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EARTH SYSTEMS: THE FOUNDATION OF FLORIDA'S LO ECOSYSTEMS ED LANE, PG 141 FRANK RUPERT, PG 149 FLORIDA GEOLOGICAL SURVEY .......... _ 1996 INTRODUCTION ECOLOGY is the study of the relationship between organisms and their environment, including the study of communities, patterns of iFe, natural cycles, relationships to each other, biogeography, and population changes. An ECOSYSTEM is a community of organisms, including humans, interacting with one another and the environment in which they live. ECOSYSTEMS EXIST IN AN INFINITE VARIETY, and on many scales, from the microscopic to planet-wide. Therefore, an ecosystem's boundaries depend on the scope of the study. Most ecosystems have a d ,er-ityj of animals and plants, all of whicn have a range of living requirements. Because of this, there is overlap among adjacent ecosystems. ECOSYSTEMS ARE NOT STATIC. They change in time and space, due to both natural and human influences. Shadow lakes, for example, tend to fill in over time with sediment and plant material; human activities can accelerate the process. Plant and animal commu- nities also change as part of the life cycle of an ecosystem. A knowledge of EARTH SYSTEMS is necessary to understand how biological and non-biological components interact to create and sustain ecosystems. The solid earth aspects of earth systems are included in the science of geology, the atmospheric components reside with meteoroloqy, and hydrogeology and hydrologe cover the aqueous parts of our physical earth system. Basic geologic knowle ge of the physical characteristics o rocks and geologic processes has direct relevance to ecosystems. ROCKS ARE THE ULTIMATE SOURCE OF MOST NUTRIENTS IN rOOD CHAINS. Weathering processes break down rocks into their constituent minerals and chemical components, forming soils and nutrients. Erosion and sediment tran-pclt processes then make them available to the biosphere. Most nutrients are recycled many times through an ecosystem before losing their usefulness to the system. The GEOLOGIC STRUCTURE of the stratigraphic formations of an area's rocks is a major factor in determin- ing the landforms, or geomorphology, of that area. The structure and stratigraphy also influence surface and ground-water conditions, such as flow, recharge and discharge areas, and location and depth of aquifers and ambient water chemistry. Some geomorphic features can create very localized microclimates and restricted ecosys- tems; steephead ravines and sinkholes are two examples. The accompanying block diagrams are generalized composites that have been constructed to illustrate the most important topo- graphic and geologic features of broad areas of the state. They do not represent all features that occur in specific areas. Vegetation has been omitted to accentuate the main geomorphic features. Florida's climate ranges from temperate and sub-tropical in the north to tropical in the south. This climatic range results in intermingled plant and animal environments and numerous overlapping ecosystems. CENOZOIC ROCKS Florida is underlain by thousands-of-feet of rocks of Precambrian, Paleozoic, Mesozoic, and Cenozoic age. However, only rocks of Cenozoic age either crop out at the surface or occur at rela- tivel9y .h'[llow depths. Figures I and Z show these relationships. Since these are the only' rocks involved in creating and sustaining Florida's ecosystems, only the9 \..,II be discussed here. Cenozoic sediments began to be laid down about 65 million years ago. Figure I shows the geological units comprising the Cenozoic strat;grapk;c column in Florida. In general, carbonate rocks (limestone and dolomite) predominate in the older sediments of the Paleogene, while siliciclastics (clays, sands, gravels) predominate in the younger sediments of the Neogene-Quaternary. Florida's present landscape is the product of sea level fluctua- tions and associated marine related processes, erosion and deposi- tional processes. Over the past 2 to 3 million years, as a result of the repeated growth and melting of continental glaciers, sea levels have fluctuated from several hundred feet above present level to several hundred feet below present sea level. The most recent fluctuation is a slow rise in sea level, causing a retreat of shorelines in many parts of the state. Siliciclastic sediments were eroded from the Appalachian Mountains to the north, blanketing the state. Rivers, waves, and cur- rents created ancient deltas, shorelines, estuaries, dunes and sand ridges. Subsequent mechanical and chemical erosion and re-deposi- tion produced Florida's present geomorphology. The more prominent of the geomorphic features are shown on Figure 5. WESTERN PANHANDLE REPRESENTATIVE PLANTS The western Florida panhandle is under- lain by thick Miocene to Recent sands and clays. Carbonates, which are common at shal- and. and d unce low depth elsewhere in riLi;dnd, en, erWl[[ lie at depths in excess of +00 feet below land sur- face. Clags in the thick overburden sediments shield this limestone from extensive dissolution. As a result, karst features, such as sinkholes, are rare to absent in the four counties of the western panhandle. The thick sands and grav- elly sands also form the primary drinking water ,I / 1#. aquifer, which due to its lithology, is named the z) Ab Sand-and-Gravel aquifer. Extending approximately 50 miles from the Alabama border on the north to the Gulf shoreline, the regon is JidJ into two maor geomorphic zones. Skirting the northern edge of the western panhandle is a broad, stream- ',- 6 die.::i-ed upland named the Western SHighlands. South of these highl,knd- is a gen- eral9 flat and ele ..3tionall[ lower zone named Sthe Gulf Coastal Lowlands. r ,, The Western Highlands consist of clajeu- S sand nill., which attain elevations up to 5 5 feet above sea level (ASL). Surface water fea- tures consist primarily of deeplg-incised I/:. Q streams. Plio-Pleistocene clayey, gravelly, quartz sands of the Citronelle formation form Sthe core of the nill Nlanketing the Western -Highlands is a relict, variably thick veneer of marine terr.-ce sands, deposited by hih-stanJinr, Pliocene ; / ./ A irSd Fleistocene seas. Local ecosystems are controlled .,,, n raiL5 r1 by the t ,oporaphy and by the lthoelt j and fBffri r*dr-l,.:'l,[] i.w the undrl in4 geologic formations. Limonite kh.1-dp.in .nd cdI ay beds, for example, commonly retard down- v.virj p.i..iti.n in manu areas, causing high _t irJinr;v water 6SnJdg joi l derived from the 1 Itron&lln rh 'rni,-ton and the undif- ferentiated terrace sands, which have been leached of any contained 1:.11. po%,,f A ). ,w,,l-Jr.:ji d i c'. ordJ ti.mo p.n,-tic.u ki hill top and L -pc u,:h :,-.,Li tLpi.::il[r .si support uplind -.indhill or pin"e -ic'.-t e~o.fsltem t I-:.n.s' ,tn rirr -tlJ irr i o I .nle "-i F'ine nflu.L .lril. ,' oi'nce th-e FIGURE 1: Florida strattgraphic column. EASTERN PANHANDLE- BIG BEND The distance from the Georgia-Florida border in the north to the Gulf of Mexico in the south is about 50 miles. This landscape is natu- rally divided into two distinct areas by the east-west oriented Cody Scarp, an ancient shoreline feature which stretches from the western panhandle almost to Jacksonville. The CodJy Scarp is a prominent fea- ture that can rise as much as 150 feet in elevation over a mile of hori- zontal distance. The higher, rolling hills north of the Cody Scarp are in the Northern Highland geomorphic province, with elevations ranging from about 500 feet above sea level (ASL) near the Georgia/Florida border to about 75 feet ASL at thk toe of the scarp. The flat region that gradually slopes down from the toe of the scarp to sea level at the Gulf is the Gulf Coastal Lowlands. The Northern Highlands are underlain by sediments of the Miocene Hawthorn Group, composed of interbedded quartz sands, clays, and carbonates. Except for a few areas where they crop out at the surface, these sediments usually underlie the younger Pliocene siliciclastic sediments of the Miccosukee formation (reddish-orange clags, silts, and sands). These heterogeneous sediments form loamy soils that are rich in nutrients. They support lush, natural vegetation, and make good farming soils. The rolling hills have been dissected by stream erosion, creating steep-walled channels in the weakl9 clay- cemented, sandy sediments. Steepheads are distinctive erosional fea- tures that result from spring-fed streams eroding headward and etch- ing narrow, steer ravines. These ravines form isolated environments that have specialized ecosystems of ferns, other sensitive plants, and amphibians. Many large lake basins are the result of dissolution of buried lime- stone, which has caused slight subsidence of the overlging siliciclastic sediments. These lakes are only a few feet deep and usually have extensive marshes or wetlands associated with them. Due to the high clay content and the thickness of the Hawthorn Group and Miccosukee Formation sediments, which act as confining units, this region has low potential for recharge to the groundwater aquifer, the Floridan aquifer system, which is developed in the underlying lime- stones. The upland ecosystems are based on the plant communities that are supported by the nutrient-rich sediments, and are dependent on the local stratigraphy and hydrogeology. Important elements in these ecosystems are the-mixed hardwood and conifer forests, containing both temperate and subtropical species. Extensive stands of large, mature hardwoods (oak, maple, beech, hickory, magnolia), conifers (pine, cedar, cypress), and palms form the foundation and protective canopy for lush undergrowth. Together, these elements create rich and varied ecosystems. The Gulf Coastal Lowlands of the eastern panhandle-big Bend area are underlain by thick sections of near-surCace carbonate rocks of oligocene age (Suwannee Limestone) and Miocene age (St. Marks Formation). These limestones have been deeply weathered by disso- lution. These limestones also extend to the north under the Northern Highlands. Karstification is largely a chemical erosion process that occurs in carbonate rocks. Because most of Florida is underlain by carbonate GULP COA$rAL 0 50 MILES i. 0 80 KM SCALE IMMOVLee RiIA Dia CT!r5S SWAI SOUTHWESTERN FLORIDA Southwestern Florida is underlain by oligocene to Recent car- has resulted in a number of J, ffcr,-rl bonates, sands, clays, and mollusk shell beds. The deeper carbonates the higher sand ridges xeric ecosys are comprised of porous Suwannee Limestone. Locally, the Suwannee well-drained quartz sands of ancient Limestone is the uppermost rock unit of the rloridan aquifer stem. These floral communities include xer Overlying the Suwannee are carbonates and interbedded sands an and typically support various specie class of the Hawthorn Group. At the eastern edge of the area, clayey, metto.A chain of large karst lakes gravelly quartz sands of the Pliocene Cy.presshead Formation overie alley, provides a tLp.ii:il sandhill uF the Hawthorn Group and form the foundation for the elevationally aquatic flora suchas bladderwort, [ higher central ridges. shallow sediments are variably-thick undifferen- narrow u and around each lake. Dr tiated Plio-Pleistocene sands and shell beds. interspersed with patches of exF In general, the region is comprised of flat-lying sandy terrain bro- Intraridge Valley between lakes. In so ken only by the higher elevations of relict, marine-eroded inland ridges blanket portions of the floor of the running parallel to the axis of the peninsula. Locally, the most promi- The broad, flat Desoto PF'l.n is nent of these ridges is named the Lake Wales Ridge, which attains ele- Hawthorn Group. Covered only by vations of 500 feet above sea level (ASL). The core of the Lake Wales sand, the cle l1 Hawthorn ced ment Ridge is comprised of clayey, gravelly, coarse quartz sands of the water. Because of the lo,w-perme.Al Pliocene Cypresshead formation. These, in turn, rest on Oligocene to however, many areas of the plain bec Pliocene si]icilastics and carbonates of the Hawthorn Grouep. The Lake duringthe rainy season. Mesic flatwc Wales Ridge is split along much of its length by a steep-walIed axial val- th non-agricultural land on the ley, referred to as the Intraridge valley. This valley probably formed by Lowlands west of the Lake Wales Rid karst dissolution of the underlying limestone. Numerous large sinkhole th flat, p l condit lakes extend along the axis of the Intraridge Valley. meto, ow shrubs, and grasses pred Extending westward from the toe of the Lake Wales Ridge is a Patches of hardwood forest ril broad, flat, sand, relict submarine plain named the DeSoto Plain. It reaches of the Peace and Mqakka ranges in elevation from about 80 feet ASL at the toe of the Lake soils the Hawthorn are hallow c Wales Ridge to about 40 feet ASL at its western boundary, where it exist. Salt marshes are developed wh, adjoins the Gulf Coastal Lowlands zone. Surface sediments are com- water near the coast. These es" uand posed largely of marine sands and interbedded shell beds. black needlerush, cordgrass, and The Gulf Coastal Lowlands extend from the western boundary of 4 r -I,,, e.. .l,,,,- ,. t-, ,., L. H I kw-N pr;n,-ipal upland tree species in prehistoric times, alonr, \ith other F species, oaks, wiregrasses, herbs, and low woody shrubs. Interspersed with the sandhll[ ecosystem on the rolling. top. phY of the Western Highlands, particularl-i along the mailer "tre courses, are upland hardwood forests and upland mixed fore Clayey9, organic, sandy sois, developed from the 5hallow-lyirng A Bluff Group and Cjtror, elle r._rmartc'n, retain more moisture than deF, sand r revZin3 and support a decJedl! mesic (moist) commun In these regions the common flora includes m.3aclia, oak, hickc beech, and various pine species. Bottomland hardwood forests z lo.pld jir, forests are developed along the major alluvial stream cou es of the region, where higher water tables and high soil JaIl organic contn provide mesic conditions. 'oistLire depAenjenft ore of water and live oak, sweetgum, ktolie., beech, pJmett nd r birch extend along the upper reaches of the Escambia River Vail Freshwater tidal swamp occupies the lower, seaward portion of valley. The southern boundary of the Western Highlands is marked a reFict marine escarpment which forms a topographic break betw thke ele.it. ,[n1 i h1i[-ir uplands to the north and the lower, eenr iiLt Gulf Coastal Lowlands zone to the south. The Gulf Coastal Lowlands comprise a .indJ, gentlu-seaw sloping plain e>tnerdinr from the southern edge of the West Highlands south to the Gulf of Mexico. elevations rise from sea leave the Gulf coast to about 50 feet ASL at the toe of the West Hihl.rndc'. The Gulf Coastal Lowlands are characterized by number -_iFc.t .jrd beach ridges, dunes and sales formed b1j hig--stand Pleistocene seas. This sandy topography is underlain by gener, clean quartz sands, creating moderate to poorlyI-drained mesic cor tions. Pine flatwoods occupy much of the inland portions of the 1c lands. Near the coast, maritime hammock flora of live oak, cabbk palm, and redbay occur in discontinuous forests, rooted in the th quartz sands of old coastal dunes. well developed dunes along modern coast support a beach dune ecosystem including sea ce cordgrass, sand spur, and mtl.-rnInw ,lrq. Behind the dunes, espec I1 on the barrier islands, is a coastal grassland ecosystem. This F treeless ecosystem is developed on clean sands and t(pi,allyI st ports only grasses, prostrate vines and other maritime herbacec flora . |
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