Front Cover
 Table of Contents
 Part 1: Tertiary mollusca from...
 Title Page
 Part 2: Tertiary mollusca from...
 Part 1: Tertiary mollusca from...
 Part 3: Fossil corals of Porto...
 Part 4: The tertiary foraminifera...
 Back Matter

Title: Scientific survey of Porto Rico and the Virgin Islands
Full Citation
Permanent Link: http://ufdc.ufl.edu/UF00091487/00005
 Material Information
Title: Scientific survey of Porto Rico and the Virgin Islands
Alternate Title: Scientific survey of Puerto Rico and the Virgin Islands
Physical Description: 19 v. : ill. ; 24 cm.
Language: English
Creator: New York Academy of Sciences
Jay I. Kislak Reference Collection (Library of Congress)
Publisher: The Academy,
The Academy
Place of Publication: New York N.Y
Publication Date: 1920-1941
Frequency: completely irregular
Subject: Scientific expeditions -- Periodicals   ( lcsh )
Natural history -- Periodicals -- Puerto Rico   ( lcsh )
Natural history -- Periodicals -- Virgin Islands of the United States   ( lcsh )
Natuurlijke historie   ( gtt )
Geologie   ( gtt )
Expedities   ( gtt )
Genre: bibliography   ( marcgt )
Spatial Coverage: Puerto Rico
United States Virgin Islands
Summary: Includes bibliographies.
Ownership: Provenance: Gift of Jay I. Kislak Foundation.
Statement of Responsibility: New York Academy of Sciences.
Dates or Sequential Designation: Vol. 1, pt. 1-
Dates or Sequential Designation: Ceased with vol. XIX, pt. 1.
General Note: Latest issue consulted: Vol. 18, pt. 4 (1952).
General Note: Kislak Ref. Collection: Vol. 18, pt. 2 (1941)-pt. 4 (1952).
 Record Information
Bibliographic ID: UF00091487
Volume ID: VID00005
Source Institution: University of Florida
Holding Location: University of Florida
Rights Management: All rights reserved by the source institution and holding location.
Resource Identifier: oclc - 01760019
lccn - 2002209050


This item has the following downloads:

VID00005 ( PDF )

Table of Contents
    Front Cover
        Front Cover 1
        Front Cover 2
    Table of Contents
        Table of Contents
    Part 1: Tertiary mollusca from Porto Rico
        Page i
        Page ii
    Title Page
        Title Page 1
        Title Page 2
    Part 2: Tertiary mollusca from the Lares District, Porto Rico
        Page 79
        Page 79a
        Page 79b
        Page 80
        Page 81
        Page 82
        Page 83
        Page 84
        Page 85
        Page 86
        Page 87
        Page 88
        Page 89
        Page 90
        Page 91
        Page 92
        Page 93
        Page 94
        Page 95
        Page 96
        Page 97
        Page 98
        Page 99
        Page 100
        Page 101
        Page 102
        Page 103
        Page 104
        Page 105
        Page 106
        Page 107
        Page 108
        Page 109
        Page 110
        Page 111
        Page 112
        Page 113
        Page 114
        Page 115
        Page 116
        Page 117
        Page 118
        Page 119
        Page 120
        Page 121
        Page 122
        Page 123
        Page 124
        Page 125
        Page 126
        Page 127
        Page 128
        Page 129
        Page 130
        Page 131
        Page 132
        Page 133
        Page 134
        Page 135
        Page 136
        Page 137
        Page 138
        Page 139
        Page 140
        Page 141
        Page 142
        Page 143
        Page 144
        Page 145
        Page 146
        Page 147
        Page 148
        Page 149
        Page 150
        Page 151
        Page 152
        Page 153
        Page 154
        Page 155
        Page 156
        Page 157
        Page 158
        Page 159
        Page 160
        Page 161
        Page 162
        Page 163
        Page 164
        Page 164a
        Page 164b
        Page 164c
        Page 164d
        Page 164e
        Page 164f
        Page 164g
        Page 164h
        Page 164i
        Page 164j
        Page 164k
        Page 164l
        Page 164m
        Page 164n
        Page 164o
        Page 164p
        Page 164q
        Page 164r
        Page 164s
        Page 164t
        Page 164u
        Page 164v
        Page 164w
        Page 164x
        Page 164y
        Page 164z
        Page 164za
        Page 164zb
        Page 164zc
        Page 164zd
        Page 164ze
        Page 164zf
        Page 164zg
        Page 164zh
        Page 164zi
        Page 164zj
        Page 164zk
        Page 164zl
        Page 164zm
        Page 164zn
        Page 164zo
        Page 164zp
        Page 164zq
        Page 164zr
        Page 164s
        Page 164t
        Page 164zu
        Page 164zv
        Page 164zw
        Page 164x
        Page 164zy
    Part 1: Tertiary mollusca from Porto Rico
        Page 1
        Page 2
        Page 3
        Page 4
        Page 4a
        Page 4b
        Page 5
        Page 6
        Page 7
        Page 8
        Page 9
        Page 10
        Page 11
        Page 12
        Page 13
        Page 14
        Page 15
        Page 16
        Page 17
        Page 18
        Page 19
        Page 20
        Page 21
        Page 22
        Page 23
        Page 24
        Page 25
        Page 26
        Page 27
        Page 28
        Page 29
        Page 30
        Page 31
        Page 32
        Page 33
        Page 34
        Page 35
        Page 36
        Page 37
        Page 38
        Page 39
        Page 40
        Page 41
        Page 42
        Page 43
        Page 44
        Page 45
        Page 46
        Page 47
        Page 48
        Page 49
        Page 50
        Page 51
        Page 52
        Page 53
        Page 54
        Page 55
        Page 56
        Page 57
        Page 58
        Page 59
        Page 60
        Page 61
        Page 62
        Page 63
        Page 64
        Page 65
        Page 66
        Page 67
        Page 68
        Page 69
        Page 70
        Page 71
        Page 72
        Page 73
        Page 74
        Page 75
        Page 76
        Page 77
        Page 78
        Page 78a
        Page 78b
        Page 78c
        Page 78d
        Page 78e
        Page 78f
        Page 78g
        Page 78h
        Page 78i
        Page 78j
        Page 78k
        Page 78l
        Page 78m
        Page 78n
        Page 78o
        Page 78p
        Page 78q
        Page 78r
        Page 78s
        Page 78t
        Page 78u
        Page 78v
        Page 78w
        Page 78x
        Page 78y
        Page 78z
        Page 78za
        Page 78zb
    Part 3: Fossil corals of Porto Rico, with descriptions also of a few recent species
        Page 165
        Page 166
        Page 167
        Page 168
        Page 169
        Page 170
        Page 171
        Page 172
        Page 173
        Page 174
        Page 175
        Page 176
        Page 177
        Page 178
        Page 179
        Page 180
        Page 181
        Page 182
        Page 183
        Page 184
        Page 185
        Page 186
        Page 187
        Page 188
        Page 189
        Page 190
        Page 191
        Page 192
        Page 193
        Page 194
        Page 195
        Page 196
        Page 197
        Page 198
        Page 199
        Page 200
        Page 201
        Page 202
        Page 203
        Page 204
        Page 205
        Page 206
        Page 207
        Page 208
        Page 209
        Page 210
        Page 211
        Page 212
        Page 213
        Page 214
        Page 215
        Page 216
        Page 217
        Page 218
        Page 219
        Page 220
        Page 221
        Page 222
        Page 223
        Page 224
        Page 225
        Page 226
        Page 227
        Page 228
        Page 229
        Page 230
        Page 231
        Page 232
        Page 233
        Page 234
        Page 235
        Page 236
        Page 236a
        Page 237
        Page 238
        Page 238a
        Page 239
        Page 240
        Page 240a
        Page 241
        Page 242
        Page 242a
        Page 243
        Page 244
        Page 244a
        Page 245
        Page 246
        Page 246a
        Page 247
        Page 248
        Page 248a
        Page 249
        Page 250
        Page 250a
        Page 251
        Page 252
        Page 252a
        Page 253
        Page 254
        Page 254a
        Page 255
        Page 256
        Page 256a
        Page 257
        Page 258
        Page 258a
        Page 259
        Page 260
        Page 260a
        Page 261
        Page 262
        Page 262a
        Page 263
        Page 264
        Page 264a
        Page 265
        Page 266
        Page 266a
        Page 267
        Page 268
        Page 268a
        Page 269
        Page 270
        Page 270a
        Page 271
        Page 272
        Page 272a
        Page 273
        Page 273a
        Page 273b
    Part 4: The tertiary foraminifera of Porto Rico
        Page 274
        Page 274a
        Page 275
        Page 276
        Page 277
        Page 278
        Page 279
        Page 280
        Page 281
        Page 282
        Page 283
        Page 284
        Page 284a
        Page 284b
        Page 285
        Page 286
        Page 287
        Page 288
        Page 289
        Page 290
        Page 291
        Page 292
        Page 293
        Page 294
        Page 295
        Page 296
        Page 297
        Page 298
        Page 299
        Page 300
        Page 301
        Page 302
        Page 303
        Page 304
        Page 305
        Page 306
        Page 307
        Page 308
        Page 309
        Page 310
        Page 311
        Page 312
        Page 313
        Page 314
        Page 315
        Page 316
        Page 317
        Page 318
        Page 319
        Page 320
        Page 321
        Page 322
        Page 323
        Page 324
        Page 325
        Page 326
        Page 327
        Page 328
        Page 329
        Page 330
        Page 331
        Page 332
        Page 333
        Page 334
        Page 335
        Page 336
        Page 337
        Page 338
        Page 339
        Page 340
        Page 341
        Page 342
        Page 343
        Page 344
        Page 345
        Page 346
        Page 347
        Page 348
        Page 349
        Page 350
        Page 351
        Page 352
        Page 353
        Page 354
        Page 355
        Page 356
        Page 357
        Page 358
        Page 359
        Page 360
        Page 361
        Page 362
        Page 363
        Page 364
        Page 365
        Page 366
        Page 367
        Page 368
        Page 369
        Page 370
        Page 371
        Page 372
        Page 373
        Page 374
        Page 375
        Page 376
        Page 377
        Page 378
        Page 379
        Page 380
        Page 381
        Page 382
        Page 383
        Page 384
        Page 385
        Page 386
        Page 387
        Page 388
        Page 389
        Page 390
        Page 391
        Page 392
        Page 393
        Page 394
        Page 395
        Page 396
        Page 397
        Page 398
        Page 399
        Page 400
        Page 401
        Page 402
        Page 403
        Page 404
        Page 405
        Page 406
        Page 407
        Page 408
        Page 409
        Page 410
        Page 411
        Page 412
        Page 413
        Page 414
        Page 415
        Page 416
        Page 417
        Page 418
        Page 419
        Page 420
        Page 421
        Page 422
        Page 423
        Page 424
        Page 425
        Page 426
        Page 427
        Page 428
        Page 429
        Page 430
        Page 431
        Page 432
        Page 433
        Page 434
        Page 435
        Page 436
        Page 437
        Page 438
        Page 439
        Page 440
        Page 441
        Page 442
        Page 443
        Page 444
        Page 445
        Page 446
        Page 447
        Page 448
        Page 449
        Page 450
        Page 450a
        Page 450b
        Page 451
        Page 452
        Page 452a
        Page 452b
        Page 453
        Page 454
        Page 454a
        Page 454b
        Page 455
        Page 456
        Page 456a
        Page 456b
        Page 457
        Page 458
        Page 458a
        Page 458b
        Page 459
        Page 460
        Page 460a
        Page 460b
        Page 461
        Page 462
        Page 462a
        Page 462b
        Page 463
        Page 464
        Page 464a
        Page 464b
        Page 465
        Page 466
        Page 466a
        Page 466b
        Page 467
        Page 468
        Page 468a
        Page 468b
        Page 469
        Page 470
        Page 470a
        Page 470b
        Page 471
        Page 472
        Page 472a
        Page 472b
        Page 473
        Page 474
        Page 474a
        Page 474b
        Page 475
        Page 476
        Page 476a
        Page 476b
        Page 477
        Page 478
        Page 478a
        Page 478b
        Page 479
        Page 480
        Page 480a
        Page 480b
        Page 481
        Page 482
        Page 482a
        Page 482b
        Page 483
        Page 484
        Page 484a
        Page 484b
        Page 485
        Page 486
        Page 487
        Page 488
        Page 489
        Page 490
        Page 491
        Page 492
    Back Matter
        Page 493
        Page 494
Full Text








Title-page ............................................................ i
C on ten ts .. .. ........................................................ iii
Dates of Publication of Parts and Editors................................ iii
Part 1. Tertiary Mollusca from Porto Rico. By C. J. MAURY. ............. 1
Part 2. Tertiary Mollusca from the Lares District, Porto Rico. By BELA
H UBBARD ............ .. .... .. .... ................................ 79
Part 3. Fossil Corals of Porto Rico, with Descriptions also of a Few Recent
Species. By H. N. CORYELL and VIOLET OHLSEN .................... 167
Part 4. The Tertiary Foraminifera of Porto Rico. By J. J. GALLOWAY and
CAROLINE E. H EMINWAY ........................................... 275

Part 4, April 21, 1941 ERICH MIAREN SCHLAIKJER



Porto Rico and the Virgin Islands

Tertiary Mollusca from Porto Rico-C. J. Maury





Porto Rico and the Virgin Islands


Tertiary Mollusca from Porto Rico-C. J. Maury

^ F7


5 D-
r.- rn

= m



Porto Rico and the Virgin Islands

VOLUME Ill-Part 2

Tertiary Mollusca from the Lares District,
Porto Rico-Bela Hubbard


The material here described was collected in making a study of
the geology of the Lares District in the summer of 1916, under the
auspices of the New York Academy of Sciences and the Insular
Government of Porto Rico. The present paper is a part of the
report on the geology of the Lares District, the completion of which
has been delayed two years by the war. As the section dealing with
the general geology and stratigraphy has not yet been completed,
the present report, dealing with the Tertiary Palaeontology, is pub-
lished in advance. It will be noted under the descriptions of the
species, that full use has been made of the material collected by Dr.
C. P. Berkey in 1914, by Dr. D). R. Semmes in 1915, and by Dr.
A. K. Lobeck in 1916, and in some cases their specimens have been
used for illustration.
No attempt will be made here to describe the stratigraphy, but the
following tabulation of the subdivisions* of the Younger Series of
the north coast is given in explanation of the horizons referred to
under the descriptions of the species.

San Juan formation

Arecibo Group

(Arecibo formation
of Berkey, Pepino
formation of Hill
and Vaughan)

(Berkey, 1915)
Pleistocene to Recent.
Quebradillas limestone (Berkey, 1915)
Upper Oligocene (Bowden)
Los Puertos limestone (Hubbard, 1917)
Upper Oligocene
Cibao Limestone (Hubbard, 1917)
AMiddle Oligocene
Lares formation (Hubbard, 1917)
Middle Oligocene
San Sebastian shale (Berkey, 1915)
Middle Oligocene

*These subdivisions were made and described by the writer in a short paper presented
before the New York Academy of Sciences, Section of Geology and Mineralogy, in February,



Acknowledgments are made to Senior Narciso Rabell, of San
Sebastian for aid in collecting fossils and for the interest which he
took in the field work; to C. J. Maury for the opportunity of com-
paring some of the material with hers; to lMiss A. L. IHcpburn,
Columbia University Library, for valuable assistance in securing
the many necessary but scattered palaeontological references; and to
Dr. A. W. Grabau for advice and assistance in describing many of
the species.
Field Numbers.
1. Lares road, north side, near K. 38 San Sebastian shale.
7. On the cart road 1.75 miles north-east of Lares. Lares limestone.
8. About 0.5 mile north of No. 7. Lares limestone.
10. About 1 mile north-east of No. S. Lares limestone.
23. About 5.5 miles north of Lares, and about 2 miles north of Central
Soller. Echinoid zone of Cibao limestone.
24. About G.5 miles north of Lares, on cart road to Camuy. at southern
boundary of the belt of pepino hills. Base of Los Puertos limestone.
41. On the cart road to Camuy, 0.25 mile south of the village of Cienega,
near the Central Alianza. Quebradillas limestone.
42. On the cart road to Camuy, 0.25 mile north of the Central Alianza.
Quebradillas limestone.
52(a). On the cart road to Collazo 1.75 miles south of Quebradillas, and
a short distance north-west of the village of San Antonio.
Quebradillas limestone.
59. Valley of the Guajataca River, 1 mile south of Planas.
Los Puertos limestone.
61. On the upland surface near the valley, and 0.3 mile south of No.
59. Los Puertos limestone.
62. About 0.6 mile south of No. 61, on the lowland just south of the belt
of pepino hills. Cibao limestone.
64. About 0.3 mile south of No. 62, on cart road to Collazo.
Cibao limestone.
65. On the cart road 2 miles north-east of Collazo. Lares limestone.
72. Lares road at K. 28.G. Sai Sebastian shale.
74. Lares road at K. 29.3. San Sebastian shale.
75. Lares road at K. 30.4. San Sebastian shale.
76. Lares road at K. 32.9. San Sebastian shale.
83. Collazo Falls section, at top of second falls below the bridge.
San Sebastian shale.


84. Collazo Falls section, at base of second falls below the bridge.
San Sebastian shale.
86. Collazo Falls section, at top of fourth falls below the bridge.
San Sebastian shale.
89. Collazo Falls section, below the sixth falls below the bridge, in the
lowest horizon exposed in the section. San Sebastian shale.
92. Lares road at K. 28.8, in ledge 12 feet above the road.
San Sebastian shale.
100. Talus at base of the cuesta north of Lares. Lares limestone.
118. On the upland surface 0.25 mile north-west of Lares plaza.
San Sebastian shale.
120. About 180 yards north-west of No. 118. San Sebastian shale.
126. Lares road, near K. 37. San Sebastian shale.
140. Base of the cuesta on west side of the Camuy River valley at the fork
of the cart road to Utuado and the new road to Arecibo
San Sebastian shale.
142. Cuesta at No. 140, in the limestone cliff overlying the basal shale.
Lares limestone.
158. On cart road to Camuy, about 500 yards north-west of K. 19.5 of the
new Arecibo-lares road. Cibao limestone.
161. About 500 yards north-west of No. 158, on the same cart road.
Cibao limestone.
162. About 600 yards north-west of No. 161, on the same cart road.
Echinoid zone, Cibao limestone
165. Near the point of emergence of the subterranean Camuy River.
Cibao limestone.
167. A short distance north of No. 165, on the west side of the Camuy
River. Cibao limestone.
174. ALout 0.5 mile north of No. 167, west side of the Camuy River on cart
road to Camuy. Los Puertos limestone.
176. About 0.75 mile north of No. 174, on the same road.
Los Puertos limestone.
179. About 0.4 mile north of No. 176, on the same road.
Los Puertos limestone.
182. About 0.6 mile northwest of No. 179, and about 0.5 mile south of the
ford where the cart road crosses to the east side of the Camuy River.
Los Puertos limestone.
191. On the automobile road between Camuy and Quebradillas, near K. 40.5.
Quebradillas limestone.
192. On the same automobile road, near K. 40.
Quebradillas limestone.
198. On the automobile road west of Quebradillas, near K. 28.7.
Quebradillas limestone.
204. Sea cliffs at the east mouth of the American Railroad tunnel at the
mouth of the Guajataca River, north coast. Quebradillas limestone.


2115. In the railroad cut 80 yards west of the railroad bridge over the
Guajataca River, and 450 yards east from the automobile road grade
crossing. Quebradillas limestone
2U9. Gunajataca River valley, west of the village of San Antonio.
Quebradillas limestone.
216. About 1.75 miles south of K. 14.7 (Isabela-Aguadilla automobile road).
on cart road near Arenales bajos. Quebradillas limestone.
217. Nearly 0.5 mile south of No. 216. Quebradillas limestone.
220. Near Arenales altos, about 2.5 miles south of No. 217, and 150 yards
east of the cart road. Los Puertos limestone.
225. On the cart road near Robles, 25 feet below the summit of the cuesta.
Top of the Lares limestone
226. Zone about 90 feet below No. 225. Lares limestone.
227. A 40 foot exposure of marl on the cart road south of No. 221, and 251
feet below the summit of the cuesta. Lares limestone.
229. About 1 mile norti of K. 17.5 (Lares road), on cart road near Hato
arriba. Base of Lares limestone.
2'0. A short distance south of No. 229, and about 50 feet lower in elevation.
Top of San Sebastian shale.
231. About 100 yards south of No. 230, at a slightly lower elevation.
Coral zone of upper San Sebastian shale.
232. About 200 yards south of No. 231, and 1,350 yards north of K. 17.-5
(Lares road), at an approximate elevation of 180 feet above the road.
A five foot exposure of marl marking the Clemeitia darieoa zone.
San Sebastian shale.
232(b). A fifteen foot exposure immediately overlying No. 232, and remarkable
for the abundance of Turritella tornata. San Sebastian shale.
233. On a small secondary cuesta a short distance south of No. 232.
San Sebastian shale.
234. South of No. 2332 on the same cuesta. San Sebastian shale.
235. South of No. 234, and 1,020 yards north of K. 17.5 (Lares road) at an
elevation of 180 feet above the road. San Sebastian shale.
236. South of No. 235, and 900 yards north of K. 17.5. San Sebastian shale.
237. South of No. 236, and 820 yards north of K. 17.5. at an elevation of 100
feet above the road. San Sebastian shale.
241. On the cart road to Camuy, 1 mile north-east of Lares, in limestone
quarry near summit of the cuesta. Lures limestone.
254-5-6. Talus from the cnesta north of Lares. Taken from the road alone
the Guajataca River, about 0.75 mile iiorth of Lares plaza.
Lares limestone.
261. Collazo Falls section, at base of second falls Ielow the bridge, and
overlying No. 84. Sun Sebastian shale.
262. Collazo Falls section. at top or third falls below the bridge.
San Sebastian shale.


263. Collazo Falls section, half way between top and bottom of the fourth
falls below the bridge. San Sebastian shale.
264. Collazo Falls section, immediately underlying No. 263.
San Sebastian shale.
265. Collazo Falls section, at base of the fourth falls below the bridge.
San Sebastian shale.
267. Collazo Falls section, near the base of the section, overlying No. 89.
San Sebastian shale.
268. Collazo Falls section, in the red sandstone at the sixth falls below the
bridge, and overlying No. 207. San Sebastian shale.
280. About 880 yards north of Moca on cart road.
Lares formation, shale faces.
281. About 1,100 yards north of Moca on cart road.
Lares formation, shale faces.
2S2. About 1,150 yards north of Moca on cart road.
Lares formation, shale faces.
287. About 1.25 miles north on Moca on cart road.
Top of the Cibao limestone.
313. Sea cliff 0.75 mile northeast of the Point Borinquen Light House, from
a stratum 55 feet above sea level. Quebradillas limestone.
314. The same cliff, in a stratum 10 feet above sea level.
Quebradillas limestone.
317. About 1.25 miles north of K. 2 (Aquadilla-Isabela automobile road), on
cart road, 350 yards north of the American Railroad grade crossing.
Quebradillas limestone.
318. A short distance south of No. 317, in railroad cut, about 2110 yards north-
east of grade crossing. Quebradillas limestone.
319. In quarry east of cart road, and about 0.6 mile north of K. 2 (Aguadilla-
Isabela road). Quebradillas limestone.
324. Small limestone outlier about 300 yards south of K. 1.7 (Lares road).
Lares limestone.
330. Southwest of bridge over Culebrines River near K. 2 (Aguadilla-Aguada
automobile road). Lares limestone.
331. Aguadilla-Aguada road, south of sugar railroad crossing near K. 2.
Lares limestone.
332. Same road, about 300 yards southwest of No. 331. Lares limestone.
333. On automobile road to Rincon, 1,150 yards west of Aguada. Exposure
on north side of road. Lares limestone.
334. Same road, 100 yards west of No. 333. Lares limestone.
335. Same road, 290 yards west oft No. 334. Lares limestone.
343. Same road, 640 yards northeast of bridge over the Rio Grande.
Lares limestone.
344. Same road, south of No. 343, and 460 yards north of bridge over the
Rio Grande. Lares limestone.


370. Exposure in sea cliff on south side of the American Railroad track at
Point Jiguero, 930 yards northeast of the lighthouse. The San Juan
formation may be seen here, plastered on the Tertiary limestone.
Lares limestone.
616. Uppermost stratum in quarry 0.5 mile south-east of Auguada, on south
side of road, and northeast of the American Railroad grade crossing.
Lares limestone.
619. Same as No. 616, but from lowest exposure in the quarry.
Lares limestone.
621. Exposure on south side of railroad tracks, about 580 yards south-east of
the Aguada railroad station. Lares limestone.
664. East side of Camuy River valley, 2.25 miles southeast of Camuy.
Quebradillas limestone.
670. East side of Camuy River valley, I mile southeast of No. 664.
Quebradillas limestone.
671. Same locality as No. 670, but from a stratum a few feet above the latter.
Quebradillas limestone.
678. East side of Camuy River valley, about 1.5 miles due south of No. 671,
and about 0.75 mile north of No. 182. Quebradillas limestone.
749(a). On cart road, about 0.5 mile north of bridge at Collazo.
Lares limestone.
757. One mile northeast of Lares, on the cuesta, in quarry near No. 244.
Lares limestone.
758. Same locality asNo. 757, but at a slightly higher horizon.
Lares limestone.
800. About 100 yards south-east of No. 24. Top of Cibao limestone.
801. In railroad cut at east end of railroad bridge over the Guajataca River,
north coast. This locality is about 50 yards east of No. 205.
Quebradillas limestone.

The following localities are recorded from field notes of I)r. C.
P. Bcrkey.
10. North of the military road in the vicinity of Carolina.
Quebradillas limestone.
77. Military road 1.5 kilometers west of Ponce. Upper Ponce formation.
(Corresponds with the Quebradillas limestone.)
78. Three kilometers west of Ponce. Upper Pouce formation.
95. Near K. 28 Lares road. San Sebastian shale.
96. A short distance west of Lares on the Lares road.
San Sebastian shale.
98. In the railroad cut near the bridge over the Guajataca River. north
coast. Quebradillas limestone.
176. Vicinity of Moca. Lares formation, shale faces.



Genus Atrina Gray

Atrina rabelli, new species

Plate X, Figure 1
Margins of shell are arenate strongly folded upward, particularly the
anterior and posterior portions; central convex zone with flat lateral slopes,
the cross-section of the valve being broadly V-shaped; surface marked by
small, regular concentric ripples which become obsolete on the lateral mar-
gins; central convex zone n'arked by two low radial ribs which die out near
the ventral margin. Length 71 mm., semi-diameter 15 mm., height 100 mm.
A single external mold was found showing only the ventral half
of the shell. It is a large, rapidly widening form of the type of
A. chipolana Dall, but is distinct from any form yet described. This
shell is named in honor of Sefior Rabell, geologist, of San Sebastian,
who has taken an active interest in the geological work of the X. Y.
Academy of Sciences and rendered every possible assistance to the
geologists in their field work.
Horizon.-Quebradillas limestone.

Genus Pecten Miller

Pecten (Pecten) laresense, new species

Plate X, Figures 2, 3
Shell small of the concavo-convex type; right valve extremely convex, left
valve shallow concave: shell equilateral: ears small and strongly ribbed
radially; right valve with twenty-seven square smooth ribs and nearly flat
interspaces of equal width, marked by fine, sharp, raised concentric threads
which rise on the sides of the ribs; left valve with similar ribs and inter-
spaces but with ribs medially grooved distally; ears of left valve with imbri-
cate radial threads; anterior ear of left valve concave and correspondingly
convex in the right; byssal notch small, ctenolium apparently lacking; sub-
margins on both valves absent, the ribs extending to the base of ear; interior
grooved and lirate ventrally; cardinal crura obscure and irregular; small
amorphous pair of auricular crura; provinculum distinct; adductor scars
markedly posterior. A right valve measures in length 20 mm., height 17 mm.,
diameter 8.5 mm.; left valve measures in length 20 mm., height 17 mm.,
concave diameter 2 mm.
This shell is distinct from any other Peeten in the Porto Rican


Type localities.-231, 232, 233.
Other localilies.-232 (b), 126>, 11b, 23G, 1.
Horizon.-San Sebastian shale. (abundant and widely dis-

Pecten (Euvola) reliquus Brown and Pilshry
variety portoricoensis, new variety
Plate X. Figure 4

Only right valves were fonid, which agree closely with the speci-
ineus of P. reliqius in the Columbia University Galunn collections.
The Porto Rican shell differs in having fewer, higher ribs with
narrow interspaces, and in living the lateral slopes faintly ribbed
instead of smooth as in the Gatun shell. The most perfect right
valve has fourteen strong rounded ribs with several faint ones on
each lateral slope. Length 3.5 mi., height 34 mmn., diameter 3 mm.
Fragments were collected showing a size of at least 50 m1m.
The left valves seem to have been easily destroyed, as none were
found in the Panama collections.
Type locality.-16,.
Other localilies.-161. 19s.
Range.-Cibao limestone to Quebradillas limeslone.

Pecten (Nodipecten) nodosus Linn6
Plate X, Figure 5
Pcclen nodosuis Linnc, Syst. Nat., No. 164, 1758.
Pecten.nmagnificus Gabb, Trans. Amer. Phil. Soc.. XV, p. 256, 1873 tnot of
Pecten (Nodipecten) nodosus Dall, Trans. Wagner inst. Sci., III, pt. 4, p. 717,
Pecten (Nodipecten) nodosus Maury. Bull. Amer. Paleont., V, No. 29, p. 1S;,
The specimens show three prominent strongly nodose ribs, one in
center of valve and two lateral. These aterniate with two lower, sub-
nodose ribs marked- l- a median and two lateral threads. The
suihnodose ribs break up into scaly threads adjacent lo tlie sil-
inargins. The imos perfect slpcimen mlleasurs in length 2!' 11mi..


height 2 iiii.. seini-diaimter 7 nun., but fragnlnils were found
showing a height of 40 1n1.
Type locality.-19s.
Other locality.-42.
Horizon.-Queblrdillas limestn ie (not abundant).

Pecten (Chlamys) collazoensis, new species

Plate XI, Figure 1
Shell small, slightly convex; surface with twenty-one to twenty-two
rounded ribs and narrower rounded interspaces; ribs ornamented with fine,
thin, erect scales extending the whole width of the rib, and on either side at
juncture with the interspace, a row of minute, densely crowded, bead-like
spinules, which are much smaller and more densely crowded than the scales
on tops of the ribs. Snbmargins and ears sculptured with a continuous series
of squamose radial threads; anterior ear (of right valve) longer than the
posterior ear, and with deep byssal notch. Length 11.5 mm., height 12.5 mm..
semi-diameter 2 mm.

No well pre-erved left valves were found. This shell is apparently
the progenitor of P. portoricoensis.
Localities.-Collazo Falls, 232, 2:;3 .
llorizoo.-Lower San Sebastian shale. (some doubtful speciimens
were found in the upper part of the San Sebastian shale.)

Pecten (Chlamys) portoricoensis, new species

Plate X f, Figures 2, 3
The typical form is a moderate sized shell (length 24.5 mm., height 26.5
mm., diameter 9 mm), equivalve, sub-equilateral; ears moderately large, un-
equal (anterior larger); byssal notch large but not deep; ctenolium incon-
spicuous; single pair of cardinal crura; valves grooved within, and scalloped
and lirate at ventral margin: surface with twenty-six to twenty-seven rounded
ribs and slightly narrower, flatly rounded interspaces; ribs ornamented with
one to three rows of fine spinose scales which in the young shells show a
tendency toward the beaded ornamentation of P. collazoensis. This feature
usually persists longer next the submargins than centrally. Following this
P. collazoensis stage, the ribs and interspaces become smoother, the latter
relatively broader; a medium thread or keel appears on each rib, and on
this keel the small scales become narrower, more spinose and more widely
spaced. When the shall is half grown, lateral keels, similarly spinose, appear
low down on the sides of the ribs. These correspond in position with the
lateral heads of the P. collazoensis stage but there is always a gap between


the two where the sides of the ribs are smooth. Finally, in the adult, an
interstitial thread, ornamented like the keels, appears in each interspace. In
the last stage of some adult specimens an additional pair of spinose lateral
keels appears one on each side of the median keel. On the lateral slopes next
the submargins the ribs break up into densely squamose threads, which,
decreasing in size, continue in unbroken series across the submargins and
the ears. The right anterior ear differs from the others in having a strongly
rugose area adjacent to the submargin, and above this the radial threads are
strong and coarsely squamose. Camplonectes striation is very plain on un-
worn specimens.
This is an extremely variable shell, apparently related to P. thie-
tides Sowerby. It is the most abundant Pecten in the Tertiary of
north side of the island and numerous specimens show intergrada-
tions between the different varieties, thus fortunately preventing the
error of making two or more species. The typical forms are es-
pecially interesting in that they show thle species to have been de-
rived from P. (Chlamys) collazoensis, which was not found in hori-
zons above the San Sebastian shale. P. portoricoensis ranges from
the San Sebastian shale through the Lares limestone, and has been
found by Dr. Berkey in the Ponce limestone on the south side of the
This shell is obviously derived from P. rollazoensis, and both might
conceivably be included under one specific name. The latter, how-
ever, retains its typical ornamentation throughout, without the
slightest tendency toward a change, whereas in P. portoricoensis the
P. collazoensis stage has become so accelerated that it is unnoticeable
in some specimens and absolutely lacking in variety reticulatis.
Type localities.-234, 235, 343.
Other localities.-232, 118, 261, 1, 332, 70, and Collazo Falls,
Stratigraphic range.-Throughout the San Sebastian shale, and
in the lower Lares limestone also found by Dr. Berkey in the Juana
Diaz shale.

Pecten (Chlamys) portoricoensis, new species

variety reticulatis

Plate XI, Figure 4
Shell, while smaller than the type, is similar in every respect, except that
In the young the ribs are crossed by line, sharp, densely but evenly spaced


concentric lamellae, which thicken slightly on crossing the ribs, the whole
producing a reticulate ornamentation. This feature is most noticeable cen-
trally, since adjacent to the submargins are the typical scales, which do not
appear until later on the ribs of the central portion of the shell. The reticu-
late ornamentation persists until the shell has reached one-quarter or one-
third adult size, and is then replaced by the typical spinose scale. The
lateral spinose keels and the interstitial threads do not appear until the shell
has reached maturity. Average size specimen measure in length 16.5 mm.,
height 18 mm., diameter 6 mm. The largest specimen measures 24 mm.
in height.
Type localities.-343, 619.
Other locality.-74.
Range -San Sebastian shale and Lares formation.

Pecten (Chlamys) portoricoensis, new species

variety grandis

Plate XI, Figures 5, 6

This shell differs from the type and from variety reticulatis
chiefly in its much greater size, and, except in gerontic individuals,
in its relatively flatter valve. Unfortunately the specimens are all
more or less worn in the umbonal region, but the reticulatis orna-
mentation seems to predominate in the young. Some specimens,
however, show in the young shell in P. collazoensis ornamentation,
especially near the submargins. In some individuals the interstitial
thread does not appear until the last stages of growth. In others it
appears early but is not spinose until the adult stage is reached.
In still others the ribs remain unornamented until the adult stage,
though this may be due partly to wear. In the largest shells there
are five threads on the ribs in the last stages of growth, and two
additional threads sometimes appear in the interspaces. The camp-
tonectes striation is obscured by the growth lines, which are es-
pecially strong in this variety.
A large flat-valved specimen measures in length 36 mm., height
39 nmm., semi-diameter 8.5 nmm. In this specimen the ribs are low
and broad. A gerontic shell from the San Sebastian shale measures,
in length 30 mm., height 32 mm., semi-diameter 8.5 mm. These geron-
tic shells show a periodic swelling and contraction of the ribs and are


notably convex. Superficially. they look like another species, e-
pecially when the ribs are worn smooth.
Type localities.-76, and K1. 29.2 Lares road.
Oiher localities.-15S, 22, 2603, 72, 344. 23:, 263 (bottom 3rd
falls). 757, 165 ?, Collazo Falls, and vicinity of Collazo (float), 140.
1, 201, 244.
Ranife.-San Sebastian shale, Lares limestone, Cibao limestone.

Pecten (Chlamys) grabaui, new species

Plate XII, Figure I

Shell transversely sub-ovate, equivalve, moderately convex with the
greatest convexity toward the umnbones'; length 27 mm., height 2(.5 mm..
diameter 11.5 mm.; umbonal third of shell smooth, remainder with twenty-
three low rounded ribs and slightly narrower shallow rounded interspaces;
following the smooth stage appear faint regular widely spaced raised con-
centric threads, strongest on the ribs, which become more distinct as the
ribs increase ventrally. These threads then become irregular, break up into
three rows of fine sharp scales on the ribs. The ribs become threaded and
grooved distally, the scales giving a squamose character to the threads. There
are as many as five of these longitudinal threads on the distal portions of
some ribs. Likewise, in the interspaces an interstitial thread appears which
is similarly squamose. Thus in the final stage of growth there is a typical
Chlamys ornamentation. Oil the lateral slopes the ribs break up into
squamose radial threads, which continue across the submargins and ears:
ears unequal, the anterior ears being longer and more strongly sculptured;
byssal notch deep; ctenolium fine; double pair of cardinal crnra: auricular
crura low, indistinct; camptonectes striation strong.

In first going over the specimens- it scented as thloulgh three or
tour species were represented, but intermediate forms made it nece-
sary to group them as variations of an exceedingly variable species.
similar to P. (Chlamys) indecisus )all, a form from the Vicksbiurg
)ligoeene. Like the latter, this shell shows variations which are
intermediate between a true Chlamys and an Anmusium. As the
lype of the species, that form is chosen which best illustrates the
greatest number of characters cimmol to the group.
Type localities.-343, 619.
Horizon.-Lares formation.


Pecten (Chlamys) grabaui, new species

variety aguadensis

Plate Xll. Figure 2
Shell nioderately convex in umbonal region, nearly flat ventrally; surfl'ace
nearly smooth, but with obsolete ribs visible chiefly as orange colored rays
in the epidermis: camptonectes striation distinct; concentric lamellae, char-
acteristic of the group, visible only under a lens. They are frequently worn
off, leaving microscopic impressed lines in their place; submargins radially
striate; ears rather strongly radially sculptured as in the other varieties;
interior of shell strongly lirate, the lirae being paired and corresponding with
the juncture of rib and interspace on the exterior. The interior is otherwise
the same as in the other varieties: etenolium distinct; length 18.5 mm.,
height 18.5 mm., semi-diameter 3.5 mm.
This interesting variety is a typical Chlamys in shape but 1mas
many of the characters of Amusium. It seems to be the primitive
member of this group, since the other varieties pass through the
smooth aguadensis stage more or less early in growth and develop
the ribbed stage.
Localities.-619, :33, :331.
Horizon.-Lares formation.

Pecten (Chlamys) grabaui, now species

variety hatoensis

Plate X[I, Figure 8
Ribs appear very early, and become high and rounded; concentric lamellae
regular and widely spaced, but becoming irregular near ventral margin
where the ribs are radially grooved and interstitial thread appears in each
interspace; submargins radially striate; ears strongly sculptured; etenolium
distinct; crura as in the type form; lirae obsolete except distally. Length
21.5 mm., height 21 mm., diameter 8.5 mm.
Some specimens do not reach the high ribbed stage, but ribs are
rather low and rounded throughout, with the concentrie lamellae
crowded ventrally and showing a tendency to alternate in strength.
Another specimen from a higher horizon and which may be a muit-
tion from this variety, has high, strong ribs, in which the irregularity
of the lamellae, grooving of ribs. and interstitial threads appear much
earlier in the growth of the shell, and in the interspaces shows as
many as thee interstitial threads in the final adult stage.


Type localilies.-332, 225.
Other localities.-3-.4, 229, 333, 287, 167 ?. 62.
Range.-I.ares formation to Cibao limestone.

Pecten (Chlamys) grabaui, new species

variety guayabensis

Plate XII, Figure 4
Shell marked by smooth submargins. The specimens can be arranged into
a series, as in variety hatoensis, varying from low to high, strong, rounded
ribs, and with concentric lamellae varying from simple and distinctly spaced,
to irregular and crowded. As in variety hatoensis, the ribs appear early,
covering the entire shell except the prodissoconch. Interior of the shell
faintly lirate throughout. Shell otherwise like the type form of the group.
The largest specimen measures in length 17 mm., height 17.5 mm., semi-
diameter 4.5 mm.
Type localities.-62, 332.
Other locality.-2S2.
Range.-Lares formation to Cibao limestone.

The probable relationship of the group is shown in the following

Table 1

Pecien (Chlainys) grabaui, new species

horizon (Iroup with smooth Group with striate
sbinmargins submargins

Cibno variety variety
limestone guayabensis hatoensis
(shell ribbed) (shell ribbed)

Larcs variety P. grabaui
formatiin aiuladen.i.s - (type form)
(shell smonith) (partially ribbed)

San Scb.xsti;:
shale ?


Pecten (Chlamys) hodgii, new species
Plate XII, Figure 5
Shell sub-ovate, sub-equilateral, strongly convex in umbonal region, slightly
convex in ventral region; ears sub-equal, deep byssal notch and strong
ctenolium; surface with twenty rounded ribs and deep, rounded, narrower
interspaces; both ribs and interspaces grooved with fine radial threads which
are densely spinose; ears and the rather broad submargins marked by sub-
spinose radial threads; inner margin strongly scalloped and interior grooved.
In the early stages of growth the ribs are not radically threaded,
and the ornamentation consists of closely spaced, minute, sharp trans-
verse scales on the ribs which join with raised concentric lamellae
in the interspaces. When the shell has reached one third full size,
the radial threads appear and the scales break up into minute densely
spaced spines, which, ventrally, become elevated, curved, and U-
shaped with concave side toward the beaks. Throughout most of
the early stages the interspaces are devoid of threads and spinules,
which appear much later than they do on the ribs. The concentric
lamellae become obsolete ventrally.
This shell is apparently closely related to an unlabelled specimen
in the Kemp collection from Gatun, which seems to be a new species.
In the latter the transverse scale stage is retained longer, the con-
rentric lamellae are noticeable throughout, and the spinules are not
long or conspicuous as in the Porto Rican shell. If they are related,
the Gatun shell would seem to be the more primitive of the two. It
may be related to P. oxygonum canalis and P. oxygonum optimum
of Brown and Pilsbry and to P. gabbi Dall. Dr. Maury has de-
scribed a similar and apparently related shell P. camuycencis from
the Quebradillas limestone. One complete right valve and some frag-
ments were found. This shell is named in honor of Dr. Edwin T.
Hodge, who studied the geology of the Coamo District, Porto Rico.':
Horizon.-Quebradillas limestone (rare).

Pecten (Aequipecten) lobecki, new species

Plate XIII, Figure 1
Shell sub-orbicular, equilateral, moderately convex, ears large, smooth,
apparently sub-equal (portion of anterior ear concealed); shell sculptured
with fourteen strong rounded ribs and sub-equal rounded interspaces; con-


centric sculpture of fine raised lamellae, strong in the interspaces, obsolete OL
tops of the ribs; fine concentric striae in the interspaces between the lamellae,
and from one to three radial threads in each interspace, which with the con-
centric lines produce a cancellate structure. The ribs are nearly smooth
apparently because of wear, but occasionally show faint radial threads
distally; submargins narrow, smooth, not depressed. Length 25 nun., height
24 mm., semi-diameter 4 mm.

A single left valve was found which resembled in some respects
P. cercadica Maury.
Hlorizon.-Lares limestone.

Pecten (Plagioctenium) rabelli, new species

Plate XIT, Figure G(

Shell suborbicular, slightly oblique in adult, sub-equivalve (right valve
slightly more convex); ears small, subequal; shell with thirteen to fourteen
strong, smooth, rounded ribs, with slightly narrower, smooth rounded inter-
spaces; right anterior ear with faint radial theads, other ears smooth; sub-
margins narrow, smooth, depressed, ctenolium distinct. The surface of the
valves presents no very marked feature to the unaided eye, but under a strong
lens, very minute, closely, evenly spaced and remarkably regular concentric
lines are visible, crossing the entire shell, including the ears. In the largest
specimens a gerontic character appears in the periodic swelling and con-
tracting of the ribs. These gerontic shells are more convex than the others.
Length 36 mm., height 34 mm., semi-diameter 8 mm. A smaller specimen
has a semi-diameter of 5 mm.

Localities.-K. 29.2-29).3 Lares road, K. 25 Ponce-Adjuntas road,
Juana Diaz, Yauco.
Horizon.-San Sebastian shale, Juana Diaz sale.

Pecten (Plagioctenium) cercadica Ma;mry

Plate XIII, Figures 2, 3
Pecten ccrcadica Maury, Bull. Amer. Paleont., V, No. 29, p. 1SS, P1. 34, fig. 11,
Shell bilaterally symmetrical, sub-equivalve (right valve of slightly
greater convexity); submargins broad, smooth except for faint growth lines;
ears large, sub-equal, nearly smooth, triangular in outline except the right
anterior, which is marked by deep byssal notch and byssal sulcus below, and
surface of ear with five imbricated radial threads; ctenolium absent ot
obscure; other ears marked by very faint radial threads or striation. strongest


near base of ear; surfaces of valves similar and marked by fourteen to seven-
teen strong rounded ribs and slightly narrower interspaces, the whole crossed
by fine close growth lines, periodically incised so as to give the effect of
obscure beading of the ribs, a feature not evident on worn specimens; valves
grooved within, but lirate only near ventral margins; single pair of strong
cardinal crura. Average size;-length 21 mm., height 21 mm., semi-diameter
3.5 minm.; largest specimen:-length 26 nmm., height 27 mm., semi-diameter
4.5 mm.

This shell resembles P. (Plugiocleuni m) andersoni Arnold of the
Pacific Coast _liocene, but is smaller. The latter species differs in
having the left valve more convex than the right. The shell here
described appears to correspond to Maury's description which was
based on a left valve only.

Type localities.-G19, 333. 324.

Other localities.--331, 227 (top 30 feet).

HIorizon.-Lares limestone.

Pecten (Plagioctenium) borinquenense, new species

Plate XIV, Figures 1, 2

Shell sub-orbicular, sub-equivalve, slightly oblique, sub-marginal slopes
slightly concave (left valve slightly less); sculpture of the two valves dis-
similar. Right valve with sixteen to nineteen strong flat ribs of T-rail cross-
section, wider than interspaces, especially in the unworn specimens where
the lateral overhang obscures the borders of the interspaces; interspaces
rounded and sculptured with raised concentric threads, very regular and
evenly spaced (one-fifth mm. apart in ventral part of shell); tops of ribs
marked by obscure median keel; concentric threads cross the ribs and are
bowed up in the form of lamellae on crossing the median keel. The ribs
are usually worn smooth showing the lamellae on top as a series of V-shaped
striations; ears sub-equal, small; hinge line less than half the length of
shell, sub-margins nearly smooth; ctenolium obscure and marked by deep
byssal groove bordering the line of teeth; byssal notch small; adductor scar
large, central; left valve with seventeen to twenty-one strong, high, rounded
ribs and sub-equal rounded interspaces; both ribs and interspaces crossed
by fine raised concentric threads, somewhat irregular and more distantly
spaced than in the right valve, and worn or obscure on crossing the ribs;
ears sub-equal and sculptured by radial grooves and incremental lines; sub-
margins as in the right valve. Interior of both valves smooth or faintly
fluted, but strongly fluted and lirate around the ventral margins; greatest
convexity in the umbonal region. The ribs of both valves are rounded in the
nepionic stage. Adult shell measures in length 32 mm., height 31 mm.,
diameter of right valve 10 mm.


This shell has ribs similar to P. diegensis Dall of the Pacific
Coast Pleistocene, but is otherwise a different shell. Maury's P.
caimitica, a single left valve from the Santo Domingo Miocene is
similar to the left valve of this shell.
Ilorizon.-Upper Quebradillas limestone.

Genus Amusium Bolten
Amusium papyraceum Gabb
Pleuronectia papyracea Gabb, Trans. Amer. Phil. Soc., V, p. 257, 1873.
Amusium papyraceum Dall, Trans. Wagner Inst. Sci., III, pt. 4, p. 718, 1898;
Idem, pt. 6, p. 1586, 1903.
Amusium papyraceum Maury, Bull. Amer. Paleout., V, No. 29, p. 190, Pl. 26,
fig. 22, 1917.
Found only in fragments, indicating a size of 35 by 36 mnm.
Localities.-118, (talus), 261.
Horizon.-San Sebastian shale.

Amusium mauryi, new species
Plate XIV, Figure 3
Shell small, thick, moderately convex, entirely smooth except for very
fine growth lines; ears sub-equal, large, sub-triangular; juncture of ears with
shell slightly depressed, but not otherwise marked off. The interior of the
valve is inaccessible, but there seem to be no internal ribs or else very
obscure ones. They do not show on the exterior, and there are no radial rays
or striations of any sort. The shell is unusually solid. It is probably related
to A. luna Brown and Pilsbry, and to A. mortoni Ravenel, but differs from
these in its small size and relatively much larger ears. Length 19 mm., height
19.5 mm., semi-diameter 3 mm.
A single left valve, which was at first included with the specimen-
of A. papyraceum. Dr. Maury, in looking over the specimens, pointed
out the mistake.
Locality.-Collazo Falls.
Horizon.-San Sebastian shale.

Amusium (Propeamusium) hollicki Manry
HIorizon.-San Sebastian shale.


Genus Spondylus Linnu
Spondylus bostrychites? Guppy
Spondylus bifrons Sowerby, Quart. Jour. Geol. Soc., VI, p. 53, 1849; not of
Goldfuss, Petref., II, p. 99, pl. 106, figs. 10a-c, 1835.
Spondylus bostrychites Guppy, Proc. Sci. Assoc. Trinidad, p. 176, 1867.
Spondylus bostrychites Gabb, Trans. Amer. Phil. Soc., XV, p. 257, 1873.
Spondylus bostrychites Dall, Trans. Wagner Inst. Sci., III, pt. 4, p. 758, 189S;
pt. 6, p. 1586, 1903.
Spondylus bostrychites Maury, Bull. Amer. Paleont., V, No. 29, p. 190, pl. 32,
fig. 4, 1917.
A small internal mold of a Spondylus, which, because of its
small hinge area and sub-equal valves, is identified doubtfully with
the above species. Length 22 umm., height 26 mm., diameter 15 mm.
Localities.-162, 23 ?, 126.
Range.-San Sebastian shale to Cibao limestone.

Spondylus gumanomocon Brown and Pilsbry
Spondylus americanis Gabb, Trans. Amer. Phil. Soc., V, p. 257, 1873; not of
Spondylus gumanomocon Brown & Pilsbry, Proc. Acad. Nat. Sci. Phila., p. 514,
1912 (footnote).
Spondylus gumanomocon Maury, Bull. Amer. Paleont., V, No. 29, p. 191, 1917.
An internal mold and a fragment of the external mold were found
of a shell which is like S. gumanomocon but somewhat smaller. It
may be a small variety or the young shell. Lower valve strongly
convex, beak long, erect; surface with strong ribs 5 mm. apart, ap-
parently spinose, and alternating with three or four small obscure
ribs. The ribs are slightly wavy; upper valve orbicular, slightly
convex; ribs known only from the internal mold but seem to be like
those of the upper valve except that they are finer; hinge line about
24: mm. in length. The internal mold shows the beak of the lower
valve to be considerably excavated. Length 55 mm., height of lower
valve 66 mm., of upper valve 54 num., diameter 29 mm.
From the proportions of the internal mold it will be seen that this
shell is less elongate than S. gumanomocon as described from Saito
Domingo. The upper valve is much flatter than the Gatun species,
S. scotti Brown & Pilsbry, but the shell is about the same size as the
latter and is similarly sculptured.
Localities.-140, 162.
Range.-San Sebastian shale to Cibao limestone.


Genus Ostrea (L) Lamarek
Ostrea antiguensis Brown
Plate XV. Figure 1
Ostrea antiguensis Brown, Proc. Acad. Nat. Sci. Phil.. p. 614, Pl. 19, fig. 7;
P1. 20, figs. 1, 5, 6, 1913.
These shells were at first regarded a, a variety of 0. hiWentss,
but on comparison, were found to be the same as Dr. Alaury's Porto
Rican specimens, id( notified by D)r. Pilsbry as 0. antiguensi.t, a-
cordingly that determination is accepted, although it is difficult to
believe that the retlexed character of the upper valve is a specific
All of the specimens here described were taken from a zone of
yellow chalk in the upper Quelradillan expo-ed in sea clitAS near
Pt. Borinquen. The abundance of these large oy-ters is remarkable.
They reach an enormous size and thickness, but the largest specimen
collected measures in length 140 nmm.. height 1:0 ium., diameter
(the two valves together) 110 nmm. The-e form- are obviously geron-
ite individuals. The lower valves are strongly curved upward
ventrallv toward the center, but are reflexed (or curved down) on
the lateral margins. Likewise, the upper valves are curved down on
the lateral margins, and strongly reflexed at the ventral margin,
which results in a strong interlocking of the two valves. The upper
valves are almost as thick as the lower ones. The upper valve.
reach their maximnnum thickness between the center and the ventral
margin, and in both valves the animal has excavated its bodv cavity
directly below the beaks by resorption of the prislatie layers at this
point with increased deposition farther on toward the ventral mar-
gin. Thus as seen in a longiludinam section of the AhIll these lavers
resemble the stratification in a migrating sand dune. The excavated
body cavity occupiis blut a miall portion of the interior of tile shell.
Other specimens were found which. although almhnt as lair'e as
those above described, are apparently not gerontic intlividual,. The
reflexed character of the upper valve is slightly or altogether absent.
and the shell does not attain a great thickness. The spinose ribs
(about 9 in number) and rounded furrow', are well -hown on these
specimens, whereas they are obsolete or extremely irregular on tlhe
gerontic forms. The adductor sear is high and nearly central in tilhe


thin shells, but well toward the anterior margin in the gerontic
shells. It is deeply impressed in the Inater. The resilifer is re-
markably small for shells of this size.
Localities.-83 : San Juan Districi (Seamnes).
HIorizon.-Upper Quebradillas limestone. Also reported from the
"Guanica shaly limestone" by Dr. Maury.

Ostrea haitensis Sowerby
0. haitensis Sowerby, Quart. Jour. Geol. Soc. London, VI, p. 53, 1850.
0. haytensis Gabb, Trans. Amer. Phil. Soc., XV, p. 257, 1873.
0. veatchii Gabb, Pal. Cal., II, p. 34, Pl. 11, fig. 59; PI. 17, fig. 21, 1869.
0. hermanni Conrad, Proc. Acad. Nat. Sci. Phila., V, p. 267, 1853.
U. vespertina Conrad, Pac. R. R. Rept., V, p. 325, Pl. 5, figs. 36-38, 1855; Gabb,
Pal. Cal., II, p. 107, 1869.
0. virginica Guppy, Quart. Jour. Geol. Soc., XXII, p. 577, 1866. Not of Gmelin.
0. virginica var. californica Marcou, Geol. No. Am., 1858.
0. haitensis Guppy, Quart. Jour. Geol. Soc. London, XXXII, p. 532, 1876.
0. haitensis Dall, Trans. Wagner Inst. Sci., III, pt. 4, p. 685, 1898. Idem, pt. 6,
p. 1586, 1903.
0. haitensis Maury, Bull. Amer. Paleont., V, No. 29, p. 182, P1. 31, figs. 1, 2,

The shells are rather thin, orbicular in outline, and strongly pli-
eate with seven or eight sharply angular ribs, very high centrally
at the ventral margin, and sharply angular furrows between. The
ribs are not noticeably spinose; resilifer shelf-like, due to excavated
beak; adductor situated in central-posterior portion of vale. The
shells do not attain a large size, the largest specimen measuring in
length 72 1mm., height 74 mm., diameter of lower valve about 30 unm.
Localities.-225, 800.
Range.-Lares limestone, Cibao limestone.

Ostrea haitensis Sowerby, variety?

Plate XV, Figures 2, 3

Some small shells were found in which the lower valves are
sculptured like 0. haaitensis, with spinose ribs, frequently angular,
while the upper valves are nearly smooth. The lateral inner mar-


gins adjacent to the beaks are finely crenulate, a feature not shown
in the specimens of 0. hailensis. Largest specimen measures in
length 36 mm., height 43 mm., diameter lower valve 7 mm. A
smaller more orbicular lower valve measures in length 22 nmm..
height 25 mm., diameter 6.5 mm. These little shells may be the
young of 0. haitensis, or a variety.
Horizon.-Cibao limestone.

Ostrea virginica Gmelin
0. virgiana of Lister and others.
0. virginica Gmelin, Syst. Nat., p. 3336, 1792; Dilwyn, Descr. Cat., 1, p. 277,
1817; Lam., Anim. s. Vert., VI, p. 207, 1819.
O. edulis Akerly, Am. Monthly Mag., II, p. 296, 1818 (not Linnd).
0. virginiana Sowerby, Genera, Ostrea, f. 2, 1822.
0. borealis Lamarck, Anim. s. Vert., VI, p. 204, 1819.
0. canandenis Lamarck, op. cit., p. 207, 1819.
0. triangularis Holmes, Proc. Elliot Soc., I, p. 29, 1856.
0. fundata Holmes, Post-Pl. Foss. S. Car., p. 11, Pl. 2, fig. 10, 1858.
0. semicylindrica Say, Jour. Acad. Nat. Sci. Phila. (1), II, p. 258, 1822.
0. virginica Gabb, Trans. Amer. Phil. Soc., XV, p. 257, 1873.
0. virginica Guppy, Quart. Jour. Geol. Soc. London, XXXII, p. 532, 1876.
0. virginica Dall, Trans. Wagner Inst. Sci., III, pt 4, p. 687, 1898.
0. virginica Maury, Bull. Amer. Paleont., V, No. 29, p. 184, 1917.
The specimens show the usual great variation, but comparison
with recent specimens of 0. virginica shows no constant differences.
The Porto Rican shells vary from elongate, subtriangular to ovate:
beak of lower valve usually pointed, straight or slightly curved
laterally; resilifer elongate in the large shells; lower valves are
frequently of extreme convexity, while the upper valves are gently
convex or flat, but very irregular; beak of lower valve excavated.
An average seized lower valve measures in length 50 mm., height
90 mm., diameter 33 mm. Some are much broader, and a few
specimens reach a much greater size and greater thickness. Colir
usually bluish, mottled with white.
Type locality.-282.
Other localities.-281, 65, 227 (top 30 ft.), 7 to S (on road-flat).
Horizon.-Upper lares limestone (very characteristic).


Ostrea sellUeformis Conrad

variety portoricoensis, new variety

Plate XIII, Figures 4, 5, 6

Shell of moderate size, pear-shaped to elongate oval, never attaining very
great thickness; valves dissimilar; lower valve with numerous radial ribs
imbricated by raised laminae; resilifer deep; margins of ligament area
marked by narrow groove which continues for some distance along the inner
margins adjacent to the beak, and is denticulate (or marked by row of small
pits) throughout; remainder of inner margin fluted by the ribs, or, in some
specimens, smooth; adductor scar large, oval, nearly central; beak more or
less curved laterally; upper valve smaller, convex to nearly flat; surface
nearly smooth or marked usually by even, widely spaced, slightly raised
concentric lamellae; beak curved laterally but less pointed than in lower
valve, and with shorter ligament area; resilifer moderately impressed or
even raised and convex in some individuals; lateral grooves indistinct or
absent; inner margin crenulate adjacent to beaks, but smooth ventrally;
adductor scar smaller than in lower valve and uniformly pear-shaped in all
specimens. An average sized shell measures, lower valve, in length 62 mm.,
height 70 mm., upper valve, in length 42 mm., height 60 mm. Diameter (both
valves) 19 mm. The upper valves of the young shells are more convex. A
typical young upper valve measures in length 23 mm., height 33 mm.,
diameter 7 mm.

Type localities.-343, 333, 176, 59, SO0.

Other localities.-61 9, 332, 64.

Range.-Lares limestone, Cibao limestone, Los Puertos limestone.

Ostrea collazica Maury

Shell sub-elongate to sub-orbicular, heavy, attaining great thickness in
large individuals; valves dissimilar; lower valve convex, with numerous
small irregular radial plications which divaricate from a median or slightly
posterior zone; plications finely imbricated by thin, slightly raised laminae;
beak strongly recurved laterally; ligamental area extremely long in large
specimens; resilium deep but becoming shallow in gerontic individuals;
margins of ligamental area marked off by narrow crenulated groove; inner
margins near the beak finely crenulated; upper valve flat, with beak similarly
recurved, usually more closely gyrate; surface smooth except for the thin
laminae; interior similar to lower valve except for an amorphous, tooth-like
process, located on the inner margin just anterior to the ligamental area,
which is transversely denticulate or marked by cross grooves. In some speci-
mens the margins of both valves are similarly marked near the beaks;
adductor scars of both valves large and situated slightly posterior. Average
size adult;-length 75 mm., height 85 mm.


Fragments indicate that the shell reaches a co(nsiderably larger
size, in which the lower valve is greatly thickened. Two of these
large lower valves were found in which the tendency of the beak to
curve laterally lias apparently been ahnost lost. the beak heing long
and pointed. An unisnal feature of this shell is the small amount, of
prismatic structure. Some of the specimens are nmad up almost
entirely of naereous laminiae 1f micaceous tliinne-.
This shell is of the 0. sellbformis di'aricata type, and the young
individuals might readily Ibe taken for a variety of 0. selllformiis.
The full grown flat upper valves with their closely gyrate beak bear
a strong resemblance to 0. raugi li Dall of the Tampa Silex beds.
Type localities.-7 to 8 (float), Lares road just east of Collazo
bridge, K. 29.2 to 29.3 Lares road.
Oliher localities.-100, 22G, 231, 233, 207, 2b. 232(b), 20G6.
Iange.-San Sebastian shale to Lares limestone. Characteristic of
the San Sebastian shale, and occuring in the lowest fossiliferous
zone of the latter.

Ostrea cahobasensis Pilsbry and Brown

variety portoricana, new variety

Plate XVI, Figure 1: Plate XVII, Figure 1
Shells reach a very large size, and are characterized by their elongate
form and long, pointed, and usually straight beak. The young shells are
much less elongate in form. The body cavity tends to remain the same size
during growth, most of the addition in height being in the beaks. The
resilifer is shallow in the lower valve, and convex in the upper valve, and in
both valves it is bordered by a broad, prominent ridge on each side. The latter
are marked off from the lamellae of the lateral slopes by a deep impressed
zone, in which there is no trace of dentition or crennlation. The beaks may
be twisted or straight, and there is considerable variation in the outline of
the shells due to the accidents of growth. The shells apparently grew in
crowded colonies. The lower valve is convex, the upper slightly convex, or
nearly flat in the younger shells. The beak of the lower valve always pro-
jects far beyond the beak of the upper valve, and in the young, is deeply
excavated within, below the resilifer. The position of the adductor scar is
variable, being rather high in the young, but travels ventrally with the
growth of the shell, until, in the largest individuals, it is located at the
ventral one-third or one-fourth of the shell. The large shells have an hour
glass-shaped body cavity, consisting of a deep round cavity immediately below
the resilifer, a contraction centrally, and a large crescent-shaped ventral
cavity which begins opposite the adductor scar, and swings around below the


scar, and close to the ventral margin. This is a fairly constant fea-
ture of the larger shells, and is strongly developed in some of the
young or half grown shells. The surface of both valves is marked
only by concentric lamellae, and shows no signs of radial fluting of
ribs. The lamellae are, however, occasionally quite irregular. This shell,
while apparently answering the description of the Haitian species, shows
differences of varietal rank. The largest specimen measures in length 100
mm., height 270 mm., diameter (both valves together) 95 mm.

Localities.-West of Guanica Mill in basal limestone a few feet
above the contact with the Older Series (Lobeck) Fragments of a
similar shell were found in the Lares limestone, but their identity
with the Gnanica -hell is by no means certain.

1Horizons.-P, nce limestone, Lares limestone (doubtful).

Genus Lithophaga Bolten

Lithophaga nigra d'Orbigny

Lithodofmus niger d'Orbigny, De la Sagra, Hist. Polit. y Nat. Isla de Cuba,
pt. 2, V, p. 351, 1845; Atlas, VIII, Pl. 28, figs. 10, 11, 1855 (Spanish Ed.).
Modiola caribira Philippi, Abbild. u. Beschr., III, p. 20. P1. 2, fig. 5, 1847.
Modiola antillarum Philippi, op. cit. p. 20, PIl. 2, fig. 4, 1847; Zeitscher., p. 116
(not of d'Orbigny); young shell.
Mytilus lithophogus Gibbes, S. Car. Cat., p. 22, 1848; not of Linn&.
Lithophagus nigra Morch, Cat. Yoldi, II, p. 56, 1853.
Lithophagus antillarutm Reeve, Conch. Icon., X, Pl. 2, fig. 7, 1857.
Lithophaga nigra Dall, Trans. Wagner Inst. Sci., III, pt. 4, p. 799, 1898; Bull.
90, U. S. Nat. Mus., p. 129, 1915.
Lithophaga nigra Maury, Bull. Amer. Paleont., V., No. 29, p. 194, 1917.

This shell is known from the Tampa Silex beds. and continues
to the recent fauna of the Antilles. The recent shell of Cuba as
described by d'Orbigny, has a length of 50 nmm. The Porto Rican
fossils differ only in size, the largest measuring in length 30 mm.,
height 90 mm., diameter S umm. The occurrence is chiefly in the
form of internal molds, but fragments of the shell show the vertical
striae which end abruptly at a transverse oblique line extending
from the beaks to the posterior ventral margin.


Horizon.-Upper Lares limestone.


Genus Leda Schumacher

Leda peltella Dall

Plate XVII, Figure 6
Leda acuta Gabb, Trans. Amer. Phil. Soc., XV, p. 255, 1873. Not L. acuta
Conrad, 1832, nor Sowerby, 1837.
Ledapeltella Dall, Trans. Wagner Inst. Sel., III, pt. 4, p. 579, Pl. 32, fig. 5;
Idem, pt. 6, p. 1586, 1903.
Leda peltella Maury, Bull. Amer. Paleont., V, No. 29, p. 161, Pl. 26, fig. 9, 1917.

One of the most abundant fossils in the Quebradillas limestone.
The posterior sulcation is always broad and distinct; anterior sulcation
narrow and often indistinct. Length 11 nun., high 6.5 mm., semi-
diameter 3 nun. No specimens were found less than 9 mm. in length.
Type locality.-204.
Other locality.-660.
Horizon.-Quebradillas limestone.

Genus Area Linn6

Area yaquensis Maury

Arca yaquensis Maury, Bull. Amer. Paleont., V, No. 29, p. 164, Pl. 30, fig. 14,

Several specimens were found of a very small Arca which closely
resembles 21aury's A. vaquensis. a somewhat doubtful species from
the Santo Domingan Oligocene. The Porto Rican shells show no
distinctive characters, and are placed with A. yaquensis because they
correspond more closely to this than to any other species. The
specimens are very numerous and all about the same size, hence it is
unlikely that they arc young individuals. Length 8 imm., height
4.5 mm.
Type locality.-204.
Other locality.-225.
Jlange.-Lares limestone to Quehradillas limestone.


Arca dariensis Brown and Pilsbry

Plate XV, Figure 5
Area dariensis Brown and Pilsbry, Proc. Acad. Nat. Sci. Phila., p. 362, Pl. 22,
fig. 10, 1911.
Umbo prominent, at anterior third of shell; surface with thirty-three flat
ribs and sub-equal flat interspaces, the whole crossed by fine raised concentric
lines, strongest on ribs and giving to the latter a nodular appearance; ribs
broaden anteriorly and posteriorly, the ten posterior and the five or six
anterior ribs medially grooved. Length 26 mm., height 16.5 mm., semi-
diameter 5.5 mm.
This Area is smaller than the Gatun species, and may be a variety
of it. It also resembles the shell described by Toula as Area cf.
consobrina Moore.
Localities.-Collazo Falls, 261?, 232(b).
Horizon.-San Sebastian shale.

Area (Scapharca) cf. donacia Dall

Plate XV, Figures 6, 7
Arca (Scapharca) donacia Dall, Trans. Wagner Inst. Sci., III, pt. 4, p. 649,
P1. 33, fig. 13, 1898.
The size and sculpture answers Dall's description of this species,
especially the raised concentric threads in the interspaces of the left
valve, a feature lacking on the right valve. It differs from the
Bowden form in the number of ribs (27), and in having a shallow
median depression, strongest in the left valve. Length 8.5 mnm.,
height 6.5 mm. A smaller shell measures in length 6.75 mm., height
4.5 mm.
Horizon.-Quebradillas limestone.

Area, species indet.

Plate XVII, Figures 2, 3
Shell small, very convex; beaks at anterior third; anterior and posterior
ends equally rounded; hinge area broad, especially anteriorly, with beaks
widely separate; surface with twenty-five broad, quadrate, rather low ribs and
equally wide, shallow rounded interspaces; central ribs medially grooved
distally; hinge margin three-fourths the length of shell and slightly raised


above hinge area; hinge area transversely grooved or striate posteriorly;
teeth fine, numerous; interior of valves unknown. Average size measures in
length 19 mm., height 10.5 mm., diameter 12 mm. Largest specimen meas-
ures in length 25 mm., height 13.6 mm., diameter 16 mram.
Type locality.-2 ,0.
Other localities.-233, 232(b).
HIorizon.-Upper San Sebastian shale.

Arca (Scapharca) collazica Maury

The material collected by the writer consists only of internal molds.
An average sized specimen measures in length 16.5 mim., height
9.5 imn., diameter 7.5 mm.
Localities.-333, 019, 64, 232(b).
Range.-Upper San Sebastian shale to (Cibao limestone. Very

Subgenus Barbatia Gray

Barbatia (Acar) reticulata Gmelin
Arca reticulata Gmelin, Syst. Nat., VI, p. 3311, 1792.
Arca reticulata Sheldon, Palaeontographica Americana, I, p. 20, P1. 4, figs.
8-12, 1916.
Barbalia (Acar) reticulata Maury, Bull. Amer. Paleont., V, No. 29, p. 166,
PI. 30, fig. 16, 1917.
A single fragment was found, sufficiently complete to identify.
Length indeterminate, height 6 nun.
HIlorizon.-Quebradillas limestone.

Barbatia ef. bonaczyi Gabb

Plate XIV, Figure 4
Barbatia bonaczyi Gabb, Trans. Amer. Phil. Soc., XV, p. 254, 1873.
Area umbonata Dall, Trans. Wagner Inst. Sci., III, pt. 4, p. 620, Pl. 38S, figs. 4,
4a, 1898. Not of Lamarck.
Barbatia of. bounaczyi Mlaury, Bull. Amer. Paleont., V, No. 29, p. 165, Pl. 30,
fig. 15. 1917.
A single external mold of a right valve resembling I?. bonaczyi
(,abb, but smaller. It is nearer the size of B. ef. bonaczyi described


by Maury, but the sculpture is coarser. It is very probably a
variety, but the specimen is too poor for complete description. The
ribs are crowded centrally and sparser at the ends of the valve.
Those on the posterior slope lose their headed character and become
scaly. Valve with shallow median sulcation. Length about 12 mm.,
height n mm.
Horizon.-Qnebradillas limestone.

Genus Glycimeris Da Costa
Glycimeris portoricoensis, new species
Plate XV, Figure (3
Shell sub-orbicular; beaks not prominent; surface with about thirty strong
sub-angular to rounded ribs and sub-angular interspaces, the whole crossed
by fine raised concentric threads, regular and evenly spaced. The ribs near
the ventral margin broaden rapidly at the expense of the interspaces, and
become round in cross-section. Interior of shell unknown. Largest specimen
measures in length 19 mm., height 17 mm., semi-diameter 4 mm.
This shell resembles G. acnticostata Sowerby, but has broader, more
rounded ribs.
Localities.-204, 182, 31S.
Range.-Upper Los Puertos limestone to Quebradillas limestone
(typical of the latter).

Glycimeris collazoensis, new species
Plate XV, Figures 4, 5
Shell sub-orbicular, beaks not prominent; hinge area short; teeth numer-
ous, in uninterrupted series; surface with thirty-four low angular ribs and
angular interspaces; each rib with fine keel or thread on the apex, and with
the appearance of a groove on either side of this median thread; inner margin
scalloped ventrally; length 15 mm., height 16 mm., semi-diameter, 4 mm.
This shell resembles G. gatunensis Toula, but the latter is smaller
and more transversely elongate. It also resembles G. porloricoensis
of the Quebradillas limestone, of which it inay possibly be the pro-
genitor. G. portoricoensis differs chiefly in its larger size, more
rounded ribs, and strong concentric growth lines.
Horizon.-Lower San Sebastian shale.


Genus Venericardia Lamarck

Venericardia scabricostata? Guppy
Cardita scabricostata Guppy, Quart. Jour. Geol. Soc. London, XXII, p. 293,
Pl. 18, fig. 10, 1866.
Cardita scabricostata Gabb, Trans. Amer. Phil. Soc., XV, p. 252, 1873.
Cardita scabricostata Guppy, Quart. Jour. Geol. Soc. London, XXXII, p. 531,
Venericardia scabricostata Dall,, Trans. Wagner, Inst. Sci., III, pt. 6, pp. 1428,
1586, 1903.
Venericardia scabricostata Maury, Bull. Amer. Paleont., V., No. 29, p. 198,
P1. 33, fig. 1, 1917.

Shell with prominent beaks; 18 to 20 high nodose ribs and nar-
rower, V-shaped interspaces which are crossed by strongly marked
growth lines; lunule broad, short, and distinct. Length about 4.5
mm., height 4 mm., semi-diameter 1.5 mm. The specimens found
are much smaller than V. scabricostata, and yet seem to be mature
shells; they may be a variety.

Horizon.-Lares limestone.

Venericardia cerrogordensis Maury, new variety
Venericardia cerrogordensis Maury, Bull. Amer. Paleont., V, No. 29, p. 199,
P1. 33, fig. 3, 1917.

A single fragment of anterior half of a left valve, which seems to
be a variety of this species, somewhat larger than the Santo Domingo
shell. About fourteen strong ribs with strongly beaded keel; inter-
spaces wider than ribs, shallow, with wavy growth lines and showing
an obscure crenulated radial thread on either side at base of rib.
Length about 27 mm., height about 23 mm., semi-diameter, etc.,
5.5 mm.
This may be a distinct species but the specimen is so incomplete
that it has been identified with the nearest form. The specimen wa<
found in a piece of road metal at K. 26.5 Lares road and its horizon
is unknown. The rock is a white limestone like the Quebradillas
and contains a mold of Chionc cf. woodwardi Guppy.


Genus Chama (Linne) Bruguierc
Chama involuta Guppy
Chama involuta Guppy, Geol. Mag. London, decade 2, I, p. 436, Pl. 17, figs. 5a-c,
1874; Quart. Jour. Geol. Soc. London, XXXII, p. 531, 1876.
Chama involuta Dall, Trans. Wagner Inst. Sci., III, pt. 6, pp. 1398, 1586, 1903.
Chama involuta Maury, Bull. Amer. Paleont., V, No. 29, p. 199, P1. 33, figs. 4,
5, 6, 1917.
The Porto Rican specimens are unusually large, but apparently
of this species. The largest specimen is fragmental, but shows a
height of 52 mm. Judging from the specimens, it attaches by either
valve. The beak is strongly prosogyrate, making over one complete
turn. The lamellae are strongly fluted, almost spinose on one
Localities.-204, 98 (Berkey).
Horizon.-Quebradillas limestone.

Chama portoricana, new species
Plate XVII, Figure 5
Beak closely prosogyrate; surface with extremely crowded, thin, high
lamellae, which on the largest specimen rise 8 mm. above the shell surface.
These lamellae are quite regular, being gently fluted over the body of the
shell, but spinose at the margin in the adult; shell marked by rounded um-
bonal ridge; inner margin finely crenulate; cardinal area pustulose; strong
bifid and crenulate cardinal tooth in the right valve, with crenulate lateral;
shell attaches by either valve, but the cardinal tooth remains in the right
valve in either case; adductor scars large; pallial line distinctly impressed.
Length 48 mm., height 35 mm., diameter of convex valve about 12 mm. A
smaller attached valve measures length 22 mm., height 18 mm., diameter
9 mm.
This shell differs from 0. congregatoides Maury in its more regular
and higher lamellae, and in its more elongate form.
Horizon.-Quebradillas limestone.

Genus Lucina (Brugui&re) Lamarck
Lucina collazoensis, new species
Plate XVIII, Figures 1, 2
Shell transversely ovate, convex, produced posteriorly, beaks being slightly
forward of center and frequently so close together that they practically touch;


hinge line long and straight, nearly nine-tenths the length of the shell. The
largest specimen, somewhat distorted so as to increase the convexity, meas-
ures in length 65 mm., height 40 mm., diameter 45 mm. A smaller shell, not
distorted, measures in length 39 mm., height 31 mm., diameter 25 mm.
This is a ponderous shell of the type of L. corpulenta Dall, but
unusual for its elongate form and long straight hinge line. It oc-
curs chiefly as internal molds of both valves intact, but the form is
often distorted by pressure. Fragments of external mold show the
surface to be nearly smooth, marked only by fine growth lines, peri-
odically incised at irregular intervals.
There is an undescribed shell in the Kemp collection from the
lower Gatun beds which is apparently this species, or very close to it.
Localities.-74, 92, 86, Collazo (float).
Horizon.-San Sebastian shale (abundant at Collazo Falls).

Lucina cf. chrysostoma (Meuschen) Philippi

Plate XVIII, Figure 3
Tellina crysostoma Meuschen, Mus. Gevers., p. 482, 1787 (typographical
Lucina chrysostoma Philippi, Abb. und Beschr. neue Conchyl., II, p. 200,
P1. 1, fig. 3, 1847.
Venus edentula Chemnitz, Conch. Cab., VII, Pl. xl, figs. 427-9, 1784; Gmelin,
Syst. Nat., VI, p. 3286, 1792; not of Linn6, Syst. Nat., ed. 10, p. 689, 1758.
Anodontia alba Link, Beschr. Rostock Samml., p. 56, 1807.
Lucina edentula Reeve, Conch. Icon., Lucina, Pl. ii, fig. 9, 1850; Hellprin,
Trans. Wagner Inst., 1, pp. 102-3, 1886.
Loripes chrysostoma Arango, Fauna M\al. Cubana, p. 257, 1878.
Lucina chrysostoma Dall, Trans. Wagner Inst., III, pt. 6, p. 1354, 1903.
Loripes edentula Gabb, Trans. Amer. Phil. Soc., XV, p. 251, 1873.
Lucina chrysostoma Maury, Bull. Amer. Paleont., V, No. 29, p. 202, Pl. 35,
fig. 2, 1917.
Resembles L. chrysosloma of the Caloosahatchie Pliocene, but the
largest and most convex specimens resemble the recent L. globosa
Forskol. Only internal molds have been found, which show lunule
impression, and posterior sulcation close to dorsal margin. The
specimens range in length 39 mm., height 36 mm., diameter 22 unm.
to length 135 mm., height 115 mm., diameter 95 mm.
Localities.-205, 174. 10, 78, 98, 176 ? (Berkey). Mona Island,
south side of Vieques Island, Guanica, and Ponce (Lobeck).


Ilorizons.-Los Puertos, Quebradillas limestones, and Ponece for-
mation. These large shells are very abundant as internal molds in
the Tertiary formations of the north and south coasts.

Genus Phacoides Blainville
Phacoides (Miltha) riocanensis Maury
Phacoides (Miltha) riocanensis Maury, Bull. Amer. Paleont., V, No. 29, p. 205,
PI. 35, fig. 7, 1917.
The Porto Rican shells are the same size as those of the Santo
Domingan Oligoeene. There seem to be no important differences.
The former shells have a slight differentiation in the concentric
lamellae, two lamellae of secondary strength alternating with each
primary one. A shallow median sulcns is a noticeable feature near
the ventral margin, but does not extend to the beak. Length about
47 mm., height about 45 nmm., semi-diameter 7.5 mnm.
Localities.-335, 204, 220, 225, 31S, 317.
Range.-Lares limestone, Cibao limestone, Los Puertos limestone,
Quebradillas limestone. This species is widespread but not abundant.

Phacoides (Miltha), species indet.

Plate XVIII, Figure 4
Shell of moderate size, convex, slightly inequalateral; hinge area narrow,
short, beaks incurved and nearly in contact; anterior margin broadly rounded,
posterior margin roundly truncate, basal margin nearly straight; posterior
sulcus close to margin; concentric sculpture of fine, even, raised lamellae,
about 1/3 mm. apart at ventral margin, and with fine growth lines in the
spaces between. Height of shell about 24 mm., length about 34 mm., semi-
diameter 7 mm.
Only a single fragment was found which was too incomplete for
an accurate description, but differed from any other species. This
shell resembles P. (Miltha) cf. smithwoodwardi Maury, a species
from the same horizon, but is more elongate than the latter, and has
the posterior suleus much closer to the margin.
Horizon.-Quebradillas limestone.


Phacoides (Miltha), species indet.

Plate XVITT, Figure 5
Shell small, orbicular, moderately convex. Sculpture of fine concentric
lamellae like P. hiillsboroense; deep radial sulcus crossing the posterior
one-quarter of shell and extending to ventral margin. Fine concentric striae
between the lamellae are visible under a lens. Length 12 mm., height 12.5
mm., semi-diameter 3.5 mm.
Only one specimen was found, the external mold of a left valve.
It may possibly be the young of a large species like P. riocanensis.
Horizon.-Quebradillas limestone.

Phacoides (Pseudomiltha) laresensis, new species

Plate XVIII, Figure 6
Shell sub-orbicular, nearly equilateral, moderately convex; surface with
concentric lamellae of the type of P. hillsboroense, but very regular instead
of wavy as in the latter; concentric striae between the lamellae; lunule
narrow, deeply impressed; escutcheon large, marked off by a narrow sulcus
which broadens at the posterior margin; concentric sculpture flexed sharply
upward over the escutcheon, where it becomes coarser; surface of shell
marked by faint radial striae; cardinal area edentulate; interior of shell
unknown. The shell is thin, and in most of the specimens is worn partially
or wholly away, leaving only the internal mold. Length 36 mm., height
34 mm., diameter 18 mm.
Localities.-262, Collazo Falls, 232(b) ?
Horizon.-San Sebastian shale.

Phacoides (Lucinisca) calhounensis Dall

Plate X, Figure 6
Phucoides (Lucinisca) calhounensis Dall, Trans. Wagner Inst. Sci., III, pt. 6,
p. 1371, Pl. 52, fig. 16, 1903.
Phacoides calhounsis Maury, Bull. Amer. Paleont., V, No. 29, p. 204, 1917.
(Typographical error.)
These shells are variable but have the same general characters a,
the Florida form. The majority of the specimens are the same size
as the latter, but occasionally attain a considerably greater size. The
greater prominence and sparcity of the concentric ridges in the urn-


bonal region is a noticeable feature. The radial ribs are finer and
somewhat crowded on the posterior slope and to a lesser extent on
the anterior slope. Dall does not mention this character but it may
be observed in his illustration. The intersections of the radials with
the concentric ribs have a tendency to form spinose nodes, especially
in the adult stage near the lateral margins. This gives the shell a
superficial resemblance to P. hispaniolana Maury, but the sculpture
of the latter is much finer. The Porto Rican shell has a large and
distinct lunule. An average size specimen measures in length 10
mnm., height 9.5 mm., semi-diameter 2.5 mm. The largest specimen
measures in length 13 mm., height 12 mm., semi-diameter 4 mm.
Type locality.-204.
Other localities.-52(a), 77(B1erkey).
Horizon.-Quebradillas limestone (abundant); Ponce limestone.

Genus Divaricella Von Martens
Divaricella prevaricata Guppy
Divaricella prevaricata Guppy, Proc. U. S. Nat. hMus., XIX, No. 1110, p. 327,
PI. 30, fig. 4, 1896.
Divaricella prevaricata Dall, Trans. Wagner Inst. Sci., III, pt. 6. pp. 1389, 1587,
Divaricella prevaricata Maury, Bull. Amer. Paleont., V., No. 29, p. 207, Pl. 35,
fig. 10, 1917.
The divaricate lines are finer than those of the Santo Domingo
shell. The single specimen found measures in length 7 mm., height
5.75 mm.
Horizon.-Quebradillas limestone.

Genus Codakia Scopoli
Codakia (Jagonia) magnoliana Dall
variety borinquenense, new variety
Plate XVII, Figure 4
A single right valve was collected. It is slightly smaller than the
shell described from the North Carolina Miocene, and the concentric
sculpture is more pronounced. The radials are slightly wavy as in


Codakia orbicularis, and increase by intercalation. They vary in
strength, parts of the shell being marked by ribs finer and more
crowded than in adjacent portions of the surface. The shell ib
noticeably inequilateral. The lunule is long and narrow. Length
10 mm., height 8 mm., semi-diameter 4 mm.
Localities.-204, 671.
Horizon.-Quebradillas limestone.

Genus Cardium (Linnd) Lamarck
Cardium (Laevicardium) of. serratum Lhinn
Cardium serratum Linn6, Syst. Nat., ed. 19, p. 680, 1758.
Cardin, 1(cvigatumn Lamarck, Anim. s. Vert., VI, pt. 1, p. 11, 1819 (not of
Born, Mus. Vind. Test., p. 47, 1780; nor of Linn6, Syst. Nat., X, p. 680,
Cardium citrinum Wood, Gen. Conch., p. 223, Pl. 54, fig. 3, 1815.
Liocardium pictuimRavenel, Proc. Acad. Nat. Sci. Phila., for 1861, p. 44, 1862.
Cardium hiatus "Meuschen" fide Krebs, W. I. Cat. Shells, p. 115, 1864.
Cardium lineatum Krebs, op. cit., not of Gmelin, 1792.
Cardium pristis Valenciennes, fide Krebs, op. cit.
Cardium oviputamen Reeve, Conch. Icon., Cardium, Pl. 7, fig. 36, 1844.
Cardium venustum Gabb, Trans. Amer. Phil. Soc., XV, p. 251, 1873.
Cardium serratum Dall, Proc. U. S. Nat. Mus., XIX, No. 1110, p. 327, 1896;
not serratum of Pennant, 1778).
Cardium serratum Dall, (in part), Trans. Wagner Inst. Sci., III, pt. 5, p. 1110,
Cardium serratum Brown and Pilsbry, Proc. Acad. Nat. Sci. Phila., p. 367, 1911.
Cardium serratum Maury, Bull. Amer. Paleont., V, No. 29, p. 212, Pl. 36, fig. S,
Numerous internal molds apparently of this species, and of about
the same size as those of the Santo Domingo Oligocene were found.
They average about 20 mm. in height. Fragments of external mold
show numerous fine thread-like radial ribh. Some specimens show a
slight radial sulcation anteriorly.
Localities.-204, 225, 333?, 64.
]?ange.-Lares limestone to Quebradillas limestone.

Cardium (Trachycardium) muricoides, new species
Plate XIX, Figure 1
Shell sub-ovate, convex; surface with about 36 spinose ribs and narrower,
rounded, transversely striate interspaces; interior of valves grooved and
scalloped at ventral margins. All ribs except the three anterior ones have a


lateral keel on each edge at the top. The three anterior r.bs have only one
such keel, placed on the anterior side. The first ten ribs (including the three
single keeled ones) are ornamnted with longitudinally ovate, long, erect, or
slightly hooked spines along the anterior keel, but spreading out over the
center of the rib. All the remaining ribs have the spines on the posterior
side. From the eleventh rib to the eighteenth inclusive, the spines are like
those of the ten anterior ribs, but commencing with the nineteenth rib
the spines posteriorly become narrower, blade-like, set longitudinally on the
rib, and curved posteriorly over the adjacent interspace. All the ribs are
flat topped, but the posterior ribs have the tops sloping posteriorly. Length
30 mm., height 32.5 mm., semi-diameter 11 mm.
This shell is twice as large as C. bowdenense Dall. The orna-
mentation especially of the rib is nearer to that of C. muricatum
than to that of any other species. It is apparently related to C.
(Trachycardium) gatunensis Toula, and is one of the early members
of the G. muricatum group.
Type locality.-179.
Other localities.-225, 24?, and (doubtfully) near Guanica
Central (Lobeck).
Range.-Lares limestone, Cibao limestone, Los Puertos limestone.

Cardium (Trachycardium) cinderellae Manry
variety alternatum, new variety
Plate XIX, Figures 2, 3, 4
Shell equivalve, slightly inequilateral; beaks high, strongly incurved;
hingd line rather long; surface with about thirty-six broad, flat ribs with
narrow interspaces; ribs appear medially grooved when worn smooth. When
perfect, they show a single row of high, vaulted, inverted V-shaped spinose
scales, which cover the width of the rib, and are somewhat unevenly spaced,
though not crowded. There is a strong tendency toward alteration of ribs
with large and ribs with small spines. This is more marked in some speci-
mens than in others. It is a feature not mentioned in Maury's description of
the Santo Domingo shell, but shows in the illustration of the latter. The
interspaces are faintly grooved radially, and strongly transversely striate;
valves internally grooved, and inner margins strongly scalloped ventrally.
Length 29 mm., height 33 mm., diameter 26 mm.
The sculpture is like C. cinderelle Maury, but the shell is larger,
shorter, and more convex. It may possibly deserve a specific name,
but the specimens are too fragmental to warrant this.
Localities.-225, 226, 280, 281, 331.
Horizon.-Lares formation.


Cardium (Trigonocardia) sambaicum Maury

variety portoricoensis, new variety

Plate XIX, Figures 5, 6

No complete shells were found, but fragments large enough to
show the outline of the shell. They seem to be quite variable. There
are some specimens in which the ribs are worn smooth as in Maury's
illustration from Santo Domingo. Some specimens show the ribs
slightly wider than the interspaces, but usually the reverse is true,
especially in the ventral portion of the shell. Fine raised concentric
threads cross the interspaces and cross the ribs between the nodes.
These threads are much finer and more crowded where they cross the
ribs. In the interspaces they are very irregular, and frequently
end short at the center of the interspace. These terminated threads
have a tendency to alternate from opposite sides of the interspace
as though shearing or faulting had displaced them. The nodes on
the ribs are high and prominent. The shells are smaller and le.s
convex than the Santo Domingo form, but the number of ribs, trun-
cation and outline of the shell are about the same. Length 13 mm.,
height 13 mm., semi-diameter 5 mm.
Horizon.-Quebradillas limestone.

Cardium, new species (aff. C. sambaicum Maury)

Plate XIX, Figure 7

Fragment indicating a shell of the size and general outline of
C. sambaicuim Maury, but with a somewhat different sculpture of the
ribs. Shell with about 25 strong ribs, high and rounded on top, and
slightly narrower U-shaped interspaces. The whole is crossed 1y
fine, close concentric threads, quite irregular. and of which every
second, third or fourth one on crossing the ribs expand into broad
semi-cone shaped scales the pieces of which are toward the beak.
These scales are irregularly spaced, and are especially crowded near


the ventral margin. Where the ribs are worn, these scales appear as
thin arched lamellae. Length of shell about 20 ninm.
Horizon.-Quebradillas limestone.

Cardium (Trigonocardia) haitense Sowerby

variety cercadicum Maury

Plate XIX, Figure 8
Cardium haitense Sowerby, Quart. Jour. Geol. Soc. London, VI, p. 52, PI. 10,
fig. 11, 1849 (in part).
Cardium (Trigonocardia) haitense variety cercadicum Maury, Bull. Amer.
Paleont., V., No. 29, p. 212, Pl. 36, fig. 6, 1917.
This shell corresponds closely to Maury's variety cercaidicum of
Santo Domingo. The truncation is sharp; ribs with inverted Y-
shaped nodules; ribs on the truncation have a few scattered spinose
nodules; ribs flat, those anterior to the truncation having the flat
tops sloping anteriorly, thus giving the ribs an asymmetric V-shaped
cross-section. There are nine ribs on the truncation, and fourteen
on the main body of the shell. Concentric threads in the interspaces,
which, in the anterior part of the shell occasionally cross the ribs.
The nodules on the ribs are rather low and indistinct except on the
rib which marks the truncation, where they are prominent. Length
10 mm., height 11 umm., semi-diameter, etc., 5 mm.
Localities.-204, 52(a), 192, 77 ? (Berkey).
Horizon.-Quebradillas limestone (characteristic), and in the
Ponce limestone.

Cardium (Trigonocardia) haitense Sowerby

variety areciboense, new variety

Plate XIX, Figure 9

Very close to C. haitense cercadicum Maury', but shorter and more
sharply truncate, and with fewer, coarser ribs which are more de-


cidedly angular. The ribs on the truncation near the dorsal margin
have regularly spaced V-shaped spines, pointing sharply upward.
The nodules and concentric threads are otherwise exactly like those
of variety cercadicum. There are six ribs on the truncation and
eleven on the remainder of the shell. Length 10 mm., height 14
mim., semi-diameter 6 mm.
Locality.-204. Also Berkey 77.
Horizon.-Abundant in the Quebradillas limestone and occurring
also in the Ponce limestone.

Genus 01ementia Gray

Clementia dariena Conrad

Plate XIX, Figures 10, 11, 12

Meretrix dariena Conrad, Pac. R. R. Rep., V, p. 328, Pl. 6, fig. 55, 1856.
Clementia dariena Gabb, Jour. Acad. Nat. Sci. Phila. (2) VIII, p. 344, Pl. 44,
fig. 16, 16a, 1881.
Clementia dariena Dall, Trans. Wag. Inst. Sci., III, pt. G, p. 1235, 1903.
Clementia dariena Toula, Jahrb. der K. K. Geol. Relchanst. Wien, LVIII,
p. 725, Pl. 27, figs. 9, 10, 1908.
Clenentia dariena Brown & Pilshry, Proc. Acad. Nat. Sci. Phila., p. 371, Pl. 28.
fig. 1, 1911.

The typical Porto Rican shells are smaller than those of the
Panama region and Costa Rica, but otherwise show no marked dif-
ferences. The Porto Rico shells are not dwarfed forms, but merely
arrested in growth, the average adult being an exact replica of a
Gatun shell when the latter is two-thirds grown. This might lead
to the supposition that the Porto Rican analogue is a primitive form
of the Gatun shell, except for the fact that the former occasionally
reaches a size as great as the average Gatun shell and is exactly
simliar, even to the loss of ribs near the ventral margin. To em-
phasize this point, an average specimen from Dr. Kemp's Gatun
collection (in the Columbia University museum) is illustrated here
together with one of the large Porto Rican shells, for comparison.
A comparison of the Porto Rican shells with available sources is
shown in the following table.








of ventral
means .

Gabb, illustration. Sapote,
Costa Rica............. 57 47.5 33.5 26-35 10

Brown & Pilsbry, illustra-
tion. Gatun............ 65 63 ? 26? 18

Toula, illustration. Gatun.. 67 56 ? 28 19

Toula, description. Gatun.. 47 38 ? ? ?

IKemp collection, specimen.
Upper Gatun........... 63 61.5 36 33 23

Kemp collection, specimen.
Lower Gatun........... 51 43 25.5 31 7

do .................. 48 44 24.5 30 10

do (young shell).. .... 27 25? 14 22? 0

Porto Rico, San Sebastian
shale at Collazo Falls....

Porto Rico, San Sebastian
calcareous shale at Robles,







do.................. 39 31 23

do0 .................. 38 33.5 20

do ........ ...... ... 38 35.5 18.5

do.................. 37.5 33 19.5

do.......... ........ 35 32.5 20

do .................. 35 32.5 17.5

do.......... ........ 35 33 17

do.................. ? 34 20

do.................. ? 33 15

do.................. ? 32 20

do .................. ? 29 17.5

do .................. ? 28 19

27 13



It will be noticed from this table that there is considerable varia-
tion in size and proportions among the specimens. This is due in
part to distortion by pressure as regards the proportions. If the
average of these measurements is taken and the height and diameter
expressed in terms of the length as unity, we have the following
Length Height Diameter Average
Gabb's illustration. S a p o t e,
Costa Rica ............. 1 0.83 0.59 57
Kemp's large specimen from
the upper Gatun........ 1 0.98 0.57 03
Kemp's smaller specimens
from the lower Gatun... 1 0.SS 0.51 42
Large Porto Rico specimen
from the Collazo district 1 0.93 0.54 48
Smaller Porto Rico specimens
from the Robles district 1 0.87 0.51 37
It will be noticed that in the larger specimens there is a relatively
greater increase in the height and diameter, due to the slight change
in shape of the shell with growth. The measurements from Gabh's
illustration are of doubtful value in this comparison.
The uniformity of size and number of ribs is a feature brought
out in the table. The Porto Rican shells have about the same number
of ribs as the Gatun shells, and in the ribbed stage are about the same
size. The Gatun shells usually, and the Porto Rican shells occa-
sionally grow beyond this stage with loss of ribs, an ontogerontic
character. The Gatun shells were longer lived, apparently living
under more favorable conditions than those of the upper San Sebas-
tian shale in Porto Rico.
Localities.-232(b), 233, Collazo Falls, S,, 84, Juana Diax.
IHorizon.-San Sebastian shale, Juana Diaz shale.

Genus Cytherea Bolten

Cytherea (Cytherea) berkeyi, new species

Plate XX, Figures 1, 2, 3

Shell large, convex, very inequilateral; right valve with two posterior
bifid and one anterior simple cardinal teeth; left valve with one middle bifid
and anterior and posterior simple cardinals; left anterior papilliform small


out distinct; adductor scars and pallial line strongly impressed, the latter
granulose; pallial sinus large, broad, rounded, ascending; inner margins
minutely crenulate; small auxiliary adductor scars, situated close to dorsal
margin between beaks and principal anterior adductor scars, and nearer to
the latter; obscure, irregular impressed line in interior of each valve, descends
vertically from anterior end of beak and dies out midway between beak and
ventral margin; surface of valves marked by numerous small radial ribs,
somewhat wavy, evenly spaced, and fairly uniform in size, but becoming
coarser over the posterior third of the shell, where they alternate in strength,
with occasionally two or three small ribs alternating with one lagrer one;
the whole cut by fine, low, sharp, raised concentric lamellae, evenly spaced
near the beaks, but gradually increasing to 2, 3, or 4 mm. apart toward the
ventral margin, where they suddenly become irregularly spaced and crowded,
apparently a gerontic character. These concentric lamellae are delicately
fluted by the radials in the umbonal region. Minute growth lines cross the
sculpture between the lamellae. The concentric sculpture is nowhere as
prominent as the radial which increases in coarseness ventrally. The largest
specimen, an internal mold collected by Dr. Berkey, measures in length 85
mm., height CS mm., diameter 48 mm.
A large shell of the same type as 0. tarquinia Dall and also resem-
bling 0. lacerata HIanley, a recent shell from China, and U. antiqua
King, recent, from Chile and Patagonia. It differs from all of these
in having the radial sculpture predominate over the concentric
Localities.-31S, 41, 78, 9S(Berkey). Bayamon and Ponce
Horizon.-Quebradillas limestone Ponce limestone.

Genus Pitaria Romer

Pitaria (Hyphantosoma) carbasea Guppy

Plate XVI, Figure 2
Cytherea (Circe) carbasea Guppy, Quart. Jour. Geol. Soc., XXII, p. 292, Pl. 18,
fig. 13, 1866.
Callista carbasea Gabb, Trans. Amer. Phil. Soc., XV., p. 250, 1873.
Pitaria (Hyphantosoma) carbasea Dall, Trans. Wagner Inst. Sci., III, pt. 6,
p. 1266, 1903.
This shell occurs abundantly as internal molds, but only one frag-
ment of external mold was found showing the delicate sculpture.
The pallial sinus is sub-angular, deep, reaching nearly to the middle
of the valve; interior of shell smooth; exterior surface marked by
fine, somewhat wavy or irregular concentric lamellae crossed by fine



divaricate lines, strongest anteriorly, and forming two zigzag zones
near the middle of the valve. These divaricate lines, when viewed
under a lens are seen to consist of rows of pits in the lamellae;
lunile large, demarked by fine sharp impressed line, and crossed
faintly by .the concentric lamellae, thus being rendered indistinct.
The shells vary considerably in size, the largest internal mold
measuring in length 32 mm., height 26 mmn., diameter 16 mm.
Type localities.-64, 225.
Other localities.-619, 233, 343, 232(b).
lange.-San Sebastian shale, Lares limestone, and Cibao lime-
stone. Abundant in the Cibao limestone.

Genus Chione Megerle von Miihlfeldt

Chione woodwardi Guppy

Plate XIII, Figure 7
Venus Woodwardi Guppy, Quart. Jour. Geol. Soc. London, XXII, p. 292, Pl. 18,
fig. 1, 1866; Idem. XXXII, p. 530, 1876.
Chione Woodwardi Dall, Trans. Wagner Inst. Sci., III, pt. 6, p. 1291, 1903.
Chione Woodwvardi lMaury, Bull. Amer. Paleont., V, No. 29, p. 218, P1. 37, fig. 6,
This is one of the most abundant fossils in the Quebradillas lime-
stone. The shells show no differences from those described from
Jamaica and Santo Domingo.
Localities-204, 217, 52(a), 670, 179, 225. Near Bayamon
(Semmes and Lobeck). Near Guanica Central (Lobeck). Vieques
Island (Lobeck). 77 (Berkey). Culenbrinas Point, Ponce, and
K. 2.4 Ponce-Adjuntas road (Mitchell).
Horizons.-Lares limestone ?, Quebradillas limestone, Ponce lime-

Chione (Lirophora) hendersoni Dall
Venus paphia Guppy, Quart. Jour. Geol. Soc. London, XXII, p. 292, 1866. Not
of Linn6.
Chione paphia Gabb, Trans. Amer. Phil. Soc., XV, p. 249, 1873.
Venus paphia Guppy, Geol. Mag., decade 2, I, p. 442, 1874, etc., ex parte.;
Quart. Jour. Geol. Soc., XXXII, p. 530, 1876.


Chione (Lirophora) hendersoni Dall, Trans. Wagner Inst. Sci., III, pt. 6,
p. 1295, Pl. 55, fig. 22, 1903.
Chione (Lirophora) henulersoni Maury, Bull. Amer. Paleont, V, No. 29, p. 219,
Pl. 37, fig. 8, 1917.
The Porto Rican shells are apparently closer to the Santo Domingo
than to the Bowden shells. The specimens show the obscure crenula-
tion at base of ribs, and the sudden thickening of the ribs over the
anterior portion of shell. There are eleven or twelve large, widely
spaced ribs. They are obsolete in the umbonal region. Interior of
shell has deeply impressed pallial line and adductor scars; pallial
sinus short and angular. An average specimen measures in length
31 mm., height 22 mm., diameter 14.5 mm. The largest measure
in length 36 mm., height 27 mm., semi-diameter S mm.
Localities.-179, 331, 343, 64, 619 ?.
Range.-Lares limestone to Los Puertos limestone.

Genus Tellina (Linn6) Lamarck

Tellina strophoidea, new species

Plate XX, Figures 4, 5
Shell moderately elongate, sub-equilateral; beaks inconspicuous; surface
with numerous elevated, sharp concentric lamellae, which, on adult specimens,
frequently merge on the posterior part of shell into fewer, coarser lamellae;
numerous radial striae can be seen under a lens in transverse light. Ven-
trally they are interrupted by the lamellae, but toward the beaks they become
more distinct and continuous, so that on the nuclear portion of the shell they
are nearly as strong as the concentric sculpture, resulting in an extremely
fine cancellate ornamentation; posterior slope with two radial folds and a
moderately deep sulcus between. The upper fold is narrower and less pro-
nounced; posterior dorsal margin marked by accentuation of the lamellae
into a slight ridge. In the adult shells the lowest fold is, in the right valve,
double, with a slight sulcus between. This is only noticeable at the distal
end of the fold. The lamellae bend sharply upward on crossing the lower
fold; lunule extremely narrow; interior of shell unknown. Length 48 mm.,
height 27 mm., semi-diameter 6 mm.
This shell is nearest to T. strophia Dall, which it resembles in
sculpture, but is considerably less elongate.
Localities.-204, 216, 98(Berkey).
Range.-Throughout the Quebradillas limestone.


Tellina portoricoensis, new species
Plate XX, Figure 6
Shell resembling T. strophoidea but with much finer concentric sculpture.
Shell sub-equilateral, slightly convex; beaks low; surface with fine, sharp,
hardly raised concentric lamellae, at first uniform and evenly spaced, but
ventrally becoming slightly irregular, with every fifth or sixth one more
prominent than the others; low, inconspicuous double fold on the umbonal
ridge, becoming almost obsolete distally, and over which the concentric
lamellae are flexed sharply upward in crossing; fine obscure growth lines
between the lamellae; faint radial rays seen only in oblique light; interior
of shell unknown. Length 42 mm., height 23 mm., semi-diameter 5 mm.
Horizon.-Quebrad illas limestone.

Tellina (Scissula) grabaui, new species
Plate XVT, Figures 3, 4
Shell small, elongate, slightly convex, sub-equilateral; anterior rounded,
posterior roundly truncate; surface with fine growth lines, periodically in-
cised, replaced on the umbonal ridge by widely spaced, sharp concentric
lamellae, which cross a faint sulcus above the umbonal ridge and, decreasing
in size and strength, continue to the dorsal margin; growth lines crossed
by faint, wavy oblique lines, becoming more distinct as they approach the
umbonal ridge, where they end suddenly, in contact with the concentric lamel-
lae. These oblique lines begin near the anterior margin and slope downward
toward the posterior. Interior of shell unknown. Length 20 mm., height
11 mm., semi-diameter 2.5 mm.
This shell is close to T. (Scissula) lampra Dall, of the Chipola
Oligocene. It is also close to T. rowlandi Toula, a species from the
Gatun formation. T'. caribea d'Orbigny seems to be the living rep-
resentative of these shells. T'. (Scissdula) cercadica Maury, a shell
from the Santo Domingo Oligocene, is a similar but less elongate
Horizon.-Qcebradillas limestone.

Tellina, species aff. T. (Angulus) atossa Dall
Plate XI, Figure 7
A single left valve of a Tellina of uncertain identity. Shell
moderately convex, showing prominent lateral teeth; surface with


fine sharp concentric lamellae, strongest near the ventral margin,
and obscure growth lines between the lamellae. The latter are ac-
centuated on the anterior slope; shallow radial sulcus located next
to the posterior dorsal margin. Interior of shell unknown. [n out-
line and general appearance, it resembles T. (Angulus) atossa of the
Tampa Silex beds. Length about 20 mm., height 14 mm., semi-
diameter 4 mm.
Ilorizon.-Quebradillas limestone.

Genus Metis H. and A. Adams

Metis trinitaria Dall

Plate X, Figure 7
Metis trinitaria Dall, Trans. Wagner Inst. Sci., III, pt. 5, p. 1041, Pl. 46, fig.
24, 1900.
cf. Tellina biplicata Guppy, Quart. Jour. Geol. Soc., XXXII, p. 530, 1876. Not
of Conrad.
Melis trinitaria Maury, Bull. Amer. Paleont., V, p. 226, 1917.
The specimens differ somewhat from Dall's description, being
more nearly equilateral, and the pallial sinuses of the two valves
being subequal and of equal height. The shell seems to be quite
variable, however. Ventral margin arcuate, with broad median sinus
in the right valve: sharp, rounded posterior fold with a second
obscure fold higher up: surface marked by growth lines and faint
radial striations, the latter fading out toward the lateral margins.
There is some variation in the size and proportions as shown by the
following measurements of internal molds:

Length 59 mm., Height 44 mm., Diam. 23 mnm.
Length 54 umm., Height 44 mm., Diam. 19 mm.
Length 48 mm., Height 39 mm., Diam. 16 mm.

Some internal molds of this shell, collected in Porto Rico in 1915
by Dr. D. R. Semmes, were shown to Dr. Dall, who identified them
as AM. trinitaria. This species is reported from Trinidad and Cuba,
and doubtfully from Santo Domingo. Specimens from Sombrero,


W. I., in the Columbia University collections seem to be of thiL'
species, but are considerably larger than the Porto Rican shells.
Localities.-204, 678, 61, 319, S01. 9S(Berkcy). San Juan
district (Semmes). Culebrinas Point (Mitchell).
Horizon.-This shell is the chief horizon marker of the Quebra-
dillas limestone along the north coast. The internal molds are found
in great numbers in many localities. It occurs rarely in the Los
Puertos limestone, and Ponce limestone.

Genus Solen Linn6

Solen (Plectosolen) collazoensis, new species
Plate XI, Figure S
Shell produced anteriorly, with beaks 7.5 mm. behind the anterior margin;
both ends roundly truncate; height slightly less in middle of shell than at
the ends; beaks inconspicuous; surface marked only by growth lines,
strengthened on anterior slope, and a narrow furrow extending from the
beaks to the anterior-ventral corner. Length 42 mm., height 11 mm., diameter
6.5 mm.
This is a small straight shell of the section Pleclosolen, having a
strong resemblance to the Eocene forms of the Paris Basin, such as
S. plagiaulax Cossmann, and S. larersinensis Lef. and Wat. It is
probably of the type of S. obliquns Sowerby. It also resembles ;1
imnall S. obliquus Spengler, but is a true Plectosolen.
The description is based on a single specimen from the black shale
at Collazo Falls. The specimen is badly fractured, though nearly
Localities.-261, Juana Diaz, Yauco.
Horizon.-San Sebastian shale, Juana Diaz shale.

Genus Psammosolen Risso

Psammosolen sancti-dominici Maury
Psammosolen saucti-dominici Maury, Bull. Amer. Paleont., V, No. 29, p. 228,
P1. 37, fig. 13, 1917.
Only two fragments were found, but the marking is distinct and
characteristic, and leaves no doubt as to the identity. One specimen
seems to be about the same size as the Sanuto omingo specimens


described by Mlaury. The other specimen is smaller. The broad
median sulcation is barely noticeable.
Horizon.-Quebradillas limestone.

Genus Semele Schumacher

Semele, species indet.

Shell equivalve, nearly equilateral; beaks low; ends of shell rounded;
anterior and posterior dorsal margins make angle of 1250; umbonal ridge
low, obscure; escutcheon long and narrow; lunule short, and same width as
the escutcheon; surface sculpture of fine, even, sharp concentric lamellae;
about fourteen to every 10 mm.; no radial striation visible. Length 25 nmm..
height m0 mm., diameter 14 mm.
This may be a new species, but the specimens are poor and
do not permit an exact description, being known only from internal
molds and a few fragments of external mold. The sculpture is finer
than that of S. claytoni 21aury. The shell resembles ,5. chipolana
Dall, but is more elongate.
Horizon.-Lares limestone.

Genus Thrachia Blainville

Subgenus Cyathodonta Conrad

Cyathodonta reedsi Maury

Shell equivalve, sub-equilateral, moderately convex; anterior dorsal slope
slowly descending, anterior rounded, posterior roundly truncate; sculpture
of about twenty-five rounded to sub-angular concentric ripples, markedly
wavy or irregular, and owing to the thinness of the shell, equally strong in
the internal molds. The ripples have a strong tendency to become oblique
on the anterior third of the shell; umbonal ridge low, rounded, beyond which
the ripples die out, leaving the posterior dorsal smooth except for growth
lines. A right valve shows a low obscure radial fold about one-third the
distance from the umbonal ridge to the dorsal margin; beaks rather high
and strongly incurved. Gerontic individuals show a complete loss of con-
centric ripples in the final stages of growth. Adult shell measures in length
27 mm., height 19 mm., semi-diameter 6 mm.
This shell is apparently nearest to C. galuensis Tonla but is more


elongate, more nearly equilateral, and the smooth posterior slope
higher and more produced. It is not as convex as the Gatun shell.
Type Locality.-204.
Other locality.-225 ?.
Lange.-Lares ? limestone to Quebradillas limestone.

Genus Corbula Lamarck
Corbula collazica Manry
Localities.-Collazo Falls, and doubtfully at 120 and 225.
Horizons.-Typical of the San Sebastian shale, and "occurs doubt-
fully as high as the Lares limestone.

Genus Teredo Linnu
Teredo incrassata Gabb
KIaphus incrassatus Gabb, Trans. Amer. Phil. Soc., XV, p. 246, 1873; Jour.
Acad. Nat. Sci. Phila., new ser., VIII, p. 342, Pl. 44, fig. 12a-e.
Teredo fistula? Guppy, Quart. Jour. Geol. Soc., XXXI I, p. 529, 1876.
Teredo incrassata Dall, Trans. Wagner Inst. Sci., III, pt. 6, p. 1587, 1903.
Teredo incrassata Maury, Bull. Amer. Paleont., V, No. 29, p. 235, Pl. 39, fig. 24,
The tubes vary in diameter, but average 20 mm. They attain
considerable length, but are usually found broken in short pieces.
Localities.-S6, 10, Collazo Falls, 758, 102, 182, 179, 332, 619,
621, 61(, 225, 23, 335, 330, 324, 74, 370, 749(a), 334.
Horizons.-San Sebastian shale, Lares limestone, Cibao limestone,
Ios Puertos limestone, Quebradillas limestone. Extremely abundant.
Outside the Lares District, this species has been found by Lobeck
on Vieques Island (south and east sides), and in basal Tertiary beds
near Yauco, P. R. It has been found by Berkey at localities 10.
and 78, and by Mitchell in the Ponce formation and Juana Diaz

Genus Dentalum Linn6
Dentalium, species indet.
Shell long, slender, nearly straight, slowly tapering, circular in cross-
section; shell thick, the aperture being only about one-half the diameter of
the tube; surface polished, and marked with numerous fine, very regular,


longitudinal threads, visible only under a hand lens and transverse light.
The threads appear, under a lens, to lbe minutely headed, but under a micro-
scope, this appears to be the result of fine raised concentric growth lines,
strongest in the interspaces between the longitudinal threads. There are
about one hundred longitudinal threads on a tube of 1 mm. diameter. The
shell structure is strongly librous, the fibrous striations appearing on the
longitudinal threads as seen under the microscope. The apex is not shown
in any of the specimens. The largest fragment, with a diameter of 1 mam.
measures 13 mm. in length. The tapering of the tube is hardly discernible
in this length.

Many fragments were found in the lower black shale of a minute
Dentalium, the largest of which is only about 1 mm. in diameter,
and the majority of which average less than 1/2 mm.
Horizon.-San Sebastian shale.

Genus Calliostoma Swainson
Calliostoma portoricoensis, new species
Plate XXT, Figure 1
Whorls seven, including a rather high protoconch of two smooth, round
whorls; remaining whorls flat; suture narrow but deeply impressed; surface
of whorls with six spirals and strongly marked oblique growth lines which
cut the spirals, giving them a wavy appearance. The five uppermost spirals
are small, sharply raised, with wider concave interspaces. The two spirals
immediately below the suture are somewhat smaller and show a tendency to
group together. The sixth spiral, immediately above the suture, is very much
larger, broad, and bluntly rounded, and forms the peripheral keel; base with
eight prominent, Lroad, flat spirals and fine raised oblique growth lines
crossing the spirals and grooves between. Aperture and umbilical area not
visible in the specimen. Leugth 11 mm., diameter 9 mm.

A single specimen was found of a Calliostoma remarkable for its
elongate conical base (more than two-thirds as long as the spire).
Horizon.-Quebradillas limestone.

Genus Turbo Linn6
Turbo fetkii, new species
Plate XXI, Figure 2
Shell small; whorls six; the first two flat topped, strongly keeled; the
remaining whorls sub-round, but with slight angulation and broad flat


shoulder, nearly one-third the axial width of the whorl; suture deeply
channelled; rounded spirals of two to three different sizes, increasing by
intercalation, with total of about twenty on the body whorl, and of these
five are very prominent; entire surface crossed by oblique incremental lines,
strongest between the spirals; spirals and incremental lines visible in the
sutural channel. Height 13 mm., diameter 11 mm.

The above description is based on a single specimen. in which the
anterior portion of the body whorl is missing. The shell is remark-
able for its extremely deep, squarely channelled sutture. The spirals
are smooth or with microscopic erenulation produced by the crossing
of the growth lines. The spiral anterior to the suture is never promi-
nent. This shell is like T. rhectogrammicus Dall, of the Florida
Pliocene, but differs from the latter chiefly in its smaller size and its
deeply channelled suture. Its suture is like that of T. sp(nileriautts
G-melen, a recent shell from the West Indies, but the latter is much
larger and has more spirals. This shell is named in honor of Dr.
C. R. Ietke, who has worked on the Geology of the eastern end of
Porto Rico.
Horizon.-Quebradillas limestone.

Turbo dominicensis Gabb

variety quebradillensis, new variety
Shell larger than the Santo Domingo variety, T. dominicensis laloi Mairy,
the Internal mold measuring in height (with tip of spire missing) 37 mm.,
maximum diameter 33 nun. Like the Santo Domingo variety, the final whorl
is angulated, as presumably are the earlier whorls, though this cannot lie
ascertained from the specimen. It is more like Gabb's type, however, in
having the majority of the spirals crenulate. On the final whorl the spiral
anterior to the suture is more strongly crenulate than the others. Anterior
to this, on the shoulder, are four minor crenulate spirals, the two central
ones being slightly more prominent than the other two. At the shoulder
angulation is a very strong, plain spiral, which projects sharply upward.
above the surface of the shoulder. The remainder of the whorl is orna-
mented with eight or more strong simple spirals, each alternating with a
single minor crenulate spiral. The basal portion of the whorl is not pre-
served. From the small portion visible in the specimen, it setms that the
ornamentation of the earlier whorls is similar, but with fewer spirals. The
outer lip is lirate for a short distance, there being thirteen lirae and a pos-
terior notch below the shoulder angulation.

This shell was found among hlie material collected by Dr. Lobeck
in the Quebradillas limestone near I ayamon. Tt consists of a single


internal mold and a portion of the external mold of the same speci-
men, which shows the sculpture of the final whorl and a small por-
lion of the spire. A few internal molds, possibly of the same species
were found in the Quenradillas limestone of the Lares district.
Locality.-Near Bayamnion (Lobeek), 204 ?
lHorizon.-Quebradillas liniestone.

Genus Liotia Gray

Liotia (Arene) coronata Dall

variety portoricoensis, new variety

Plate XXI, Figure 3
Shell small, five whorls, including flat, smooth naticoid protoconch of
two whorls. Two spiral keels appear on the third (or first post-nuclear)
whorl. A third spiral appears on the fourth whorl just above the suture;
suture deeply channelled; shoulder of third and fourth whorls marked by
fine beaded spiral ridge immediately below the suture. Fine growth lines
crossed the shoulder and produce a slight beading of the uppermost spiral
keel; final whorl smooth except for strongly incised growth lines, which are
stronger than the almost obsolete spirals. The final whorl tends to lose the
coiling habit, becoming almost free from the preceding whorl. Only a single
specimen was found, and the aperture and base of the shell are unknown.
Height 5 mm., diameter 5 mm.
The first four whorls of this shell are quite similar to Dall'
Tampa species. In the Porto Rican shell the longitudinal wrinkles.
characteristic of the Tampa shell, become obsolete on the fourth
wvhorl, but reappear strongly on the last whorl. The Porto Rican
variety seems to have been derived from the Tampa species, and is
an interesting example of a phylogerontic form.
Ilorizon.-Quebradillas limestone.

Genus Xenophora Fischer de Waldheim

Xenophora conchyliophora (Born)
Trochus conchyliophorus Born, Mus. Caes. Vind. p. 333, 1778.
Trochus agglutinans Lamarck. Anim. sans Vert., VII, XIV.
Phorus agglutinans Gabb, Trans. Amer. Phil. Soc., XV, p. 241, 1873,


Xenophora conchyliophora Dall, Trans. Wagner lust. Sci., Ill, pt. 2, pp. 360-
362, 1892; Bull. 90, U. S. Nat. Mus., p. 105, Pl. 15, figs. 1, 3, 1915.
Xenophora cunchyliophora Alaury, Bull. Amer. Paleont.. V, No. 29, p. 133,
Pl. 23, fig. 7, 1917.
The Porto Rican specimens show no differences from the typical
forms of this species.
Localities.-204, 225, 24.
Range.-Lares limestone to Quebradillas limestone. Very abun-
dant in the Quebradillas.

Genus Neritina Lamarck
Neritina (chipolana var.?) collazoensis, new species
Plate XXT, Fizgre 4
Shell small, fragile, polished, with low spire, and apparently three whorls.
A single specimen was obtained, in which the spire has been crushed and
the aperture is not visible. The outline of the shell and the surface mark-
ings are like N. chipolana Dall, but with the markings more crowded and
numerous, and all being discontinuous. These lines show, however, the same
arrangement in groups of three or four, with zig-zags and curves. The curves
are frequently convex in opposite directions in adjacent lines, leaving circular
areas between the lines, a feature which is also characteristic of the Chipola
species, judging from Dall's illustration (Wagner Inst., III, pt. 4, Pl. 23,
fig. 19, 1898). The lines are very narrow, and a dark reddish purple in
color. The shell is slightly smaller than the Chipolan form, the height
being 4 mm. and the maximum diameter 4.5 mm. In places where the epi-
dermis has been chipped off, exceedingly fine and very regular growth lines
may be seen with a lens.
This shell was obtained from a stratum of lignitic shale in the
Collazo Falls section, and is associated with fossil leaves and minute,
thin-shelled bivalves, apparently fresh water forms. This stratum
is intercalated in marine beds, and is of interest in showing' the con-
ditions existing in this locality during deposition of the San Seba,-
tian shale.
Locality.-2 61.
Ilorizon.-San Sebastian shale.

Genus Hipponyx Defrance
Hipponyx portoricoensis, new species
Plate XXT, Figures 5, 6
Shell small, obliquely conical; apex at or near posterior extremity, often
overhanging, and slightly recurved; surface with irregular bifurcating radial


ribs and less prominent fluted concentric lamellae; Interior with faint but
large horse-shoe shaped scar.
This shell is very variable, but in all specimens the apex is blunt
and rounded. Some specimens show hardly any recurvature to the
apex, while in others this feature is marked, and in all cases it is
to be left. A strongly recurred specimen measures in diameter of
aperture about 7 mm., height of apex above base 1 mm. A low flat
shell without curvature measures in diameter 10 mm., height of apex
3 mm. The largest specimen measures in diameter 11 mm., height
of apex 6.5 mm.
Localities.-204, 220, 23 ?.
Range.-Los Puertos limestone to Quebradillas limestone.

Genus Calyptraea Lamarck
Calyptrsea cf. centralis Conrad
Infundibulam centralis Conrad, Am. Jour. Sci., XLI, p. 348, 1841; Med. Tert.,
p. 80, Pl. 45, fig. 5, 1845.
Trochita centralis Emmons, Geol. N. C., p. 276, fig. 193, 1858.
Infundibulum concentricum H. C. Lea, Trans. Amer. Phil. Soc., (2), IX, p. 249,
P1. 35, fig. 39, 1845 (young shell).
Crochita collinsii Gabb, Jour. Acad. Nat. Sci. Phila., (2), VIII, p. 342, Pl. 44,
figs. 11, lla, 1875.
Infundibulum candeanum d'Orbigny, Moll. Cuba, II, p. 190, Pl. 24, figs. 28, 29,
Galerus candeanits Dall, Hemphill's Shells, p. 335, 1883.
Galerus parvulus Dunker, Jahrb. Deut. Mal. Ges., p. 244, 1875.
Calyptrea candeana Dall, Blake Rept., p. 284, 1889.
Calyptrea centralis Dall, Trans. Wagner Inst. Sci., III, pt. 2, p. 353, 1892.
A single external mold was found which is apparently this species.
Diameter of base 8 mm., height of apex 4 mm.
Horizon.-Quebradillas limestone.

Genus Crucibulum Schumacher
Crucibulum auricula Gmelin
variety portoricoensis, new variety
Plate XXI, Figures 7, 8
Cup typical of C. auricula Gmelin. Surface with fine, Irregular, dichoto-
mus radial lines, crossed by fine wavy growth lines. Form varies from


high, acute apex to low apex with broad base. Cup uniform in size and
shape in all the specimens. A shell with low apex measures in diameter of
base 20 mm. height of apex 7 mm.
Localities.-204. (also Berkey 9S).
Horizon.-Quebradillas limestone (fairly abundant).

Crucibulum auricula? variety
Plate XXI, Figure 9
This is probably one of the many variations of C. auricula. Only
a few fragmental specimens were obtained, and the interior is un-
Apex sub-central; coarse radial ribs, somewhat irregular, bifur-
cating and trifurcating. Diameter 14 mm., height of apex 6 mm.
Horizon.-Quebradillas limestone.

Crucibulum (Dispotaea) collazum, new species
Plate XXI, Figure 10
Shell small, smooth; apex eccentric, sharp, strongly gyrate in the normal
direction; basal margin sub-circular, with large rounded sinus on the side
below the apex. On either side of the sinus the margin laiaes, or flattens
out, and is thickened, being analogous to the thickened outer lip or varix of
coiled gastropods; cup large, deep, and about one-half free from the shell
wall. Most of the free half has been broken away in the specimen illustrated,
but when perfect, it has a tortuous, or S-shaped curvature, and overhangs the
apex cavity, being suggestive of the internal appendage of a Calyptrwa.
Diameter of aperture 7 mm., height of apex 4 mm.
Two specimens were collected, both showing the marginal sinus,
which is therefore assumed not to be an accident of growth. The
shell illustrated appears to be an adult specimen. The other shell
is smaller, with a correspondingly smaller marginal sinus.
Localities.-S4, 261.
Ilorizon.-San Sclastian shale.

Genus Natica Scopoli
Natica canrena? (Linne) \loorch
Nerita canrena (Linnd in part) Auct., Moerch, Mlalak. Blatt., XXIV, p. 62, 1877.
Natica canrena Gabb, Trans. Amer. Phil. Soc., XV., p. 223, 1873.


Natica canrena Guppy, Quart. Jour. Geol. Soc., XXXII, p. 518, 1876.
Natica canrena Dall, Trans. Wagner Inst. Sci., III, pt. 2, pp. 364-365, 1892.
Natica canrena Brown and Pilsbry, Proc. Acad. Nat. Sci. Phila., p. 508, 1912.
Natica canrena Maury, Bull. Amer. Paleont., V, No. 29, p. 134, Pl. 23, fig. 10,
A small Natica was found which is probably the young of N.
can rena. The specimen shows on the last whorl the faint oblique
striations, which start at the suture. Diameter 7 mm. Several in-
ternal molds, possibly of this species, were found in the Cibao lime-
stone at locality 22. The largest measures in height 22 mm., diameter
25 mm.
Ilorizon.-Quebradillas limestone.

Natica (Ampullina?) collazoensis, new species
Plate XXI, Figures 11, 12
Shell large, moderately high spired; suture deeply impressed; whorls six
to eight; umbilicus partly covered by callus, or entirely open in some speci-
mens, in which the callus may have been broken away. Some specimens
show the last whorl strongly flattened below the suture, while in others the
whorl is evenly rounded. This is probably a sex difference; shell thin; sur-
face with numerous fine, impressed spiral lines, and much finer, almost
microscopic growth line threads, periodically incised, marking a resting stage
in the growth of the shell. The growth lines are remarkable for their even-
ness. Largest specimen measures in height 66 mm., diameter, 52 mm.
The shell in size and general appearance resembles Ampullina
amphora IIeilprin, and is probably related to the large Eocene forms.
It has some resemblance to Natica species indet. Brown & Pilsbry
(Proc. Acad. Nat. Sci. Phila., p. 360, Pl. 25, figs. 1, 2, 1911), a
Gatun shell.
Localities.-262, 265, 86 (base of falls), Collazo Falls,. 264 (bot-
tom 3rd falls), 83, 235.
Horizon.-San Sebastian shale. (very abundant).

Genus Epitonium Bolten
Epitonium (Cirsotrema) collazoensis, new species
Plate XXII, Figures 1, 2
Shell with three and one-half whorls measures In height 25 mm., maximum
diameter 13 mm.; whorls rounded; suture distinct, but not deep; whorls
with about twenty-four slightly oblique, moderately strong longitudinal ribs


and two opposite rows of sub-continuous varices, the last varix being at the
outer lip; longitudinal ribs fluted by another series of oblique ribs of equal
strength, descending to the left at about 45, continuing from suture to suture,
and being more or less interrupted by the longitudinal series of ribs. At
base of body whorl, both series of ribs terminate against a strong, rugose or
rope-like spiral, which encircles the umbilical area, and is interrupted only
by the varices. Over the entire surface of the shell is a fine microscopic cloth-
like texture composed of three intersecting series of threads, one series
vertical, another parallel with the 450 ribs, and the third series descending
to the right at about 45. The three series are named in order of their
relative strength, the vertical series being strongest, and locally obscuring
the other two series. Aperture circular. In general appearance the shell is
typical of the high spired forms of this group.
A single decollate specimen was found of an Epitonium which is
remarkable for its microscopic texture, which is somewhat of the
nature of that of E. text vestitum Maury, but the shell is quite
different from the latter. Apparently the nearest relatives of thi<
shell are to be found in the Paris Basin beds. Two such forms, very
simihr., though smaller, are Cirsotrenm subspinosum (Grat.) iof the
Aquitanien, and C. bourgeois do Boury, of the HIelvetian. This
relationship is of considerable significance in considering the age
of the San Sebastian shale.
Locality.-K. 29.2-3 Lares road.
Horizon.-San Sebastian shale.

Genus Turritella Lamarck

Turritella, tornata Guppy

variety portoricoensis, now variety

Phlte XX Figures 13, 14
Shell corresponds with Guppy's description, except that in the last few
whorls the anterior heel is double, and there are frequently as many as four
spiral threads in the median concavity. There are also from one to two
threads on the shoulder and likewise on the slope below the lower keel. The
base is spirally threaded and marked by accentuated growth lines. The
beading of the spirals is lacking or obscure, due probably to the poor state
of preservation of the specimens. All specimens are decollate. The largest
has a maximum diameter of 14 mm. and height (4 whorls) of 2G mm.; the
average sized specimen has a maximum diameter of 10 mm.
This species is extremely abundant in the San Sebastian shale.
Nearly all specimens are poorly preserved casts or internal molds,
and the ornamnentalion is difficult to make out.


Type locality.-232(b).
Other localities.-225, 331, 333, 343, Collazo Falls, 226, 262, S3,

Range.-San Sebastian shale to Lares limestone.

Turritella planigyrata Guppy

T. planiyyrata Guppy, Proc. Sci. Assoc. Trinidad, p. 169, 1867; Geol. Mag., p.
408, Pl. 18, fig. 5, 1874.
T. planigyrata Gabb, Trans. Amer. Phil. Soc., XV, p. 240, 1873.
T. planigyrata Guppy, Quart. Jour. Geol. Soc., XXXII, p. 519, 1876.
T. planigyrata Maury, Bull. Amer. Paleont., V, No. 29, p. 129, P1. 22, fig. 14,
A fragment comprising the last three whorls. Height 14 mm.,
maximum diameter 8.5 mm. The specimen agrees with the de-
scriptions by Guppy and Gabb, and there seems to be no doubt of
the identity.
Horizon.--San Sebastian shale.

Turritella mitchelli, new species

Plate XX II. Figure 3

A single decollate specimen with seven and one-half whorls. Shell large,
high spired (apical angle 5); whorls flat and continuous; suture indistinct
or invisible; whorls marked by three prominent crenulated spirals and
minor, minutely beaded spiral threads; the whole surface covered by minute
spiral striae and longitudinal growth lines, the latter producing the crenu-
lation in the spirals. In front of each suture is a flat area, less than one-third
the axial width of the whorl, marked by two to three obscure spiral threads;
next in order come a single primary spiral, followed by two fairly prominent
secondary spirals in the median portion of the whorl. The anterior third
of the whorl is marked by the two most prominent primary spirals with a
narrow channel between them, giving the effect of a doubled spiral band.
The anterior member of this pair is slightly larger than the other on the
last two whorls. Of the microscopic sculpture above mentioned, the sairal
striae are the most prominent on the primary ridges, and obsolete in the
interspaces, whereas the reverse is true of the growth lines, which results in
the strongest beading being on the minor spirals in the interspaces. The
base is sulcated below the anterior primary spiral, and is moderately convex
anterior to this sulcus. The surface of the base is marked only by the spiral
striae and growth lines, which here are markedly wavy or vermicular. The


suture is distinct on the last two whorls, but on the spire it is invisible,
being concealed by the anterior primary spiral, which overhangs it. Height
59 mm., maximum diameter 22 mm.
As the protoconch and upper portion of the spire are missing, it
is difficult to trace the relationship of this shell with other species.
The succession and arrangement of spirals is similar to T. halensis
Dall, a shell from the Flint River Oligocene. The finer sculpture
and beading resemble T. altilira Conrad, of the Gatun formation,
and T. calostemma Pilsbry & Brown, of the Oligocene of Haiti.
This shell is undoubtedly a member of the T. lornata-T. altilira
group, and is probably closest to T. calostemma. Berkey found a
similar, and apparently related shell from the basal Tertiary near
Juana Diaz. It differs in having four equally strong major spirals.
This shell is named in honor of Dr. G. J. Mitchell.
Horizon.-San Sebastian shale.

Turritella portoricoensis, new species

Plate XXI, Figure 15
Shell small; spire acute; about ten rounded whorls with about fifteen fine
spiral threads with sub-equal intervals; suture distinct, but not channelled
until the last three whorls; whorls at first evenly rounded with spiral
threads of equal strength, but later becoming angulated by accentuation of
spiral threads into a keel at periphery of whorl and another at anterior end
of whorl, overhanging the suture; first three whorls loosely coiled; proto-
conch missing on all specimens. The angulation begins at about the seventh
whorl. A specimen with nine whorls measures in height 13 mm., maximum
diameter 4 mm.
The young of this shell has the whorls of T. plcbeia Say, and in
the adult, the whorls are like those of T. wquistriata Conrad, in
some respects. The later whorls also resemble T. subannulata acro-
pora Dall, but are not quite as angular as in the latter species. The
nearest Antillean forms are T. planigyrata Guppy, of Santo Domingo,
and T. gatunensis Conrad, of Panama. The Porto Rican shell is
probably a less specialized member of this group.
Horizon.-Quebradillas limestone.


Turritella berkeyi, new species

Plate XXII, Figure 4

Shell extremely high spired, slender; whorls twenty-one; earliest whorls
simple, with two keels of equal size; anterior and posterior slopes of whorls
straight and of equal width; simple rounded median depression between the
two keels; last ten whorls with the posterior keel becoming double and the
shoulder of the whorl becoming gently convex, features which are quite promi-
nent in the adult stage; beginning with the eighth or ninth whorl an obscure
spiral appears on the anterior extermity of the whorl just above the suture
and partially overlapped by the succeeding whorl; surface throughout is
lacking in ornamentation. Height 73 mm., maximum diameter 11 mm.
This shell may possibly be a primitive form of T. perattennuata.
Heilprin. It resembles T. alcida Dall, but is very much more
slender. Other somewhat similar forms are T. altilirata Conrad, and
T. variabilis variety alticosta Conrad.
Horizon.-Quebradillas limestone.

Genus Petaloconchus Lea

Petaloconchus domingensis Sowerby

Petaloconchus domingensis Sowerby, Quart. Jour. Geol. Soc., VI, p. 51, Pl. 10,
fig. 8, a, b, c, 1849.
Petaloconchus sculpturatus Gabb, Trans. Amer. Phil. Soc., XV, p. 240, 1873.
Not of Lea.
Petaloconchus sculpturatus Guppy, Quart. Jour. Geol. Soc., XXXII, p. 519, 1876.
Vermetus (Petaloconehus) sculpturatus Dall (in part), Trans. Wagner Inst.
Sci., III, pt. 2, pp. 305, 306, 1892.
Petaloconchus domingensis Brown and Pilsbry, Proc. Acad. Nat. Set. Phila.,
p. 359, 1911.
Petaloconchus domingensis Maury, Bull. Amer. Paleont., V, No. 29, p. 128, P1.
22, fig. 11, 1917.
A single specimen was found, showing part of the coiled portion
of the shell. There seems to be no doubt as to the identity. The
fragment with three whorls measures in height 8 mm., diameter 6
mm., diameter of whorl about 3 mm.
Horizon.-Quebradillas limestone.


Petaloconchus? collazoensis, new species

Plate XXI, Figure 16
Tube of small diameter (2.5 mm.); surface smooth except for faint
growth lines; tip of spire missing in specimens, but one specimen shows a
loosely, irregularly coiled cylindrical spire of five whorls; diameter of spire
10 mm.; whorls are in contact, but coiled with a 5 mm. radius, leaving large
umbilical opening, the whole resembling the coiled steam pipes of a cylindrical
boiler. The specimens show that the coiling habit ceases abruptly, the tubes
then growing in all directions, almost tying knots.
Localities.-263, Collazo Falls.
Horizon.-San Sebastian shale.

Genus Pyramidella Lamarck

Pyramidella portoricoensis, new species

Plate XXI, Figure 17
Shell small, conic, smooth, rather low spired; seven flat whorls with
sharp median spiral offset, forming, with the sharply beveled shoulder of
the overlapping whorl, a broad, square sutural channel. The median offset,
or angulation, continues around the body whorl; suture of earlier whorls is
narrow; large circular perforation extends the entire length of the axis, and
is very conspicuous as seen from the base; columella tri-plicate; posterior
plication large and distinct; the other two small and obscure, of which the
anterior one is situated at the extreme base of the columella; aperture oval;
outer lip simple; height 4.75 mm., diameter 2 mm.
This shell resembles P. (Longchwus) forulata Guppy, but its
umbilicus places it with the typical group of Pyramidella. The
suture is of the type of P. semicanaliculatus Maury.
Horizon.-Quebradillas limestone.

Genus Modulus Gray

Modulus modulus Linn6

variety basileus Guppy
Monodonta busilea Guppy, Geol. 3lag., new ser. decade 2, I, p. 434, Pl. 16, fig.
2, 1874.
Modulus basilica Dall. Trans. Wagner Inst. Sci., III, pt. 2, p. 294, 1892.
Modulus modulus var. boasilea Dall, Trans. Wagner Inst. Sci., III, pt. 6, p. 1585,


This shell is apparently the same as that described by Guppy from
the Bowden. It agrees with Guppy's description, especially in its
low spire, and tuberculated, carinate whorls. On the lower portion
of the final whorl, however, it apparently differs somewhat from the
Bowden shell in having spirals of variable size, the primaries alter-
anting with one or more fine secondary threads. Judging from
Guppy's illustration, the aperture of the Porto Rican shell is broader.
The outer lip is denticulate, but the interior is not preserved, so as
to show whether the lirate character exists or not. Height of shell
is 14 mm., diameter 13 mm.
Apparently the modern representative of this shell is M. lentic1u-
laris (Chemin.) Auct., which according to Tryon is synonymous with
illM. modulus Linn6.
Locality.-East side of Vieques Island (Collected by A. K.
Horizon.- Quebradillas limestone.

Genus Bittium (Leach M. S.) Gray
Bittium species indet.

Plate XXI, Figure 18
Whorls six (or more), with three raised spirals crossed by sharp ribs
of equal height, and about twelve to a whorl; prominent nodes produced at
intersections of ribs and spirals; suture deeply impressed; height (6 whorls)
11 mm., diameter 5 mm.
A single specimen, too imperfect to identify or describe fully.
Horizon.-Qucbradillas limestone.

Genus Cerithium Adanson
Cerithium portoricoensis, new species
Plate XXII, Figures 5, G
Shell small, slender; protoconch unknown; whorls twelve or thirteen,
rounded, with obscure varices of irregular occurrence on the last six whorls,
and usually a prominent varix on the body whorl; first three whorls with
three primary spirals crossed by about twelve narrow rounded ribs, more
prominent than the spirals, and slightly nodose where they cross the latter,
the whole producing a cancellate structure with rectangular pits; a fourth


primary spiral appears on the fourth whorl; beginning with the sixth whorl,
secondary, less prominent spiral threads appear, two immediately above the
suture, and one between each of the four primaries. The two uppermost
secondaries do not appear until the seventh whorl; on last four whorls the ribs
have increased to about twenty in number, are irregular, low, and obscure, be-
ing marked chiefly by the lines of nodes where they cross the primary spirals.
The ribs usually become obsolete on the anterior two-thirds of the body
whorl; aperture short, slightly more than half the length of the body whorl;
posterior canal small, U-shaped; anterior canal not quite covered by inner lip;
outer lip slightly thickened; three average specimens have the following
Height 20 mm., Maximum Diameter 6 mm.
Height 17 mm., Maximum Diameter 5 mm.
Height 16 mm., Maximum Diameter 5 mm.

This shell has a superficial resemblance to C. coccodes Dall, and
to Clava chipolana Dall.

Horizon.-Quebradillas limestone (abundant).

Cerithium quebradillensis, new species

Plate XXI, Figures 7, S

A single decollate specimen with six whorls: earliest whorls roundly
angular, with shoulder and peripheral keel; last three whorls with about 12
high, rounded ribs, sub-nodose at the keel; all whorls with primary spirals
and secondary spiral threads; last four whorls with very fine tertiary spiral
threads; earliest whorls show two closely grouped primary spirals at the
keel, the upper one slightly stronger and forming the periphery of the whorl;
a third primary occurs at (just posterior to) the suture; these primaries
persist throughout, and a fourth primary spiral occurs on the base, being
covered in the preceding whorls. It lies close to the suture; the secondary
spirals at first form a group of three on the shoulder, another group of three
below the keel, and a single one between the primary spirals of the keel. The
three on the shoulder are markedly stronger than the other secondaries, a
feature which persists throughout; on the third whorl (of the decollate
specimen) the tertiary spiral threads appear, the first just above the sutural
primary, the next ones intercalated between the secondaries In the group
below the keel. The tertiaries do not appear on the shoulder until the fourth
whorl; the shoulder, adjacent to the suture, is, in all whorls, marked by a
rather wide band, at first smooth, but later marked by from one to six tertiary
threads; all whorls are marked by fine, raised growth lines, or longitudinal
threads, which produce a minute beading on crossing the spirals, and which
in the earlier whorls, produce with the spirals a fine cancellate ornamenta-
tion; the primary spirals of the later whorls have a secondary superimposed
thread, which is beaded by the growth lines; the base is slightly concave,.


and is sculptured with spirals of four sizes, more or less alternating. The
single primary, above described, is sculptured with numerous minute spiral
threads; pillar smooth, with no callus, anterior notch small, short, recurved;
slight suggestion of a posterior notch; aperture ovate. Height 32 mm.,
diameter 15 mm.
The most striking feature of this shell is the sudden change in
the character of ornamentation of the whorls, with the appearance
of the ribs. This final ribbed sculpture resembles that of C. calli-
soma Dall, from the Caloosahatchie Pliocene, but the Porto Rican
shell differs from the latter in its concave base and general turritel-
olid aspect. It is apparently not closely related to any species of the
Antillean region.
Horizon.-Quebradillas limestone.

Cerithium (Campanile) collazum, new species

Plate XXIII, Figures 1, 2

Shell large, conic, rapidly enlarging; whorls flattened to slightly rounded,
with a median concavity in the later whorls of adult individuals; suture
deeply impressed; each whorl with a flat, narrow shelf or shoulder pro-
jecting beyond the proceeding whorl; surface of whorls with medium fine,
nodose spirals like those of C. halense Dall, but with different arrangement.
Just behind the suture are one to two very finely nodose spiral threads, one
of which is frequently obscured by the overlapping whorl. Behind these
is the largest spiral, ornamented with small round nodes about 2 mm. apart.
The remainder of the whorl is ornamented with finer spirals, nodose, and
alternating in size. The median concavity of the later whorls comes im-
mediately posterior to the large noded spiral above described. The Inter-
spaces are slightly narrower than the spirals; base flattened, slightly convex,
smooth; aperture roundly quadrate; columella stout, with single prominent
fold, anterior notch short, large, recurved. A large decollate specimen with
six whorls measures in height 48 mm., maximum diameter 28 mm.
An interesting feature of this shell is the presence in the adult
stage of an impressed spiral line near the periphery of the base,
which marks the line of juncture with each succeeding whorl. This
is an adult character, and is lacking in the earlier whorls. It is
apparently an illustration of the inheritance of an acquired charac-
ter. Another interesting feature is the change in form of the whorl
from rounded in youth to flattened, and finally to a concave form
in the final stages of old individuals.


It would seem that this shell is related to some of the large, coni-
cal forms from the Flint River Oligocene, such as C. halense Dall,
as its ornamentation is quite similar to the latter. It differs notably
in the shape of its whorls, and in this latter respect, is more like the
large species of Campanile from the Paris Basin Eocene. Maiiry
has described the internal mold of a larger, but similar shell from the
San Sebastian shale Campanile (Portoricia) laricum Maury, which
may be a related species or a variety. Maury's species differs chiefly
in its flatter and more concave whorls, as well as in its considerably
greater size.
Localities.-262, Collazo Falls, 142 ?, 236. 749(a).
Range.-San Sebastian shale to lower Lares limestone.

Genus Strombus Linn6
Strombus proximus? Sowerby
Stromibus proximnts Sowerby, Quart. Jour. Geol. Soc.. VI, p. 4S, Pl. 9, fig. S,
Strombus pugilis Gabb (in part), Trans. Amer. Phil. Soc., XV, p. 233, 1873.
Not pugilis Linne. Exclude other synonyms.
Strombus proximus Guppy, Quart. Jour. Geol. Soc., XXXII, p. 521, 1876.
Strombus pugilis Dall (in part), Trans. Wagner Inst. Sci., III, pt. 1, p. 177,
1890. Not pu.ilis Linn6.
St rombus proximus Maury, Bull. Amer. Paleont., V, No. 29, p. 119, Pl. 20, figs,
4, 5, 1917.
Only incomplete specimens were found, which are identified doubt-
fully with this species.
Localities.-20-1, ,lona Island (Lobeck).
[lorizon.-Qncbradillas limestone. Ponce limestone.

Strombus bifrons? Sowerby

A single specimen was found among the material olleced froiii
1he Quebradillas limestone near iavaminon by O)r. Lobcck. The in-
leriial mold shows that the shell is the same shape as S. bifrons, and
measures in height (with tip of spire missing) 32 m1m.. diameter
10 mm. It is apparently an immature specimen., Ifr the outer lip
does not flare to any extent, and the anterior notch is not noticeably
recurred. The ornamentation consists of spirals like those of S.
bifrons, but with Ilie niodus or spines rather obsolete on the final


whorl, in which respect it resembles S. gatunenisis Toula. S. bifrons
is characteristic of the Aphera and Sconsia formation-s in Santo
Locality.-Xear Bayamon (L.obeck).
Horizon.-Quebradillas limestone.

Genus Orthaulax Gabb
Orthaulax gabbi? Dall
Plate XXIII, Figure 3

A single specimen was obtained from the sea cliff at the month of the
Guajataca River in a stratum of very resistant limestone. The speci-
men had been exposed to the polishing action of the waves, and the
original coloring, a light mottled brown, is preserved on the body of
the shell near the aperture. The surface of the body whorl and
spire are marked by the numerous faint longitudinal growth line
characteristic of the genus. Viewed from above, the outline is sub-
circular, with a slight triangularity as in 0. pvgnax HIeilprin and
0. gabbi Dall. The apical angle is about the same as that of 0.
gabbi, but considerably more acute than that of 0. aguadilleasis
Maury, a shell apparently from the same horizon. A marked sub-
angularity of the body whorl is very similar to that of 0. gabbi.
Above this angulation the whorl is strongly convex; below it, the
whorl is concave, with a slight convexity midway between the angu-
larity and the anterior end of the shell. The angularity is strongest
at the aperture, on the inner lip, but is obsolete over the final two
thirds of the body whorl. A slight shoulder is developed above the
angulation on the final portion of the body. Height of shell 65 mm.,
diameter 38 mm.
This shell was examined by Dr. Dall, who states that while the
specimen is too poor for positive identification, it is probably 0.
gabbi, a shell characteristic of the Chipola horizon. A comparison
with 0. aguadillensis Maury, a fragmental specimen, shows that the
shell here described is quite a different species.
Horizon.-Quebradillas limestone.


Orthaulax portoricoensis, new species

Plate XXV, Figures 1, 2, 3, 4, 5

Under this heading are included all the large Strombus-like in-
ternal molds, which are among the most abundant forms to be found
in the Quebradillas limestone. One of these internal molds was
figured by Semmes, in his report on the San Juan District, as
Strombus, species indet. The internal molds, taken by themselves,
are generically indeterminate, but resemble the molds of a large
Strombus. Fortunately, a few of these internal molds were found
with portions of the external mold adhering to the specimens, which
proves beyond a doubt that these large shells belong to the genus
Orthaulax. They attain a size much greater than any species of
Orthaulax yet known.
The specimens illustrated by figures 1 and 2 were examined by
Dr. Dall, who states that:

1. Figure 1 is undoubtedly an Orthaulax, possibly the same
species as Figure 2, but having a more acute or pointed

2. Figure 2 is also an Orthaulax, new species, almost cer-
tainly the same species as an undescribed form from
Santa Cruz.

3. Neither of these specimens is the same species as the
shell described in this report as 0. gabbi? Dall.

A large number of specimens, in addition to the two examined by
Dr. Dall, show considerable variation in size and elevation, or acute-
ness of the spire. The fact that all these variations can be found
in shells collected from a single fossiliferous pocket in the limestone,
would indicate that they arc for the most part variations of a single
species, although more than one species of Orthaulax occur in the
Quebradillas formation.
The apical angle as measured from the external molds, varies from
780 to 90 (largest specimen) ; and from the internal molds, varies
from 600 to 850. The following are measurements taken from a few
of the internal molds:



Height Diameter Apical Angle

1 .......... 155 mm
R .......... 115
3 . . . . . . . . . .
4........ ..........
5... (Fig. 1) .........
6...(Fig. 2) 80
7 .......... 75
8 .......... 70
9 .......... 70
10.......... 60

......... 105 m m .........
... ... ... S O .. ... ..

.. .... ... 60 ... .... ..
......... 4 0 .........
. ... ... .. 4 0 ...... ..

. . . . . 50 .. . . .
. ...... .. 4 3 .... .....
. .... .. .. 4 0 .. .....
.. .. ... 35 .. ... ..


From these measurements, it will be observed that the smaller
shells have a higher, more acute spire. This may be a varietal dif-
ference. Figures 3 and 4 show the internal mold of one of these
smaller shells in which the final whorl has begun to envelop the
spire, thus indicating that all of these small shells may be immature
individuals. Figure 1 shows the internal mold of the spire, and the
external mold of the enveloping dome of an adult shell. The curious
openwork structure replacing the shell wall is the cast of the burrow-
ing of a sponge or some other organism. This boring animal seems
to have taken a special liking to the thick-shelled Orthaulax and other
ponderous gastropods of the Quebradillas formation. The burrows
became infiltrated with lime carbonate, which solidified with the
surrounding limestone. The shell structures were then removed by
solution, leaving the casts of these burrows in the form of a complex
network resembling calcareous algae. Figure 5 shows the dome of
an adult shell of the low spired type. and illustrating the degree to
which the shell had been riddled with globular chambers, frequently
connected by long worm-like passages.

Localities.-204, 41, 52(a), 801, and numerous other localities
not recorded. An excellent specimen was collected by Dr. A. K.
Lobeck on the east coast of Vieques Island.

Horizon.-Quebradillas limestone, Los Puertos limestone. Char-
acteristic of the Quebradillas of the entire north coast of Porto Rico,
as shown by the collections of Berkey and Semmes.


Genus Cypraea Linn6
Cypraea spurcoides? Gabb
Cyprwa spurcoides Gabb, Trans. Amer. Phil. Soc., V, p. 235, 1873.
Cyprara spurcoides Alaury, Bull. Amer. Paleont., V, No. 29, p. 115, PI. 19, figs.
7, 8, 9, 1917.
Apparently the same as the form described from Santo Domingo,
except that the crenulations of the two lips are more nearly equal
in number. Height 24 mm., maximum diameter 18 mm.
Locality.-20 1.
Hoiizon.-Quebradillas limestone.

Cyproea sancti-sebastiani Maury
Localities.-262, 232(b), 263 ?, 233, 2305, 237, Juana Diaz.
Horizon.-San Sebastian shale, Juana Diaz shale.

Genus Cassis Lamarck
Cassis, species indet.
Plate XXTT. Figure 9
internal molds of a shell which may possibly be a new specie-.
The distinguishing feature is the presence of a strong varix on the
body whorl on the side opposite the aperture. Outer lip simple, ap-
parently not thickened. The specimens are uniform in size and
appear to be adult. The external sculpture, judging from impres-
sions on the internal molds, coin.,ists of from eighteen to twenty broad
bpirals on the last whorl, with correspondingly fewer spirals on the
earlier whorls. Height 30 mun., diameter 23 mim.
This shell is apparently of the type of Phalium ahlrichli Iall, but
is not quite as broad as the latter.
Localities.-232 (b), 23G.
Ilorizoln.-San Sebastlian shale.

Genus Dolium Lamarck
Subgenus Malea Valenciennes
Malea camura Guppy
Malca camera Guppy, Quart. .lour. Geol. Soc., XXII, p. 287, P). 17, fig. 9, 1866.
.Itoala ringenis Gabb (in part), Trans. Amer. Phil. Soc., XV. p. 223, 1873.

14 S


Malca camera Guppy, Quart. Jour. Geol. Soc., XXXII, p. 525, 1876.
Malea camera Dall, Trans. Wagner Inst. Sci., III, pt. 6, p. 1584, 1903.
Malea camura Maury, Bull. Amer. Paleont., V, No. 29, p. 112, Pl. 19, fig. 3, 1917.
The specimens average about 37 mm. by 27 mm. Only internal
molds were found with a few fragments of external mold.
Locality.-204; and Berkey 98.
Horizon.-Qiebradillas limestone (abundant).

Genus Columbella Lamarck

Subgenus Strombina Moerch

Strombina portoricana, new species

Plate XXTII, Figure 4
Protoconch of two whorls, rather large, smooth, naticoid; post-nuclear
whorls six to seven, moderately convex; suture distinct, but not channelled,
and becoming obscure on last whorl; surface of whorls smooth except for
faint growth striae, and by hump-like varices, or swellings, occurring as early
as the third post-nuclear whorl; body whorl with spiral lines, alternating in
strength, on the pillar and lower part of -whorl, but very faint over the
remainder of the whorl, and more closely spaced near the suture and on the
pillar. A specimen with eight whorls and base of pillar missing measures
in height 19.5 mm., diameter 9 mm. The largest specimen measures in height
33 mm., diameter 17 mm.
No complete specimens were found, and none showing the lip or
aperture. This shell is apparently related to the group represented
by S. lessepsiana Brown & Pilsbry, S. prisma Pilsbry & Johnson,
and S. cyphonolus Pilsbry & Johnson.
Locality.-204. /
IHorizon.-Quebradillas limestone. -

Genus Phos Montfort .

Phos costatus Gabb

Plate XXIT, Figure 10
Phos costatus Gabb, Trans. Amer. Phil. Soc., XV, p. 212, 1873.
Phos costatus Maury, Bull. Amer. Paleont., V, No. 29, p. 88, Pl. 14, figs. 13, 14,
The specimens correspond closely to those figured by Maury from
Santo Domingo. Protoconch not preserved in any of the specimens.


Ai1 adult specimen measures i14 height 29 mm., diameter 14 inm.
Number of whorls eight.
Horizon.-Qnehbradillas limestone (abundant).

Phos elegans Guppy

variety portoricoensis, new variety

Plate XXII, Figure 11

Shell small (height about 20 mm.), high spired; whorls rounded, with
about fourteen rather sharply rounded ribs, crossed by primary, and fre-
quently alternating smaller secondary spirals; small tubercules at inter-
sections of spirals and ribs; occasionally two or three longitudinal threads
between the ribs which do not cross the spirals; ribs irregularly spaced,
especially on the body whorl, where a varix is formed at or near the aperture;
ribs on the spire frequently vary in size, but do not form varices.
All the specimens are fragmental, but sufficient to show the entire
shell except the protoconch. This variety is probably closest to the
Santo Domingan occurrence of the species.
Horizon.-Quebradillas limestone.

Genius Murex Linn6

Murex (Phyllonotus) cornurectus Guppy

M. (Chicorcus) n etacerus Gabb, Trans. Amer. Phil. Soc., XV, p. 202, 1873.
Not M3. teguaccrus Sowerby.
l1. coruarectis Guppy, Quart. Jour. Geol. Soc., XXXII, p. 521, Pl. 2S, fig. 4,
Ml. (P'hyllonotus) cornturectus 2Maury, Bull. Amer. Paleont., V, No. 29, p. 103,
Pl. 16, figs. 9, 10, 1917.
An internal mold. together with a portion of the external mold.
was found of a large lMurex which seems to be this species. Height
(of last three whorls) SO min.. diameter (of internal mold) 45 mm.
Horizon.-Quebradillas limestone.


Genus Alectrion Montfort
Alectrion gurabensis A\laury
variety portoricoensis, new variety
variety varicum, new variety
Plate XX11, Figures 12, 11, 14, 15
Plate XXIV, Figure 19

The Arecibo limestone contains an abundance of a small shell
which is like 1. gurabensis Maury, a Santo Domingan form, but
which has some very evident differences from the latter. These
shells fall into two distinct varieties, one with coarse ribs, the other
with liner, more numerous ribs and a pronounced varix. The former
arc here called variety portoricoensis, the latter variety varicum.
The former is by far the most abundant of the two. Both forms are
smaller and have fewer whorls than the Santo Domingo type, but
they are not young shells as shown by the uniformity in size among a
large number of specimens. No specimen of either variety was found
which measures more than 8 mm. in height.
The following table shows the relation of the two varieties to the
type form from Santo Domingo, as described by Maury (Bull. Amer.
Palcont., V, No. 29, p. 91, Pl. 15, fig. 21, 1917):
H orizon.-Qnebradillas limestone.

Genus Fusus Lamarck
Fusus henekeni Sowerby
Plate XVII, Figure 5
Fusius henekeri Sowerby, Quart. Jour. Geol. Soc., VI, p. 49, 1849.
Fusus henekeni Gabb, Trans. Amer. Phil. Soc., XV, p. 204, 1873.
Fusis lhenekeni Guppy, Geol. Mag., p. 439, 1874; Quart. Jour. Geol. Soc.,
XXXII, p. 524, P1. 28, fig. 6 (type), fig. 2 (var. haitensis), 1876.
Fusius henekeri Dall, Trans. Wagner Inst. Sci., III, pt. 6, p. 1584, 1903.
Fusus henekeni Grabau, Smithsonian Misc. Coll., XLIV, No. 1417, p. 19, 1904.
Fusus henekeni Mlaury, Bull. Amer. Paleont., V, No. 29, p. 78, P1. 12, fig. 1,
A single specimen was found, with protoconch and body whorl
missing. Only primary and secondary spirals show on the specimen.



A. gurabensis Maury A. gurabeisis var. A. gurabensis var.
Characters Santo Domingo portoricensis varicum
Porto Rico Porto Rico

Height 11 mm. Height 7 mm. Height S nun.
Size Diameter 6 mm. Diameter 4.5 mm. Diameter 4.5 mm.

whorls 3 smooth 2-2.5 2?

whorls 6 5 5

Character of Con vex, angulated Convex, sub-angular Earlier whorls sub-
post nuclear and shouldered; ribs and shouldered. angular-the last
whorls narrow, round ; two are rounded.
wider interspaces.

Suture Wavy Wavy Slightly wavy

Ribs on body 11 12 13-14
whorl Slightly oblique Oblique

Spirals on spire. Penultimate whorl Penultimate whor 1 3rd post-nuclear
3 strong, widely same as in the San- 1 weak posterior
spr ced to Domingo shell 3 strong median
3 weaker above 2 weak anterior
3 weaker below
3 weak posterior
3 strong median
4 weak anterior

Spirals on body 3 or 4 weak poster, r 2-3 weak posterior 3 weak posterior
3 strong median 3-4 strong median 3 strong median
12 weak anterior- 10-13 weak anterior 12 weak anterior

Spirals on pillar S 6-7, arrangement-2
strong alternate
with 2 weak ?

Aperture Round Sub-Round 1Round

Outer lip 8 lirae Thin T h i c k o n e d by
15 lirae, alternate strong varix
long and short

Inner lip Posterior denticle Smooth, moderately Smooth, moderately
thick callus, thick callus.
Posterior denticle Posterior denticle
absent or obscure.

Anterior canal.. Well defined Broad and deep


This is probably a small variety, otherwise very close to the type form,
but more specimens are required for a complete description. The
spire of eight whorls measures in height 20 mm., maximum diameter
8.5 mm.
Horizon.-Quebradillas limestone.

Genus Turbinella Lamarck

Turbinella chipolana Dall

variety precursor, new variety

Plate XXIII, Figures 6, 7

Shell small, fusiform, consisting of three and one-half nuclear whorls and
three post-nuclear whorls; protoconch like T. chipolana Dall and T. regina
Heilprin, in having a swollen tip, succeeded by smaller nuclear whorls. The
tip consists of one and one-half volutions, is transversely ovate, and is
peculiar in having a shallow median constriction. The two succeeding
nuclear whorls considerably smaller in diameter, the first being gently
convex, the last nearly flat. The latter is succeeded abruptly by sculptured
whorls with seven rounded, not nodose ribs, and prominent spirals, of which
there are four on the first two, and about twenty-three on the final whorl
and pillar. The spirals are separated by wider interspaces. The whorls
are marked by a prominent, narrow and wavy sub-sutural band or shoulder,
sculptured with at first one and later with two spirals. The interspace
immediately below the shoulder is strongly impressed. The spirals are of
equal strength, except on the pillar, where faint intercalated threads appear.
The entire post-nuclear surface is marked by fine raised longitudinal growth
lines, strongest in the interspaces between the spirals. Height of shell
17.5 mm., maximum diameter 7 mm.; height of protoconch 31/2 mm., diameter
of tip of protoconch 2% mm.
This shell is probably the ancestor of T. chipolana Dall. It is
apparently adult, its small size being indicative of its primitive
character. The same is true of variety areriboense, a mutation
from the San Sebastian shell occurring in the Quebradillas lime-
stone. All the material available was gone over thoroughly with
the purpose of trying to find some molds or fragments which would
show that these shells reach a larger size, but the results were nega-
tive. It seems, therefore, safe to assume that the primitive Turbi-
nellas were small forms, which did not reach their maximum size


until the Pliocene. Analogous cases may be seen in smie of the
genetic groups of Fusus (Grabau, A. W. Phyl. Fusus, Smithson.
Misel. Coll., XLIV, No. 1417, 191041.


Horizon.-San Sebastian shale.

Turbinella chipolana Dall

variety arediboense, new variety

Plate XXITT, Figure <

A single specimen in which the lower half of the body whorl is missing.
Protoconch like that of variety precursor, but without median construction
of the tip, and with the third whorl almost entirely concealed by the over-
lapping of the first post-nuclear whorl, giving the appearance of a two-
whorl protoconch. The tip is slightly larger (diameter 3 mm.) than that
of precurs-or, and slightly more swollen when compared with the succeeding
whorls. The whorl following the tip is nearly flat, whereas in precursor it
is noticeably convex. The first post-nuclear whorl has ribs, spirals, and sub-
sutural band, or shoulder, as in precursor and represents the precursor stage.
On the next and following whorls, however, the ribs become nodose, and
on later whorls there are more spirals Ihan in preccursor, with occasionally
a fine intercalated spiral. The longitudinal growili lines are similar to
those of precursor, but show a tendency to alternate in strength. The sub-
sutural hand, or shoulder, is prominent in the first two post-nuclear whorls
and marked by a single spiral; on the third whorl it is inconspicuous, and
marked by two spirals, as in the later whorls of precuirsor. while on lhe
fourth and fifth (or final) whorls it has disappeared altogether. This fact
is interesting in showing the tendency to lose a primitive character which
has apparently been completely lost in the Chipolan shell. Height of frag-
mient 20 mni., maximum diameter t11 mn.

This shell is a mutintion from variety precursor, the precursor, (or
simple ribbed stage being accelerated and limited to the first post-
nuclear whorl. The Inter whorls have nodose ribs and repreount
what may be termed the arecibocese stage. This shell is best inter-
preted as a local mu ation not in the direct line leading to 7'. dchipo-
lana and T. regina.
Ilorizon.-Qnebradillas limestone.



Genus Xancus Bolten

Xancus validus Sowerby
Turbinellus validus Sowerby, Quart. Jour. Geol. Soc., VI, p. 50, 1849.
Turbinella valida Gabb, Trans. Amer. Phil. Soc., XV, p. 218, 1873.
Turbinellus validus Guppy, Geol. Mag., p. 438, 1874; Quart. Jour. Geol. Soc.,
XXXII, p. 523, 1876.
Turbinella scolymus Tryon (in part) Man. Couch., IV, p. 70, 1882. Not the
recent shell.
Turbinella validus Dall, Trans. Wagner Inst. Sci., III, pt. 1, p. 99, 1890.
Turbinella scolymus Dall (in part), Trans. Wagner Inst. Sci., Ill, pt. 1, p. 97,
Xancus validus Maury, Bull. Amer. Paleont., V, No. 29, p. 83, Pl. 13, fig. 5, 1917.

Apparently rare. Only a few fragments were found, showing the
Horizon.-Quebradillas limestone.

Xancus, new species?

Plate XXI[I, Figure 9
Known only from fragments. Spire unknown; shell apparently like
X. validus Sowerby and X. wilsoni Conrad, but differing chiefly in the greater
number of tubercles, or spines, which are along a contracted zone below the
rounded shoulder. There are four high, strong columellar folds, the
anterior one being smaller than the others. The lower portion of the body
whorl is sculptured with strong spirals as in X. validus. The exterior of the
remainder of the shell is unknown, except for the presence of the tubercles.
Height of body whorl 67 mm., maximum diameter about 50 mm.

Locality.-244, 254-5-6.
Horizon.-Lares limestone.

Genus Mitra Lamarck

Mitra henekeni Sowerby

Plate XXIV, Figure 1
Mitra henekeri Sowerby, Quart. Jour. Geol. Soc., VI, p. 46, Pl. 9, fig. 5, 1849.
Mitra henekeni Gabb, Trans. Amer. Phil. Soc., XV., p. 219, 1S73.
Mitra henekeni Guppy (in part), Quart. Jour. Geol. Soc., XXXII, p. 528, 1876.
.llitra henekeni Maury, Bull. Amer. Paleont.. V, No. 29, p. 74, Pl. 12, figs. 5, 5a,


A single fragmental specimen of a young shell. Protoconch miss-
ing. Seven whorls. Height 20 mm., diameter 7 mm.
Horizon.-Quebradillas limestone.

Mitra symmetrica Gabb
Mitra symmetrica Gabb, Trans. Amer. Phil. Soc., XV, p. 220, 1873.
Mitra symmetricus Maury, Bull. Amer. Paleont., V, No. 29, p. 75, 1917.
A single fragmental specimen, consisting of the last two whorls with
anterior portion missing. Columellar folds not shown; whorls flattened,
slightly convex; suture distinct, but not impressed; aperture elongate-ovate;
surface with flattened spirals and much narrower grooves between, which
are marked by growth lines. Height 25 mm., diameter 12 mm.
It has been suggested that M. symmetrica may be the young of 3V.
titan Gabb.
Horizon.-Lares formation.

Genus Olivella Swainson
Olivella muticoides Gabb
variety portoricoensis, new variety
Plate XXTV, Figures 2, 3
The shells vary from high to low spired, and also in the number
of columellar plications. They include the types represented by
0. muticoides and 0. muticoides variety canaliculata of Santo
Domingo, and there are all gradations between. The sutural canal
is extremely broad in all specimens; columellas callus very thick;
three to four columellar plications, with several faint ones posterior
to these; largest specimen measures in height 12 mm., diameter
4.5 mm.
Localities.-2 04, 671; Aguila Point, K 75.2 Ponce-Penuelas road
Horizon.-Quebradillas limestone (extremely abundant); and
Ponce limestone.


Olivella portoricoensis, new species
Plate XXIV, Figure 4
Shell small, smooth, moderately high spired; whorls about six; protoconch
missing on all specimens; suture distinct, impressed below upper margin of
the succeeding whorl, leaving a gap between top of whorl and surface of the
preceding whorl; a raised spiral thread appears above the suture, barely
showing above the upper margin of the overlapping whorl; columella with
two spiral grooves; callus moderately thick; anterior notch shallow, incon-
spicuous. Height of shell 12 mm., diameter 5 mm.
The distinguishing feature of this shell is the spiral thread at the
suture. This thread is not continued on the body whorl.
Localities.-204, 216.
Horizon.-Quebradillas limestone.

Genus Cancellaria Lamarck
Cancellaria hevescens Guppy

Plate XXIV, Figures 5, 6
Cancellaria 7lvescens Guppy, Quart. Jour. Geol. Soc., XXII, p. 289, Pl. 17,
fig. 12, 1866.
Cancellaria lwvescens Gabb, Trans. Amer. Phil. Soc., XV, p. 239, 1873.
Cancellaria larvescens Mauri, Bull. Amer. Paleont., V, No. 29, p. 64, P1. 10,
fig. 6, 1917.
Cancellaria 7evescens Dall, Trans. Wagner Inst. Sci., III, pt. 1, p. 43, 1890.
The specimens are smaller than the Jamaican shells, and show
all variations between complete loss of cancellation on the body whorl
and merely a diminution of strength of cancellation near the aper-
ture. In the latter cases, the spirals continue, but the ribs are ir-
regular in occurrence and indistinct or absent for considerable in-
tervals. In some shells the ribs on the body whorl become merely
irregular crowded growth lines; callus very thick; columella with at
least three prominent plaits; aperture not widening anteriorly as
does that of the Jamaican shell. An average shell measures in height
23 mm., diameter 14 mm. The largest complete specimen measures
26 mm. by 15 mm., while a fragment indicates that the shells attain
a size slightly larger than this.
Horizon.-Quebradillas limestone (abundant).


Genus Turris Bolten
Turris albida Perry
variety haitensis Sowerby
Plate XXIV, Figure 7
Pleurotoma haitensis Sowerby, Quart. Jour. Geol. Soc., VI, p. 50, 1849.
Turris albida var. haitensis Maury, Bull. Amer. Paleont., V, No. 29, p. 50,
PI. S, fig. 4, 1917.
A few specimens were collected which agree closely with this
variety. The largest specimen (spire with body whorl missing)
measures in height 22 imm., diameter S mim.
Ilorizon.-Quebradillas limestone.

Turris albida Perry
variety cf. virgo Lamarck
1'leurotoma virgo Lamnarek, Anim. s. Vert., VII, p. 94, 1822.
1'leurotoma virgo Moore, Quart. Jour. Geol. Soc.. IX, p. 130, 1853.
Turris (Surcula) virgo Gabb, Trans. Amer. Phil. Soc., XV, pp. 206, 20t, 1873.
7'Trris albida var. rirgo Maury, Bull. Amer. Paleont. V, No. 29, p. 50, P1. 8,
fig. 6, 1917.
A single fragment showing portions of the last two whorl:. and
indicating a shell of about 60 mm. height. An unusual feature is
the absence of secondary spirals. It seem to bh an adult form which
has retained the form of the young variety cirgo. Another special
feature is the co rseness of the spiral, the prominent median keel
being especially broad and thick, resembling the keels of Erphora
,1uadricosiata. The suture is indistinct. Interspaces between spirals
marked only by the characteristic oblique growth lines.
florizon.-Q clbradillas limestone.

Genus Drillia Gray
Drillia consors Sowerby
variety portoricoensis, new variety
Plate X XIV, Figures s, )9
Shell small, moderately slender; whorls about nine; sutural band as in
tli type, but nearly half the axial width of the whorl; sculpture of from



four to five prominent spirals, and about 18 sharp ribs which do not cross
the sub-sutural band, the latter being smooth, and concave on either side of
the spiral ridge which divides the band. The concavity below this spiral
ridge is over twice the width of that above the ridge, and much more deeply
concave. Height 15 mm., diameter 5 mm.

This shell is close to D. consors, but has some distinctive differ-
cnces. One of the specimens shows the protoconch fairly well. Tt
is rather large, smooth, naticoid, and apparently consists of two
whorls. The first post-nuclear whorl is sculptured with ribs only,
the spirals being too fine to observe (if present at all) and the sub-
sutural band not appearing until the second sculptured whorl.
Locality.-204; Berkey 98.
Horizon.-Quebradillas limestone.

Drillia grabaui, new species
Plate XXTV, Figure 10
Shell small, slender; decollate specimen (5 whorls) measures in height
14 mm., diameter 4 mm.; protoconch of two smooth, rather large, globular
whorls; post-nuclear whorls with sharp, widely spaced ribs and fine, flattened
spiral threads, not crossing the ribs and slightly arcuate between the ribs;
sub-sutural band like that of D. consors, but narrower and wavy, or fluted,
and marked with faint spiral threads and arcuate growth lines indicating
a fairly deep, semi-circular posterior sinus. On the penultimate whorl, there
are six strong spirals, with three additional ones just below the band, which
are fine, crowded, and about the same strength as those on the band. On the
body whorl there are twenty-two spirals, and the ribs extend well toward the
anterior portion of the shell before dying out. The ribs, spirals, and band
appear together immediately following the protocouch stage.

This shell has a slight resemblance to D. maonorisparuin lMaurvy,
of the Santo Domingan Oligocene, but differs in its slightly larger
size, more slender form, different protoconeh, and absence of mny
varix. It probably belongs to the D. consors group.
Horizon.-Quebradillas limestone.

Drillia portoricoensis, new species
Plate XXIV, Figures 11, 12
Shell with ten whorls, including protoconch of two smooth, globular
volutions; post-nuclear whorls sculptured with low, straight longitudinal ribs
and faint spirals, visible only between the ribs, except on the anterior half


of the body whorl, where they are much stronger, and slightly nodose where
they cross the ribs; sub-sutural band of the D. consors type, with dividing
spiral ridge prominent in the earlier whorls, but becoming low and incon-
spicuous in the adult; band slightly wrinkled where it laps over the ribs
of the preceding whorl; lower half of band slightly concave, and smooth
except for faint, microscopic spiral lines, and growth lines indicating a broad,
sub-angular posterior sinus; adult shells with a strong, thick varix at the
outer lip. Height of shell 10 mm., diameter (excluding varix) 5.5 mm.
This shell is easily recognizable from its conspicuous varix, and
its almost obsolete sculpture.
Horizon.-Quebradillas limestone.

Drillia semmesi, new species

Plate XXIV, Figure 13
Shell small, broader and lower spired than is usual in members of this
genus; whorls seven, including a large protoconch of two smooth, rounded
whorls, the first small, the second considerably larger; first post-nuclear
whorl sculptured with about thirteen sharp ribs extending from suture to
suture, and crossed by three raised spiral ridges, producing a cancellate
sculpture; all post-nuclear whorls sharply shouldered, and the ribs, on crossing
the shoulder, are strongly arched backward, a feature most noticeable in the
later whorls; shoulders of the last four whorls are marked by obscure spirals;
later whorls retain the three primary spirals, and on the fourth whorl,
smaller intercalated secondary spirals appear. The uppermost primary spiral
marks the shoulder angulation. The body whorl has about twelve prominent
(or "primary") spirals, with some of the interspaces marked by one or two
faint (or "secondary") spirals. All post-nuclear whorls show longitudinal
growth lines, arcuate on the shoulder, indicating a wide U-shaped posterior
notch. Aperture and anterior portion of shell unknown. Length of most
complete specimen, with lower portion of body whorl missing, 8 mm., diameter
4 mm.
This shell is named in honor of Dr. D. R. Semmes, in recognition
of his work on the San TTuan District of Porto Rico.
Ilorizon.-Quebradillas limestone.

Genus Cythara Schumacher

Cythara of. elongata Gabb
Mangilia elongata Gabb, Trans. Amer. Phil. Soc., XV, p. 211, 1S73; Jour. Acad.
Nat. Sci. Phila., VIII, p. 351, Pl. 46, fig. 34, 1881.


Cythara elongata Dall, Trans. Wagner Inst. Sci., III, pt. 6, p. 1583, 1903.
Cythara elongata Maury, Bull. Amer. Paleont., V, No. 29, p. 59, Pl. 9, fig. 12,
A single poorly preserved specimen, which seems to be this species
or one very closely related. Height 7.5 mm., diameter 3 mm.
Horizon.-Quebradillas limestone.

Genus Terebra Adanson

Terebra quebradillensis, new species

Plate XXIV, Figure 14
Shell small, slender; protoconch of two smooth globular whorls, slightly
larger than the succeeding post-nuclear whorl; twelve post-nuclear whorls,
rounded as in T. berlinerw Maury; sub-sutural band nearly one-third the
axial width of the whorl, marked off by deep, narrow furrow, and faintly
striated by one or two spiral threads as in T. wolfgangi Toula; about
eighteen arcuate ribs on each whorl, which cross the furrow and sub-sutural
band without interruption or change of direction; remainder of whorl below
sub-sutural band marked by about nine spiral threads (sixteen on last whorl)
which vary in size. A prominent spiral is situated next to the furrow. The
rest are finer, and crowded together in groups of two or three. In the earlier
whorls they are more uniform in size and distribution; and fewer in number;
columella short and non-plicate. The' largest specimen measures in height
18 mm., diameter 4 mm.
This shell resembles T. berlinerce Maury in its convex whorls
and the character of the spiral threads. It differs in its smaller
size and wider sub-sutural band. It resembles T. wolfgangi Toula
in its bisected or trisected sub-sutural band.
Localities.-204, and near Ponce (Mitchell).
Horizon.-Quebradillas limestone, Ponce limestone.

Genus Conus Linn6

Conus catenatus Sowerby

Plate XXIV, Figure 15
Conus catenatus Sowerby, Quart. Jour. Geol. Soc., VI, p. 45, Pl. 9, fig. 2, 1S49.
Conus interstinctus Guppy, Quart. Jour. Geol. Soc., XXII, p. 288, Pl. 16, fig. 3,
Conus catenatus Gabb, Trans. Amer. Phil. Soc., XV, p. 230, 1873.


Conus catenati s Guppy, Quart. Jour. Geol. Soc., XXXII, p. 527, 1876.
Conus catenatus Maury, Bull. Amer. Paleont., V, No. 29, p. 38, PI. 5, fig. 4;
Pl. 6, figs. 1, 2, 1917.
The specimens vary from concave to only slightly concave spire.
from high to moderately low spire, and there is considerable range
in size. Upper whorls marinated; last five whorls rounded on shoulder
and broadly channelled above, with faint arenate growth lines crossing
the channels; posterior third of body whorl smooth; anterior two-
thirds marked by spiral threads which alternate with much finer.
obscure threads. There are many variations, some of the specimens
resembling G. cercadensis Alaury, other resembling 0. kitleredgei
Maury, but all evidently variations of 0. catenatus. A typical shell
measures in height 31 iron., diameter 17 mm.
Locality.-204. Also found by Berkey in the vicinity of Quc-
Horizon.-Quebradillas limestone.

Conus of. marginatus Sowerby

Plate XXIV, Figure 16

Conus marginatus Sowerby, Quart. Jour. Geol. Soc., VI, p. 44, 1849.
Conus marginalits Gabb, Trans. Anier. Phil. Soc., XV, p. 230, 1873.
Conns marginatus Guppy, Quart. Jour. Geol. Soc., XXXII, p. 528, Pl. 29, fig. 5,
Conts (Chelycownts) marginatus Cossmann, Jour. de Conch., LXI. pi. 44-46.
PI. 3, figs. 14, 15, 1913.
Conns marginatus Maury, Bull. Amer. Paleont., V, No. 29, p. 46, Pl. 7, fig. 11,
A small conus which has features in conunon with 0. marginatus
and may be the young of the latter. Whorls eight, including two
nuclear; spire moderately high, scarcely concave; whorls marinate
except last whorl, which is rounded at the shoulder; whorls of the
spire flat above, while the final whorl is slightly channelled above;
posterior half of body smooth; anterior half with twelve smooth,
flat, spiral bands separated by deep grooves marked by raised longi-
tudinal threads, which are limited to the grooves and do not eros? the
spiral hands. THeight 7.5 miim., diameter 4 111111.
Horizon.-Quebradillas limestone.


Genus Bullaria Rafinesque

Bullaria paupercula Sowerby
Bulla pauperccla Sowerby, Quart. Jour. Geol. Soc., VI, p. 52. 184l0.
Bulla paupercala Gabb, Trans. Amer. Phil. Soc., XV, p. 246, 1873.
Rulla paupercula Guppy, Geol. Mag., p. 437, 1874; Quart. Jour. Geol. Soc.,
XXXII, p. 518, 1876.
Balla paupercula Dall, Trans. Wagner Inst. Sci., III, pt. 1, p. 18, 1890.
Bullastriata Dall, Trans. Wagner Inst. Sci., III, pt. 6, p. 1583, 1903.
Bullaria paupercula Maury, Bull. Amer. Paleont., V, No. 29, p. 18, Pl. 3, fig. 8,
The specimens are internal molds, but their general outline seem-
sufficient to identify them with this species. Fragments of external
mold show faint spiral grooves on the anterior part of the shell.
Spire depressed. Height 15 mm., diameter 10 mm.
Localities.-204, 664, 235 ?.
Horizon.-Quebradillas limestone, Ponce limestone.

Bullaria portoricoensis, new species

Plate XXIV, Figure 17
Shell cylindrical, length nearly twice the diameter; spire sunken, imper-
forate; aperture longer than shell, extending above the truncate apex; surface
with fine, slightly wavy impressed spiral lines, covering all of the shell except
the summit, and crossed by slightly curved growth lines. An average
specimen measures in height 20 mm., diameter 11.5 mm.
The most striking characteristic of this shell is its cylindrical form.
The young shell is more globose in form, resembling one of the
elongate forms of B. granosa.
Horizon.-Quebradillas limestone.

Bullaria granosa Sowerby

Plate XXIV, Figure 18
BRulla granosa Sowerby, Quart. Jour. Geol. Soc., VI, p. 51, Pl. 10, fig. 10, 1849.
Bulla granosa Gabb, Trans. Amer. Phil. Soc., XV, p. 246, 1873.
Bulla granosa Guppy, Geol. AMag., p. 437, 1874; Quart. Jour. Geol. Soc., XXXII,
p. 518, 1876.
Bullaria granosa Maury, Bull. Amer. Paleont., V, No. 29, p. 20, Pl. 3, fig. 10,


The specimens are slightly more globular than the Santo Domingo
shells. There is a great range in size, but the specimens fall into
two groups, one of a large size averaging 20 mm. in height and
16.5 mm. in diameter, and the other a small size averaging 14 mm.
in height and 11 mm. in diameter. The small shells have a large
range in size and proportions, some of the measurements being:
Height Diameter
17 mm. 15 mm.
11 mm. S mim.
16.5 mm. 12.5 mm.
11.5 mm. 10 mm.
The surface sculpture of the shells agrees closely with the shells
from the Santo Domingo Oligocene.
Localities.-204, 209 ?, 216, 52(a), 191, 225, 678. San Juan
district (Semmes). Culebrinas Point? (Mitchell).
Horizons.-Lares limestone to Quebradillas limestone, Ponce lime-
stone? Extremely abundant in the Quebradillas limestone.


Fig. 1. Alrina rabelli n. s.; nat. size; external mold.
Fig. 2. Pecten (Pecten) laresense n. s.; x 1.6; left valve.
Fig. 3. Pecten (Pecten) laresense n. s. ; x 2; right valve.
Fig. 4. Pecten (Euvola) reliquus Brown and Pilsbry, var. porto-
ricoensis n. var.; x 2.3.
Fig. 5. Pecten (Nodipecten) nodosus Linn6; x 1.8.
Fig. 6. Phacoides (Lucinisca) calhounensis Dall; x 2.6; immature
shell; gutta-percha cast.
Fig. 7. Metis trinitaria Dall; x 0.75; internal mold, left valve.




/.' ~
-~, /




->. .-1 '
* -^

University of Florida Home Page
© 2004 - 2010 University of Florida George A. Smathers Libraries.
All rights reserved.

Acceptable Use, Copyright, and Disclaimer Statement
Last updated October 10, 2010 - - mvs