• TABLE OF CONTENTS
HIDE
 Front Cover
 Copyright
 Main






Group Title: Distributional patterns of southeastern mayflies (Ephemeroptera) (FLMNH Bulletin v.22, no.1)
Title: Distributional patterns of southeastern mayflies (Ephemeroptera)
CITATION THUMBNAILS PAGE IMAGE ZOOMABLE
Full Citation
STANDARD VIEW MARC VIEW
Permanent Link: http://ufdc.ufl.edu/UF00095839/00001
 Material Information
Title: Distributional patterns of southeastern mayflies (Ephemeroptera)
Physical Description: 55 p. : map ; 23 cm.
Language: English
Creator: Berner, Lewis
Donor: unknown ( endowment )
Publisher: Florida State Museum, University of Florida
Place of Publication: Gainesville, Fla.
Publication Date: 1977
Copyright Date: 1977
 Subjects
Subject: Mayflies -- Geographical distribution -- Southern States   ( lcsh )
Insects -- Geographical distribution   ( lcsh )
Insects -- Southern States   ( lcsh )
Genre: bibliography   ( marcgt )
government publication (state, provincial, terriorial, dependent)   ( marcgt )
non-fiction   ( marcgt )
 Notes
Bibliography: Bibliography: p. 53-55.
General Note: Bulletin of the Florida State Museum, Biological science, volume 22, number 1
Statement of Responsibility: Lewis Berner.
 Record Information
Bibliographic ID: UF00095839
Volume ID: VID00001
Source Institution: University of Florida
Holding Location: University of Florida
Rights Management: All rights reserved by the source institution and holding location.
Resource Identifier: oclc - 04000461
lccn - 78621451

Table of Contents
    Front Cover
        Front Cover
    Copyright
        Copyright
    Main
        Page 1
        Page 2
        Page 3
        Page 4
        Page 5
        Page 6
        Page 7
        Page 8
        Page 9
        Page 10
        Page 11
        Page 12
        Page 13
        Page 14
        Page 15
        Page 16
        Page 17
        Page 18
        Page 19
        Page 20
        Page 21
        Page 22
        Page 23
        Page 24
        Page 25
        Page 26
        Page 27
        Page 28
        Page 29
        Page 30
        Page 31
        Page 32
        Page 33
        Page 34
        Page 35
        Page 36
        Page 37
        Page 38
        Page 39
        Page 40
        Page 41
        Page 42
        Page 43
        Page 44
        Page 45
        Page 46
        Page 47
        Page 48
        Page 49
        Page 50
        Page 51
        Page 52
        Page 53
        Page 54
        Page 55
        Page 56
        Page 57
        Page 58
Full Text








of the
FLORIDA STATE MUSEUM
Biological Sciences


Volume 22 1977 Number 1




DISTRIBUTIONAL PATTERNS OF SOUTHEASTERN
MAYFLIES (EPHEMEROPTERA)


LEWIS BERNER


UNIVERSITY OF FLORIDA


GAINESVILLE









Numbers of the BULLETIN OF THE FLORIDA STATE MUSEUM, BIOLOGICAL SCIENCES,
are published at irregular intervals. Volumes contain about 300 pages and are not necessarily
completed in any one calendar year.









OLIVER L. AUSTIN, Editor

RHODA J. RYBAK, Managing Editor


Consultants for this issue:

RICHARD FRANZ

CLIFFORD JOHNSON













Communications concerning purchase or exchange of the publications and all manuscripts
should be addressed to the Managing Editor of the Bulletin, Florida State Museum, Museum
Road, University of Florida, Gainesville, Florida 32611.


Publication date: May 6, 1977


This public document was promulgated at an annual cost of $2,291.45 or
$2.291 per copy. It makes available to libraries, scholars, and all interested
persons the results of researches in the natural sciences, emphasizing the
Circum-Caribbean region.


Price: $2.30























DISTRIBUTIONAL PATTERNS OF SOUTHEASTERN
MAYFLIES (EPHEMEROPTERA)I

LEWIS BERNER2


SYNOPSIS: Distributional records of mayflies (Ephemeroptera) occurring in the southeastern
states of Alabama, Florida, Georgia, Louisiana, Mississippi, North Carolina, South Carolina,
and Tennessee are given along with the stage of development of the insect and the month it was
collected. Previously published state records are summarized and each species is assigned physi-
ographic provinces to help the user of the data relate distribution to terrain.

TABLE OF CONTENTS


INTRODUCTION ......................................... ............... ..................... .................................. 2
ACKNOWLEDGMENTS AND EXPLANATIONS ..................................................................................... 2
PHYSIOGRAPHIC PROVINCES AND ASSIGNED COUNTIES .................................................................. 2
D ISCUSSION ....... ................... ........ ............................. .......................................... ............ 5
SYSTEMATIC LISTING OF SOUTHEASTERN GENERA................................................................. 10
DISTRIBUTIONAL RECORDS
S ip h lo n u rid a e .......................................................... .. ....... ... ... ................................................ 10
M e treto p o d id a e .................................................... .................................. .... .......................... 15
B a e tid a e ....................................... .......................... ............. ............................................... 1 5
O lig o n eu riid ae .................... .................................................................. ....... ...... .... .......... 23
H eptageniidae ............................ ................................. ....... ........... ............ 23
N eoep h em eridae ............................................................... ............................................... 33
C aen idae ........... ................................................................................................. ...... . ... 33
Baetiscidae ... ................................................................. ....... ........ ................ 35
L ep to p h leb iid ae ...................................... .... .................. ...... ............ ................. ............. ...... 3 6
E p h em erellid ae ...................................................... ................... ................ ...................... 4 1
T rico ry th id a e ..................................................................................................... .................. 46
B ehningiidae .................................... .......... .... ........................... ... ....... ............ .. 48
P o tam an th id ae ............................ .............................................. ................... .................... 48
Ephem eridae ....................... ............................................. ..................... ............ 49
P o ly m itarcid ae ....................................................................................... ............. .................. 52
R E F E R E N C E S ........................................ ....... ............................................................................... 53


' Florida Agricultural Experiment Station Journal Series No. 5948.
* The author is a Professor in the Department of Zoology, University of Florida, Gainesville, Florida 32611,
and a Research Associate of the Florida State Collection of Arthropods. Manuscript accepted March 27,
1975.
BERNER, LEWIS. 1977. Distributional Patterns of Southeastern Mayflies (Ephemeroptera).
Bull. Fla. State Mus., Biol. Sci. Vol. 22(1): 1-56.







BULLETIN FLORIDA STATE MUSEUM


INTRODUCTION

For many years I have been accumulating records of southeastern may-
flies, many of which have never been published. Because of the increasing
interest that has been generated in these insects in recent years and their re-
lationship to water quality studies, I feel that it is now appropriate to bring
these data together for the use of aquatic biologists of the eight southeastern
states included in this paper.
During the years 1953-1961, I received support from the Public Health
Service (Grant No. RG4058) to make a detailed study of mayflies of the south-
east. Because of distances involved and time limitations, I confined my work
to the following states: Alabama, Florida, Georgia, Louisiana (eastern half),
Mississippi, North Carolina, South Carolina, and Tennessee. Admittedly the
delimitation was arbitrary, for certainly other states or parts of them should
have been included. Hopefully this publication will stimulate scientists work-
ing in Virginia, Kentucky, Arkansas, Texas and other contiguous states, usually
considered as southeastern, to augment these records.
The temptation to include systematic and ecological notes in this paper
is great but time and space do not allow such broadening of the subject. Some
of my later papers will be concerned with these matters. Space limitations
prevent listing of specific localities but the county and month of collection
are given.

ACKNOWLEDGMENTS AND EXPLANATIONS
Unless otherwise specified, all collections were made by me, my assistants, especially Dr.
C. D. Hynes, or friends, and all specimens and records are deposited in the Florida State Col-
lection of Arthropods of the Division of Plant Industry. I am indebted to the Highlands Biological
Station, North Carolina, for its support of my studies over many years.
For the sake of completeness, I have included all published records available to me. In the
late 1950's I was given the opportunity to study the mayfly collections of the Illinois Natural
History Survey, the Academy of Natural Sciences of Philadelphia, the Fattig Collection of Emory
University, and the Clemson College Entomological Collection. At the time of my visits to these
institutions I was authorized to include their southeastern records in my publications. Each such
record is appropriately recognized by a superscript as follows: 1. Illinois Natural History Survey;
2. Academy of Natural Sciences of Philadelphia; 3. Fattig Collection; 4. Clemson College.
State records published prior to 1935 are not listed unless the book, "The Biology of May-
flies" by Needham, Traver, and Hsu, failed to include specific references to them. To avoid
lengthy citations in this section, generally only the initials of the author or authors and the date
of publication are given. In the list of references at the end of this paper the symbols used are
given with the complete citation.
To allow the reader to make some estimate of time of year when adults may be on the wing,
I have designated the stage collected, using the following symbols: N = nymph, A = adult.


PHYSIOGRAPHIC PROVINCES AND ASSIGNED COUNTIES

I consider it desirable to relate patterns of distribution in terms of the
physiographic regions from which the records come. To allow easy placement


Vol. 22, No. 1







BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


of species records in physiographic provinces, I have assigned counties of each
state as nearly as possible using the criteria generally recognized for distin-
guishing them. The provinces, shown in figure 1, are: (1) Coastal Plain (CP)-
altitudes below 500 feet with more than half the plain being below 100 feet
in elevation. (2) Piedmont (P)-altitudes range from 500 feet at the Fall Line
to about 1000 feet at the base of the mountains to the west. (3) Blue Ridge (BR)
-narrow, varying from 5 to 50 miles in width. Easternmost ridge of the Appa-
lachian Highlands and rising 1000-5000 feet above the Piedmont. (4) Valley
and Ridge (VR)-much folded mountains with thrust faulting in the southern
part. (5) Appalachian Plateaus (AP)-almost equal in area to the Piedmont,
Blue Ridge, and Valley and Ridge provinces combined. An elevated tract of
nearly horizontal or gently folded strata with altitudes from 1000 feet along
the western edge to about 3000 feet to the east. (6) Interior Low Plateaus (ILP)
-part of a broad anticline.


COASTAL PLAIN
ALABAMA: Baldwin, Barbour, Bullock, Butler, Choctaw, Clarke, Coffee, Conecuh, Covington,
Crenshaw, Dale, Dallas, Escambia, Geneva, Greene, Hale, Henry, Houston, Lamar, Lowndes,
Macon, Marengo, Mobile, Monroe, Montgomery, Perry, Pickins, Pike, Russell, Sumter, Washing-
ton, Wilcox.
FLORIDA: All counties
GEORGIA: Appling, Atkinson, Bacon, Baker, Ben Hill, Berrien, Bibb, Bleckley, Brantley, Brooks,
Bryan, Bulloch, Burke, Calhoun, Camden, Candler, Charlton, Chatham, Chattahoochee, Clay,
Clinch, Coffee, Colquitt, Cook, Crawford, Crisp, Decatur, Dodge, Dooly, Dougherty, Early,
Echols, Effingham, Emanuel, Evans, Glynn, Grady, Harris, Houston, Irwin, Jeff Davis, Jefferson,
Jenkins, Johnson, Lanier, Laurens, Lee, Liberty, Long, Lowndes, McIntosh, Macon, Marion,
Miller, Mitchell, Montgomery, Muscogee, Peach, Pierce, Pulaski, Quitman, Randolph, Rich-
mond, Schley, Screven, Seminole, Stewart, Sumter, Talbot, Tattnall, Taylor, Telfair, Terrell,
Thomas, Tift, Toombs, Treutlen, Turner, Twiggs, Ware, Washington, Wayne, Webster, Wheeler,
Wilcox, Wilkinson, Worth.
LOUISIANA: All counties.
MississiPPI: All counties.
NORTH CAROLINA: Beaufort, Bertie, Bladen, Brunswick, Camden, Carteret, Chowan, Columbus,
Craven, Cumberland, Currituck, Dare, Duplin, Edgecombe, Gates, Greene, Halifax, Harnett,
Hertford, Hoke, Hyde, Johnston, Jones, Lenoir, Martin, Nash, New Hanover, Northampton,
Onslow, Pamlico, Pasquotank, Pender, Perquimans, Pitt, Richmond, Robeson, Sampson, Scot-
land, Tyrrell, Washington, Wayne, Wilson.
SOUTH CAROLINA: Aiken, Allendale, Bamberg, Barnwell, Beaufort, Berkeley, Calhoun, Charles-
ton, Clarendon, Colleton, Darlington, Dillon, Dorchester, Florence, Georgetown, Hampton,
Horry, Jasper, Lee, Lexington, Marion, Marlboro, Orangeburg, Richland, Sumter, Williamsburg.
TENNESSEE: Benton, Carroll, Chester, Crockett, Decatur, Dyer, Fayette, Gibson, Hardeman,
Hardin, Haywood, Henderson, Henry, Lake, Lauderdale, McNairy, Madison, Obion, Shelby,
Tipton, Weakley.


PIEDMONT PROVINCE
ALABAMA: Autauga, Chambers, Chilton, Clay, Elmore, Lee, Randolf, Tallapoosa.
GEORGIA: Baldwin, Barrow, Butts, Carroll, Cherokee, Clark, Clayton, Cobb, Columbia, Coweta,
DeKalb, Douglas, Elbert, Fayette, Franklin, Fulton, Glascock, Greene, Gwinnett, Hall, Hancock,
Haralson, Hart, Heard, Henry, Jackson, Jasper, Jones, Lamar, Lincoln, McDuffie, Madison,


1977













































Figure 1.-Physiographic provinces of the southeastern United States.







BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


Meriwether, Monroe, Morgan, Newton, Oconee, Oglethorpe, Paulding, Pike, Putnam, Rock-
dale, Spaulding, Taliaferro, Troup, Upson, Walton, Warren, Wilkes.
NORTH CAROLINA: Alamance, Anson, Cabarrus, Caswell, Chatham, Cleveland, Davidson, David,
Durham, Forsyth, Franklin, Granville, Guilford, Iredell, Lee, Mecklenburg, Montgomery, Moore,
Orange, Person, Randolph, Rockingham, Rowan, Stanly, Stokes, Surry, Union, Vance, Wake,
Warren, Yadkin.
SOUTH CAROLINA: Abbeville, Aiken, Anderson, Cherokee, Chester, Chesterfield, Edgefield, Fair-
field, Greenville, Greenwood, Kershaw, Lancaster, Laurens, Lexington, McCormick, Newberry,
Richland, Saluda, Spartanburg, Union, York.

BLUE RIDGE PROVINCE
GEORGIA: Banks, Habersham, Lumpkin, Rabun, Stephens, Towns, Union, White.
NORTH CAROLINA: Alexander, Alleghany, Ashe, Avery, Buncombe, Burke, Caldwell, Catawba,
Cherokee, Clay, Cleveland, Gaston, Graham, Haywood, Henderson, Jackson, Lincoln, Mc-
Dowell, Macon, Madison, Mitchell, Polk, Rutherford, Surry, Swain, Transylvania, Watauga,
Wilkes, Yancey.
SOUTH CAROLINA: Greenville, Oconee, Pickens
TENNESSEE: Blount, Carter, Cocke, Greene, Johnson, Monroe, Polk, Rhea, Sevier, Sullivan,
Unicoi, Washfington.

VALLEY AND RIDGE PROVINCE
ALABAMA: Bibb, Calhoun, Cleburne, Coosa, Saint Clair, Shelby, Talladega.
GEORGIA: Bartow, Catoosa, Chattooga, Dawson, Fannin, Floyd, Forsyth, Gilmer, Gordon,
Murray, Pickins, Polk, Walker, Whitfield.
TENNESSEE: Bradley, Claiborne, Grainger, Hamblen, Hamilton, Hancock, Hawkins, Jefferson,
Knox, Loudon, McMinn, Meigs, Rhea, Roane, Union.

APPALACHIAN PLATEAUS
ALABAMA: Blount, Cherokee, Cullman, DeKalb, Etowah, Fayette, Franklin, Jackson, Jefferson,
Marion, Marshall, Morgan, Tuscaloosa, Walker, Winston.
GEORGIA: Dade
TENNESSEE: Anderson, Bledsoe, Campbell, Coffee, Cumberland, DeKalb, Fentress, Franklin,
Grundy, Marion, Moore, Morgan, Overton, Pickett, Putnam, Scott, Sequatchie, Van Buren,
Warren, White.

INTERIOR Low PLATEAUS

ALABAMA: Colbert, Lauderdale, Lawrence, Limestone, Madison
TENNESSEE: Bedford, Cannon, Cheatham, Clay, Davidson, DeKalb, Dickson, Giles, Hickman,
Houston, Humphreys, Jackson, Lawrence, Lewis, Lincoln, Macon, Marshall, Maury, Mont-
gomery, Perry, Robertson, Rutherford, Smith, Stewart, Sumner, Trousdale, Wayne, Williamson,
Wilson.



DISCUSSION

The geographic distribution of mayflies is a reflection of ecologically
adaptive features developed by the nymphs and are the ones most severely
limiting the spread of the species. The southeastern mayfly fauna is a large
one with more than 250 species collected in the states included in this study.






BULLETIN FLORIDA STATE MUSEUM


The greatest variety occurs in the regions with the most varied aquatic habi-
tats, particularly those with a wide range of stream types.
The southeastern United States is well-endowed with waterways and these
have served as the pathways for the spread of species into all suitable stream
systems. The bulk of the species are inhabitants of flowing waters in the im-
mature stages and the most diverse habitats are located in the uplands.
Table 1 summarizes the distributional patterns of the various genera based
on their presumed area of origin. Following the concepts of Dr. George F.
Edmunds et al. (1976) with regard to Ephemeropteran origins, mid-austral
genera are derived from South America or South American ancestors and ex-
tend through Central America and Mexico into the southern part of the
United States. Some of these mid-austral genera have frequently moved into
the central United States and central Canada via major river systems. The
upper austral genera are clearly of southern origin but have a few species that
extend their ranges northward into Canada. The boreal genera are Holarctic
or Nearctic, the latter being closely related to Holarctic forms. The mid-
boreal genera are generally distributed southwards in the United States and
their immatures are adapted to living in cool waters. The lower boreal genera
range southwards into Mexico and sometimes into South America.
The austral element in the southeast appears to have been derived by
movement northwards from Mexico and through the warm-water streams of
the southern Coastal Plain. Boreal elements left in the warm waters of the
southern Coastal Plain are probably relicts of Pleistocene glaciation. The few
austral species in the genera Callibaetis, Paracloeodes, Homoeoneuria, Lep-
tohyphes, Tricorythodes, Hexagenia, Pentagenia, and Tortopus moved into
the United States via Mexico and spread widely. Of the eight, Paracloeodes
and Homoeoneuria seem to be the most highly restricted. None of the genera,
other than Callibaetis and Hexagenia, have succeeded in diversifying ex-
tensively as they moved northwards and they are represented by only one
or a few northern species. There seems to have been no movement from the
islands of the Antilles into the southeastern United States, likely because of
the lack of suitable habitats for the immatures in Florida, the most probable
land mass to be colonized from the Greater Antilles, other than Central
America or Mexico.
Table 2 lists the genera associated with each of the southeastern physio-
graphic provinces. The greatest number, by far, occurs in all six divisions.
Most of those of limited distribution have an austral origin. The Blue Ridge
province with its multitude of fast-flowing, cool streams has the largest num-
ber of species, while the slow-flowing waters of that part of the Interior Low
Plateaus studied has the fewest.


Vol. 22, No. 1














TABLE 1.-GEOGRAPHIC DISTRIBUTION OF THE MAYFLIES OF THE SOUTHEASTERN UNITED STATES.


Mid-Boreal


Acanthametropus.
Ameletus
Anepeorus
Baetisca
Centroptilunm
Cinygmula
Cloeon
Dolania
Epeorus
Ephemera
Ephenierella
Ephoron
Habrophlebia
Habrophlebiodes


Ileterocloeon
Leptophlebia
Litobrancha
Neoephemera
Paraleptophlebia
Potamnanthus
Pseudiron
Pseudocloeon
Rhithrogena
Siphlonurus
Siphloplecton
Spinadis
Stenacron


Mid-Austral



Homnoeoneuria
Leptohyphes
Paracloeodes
Pentagenia
Tortopus


Upper Austral



Callibaetis
Hlexagenia
Tricorythodes


Lower Boreal



Brachycercus
Caenis
Choroterpes
Heptagenia
Isonychia
Stenonemna


Cosmopolitan


Bactis











TABLE 2.-DISTRIBUTION OF SOUTHEASTERN GENERA OF MAYFLIES AS RELATED TO PHYSIOGRAPHIC PROVINCES IN WHICH THEY OCCUR.


GENUS Coastal Plain Piedmont Blue Ridge Valley and Ridge Appalachian Plateaus Interior Low Plateaus



Acanthametropus X
Anepeorus X
Dolania X
Litobrancha X
Spinadis X
Tortopus X

Homoeoneuria X X
Leptohyphes X X
Pentagenia X X

Baetisca X X X
Choroterpes X X X
Cinygmula X X X
Habrophlebia X X X
Heterocloeon X X X
Pseudiron X X X
Siphloplecton X X X

Ameletus X X X X
Brachycercus X X X X
Epeorus X X X X
Neoephemera X X X X
Paracloeodes X X X X









Calibaetis X X X X X
Ephoron X X X X X
Habrophlebiodes X X X X X
Leptophlebia X X X X X
Potamanthus X X X X X
Siphlonurus X X X X X

Baetis X X X X X X
Caenis X X X X X X
Centroptilum X X X X X X
Cloeon X X X X X X
Ephemera X X X X X X
Ephemerella X X X X X X
Heptagenia X X X X X X
Hexagenia X X X X X X
Isonychia X X X X X X
Paraleptophlebia X X X X X X
Pseudocloeon X X X X X X
Rhithrogena X X X X X X
Stenacron X X X X X X
Stenonema X X X X X X
Tricorythodes X X X X X X

Total no. genera 38 33 33 25 22 22
Total no. species 111 93 150 85 63 49








BULLETIN FLORIDA STATE MUSEUM

SYSTEMATIC LISTING OF SOUTHEASTERN GENERA3


Heptagenioidea
Siphlonuridae
Siphlonurinae
Ameletus
Siphlonurus
Acanthametropodinae
Acanthametropus
Isonychiinae
Isonychia
Metretopodidae
Metretopodinae
Siphloplecton
Baetidae
Baetinae
Baetis
Callibaetis
Centroptilum
Cloeon
Heterocloeon
Paracloeodes
Pseudocloeon
Oligoneuriidae
Oligoneuriinae
Homoeoneuria
Heptageniidae
Pseudironinae
Pseudiron
Heptageniinae
Cinygmula
Epeorus
Heptagenia
Rhithrogena
Stenacron
Stenonema
Spinadinae
Spinadis
Anepeorinae
Anepeorus
Caenoidea
Neoephemeridae
Neoephemerinae
Neoephemera


Vol. 22, No. 1


Caenidae
Caeninae
Brachycercus
Caenis
Prosopistomatoidea
Baetiscidae
Baetiscinae
Baetisca
Leptophlebioidea
Leptophlebiidae
Leptophlebiinae
Choroterpes
Habrophlebia
Habrophlebiodes
Leptophlebia
Paraleptophlebia
Ephemerellidae
Ephemerellinae
Ephemerella
Tricorythidae
Leptohyphinae
Leptohyphes
Tricorythodes
Ephemeroidea
Behningiidae
Behningiinae
Dolania
Potamanthidae
Potamanthinae
Potamanthus
Ephemeridae
Ephemerinae
Ephemera
Hexagenia
Litobrancha
Pentagenia
Polymitarcidae
Polymitarcinae
Ephoron
Campsurinae
Tortopus


SUPERFAMILY HEPTAGENIOIDEA
FAMILY SIPHLONURIDAE
SUBFAMILY SIPHLONURINAE
GENUS Ameletus EATON
Amneletus lineatus Traver
Physiographic provinces: BR, VR, AP, ILP.
Previously published state records: N.C., NTH, 1935; Tenn., WB, 1949.


'Higher categories after Dr. George F. Edmunds, Jr.








BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


Additional records: NORTH CAROLINA, McDowell Co., April (N,A). TENNESSEE, Cumberland
Co., April (N); Fentress Co., May (A); Grainger Co., April (N,A); Lincoln Co., April (N);
Marion Co., April (N); Overton Co., April (N); Sevier Co., April (A).
Ameletus species
Physiographic provinces: BR, VR, AP.
Additional records: ALABAMA, DeKalb Co., April (A). NORTH CAROLINA, Avery Co., February
and April (N); Clay Co., April (N); Macon Co, May (N)'; Orange Co., February and May
(N); Swain Co., April, May, and December (N); Transylvania Co., March and June (N);
Yancey Co., February, May, and July (N). TENNESSEE, Meigs Co., April (N); Sevier Co.,
April, May (N), July (A).



GENUS Siphlonurus EATON
Siphlonurus decorus Traver
Physiographic provinces: CP
Previously published state records: N.C., NTH, 1935.
Additional records: SOUTH CAROLINA, Orangeburg Co., April (A).
Siphlonurus luridipennis (Burmeister)
Previously published state records: Carolina, Bur., 1839
Additional records: None
Siphlonurus marginatus Traver
Physiographic provinces: P
Previously published state records: N.C., NTH, 1935
Additional records: None
Siphlonurus mirus Eaton
Physiographic provinces: P, BR
Previously published state records: GEORGIA, Cowetta Co., April (A)3. NORTH CAROLINA,
Macon Co., June (A). SOUTH CAROLINA, Greenville Co., March and April (A)'; Newberry
Co., April (N)'.
Siphlonurus quebecensis (Provancher)
Physiographic provinces: P, BR, VR, AP
Previously published state records: N.C., S.C., NTH, 1935; Tenn., WB. 1949.
Additional records: NORTH CAROLINA, Wake Co., April (A). SOUTH CAROLINA, Anderson Co.,
April (A)'; Greenville Co., April (A)'. TENNESSEE, Cumberland Co., April (N); Fentress
Co., May (A); Greene Co., May (A).
Siphlonurus species
Physiographic provinces: P, BR, VR, AP
Additional records: GEORGIA, Catoosa Co., March (N)3; Clayton Co., March (N)3; Fulton Co.,
March (A)3; Towns Co., April (A)3. SOUTH CAROLINA, Newberry Co., April (A)'; Greenville
Co., April (N)'. TENNESSEE, Cumberland Co., April (N); Fentress Co., May (A); Greene
Co., May (A).



SUBFAMILY ACANTHAMETROPODINAE
GENUS Acanthametropus TSHERNOVA
Acanthametropus species
Physiographic provinces: CP
Previously published state records: Ga.-S.C. state line, EAP, 1963
Additional records: None







BULLETIN FLORIDA STATE MUSEUM


SUBFAMILY ISONYCHIINAE
GENUS Isonychia EATON
Isonychia annulata Traver
Physiographic provinces: P
Previously published state records: N.C., NTH, 1935
Additional records: None
Isonychia aurea Traver
Physiographic provinces: BR, VR
Previously published state records: N. C., NTH, 1935, TH, 1941
Additional records: GEORGIA, Fannin Co., July (A)3;Habersham Co., June (A), July (A),; Lump-
kin Co., July (A)'; Rabun Co., August (N,A). NORTH CAROLINA, Jackson Co., August (N,A);
Macon Co., June (A), July and August (N,A), September (A); Swain Co., August (N).
SouTH CAROLINA, Oconee Co., October (N)'. TENNESSEE, Sevier Co., July (A).
Isonychia bicolor (Walker)
Physiographic provinces: CP
Previously published state records: N. C., NTH, 1935 (as I. albomanicata); S. C., Bks, 1953.
Additional records: Mississippi, Lauderdale Co., June (A).
Isonychia christina Traver
Physiographic provinces: ILP
Previously published state records: None
Additional records: TENNESSEE, Cheatham Co., July (A); Hickman Co., July (A).
Isonychia circe Traver
Physiographic provinces: P, BR, VR, AP
Previously published state records: Ga., NTH, 1935; Ala., Try., 1937.
Additional records: ALABAMA, Bibb Co., June (A); Walker Co., June (N,A). GEORGIA, Chero-
kee Co., August (A); Coweta Co., June (A)3; Fulton Co., August (A)3; Habersham Co.,
August (A)'; Lumpkin Co., August (N,A); Walton Co., August (A)3.
Isonychia diverse Traver
Physiographic provinces: VR
Previously published state records: Tenn., NTH, 1935.
Additional records: None.
Isonychia fattigi Traver
Physiographic provinces: CP, P, BR, VR
Previously published state records: Ala., Try., 1937; Fla., Sch, 1967; Ga., NTH, 1935; Tenn.,
WB, 1949.
Additional records: ALABAMA, Bibb Co., June (N,A); Macon Co., June (A); Perry Co., June
(A). FLORIDA, Escambia Co., August (A); Gadsden Co., June (A); Santa Rosa Co., June
(A). GEORGIA, Baker Co., March, April, and September (A); Cherokee Co., June (A); De-
catur Co., June (A), November (N); Fulton Co., June (A)'; Lumpkin Co., July (A); Walton
Co., June (N,A)3; White Co., July (A)'; Whitfield Co., June (A)3. Mississippi, Itawamba
Co., July (A); Lauderdale Co., July (N,A). TENNESSEE, Cocke Co., June (A); Sevier Co.,
August (A).
Isonychia georgiae McDunnough
Physiographic provinces: P, BR.
Previously published state records: Ga., McD., 1931, NTH, 1935; N.C., Wy., 1968.
Additional records: GEORGIA, Cherokee Co., (N,A); Rabun Co., June (A)3; Union Co.4 June
(A)3; White Co., July (A)'.
Isonychia harper Traver
Physiographic provinces: BR.
Previously published state records: none.
Additional records: NORTH CAROLINA, Macon Co., August (A).


Vol. 22, No. 1







BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


Isonychia matilda Traver
Physiographic provinces: BR.
Previously published state records: none.
Additional records: NORTH CAROLINA, Macon Co., August (A).
Isonychia notata Traver
Physiographic provinces: BR.
Previously published state records: N. C., NTH, 1935, Try., 1937. H, 1941
Additional records: GEORGIA, Rabun Co., July (N). NORTH CAROLINA, Swain Co., June (A).
Isonychia obscura Traver
Physiographic provinces: BR.
Previously published state records: N. C., NTH, 1935; Tenn., WB, 1949.
Additional records: NORTH CAROLINA, Haywood Co., July (A) TENNESSEE, Greene Co.,
May (A).
Isonychia pacoleta Traver
Physiographic provinces: BR.
Previously published state records: N. C., NTH, 1935; Tenn., WB, 1949.
Additional records: TENNESSEE, Greene Co., March (N).

Isonychia pictipes Traver
Physiographic provinces: CP, P, BR.
Previously published state records: Fla., Ber., 1950, 1958, Sch, 1967; Ga., NTH, 1935.
Additional records: ALABAMA, Dallas Co., June (A); Escambia Co., June (A); Macon Co.,
June (A); Perry Co., June (N,A); Washington Co., May (A). FLORIDA, Gadsden Co., April
(N,A), June (A); Gilchrist Co., October (A); Liberty Co., March, April, and June (N).
GEORGIA, Baker Co., March (A); Cherokee Co., June and August (A); Decatur Co., March
and April (N), June (A); Walton Co., June (A)'. MississiPPi, Itawamba Co., July (A);
Lowndes Co., June (A); Monroe Co., July (A); Pike Co., June (A). SouTH CAROLINA,
Oconee Co., August (A).
Isonychia rufa McDunnough
Physiographic provinces: CP.
Previously published state records: none.
Additional records: MISSISSIPPI, Lawrence Co., August (A); Pike Co., August (A).

Isonychia sadleri Traver
Physiographic provinces: BR.
Previously published records: N. C., Try., 1937; Tenn., WB, 1949.
Additional records: NORTH CAROLINA, Watauga Co., June (N). TENNESSEE, Greene Co.,
March, April (N), May (N,A); Sevier Co., June (A); Unicoi Co., June (A).
Isonychia sayi Burks
Physiographic provinces: CP.
Previously published state records: none.
Additional records: FLORIDA, Okaloosa Co., May (A); Santa Rosa Co., May (A); Walton Co.,
April (A).
Isonychia serrata Traver
Physiographic provinces: BR, AP.
Previously published state records: N. C., NTH, 1935, H, 1941
Additional records: GEORGIA, Rabun Co., June (A)'. NORTH CAROLINA, Macon Co., August
(A). TENNESSEE, Fentress Co., June, July (A).

Isonychia sicca sicca (Walsh)
Physiographic provinces: VR.
Previously published state records: none.
Additional records: GEORGIA, Whitfield Co., August (A)'.








BULLETIN FLORIDA STATE MUSEUM


Isonychia similis Traver
Physiographic provinces: P.
Previously published state records: N. C. and S. C., NTH, 1935; N. C., H, 1941
Additional records: SOUTH CAROLINA, Greenville Co., no date (A)*.
Isonychia species
Physiographic provinces: CP, P, BR, VR, AP, ILP.
Records: ALABAMA, Dallas Co., June (N); DeKalb Co., April, July (N); Escambia Co., April
(N); Henry Co., June (N); Randolph Co., June (N); St. Clair Co., July (N). GEORGIA, Baker
Co., May, October (N); Baldwin Co., May, July (N)3; Barrow Co., July (N,A); Bartow Co.,
May (N)3; Bibb Co., June (N)3, July (N); Burke Co., June (N); Calhoun Co., June (A)3;
Carroll Co., March (N); Catoosa Co., May, August (N)3; Chattooga Co., June (N)3; Chero-
kee Co., June, August (N,A); Clarke Co., April (N)3; Cobb Co., November (A); Dawson
Co., June (N,A); Decatur Co., June (N,A), August (N); Early Co., February (N); Effingham
Co., July (N); Emmanuel Co., May (N)3; Fannin Co., May, June (N)3; Floyd Co., June
(N); Forsyth Co., June (N); Fulton Co., May, June, July, September (A)3; Gilmer Co.,
June (N)3; Gordon Co., May, June (A)3; Green Co., August (N); Hall Co., June, July (N)3;
Hart Co., September (N); Heard Co., July (N); Jackson Co., July (N); Johnson Co., August
(N); Lowndes Co., March (N); Lumpkin Co., April (N), June (A)3, July (A), August (A),
September (N)3; Madison Co., July (N); Monroe Co., July (N); Murray Co., August (N);
Muscogee Co., April (N)3; Pickens Co., June (N), July (N)3; Rabun Co., July (N); Screven
Co., July (N); Stephens Co., July (A)3; Towns Co., May (N)3; Troup Co., July (N); Union
Co., July (N)3; Walker Co., April, July (N); Walton Co., June (N)3; White Co., June, July,
August (N)3; Wilkes Co., July (N); Worth Co., November (N). LOUISIANA, Natchitoches
Par., June (A); Red River Par., June (A); Tangipahoa Par., August (N); Vernon Par., June
(N). MIssissiPPI, Harrison Co., August (N); Lamar Co., August (N); Marion Co., August
(N). NORTH CAROLINA, Ashe Co., July (N,A); Avery Co., February, August (N); Caldwell
Co., July, August (N); Caswell Co., July (N); Davidson Co., July (N); Forsyth Co., June (N);
Graham Co., August (N); Haywood Co., August (A); Henderson Co., June (N); Jackson
Co., July, August (N); McDowell Co., April (N); Macon Co., June, July, August (N.A);
Polk Co., July (N); Randolf Co., July (N); Swain Co., April, May, June, July, August (N);
Transylvania Co., May, June, July (N), August (N,A); Union Co., July (N); Wake Co.,
February, May, December (N); Watauga Co., February, June, August (N), July (A); Wilkes
Co., June (N); Yadkin Co., July (N). SOUTH CAROLINA, Aiken Co., July (N); Anderson Co.,
July (A); Edgefield Co., July (N); Fairfield Co., April (N); Greenville Co., July (N), August
(N,A); Lancaster Co., June (N); Laurens Co., April (N); Newberry Co., April (N); Mc-
Cormick Co., July (N); Oconee Co., August (N); Pickens Co., August (N). TENNESSEE,
Blount Co., April, June (N); Franklin Co., April (N); Greene Co., May (N,A); Hamilton
Co., May, December (N); Henderson Co., June (N); Lincoln Co., July (N); Morgan Co.,
July (N); Robertson Co., June (N); Rutherford Co., July (N); Sevier Co., April-August,
December (N); Sumner Co., June (N).
Isonychia thalia Traver
Physiographic provinces: P, BR.
Previously published state records: N. C., NTH, 1935, H, 1941
Additional records: GEORGIA, Rabun Co., July (A)': NORTH CAROLINA, Rockingham Co.,
July (N,A). South Carolina, Oconee Co., July (A).
Isonychia tuscalanensis Berner
Physiographic provinces: BR.
Previously published state records: Tenn., WB., 1949.
Additional records: none.


Vol. 22, No. 1







BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


FAMILY METRETOPODIDAE
SUBFAMILY METRETOPODINAE
GENUS Siphloplecton CLEMENS

Siphloplecton basale (Walker)
Physiographic provinces: CP
Previously published state records: N.C., NTH, 1935
Additional records: SOUTH CAROLINA, Barnwell Co., April (A)*
Siphloplecton signatum Traver
Physiographic provinces: BR
Previously published state records: N. C., NTH, 1935
Additional records: GEORGIA, Rabun Co., April (A)3
Siphloplecton species
Physiographic provinces: CP
Additional records: FLORIDA, Okaloosa Co., May (A). GEORGIA, Lowndes Co., March (N,A)
Siphloplecton speciosum Traver
Physiographic provinces: CP, P.
Previously published state records: Fla., Ber., 1950 and 1958, PJ, 1973, Sch, 1967; Ga., NTH,
1935.
Additional records: ALABAMA, Baldwin Co., December (N). FLORIDA, Escambia Co., January
(N); Gadsden Co., April (A). GEORGIA, Bibb Co., April (A)3; Fayette Co., March (A)';
Lowndes Co., March (N,A).



FAMILY BAETIDAE
SUBFAMILY BAETINAE
GENUS Baetis LEACH

Baetis amplus (Traver)
Physiographic provinces: CP, P, BR, VR, AP.
Previously published state records: N. C., Tray., 1932 and 1937.
Additional records: ALABAMA, Bibb Co., June (N). GEORGIA, Dougherty Co., September (N)2;
Lumpkin Co., August (N); Rabun Co., May (N). MIssIssIPPI, Lauderdale Co., June (N).
NORTH CAROLINA, Clay Co., April (N). TENNESSEE, Cumberland Co., April (N,A); Marion
Co., April (N).
Baetis australis Traver
Physiographic provinces: CP, P, AP.
Previously published state records: Ala., Ber., 1958; Fla., Ber., 1950 and 1958, PJ, 1973, Sch,
1967; Ga., NTH, 1935, Ber., 1958; La., Bick, 1953; N. C., Trav., 1932.
Additional records: ALABAMA, Baldwin Co., June (N); Blount Co., June (N); Escambia Co.,
April (N); Houston Co., February (N); Mobile Co., June (N). FLORIDA, Alachua Co.,
December (A); Columbia Co., November (N); Gulf Co., April (A); Hamilton Co., February
(N); Levy Co., November (N); Madison Co., March (N); Walton Co., April (N), April
(A). GEORGIA, Early Co., February (N); Peach Co., April (N). LOUISIANA, Tangipahoa
Par., June (N). MIssISSIPPI, Stone Co., August (N). NORTH CAROLINA, Rockingham Co.,
July (N). SOUTH CAROLINA, Aiken Co., May (A)2, July (A)2.
Baetis cingulatus McDunnough
Physiographic provinces: BR.
Previously published state records: N. C., Trav., 1937.
Additional records: NORTH CAROLINA, Jackson Co., August (A); Macon Co., August, Sep-
tember (A); Swain Co., June (A).







BULLETIN FLORIDA STATE MUSEUM


Baetis ephippiatus Traver
Physiographic provinces: CP, P, VR, AP.
Previously published state records: Fla., Ber., 1950 and 1958, PJ, 1973, Sch, 1967; Ga., NTH,
1935, Ber., 1958.
Additional records: ALABAMA, Baldwin Co., June (N); Coosa Co., June (N); Escambia Co.,
June (N); Macon Co., June (N); Mobile Co., June (N,A); Perry Co., June (N); Walker Co.,
June (N,A); Washington Co., May (N,A), August (N); Winston Co., August (N). FLORIDA,
Bay Co., April (N); Escambia Co., October (N); Santa Rosa Co., August (N). GEORGIA,
Baker Co., May (N); Floyd Co., June (A)', August (A)3; Muscogee Co., June, July (A).
LOUISIANA, Tangipahoa Par., August (N). MIssIssIPPI, Amite Co., June (N); Calhoun Co.,
July (N); Clay Co., July (N); Copiah Co., August (N); Holmes Co., July (N); Lamar Co.,
May (N), August (N,A); Lawrence Co., August (N); Marion Co., May (A), August (N);
Monroe Co., July (N); Pike Co., June (A), August (N); Smith Co., August (N); Sunflower
Co., July (N). SOUTH CAROLINA, Aiken Co., July (N); Darlington Co., July (A).
Baetis flavistriga McDunnough
Physiographic provinces: BR.
Previously published state records: Tenn., WB, 1949.
Additional records: NORTH CAROLINA, Macon Co., August (A). TENNESSEE, Greene Co., May
(A); Sevier Co., July (A).
Baetis frondalis McDunnough
Physiographic provinces: BR.
Previously published state records: Tenn., WB, 1949.
Additional records: Greene Co., April, May, September (A).
Baetis hageni Eaton
Physiographic provinces: BR, VR, AP.
Previously published state records: Tenn., WB, 1949 (listed as B. parvus)
Additional records: TENNESSEE, Claiborne Co., August (N); Cumberland Co., July (A); Greene
Co., September (A).
Baetis harti McDunnough
Physiographic provinces: CP.
Previously published state records: none.
Additional records: TENNESSEE, Benton Co., June (A).
Baetis incertans McDunnough
Physiographic provinces: BR, VR.
Previously published state records: N. C., Trav., 1937; Tenn., WB, 1949.
Additional records: TENNESSEE, Claiborne Co., July (N,A); Greene Co., March, April (A);
Sevier Co., August (N), October (A); Unicoi Co., August (A).
Baetis intercalaris McDunnough
Physiographic provinces: CP, P, VR, AP, ILP.
Previously published records: Ala., Ber., 1958; Fla., Ber., 1950, 1958, PJ, 1973, Sch, 1967;
Ga., Ber., 1958.
Additional records: ALABAMA, Bibb Co., June (N,A); Blount Co., June (A); Chilton Co., June
(N); Coosa Co., June (N); Covington Co., July (N); Dale Co., June (N); Dallas Co., April
(A), June (N); Elmore Co., June (N,A); Houston Co., June (N); Lauderdale Co., June (A);
Macon Co., April, June (N); Shelby Co., June (N); Tuscaloosa Co., June (N). FLORIDA,
Escambia Co., October (N,A); Gadsden Co., April (N); Liberty Co., March, June (N).
GEORGIA, Baker Co., November (N); Bibb Co., July (N); Cherokee Co., August (A); De-
catur Co., March (N); Grady Co., November (N); Muscogee Co., June, July (A). LOUISIANA,
Tangipahoa Co., August (N). MISSISSIPPI, Jasper Co., April (N); Lafayette Co., July (N);
Lauderdale Co., July (N,A); Perry Co., August (N). NORTH CAROLINA, Macon Co., August
(A). TENNESSEE, Campbell Co., August (A); Davidson Co., October (N).
Baetis levitans McDunnough
Physiographic provinces: BR.


Vol. 22, No. 1








BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


Previously published state records: Tenn., WB, 1949.
Additional records: TENNESSEE, Greene Co., May, September (A).
Baetis pallidulus McDunnough
Physiographic provinces: BR.
Previously published state records: none.
Additional records: TENNESSEE, Sevier Co., July (A).
Baetis propinquus (Walsh)
Physiographic provinces: CP, VR.
Previously published state records: Ala., Trav., 1937; Fla., Ber., 1950 and 1958, Sch, 1967.
Additional records: ALABAMA, Dallas Co., June (A). FLORIDA, Gadsden Co., June (A). Missis-
SIPPI, Leflore Co., July (A); Lowndes Co., June (A). SOUTH CAROLINA, Darlington Co.,
July (A). TENNESSEE, Hamilton Co., August (A).
Baetis pygmaeus (Hagen) group
Physiographic provinces: BR, VR, AP.
Previously published state records: none
Additional records: NORTH CAROLINA, Jackson Co., August (N); Macon Co., August (N);
Swain Co., April (N). TENNESSEE, Claiborne Co., August (N); Greene Co., March, April
(N); Morgan Co., July (N); Van Buren Co., July (N).
Baetis species
Physiographic provinces: CP, P, BR, VR, AP, ILP.
Additional records: ALABAMA, DeKalb Co., April, May, July (N); Tuscaloosa Co., April-
July (N). GEORGIA, Baker Co., October (N); Burke Co.,June (N); Carroll Co., March
(N); Cherokee Co., June (N,A), July, October (N), August (N,A); Coweta Co., June (A)3;
Dawson Co., June (N); Decatur Co., March, August, October (N), June (A); Floyd Co.,
June (N); Forsyth Co., June (N); Habersham Co., April (A)'; Hart Co., November (N);
Lumpkin Co., April, August (N); Monroe Co., July (N); Muscogee Co., June (N); Peach
Co., April (N); Pickens Co., June (N); Rabun Co., April, July, August (N); Toombs Co.,
August (N); Walker Co., April (N); White Co., August (A); Wilkes Co., (N). MississIPPI,
Harrison Co., August (N); Lauderdale Co., July (N,A); Marion Co., August (A); Perry Co.,
August (N). NORTH CAROLINA, Avery Co., April (N); Buncombe Co., May (N); Clay Co.,
April (N); Jackson Co., June (N,A), August (N); Macon Co., May (N)', July, August (N);
Swain Co., April-July (N); Transylvania Co., March, July (N); Wake Co., February, May
(N); Yancey Co., May, July, August (N). TENNESSEE, Blount Co., April (N); Franklin Co.,
April (N); Greene Co., May (A), August (N); Hamilton Co., May, December (N); Hender-
son Co., June (N); Lincoln Co., July (N); Morgan Co., July (N); Robertson Co., June (N);
Rutherford Co., July (N); Sevier Co., May, June, December (N); Sumner Co., June (N);
Van Buren Co., July (N).
Baetis spiethi Berner
Physiographic provinces: CP, P, VR, AP.
Previously published state records: Fla., Ber., 1950 and 1958, PJ, 1973, Sch, 1967; Ga., Ber.,
1958.
Additional records: ALABAMA, Baldwin Co., June (N); Bibb Co., June (N); Conecuh Co.,
June (N); Dallas Co., June (N); Escambia Co., April, June (N); Mobile Co., June (N,A);
Tuscaloosa Co., June (N); Walker Co., June (N,A). FLORIDA, Alachua Co., February (A);
Baker Co., February (N); Clay Co., January (N); Columbia Co., November (A); Escambia
Co., October (N); Hamilton Co., March (N); Hernando Co., February (A); Jackson Co.,
April (A); Liberty Co., March (A); Madison Co., January (N); Walton Co., April (N).
GEORGIA, Baker Co., May (N), October (N); Charlton Co., February (N); Decatur Co.,
June (N); Early Co., February (N); Grady Co., November (N); Lowndes Co., March (N,A);
Macon Co., November (N); Peach Co., April (N,A); Worth Co., November (N). LOUISIANA,
St. Tammany Par., November (N); Tangipahoa Par., June (N). MIssIssIPPI, Amite Co.,
June (N); Harrison Co., December (N); Jasper Co., April (N); Lafayette Co., July (N,A);
Lamar Co., May (N); Marion Co., August (N); Perry Co., August (N); Pike Co., August
(N); Stone Co., August (N). SOUTH CAROLINA, Aiken Co., July (N); Lexington Co., April
(N); Orangeburg Co., October (N).







BULLETIN FLORIDA STATE MUSEUM


Baetis spinosus McDunnough
Physiographic provinces: CP, P, BR, VR, AP.
Previously published state records: Ala., Ber., 1958; Fla., Ber., 1950 and 1958, PJ, 1973, Sch,
1967; Ga., NTH, 1935, Ber., 1958.
Additional records: ALABAMA, Baldwin Co., June (N); Bibb Co., June (N); Chilton Co., June
(N); Coosa Co., June (N); Covington Co., July (N); Dale Co., June (N); Dallas Co., June
(N); Elmore Co., June (N); Escambia Co., April (A), June (N); Henry Co., June (N,A);
Houston Co., June (N); Macon Co., April (N); Marion Co., June (N); Mobile Co., June
(N,A); Pike Co., June (N); Randolf Co., June (N); Sumter Co., April (N); Tuscaloosa Co.,
June (N); Washington Co., June (N). FLORIDA, Alachua Co., January, December (A), Octo-
ber (N); Baker Co., February (N); Columbia Co., November (A); Escambia Co., October
(N); Gadsden Co., April (N), June (A); Hamilton Co., November (N); Levy Co., November
(N); Liberty Co., March (N,A); Madison Co.. January (N); Walton Co., April (N). GEORGIA,
Barrow Co., July (N); Berrian Co., March (N); Bibb Co., July (N); Burke Co., August (N);
Charlton Co., February (N); Cherokee Co., June (N,A); Decatur Co., March, June (N);
Dougherty Co., November (N); Early Co., February (N); Fulton Co., June (A)'; Lowndes
Co., March (N); Macon Co., April (N); Muscogee Co., June (N); Worth Co., November
(N). LOUISIANA, Tangipahoa Par., August (N). MissIssIPPI, Claiborne Co., April (N); Jasper
Co., April (N); Lafayette Co., July (N); Marshall Co., July (N); Monroe Co., July (N); Perry
Co., August (N); Rankin Co., April (N). SOUTH CAROLINA, Aiken Co., May (A)2; Allendale
Co., May (N)'; Edgefield Co., April (N); Lancaster Co., June (N); Lexington Co., April
(N); Orangeburg Co., April (A); Pickens Co., August (N).
Baetis vagans McDunnough
Physiographic provinces: BR.
Previously published state records: none.
Additional records: NORTH CAROLINA, Jackson Co., August (A); Macon Co., August (A).

GENUS Callibaetis EATON
Callibaetis floridanus Banks
Physiographic provinces: CP.
Previously published state records: Ala., Ber., 1958; Fla., Banks, 1900, Ber., 1950 and 1958,
Sch, 1967; Ga., Ber. 1958.
Additional records: ALABAMA, Houston Co., June (N,A). FLORIDA, Alachua Co., January
(A), May (A), June (A), August (A), October (A); Broward Co., November (A); Collier
Co., April (A), September (A); Dade Co., June (A); Hernando Co., February (A); Indian
River Co., April (A); Jackson Co., April (A); Monroe Co., February (A), March (A), October
(A); December (A); Polk Co., April (A); St. Lucie Co., October (A). GEORGIA, Decatur Co.,
April (N), August (A)'; Thomas Co., May (N). LOUISIANA, East Baton Rouge Par., June
(A); Natchitoches Par., July (A); Orleans Par., January, December (A); Plaquemines Par.,
December (A); St. Tammany Par., January (A). Mississippi, Sunflower Co., July (A);
Tallahatchee Co., July (A).
Callibaetis fluctuans (Walsh)
Physiographic provinces: CP.
Previously published state records: None
Additional records: LOUISIANA, Lafayette Co., February, July (A).
Callibaetis pretiosus Banks
Physiographic provinces: CP.
Previously published state records: Fla., Ber., 1950, PJ, 1973.
Additional records: ALABAMA, Baldwin Co., June (N). FLORIDA, Alachua Co., April (A); Bre-
vard Co., March (A); Collier Co., April (A); Hernando Co., February (A); Highlands Co.,
May (A); Lee Co., September (A); Leon Co., April (A); Monroe Co., January (A); Polk
Co., February (A), April (A); Seminole Co., February (N,A). GEORGIA, Berrian Co., March
(N); Echols Co., February (N); Lowndes Co., October (N); Seminole Co., June (N). NORTH
CAROLINA, Wayne Co., September (A). TENNESSEE, Shelby Co., June (A).


Vol. 22, No. 1








BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


Callibaetis species
Physiographic provinces: CP, P, BR, VR, ILP.
Additional records: ALABAMA, Baldwin Co., December (N); Bibb Co., June (N,A); Houston
Co., June (A); Macon Co., June (N); Mobile Co., June (N); Pike Co., June (N); Shelby
Co., June (N). GEORGIA, Burke Co., August (N); Calhoun Co., July (N); Camden Co.,
September (N); Clayton Co., September (A)3; Decatur Co., March (A); Johnson Co.,
August (N); Lowndes Co., February (A), March (A); Richmond Co., July (A); Thomas
Co., June (N). LOUISIANA, Acadia Par., April (N); Ascension Par., December (N);
East Feliciana Par., January (N); Lafayette Par., May (N); Natchitoches, April, June, July
(A); Orleans Par., June (N). NORTH CAROLINA, Henderson Co., July (N); Macon Co., June
(A), July (N,A), August (A); Transylvania Co., May (N). SOUTH CAROLINA, Aiken Co.,
May, June, July (A)2; Marlboro Co., July (N). TENNESSEE, Davidson Co., June (N); Greene
Co., March (N); Humphreys Co., July (N); Williamson Co., May (N).


GENUS Centroptilum EATON
Centroptilum fragile McDunnough
Physiographic provinces: BR.
Previously published state records: None
Additional records: NORTH CAROLINA, Jackson Co., August (A).
Centroptilum hobbsi Berner
Physiographic provinces: CP, VR, AP.
Previously published state records: Ala., Ber., 1958; Fla., Ber., 1950 and 1958, Sch, 1967;
Ga., Ber., 1958.
Additional records: ALABAMA, Barbour Co., June (N); Bibb Co., June (N); Coosa Co., June
(N); Macon Co., June (N); Perry Co., June (A); Walker Co., June (A). FLORIDA, Escambia
Co., August (A); Liberty Co., June (A); Santa Rosa Co., August (N,A). GEORGIA, Baker
Co., May (N), November (N). MISSISSIPPI, Copiah Co., August (N); Lamar Co., August (A).
Centroptilum intermediate McDunnough
Physiographic provinces: P, BR.
Previously published state records: None
Additional records: GEORGIA, Cherokee Co., June (N,A). NORTH CAROLINA, Haywood Co.,
July (A); Macon Co., September (A).
Centroptilum simile McDunnough
Physiographic provinces: BR.
Previously published state records: None.
Additional records: NORTH CAROLINA, Swain Co., June (A).
Centroptilum species
Physiographic provinces: CP, P, BR, VR, AP, ILP.
Additional records: ALABAMA, Calhoun Co., July (A); DeKalb Co., May (N), July (N). FLORIDA,
Okaloosa Co., May (A). GEORGIA, Cherokee Co., June (N), August (A); Dawson Co., June
(N); Johnson Co., August (N); Monroe Co., July (N); Towns Co. June (A)3. LOUISIANA,
Tangipahoa Par., October (N). MississIPpi, Forrest Co., August 22, 1954 (A); Lamar Co.,
May (N), August (A); Marion Co., August (N). NORTH CAROLINA, Caldwell Co., July (A);
Cumberland Co., July (A); Graham Co., August (N); Granville Co., July (A); Jackson Co.,
August (N,A); Macon Co., June (A), August (N,A); Person Co., July (N,A); Robson Co.,
August (A); Swain Co., August (N). SOUTH CAROLINA, Aiken Co., July (A); Edgefield Co.,
April (N); Lexington Co., April (N); Oconee Co., August (N). TENNESSEE, Benton Co.,
June (A); Cooke Co., June (A); Cumberland Co., July (N); Greene Co., May (A); Hamilton
Co., August (A); Houston Co., June (N); Lincoln Co., July (N); Robertson Co., June (N);
Sevier Co., August (N,A).
Centroptilum viridocularis Berner
Physiographic provinces: CP, P, VR.


1977








BULLETIN FLORIDA STATE MUSEUM


Previously published state records: Fla., Ber., 1950 and 1958, PJ, 1973, Sch, 1967; Ga., Ber.,
1958.
Additional records: ALABAMA, Baldwin Co., June (N); Bibb Co., June (N); Elmore Co., June
(N); Macon Co., June (N). FLORIDA, Alachua Co., June (A); Escambia Co., October (N);
Liberty Co., June (N); Polk Co., January-April (A), June-August (A), October (A), May-
August (N). GEORGIA, Decatur Co., June (A). MISSISSIPPI, Claibome Co., April (N).

GENUS Cloeon LEACH
Cloeon alamance (Traver)
Physiographic provinces: BR, AP.
Previously published state records: Ga., NTH, 1935; N. C., Tray., 1932; Tenn., WB, 1949
(as Neocloeon alamance).
Additional records: NORTH CAROLINA, Macon Co., July and August (N,A); Swain Co., June
(A), August (A). TENNESSEE, Cumberland Co., April (N); Greene Co., March (N); Sevier
Co., May (A)'.
Cloeon rubropictum McDunnough
Physiographic provinces: CP, VR.
Previously published state records: Fla., Ber., 1950, PJ, 1973, Sch, 1967.
Additional records: ALABAMA, Covington Co., July (N); Dallas Co., June (N); Macon Co.,
June (N); Shelby Co., June (N). GEORGIA, Baker Co., May (N); Early Co., February (N).
MISSISSIPPI, Claibome Co., April (A).
Cloeon simplex McDunnough
Physiographic provinces: CP
Previously published state records: None.
Additional records: LOUISIANA, East Baton Rouge Par., June (A).
Cloeon species
Physiographic provinces: CP, P, BR, VR, AP, ILP
Additional records: ALABAMA, Baldwin Co., June (N); Bibb Co., June (N); Calhoun Co., July
(A); Covington Co., July (N); Cullman Co., June (N); Dallas Co., June (A); DeKalb Co.,
July (N); Elmore Co.; June (N); Houston Co., February (N); Lauderdale Co., June (N);
Macon Co., April (N); Perry Co., June (A); Randolf Co., June (N); St. Clair Co., June (N).
GEORGIA, Bibb Co., (N); Early Co., February (N); Floyd Co., June (A); Muscogee Co.,
June (N); Richmond Co., July (A). LOUISIANA, Washington Co., August (N). MississipPI,
Forrest Co., August (N,A); Harrison Co., August (N); Lafayette Co., July (N); Lamar Co.,
August (A). NORTH CAROLINA, Cabarrus Co., July (N,A); Davidson Co., (N); Graham Co.,
August (N); Jackson Co., June (A), August (N,A), September (A); Lincoln Co., July (N);
Macon Co., June (A), July-August (A), August (N); Person Co., July (N,A); Randolf Co.,
July (N); Swain Co., August (A); Union Co., July (A). SorrTH CAROLINA, Edgefield Co.,
April (N); Fairfield Co., April (N); Laurens Co., April (N,A) TENNESSEE, Benton Co.,
June (N,A); Fentress Co., May (A); Greene Co., April (A), May (N,A); Lincoln Co., April
(N); Morgan Co., July (N,A); Polk Co., August (A); Robertson Co., June (N); Sevier Co.,
July (A); Van Buren Co., July (N).

GENUS Heterocloeon McDUNNOUGH
Heterocloeon bemeri (Mfiller-Liebenau)
Physiographic provinces: P
Previously published state records: Ga., M.-L., 1974.
Additional records: None.
Heterocloeon curiosum (McDunnough)
Physiographic provinces: CP, P, BR.
Previously published state records: Ga., M.-L., 1974 (as Rheobaetis traverae); N. C., Ber.,
1949.


Vol. 22, No. 1







BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


Additional records: GEORGIA, Baker Co., (N); Cherokee Co., June (N). NORTH CAROLINA,
Macon Co., July (A), August (N,A). SouTH CAROLINA, Spartanburg Co., August (N).
Heterocloeon peters (Mfiller-Liebenau)
Physiographic provinces: P
Previously published state records: Ga., M.-L., 1974.
Additional records: None.


GENUS Paracloeodes DAY

Paracloeodes species
Physiographic provinces: CP, P, VR, AP.
Previously published state records: None.
Additional records: ALABAMA, Bibb Co., June (N); Coosa Co., June (N); DeKalb Co., July
(N); Perry Co., June (N). GEORGIA, Bibb Co., July (N,A); Monroe Co., July (N). MIssis-
sipPI, Calhoun Co., July (N); Copiah Co., August (N); Franklin Co., June, August (N);
Smith Co., August (N).


GENUS Pseudocloeon KLAPALEK
Pseudocloeon alachua Berner
Physiographic provinces: CP.
Previously published state records: Fla., Ber., 1950, Sch, 1967.
Additional records: FLORIDA, Levy Co., November (N). MIssIssIPPI, Lowndes Co., June (A).
Pseudocloeon bimaculatus Berner
Physiographic provinces: CP, P.
Previously published state records: Fla., Ber., 1950, Sch, 1967.
Additional records: ALABAMA, Baldwin Co., June (N); Escambia Co., June (N,A); Macon Co.,
June (A); Mobile Co., April (N), June (N,A). FLORIDA, Santa Rosa Co., August (N).
GEORGIA, Cherokee Co., June (N); Early Co., February (N); Monroe Co., July (N); Mus-
cogee Co., June (A); Peach Co., April (N). MississiPPi, Lawrence Co., August (N); Perry
Co., August (N); Stone Co., August (N); Sunflower Co., July (N,A). SOUTH CAROLINA,
Edgefield Co., April (N); Lexington Co., April (N).
Pseudocloeon carolina (Banks)
Physiographic provinces: P, BR, VR, AP, ILP.
Previously published state records: N. C., Banks, 1924, NTH, 1935, Trav., 1937, H, 1941.
Additional records: ALABAMA, Bibb Co., June (N); Blount Co., June (N); Coosa Co., June (N);
Lauderdale Co., June (N); Walker Co., June (N). GEORGIA, Lumpkin Co., April (A)'; Rabun
Co., May (N); Walker Co., April (N). NORTH CAROLINA, Jackson Co., June (N); Macon Co.,
August (N,A); Person Co., July (N); Polk Co., July (N); Swain Co., April (A), June (N,A),
July (A), July (N), August (N). TENNESSEE, Blount Co., June and December (N); Robertson
Co., June (N); Sevier Co., May, August, October (N).
Pseudocloeon cingulatum McDunnough
Physiographic provinces: BR, ILP.
Previously published state records: None
Additional records: NORTH CAROLINA, Jackson Co., August (A). TENNESSEE, Lincoln Co.,
July (A).
Pseudocloeon dubium (Walsh)
Physiographic provinces: CP, BR, VR, AP.
Previously published state records: Ala., Trav., 1937; N. C., NTH 1935, Tray., 1937.
Additional records: ALABAMA, Bibb Co., June (N); Covington Co., July (N); Marion Co.,
June (N); Perry Co., June (N). GEORGIA, Habersham Co., April (A)'. NORTH CAROLINA,
Jackson Co., August (A); McDowell Co., April (N,A).







BULLETIN FLORIDA STATE MUSEUM


Pseudocloeon etowah Traver
Physiographic provinces: VR
Previously published state records: Ga., NTH, 1935.
Additional records: GEORGIA, Catoosa Co., May (A)'; Floyd Co., June (A).
Pseudocloeon futile McDunnough
Physiographic provinces: CP.
Previously published state records: None.
Additional records: MississiPPi, Lamar Co., August (A).
Pseudocloeon parvulum McDunnough
Physiographic provinces: CP, P, BR, VR.
Previously published state records: Fla., Ber., 1950 and 1958, Sch, 1967.
Additional records: ALABAMA, Bibb Co., May (N); Coosa Co., June (N); Macon Co., April
(N). GEORGIA; Richmond Co., August (A). MississiPPi, Amite Co., June (N); Claibome
Co., April (N); Copiah Co., August (N); Jasper Co., April (N). SOUTH CAROLINA, Edgefield
Co., April (N); Fairfield Co., April (N); Greenville Co., August (N); Laurens Co., April
(N); Newberry Co., April (N).
Pseudocloeon punctiventris McDunnough
Physiographic provinces: CP, P, BR, VR
Previously published state records: Fla., Ber., 1950 and 1958, PJ, 1973, Sch, 1967; Ga., Ber.,
1958.
Additional records: ALABAMA, Blount Co., June (N); Coosa Co., June (N); Dekalb Co., April
(N); Macon Co., April (N); Marion Co., June (N); Sumter Co., April (N); Walker Co.,
June (N,A). FLORIDA, Clay Co., January (N); Madison Co., January (N). GEORGIA, Baker
Co., May (N), November (N,A); Berrian Co., March (N); Chattahoochee Co., April (N);
Cherokee Co., June (N,A). Jones Co., April (N); Lowndes Co., March (N); Lumpkin
Co., August (N,A); Muscogee Co., June (N). MississiPPi, Monroe Co., July (N,A). NORTH
CAROLINA, Forsyth Co., June (N); Person Co., July (N). SOUTH CAROLINA, Edgefield Co.,
July (N); Greenville Co., August (N); Orangeburg Co., April (A); Richland Co., July (A).
TENNESSEE, Sevier Co., August (A).
Pseudocloeon rubrolaterale McDunnough
Physiographic provinces: BR.
Previously published state records: None.
Additional records: TENNESSEE, Sevier Co., July (A), August (A).
Pseudocloeon species
Physiographic provinces: CP, P, BR, VR, AP.
Additional records: ALABAMA, Dekalb Co., May (N). FLORIDA, Okaloosa Co., May (A).
GEORGIA, Burke Co., June (N); Cherokee Co., June (N,A), July and August (N); Dawson
Co., June (N,A); Fulton Co., April (N); Lowndes Co., March (A); Lumpkin Co., April
(N); Rabun Co., August (N); Richmond Co., April (A). LOUISIANA, Natchitoches Par.,
March (N). MississiPPi, Lamar Co., May (N); Marion Co., August (A). NORTH CAROLINA,
Buncombe Co., May (N); Caldwell Co., July (N); Clay Co., April (N); Davidson Co., July
(N); Graham Co., August (N,A); Jackson Co., June (A), August (N); Macon Co., June and
July (A), August and September (N,A); Orange Co., February (N); Person Co., July (A);
Polk Co., July (N); Randolf Co., July (N); Swain Co., May (N)', June and July (N,A), August
(A), Transylvania Co., March, June, July and August (N); Wake Co., February and May
(N); Yadkin Co., February and July (N). SOUTH CAROLINA, Aiken Co., May-August (A)2,
July (N); Edgefield Co., April (N,A); Fairfield Co., April (N); Greenville Co., August (N);
Greenwood Co., April (A); Laurens Co., April (N); Oconee Co., August (N,A); Orange-
burg Co., April (A), October (N). TENNESSEE, Hamilton Co., May (N); Morgan Co., July
(N); Sevier Co., April-June (N); Van Buren Co., July (N).


Vol. 22, No. 1







BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


FAMILY OLIGONEURIIDAE
SUBFAMILY OLIGONEURIINAE
GENUS Homoeoneuria EATON
Homoeoneuria dolani Edmunds, Berner, and Traver
Physiographic provinces: CP.
Previously published state records: Fla., PJ, 1973; Ga., EBT, 1958; S. C., EBT, 1958.
Additional records: None.
Homoeneuria species
Physiographic provinces: CP, P.
Previously published state records: Fla., EBT, 1958; Miss., EBT, 1958; S. C., EBT, 1958.
Additional records: FLORIDA, Santa Rosa Co., August (N).

FAMILY HEPTAGENIIDAE
SUBFAMILY PSEUDIRONINAE
GENUS Pseudiron McDUNNOUGH

Pseudiron centralis McDunnough
Physiographic provinces: CP
Previously published state records: None
Additional records: ALABAMA, Dallas Co., April (A). FLORIDA, Walton Co., April (A). TEN-
NESSEE, Shelby Co., June (A).
Pseudiron meridionalis Traver
Physiographic provinces: CP, P.
Previously published state records: Fla., PJ, 1973; Ga., NTH, 1935.
Additional records: None
Pseudiron species
Physiographic provinces: CP, BR
Additional records: FLORIDA, Santa Rosa Co., March (N). MississiPPi, Leflore Co., June (A).
SOUTH CAROLINA, Allendale Co., May (N)'. TENNESSEE, Unicoi Co., May (A).

SUBFAMILY HEPTAGENIINAE
GENUS Cinygmula MCDUNNOUGH

Cinygmula species
Physiographic provinces: BR.
Additional records: NORTH CAROLINA, Clay Co., April (N); Macon Co., May (N)'; Swain Co.,
May (N)', April and May (N).
Cinygmula subaequalis (Banks)
Physiographic provinces: BR, AP, ILP.
Previously published state records: N. C., Banks, 1914, Trav., 1937; Tenn., WB, 1949 (Last
two as C. atlantica).
Additional records: GEORGIA, Rabun Co., July (A)3. NORTH CAROLINA, Clay Co., April (N);
McDowell Co., April (N); Swain Co., June (A). TENNESSEE, Cocke Co., April (A); Dick-
son Co., June (N); Houston Co., June (N); Marion Co., April (N); Moore Co., April (N).

GENUS Epeorus EATON
Epeorus dispar (Traver)
Physiographic provinces: BR.
Previously published state records: N. C., Tray., 1933 and 1937, H, 1941; S. C., NTH, 1935.







BULLETIN FLORIDA STATE MUSEUM


Additional records: NORTH CAROLINA, Jackson Co., August (A,N); Macon Co., August (A).
SOUTH CAROLINA, Greenville Co., July (A)*, June (A)'. TENNESSEE, Unicoi Co., August,
October (A).
Epeorus pleuralis (Banks)
Physiographic provinces: BR
Previously published state records: N. C., Tray., 1937 (as E. confusus)
Additional records: GEORGIA, Rabun Co., May (A)'. TENNESSEE, Sevier Co., July (A,N), April,
May (A)3; Unicoi Co., June (A).
Epeorus rubidus (Traver)
Physiographic provinces: BR, VR.
Previously published state records: N. C., Tray., 1933 and 1937 H, 1941; Tenn., WB, 1949.
Additional records: GEORGIA, Lumpkin Co., August (A)'. NORTH CAROLINA, Jackson Co., June
(N,A). SOUTH CAROLINA, Pickens Co., May (A)'. TENNESSEE, Cocke Co., June (A); Greene
Co., May (N); Hawkins Co., July (N).
Epeorus species
Physiographic provinces: CP, P, BR, VR.
Additional records: ALABAMA, Calhoun Co., July (N). GEORGIA, Burke Co., June (N); Carroll
Co., March (N); Cherokee Co., June (N); Dawson Co., June (N,A); Fannin Co., May (N)3;
Habersham Co., April (A)'; Lumpkin Co., August (N), April (N); Pickens Co., June (N);
Rabun Co., May (N)3, April (N); August (N), White Co., June, July (N)3. NORTH CAROLINA,
Avery Co., February, April (N); Buncombe Co., May (N); Clay Co., April (N); Macon Co.,
July, August (N); McDowell Co., May, August (N); Swain Co., June, July, December (N);
Transylvania Co., March, May, July (N); Watauga Co., February (N); Yancey Co., May,
June, July (N). SOUTH CAROLINA, Aiken Co., May (A); Oconee Co., July (A), August (N).
TENNESSEE, Blount Co., April, June (N); Greene Co., May (N)', August (N); Sevier Co.,
April (A)', April-August, December (N).
Epeorus subpallidus (Traver)
Physiographic provinces: BR.
Previously published state records: N. C., Tray., 1937, H, 1941
Additional records: NORTH CAROLINA, Macon Co., August (A). TENNESSEE,
Sevier Co., June (A).


GENUS Heptagenia WALSH

Heptagenia aphrodite McDunnough
Physiographic provinces: P, BR, VR, AP, ILP.
Previously published state records: Ga., NTH, 1935; N.C., NTH, 1935, Tray., 1937, H,
1941.
Additional records: ALABAMA, Bibb Co., June (N); Coosa Co., June (N); Lauderdale Co.,
June (A); Walker Co., June (A). GEORGIA, Catoosa, June (A)', July (A)3; Floyd Co.,
June (A)3; Fulton Co., July (A)'; White Co., July (A)3, June, July (A); Whitfield Co.,
June, July, August, September (A)3. NORTH CAROLINA, Macon Co. September (A).
TENNESSEE, Cheatham Co., July (A); Davidson Co., June (A); Dickson Co., July (N,A);
Lincoln Co., July (N); Rutherford Co., July (N); Sevier, October (N).
Heptagenia cruentata Walsh
Physiographic provinces: CP.
Previously published state records: None.
Additional records: TENNESSEE, Shelby Co., June (A).
Heptagenia dolosa Traver
Physiographic provinces: CP, P, BR, VR.
Previously published state records: Ga., NTH, 1935.
Additional records: GEORGIA, Bartow Co., August (A)'; Cherokee Co., June (N,A), August


Vol. 22, No. 1







BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


(A); Cobb Co., July (A); Dougherty Co., June (A)'; Lumpkin Co., July (A), August
(N); Muscogee Co., June, July (A); Pickens Co., June (N); Stephens Co., July (A)'.
NORTH CAROLINA, Forsyth Co., June (N); Randolf Co., July (N). SOUTH CAROLINA,
Greenville Co., June (A)'.
Heptagenia flavescens (Walsh)
Physiographic provinces: CP, P, VR, AP
Previously published state records: Fla., Ber., 1953 and 1958, Sch, 1967; Ga., NTH, 1935,
Ber., 1958.
Additional records: ALABAMA, Dallas Co., April (A), Dekalb Co., July (N). FLORIDA, Es-
cambia Co., March, November (N); Walton Co., June (A). GEORGIA, Decatur Co.,
June (A)3; Floyd Co.,June (A); Fulton Co. June (A)3; Richmond Co., April (A). LOUISIANA,
East Baton Rouge Co., May (A); Natchitoches Co., May (A), April (A). SOUTH CAROLINA,
Orangeburg Co., April (A).
Heptagenia hebe McDunnough
Physiographic provinces: BR, VR, AP, ILP
Previously published state records: Tenn., WB, 1949.
Additional records: ALABAMA, Blount Co., June (N); Cullman Co., June (N); Marion Co.,
June (N). GEORGIA, Catoosa Co., June (A)'3 Rabun Co., June (A)3. NORTH CAROLINA,
Avery Co., August (A); Jackson Co., August (A); Macon Co., September (A), August (A);
Watauga Co., July (N). TENNESSEE, Blount Co., June (A); Cumberland Co., July (N);
Grainger Co., July (N,A); Greene Co., May (A); Hamilton Co., July (A); Lincoln Co., July
(A); Sevier Co., August, July (A).
Heptagenia julia Traver
Physiographic provinces: P, BR.
Previously published state records: N. C., NTH, 1935, Tray. 1937, H, 1941; S. C. NTH, 1935.
Additional records: GEORGIA, Towns Co., May (A)3, NORTH CAROLINA, Jackson Co., August
(A), August (N). SOUTH CAROLINA, Greenville Co., June (A)*.
Heptagenia juno McDunnough
Physiographic provinces: P, BR, VR, AP, ILP.
Previously published state records: N. C., Tray., 1937, H, 1941.
Additional records: ALABAMA, Dekalb Co., July (N). GEORGIA, Lumpkin Co., June (A)3.
NORTH CAROLINA, Davidson Co., July (N); Jackson Co., June (A), Person Co., July (N).
TENNESSEE, Hawkins Co., July (N); Robertson Co., June (N); Sevier Co., October (N),
June (A).
Heptagenia maculipennis Walsh
Physiographic provinces: CP, P, VR
Previously published state records: Tenn., NTH, 1935
Additional records: ALABAMA, Bibb Co., May (N); Dallas Co., June (A); Perry Co., June (A).
MIssIssIPPI, Lamar Co., August (A); Lawrence Co., August (A), Lowndes Co., June (A);
Marion Co., May (A); Pike Co., June (A). SOUTH CAROLINA, Chesterfield Co., July (A).
Heptagenia marginalis Banks
Physiographic provinces: CP, P, BR, VR, AP
Previously published state records: N. C., NTH, 1935, Trav., 1937, H, 1941
Additional records: ALABAMA, Dekalb Co., July (N). GEORGIA, Bartow Co., June (A)'. Chero-
kee Co., June (A); Dougherty Co., September (A); Gilmer Co., July (A)3; Habersham Co.,
June (A)3; Upson Co., June (A)3. NORTH CAROLINA, Davidson Co., July (N), Macon Co.,
June (A), August (A); Swain Co., August (A). SOUTH CAROLINA, Edgefield Co., July (N);
Greenville Co., August (N); Oconee Co., August (N). TENNESSEE, Fentress Co., June (A).
Heptagenia minerva McDunnough
Physiographic provinces: BR.
Previously published state records: Ala., Trav., 1937.
Additional records: TENNESSEE, Sevier Co., July (A).


1977







BULLETIN FLORIDA STATE MUSEUM


Heptagenia persimplex McDunnough
Physiographic provinces: CP, P.
Previously published state records: None
Additional records: ALABAMA, Macon Co., June (A). GEORGIA, Cherokee Co., June (A).
TENNESSEE, Shelby Co., June (A).
Heptagenia pulla (Clemens)
Physiographic provinces: BR, AP.
Previously published state records: None
Additional records: ALABAMA, Cullman Co., June (N,A). NORTH CAROLINA, Swain Co., August
(A); Watauga Co., August (A).
Heptagenia species
Physiographic provinces: CP, P, BR, VR, AP, ILP.
Additional records: ALABAMA, Randolph Co., June (N). FLORIDA, Escambia Co., October
(N), March (N). GEORGIA, Burke Co., November, December (N), June (N); Chattooga Co.,
June (A)3, April (N); Cherokee Co., June (N,A); Cobb Co., August (N), November (N);
Dawson Co., June (N); Fulton Co., April (N); Lumpkin Co., July (A)3, April (N); Rabun
Co., April (N); Talbot Co., July (A)3; White Co., August (A), July (A). MISSISSIPPI, Forrest
Co., August (A), Lamar Co., May (N); Marion Co., August (N). NORTH CAROLINA, Avery
Co., February (N); Clay Co., April (N); Jackson Co., August (N,A); Macon Co., June, July
(A), August (N); McDowell Co., August (N); Orange Co., May (N); Swain Co., May, June
(N). SOUTH CAROLINA, Aiken Co., June (N); Fairfield Co., April (N); Greenville Co., July
(A)4. TENNESSEE, Blount Co., May (N); Giles Co., July (N); Greene Co., August (N); Hamil-
ton Co., May (N); Robertson Co., July (N); Sevier Co., May, June (N); Sumner Co., June
(N), Unicoi Co., September (A).
Heptagenia spinosa Traver
Physiographic provinces: BR
Previously published state records: N. C., NTH, 1935, Tray. 1937.
Additional records: NORTH CAROLINA, Swain Co., August (A).
Heptagenia thetis Traver
Physiographic provinces: P, BR, VR.
Previously published state records: Ga., Tray., 1937; N. C., NTH, 1935, Tray., 1937, H, 1941.
Additional records: GEORGIA, Cowetta Co., June (A)3; Lumpkin Co., June (A)3; Whitfield
Co., May (A)3. NORTH CAROLINA, Swain Co., July (N). SOUTH CAROLINA, Oconee Co.,
August (N). TENNESSEE, Greene Co., August (N).
Heptagenia townesi Traver
Physiographic provinces: BR.
Previously published state records: S. C., NTH, 1935.
Additional records: GEORGIA, Rabun Co., August (A). NORTH CAROLINA, Jackson Co., August
(A), Macon Co., July (A). SOUTH CAROLINA, Greenville Co., June (A)'. TENNESSEE, Sevier
Co., July (A).

GENUS Rhithrogena EATON
Rhithrogena amica Traver
Physiographic provinces: BR.
Previously published state records: N. C., Tray., 1937.
Additional records: GEORGIA, Towns Co., June (A)3.
Rhithrogena exiles Traver
Physiographic provinces: BR.
Previously published state records: N. C., NTH, 1935.
Additional records: GEORGIA, Rabun Co., June (A)3. NORTH CAROLINA, Macon Co., August
(N); Swain Co., May (N)', August (N). TENNESSEE, Blount Co., June (N).


Vol. 22, No. 1








BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


Rhithrogena fasciata Traver
Physiographic provinces: BR.
Previously published state records: N. C., Trav., 1933 and 1937, H, 1941.
Additional records: GEORGIA, Rabun Co., June (A)3; White Co., July, August (A).
Rhithrogena fuscifrons Traver
Physiographic provinces: BR, VR.
Previously published state records: N. C., NTH, 1935, H, 1941.
Additional records: GEORGIA, Fannin Co., July (A)3; NORTH CAROLINA, Jackson Co., June (A).
Rhithrogena pellucida Daggy
Physiographic provinces: CP.
Previously published state records: None
Additional records: ALABAMA, Perry Co., June (A).
Rhithrogena rubicunda Traver
Physiographic provinces: BR
Previously published state records: N. C., Tray., 1937.
Additional records: NORTH CAROLINA, Jackson Co., August (A).
Rhithrogena species
Physiographic provinces: CP, BR, VR, AP, ILP.
Additional records: ALABAMA, Marion Co., June (N). GEORGIA, Lumpkin Co., April (N);
Rabun Co., August (N); Whitfield Co., May (N)3. MississiPpi, Lamar Co., May (N). NORTH
CAROLINA, Jackson Co., August (N); Macon Co., August (N); Swain Co., June (N); Transyl-
vania Co., August (N), Yancey Co., July (N). TENNESSEE, Campbell Co., June (N)'; Dick-
son Co., June (N); Humphreys Co., July (N); Sevier Co., June (N)'; Wayne Co., July (N).

Rhithrogena uhari Traver
Physiographic provinces: CP, P, BR.
Previously published state records: N. C., NTH, 1935.
Additional records: ALABAMA, Perry Co., May (N). GEORGIA, Rabun Co., May (A)3. NORTH
CAROLINA, Transylvania Co., August (N). SOUTH CAROLINA, Edgefield Co., April (N).


GENUS Stenacron JENSEN
Stenacron canadense (Walker)
Physiographic provinces: BR
Previously published state records: None
Additional records: GEORGIA, Rabun Co., May (A)3.
Stenacron carolina (Banks)
Physiographic provinces: BR
Previously published state records: N. C., Banks, 1914, NTH, 1935, Tray., 1937, H, 1941,
L, 1974 a, b; S. C., NTH, 1935 L, 1974b; Tenn.; McD., 1925, NTH, 1935, 1974b.
Additional records: GEORGIA, Rabun Co., May (A)3; White Co., June (A)'.
Stenacron floridense (Lewis)
Physiographic provinces: CP
Previously published state records: Fla., L, 1974a and 1974b.
Additional records: None.
Stenacron gildersleevei (Traver)
Physiographic provinces: BR
Previously published state records: Tenn., L, 1974b
Additional records: None.
Stenacron heterotarsale (McDunnough)
Physiographic provinces: BR







BULLETIN FLORIDA STATE MUSEUM


Previously published state records: N. C., NTH, 1935; Tenn., WB, 1949.
Additional records: ALABAMA, Conecuh Co., June (A); GEORGIA, Cherokee Co., June (A);
Fannin Co., May (A)3; Peach Co., April (A); Rabun Co., August (A). NORTH CAROLINA,
Macon Co., August (A). SOUTH CAROLINA, Oconee Co., August (A). TENNESSEE, Greene
Co., April (N), May (A), September (A); Lincoln Co., July (A); Sevier Co., August (A).
Stenacron interpunctatum (Say)
Physiographic provinces: CP, P, BR, VR, AP, ILP.
Previously published state records: Ala., Trav., 1937; Fla., Ber., 1950 (as Stenonema proxi-
mum), 1958, PJ, 1973, Sch, 1967, L, 1974b; Ga., NTH, 1935, Ber., 1958; N. C., L, 1974b; Tenn.,
NTH, 1935.
Additional records: ALABAMA, Baldwin Co., April, June, December (N); Bibb Co., May (N),
June (A); Calhoun Co., July (N,A); Cherokee Co., July (N,A); Clay Co., June (N,A); Col-
bert Co., June (A); Dallas Co., June (N); Dekalb Co., April, May, July (N); Elmore Co.,
June (N); Escambia Co., June (N,A); Lauderdale Co., June (A), July (N); Mobile Co., June
(N,A); St. Clair Co., June (N,A); Tallapoosa Co., June (A); Tuscaloosa Co., April (N), May
(N); Walker Co., June (N,A); Winston Co., August (N,A). FLORIDA, Alachua Co., October
(A); Baker Co., February (N); Columbia Co., November (A); Gadsden Co., June (A);
Hamilton Co., July (A); Hardee Co., June (N); Hernando Co., February (A); Highlands
Co., January (A); Jackson Co., April (A); Levy Co., November (N); Santa Rosa Co., August
(N); Walton Co., April (N). GEORGIA, Baker Co., May (N,A), November (N); Bartow Co.,
August (A)'; Burke Co., September (N), December (N); Catoosa Co., June (A)'; Cherokee
Co., August (A); Cobb Co., August (N); Dawson Co., June (A); Decatur Co., June (A);
August (A)'; Early Co., October (N); Fannin Co., June (A)3; Floyd Co., June (A), June and
August (A)3, July (A)'; Jackson Co., July (N); Lowndes Co., February (A); Lumpkin Co.,
July (A), August (N); Monroe Co., July (N); Muscogee Co., June (A); Schley Co., April
(A)'; Screven Co., September (N); Thomas Co., July (A)'; Toombs Co., August (N); Whit-
field Co., June and July (A)3, September (A)'. LOUISIANA, Tangipahoa Par., June (N).
MISSISSIPPI, Benton Co., June (N); Clark Co., August (N); Franklin Co., August (N);
Harrison Co., August (N); Holmes Co., July (N); Itawamba Co., July (N,A); Jasper Co.,
April (A); Lafayette Co., July (N,A); Lamar Co., May (N), August (A); Lauderdale Co.,
July (N,A); Lawrence Co., August (N,A); Leflore Co., June (A); Lowndes Co., June (A);
Marion Co., August (N); Marshall Co., July (N); Monroe Co., July (A); Perry Co., August
(N); Pike Co., August (N); Quitman Co., July (N); Sunflower Co., July (N,A). NORTH
CAROLINA, Avery Co., February (N); Davidson Co., July (N,A); Jackson Co., August (A);
Macon Co., June-July (A); Surry Co., February (N); Union Co., July (A). SOUTH CAROLINA,
Aiken Co., May (A), July (N); Newberry Co., April, August (N); Oconee Co., August (A);
Orangeburg Co., April (A); TENNESSEE, Cheatham Co., July (A); Davidson Co., June,
July (A); Dekalb Co., June, July (N,A); Greene Co., August (N); Hamilton Co., December
(N); Hawkins Co., August (N); Hickman Co., June (A); Humphreys Co., June (A); Obion
Co., June (A); Polk Co., August (N,A); Sevier Co., June, July, August (A); Union Co.,
August (N); Wayne Co., July, August (N,A); Williamson Co., May (N).
Stenacron pallidum (Traver)
Physiographic provinces: P, BR
Previously published state records: N. C., Trav., 1933, 1937, L, 1974b.
Additional records: None.


GENUS Stenonema TRAVER

Stenonema alabamae Traver
Physiographic provinces: AP, ILP.
Previously published state records: Ala., Trav., 1937.
Additional records: ALABAMA, Lauderdale Co., June (A); Tuscaloosa Co., June (A).
Stenonema annexum Traver
Physiographic provinces: CP, P, BR, VR, ILP.


Vol. 22, No. 1







BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


Previously published state records: Ga., L, 1974b; N. C., NTH, 1935, L, 1974b; S. C., NTH,
1935; Tenn., WB, 1949.
Additional records: ALABAMA, Lauderdale, June (A). GEORGIA, Calhoun Co., April (A)';
Cherokee Co., August (A); Fulton Co., September (A)'; Houston Co., March (A)'; Jackson
Co., July (N,A); Lumpkin Co., August (A); Walton Co., June (A)'; Wiles Co., July (N,A).
NORTH CAROLINA, Davidson Co., July (N,A); Jackson Co., June (N); Lincoln Co., July
(A); Randolf Co., July (A); Surry Co., June (A); Union Co., July (A). SOUTH CAROLINA,
Edgefield Co., July (N,A); Greenville Co., July (A)'; Lexington Co., April (A). TENNESSEE,
Greene Co., May (A); Hickman Co., May (A); Sevier Co., July (A); Union Co., August
(A).
Stenonema bipunctatum (McDunnough)
Physiographic provinces: CP, BR, VR, AP, ILP.
Previously published state records: None.
Additional records: ALABAMA, Bibb Co., June (A); Cullman Co., June (A); Dallas Co., June
(A); Perry Co., June (A); Walker Co., June (A). GEORGIA, Floyd Co., June (A); Muscogee
Co., June (A). MISSIssIPPI, Lamar Co., August; Lawrence Co., August (A); Lowndes Co.,
June (A); Pike Co., August (A). NORTH CAROLINA, Macon Co., July (A). TENNESSEE, Chea-
tham Co., July (A); Lincoln Co., July (A).
Stenonema carlsoni Lewis
Physiographic provinces: P, BR
Previously published state records: Ga., L, 1974 a, b; S. C., L, 1974 a, b.
Additional records: None.
Stenonema exiguum Traver
Physiographic provinces: CP, P, BR, VR, AP, ILP.
Previously published state records: Ala., Trav., 1937, Ber., 1958, L, 1974b; Fla., Ber., 1950,
1958, PJ, 1973, Sch, 1967; Ga., NTH, 1935, Ber., 1958, L, 1974b; N. C., NTH, 1935; Wy.,
1950, L, 1974b.
Additional records: ALABAMA, Baldwin Co., April (N); Barbour Co., June (N); Bibb Co., May
(N), June (N,A); Blount Co., June (A); Cherokee Co., July (N,A); Cullman Co., June (A);
Dallas Co., June (A); Escambia Co., April (N), June (A); Houston Co., February, June
(N); Lauderdale Co., June (A); Macon Co., June (N,A); Mobile Co., June (N,A); Perry
Co., May (N), June (A); Talladega Co., June (A); Tuscaloosa Co., May (A); Washington Co.,
May (A). FLORIDA, Alachua Co., October (A); Clay Co., January (N); Columbia Co., No-
vember (A); DeSoto Co., February-April, November (N); Escambia Co., March (N),
August (A), October (N,A); Gadsden Co., April (N,A), June (A); Hernando Co., February
(A); Jackson Co., April (A); Marion Co., July (A); Santa Rosa Co., June (A); Walton Co.,
April (A). GEORGIA, Baker Co., October (N,A); Cherokee Co., June (N,A), July (A), August
(N,A); Decatur Co., March (A); June (N,A); Dougherty Co., September (A)'; Echols Co.,
February (N); Floyd Co., June (A), July (A)', August (A); Fulton Co., June (A)3; Johnson
Co., August (N); Lowndes Co., March (N,A); Muscogee Co., June (A), July (A); Richmond
Co., August (A); Thomas Co., June, July (A)'; Troup Co., July (N). LOUISIANA, Natchi-
toches Par., April (A), May (A); Tangipahoa Par., August (N); Washington Par., June (A).
MISSISSIPPI, Amite Co., June (N); Clark Co., August (N); Copiah Co., August (N); Franklin
Co., August (N); Green Co., May (N); Harrison Co., August (N); Itawamba Co., July (N,A);
Lamar Co., May (N); Lauderdale Co., June, July (A); Lawrence Co., August (N); Leflore
Co., June (A); Lowndes Co., June (A); Marion Co., May (A), August (N); Monroe Co.,
July (N); Perry Co., August (N); Pike Co., August (N,A); Sunflower Co., July (A). NORTH
CAROLINA, Surry Co., June (A). SOUTH CAROLINA, Aiken Co., May-July (A); Kershaw Co.,
July (N); Newberry Co., August (N,A). TENNESSEE, Hamilton Co., August (A); Lincoln Co.,
July (A); Sevier Co., August (A).

Stenonema femoratum (Say)
Physiographic provinces: P, BR, VR, AP, ILP.
Previously published state records: Ga., NTH, 1935; S. C., NTH, 1935.
Additional records: ALABAMA, Blount Co., June (N); Cullman Co., (N,A); Walker Co., June







BULLETIN FLORIDA STATE MUSEUM


(A). GEORGIA, Floyd Co., March (A); Whitfield Co., May, July (A)3. SOUTH CAROLINA,
Greenville Co., July (A)'; Lexington Co., May (A)4; Oconee Co., May (A),. TENNESSEE,
Davidson Co., May (N), July (A); October (N).
Stenonema integrum (McDunnough)
Physiographic provinces: CP, P, BR, VR, ILP.
Previously published state records: Fla., L, 1974b; Ga., NTH, 1935; N. C., NTH, 1935 (as
S. bellum), L, 1974b; S. C., L, 1974b.
Additional records: ALABAMA, Bibb Co., June (A); Dallas Co., June (A); Elmore Co., June
(N,A). GEORGIA, Catoosa Co., July, August (A)3; Cherokee Co., June (A); Dawson Co.,
June (A); Floyd Co., June, August (A)'; Lowndes Co., February (A); Monroe Co., July
(A); Muscogee Co., July (A); Richmond Co., June (A); Stephens Co., July (A)'; Union Co.,
July (A)3; Walton Co., August (A)'; White Co., August (A), September (A)'; Whitfield
Co., June-August (A)3. LOUISIANA, East Baton Rouge Par., June (A), September (A);
Natchitoches Par., (A). MISSIsSIPPI, Lauderdale Co., June (A); Leflore Co., June (A);
Lowndes Co., June (A); Marion Co., May (A). NORTH CAROLINA, Caldwell Co., July (N,A);
Davidson Co., July (A); Forsyth Co., June (A); Lincoln Co., July (A); Person Co., July
(A); Rockingham Co., July (A); Transylvania Co., August (N,A). SOUTH CAROLINA, Aiken
Co., May (A); Anderson Co., July (N,A); Greenville Co., June (A),, August (N,A), Septem-
ber (A)'; Newberry Co., August (N,A); Oconee Co., July (A); August (A). TENNESSEE,
Humphreys Co., June (A); Polk Co., August (A); Sevier Co., August (A); Shelby Co., June
(A); Williamson Co., July (A).

Stenonema ithaca (Clemens and Leonard)
Physiographic provinces: BR, VR, AP, ILP
Previously published state records: Ga., NTH, 1935, N. C., NTH, 1935, Tray., 1937, H, 1941;
S. C., NTH, 1935; Tenn., WB, 1949.
Additional records: ALABAMA, Lauderdale Co., July (N); GEORGIA, Fannin Co., June (A)3;
Lumpkin Co., May (A)'; Towns Co., June (A)'; Union Co., June (A)3. NORTH CAROLINA,
Clay Co., June (A); Haywood Co., June (A); Jackson Co., June, August (A); Macon Co.,
June, July, August (A); Swain Co., June (A); Watauga Co., June (N); August (N), August
(A); Yancey Co., July (A). SOUTH CAROLINA, Greenville Co., April, May, September (A)'.
TENNESSEE, Fentress Co., May (A); Greene Co., May (A), July (A), August (N); Hawkins
Co., August (N); Lincoln Co., July (N); Montgomery Co., July (N); Moore Co., July (N);
Polk Co., August (A); Robertson Co., July (N); Sevier Co., July-August (A); Unicoi Co.,
August (N,A); Wayne Co., July (N).
Stenonema metriotes Burks
Physiographic provinces: CP, P, BR
Previously published state records: None
Additional records: LOUISIANA, East Baton Rouge Par., June (A). MISSISSIPPI, Adams Co.,
June (A). TENNESSEE, Humphreys Co., June (A); Shelby Co., June (A).
Stenonema nepotellum McDunnough
Physiographic provinces: CP, ILP.
Previously published state records: Tenn., L, 1974b.
Additional records: NORTH CAROLINA, Cumberland Co., July (A); Macon Co., June (A);
Rockingham Co., July (A); Wilkes Co., June (A). SOUTH CAROLINA, Lexington Co., April
(A).TENNESSEE, Sevier Co., October (N).
Stenonema pudicum (Hagen)
Physiographic provinces: P, BR, VR, AP.
Previously published state records: N. C., NTH, 1935, Tray. 1937, H, 1941, L, 1974b; S. C.,
NTH, 1935; Tenn., WB, 1949.
Additional records: GEORGIA, Clarke Co., April (A)'; Habersham Co., April (A)'. NORTH
CAROLINA, Clay Co., April (N); Jackson Co., June (A), August (N,A); McDowell Co., April
(N,A); Macon Co., June (N,A), July (A), August (N,A); Polk Co., July (N,A); Swain Co.,
April (N), June (N,A), August (A); Transylvania Co., August (A). SOUTH CAROLINA, Green-


Vol. 22, No. 1







BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


ville Co., February (N)', May-July, September (A)'; Oconee Co., August (N); Pickens Co.,
May (A)'. TENNESSEE, Blount Co., June (A); Cocke Co., April, June (A); Cumberland Co.,
July (N); Greene Co., March (N), May (A); Marion Co., April (N); Morgan Co., July (N);
Rhea Co., April (N); Sevier Co., April, May, July, August (A); Unicoi Co., May, June,
August (A).
Stenonema quinquespinum Lewis
Physiographic provinces: VR
Previously published state records: Ga., L, 1974a, b.
Additional records: None.
Stenonema rubrum (McDunnough)
Physiographic provinces: P, BR, VR, ILP.
Previously published state records: Ga., NTH, 1935; N. C., L, 1974b; S. C., 1935, NTH.
Additional records: GEORGIA, Dawson Co., June (A); Lumpkin Co., July (A). NORTH CARO-
LINA, Swain Co., August (N). SOUTH CAROLINA, Greenville Co., April (A)*; Laurens Co.,
August (A). TENNESSEE, Houston Co., June (A); Sevier Co., August (A).

Stenonema scitulum Traver
Physiographic provinces: CP, VR, ILP
Previously published state records: None
Additional records: TENNESSEE, Benton Co., June (N,A); Granger Co., July (N); Hawkins Co.,
August (A); Hickman Co., June (A); Humphreys Co., June (N,A); Lincoln Co., July (N,A);
Montgomery Co., July (N,A); Robertson Co., June (N).

Stenonema smithae Traver
Physiographic provinces: CP, P, VR, AP
Previously published state records: Ala., Trav., 1937, Ber., 1958, L, 1974b; Fla., Ber., 1950,
1958, PJ, 1973, Sch., 1967, L, 1974b; Ga., Ber., 1958.
Additional records: ALABAMA, Baldwin Co., April, June (N,A); Bibb Co., June (A); Conecuh
Co., June (A); Dale Co., June (N); Dallas Co., June (A); Escambia Co., June (N,A); Houston
Co., February (N); Macon Co., June (A); Marion Co., July (N); Mobile Co., June (N,A);
St. Clair Co., June (A); Tuscaloosa Co., May (A); Walker Co., June (A). FLORIDA, Alachua
Co., January (N,A), April (A), November (A); Baker Co., February (N); Clay Co., January
(A); Columbia Co., November (A); Escambia Co., October (A); Gadsden Co., April (N,A),
June (A); Hamilton Co., February, November (N); Levy Co., November (N); Liberty Co.,
March (N,A), June (N); Madison Co., January (N); Nassau Co., February (A); Okaloosa
Co., April, May (A). GEORGIA, Baker Co., May (N), November (N,A); Charlton Co., Febru-
ary (N); Cherokee Co., June and August (A); Clinch Co., November (A); Early Co., Febru-
ary (N): Echols Co., February (A), November (N); Lowndes Co., March (N,A); Monroe Co.,
July (N,A); Muscogee Co., June (A). LOUISIANA, Tangipahoa Par., June (N). MIssIssIPPI,
Jasper Co., April (A); Lafayette Co., July (N,A). SOUTH CAROLINA, Aiken Co., May (A).

Stenonema species
Physiographic provinces: CP, P, BR, VR, AP, ILP.
Additional records: ALABAMA, Baldwin Co., April, June (N,A); Bibb Co., June (A); Conecuh
Co., June (A); Dale Co., June (N); Dallas Co. June (A); Dekalb Co., July (N); Elmore Co.,
April (N); Escambia Co., April (N); Henry Co., June (N); Randolph Co., June (N); St. Clair
Co., July (N); Tuscaloosa Co., April, May, and June (N), May (A). FLORIDA, Escambia
Co., March (N); Gadsden Co., April (A); Liberty Co., March (N); Okaloosa Co., May (A);
Santa Rosa Co., August (N); Walton Co., April (A). GEORGIA, Cherokee Co., June (A),
August (N); Cobb Co., August (N); Decatur Co., August (N); Forsyth Co., June (N); Macon
Co., April (N); Peach Co., April (N); Rabun Co., July (N), August (A); Sumter Co., August
(N); Toombs Co., August (N); Walker Co., April and July (N); Ware Co., April (N). MissIs-
SIPPI, Lamar Co., May and August (N); Marion Co., August (N); Stone Co., August (N).
NORTH CAROLINA, Buncombe Co., May (N); Clay Co., April (N); Granville Co., December
(N); Jackson Co., August (N); Macon Co., June (A), May-August (N); McDowell Co., May
and August (N); Moore Co., December (N); Orange Co., February and May (N); Surry


1977







BULLETIN FLORIDA STATE MUSEUM


Co., February (N); Swain Co., April-June (N)'; Transylvania Co., March-July (N); Wake
Co., February, May, December (N); Watauga Co., February (N). SOUTH CAROLINA,
Chesterfield Co., December (N); Edgefield Co., April and July (N); Lexington Co., April
(N); Oconee Co., August (A). TENNESSEE, Blount Co., June (N); Davidson Co., July (A),
October (N); Giles Co., July (N); Greene Co., May (A), August (N); Hamilton Co., May
(N); Henderson Co., June (N); Meigs Co., April (N); Morgan Co., July (N); Robertson
Co., June (N); Rutherford Co. July (N); Sevier Co., April, June-August, December (N);
Sumner Co., June (N); Van Buren Co., July (N).
Stenonema terminatum (Walsh)
Physiographic provinces: BR
Previously published state records: None
Additional records: NORTH CAROLINA, Macon Co., July (A)
Stenonema tripunctatum (Banks)
Physiographic provinces: CP, P, BR, VR, AP, ILP.
Previously published state records: Ala., Spieth, 1947; N. C., Spieth, 1947.
Additional records: ALABAMA, Calhoun Co., July (N); Marion Co., June (N). GEORGIA, Rabun
Co., July (A)3. LOUISIANA, Natchitoches Par., March (N), May (A). NORTH CAROLINA,
Cabarrus Co., July (N); Davidson Co., July (N,A). SOUTH CAROLINA, Kershaw Co., April
(N). TENNESSEE, Fentress Co., May (A); Hamilton Co., December (N); Lincoln Co., April
(N); Moore Co., April (N); Union Co., May (N).
Stenonema varium Traver
Physiographic provinces: CP, BR, ILP
Previously published state records: Ga., Ber., 1958; N. C., NTH, 1935, H, 1941; S. C., NTH,
1935
Additional records: GEORGIA, Baker Co., March (A); Crawford Co., March (A)'; Dougherty
Co., November (N,A); Lumpkin Co., May (A)'; Worth Co., November (N). NORTH CARO-
LINA, Macon Co., June (A), August (A). SOUTH CAROLINA, Greenville, June (A)'. TEN-
NESSEE, Lincoln Co., April (A).

Stenonema vicarium (Walker)
Physiographic provinces: Br
Previously published state records: N. C., NTH, 1935, H, 1941.
Additional records: NORTH CAROLINA, Macon Co., July (A); Swain Co., June (A).



SUBFAMILY SPINADINAE
GENUS Spinadis EDMUNDS AND JENSEN

Spinadis wallacei Edmunds and Jensen
Physiographic provinces: CP
Previously published state records: Ga., EJ, 1974.
Additional records: None.



SUBFAMILY ANEPEORINAE
GENUS Anepeorus McDUNNOUGH

Anepeorus simplex (Walsh)
Physiographic provinces: P
Previously published state records: Ga., NTH, 1935.
Additional records: None


Vol. 22, No. 1







BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


SUPERFAMILY CAENOIDEA
FAMILY NEOEPHEMERIDAE
SUBFAMILY NEOEPHEMERINAE
GENUS Neoephemera McDUNNOUGH
Neoephemera compressa Berner
Physiographic provinces: CP
Previously published state records: Fla., Ber., 1956, Sch, 1967; Ga., Ber., 1956 and 1958.
Additional records: None.
Neoephemera purpurea (Traver)
Physiographic provinces: BR, AP
Previously published state records: N. C., NTH, 1935, Trav., 1937, H, 1941.
Additional records: GEORGIA, Rabun Co., July (N). NORTH CAROLINA, Jackson Co., August
(N); Macon Co., August (N); Polk Co., July (N); Swain Co., August (N); Transylvania Co.,
June and August (N). SOUTH CAROLINA, Greenville Co., June (A)'. TENNESSEE, Morgan
Co., July (N).
Neoephemera young Berner
Physiographic provinces: CP, P.
Previously published state records: Fla., Ber., 1950 (as Orieanthus sp. no. 1 Tray.), 1953, 1956,
and 1958, PJ, 1973, Sch, 1967; Ga., Ber., 1953, 1956, and 1958; S. C., Ber., 1956.
Additional records: ALABAMA, Elmore Co., April (N); Escambia Co., April; Macon Co., April
(N). FLORIDA, Liberty Co., March (N); Okaloosa Co., May (N); Santa Rosa Co., August
(N).

FAMILY CAENIDAE
SUBFAMILY CAENINAE
GENUS Brachycerus CURTIS
Brachycercus flavus Traver
Physiographic provinces: CP
Previously published state records: La., NTH, 1935.
Additional records: None
Brachycercus maculatus Berner
Physiographic provinces: CP
Previously published state records: Fla., Ber.; 1950.
Additional records: FLORIDA, DeSoto Co., August (N); Hamilton Co., July (A).
Brachycercus nitidus (Traver)
Physiographic provinces: P, BR
Previously published state records: Fla., WR, 1955; N. C., Trav., 1932.
Additional records: GEORGIA, Cherokee Co., June (N,A); Rabun Co., July (N). NORTH CARO-
LINA, Jackson Co., July (N); Transylvania Co., June (N).
Brachycercus prudens (McDunnough)
Physiographic provinces: CP
Previously published state records: None
Additional records: ALABAMA, Perry Co., June (A)
Brachycercus species
Physiographic provinces: CP, P
Additional records: ALABAMA, Dallas Co., June (A). FLORIDA, Escambia Co., August (N,A),
August (A), October (N); Okaloosa Co., August (N). GEORGIA, Baker Co., May (N); Chero-
kee Co., June (N); Monroe Co., July (A). MIssIssIPPI, Harrison Co., August (N); Lawrence
Co., August (A); Lowndes Co., June (A).


1977







BULLETIN FLORIDA STATE MUSEUM


Brachycercus species A
Physiographic provinces: CP, P, VR
Previously published state records: Fla., Ber., 1950, 1958, Sch, 1967.
Additional records: ALABAMA, Macon Co., April (A). FLORIDA, Liberty Co., March (N), April
(A), June (N,A); Santa Rosa Co., August (N). GEORGIA, Dawson Co., June (N); Decatur
Co., June. MISSISSIPPI, Covington Co., August (N). SOUTH CAROLINA, Aiken Co., May (N)2.


GENUS Caenis STEPHENS
Caenis amica Hagen
Physiographic provinces: CP, P, BR, AP, ILP
Previously published state records: N. C., NTH, 1935.
Additional records: GEORGIA, Rabun Co., April (A)3. SOUTH CAROLINA, Beaufort Co., August
(A)'; Spartanburg Co., June (A). TENNESSEE, Dekalb Co., July (A); Sevier Co., July (A).
Caenis anceps Traver
Physiographic provinces: AP, VR
Previously published state records: None
Additional records: ALABAMA, Cherokee Co., July (A). TENNESSEE, Campbell Co., August
(A); Hamilton Co., August (A).
Caenis diminuta Walker
Physiographic provinces: CP, P, VR
Previously published state records: Fla., Walker, 1853, NTH, 1935, Ber., 1950 and 1958,
PJ, 1973, Sch, 1967; Ga., NTH, 1935, Ber., 1958.
Additional records: ALABAMA, Bibb Co., June (A); Cleburne Co., July (A); Macon Co., April
June (A); Mobile Co., June (N); Perry Co., June (A). FLORIDA, Alachua Co., July (A);
Broward Co., November (A); Columbia Co., October (A), November (N); DeSoto Co.,
August (N); Escambia Co., August (A); Gadsden Co., March (A), April (A), June (A); Gulf
Co., April (A); Hamilton Co., March (A); Hernando Co., February (A); Highlands Co.,
April (A); Jackson Co., April (A); Lee Co., September (A); Levy Co., September (A);
Monroe Co.,January (A), February (A); Nassau Co., February (A) Okaloosa Co., May (A);
Santa Rosa Co., June, August (A). GEORGIA, Charlton Co., February (N); Clinch Co.,
November (A); Decatur Co., June (A), July (A)3; Echols Co., February (N); Lowndes Co.,
March (A); Thomas Co., May (N). LOUISIANA, Orleans Co., June (A). MISSISSIPPI, Forrest
Co., August (N); Itawamba Co., July (A); Lafayette Co., July (A); Lawrence Co., August
(A); Lauderdale Co., April, June, July (A); Monroe Co., July (A); Pike Co., June, August
(A); Sunflower Co., July (A); Tallahatchie Co., July (A). NORTH CAROLINA, Wayne Co.,
June (A), September (A). SOUTH CAROLINA, Aiken Co., March2, April2, May2, June2, July2,
August2, October2 (A); Chesterfield Co., July (A); Darlington Co., July; Greenwood Co.,
April (A); Orangeburg Co., April (A); Richmond Co., April (A), July (A). TENNESSEE, Obion
Co., June (A).
Caenis hilaris (Say)
Physiographic provinces: CP, P, VR, AP
Previously published state records: Ala., Ber., 1958; Fla., Ber., 1950 and 1958, PJ, 1973, Sch,
1967; Ga., Ber., 1958.
Additional records: ALABAMA, Barbour Co., June (N); Bibb Co., June (A); Cherokee Co.,
July (A); Coosa Co., June (N); Dallas Co., June (N,A); DeKalb Co., June (N); Escambia
Co., June (N); Macon Co., June (N,A); Perry Co., June (N,A); Tuscaloosa Co., June (N);
Walker Co., June (N,A). FLORIDA, Escambia Co., August (A); Gadsden Co., June (A);
Liberty Co., June (N); Santa Rosa Co., June (A). GEORGIA, Cherokee Co., June (N,A),
August (A); Decatur Co., June (A). LOUISIANA, Orleans Co., April (A); Union Co., July
(A). MississiPPI, Copiah Co., August (N); Covington Co., August (N); Forrest Co., August
(A); Franklin Co., June, August (N); Lamar Co., August (A); Lauderdale Co., June (A),
July (A); Lawrence Co., August (A); Lowndes Co., June (A); Pike Co., June, August (A);
Smith Co., August (N); Sunflower Co., July (A). NORTH CAROLINA, Johnston Co., Septem-
ber (A); Wayne Co., June (A). SOUTH CAROLINA, Aiken Co., June2, July2, August2 (A);
Chesterfield Co., July (A); Spartanburg Co., June (A). TENNESSEE, DeKalb Co., July (A).


Vol. 22, No. 1







BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


Caenis jocosa McDunnough
Physiographic provinces: CP, VR, AP, ILP
Previously published state records: Ala., Trav., 1937.
Additional records: ALABAMA, Bibb Co., June (A); Lauderdale Co., June (A); Walker Co.,
June (A). LOUISIANA, Natchitoches Co., March (A), April (A). MississiPPi, Coahoma Co.,
July (N,A); Lafayette Co., July (N,A).
Caenis ridens McDunnough
Physiographic provinces: CP, VR, AP, ILP
Previously published state records: Ala., Trav., 1937.
Additional records: LOUISIANA, (county unknown) July (A). TENNESSEE, Davidson Co., July
(A); Sevier Co., July (A), August (N,A).
Caenis simulans McDunnough
Physiographic provinces: CP, AP
Previously published state records: None
Additional records: LOUISIANA, Calcasieu Par., March (A); Lafayette Par., March, October,
November (A). TENNESSEE, Fentress Co., May (A).

Caenis species
Physiographic provinces: CP, P, BR, VR, AP, ILP
Additional records: GEORGIA, Baker Co., May (N); Burke Co., August, September (N); Dawson
Co., June (N); Effingham Co., July (N); Jackson Co., July (N); Johnson Co., August (N);
Seminole Co., June (N). LOUISIANA, Lafayette Co., October (A), November (N,A); Natchi-
toches Co., July (A); Washington Co., August (N). MississiPPi, Lamar Co., May (N),
August (N). NORTH CAROLINA, Cabarrus Co., July (N); Davidson Co., July (N); Granville
Co., July (N); Haywood Co., August (N); Henderson Co., June (N); Macon Co., July,
August (N); Person Co., July (N); Polk Co., July (N); Randolf Co., July (N); Rockingham
Co., July (N); Transylvania Co., August (N); Wilkes Co., June (N). SOUTH CAROLINA, Aiken
Co., July (N); Anderson Co., July (N); Edgefield Co., April (N), July (N); Fairfield Co.,
April (N); Greenville Co., August (N); Kershaw Co., July (N); Lancaster Co., June (N),
July (N); Laurens Co., April (N); Marlboro Co., July (N); Orangeburg Co., October (N);
Pickens Co., August (N). TENNESSEE, Benton Co., June (N); Davidson Co., October (N);
Giles Co., July (N); Lincoln Co., July (N); Rutherford Co., July (N).

SUPERFAMILY PROSOPISTOMATOIDEA
FAMILY BAETISCIDAE
SUBFAMILY BAETISCINAE
GENUS Baetisca WALSH

Baetisca becki Schneider and Berner
Physiographic provinces: CP
Previously published state records: Fla., SB, 1963, PJ, 1973, PP, 1971, PP, 1974, Sch, 1967.
Additional records: FLORIDA, Escambia Co., May (N); Okaloosa Co., May (A); Santa Rosa
Co., (N).
Baetisca callosa Traver
Physiographic provinces: CP.
Previously published state records: Miss., SB., 1963.
Additional records: None
Baetisca carolina Traver
Physiographic provinces: P, BR.
Previously published state records: N. C. Trav. 1931 and 1937, H, 1941 (as B. thomsenae),
Ber., 1955; Tenn., WB, 1949 (as B. thomsenae).
Additional records: GEORGIA, Cherokee Co., June (N). NORTH CAROLINA, Buncombe Co., May
(N); Henderson Co., March (N); Jackson Co., June (N); Macon Co., June (A), August (N);
McDowell Co., April, May (N); Polk Co., July (N); Surry Co., February (N); Transylvania


1977







BULLETIN FLORIDA STATE MUSEUM


Co., May, June, (N), August (N); Yadkin Co., May (N). TENNESSEE, Greene Co., March
(A), April (N), May (N).
Baetisca escambiensis Berner
Physiographic provinces: CP.
Previously published state records: Fla., Ber., 1955, PJ, 1973, Sch, 1967.
Additional records: FLORIDA, Escambia Co., October (N), November (A); Okaloosa Co.,
August (N); Santa Rosa Co., August (N).
Baetisca gibbera Berner
Physiographic provinces: CP.
Previously published state records: Fla., Ber., 1955, PJ, 1973, Sch, 1967; Ga., Ber., 1955 and
1958.
Additional records: FLORIDA, Clay Co., January (N); Escambia Co., October (N); Holmes Co.,
September (N). GEORGIA, Baker Co., November, (N); Echols Co., February (N); Merri-
weather Co., September (N). MississiPPI, Itawamba Co., July (N). SOUTH CAROLINA,
North Anna River, September (N)2.
Baetisca obesa (Say)
Physiographic provinces: CP, P.
Previously published state records: Fla., Ber., 1953, 1955 and 1958, PJ, 1973, Sch, 1967; Ga.,
NTH, 1935, Ber., 1955; Miss., Ber., 1955.
Additional records: FLORIDA, Escambia Co., April (N); Gulf Co., April (N); Hamilton Co.,
February (N), March (N,A); Madison Co., January (N), March (A); Okaloosa Co., March
(N), April (N); Santa Rosa Co., April (N). GEORGIA, Cowetta Co., April (N)3; Echols Co.,
February (N); Lowndes Co., February (A), March (N,A). MississipPI, Rankin Co., April
(A). SOUTH CAROLINA, Aiken Co., May (N)'; Chesterfield Co., May (A)'; Orangeburg Co.,
April (A).
Baetisca rogersi Bemer
Physiographic provinces: CP, P.
Previously published state records: Ala., Ber., 1958; Fla., Ber., 1950, 1955, and 1958, PI,
1973, Sch, 1967; Ga., Ber., 1955 and 1958, PP, 1974; S. C., PP, 1974.
Additional records: ALABAMA, Barbour Co., June (N); Escambia Co., March (N); April (N),
May (N), June (A); Washington Co., May (N). FLORIDA, Escambia Co., January (N), April
(N); Gadsden Co., April (N); Liberty Co., March (N), April (N,A); Okaloosa Co., March
(N); Santa Rosa Co., August (N), September (N). GEORGIA, Cherokee Co., June (N); De-
catur Co., March (N); Peach Co., April (N). SOUTH CAROLINA, Aiken Co., January (N)',
May (N)2; Lexington Co., April (N).
Baetisca species
Physiographic provinces: CP.
Additional records: SOUTH CAROLINA, Aiken Co., May (N)3 (identified as B. laurentina by
S. Roback); Orangeburg Co., October (N).



SUPERFAMILY LEPTOPHLEBIOIDEA
FAMILY LEPTOPHLEBIIDAE
SUBFAMILY LEPTOPHLEBIINAE
GENUS Choroterpes EATON

Choroterpes basalis (Banks)
Physiographic provinces: VR
Previously published state records: None
Additional records: GEORGIA, Walker Co., August (A).


Vol. 22, No. 1








BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


Choroterpes hubbelli Berner
Physiographic provinces: CP
Previously published state records: Fla., Ber., 1950, Sch, 1967.
Additional records: ALABAMA, Dallas Co., June (N). FLORIDA, Alachua Co., August (A); Glades
Co., October (N); Hamilton Co., July (A). GEORGIA, Thomas Co., May (N), June, July
(A)'; Toombs Co., August (N). LOUISIANA, Tangipahoa Par., June (N,A). MISSISSIPPI,
Forrest Co., August (A); Harrison Co., August (N); Lamar Co., May (N), August (N,A);
Lauderdale Co., July (A).
Choroterpes species
Physiographic provinces: CP, ILP
Additional records: GEORGIA. Colquitt Co., (N); Tift Co., October (N). LOUISIANA, St. Helena
Par., August (N). MissIssIPPI, Clay Co., August (N); Harrison Co., August (N); Marion
Co., August (N). TENNESSEE, Davidson Co., June, July (A), October (N); Giles Co., July
(N); Lincoln Co., July (N); Robertson Co., June (N); Rutherford Co., July (N); Williamson
Co., July (A).


GENUS Habrophlebia EATON
Habrophlebia species
Physiographic provinces: BR
Additional records: NORTH CAROLINA, Macon Co., June and July (N); McDowell Co., August
(A).
Habrophlebia vibrans Needham
Physiographic provinces: CP, P, BR
Previously published state records: Fla., Ber., 1950, PJ, 1973, Sch, 1967; N. C., Banks, 1914
(as H. jocosa), NTH, 1935 (as H. pusilla); S. C., NTH, 1935 (as H. pusilla).
Additional records: ALABAMA, Escambia Co., April (A); Tuscaloosa Co., May (A). GEORGIA,
DeKalb Co., June (A)3; Peach Co., April (A); NORTH CAROLINA, Haywood Co., July (A);
Jackson Co., July (A); Macon Co., July (A), June (A); Swain Co., June (A). SOUTH CARO-
LINA, Greenville Co., April (A), August (A)'; Lexington Co., April (A). TENNESSEE, Sevier
Co., June (A), December (N).


GENUS Habrophlebiodes ULMER
Habrophlebiodes americana (Banks)
Physiographic provinces: CP, P, BR, VR, AP
Previously published state records: N. C., NTH, 1935, Trav., 1937 (as H. betteni); S. C., NTH,
1935; Tenn., WB, 1949.
Additional records: ALABAMA, Tuscaloosa Co., May (A). GEORGIA, DeKalb Co., June (A)';
Gordon Co., June (A)3; Houston Co., May (A)3, White Co., June (A)'. NORTH CAROLINA
Clay Co., April (N); Haywood Co., July (A); Jackson Co., June, August, September (A);
Macon Co., June, July, August (A), July (N); Yancey Co., (A). SOUTH CAROLINA, Green-
ville Co., May, June, September (A)*; Oconee Co., July (A). TENNESSEE, Benton Co., June
(A); Greene Co., April, May, June (A); Sevier Co., May, June (A)', June (A).
Habrophlebiodes brunneipennis Berner
Physiographic provinces: CP
Previously published state records: Ala., Ber., 1958; Fla., Ber., 1950 and 1958, PJ, 1973, Sch,
1967; Ga., Ber., 1958.
Additional records: ALABAMA, Conecuh Co., June (N); Escambia Co., June (N); Mobile Co.,
June (N). FLORIDA, Alachua Co., January (A); Gadsden Co., June (N); Liberty Co., March
(N); June (A). GEORGIA, Decatur Co., March (N,A), April (A). SOUTH CAROLINA, Aiken
Co., May (A).








BULLETIN FLORIDA STATE MUSEUM


Habrophlebiodes celeteria Berner
Physiographic provinces: BR
Previously published state records: Tenn., Ber., 1975.
Additional records: None.
Habrophlebiodes species
Physiographic provinces: CP, P, BR, VR, AP.
Additional records: ALABAMA, Calhoun Co., July (A); DeKalb Co., 1954 (N); Elmore Co.,
April (N); Escambia Co., April (N,A); Henry Co., June (N); Randolph Co., June (N); Tusca-
loosa Co., April, June (N). GEORGIA, Dawson Co., June (N); Forsyth Co., June (N); Macon
Co., April (N); Peach Co., April (N); Rabun Co., July, August (N). LOUISIANA, Tangipahoa
Co., June (N). MississiPPi, Perry Co., June (N); Stone Co., August (N). NORTH CAROLINA,
Avery Co., February (N); Buncombe Co., May (N); Macon Co., May-July (N); Orange Co.,
May (N); Swain Co., May, June (N)'. Transylvania Co., May (N); Yancey Co., May and
July (N). SOUTH CAROLINA, Edgefield Co., April, July (N); Lexington Co., April (N);
Orangeburg Co., April (A). TENNESSEE, Benton Co., June (A); Blount Co., June (N); Greene
Co., April, May, June (A); Sevier Co., June (A)', May, June (A), December (N); Unicoi Co.,
September (A).


GENUS Leptophlebia WESTWOOD
Leptophlebia austrina (Traver)
Physiographic provinces: BR
Previously published state records: N. C., NTH, 1935.
Additional records: GEORGIA, Rabun Co., April (A)3. SOUTH CAROLINA, Greenville Co., April,
March (A)'.
Leptophlebia bradleyi Needham
Physiographic provinces: CP
Previously published state records: Fla., Ber., 1950, Sch., 1967; Ga., Need., 1932, NTH, 1935,
Ber., 1958, 1975.
Additional records: ALABAMA, Baldwin Co., December (N). FLORIDA, Alachua Co., January,
February (A); Bay Co., February (A); Clay Co., January (A); Hamilton Co., November
(N). GEORGIA, Bryan Co., December (N); Coffee Co., March (N,A); Dooley Co., December
(A); Echols Co., November (N); Houston Co., March (A); Lowndes Co., February (N,A).
LOUISIANA, St. Tammany Par., November (N).
Leptophlebia collina (Traver)
Physiographic provinces: CP, P, BR
Previously published state records: N. C., NTH, 1935; S. C., NTH, 1935.
Additional records: FLORIDA, Baker Co., March (A). GEORGIA, Fulton Co., March (A)3;
Lowndes Co., February (A); Stephens Co., May (A)'. SOUTH CAROLINA, Anderson Co.,
April (A)'; Greenville Co., March-April (A)'.
Leptophlebia cupida (Say)
Physiographic provinces: P, BR, VR, ILP
Previously published state records: N. C., Eaton, 1884; Tenn., WB, 1949.
Additional records: GEORGIA, Paulding Co., March (A)'; Rabun Co., May (A)3. TENNESSEE,
Greene Co., February, March (N); Hawkins Co., April (A); Knox Co., March (N); Lincoln
Co., April (N,A).
Leptophlebia grandis (Traver)
Physiographic provinces: P
Previously published state records: N. C., NTH, 1935
Additional records: None
Leptophlebia intermedia (Traver)
Physiographic provinces: CP, P


Vol. 22, No. 1







BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


Previously published state records: Ala., Ber., 1958, PJ, 1973, Sch, 1967; Fla., Ber., 1950
and 1958; Ga., Ber., 1958; N. C., NTH, 1935.
Additional records: GEORGIA, Berrien Co., March (A); Charlton Co., February (N,A);
Dougherty Co., November (N); DeKalb Co., April (A)3; Grady Co., November (N);
Lowndes Co., February (A), March (A); Macon Co., April (N,A); Thomas Co., May (N).
SOUTH CAROLINA, Chesterfield Co., December (N).
Leptophlebia johnsoni McDunnough
Physiographic provinces: P
Previously published state records: N. C., NTH, 1935 (As L. gracilis Trav.).
Additional records: None
Leptophlebia species
Physiographic provinces: CP, P, BR, VR.
Additional records: ALABAMA, Houston Co., February (N); FLORIDA, Okaloosa Co., May (A).
GEORGIA, Burke Co., April (N)3; Catoosa Co., March (N)'; Clayton Co., March (N)3;
Columbia Co., April (N)'; Crawford Co., March (A)'; DeKalb Co., April (N)3; Fayette
Co., March (A)'; Fulton Co., April (N)3; Hall Co., March (N); Henry Co., March (N)3;
Lowndes Co., March (N)3; Richmond Co., April (N)3; Sumter Co., December (N). LOUISI-
ANA, St. Tammany Par. March (N); Tangipahoa Par., June (N). MississIPPI, Harrison
Co., December (N). NORTH CAROLINA, McDowell Co., April (N,A); Orange Co., Febru-
ary (N). SOUTH CAROLINA, Greenville Co., March, June (N)'; Lexington Co., April (N,A).


GENUS Paraleptophlebia LESTAGE
Paraleptophlebia adoptive (McDunnough)
Physiographic provinces: BR
Previously published state records: N. C., Travy., 1937, H, 1971.
Additional records: GEORGIA, Lumpkin Co., April (N); Towns Co., June (A)3
Paraleptophlebia assimilis (Banks)
Physiographic provinces: BR
Previously published state records: N. C., NTH, 1935; S. C., NTH, 1935.
Additional records: GEORGIA, Union Co., May (A)3; White Co., June (A)'. NORTH CAROLINA,
Clay Co., April (A); Macon Co., July (A). SOUTH CAROLINA, Greenville Co., April (A)4.
Paraleptophlebia debilis (Walker)
Physiographic provinces: BR
Previously published state records: N. C., Travy., 1932, H, 1941; S. C., PCR, 1967.
Additional records: NORTH CAROLINA, Watauga Co., August (A)
Paraleptophlebia georgiana Traver
Physiographic provinces: VR.
Previously published state records: Ga., NTH, 1935.
Additional records: GEORGIA, Whitfield Co., May and June (A)3.
Paraleptophlebia guttata (McDunnough)
Physiographic provinces: BR, VR, ILP.
Previously published state records: N. C., NTH, 1935, Tray., 1937, H, 1941; Tenn., WB, 1949.
Additional records: GEORGIA, Rabun Co., June (A)3; White Co., June (A)'. NORTH CAROLINA,
Avery Co., August (A); Haywood Co., July (A); Jackson Co., August (N,A), September
(A); Macon Co., June, July, August (A); Swain Co., August (A). SOUTH CAROLINA, Oconee
Co., July (A). TENNESSEE, Claiborne Co., July (A); Davidson Co., May (A); Dickson Co.,
June(N); GreeneCo., May,June(A); HickmanCo.,June(N,A); HoustonCo.,June(N,A); Sevier
Co., June (A); Wayne Co., July (N,A).
Paraleptophlebia jeanae Berner
Physiographic provinces: CP, P








BULLETIN FLORIDA STATE MUSEUM


Previously published state records: None
Additional records: ALABAMA, Sumter Co., April (A). SOUTH CAROLINA, Edgefield Co., April
(A).

Paraleptophlebia moerens (McDunnough)
Physiographic provinces: VR
Previously published state records: N. C., NTH, 1935.
Additional records: GEORGIA, Gilmer Co., June (A)3. TENNESSEEE, Claiborne Co., July (A).

Paraleptophlebia mollis (Eaton)
Physiographic provinces: BR
Previously published state records: N. C., Eaton, 1884, Tray., 1937.
Additional records: None.

Paraleptophlebia species
Physiographic provinces: CP, P, BR, VR, AP, ILP.
Additional records: ALABAMA, DeKalb Co., April, May, July (N); Elmore Co., April (N);
Tuscaloosa Co., April (N). GEORGIA, Crawford Co., May (A)3; Dawson Co., June (N,A);
Hancock Co., August (N,A); Heard Co., July (N); Lumpkin Co., April (A)', June (A)3;
Union Co., June (A)3. MIssIssIPPI, Claibome Co., April (N); Franklin Co., August (N);
Lafayette Co., July (N); Monroe Co., July (N); Stone Co., August (N). NORTH CAROLINA,
Avery Co., April (N); Jackson Co., June and August (A); Macon Co., July, August, and
September (N), June, July, and August (A); McDowell Co., August (A); Orange Co., May
(N); Pender Co., March and April (A); Person Co., July (A); Swain Co., April, August,
December (N), June (N,A), August (A); Transylvania Co., March and June (N); Watauga
Co., July (A). SOUTH CAROLINA, Edgefield Co., April, July (N); Laurens Co., April (A);
Newberry Co., April (A); Oconee Co., July (A). TENNESSEE, Blount Co., June (N); Greene
Co., August (N); Henderson Co., June (N); Robertson Co., June (N); Sevier Co., May
(N)', May-August, December (N).

Paraleptophlebia swannanoa (Traver)
Physiographic provinces: CP, BR, VR.
Previously published state records: N. C., NTH, 1935, Tray., 1937, H, 1941; S. C., NTH,
1935.
Additional records: GEORGIA, Fannin Co., June (A)3; White Co., June (N)'. NORTH CAROLINA,
Haywood Co., July (A). SOUTH CAROLINA, Greenville Co., October (A)'; Orangeburg Co.,
April (N,A).

Paraleptophlebia volitans (McDunnough)
Physiographic provinces: CP, P, BR, VR, AP.
Previously published state records: Fla., Ber., 1950 and 1958, PJ, 1973, Sch, 1967; Ga., NTH,
1935.
Additional records: ALABAMA, Baldwin Co., June (N); Conecuh Co., June (N); Covington
Co., June, July (N); Dallas Co., June (N); DeKalb Co., July (N); Elmore Co., April (N);
Escambia Co., April, June (N,A); Mobile Co., June (N,A); Monroe Co., April (N). FLORIDA,
Columbia Co., November (N); Escambia Co., October (N); Gadsden Co., April (N); Santa
Rosa Co., August (N). GEORGIA, Crawford Co., May (A)'; Early Co., February (N); Echols
Co., February (N); Gordon Co., June (A)'; Lanier Co., August (A)'; Lowndes Co., March
(N,A); Macon Co., April (N,A); Peach Co., April (A), June (A)3; Schley Co., April (A)';
Taylor Co., April (A)'. LOUISIANA, Tangipahoa Par., June (A). MIssIssIPPI, Lafayette
Co., July (A); Lamar Co., May (A), August (A); Marion Co., August (A). NORTH CAROLINA,
Transylvania Co., July (A). SOUTH CAROLINA, Aiken Co., May (A), July (N); Greenville
Co., June (A)'; Kershaw Co., July (N); Lexington Co., April (N,A); Orangeburg Co., April
(A).


Vol. 22, No. 1








BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


FAMILY EPHEMERELLIDAE
SUBFAMILY EPHEMERELLINAE
GENUS Ephemerella WALSH
Ephemerella alleghenensis Traver
Physiographic provinces: BR
Previously published state records: N. C., AE, 1962.
Additional records: GEORGIA, Lumpkin Co., July, August (N). NORTH CAROLINA, Graham Co.,
August (N); Macon Co., August (N); McDowell Co., August (N); Swain Co., August (N).
TENNESSEE, Unicoi Co., August (N).
Ephemerella argo Burks
Physiographic provinces: CP
Previously published state records: Ga., S. C., PCR, 1967.
Additional records: None
Ephemerella attenuate McDunnough
Physiographic provinces: CP, P, BR, VR, AP.
Previously published state records: Ala., AE, 1961; Fla., Ber., 1950 and 1958 (as E. hirsuta),
PJ, 1973, Sch, 1967 (as E. h irsuta); Ga., AE, 1961; S. C., AE, 1961.
Additional records: ALABAMA, Coosa Co., June (N); Escambia Co., April (N); Macon Co.,
April (N); Walker Co., June (N). FLORIDA, Escambia Co., March (N); Liberty Co., April
(N). GEORGIA, Cherokee Co., June (N); Dawson Co., June (N); Macon Co., April (N); Peach
Co., April (N). NORTH CAROLINA, Macon Co., July (N). SOUTH CAROLINA, Aiken Co., May
(A); Edgefield Co., April (N); Lexington Co., April (N).
Ephemerella bemeri Allen and Edmunds
Physiographic provinces: P
Previously published state records: Ga., AE, 1965
Additional records: None
Ephemerella bicolor Clemens
Physiographic provinces: BR, ILP
Previously published state records: N. C., AE, 1963; Tenn., WB, 1949.
Additional records: TENNESSEE, Greene Co., April (N); Lincoln Co., April (N).
Ephemerella carolina Berner and Allen
Physiographic provinces: BR
Previously published state records: N. C., BA, 1961, AE, 1963; Tenn., AE, 1963.
Additional records: NORTH CAROLINA, Macon Co., July, August (N); Polk Co., July (N); Swain
Co., June, August (N); Transylvania Co., July (N).
Ephemerella catawba Traver
Physiographic provinces: P, BR, VR, AP
Previously published state records: Ga., AE, 1965; N. C., NTH, 1935, Trav., 1937; Tenn.,
WB, 1949, AE, 1965.
Additional records: GEORGIA, Cherokee Co., June (N); Clark Co., April (N)'; Fulton Co.,
April (N); Rabun Co., May (N)3; Union Co., May (N)'. Walker Co., April (N). NORTH
CAROLINA, Avery Co., February (N); Buncombe Co., April (N)', May (N); Haywood Co.,
July (N); Jackson Co., April (N)', June (N); Macon Co., May, July, August (N), September
(A); McDowell Co., April (N); Polk Co., July (N); Swain Co., May (N)', April, May, August
(N); Transylvania Co., June (N); Yancey Co., July (N). SOUTH CAROLINA, Greenville Co.,
August (N). TENNESSEE, Cocke Co., April (N); Cumberland Co., April (N); Greene Co.,
May (N); Marion Co., April (N); Sevier Co., May, June (N)'.
Ephemerella choctawhatchee Berner
Physiographic provinces: CP
Previously published state records: Fla., Ber., 1950 and 1958, AE, 1965, Sch, 1967; Ga., Ber.,
1958.







BULLETIN FLORIDA STATE MUSEUM


Additional records: FLORIDA, Gadsden Co., April (N); Liberty Co., March (A,N), April (A).
GEORGIA, Burke Co., January (N)2; Decatur Co., March (A,N); Early Co., February (N).
SOUTH CAROLINA, Aiken Co., May (N)2; Allendale Co., May (N)2.
Ephemerella conestee Traver
Physiographic provinces: BR, AP.
Previously published state records: N. C., NTH, 1935, H, 1941, AE, 1962; Tenn., NTH, 1935,
AE, 1962.
Additional records: ALABAMA, DeKalb Co., April (N). GEORGIA, Lumpkin Co., July (N)3;
Rabun Co., (N). NORTH CAROLINA, Avery Co., August (N); Jackson Co., August (N); Macon
Co., June, July, August, September (N); McDowell Co., August (N); Swain Co., August
(N); Transylvania Co., June, July, August (N); Watauga Co., July (N); Yancey Co., July,
August (N). SOUTH CAROLINA, Oconee Co., August (N). TENNESSEE, Greene Co., August
(N).
Ephemerella cornuta Morgan
Physiographic provinces: BR
Previously published state records: N. C., NTH, 1935, Trav., 1937, AE., 1962; Tenn., Burks,
1953.
Additional records: GEORGIA, Tray Mt., July (A)3. NORTH CAROLINA, Jackson Co., April (N)';
Macon Co., June (N); Swain Co., May (N)', July (N), June (A,N), August (N); Transylvania
Co., June, July (N). TENNESSEE, Sevier Co., May, June (N)'.

Ephemerella cornutella McDunnough
Physiographic provinces: BR, VR.
Previously published state records: Ga., NTH, 1935; N. C., NTH, 1935, Trav., 1937, H, 1941,
AE, 1962; Tenn., WB, 1949, AE, 1962.
Additional records: GEORGIA, Rabun Co., May (N); White Co., June (A)'. NORTH CAROLINA,
Macon Co., August (N); McDowell Co., May (N); Swain Co., May-July (N); Transylvania
Co., July (N); Watauga Co., June, August (N). TENNESSEE, Blount Co., June (N); Greene
Co., May (N); Hawkins Co., July (N); Sevier Co., June (N).

Ephemerella coxalis McDunnough
Physiographic provinces: BR, VR
Previously published state records: Ga., NTH, 1935; N. C., NTH, 1935, Trav., 1937; Tenn.,
WB, 1949.
Additional records: GEORGIA, Catoosa Co., July (N)3. NORTH CAROLINA, Henderson Co., June
(N)'. TENNESSEE, Greene Co., March, May (N).

Ephemerella crenula Allen and Edmunds
Physiographic provinces: BR
Previously published state records: N. C., AE, 1965; Tenn., AE, 1965.
Additional records: NORTH CAROLINA, Swain Co., May (N); Yancey Co., May (N).

Ephemerella deficiens Morgan
Physiographic provinces: CP, P, BR, VR, AP, ILP
Previously published state records: Ala., BA, 1961; Fla., Ber., 1950 (as Ephemerella sp. A),
BA, 1961, PJ, 1973, Sch, 1967; Ga., Burks, 1953, BA, 1961; Miss., BA, 1961; N. C., NTH,
1935, Burks, 1953, BA, 1961; S. C., BA, 1961, PCR, 1967; Tenn., WB, 1949, BA, 1961.
Additional records: ALABAMA, Bibb Co., June (A,N), May (N); Coosa Co., June (N); Covington
Co., July (N); Dallas Co., June (N); Elmore Co., June (N); Lauderdale Co., June, July
(N); Macon Co., June (N); Mobile Co., June (N); Shelby Co., June (N); Tuscaloosa Co.,
June (N); Walker Co., June (N). FLORIDA, Gadsden Co., April (N); Liberty Co., March
(N). GEORGIA, Cherokee Co., June, August (N); Dawson Co., June (N); Decatur Co., April,
August (N); Early Co., February (N); Peach Co., April (N); Pickens Co., June (N); Pike
Co., April (N)3; Rabun Co., July (N); Wilkes Co., July (N). MISSIsSIPPI, Lauderdale Co.,
July (N); Pike Co., August (N). NORTH CAROLINA, Caldwell Co., July (N); Davidson Co.,
July (N); Jackson Co., June (N); Macon Co., August (N); Swain Co., May (N)', June (N,A);


Vol. 22, No. 1








BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


Transylvania Co., June (N), July (A). SOUTH CAROLINA, Aiken Co., May (N); Edgefield
Co., July (N); Fairfield Co., April (N); Greenville Co., August (N). TENNESSEE, Blount
Co., June (N); Hawkins Co., August (N); Polk Co., August (N); Sevier Co., May (N)', July
(A), August (N); Unicoi Co., August (N).
Ephemerella doris Traver
Physiographic provinces: CP, P, BR, VR
Previously published state records: Ala., Tray., 1937; Ga., NTH, 1935, AE, 1963; N. C., NTH,
1935, Tray., 1937, H, 1941, AE, 1963; Tenn., WB, 1949.
Additional records: ALABAMA, Baldwin Co., June (N); Bibb Co., March (N)3; Cherokee Co.,
June (N); Lowndes Co., March (N)3; Thomas Co., April (N)3. LOUISIANA, Lincoln Co.,
February, March, 1952 (N). NORTH CAROLINA, McDowell Co., April (N); Orange Co.,
May (N). SOUTH CAROLINA, Aiken Co., May (A); Allendale Co., January (N)2; Kershaw
Co., April (N); Orangeburg Co., April (A). TENNESSEE, Cocke Co., April (N); Hardin Co.,
June (N).
Ephemerella dorothea Needham
Physiographic provinces: P, BR
Previously published state records: Fla., Sch, 1967; N. C., McD., 1931, NTH, 1935, Tray.,
1937, AE, 1965; S. C., NTH, 1935; Tenn., McD., 1931, NTH, 1935, WB, 1949, AE, 1965.
Additional records: GEORGIA, Union Co., May (A)3. NORTH CAROLINA, Avery Co., April (N);
Buncombe Co., May (N); Macon Co., July, August (N); McDowell Co., May (N); Swain
Co., April, May (N); Transylvania Co., May, June, July (N). SOUTH CAROLINA, Edgefield
Co., April (A,N); Fairfield Co., April (N); Greenville Co., May, June, September (A)';
Laurens Co., April (N); Newberry Co., April (N). TENNESSEE, Cocke Co., June (A).
Ephemerella frisoni McDunnough
Physiographic provinces: ILP
Previously published state records: Ala., BA, 1961
Additional records: None
Ephemerella funeralis McDunnough
Physiographic provinces: BR, VR, ILP
Previously published state records: Ga., NTH, 1935; N. C., Trav., 1937, H, 1941, EA, 1963;
S. C., NTH, 1935; Tenn., AE, 1963.
Additional records: NORTH CAROLINA, Haywood Co., July (N); Macon Co., January (N); Mc-
Dowell Co., April (N). TENNESSEE, Cheatham Co., April (N); Knox Co., February (N);
Sevier Co., May (N)', December (N).
Ephemerella hispida Allen and Edmunds
Physiographic provinces: BR
Previously published state records: N. C., AE, 1965; Tenn., AE, 1965.
Additional records: NORTH CAROLINA, Swain Co., May (N).
Ephemerella inconstans Traver
Physiographic provinces: P, BR
Previously published state records: N. C., Tray., 1932, NTH, 1935 (as E. vernalis). H, 1941,
AE, 1965; Tenn., WB, 1949 (As E. vernalis).
Additional records: GEORGIA, Cobb Co., March (N); Fulton Co., April (A)3; Greene Co.,
April (A)3. NORTH CAROLINA, Jackson Co., June (A); Macon Co., June (N). TENNESSEE,
Greene Co., April (A).
Ephemerella invaria (Walker)
Physiographic provinces: BR, VR, AP
Previously published state records: Ala., AE, 1965; Ga., AE, 1965; N. C., AE, 1965; Tenn.,
AE, 1965.
Additional records: GEORGIA, Rabun Co., May (N). NORTH CAROLINA, Swain Co., April (N).
TENNESSEE, Blount Co., April (N); Grainger Co., April (N); Greene Co., August (N); Moore
Co., April (N); Sevier Co., May (N)'.








BULLETIN FLORIDA STATE MUSEUM


Ephemerella lata Morgan
Physiographic provinces: BR
Previously published state records: N. C., NTH, 1935, Tray., 1937, AE, 1962; Tenn., WB,
1949, Burks, 1953.
Additional records: GEORGIA, Lumpkin Co., July (A)3. NORTH CAROLINA, Macon Co., July
(N); Transylvania Co., August (N).
Ephemerella lita Burks
Physiographic provinces: BR
Previously published state records: N. C., AE, 1962
Additional records: NORTH CAROLINA, Macon Co., June, July (N).
Ephemerella longicornis Traver
Physiographic provinces: BR
Previously published state records: N. C., NTH, 1935, Trav., 1937, H, 1941, AE, 1962; Tenn.,
AE, 1962.
Additional records: NORTH CAROLINA, Macon Co., July (N); Transylvania Co., June (N);
Yancey Co., July (N).
Ephemerella lutulenta Clemens
Physiographic provinces: BR, AP, ILP
Previously published state records: N. C., NTH, 1935; Tenn., WB, 1949.
Additional records: NORTH CAROLINA, Swain Co.; April (N), May (N)'. TENNESSEE, Cumber-
land Co., April (N); Davidson Co., May (N); Putnam Co., April (N).
Ephemerella minimella McDunnough
Physiographic provinces: P, BR, VR, ILP
Previously published state records: N. C., NTH, 1935; Tenn., WB, 1949.
Additional records: NORTH CAROLINA, Macon Co., August (N). SOUTH CAROLINA, Lexington
Co., April (N). TENNESSEE, Cheatham Co., April (N); Grainger Co., April (N); Greene
Co., April (N); Meigs Co., April (N).
Ephemerella needhami McDunnough
Physiographic provinces: P, AP
Previously published state records: None
Additional records: ALABAMA, DeKalb Co., April (N). SOUTH CAROLINA, Edgefield Co.,
April (N).
Ephemerella prudentalis McDunnough
Physiographic provinces: CP
Previously published state records: N. C., AE, 1963
Additional records: None
Ephemerella rossi Allen and Edmunds
Physiographic provinces: BR
Previously published state records: Tenn., AE, 1965
Additional records: NORTH CAROLINA, Swain Co., April May (N); Transylvania Co., May,
July (N); Yancey Co., July (N). TENNESSEE, Blount Co., June (N); Sevier Co., May (N).
Ephemerella rotunda Morgan
Physiographic provinces: CP, P, BR, AP
Previously published state records: Fla., Ber., 1958, Sch, 1967; N. C., NTH, 1935, Trav.,
1937, H, 1941, Burks, 1953, AE, 1965.
Additional records: FLORIDA, Gadsden Co., March (A); Liberty Co., March (N). GEORGIA,
Rabun Co., June (A)3. NORTH CAROLINA, Buncombe Co., May (N); Macon Co., April (N);
McDowell Co., April (N); Transylvania Co., March (N); Watauga Co., February (N);
Yadkin Co., February (N). SOUTH CAROLINA, Aiken Co., May (N). TENNESSEE, Cumber-
land Co., July (N); Sevier Co., April (N).
Ephemerella septentrionalis McDunnough
Physiographic provinces: BR


Vol. 22, No. 1








BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


Previously published state records: N. C., Trav., 1937, AE, 1965.
Additional records: GEORGIA, May (A)3. NORTH CAROLINA, Buncombe Co., May (N); Mc-
Dowell Co., April (N); Transylvania Co., May (N).
Ephemerella serrata Morgan
Physiographic provinces: BR
Previously published state records: N. C., NTH, 1935, AE, 1963.
Additional records: NORTH CAROLINA, Macon Co., July (A), Swain Co., July (N,A), August
(A).
Ephemerella serratoides McDunnough
Physiographic provinces: CP, P, BR, VR, AP, ILP
Previously published state records: N. C., NTH, 1935, AE, 1963; Tenn., AE, 1963.
Additional records: ALABAMA, Coosa Co., June (N); Shelby Co., June (N). GEORGIA, Baker
Co., May (N); Cherokee Co., June, July, August (N), Lumpkin Co., July, August (N),
Pickens Co., June (N). NORTH CAROLINA, Graham Co., August (N); Macon Co., August
(N); Polk Co., July (N); Randolf Co., July (N); Swain Co., June, July (A), August (N); Tran-
sylvania Co., June (N); Union Co., July (N). SOUTH CAROLINA, Greenville Co., August (N).
TENNESSEE, Franklin Co., April (N); Robertson Co., June (N); Sevier Co., July (N); Unicoi
Co., August (N).
Ephemerella simplex McDunnough
Physiographic provinces: CP, P, BR, AP
Previously published state records: Fla., Ber., 1958; Ga.-S. C., PCR, 1967; N. C., NTH,
1935, Burks, 1953, AE, 1962.
Additional records: ALABAMA, Macon Co., April (N); Tuscaloosa Co., April (N), May (A,N).
FLORIDA, Liberty Co., March (N). GEORGIA, Macon Co., April (N). NORTH CAROLINA,
Jackson Co., June (N); Macon Co., June, August (N); Swain Co., May (N)', June (N); Wake
Co., May (N). SOUTH CAROLINA, Edgefield Co., April (A,N); Fairfield Co., April (N);
Laurens Co., April (N); Lexington Co., April (N).
Ephemerella sordida McDunnough
Physiographic provinces: CP, BR, AP
Previously published state records: N. C., AE, 1963
Additional records: ALABAMA, Perry Co., May (N); Walker Co., June (N). NORTH CAROLINA,
Macon Co., July (A). TENNESSEE, Cumberland Co., July (N).
Ephemerella species
Physiographic provinces: CP, BR, AP
Previously published state records: N. C., AE, 1963
Additional records: FLORIDA, Liberty Co., March (N). GEORGIA, Baker Co., March (A); Bibb
Co., April (N)'; Burke Co., June (N); Carroll Co., March (N); Cherokee Co., June (N,A),
August (A); Dawson Co., June (N); Fulton Co., March, April (A)3; Gilmer Co., June
(A)'; Habersham Co., August (A)'; Lumpkin Co., August (N); Pickens Co., June (N);
Rabun Co., April (N), July (N), August (N); Richmond Co., April (A); Sumter Co.,
August (N); White Co., July (A). NORTH CAROLINA, Clay Co., April (N); Jackson Co.,
August (A); Macon Co., June (N,A), July (N), August (A); Swain Co., May, June (N)',
April, December (N); Transylvania Co., July (N); Wake Co., May (N); Yancey Co., May
(N). SOUTH CAROLINA, Aiken Co., May (A); Oconee Co., August (N); Orangeburg Co.,
April (A), October (N); Pickens Co., April (N)'. TENNESSEE, Fentress Co., May (A); Greene
Co., May (A); Sevier Co., May (N)', April, June, December (N).
Ephemerella spiculosa Bemer and Allen
Physiographic provinces: BR
Previously published state records: N. C., BA, 1961, AE, 1963; Tenn., BA, 1961, AE, 1963.
Additional records: NORTH CAROLINA, Macon Co., July (N). TENNESSEE, Greene Co., August
(N).
Ephemerella subvaria McDunnough
Physiographic provinces: BR


1977








BULLETIN FLORIDA STATE MUSEUM


Previously published state records: None
Additional records: NORTH CAROLINA, Swain Co., April (N).

Ephemerella temporalis McDunnough
Physiographic provinces: BR
Previously published state records: Ga., NTH, 1935; N. C., NTH, 1935, Trav., 1937, AE,
1963; S. C., AE, 1963.
Additional records: GEORGIA, Habersham Co., July (A)', NORTH CAROLINA, Macon Co., July
(N).
Ephemerella trilineata Berner
Physiographic provinces: CP, VR, AP
Previously published state records: Fla., Ber., 1950 and 1958, AE, 1963, PJ, 1973, Ga., Sch,
1967; Ber., 1958.
Additional records: ALABAMA, Escambia Co., April, June (N); Mobile Co., June (N), Tusca-
loosa Co., May, June (N). FLORIDA, Alachua Co., January (N), April (A); Baker Co., Febru-
ary (N); Bay Co., April (N); Clay Co., January (N); Columbia Co., November (N); Escambia
Co., March, April (N); Gadsden Co., March, April (N,A); Gulf Co., April (N,A); Jackson
Co., April (A); Lake Co., December (N); Liberty Co., March (A); Madison Co., January
(N); Manatee Co., February (N); Okaloosa Co., April (A), March, April (N), May (A);
Walton Co., April (A), April (N). GEORGIA, Berrian Co., March (N); Burke Co., May (A)2,
October, November, December (N); Charlton Co., February (N); Chattachoochee Co.,
April (N); Decatur Co., April (N); Houston Co., April (A); Lowndes Co., February (A),
March (A,N); Whitfield Co., May (A)3. LOUISIANA, Lamar Co., May (N); Tangipohoa Co.,
June (N); Washington Co., August (N). MISSISsIPPI, Lamar Co., May (N). SOUTH CARO-
LINA, Orangeburg Co., April (A).
Ephemerella tuberculata Morgan
Physiographic provinces: BR
Previously published state records: N. C., NTH, 1935, Trav., 1937 (as E. cherokee), Burks,
1953, AE, 1962; Tenn., Burks, 1953, AE, 1962.
Additional records: GEORGIA, Cherokee Co., June (N); Lumpkin Co., April (N); Union Co.,
June (A)3. NORTH CAROLINA, Jackson Co., June (N); Macon Co., July (N); Swain Co.,
August (N); Transylvania Co., July (N). TENNESSEE, Sevier Co., August (N).
Ephemerella walker Eaton
Physiographic provinces: BR
Previously published state records: None
Additional records: NORTH CAROLINA, Buncombe Co., April (N)' (identified as E. simplex by
Burks)
Ephemerella wayah Traver
Physiographic provinces: BR
Previously published state records: N. C., NTH, 1935, TH, 1941, AE, 1962
Additional records: GEORGIA, Rabun Co., July (A)"; NORTH CAROLINA, Macon Co., June (N);
Swain Co., June, July (N); Transylvania Co., June (N); Yancey Co., July (N). TENNESSEE,
Blount Co., June (N).


FAMILY TRICORYTHIDAE
SUBFAMILY LEPTOHYPHINAE
GENUS Leptohyphes EATON
Leptohyphes dolani Allen
Physiographic provinces: CP, P
Previously published state records: Ga.-S. C. state line, A, 1967.
Additional records: GEORGIA, Burke Co., August (N), October (N)2; Effingham Co., July (N).
SOUTH CAROLINA, Aiken Co., July, September (N)2; Newberry Co., August (N).


Vol. 22, No. 1








BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


Leptohyphes robacki Allen
Physiographic provinces: CP
Previously published state records: Ga.-S. C. state line, A, 1967.
Additional records: None

GENUS Tricorythodes ULMER
Tricorythodes albilineatus Berner
Physiographic provinces: CP, VR
Previously published state records: Ala., Ber., 1958; Fla., Ber., 1950 and 1958, PJ, 1973, Sch,
1967; Ga., Ber., 1958.
Additional records: ALABAMA, Bibb Co., Escambia Co., June (N); Henry Co., June (N).
FLORIDA, Alachua Co., October (N); Escambia Co., October (A); Hamilton Co., July (A);
Levy Co., November (N); Liberty Co., June (A); Santa Rosa Co., August (N). GEORGIA,
Burke Co., July-September (N); Decatur Co., June (A); Effingham Co., July (N); Monroe
Co., July (N,A); Muscogee Co., June (N).
Tricorythodes allectus (Needham)
Physiographic provinces: BR
Previously published state records: None
Additional records: NORTH CAROLINA, Macon Co., August (A).
Tricorythodes atratus (McDunnough)
Physiographic provinces: BR
Previously published state records: None
Additional records: TENNESSEE, Sevier Co., August (A).
Tricorythodes species
Physiographic provinces: CP, P, BR, VR, AP, ILP
Additional records: ALABAMA, Bibb Co., May, June (N); Chilton Co., June (N); Coosa Co.,
June (N); Covington Co., July (N); Dallas Co., June (N); DeKalb Co., July (N); Elmore
Co., June (N); Lauderdale Co., June (N); Macon Co., June (N); Marion Co., June (N);
Perry Co., May, June (N); Randolf Co., June (N); Walker Co., June (N). GEORGIA, Baker
Co., May (N,A); Bibb Co., July (N); Cherokee Co., June, July, August (N); Dawson Co.,
June (N); Decatur Co., July (N)3, August (N); Hancock Co., August; Hart Co., September
(N); Heard Co., July (N); Lumpkin Co., August (N); Monroe Co., July (N); Newton Co.,
July (N); Wilkes Co., July (N). LOUISIANA, St. Helena Par., August (N); Tangipahoa Par.,
August (N). Mississippi, Amite Co., June (N); Clark Co., August (N); Clay Co., July (N);
Copiah Co., August (N); Franklin Co., June, August (N); Harrison Co., August (N); Ita-
wamba Co., July (N); Lafayette Co., July (N); Lamar Co., August (N); Lauderdale Co.,
June, July (N); Lawrence Co., August (N); Lowndes Co., June (N); Monroe Co., July (N);
Pike Co., August (N); Rankin Co., April (N); Smith Co., August (N); Sunflower Co., July
(N). NORTH CAROLINA, Caswell Co., July (N); Davidson Co., July (N); Forsyth Co., June
(N); Graham Co., August (N); Macon Co., August (N,A); Person Co., July (A); Transylvania
Co., August (N); Wilkes Co., June (N). SOUTH CAROLINA, Aiken Co., July (N); Edgefield
Co., July (N); Lancaster Co., June, July (N); Laurens Co., August (N); McCormick Co., July
(N); Newberry Co., August (N); Oconee Co., August (N); Orangeburg Co., October (N);
Pickens Co., August (N); Spartanburg Co., August (N). TENNESSEE, Dickson Co., June
(N,A); Giles Co., July (N); Greene Co., August (N); Houston Co., July (N); Lincoln Co.,
July (N); Moore Co., July (N); Robertson Co., June, July, August (N); Rutherford Co.,
July (N); Sumner Co., June (N); Warren Co., June (N).
Trichorythodes stygiatus McDunnough
Physiographic provinces: BR
Previously published state records: None
Additional records: NORTH CAROLINA, Macon Co., August (A).
Tricorythodes texanus Traver
Physiographic provinces: CP


1977







BULLETIN FLORIDA STATE MUSEUM


Previously published state records: None
Additional records: MISSISSIPPI, Marion Co., August (N,A).


SUPERFAMILY EPHEMEROIDEA
FAMILY BEHNINGIIDAE
SUBFAMILY BEHNINGIINAE
GENUS Dolania EDMUNDS AND TRAVER

Dolania americana Edmunds and Traver
Physiographic provinces: CP
Previously published state records: Fla., PJ, 1973, Sch, 1967; S. C., ET, 1959, PCR, 1967.
Additional records: FLORIDA, Okaloosa Co., May (N,A), August (N); Santa Rosa Co., February
August (N). SOUTH CAROLINA, Aiken Co., (N)'; Allendale Co., no date (N)2.


FAMILY POTAMANTHIDAE
SUBFAMILY POTAMANTHINAE
GENUS Potamanthus PICTET

Potamanthus distinctus Traver
Physiographic provinces: BR, VR
Previously published state records: Ga., McC., 1975; N. C., Tray., 1937; Tenn., McC., 1975.
Additional records: GEORGIA, Chatooga Co., June (A)'; Fannin Co., June (A)'; White Co.,
July and August (A). NORTH CAROLINA, McDowell Co., April (N), Watauga Co., July
(A). TENNESSEE, Sevier Co., July (A), August (N,A), August (A).
Potamanthus myops (Walsh)
Physiographic provinces: ILP
Previously published state records: Tenn., McC., 1975.
Additional records: TENNESSEE, Davidson Co., June (A)
Potamanthus neglectus Traver
Physiographic provinces: CP, ILP
Previously published state records: Miss., McC., 1975; Tenn., McC., 1975.
Additional records: None.
Potamanthus rufous Argo
Physiographic provinces: VR
Previously published state records: Tenn., McC., 1975.
Additional records: TENNESSEE, Hamilton Co., August (A).
Potamanthus species
Physiographic provinces: P, BR, VR, ILP
Additional records: GEORGIA, Catoosa Co., June (A)'; Cherokee Co., June (N); Hall Co., July
(A)3; Pickens Co., June (N). NORTH CAROLINA, Watauga Co., February (N). TENNESSEE,
Rutherford Co., July (N); Unicoi Co., August (N).
Potamanthus verticis (Say)
Physiographic provinces: CP, ILP
Previously published state records: Miss., McC., 1975; Tenn., NTH, 1935.
Additional records: MississiPPi, Lawrence Co., August (A); Lowndes Co., June (A); Pike
Co., August (A). TENNESSEE, Cheatham Co., July (A); Davidson Co., June (A); Lincoln
Co., July (A).


Vol. 22, No. 1







BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


FAMILY EPHEMERIDAE
SUBFAMILY EPHEMERINAE
GENUS Ephemera LINNAEUS
Ephemera blanda Traver
Physiographic provinces: BR, VR
Previously published state records: Ga., NTH, 1935; N. C., NTH, 1935; S. C., NTH, 1935.
Additional records: GEORGIA, Dawson Co., June (N); Gilmer Co., July (A)', June (A)3; Haber-
sham Co., July (A)'; Lumpkin Co., August (A); Rabun Co., June (A)3; Union Co., July
(A)3; White Co., July (A)', July, August (A). NORTH CAROLINA, Macon Co., July (A), June
(N,A); McDowell Co., April (N); Swain Co., August (N). SOUTH CAROLINA, Greenville
Co., May, June (A)'.
Ephemera guttulata Pictet
Physiographic provinces: BR, AP
Previously published state records: Ga., McC., 1975; N. C., NTH, 1935, Trav., 1937; S. C.,
McC., 1975; Tenn., Kenn., 1926, WB, 1949.
Additional records: GEORGIA, Lumpkin Co., May (A)3. NORTH CAROLINA, Swain Co., August
(N). TENNESSEE, Cumberland Co., July (N); Greene Co., May (N,A); Morgan Co., July
(N); Sevier Co., August (N); Unicoi Co., June (A), August (N).
Ephemera simulans Walker
Physiographic provinces: P
Previously published state records: Fla., Ber., 1950 (very doubtful record); S. C., McC., 1975.
Additional records: SOUTH CAROLINA, Fairfield Co., April (A).
Ephemera species
Physiographic provinces: CP, BR, VR, AP, ILP
Additional records: ALABAMA, Winston Co., August (N). GEORGIA, Fannin Co., June (N)3;
Screven Co., July (N). NORTH CAROLINA, Buncombe Co., May (N); Macon Co., July (N);
Polk Co., July (N); Transylvania Co., June (N); Watauga Co., February, July, August
(N). SOUTH CAROLINA, Oconee Co., August (N). TENNESSEE, Lincoln Co., April (N);
Marion Co., April (A)', April (A); Sevier Co., December (N).
Ephemera varia Eaton
Physiographic provinces: P, BR, AP, ILP
Previously published state records: Ga., McC., 1975; N. C., McC., 1975; Tenn., WB, 1949.
Additional records: GEORGIA, Rabun Co., July August (N). NORTH CAROLINA, Jackson Co.,
July (A), August (N,A); Macon Co., June-August (N), June, August (A); Randolf Co., July
(N); Watauga Co., July (A), June (N). TENNESSEE, Cumberland Co., July (N); Dickson Co.,
July (N); Greene Co., May (N,A), June, July, September (A); Rutherford Co., July (N);
Sevier Co., April (N); Unicoi Co., June (A); Wayne Co., July (N).


GENUS Hexagenia WALSH
Hexagenia atrocaudata McDunnough
Physiographic provinces: BR., VR
Previously published state records: Ga., NTH, 1935; N. C., NTH, 1935; S. C., PCR, 1967
(doubtful identification); Tenn., WB, 1949.
Additional records: GEORGIA, Bartow Co., August (A)3; Floyd Co., August (A)3; NORTH CARO-
LINA, Watauga Co., August (A). TENNESSEE, Unicoi Co., August (A).
Hexagenia bilineata (Say)
Physiographic provinces: CP, P, BR, VR, AP, ILP
Previously published state records: Ala., Trav., 1937, Spieth, 1941; Fla., Ber., 1950 and 1958,
Sch, 1967; Ga., NTH, 1935, Spieth, 1941, Ber., 1958; La., Eaton, 1883, Spieth, 1941; Miss.,


1977







BULLETIN FLORIDA STATE MUSEUM


NTH, 1935, Spieth, 1941, Ber., 1957; N. C., McC., 1975; S. C., PCR, 1967 (questionably),
McC., 1975; Tenn., NTH, 1935, Spieth, 1941, WB, 1949.
Additional records: ALABAMA, Baldwin Co., June (A); Chilton Co., June (A); Clarke Co.,
May (A); Colbert Co., August (A)2, Lauderdale Co., August (A)', July (A), June (A);
Washington Co., May (A). FLORIDA, Gadsden Co., June (A); Jackson Co., July (A).
GEORGIA, Catoosa Co., June (A)3; Cherokee Co., June (A); Clark Co., July (A); Cobb Co.,
July (A)'; Crawford Co., June (A)3; Decatur Co., June (A); Dougherty Co., June, July (A)',
Floyd Co., June, July (A)'; Fulton Co., July, August (A)', July (A)3; Gordon Co., July (A)';
Habersham Co., July (A)'; Harris Co., July (A)'; Macon Co., July (A); Screven Co., July
(A); Taylor Co., June (A)3; Upson Co., June (A)3; LOUISIANA, Beauregard Par., June (A);
East Baton Rouge Par., April and May (A); Tangipahoa Par., June (A). MISSISSIPPI,
Lawrence Co., August (A); Monroe Co., July (A). NORTH CAROLINA, Macon Co., June
(A), July (A); Swain Co., August (A). SOUTH CAROLINA, Abbeville Co., June (A)'; Aiken
Co., July (N,A)*; Anderson Co., June (A)'. TENNESSEE, Benton Co., August (A); David-
son Co., July (A); Fentress Co., July (A); Greene Co., June, July, August, (A); Hamblen
Co., August (A); Hamilton Co., August (A); Hardin Co., July (A); Humphreys Co., June
(A); Lincoln Co., July (A).
Hexagenia limbata occulta (Walker)
Physiographic provinces: BR
Previously published state records: N. C., Spieth, 1941.
Additional records: None
Hexagenia limbata venusta Eaton
Physiographic provinces: CP, P, AP, VR, ILP
Previously published state records: Tenn., Spieth, 1941.
Additional records: ALABAMA, Walker Co., June (A). GEORGIA, Bartow Co., June (A)'; De-
catur Co., April, June (A)'; Fannin Co., July (A)3; Floyd Co., June (A)'; Monroe Co.,
September (A)'. LOUISIANA, East Baton Rouge Par., May (A); Vermillion Par., July (A).
MIssIssIPPI, Laurence Co., August (A). TENNESSEE, Humphreys Co., June (A); Sumner Co.,
June (N).
Hexagenia munda elegans Traver
Physiographic provinces: CP, P, BR, VR, AP, ILP
Previously published state records: Ala., Spieth, 1941, Ber., 1958; Fla., NTH, 1935 (as H.
weewa), Spieth, 1941, Ber., 1950 and 1958, Sch, 1967; Ga., NTH, 1935, Travy., 1937 (as H.
kanuga), Spieth, 1941; S. C., Trav., 1937 (as H. kanuga), Spieth, 1941.
Additional records: ALABAMA, Baldwin Co., August (N); Barbour Co., June (N); Bibb Co.,
June (A); Cherokee Co., July (A); Chilton Co., June (A); Covington Co., July (N); Elmore
Co., June (A); Henry Co., June (N); Lauderdale Co., August (A)'; Macon Co., June (A);
Perry Co., June (A); Pike Co., June (N); Walker Co., June (A). FLORIDA, Alachua Co.,
May-October (A), October (N); Columbia Co., April (N), May (A); Escambia Co., August
(A), October (N); Gadsden Co., April (N), June (A); Hillsborough Co., June (A); Levy Co.,
July (A); Liberty Co., March, June (N); Okaloosa Co., April, May, July (A); Santa Rosa Co.,
June, August (A); Walton Co., April, July, August, October (A). GEORGIA, Bartow Co.,
August (A)'-'; Bibb Co., August (A)'; Clarke Co., June, August (A)'; Clayton Co., June
(A)3; Crawford Co., June (A)3; Dougherty Co., July (A)', November (N); Decatur Co.,
June-August (A) Floyd Co., August (A)3; Fulton Co., July, August (A)3; Gordon Co., July
(A)'; Grady Co., November (N); Jefferson Co., July (A)'; Lowndes Co., March (N); Lump-
kin Co., August (N); Macon Co., July (A); Newton Co., July (N); Pickens Co., June (N);
Rabun Co., June (A)3; Screven Co., July (A); Ware Co., April (N); White Co., July, August
(A). LOUISIANA, East Baton Rouge Par., April, May (A); St. Tammany Par., June (A);
Tangipahoa Par., August (N). MississiPPI, Clark Co., August (N); Forrest Co., August
(A); Lamar Co., June, August (A); Lauderdale Co., June, July (N,A); Marion Co., May (A).
NORTH CAROLINA, Transylvania Co., August (A); Wake Co., May (A). SOUTH CAROLINA,
Aiken Co., July, August (A)2; Florence Co., May (A)'; Lexington Co., April (N); Oconee
Co., August (N). TENNESSEE, Cheatham Co., July (A); Davidson Co., July (A); DeKalb
Co., July (A); Greene Co., July (A); Hamblen Co., August (A); Hamilton Co., August (A);
Lincoln Co., July (A); Robertson Co., June (N).


Vol. 22, No. 1







BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


Hexagenia munda marilandica Traver
Physiographic provinces: CP, P, BR, VR, AP, ILP
Previously published state records: Ala., Spieth, 1941; Fla., Spieth, 1941, Ber., 1950 and 1958,
PJ, 1973, Sch, 1967; Ga., NTH, 1935 (as 1H. carolina and H. marilandica), Spieth, 1941,
Ber., 1958; N. C., NTH, 1935 (as H. carolina and H. rosacea), Trav., 1937 and H, 1941 (as
H. carolina), Spieth, 1941; S. C., Trav., 1937 (as H. carolina), Spieth. 1941.
Additional records: ALABAMA, Elmore Co., June (N,A); Talladega Co., June (A); Tallapoosa
Co., June (N). FLORIDA, Liberty Co., March (N); GEORGIA, Bartow Co., July (A)'; Bibb
Co., July (A); Calhoun Co., June (A)'; Cherokee Co., June, August (A), June (N), July (A)';
Clayton Co., June (A)'; Cobb Co., July (A); Coweta Co.,June (A)'; Crawford Co., June
(A)'; Dawson Co., June (N,A); Floyd Co., (A); Fulton Co., June, July, September (A)';
Gordon Co., June (A)'; Gwinnet Co., July (A)'; Johnson Co., August (N); Lumpkin Co.,
August (A)'; Murray Co., July (A); Muscogee Co., June (A); Oconee Co., June (A):; Polk
Co., June (A)'; Rabun Co., July (N); Stephens Co., July (A)"'; Taylor Co., July, August
(A)'; Thomas Co., June (A)'; Troup Co., June (A)'; Upson Co., June (A)'; Walton Co., June
(A)'; White Co., July, August (A). NORTH CAROLINA, Granville Co., July (N); Jackson Co.,
July (N); Macon Co., June (N,A), July (A), August (N); Rockingham Co., July (A); Swain
Co., June, July (A); Transylvania Co., August (A); Wiles Co., June (N). SOUTH CAROLINA,
Aiken Co., June, July (A)"; Anderson Co., July (N); Greenville Co., June (N)', July (N);
Oconee Co., August (N); Spartanburg Co., June (A).

Hexagenia munda munda Eaton
Physiographic provinces: BR, VR, ILP
Previously published state records: N. C., Spieth, 1941.
Additional records: ALABAMA, Cleburne Co., July (A). TENNESSEE, Davidson Co., July (A).

Hexagenia munda orlando Traver
Physiographic provinces: CP
Previously published state records: Ala., Trav., 1937; Fla., NTH, 1935, Spieth, 1941, Ber.,
1950.
Additional records: FLORIDA, Bradford Co., May (A); Clay Co., July (N), June, August (A); High-
lands Co., March, April, July, October (A); Hillsborough Co., October (A), December
(N); Lake Co., April (A); Marion Co., September (A); Polk Co., March (N), April (A);
Putnam Co., June (A).

Hexagenia rigida McDunnough
Physiographic provinces: VR, ILP
Previously published state records: Tenn., McC., 1975
Additional records: TENNESSEE, Davidson Co., June, July (A); Hamblen Co., August (A);
Hamilton Co., August (A).

Hexagenia species
Physiographic provinces: CP, P, BR, VR, AP, ILP
Additional records: ALABAMA, Elmore Co., April (N). FLORIDA, Gadsden Co., April (N);
Liberty Co., April (N). GEORGIA, Bartow Co., June, July, August (A)'; Burke Co., May (N)2,
September (N); Calhoun Co., August (N); Colquitt Co., October (N); Chattahoochee Co.,
April (N); Cherokee Co., June, July (A)', August (N); Crawford Co., June (A)'; Cowetta
Co., June (N)'; Decatur Co., June (A), June, October (N), August (A)'; Dooley Co., Novem-
ber (N); Early Co., October (N); Floyd Co., June (A)3; Fulton Co., June, July (A)3; Gilmer
Co., June (A)'; Gordon Co., July (N)'; Gwinnett Co., June, July (A)'; Habersham Co., July
(A)'; Hall Co., July (A)'; Lumpkin Co., July, September (A)3; Miller Co., June (N)'; Monroe
Co., (N); Murray Co., August (N); Peach Co., April (N); Pickins Co., June (A)'; Rabun
Co., July (N); Richmond Co., June (A)'; Sumter Co., June (N)'; Troup Co., June (A)'; Union
Co., July (A)'; Walton Co., August (A)'; White Co., July (N,A)'; Whitfield Co., August
(A)'. NORTH CAROLINA, Davidson Co., July (N); Henderson Co., July (N); McDowell Co.,
July, August (N); Transylvania Co., July (N); Watauga Co., August (N). SOUTH CAROLINA,
Aiken Co., January, May (N)'; Allendale Co., May (N)'; Greenville Co., August (A)',
August (N); Kershaw Co., July (A)'; Pickens Co., July (N).







BULLETIN FLORIDA STATE MUSEUM


GENUS Litobrancha MCCAFFERTY
Litobrancha recurvata (Morgan)
Physiographic provinces: BR
Previously published state records: N. C., Spieth, 1941, Burks, 1953; Tenn., WB, 1949.
Additional records: NORTH CAROLINA, Avery Co., August (N); Macon Co., June, July (N);
Watauga Co., August (N). TENNESSEE, Greene Co., March (N).

GENUS Pentagenia WALSH
Pentagenia vittigera (Walsh)
Physiographic provinces: CP, BR
Previously published state records: Ala., Tray., 1937; Fla., Ber., 1950 and 1958, Sch, 1967;
Ga., McC., 1975; La., McC., 1975; Miss., McC., 1975; Tenn., NTH, 1935.
Additional records: ALABAMA, Dallas Co., May, June (A); Washington Co., May (A). FLORIDA,
Gadsden Co., June (A). GEORGIA, Baker Co., no date (A)'; Effingham Co., July (N).
LOUISIANA, East Baton Rouge Par., September (A). MISSISSIPPI, Lawrence Co., August
(A). TENNESSEE, Shelby Co., June (A).

FAMILY POLYMITARCIDAE
SUBFAMILY POLYMITARCINAE
GENUS Ephoron WILLIAMSON
Ephoron leukon Williamson
Physiographic provinces: CP, P, BR, AP, ILP
Previously published state records: Ga., McC., 1975; Miss., McC., 1975; N. C., McC., 1975;
S. C., McC., 1975; Tenn., NTH, 1935, WB, 1949.
Additional records: GEORGIA, Cherokee Co., June (N), August, July (A); Cobb Co., July (A).
MIssIssIPPI, Covington Co., August (N); Forrest Co., August (A); Itawamba Co., July
(N,A); Lauderdale Co., June, July (N); Lowndes Co., June (A); Monroe Co., July (A).
NORTH CAROLINA, Caldwell Co., July (N); Randolf Co., July (N). SOUTH CAROLINA, Aiken
Co., July (A)2, May (N)2; Chesterfield Co., July (A); Darlington Co., July (A). TENNESSEE,
Cheatham Co., July (A); Fentress Co., August (A); Giles Co., July (N); Greene Co., July-
August (A); Lincoln Co., July (A); Robertson Co., July (N,A), June (N); Rutherford Co.,
July (N); Sevier Co., August (A); Unicoi Co., July (A).
Ephoron species
Physiographic provinces: CP, AP
Additional records: ALABAMA, Perry Co., June (N); Walker Co., June (N).

SUBFAMILY CAMPSURINAE
GENUS TortOpus NEEDAM AND MURPHY
Tortopus incertus Traver
Physiographic provinces: CP
Previously published state records: Ala., NTH, 1935; Fla., Ber., 1950 and 1958; Ga., NTH,
1935, Ber., 1958, Ga.-S. C. state lines, SBH, 1959, PCR, 1967; La., Ber., 1959; Miss., Ber.,
1959; S. C., Ber., 1959.
Additional records: FLORIDA, Escambia Co., August (A). GEORGIA, Bibb Co., August (A); De-
catur Co., July, August (A)'; Doughtery Co., no date (A)3; Effingham Co., July (N,A);
Richmond Co., no date (A). MIssIssIPPI, Forrest Co., August (A); Itawamba Co., August
(A); Monroe Co., July (A); Pike Co., August (A); Smith Co., August (A). SOUTH CAROLINA,
Aiken Co., July, August (A)2; Allendale Co., July, August (N)2.


Vol. 22, No. 1







BERNER: MAYFLY DISTRIBUTIONAL PATTERNS


REFERENCES'

(A) Allen, R. K. 1967. New species of new world Leptohyphinae (Ephemeroptera: Tri-
corythidae). Canad. Entomol. 99(4):350-375, 64 figs.
(AE) Allen, R. K., and G. F. Edmunds, Jr. 1961. A revision of the genus Ephemerella (Ephe-
meroptera: Ephemerellidae) III. The subgenus Attenuatella. Jour. Kans. Entomol.
Soc. 34(4):161-173, 15 figs.
(AE) 1962. A revision of the genus Ephemerella (Ephemeroptera: Ephemerel-
lidae) IV. The subgenus Dannella. Jour. Kans. Entomol. Soc. 35(3):333-338, 13
figs.
(AE) 1962. A revision of the genus Ephemerella (Ephemeroptera, Ephemerel-
lidae) V. The subgenus Drunella in North America. Misc. Publ. Entomol. Soc.
Amer. 3(5): 147-179, 70 figs.
(AE) 1963. A revision of the genus Ephemerella (Ephemeroptera: Ephemerellidae)
VI. The subgenus Serratella in North America. Ann. Entomol. Soc. Amer. 56(5):
583-600, 68 figs.
(AE) 1963. A revision of the genus Ephemerella (Ephemeroptera: Ephemerel-
lidae) VII. The subgenus Eurylophella. Canad. Entomol. 95(6):597-623, 58 figs.
(AE) 1965. A revision of the genus Ephemerella (Ephemeroptera: Ephemerel-
lidae). VIII. The subgenus Ephemerella in North America. Misc. Publ. Entomol.
Soc. Amer. 4(6):244-282, 81 figs.
Banks, N. 1900. New genera and species of Nearctic neuropteroid insects. Trans. Amer.
Entomol. Soc. 26:239-259.
__ 1914. New neuropteroid insects, native and exotic. Proc. Acad. Natur. Sci.
Philad. 66:608-632, pls. I-IV.
___ 1924. Descriptions of new neuropteroid insects. Bull. Mus. Comp. Zool.
65(12):421-455, pls. I-IV.
(Ber) Berner, L. 1950. The mayflies of Florida. U. Fla. Press, Gainesville, Bio. Sci. Ser. xii +
267. Front., 88 figs., 24 pls., 19 maps.
(Ber) 1953. New mayfly records from Florida and a description of a new species.
Fla. Entomol. 36(4): 145-149, 3 figs.
(Ber) 1955. The southeastern species of Baetisca (Ephemeroptera: Baetiscidae).
Quart. Jour. Fla. Acad. Sci. 18(1):1-19, 23 figs.
(Ber) 1956. The genus Neoephemera in North America (Ephemeroptera: Neo-
ephemeridae). Ann. Entomol. Soc. Amer. 49(1):33-42, 26 figs.
(Ber) .1958. A list of mayflies from the lower Apalachicola River drainage. Quart.
Jour. Fla. Acad. Sci. 21(1):25-31, 1 fig.
(Ber) 1975. The mayfly family Leptophlebiidae in the southeastern United States.
Fla. Entomol. 58(3):137-156, 29 figs.
(BA) Berner, L., and R. K. Allen. 1961. Southeastern species of the mayfly subgenus Serratella
(Ephemerella: Ephemerellidae). Fla. Entomol. 44(4): 149-158, 10 figs.
Bick, G. H., L. E. Hornuff, and E. N. Lambremont. 1953. An ecological reconnaissance
of a naturally acid stream in southern Louisiana. Jour. Tenn. Acad. Sci. 28(3):221-
231, 2 figs.
Burks, B. D. 1953. The mayflies, or Ephemeroptera, of Illinois. Bull. III. Nat. Hist.
Surv., Art. 1, 26:1-216, 395 figs.
(Bur) Burmeister, H. C. C. 1839. Handbuch der Entomologie (Neuroptera). 2:757-1050.
Eaton, A. E. 1883-1888. A revisional monograph of recent Ephemeridae or mayflies.
Trans. Linn. Soc. London. Second Ser. Zool. 3:1-352, 65 p1s.
(EAP) Edmunds, G. F., R. K. Allen, and W. L. Peters. 1963. An annotated key to the nymphs
of the families and subfamilies of mayflies (Ephemeroptera). Univ. Utah Biol.
Series. 13(1):1-49, 78 figs.
(EBT) Edmunds, G. F., L. Berner, and J. R. Traver. 1958. North American mayflies of the
family Oligoneuriidae. Ann. Entomol. Soc. Amer. 51(5):375-382, 31 figs.

Abbreviations used under the heading "Previously published state records" are shown in parentheses to
the left of each reference.







BULLETIN FLORIDA STATE MUSEUM


(EJ) Edmunds, G. F., and S. L. Jensen. 1974. A new genus and subfamily of North American
Heptageniidae (Ephemeroptera). Proc. Entomol. Soc. Washington. 76(4):495-497.
Edmunds, G. F., Jr., S. L. Jensen, and L. Berner. 1976. The mayflies of North and Central
America. University of Minnesota Press, Minneapolis. In press.
(ET) Edmunds, G. F., and J. R. Traver. 1959. The classification of the Ephemeroptera. I.
Ephemeroidea: Behningiidae. Ann. Entomol. Soc. Amer. 52(1):43-51, 32 figs.
(H) Howell, T. 1941. Some aquatic insects from the Great Smoky Mountains National Park.
Jour. Tenn. Acad. Sci. 16(4):406-407.
(H) 1941. Notes on Ephemeroptera and aquatic Diptera of western North
Carolina. Jour. Elisha Mitchell Sci. Soc. 57(2):306-317.
(Kenn) Kennedy, C. H. 1926. The nymphs of Ephemera guttulata Pictet with notes on the
species. Canad. Entomol. 58:61-63, 1 fig.
(L) Lewis, P. A. 1974a. Three new Stenonema species from eastern North America (Hepta-
geniidae: Ephemeroptera). Proc. Entomol. Soc. Washington. 76(3):347-355, 19 figs.
(L) 1974b. Taxonomy and ecology of Stenonenma mayflies (Heptageniidae:
Ephemeroptera). U. S. Environ. Protec. Agency, Environmental Monitoring Series,
EPA-670/4-74-006. vii + 81 p., 203 figs.
(McC) McCafferty, W. P. 1975. The burrowing mayflies (Ephemeroptera: Ephemeroidea) of
the United States. Trans. Amer. Entomol. Soc. 101:447-504, 51 figs.
(McD) McDunnough, J. 1931. The genus Isonychia. Canad. Entomol. 63:157-163, pl. 8.
(McD) 1931. The eastern North American species of the genus Ephemerella and
their nymphs. Canad. Entomol. 63:187-197,201-216, pls. 11-14.
(M.-L.) Milller-Liebenau, I. 1974. Rheobaetis: a new genus from Georgia (Ephemeroptera:
Baetidae). Ann. Entomol. Soc. Amer. 67(4):555-567, 27 figs.
(Need) Needham, J. G. 1932. Three new American mayflies. Canad. Entomol. 64:273-276, 1 fig.
(NTH) Needham, J. G., J. R. Traver, and Yin-Chi Hsu. 1935. The biology of mayflies, with a
systematic account of North American species. Comstock Publ. Co., Ithaca. xvi +
759 p., 40 pls., 168 figs.
(PCR) Patrick, R., J. Cairns, and S. S. Roback. 1967. An ecosystematic study of the fauna and
flora of the Savannah River. Proc. Acad. Nat. Sci. Phila. 118:109-407.
(PP) Pescador, M. L., and W. L. Peters. 1971. The imago of Baetisca becki Schneider and
Berner (Baetiscidae: Ephemeroptera). Fla. Entomol. 54(4):329-334, 3 figs.
(PP) 1974. The life history and ecology of Baetisca rogersi Berner (Ephemerop-
tera: Baetiscidae). Bull. Fla. State Mus., Biol. Sci. 17(3):151-209, 25 figs.
(PJ) Peters, W. L., and J. Jones. 1973. Historical and biological aspects of the Blackwater
River in northwestern Florida. Proc. 1st Intern. Conf. on Ephemeroptera. E. J.
Brill, Leiden. pp. 241-253, pls. I-III.
(Sch) Schneider, R. F. 1962. Seasonal succession of certain invertebrates in a northwestern
Florida lake. Quart. Jour. Fla. Acad. Sci. 25(2):127-141, 5 figs.
(Sch) 1967. Mayfly nymphs from northwestern Florida. Quart. Jour. Fla. Acad.
Sci. 29(3):202-206.
(SB) Schneider, R. F., and L. Berner. 1963. A new southeastern species of Baetisca (Ephe-
meroptera: Baetiscidae). Fla. Entomol. 46(2):183-187, 6 figs.
(SBH) Scott, D. C., L. Berner, and A. Hirsch. 1959. The nymph of the mayfly genus Tortopus
(Ephemeroptera: Polymitarcidae). Ann. Entomol. Soc. Amer. 52(2):205-213, 14
figs.
Spieth, H. T. 1941. Taxonomic studies on the Ephemeroptera. II. The genus Hexagenia.
Amer. Midl. Natur. 26:233-280, 12 maps, 50 figs.
1947. Taxonomic studies on the Ephemeroptera. IV. The genus Stenonerna.
Ann. Entomol. Soc. Amer. 40(1):87-122, pls. 1-2, figs. 29-31.
(Trav) Traver, J. R. 1931. The ephemerid genus Baetisca. Jour. New York Entomol. Soc. 39:
45-66, pls. 5-6.
(Trav) __ 1932-33. Mayflies of North Carolina. Jour. Elisha Mitchell Sci. Soc. 47(1):
85-161, pls. 5-12. (2):163-236. 48(2):141-206, pl. 15.
(Trav) 1937. Notes on the mayflies of the southeastern states (Ephemeroptera).
Jour. Elisha Mitchell Sci. Soc. 53(1):27-86, pl. 6.


Vol. 22, No. 1








1977 BERNER: MAYFLY DISTRIBUTIONAL PATTERNS 55


Walker, Francis. 1853. List of the specimens of neuropterous insects in the collection
of the British Museum. Part III. (Termitidae-Ephemeridae). Brit. Mus. pp. 477-585.
(Wy) Wray, D. L. 1959. Insects of North Carolina. Second supplement. N. C. Dept. Agricul.
pp. 1-59, 1 fig.
(Wy) 1967. Insects of North Carolina. Third supplement. N. C. Dept. Agricul.
pp. 1-181, 1 fig.
(WB) Wright, M., and L. Berner. 1949. Notes on the mayflies of eastern Tennessee. Jour.
Tenn. Acad. Sci. 24(4):287-298.
(WR) Wurtz, C. B., and S. S. Roback. 1955. The invertebrate fauna of some Gulf Coast rivers.
Proc. Acad. Nat. Sci. Phila. 107:167-206.

















Contributions to the BULLETIN OF THE FLORIDA STATE MUSEUM, BIOLOGICAL SCI-
ENCES SERIES, may be in any field of biology. Manuscripts dealing with natural history of
systematic problems involving the southeastern United States or the New World tropics are
solicited especially. Manuscripts should be of medium length-circa 35 to 200 pages (10,500-
16,000 words). Examination for suitability is made by an Editorial Board.
The BULLETIN is distributed worldwide through institutional subscriptions and exchanges.
It is considered the responsibility of the author to distribute his paper to all interested individ-
uals. To aid in this the authors) receives) 50 copies free, and he may purchase additional
separates at cost if ordered when page proof is returned. The author is also responsible for any
charges incurred for alterations made by him on galley or page proofs. The Museum will send
an invoice to the author for this amount upon completion of publication.


PREPARATION OF MANUSCRIPT

Contributors should consult recent numbers of the BULLETIN for preferred style and format.
Highly recommended as a guide is the CBE Style Manual, 3rd Edition, 1972 (Washington, D.C.,
Amer. Inst. Biol. Sci.).
MSS must be submitted in duplicate (please no onionskin) and satisfy the following minimal re-
quirements: They should be typewritten, double-spaced (especially tables, figure captions, and
"Literature Cited"), on one side of numbered sheets of standard (8-1/2 x 11 in.) bond paper,
with at least one-inch margins all around. Figure legends and tables should be typed on separate
sheets. All illustrations are referred to as figures. They must comply with the following stan-
dards: Photographs should be sharp, with good contrast, and printed on glossy paper. Drawings
should be made with dense black waterproof ink on quality paper or illustration board. All il-
lustrations should have a cover sheet. All lettering will be medium weight, sans-serif type (e.g.
Futura Medium, News Gothic) in cutout, dry transfer, or lettering guide letters. Make allow-
ance so that after reduction no lowercase letter will be less than 1 mm high (2 mm is preferred)
nor any capital letter greater than 5 mm high. The maximum size for illustrations is 8-5/8 x 14 in.
(twice typepage size); illustrations should not be less than typepage width (4-5/16 in.). Designate
the top of each illustration and identify on the back with soft pencil by author's name, MS title,
and figure number.
All manuscripts not submitted in BULLETIN format will be returned to the author for retyping.



Manuscripts and all editorial matters should be addressed to:
Managing Editor of the BULLETIN
Florida State Museum
Museum Road
University of Florida
Gainesville, FL 32611







University of Florida Home Page
© 2004 - 2010 University of Florida George A. Smathers Libraries.
All rights reserved.

Acceptable Use, Copyright, and Disclaimer Statement
Last updated October 10, 2010 - - mvs