelis_9356_22____________________X______) Leucochimona_7617_87_______________________X)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((Uraneis_hyalina____________________ANTS__O_o__o Adelotypa_7807_37_________________ANTS__O_o__o) (((((((((((((((((((((((((((Protonymphidia_3948_59_________ANTS__O_o__o Esthemopsis_7813_32) ((((((Amarynthis_5220_51 Cremna_5260_53) Panara_3932_56) (Melanis_7612_30_________________________X______ Napaea_m17_89_______________________________)) Styx_infernalis) (Echenais_7168_48 Euselasia_8816_01_________________________X)) (Cyrenia_3992_63 Hyphilaria_5239_91))) Ancyluris_6834_45) Phaenochitonia_6926_47) Pachythone_9050_18) Cariomothis_6518_41) ((Alesa_8380_84____________________ANTS__O_o__o Symmachia_8849_08___________________________) (Roeberella_7457_36 Zelotea_4057_79))) (Mesosemia_8822_02_________________________X Themone_7643_34)) Leucochimona_9306_27_______________________X) (((((((((Eurybia_9363_20_________________ANTS__O_o__o T_Poritia_erycinoides) ((Menander_7339_49_________________ANTS__O_o__o ((Ithomiola_5924_82 (Leucochimona_9302_33_______________________X (Mesene_7450_28_________________________X______ Euselasia_orfita____________________________X))) (Metacharis_9347_23__________________________ Semomesia_5223_52_________________________X))) Pandemos_8572_97)) Ithomeis_7773_35) Caria_7625_31_______________________________) Mesosemia_6516_85_________________________X) Emesis_mandana___________________________X_____) (Calospila_8532_96_________________ANTS__O_o__o Leucochimona_7617_87_______________________X)) Isapis_6815_44) Mesophthalma_6327_88_____________________X)) Monethe_3977_61) Hamearis_lucina) Charis_9365_19_______________________________) Calephelis_9356_22____________________X______) Lepricornis_5041_80) Theope_7789_65__________________ANTS__O_o__o) Detritivora_9327_25) Theope_9026_5____________________ANTS__O_o__o) Anteros_7097_50______________________________) Stalachtis_6422_39) (Juditha_9315_26__________________ANTS__O_o__o Crocozona_3995_62)) (Methone_8523_95 Riodina_lysippus)) Apodemia_mormo) ((((Setabis_6330_38_________________ANTS__O_o__o (Lemonias_3950_55_________________ANTS__O_o__o Perophthalma_9330_24_______________________X)) Thysanota_5241_81_________________ANTS__O_o__o) Chamaelimnas_3942_57) Rhetus_3973_60)) Necryia_5966_93) (((Thisbe_6766_43___________________ANTS__O_o__o T_Miletus_ancon) (Brachyglemis_3956_58 Parcella_4012_64)) (Emesis_6837_46_______________________________ Exoplisia_6433_40))) Argyrogrammana_6709_42)) Helicopis_5318_54) Sarota_9358_21) (Synargis_7605_29_________________ANTS__O_o__o Calydna_6346_83)) Ourocnemis_4621_90___________________________)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((Uraneis_hyalina____________________ANTS__O_o__o (((((Adelotypa_7807_37_________________ANTS__O_o__o Theope_7789_65__________________ANTS__O_o__o) Calephelis_9356_22____________________X______) Esthemopsis_7813_32) Caria_7625_31_______________________________) Cariomothis_6518_41)) ((((Protonymphidia_3948_59_________ANTS__O_o__o Napaea_m17_89_______________________________) (Emesis_6837_46_______________________________ Perophthalma_9330_24_______________________X)) ((((((Juditha_9315_26__________________ANTS__O_o__o Necryia_5966_93) (Mesosemia_6516_85_________________________X Pandemos_8572_97)) Symmachia_8849_08___________________________) Phaenochitonia_6926_47) Lepricornis_5041_80) Helicopis_5318_54)) ((Alesa_8380_84____________________ANTS__O_o__o Argyrogrammana_6709_42) Semomesia_5223_52_________________________X))) ((((((((Calospila_8532_96_________________ANTS__O_o__o Panara_3932_56) Isapis_6815_44) Exoplisia_6433_40) Thysanota_5241_81_________________ANTS__O_o__o) Monethe_3977_61) Cyrenia_3992_63) Zelotea_4057_79) Mesene_7450_28_________________________X______)) ((((((((((((((((((((Setabis_6330_38_________________ANTS__O_o__o Euselasia_8816_01_________________________X) Leucochimona_9306_27_______________________X) Amarynthis_5220_51) Metacharis_9347_23__________________________) Anteros_7097_50______________________________) (Hamearis_lucina T_Poritia_erycinoides)) Brachyglemis_3956_58) (Methone_8523_95 Roeberella_7457_36)) ((Detritivora_9327_25 Apodemia_mormo) Melanis_7612_30_________________________X______)) (((Menander_7339_49_________________ANTS__O_o__o Mesophthalma_6327_88_____________________X) Euselasia_orfita____________________________X) (Calydna_6346_83 Parcella_4012_64))) Themone_7643_34) Rhetus_3973_60) (Cremna_5260_53 Crocozona_3995_62)) (Hyphilaria_5239_91 Emesis_mandana___________________________X_____)) Mesosemia_8822_02_________________________X) (Lemonias_3950_55_________________ANTS__O_o__o Riodina_lysippus)) Leucochimona_7617_87_______________________X) (Chamaelimnas_3942_57 (Ithomiola_5924_82 Stalachtis_6422_39))) Pachythone_9050_18) Charis_9365_19_______________________________)) Synargis_7605_29_________________ANTS__O_o__o) T_Miletus_ancon) Theope_9026_5____________________ANTS__O_o__o) Thisbe_6766_43___________________ANTS__O_o__o) (Sarota_9358_21 Styx_infernalis)) Eurybia_9363_20_________________ANTS__O_o__o) Ancyluris_6834_45) Leucochimona_9302_33_______________________X) (Echenais_7168_48 Ourocnemis_4621_90___________________________)) Ithomeis_7773_35)*
(T_Curetis_bulis_______________________ANTS__O_o__o (((((((((((((Uraneis_hyalina____________________ANTS__O_o__o Sarota_9358_21) Parcella_4012_64) Brachyglemis_3956_58) Adelotypa_7807_37_________________ANTS__O_o__o) Mesophthalma_6327_88_____________________X) Calydna_6346_83) Amarynthis_5220_51) (Setabis_6330_38_________________ANTS__O_o__o (Lemonias_3950_55_________________ANTS__O_o__o Lepricornis_5041_80))) Pachythone_9050_18) ((Thysanota_5241_81_________________ANTS__O_o__o ((Calephelis_9356_22____________________X______ Exoplisia_6433_40) ((Esthemopsis_7813_32 Napaea_m17_89_______________________________) (Melanis_7612_30_________________________X______ Zelotea_4057_79)))) T_Poritia_erycinoides)) (Caria_7625_31_______________________________ Necryia_5966_93)) Metacharis_9347_23__________________________) ((((((((((((((((((((Protonymphidia_3948_59_________ANTS__O_o__o (Anteros_7097_50______________________________ Apodemia_mormo)) Leucochimona_9302_33_______________________X) Phaenochitonia_6926_47) Calospila_8532_96_________________ANTS__O_o__o) Hyphilaria_5239_91) (((Theope_7789_65__________________ANTS__O_o__o Cremna_5260_53) Argyrogrammana_6709_42) ((Juditha_9315_26__________________ANTS__O_o__o Charis_9365_19_______________________________) Echenais_7168_48))) (((((Menander_7339_49_________________ANTS__O_o__o ((((Synargis_7605_29_________________ANTS__O_o__o Crocozona_3995_62) (Cyrenia_3992_63 Panara_3932_56)) (Rhetus_3973_60 (Stalachtis_6422_39 Themone_7643_34))) (Helicopis_5318_54 Mesosemia_6516_85_________________________X))) Symmachia_8849_08___________________________) Thisbe_6766_43___________________ANTS__O_o__o) (Cariomothis_6518_41 Euselasia_orfita____________________________X)) Chamaelimnas_3942_57)) Ithomeis_7773_35) (Leucochimona_7617_87_______________________X Styx_infernalis)) Emesis_mandana___________________________X_____) ((((Detritivora_9327_25 Emesis_6837_46_______________________________) Roeberella_7457_36) Semomesia_5223_52_________________________X) Riodina_lysippus)) (Theope_9026_5____________________ANTS__O_o__o Euselasia_8816_01_________________________X)) Eurybia_9363_20_________________ANTS__O_o__o) ((Alesa_8380_84____________________ANTS__O_o__o Ancyluris_6834_45) Isapis_6815_44)) (Monethe_3977_61 Perophthalma_9330_24_______________________X)) T_Miletus_ancon) Hamearis_lucina) Ourocnemis_4621_90___________________________) (Leucochimona_9306_27_______________________X (Methone_8523_95 Pandemos_8572_97))) Mesosemia_8822_02_________________________X)) (Ithomiola_5924_82 Mesene_7450_28_________________________X______));

procedure /;