• TABLE OF CONTENTS
HIDE
 Front Cover
 Table of Contents
 Introduction
 Results: genus incania
 Genus cyclonenia
 Genus bequaertineia
 Genus hemicena
 Genus temesa
 Genus incaglaia
 Genus cylindronenia
 Genus andinia
 Genus brevinenia
 Ecological remarks
 Acknowledgement
 Literature cited
 Back Cover






Group Title: Bulletin of the Florida Museum of Natural History
Title: New South American clausiliidae from the collections of The Florida Museum of Natural History
CITATION THUMBNAILS PAGE IMAGE ZOOMABLE
Full Citation
STANDARD VIEW MARC VIEW
Permanent Link: http://ufdc.ufl.edu/UF00087418/00001
 Material Information
Title: New South American clausiliidae from the collections of The Florida Museum of Natural History (Gastropoda, Clausiliidae, Neniinae)
Series Title: Bulletin of the Florida Museum of Natural History ; vol. 45, no. 2
Physical Description: p. 45-62 : ill. ; 28.
Language: English
Creator: Neubert, Eike
Nordsieck, Hartmut
Publisher: University of Florida
Place of Publication: Gainesville, FL
Publication Date: 2005
Copyright Date: 2005
 Subjects
Subject: Clausiliidae -- South America   ( lcsh )
Genre: bibliography   ( marcgt )
non-fiction   ( marcgt )
 Notes
Summary: The material described here contains 32 taxa of Neniinae including sixteen new species, all but one of which were found to live in areas dominated by limestone substrata.
Bibliography: Includes bibliographical references (p. 59).
General Note: Cover title.
Statement of Responsibility: Eike Neubert and Hartmut Nordsieck.
 Record Information
Bibliographic ID: UF00087418
Volume ID: VID00001
Source Institution: University of Florida
Holding Location: University of Florida
Rights Management: All rights reserved by the source institution and holding location.
Resource Identifier: oclc - 62862662
issn - 0071-6154 ;

Table of Contents
    Front Cover
        Page i
        Page ii
    Table of Contents
        Page 45
    Introduction
        Page 46
    Results: genus incania
        Page 47
    Genus cyclonenia
        Page 48
    Genus bequaertineia
        Page 49
    Genus hemicena
        Page 50
    Genus temesa
        Page 51
    Genus incaglaia
        Page 52
    Genus cylindronenia
        Page 53
    Genus andinia
        Page 54
    Genus brevinenia
        Page 55
        Page 56
    Ecological remarks
        Page 57
    Acknowledgement
        Page 58
    Literature cited
        Page 59
        Page 60
        Page 61
        Page 62
        Page 65
    Back Cover
        Page 66
Full Text





-FLORIDA
_MUSEUM
OF NATURAL HISTORY*


BULLETIN


NEW SOUTH AMERICAN CLAUSILIIDAE FROM THE
COLLECTIONS OF THE FLORIDA MUSEUM OF NATURAL
HISTORY (GASTROPODA, CLAUSILIIDAE, NENIINAE)




Eike Neubert and Hartmut Nordsieck


Vol. 45, No. 2, pp. 45-62


UNIVERSITY OF FLORIDA


2005


GAIN ESVILLEE







The FLORIDA MUSEUM OF NATURAL HISTORY is Florida's state museum of natural history, dedicated to
understanding, preserving, and interpreting biological diversity and cultural heritage.

The BULLETIN OF THE FLORIDA MUSEUM OF NATURAL HISTORY is a peer-reviewed publication
that publishes the results of original research in zoology, botany, paleontology, and archaeology. Address all inquiries
to the Managing Editor of the Bulletin. Numbers of the Bulletin are published at irregular intervals. Specific volumes
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last issue in that volume.



Richard Franz, Managing Editor
Erika H. Simons, Production



Bulletin Committee
Richard Franz, Chairperson
Ann Cordell
Sarah Fazenbaker
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ISSN: 0071-6154

Publication Date: December 15, 2005






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Managing Editor of the BULLETIN
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NEW SOUTH AMERICAN CLAUSILIIDAE FROM THE COLLECTIONS OF
THE FLORIDA MUSEUM OF NATURAL HISTORY (GASTROPODA,

CLAUSILIIDAE, NENIINAE)





Eike Neubert' and Hartmut Nordsieck2





ABSTRACT

The material described here contains 32 taxa of Neniinae. The following new genus is described: Brevinenia n. gen. The following
species or subspecies are new to science: Bequaertinenia delicate n. sp., Columbinia (Steatonenia) pachygastris n. sp.,
Cyclonenia albosuturalis n. sp., Cyclonenia gibber n. sp., Cvlindronenia violacea n. sp., Hemicena polinskiana ancashensis
n. ssp., Incaglaia leucostoma n. sp., Incania papillosa papillosa n. sp. n. ssp., Incania papillosa imbecilla n. sp. n. ssp.,
Incania platystoma n. sp., Steeriana cajamarcana sororminor n. ssp., Steeriana sorormajor n. sp., Temesa gradata n. sp.,
Temesa pilsbrvi perfectecostata n. ssp., Zilchiella palatalis n. sp., Zilchiella scala n. sp. All species except Incaglaia huanucensis
were found to live in areas dominated by limestone substrata. Several species seem to persist even when the vegetation type
changed due to human activities.


Key words: Gastropoda, Pulmonata, Clausiliidae, new species, Peru.

TABLE OF CONTENTS


Introduction ........................................................................ .....................................46
M materials and M ethods..................................................... ................................ 46
R results ............................. ........................................................... ............... ........ ... ..46
G enus C olum binia ..................................................... ................................ 46
G enus Incania ................................................................... ...........47
G enus Cyclonenia ................................................... .................................. 48
G enus Bequaertinenia .......................................................... .................... 49
G enus H em icena ...................................................... .................................. 50
G enus P eruinia ...................................................................... .................... 50
Genus Pseudogracilinenia ...................................................... ....................50
G enus Tem esa .......................................................................... .................... 5 1
G enus Incaglaia ...................................................... .................................. 52
G enus Cvlindronenia .......................................................... ..................... 53
G enus A ndinia .......................................................................... ................... 54
G enus Steeriana ................................................................... ..................... 54
G enus B revinenia .................................................................. .................... 55
G enus Z ilchiella ....................................................... .................................. 56
Ecological Rem arks ............................... ............................................................ 57
Acknowledgments ............................... ............................... ............. ........ 58
L literature C ited. ........................................................... .................................. ..... 59







I Wiesenstrasse 2, D-79410 Badenweiler, Germany, .
SRathenaustrasse 8, D-65326 Aarbergen-Rickershausen, Germany, .

Neubert, E., & H. Nordsieck. 2005. New South American Clausiliidae from the collections of the Florida Museum of Natural History (Gastropoda,
Clausiliidae, Neniidae). Bull. Florida Mus. Nat. Hist. 45(2): 45-62








INTRODUCTION
The knowledge on Neniinae, the subfamily ofClausiliidae
living in the neotropical realm, is still fragmentary. Large
parts of the potentially inhabited area are unknown. This
is especially true for Colombia, Venezuela, Ecuador and
Bolivia. The Neniinae of Peru are better understood since
they have been intensively studied by Weyrauch. In the
beginning of his engagement in Clausiliidae, Weyrauch
gave his material to Pilsbry (1945, 1949a, b), who de-
scribed several species from these collections. Later,
Weyrauch started a cooperation with Zilch and published
several articles on the Neniinae alone or together with
Zilch (Zilch, 1954; Weyrauch, 1957, 1963, 1964). His
collection of preserved animals as well as material from
additional sources was then used by Loosjes (Loosjes
& Loosjes-van Bemmel, 1966). Loosjes dissected many
species and used the morphology of the radula and the
genital organs altering Zilch's (1960) taxonomy of the
group, which was based on shell characters only. Re-
cently, the material of Loosjes (Loosjes & Loosjes-van
Bemmel, 1984, 1989) has been reinvestigated by
Nordsieck (1999), who also described new species from
the collection of Loosjes. The present study was neces-
sary to integrate a considerably high number of species
new to science, which were collected by F. G. Thomp-
son in northern Peru, especially in the departamentos of
Cajamarca, Amazonas, and Huanuco.

MATERIALS AND METHODS
Abbreviations: AH = height of aperture; AD = diameter
of aperture; D = diameter of shell; depto. =
departamento; H = height of shell; R, = rib number on 1
mm of the penultimate whorl; W = number of whorls.
Terms of the clausilial apparatus follow Pilsbry (1949b).
Collections:
ANSP = Academy of Natural Sciences Philadelphia
UF = Florida Museum of Natural History, Gainesville,
USA (F. G. Thompson = FGT).
NEUB = private collection Neubert.
NHM = Natural History Museum, London.
SMF = Senckenberg Museum, Frankfurt, Germany (R.
Janssen).

RESULTS
Genus Columbinia Polinski, 1924
1924 Nenia (Columbinia) Polinski, Bulletin de l'Acad&-
mie Polonaise des Sciences et des Lettres Cracovie,
(B) 1924: 743.

Subgenus Columbinia (Columbinia)
Columbinia (Columbinia) bryantwalkeri (Pilsbry, 1922)


BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 45(2)

1922 Nenia bryantwalkeri Pilsbry, The Nautilus,
35 (3): 95, pl. 2, figs. 4, 5, 12-14 [Peru, prov. of Hual-
laga].
Material.- UF 268765: Depto. Huanuco, 4.7 km S of
Tingo Maria, 750 m alt., FGT-1094.
Note.- The collection of Thompson contains
another specimen from a locality nearby, which is not
the same as C. bryantwalkeri. It is bigger (H = 19.9
mm, D = 4.6 mm) and more densely ribbed than the
latter. The preservation state of this specimen does not
allow a further judgment on its taxonomic status.
Material.- UF 268772: Depto. Huanuco, 9.2 km
S of Tingo Maria, 800 m alt., FGT-1102.

Columbinia (Steatonenia) Pilsbry, 1926
1926 Nenia (Steatonenia) Pilsbry, Proceedings of the
Academy of Natural Sciences Philadelphia, 78: 9.
Note.- The subgenus Steatonenia was redefined
by Nordsieck (1999: 172) and comprises the
species of Columbinia with ventricose shells.

Columbinia (Steatonenia) pachygastris n. sp.
(Fig. 1)
Diagnosis.- A Columbinia (Steatonenia) spe-
cies with moderately thick apical part of the shell, roundish
aperture and posterior lower palatal plica. Differs from
C. (S.)juninensis (M. Smith, 1943) by the more slen-
der apical part of the shell; aperture more roundish; in-
ferior lamella more spirally ascending; posterior lower
palatal plica more distinct; clausilium plate less pointed.
In comparison with C. (S.) bartletti (H. Adams, 1866)
and similar species shell with more slender apical part;
sculpture simple; inferior lamella more spirally ascend-
ing; posterior lower palatal plica present.
Description.- Shell with a blunt broadly conical
apical part, suture shallow; yellowish brown; teleoconch
whorls + regularly and obliquely rib-striated, somewhat
more coarsely on the neck; neck rounded; aperture de-
tached and protruding, obliquely oval or roundish-oval,
white inside, peristome somewhat expanded; superior
lamella high, continuous with the spiral lamella by a +
marked curve, occasionally somewhat elongated by its
side; inferior lamella relatively high in position, straightly
ascending with a weak spiral, of different height inwards,
in front ending on the columellar edge with a weak or
without thickening; subcolumellar lamella descending to
the lunellar; lunellar dorsal-dorsolateral in position, prin-
cipal plica ending dorsolaterally or more laterally; upper
palatal plica divergent from principal plica, continuous
with the lunella by a curve, posterior lower palatal plica
present, connected with the lunella by an obtuse angle;
clausilium plate in oblique view fully visible, relatively





NEUBERT & NORDSIECK: New South American Clausiliidae

broad, leaving a gap at the palatal side, distally blunt, wih
an inconspicuous outer corner. Spiral lamella penetrat-
ing inwards to a dorsolateral position, inferior lamella to
a ventral one (one specimen examined).
Type Material.- Peru: Depto. Piura, 23 miles E
of Canchaque on road to Huancabamba, 9400 ft. alt.,
K. Campbell leg., 11.04.1970. Holotype UF 268818.
Paratypes UF 298618 (1+1 fragment). Paratypes UF
268820 (2 complete shells, 6 fragments with aperture),
SMF 311954/1, same locality, 18.04.1970.
Measurements.- Holotype: H = 27.45 mm;
D = 6.85 mm; AH = 6.35 mm; AD = 6.3 mm; W = 8.
Paratypes (in mm, n = 4): H = 25.2 28.2 ; D = 6.4
7.3; W: 73/4- 82.
Etymology.- The specific epithet is derived from
Greek pachys (= thick), and Greek gastron (= venter,
belly).

Genus Incania Polinski, 1922
1922 Nenia (Incania) Polinski, Bulletin de l'Academie
Polonaise des Sciences et des Lettres Cracovie,
(B) 1921: 125.

Incania platystoma n. sp.
(Fig. 10)
Diagnosis.- A small-sized Incania species with
flared aperture with thickened lip and complete clausilial
apparatus with posterior lower palatal plica. In compari-
son with other Incania species superior lamella sepa-
rated from spiral lamella; posterior lower palatal plica
present.
Description.- Shell with relatively thick conical
apical part, suture shallow; yellowish reddish-brown;
teleoconch whorls densely rib-striated, more densely on
the lower whorls, on the neck more coarsely, riblets in
part white, at the suture patches of white riblets alter-
nating with non-white weakly sculptured ones, on the
upper whorls patches of white riblets in part giving rise
to protractive rib-folds; neck rounded, longitudinally en-
larged (in part with indistinct dorsal keel); aperture ad-
nate, oblong rhombic-oval, basally somewhat pointed,
brown inside, peristome much expanded and except in-
terlamellar part thickened, white, interlamellar covered
by a transparent callus; superior lamella high, widely
separated from the spiral lamella; inferior lamella steeply
ascending, low to moderately high inwards, in front end-
ing or bent on the columellar edge with a thickening,
occasionally continued to the peristome; subcolumellar
lamella descending to the lunellar; lunellar dorsolaterally
situated, principal plica ending about laterally, in front
far from the peristome; upper palatal plica continuous
with the oblique lunella by a curve or an angle, posterior


lower palatal plica present, short to nearly reduced, con-
nected with the lunella by an obtuse angle; clausilium
plate in oblique view nearly fully or fully visible, occa-
sionally leaving a gap at the palatal side, moderately broad,
distally rounded, in part with an inconspicuous outer cor-
ner. Spiral lamella penetrating inwards to a lateral posi-
tion, inferior lamella to the beginning of the penultimate
whorl (one specimen examined).
Type Material.- Peru: Depto. Amazonas, 11 km
SSW of Leimebamba, 2810 m alt., F. G. Thompson leg.,
25.04.1972 (FGT-1835). Holotype: UF 268805.
Paratypes: UF 298616 (6), SMF 311950/3. UF 268803:
paratype specimen ex UF 268805, radula removed by F.
G. Thompson.
Measurements.- Holotype: H 13.15 mm;
D = 3.25 mm; AH = 4.5 mm; AD = 3.2 mm; W = 7/2.
Paratypes (in mm, n = 9): H = 12.9 15.2; D = 3.3 -
3.6; W: 7/4 8.
Etymology.- The name platystoma is derived
from Greek plates (= broad, flat) and stoma (= mouth),
referring to the flat and recurved aperture.

Incania papillosa n. sp.
Incania papillosa papillosa n. sp. n. ssp.
(Fig. 8)
Diagnosis.--A medium-sized Incania species with
characteristic papillae and reduced clausilial apparatus.
Description.- Shell with a relatively slender coni-
cal or somewhat attenuated apical part, suture shallow;
yellowish reddish- to greyish-brown; teleoconch whorls
densely rib-striated, in part more densely on the lower
whorls, on the neck more coarsely, riblets in part white,
at the suture a brighter ribbon with white riblets, espe-
cially at the base of the upper whorls papilla-like nodes
as basal parts of indistinct rib-folds; neck rounded, some-
what longitudinally enlarged, umbilicus open; aperture
detached, occasionally adnate, oblong oval to rhombic-
oval, brown inside, peristome somewhat expanded, white;
superior lamella reduced to a weak callus at the peris-
tome; inferior lamella s-like ascending, ending or bent
on the columellar edge, not continued to the peristome;
other lamellae and plicae and clausilium missing. Infe-
rior lamella penetrating inwards to a lateral or ventrolat-
eral position (three specimens examined).
Type material.- Peru: Depto. Amazonas, 24 km
SSW of Leimebamba, 3370 m alt., F. G. Thompson leg.,
25.04.1972 (FGT-1834). Holotype: UF 268801.
Paratypes: UF 298614 (22 + 3 fragments), SMF 311951/
8, NEUB (3). UF 268802: paratype specimen ex UF
268801, radula removed by F. G. Thompson.
Measurements.- Holotype: H = 18.7 mm; D = 4.2
mm; AH = 5.1 mm; AD = 3.3 mm; W = 9/4. Paratypes







(in mm, n= 31): H = 16.5 21.0; D = 3.8 4.7; W (n=
20): 9- 10/4.
Etymology.- This species is calledpapillosa from
Latin papilla (= small wart), because of the suprasutural
papillae of its shell.

Incania papillosa imbecilla n. ssp.
(Fig. 9)
Diagnosis.- In comparison with I. p. papillosa
shell smaller, superior lamella well-developed, but infe-
rior lamella reduced.
Description.- Smaller and more ventricose than
I. p. papillosa; upper whorls with rib-folds, forming basal
nodes; aperture adnate; superior lamella well-developed,
short; inferior lamella reduced, minute s-like on the thick-
ened columellar edge.
Type Material.- Peru: Depto. Amazonas, 11 km
SSW of Leimebamba, 2810 m alt., F. G. Thompson leg.,
25.04.1972 (FGT-1835). Holotype: UF 298626. Paratype:
SMF 311964/1.
Measurements.- Holotype: H = 13.2 mm; D = 3.9
mm; AH = 3.85 mm; AD = 2.8 mm; W = 9. Paratype
(in mm): H = 12.6; D = 3.5; W: 8.
Etymology.- This subspecies is called imbecilla
from Latin imbecillus (= weak, weakened) for the re-
duced inferior lamella.

Genus Cyclonenia Nordsieck, 1999
1999 Cyclonenia Nordsieck, Basteria, 63 (4/6): 172.

Cyclonenia gibber n. sp.
(Fig. 2)
Diagnosis.- A Cyclonenia species with promi-
nent dorsal keel, brown aperture and with posterior but
without anterior lower palatal plica. In comparison with
C. cyclostoma (L. Pfeiffer, 1850) sculpture stronger (rib-
striation less fine and undulate); dorsal keel more promi-
nent; superior lamella constantly connected with spiral
lamella (without a curve, spiral lamella high); inferior
lamella lower.
Description (of type series).- Shell with some-
what attenuated apical part, suture shallow; reddish-
brown, apical part darker; teleoconch whorls densely
rib-striated, lower whorls in part more densely, on the
neck more coarsely, riblets in part white, at the suture
patches of white riblets alternating with non-white
weakly sculptured ones; neck with prominent dorsal keel,
divergent from the indistinct continued basal edge of the
body whorl, fading before the peristome, with neck fur-
row and sutural bulge, basal keel ill-defined; aperture
detached and protruding, obliquely oval, brown inside;
peristome expanded, lighter in colour; superior lamella


BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 45(2)

high, continuous with the spiral lamella without a curve;
inferior lamella straightly ascending with a weak spiral,
low to moderately high inwards, in front bent on the col-
umellar edge with a thickening; subcolumellar lamella
descending to the lunellar; lunellar dorsal-dorsolateral to
dorsolateral in position, principal plica ending nearly lat-
erally to ventrolaterally; upper palatal plica of different
length and divergent, continuous with the straight lunella
by a curve or an angle, posterior lower palatal plica
present, connected with the lunella by an obtuse angle;
clausilium plate in oblique view fully visible, relatively
narrow, leaving a gap at the palatal side, in part with an
outer corner.
Spiral lamella penetrating inwards to a nearly lateral
position, inferior lamella to a nearly ventral or ventral
one (three specimens examined).
Type Material.- Peru: Depto. Amazonas, 24 km
SSW of Leimebamba, 3370 m alt., F. G. Thompson leg.,
24.04.1972 (FGT-1832). Holotype: UF 268799.
Paratypes: UF 298620 (36 + some fragments), SMF
311955/5, NEUB (2).
Additional Material.- UF 268804: 11 km SSW of
Leimebamba, 2810 m alt., FGT-1835.
Measurements.- Holotype: H = 25.9 mm; D = 5.3
mm; AH = 5.3 mm; AD = 4.8 mm; W = 103/4. Paratypes
(in mm, n = 32): H = 19.9 25.4; D = 4.8 5.7; W
(n =20): 9/4- 11; R, (n =20): 11 18, mean 15.1. 11
km SSW of Leimebamba (in mm, n = 3): H = 19.1
20.2; D = 4.1 4.5; W: 9/2 93/4.
Etymology.- This species is called gibber from
Latin gibber (= humped).

Cyclonenia sanmarcos Grego & Szekeres, 2004
(Fig. 3)
2004 Cyclonenia sanmarcos Grego & Szekeres, Schrif-
ten zur Malakozoologie, 21: 7, fig. 3.
Description.-Shell with conical or attenuated api-
cal part, suture shallow; yellowish reddish-brown, apical
part in part darker; teleoconch whorls regularly densely
rib-striated, lower whorls more densely, on the neck more
coarsely, except at the suture riblets hardly white, at the
suture patches of white riblets alternating with non-white
weakly sculptured ones, on the lower whorls a spiral
sculpture visible; neck nearly rounded, with broad basal
keel and more distinct dorsal keel, both weakly devel-
oped, neck furrow and sutural bulge indistinct; aperture
detached and protruding, obliquely broad-oval, brown
inside, peristome expanded, white; superior lamella high,
continuous with the spiral lamella by a marked curve,
distinctly elongated by its side; inferior lamella straightly
ascending, in part with a weak spiral, low to moderately
high inwards, in front ending or bent on the columellar





NEUBERT & NORDSIECK: New South American Clausiliidae

edge with a thickening; subcolumellar lamella descend-
ing to the lunellar (as far as anterior lower palatal plica);
lunellar dorsal-dorsolateral to dorsolateral in position,
principal plica ending nearly laterally or laterally; upper
palatal plica continuous with lunella by a curve, anterior
and posterior lower palatal plicae present, both connected
with lunella, anterior lower palatal plica of different length,
posterior one long; clausilium plate in oblique view fully
visible, leaving a gap at the palatal side, relatively broad,
with an outer corner. Spiral lamella penetrating inwards
to a nearly lateral position, inferior lamella to a ventral
one (one specimen examined).
Material.- UF 268810, 268811, 298615, SMF
311956/1: Depto. Amazonas, 10 km SE of Tingo, 1820
malt., FGT-1839.
Measurements. FGT-1839 (in mm, n = 7):
H = 24.6 26.9; D = 6.0 6.6; W = 11 12; R, (n = 6):
11 15, mean 12.8.
Note. In comparison with C. cyclostoma (L.
Pfeiffer, 1850) shell bigger; sculpture different (rib-stria-
tion less undulate); dorsal keel weaker; inferior lamella
more spirally ascending; anterior lower palatal plica
present.

Cvclonenia albosuturalis n. sp.
(Fig. 15)
Diagnosis.- A Cvclonenia species which in com-
parison with other Peruvian species of the genus is char-
acterized by its small size, weakly developed dorsal keel
and deeply situated lunellar.
Description.- Shell with broad conical apical part,
suture of upper whorls deep, of lower whorls shallow;
yellowish reddish-brown or reddish-brown; teleoconch
whorls densely rib-striated, on the neck somewhat more
coarsely, riblets in part white, especially above and be-
low at the suture, at the suture patches of white riblets
alternating with non-white weakly sculptured ones, a
more or less pronounced white sutural thread present;
neck nearly rounded, with swollen basal keel and fili-
form dorsal keel continuing the indistinct basal edge of
the body whorl, both weak, sutural bulge distinct; ap-
erture detached and protruding, obliquely oval to oval-
piriform, brown inside, peristome somewhat expanded,
lighter in colour; superior lamella continuous with the
spiral lamella without a curve, in part elongated by its
side, lamellae at the connection lowered; inferior lamella
straightly ascending, of different height inwards, in front
ending or indistinctly bent on the columellar edge with
a thickening; subcolumellar lamella descending to the
lunellar, in oblique view hardly visible; lunellar
dorsolaterally or nearly laterally situated, principal plica
ending laterally or nearly ventrolaterally; upper palatal


plica continuous with lunella by a curve or an angle, pos-
terior lower palatal plica present, of different strength,
connected with the lunella by an obtuse angle; clausilium
plate in oblique view only partly visible, occasionally
nearly fully, with or without an indistinct outer corner.
Spiral lamella penetrating inwards to a dorsolateral posi-
tion, inferior lamella to the beginning of the penultimate
whorl (three specimens examined).
Type Material.- Peru: Depto. Cajamarca, 2.6 km
NE of Encafiada, 3650 m alt., F. G. Thompson leg.,
22.04.1972 (FGT-1815). Holotype: UF 268788.
Paratypes: UF 298609 (75 + apical fragments), SMF
311958/8, NEUB (4).
Measurements.- Holotype: H = 13.55 mm;
D = 3.5 mm; AH = 3.4 mm; AD = 2.75 mm; W = 8V2.
Paratypes (in mm, n = 60): H = 11.4 -14.7; D = 3.2 -
3.6; W (n= 15):8 9; R (n = 20): 13 20, mean 16.4.
Etymology.- This species is called albosuturalis
from Latin albus (= white) and sutura (= suture).

Genus Bequaertinenia Weyrauch, 1964
1964 Bequaertinenia Weyrauch, Acta Zoologica Lillo-
ana, 20: 150.
Note.- Supplemented diagnosis of the genus see
Nordsieck (1999: 177).

Bequaertinenia delicate n. sp.
(Fig. 14)
Diagnosis.- A slender Bequaertinenia species
with rib-folds on the whole teleoconch and pronounced
dorsal keel. In comparison with B. bequaerti (Weyrauch,
1957) shell more slender and delicate; with rib-folds and
more distinct sutural patches.
Description. Shell with attenuated apical part,
suture deep; yellowish reddish-brown; teleoconch whorls
densely rib-striated, lower ones in part more densely, on
the neck less densely, riblets more or less white, at the
suture patches of white riblets alternating with non-white
weakly sculptured ones, on the lower or on all whorls
the patches of white riblets giving rise to protractive rib-
folds; neck with indistinct basal keel and prominent fili-
form dorsal keel continuing the basal edge of the body
whorl, sutural bulge pronounced behind; aperture much
protruding, roundish-oval to rhombic-oval, brownish in-
side, peristome somewhat expanded, lighter in colour;
superior lamella high, continuous with the spiral lamella
by a weak curve, in part elongated by its side; inferior
lamella straightly ascending, moderately high to high in-
wards, in front ending or bent on the columellar edge;
subcolumellar lamella descending to the lunellar, in ob-
lique view hardly visible; lunellar dorsolaterally or nearly
laterally situated, principal plica ending laterally or nearly







ventrolaterally; upper palatal plica continuous with the
lunella by a curve or an angle, posterior lower palatal
plica present, of different length, connected with the
lunella by an angle; clausilium plate in oblique view nearly
fully visible, moderately broad, distally rounded, + with
an outer corer. Spiral lamella penetrating inwards to a
dorsolateral position, inferior lamella to a ventral one (one
specimen examined).
Type Material.- Peru: Depto. Cajamarca, 13 km
by road E of Celendin, 2980 m alt., F. G. Thompson leg.,
23.04.1972 (FGT-1822). Holotype: UF 268795.
Paratypes: UF 298622 (3), SMF 311957/1.
Measurements.- Holotype: H = 17.0 mm; D = 2.9
mm; AH = 3.15 mm; AD = 2.7 mm; W = 10/4. Paratypes
(in mm, n = 2): H = 14.3, 14.0; D = 3.0, 3.0; W = 9/4;
R, = 17, 19, 20.
Etymology.-This species is named from Latin
delicatus (= delicate).

Genus Hemicena Pilsbry, 1949
1949 Nenia (Hemicena) Pilsbry, Proceedings of the
Academy of Natural Sciences Philadelphia, 101: 229.

Hemicena polinskiana (Pilsbry, 1949)
1949 Nenia [Hemicena] polinskiana Pilsbry, Proceed-
ings of the Academy of Natural Sciences Philadel-
phia, 101: 229, pl. 20 figs. 1-lb.

Hemicena polinskiana ancashensis n. ssp.
(Fig. 7)
Diagnosis.- In comparison with H. p.
polinskiana (fig. 6) shell smaller and more coniform;
clausilial apparatus more reduced (no rudimentary spi-
ral lamella, inferior lamella more weakly developed).
Description.- Shell conical, apical part conical to
somewhat attenuated; yellowish-brown; teleoconch
whorls densely rib-striated, more coarsely on the neck,
at the suture nodes formed by bundles of white rib-striae,
giving rise to indistinct protractive rib-folds, rib-folds on
the neck strong; neck rounded; aperture detached, oc-
casionally adnate, broadly oval, yellowish-white within,
peristome expanded; superior lamella short, of different
strength; inferior lamella low, steeply ascending on the
columellar edge or reduced; other lamellae and plicae
and clausilium missing.
Type Material.- Peru: Depto. Ancash, Llupash,
3310m alt., F. G. Thompson leg., 18.03.1972 (FGT-1757).
Holotype: UF 210212. Paratypes: UF 298621 (11), SMF
323435/4.
Measurements.- Holotype: H = 18.1 mm;
D = 4.55 mm; AH = 4.55 mm; AD = 3.5 mm; W = 9/4.
Paratypes (in mm, n = 15): H = 16.4- 19.7; D= 4.3-


BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 45(2)

5.0; W = 8/4- 10/4.
Etymology.- This subspecies is called
ancashensis for its type locality which is situated in
Depto. Ancash.
Note.- Florida State Museum and SMF house
specimens of H. p. polinskiana (fig. 6) collected by
Weyrauch (UF 203136/5, 118936/6, SMF 62672/7,
89492/5, 199116/1). The new subspecies was compared
with the holotype and the above listed voucher speci-
mens from the type locality (Tapacocha, Rio Fortaleza,
at 3000 3500 m, on the western slope of the Andes
(inland from Pativilca in the northwestern part of Depto.
Lima)). From these lots it is evident that the spiral lamella
of the nominal subspecies (Pilsbry, 1949: 230) is some-
times very low and nearly invisible.

Genus Peruinia Polinski, 1922
1922 Nenia (Peruinia) Polinski, Bulletin de l'Acaddmie
Polonaise des Sciences et des Lettres Cracovie,
(B) 1921: 125.

Peruinia tingamariae (Pilsbry, 1922)
1922 Nenia flachi tingamariae Pilsbry, The Nautilus,
35 (3): 94, pl. 2, fig. 3 [Peru, Tinga Maria].
1957 Peruinia albicolor Weyrauch, Archiv fur Mollus-
kenkunde, 86 (1/3): 13, pl. 1, fig. 11 [Peru: Near en-
trance of a cave on the left side of Rio Tulumayo, af-
fluent of Rio Huallaga, near to the bridge at the road
from Huanuco to Pucallpa, 23 km NE of Tingo Maria,
670 m alt.].
Material.- UF 268776, SMF 323449/7: Depto.
Huanuco, 16.6 km NE ofTingo Maria, 800 m alt., FGT-
1106 (= P. albicolor); UF 268774: 4.6 km N of Tingo
Maria, 750 m alt., FGT-1103; UF 268767: 1.5 km SW of
Tingo Maria, 750 m alt., FGT-1096; UF 268763, 268764,
SMF 323448/10: 4.7 km S of Tingo Maria, 750 m alt.,
FGT-1094; UF 268769: 7.9 km S of Tingo Maria, 750 m
alt., FGT-1099; UF 268771, SMF 323450/3: 9.2 km S of
Tingo Maria, 800 m alt., FGT-1102; UF 268778:
Cayumba, 850 m alt., FGT-1110.
Note.- Because of the differences in genital mor-
phology (Loosjes and Loosjes-van Bemmel, 1966: figs.
6-10) P. tingamariae is separated from P. peruana
(Troschel, 1847) as an independent species. P. albicolor
is regarded as a colour form of P. tingamariae. Besides
the whitish shell colour in P. albicolor, no essential char-
acter states can be found to separate these two nominal
taxa.

Genus Pseudogracilinenia Loosjes & Loosjes-van
Bemmel, 1984
1984 Pseudogracilinenia Loosjes and Loosjes-van Be-





NEUBERT & NORDSIECK: New South American Clausiliidae

mmel, Zoologische Verhandelingen, Leiden, 212:10.

Pseudogracilinenia huallagana (Pilsbry, 1949)
1949 Nenia [Gracilinenia] huallagana Pilsbry, Proce-
edings of the Academy of Natural Sciences Philadel
phia, 101: 231, pl. 22, figs. 4-4f[Peru, Tingo Maria, in
a dense and rocky thicket at the west end of the Rio
Huallaga bridge to the airport].

Pseudogracilinenia huallagana huallagana
(Fig. 17)
Material.- UF 268766, SMF 323440/10: Depto.
Huanuco, 4.7 km S of Tingo Maria, 750 m alt., FGT-
1094; UF 268768: 5 km W of Tingo Maria, 750 m alt.,
FGT-1097.

Pseudogracilinenia huallagana amoena n. ssp.
(Fig. 18)
Diagnosis.- In comparison with P. h. huallagana
(fig. 17) shell bigger, more densely ribbed; superior lamella
higher.
Description.- Shell decollated, apical part at-
tenuated, suture deep, whorls more or less shouldered;
yellowish-brown; uppermost whorls of teleoconch
densely rib-striated, following whorls densely ribbed,
riblets in part white, patches of white riblets alternating
with non-white weakly sculptured ones, patches on lower
whorls with more riblets and extended over the whole
whorl, sculpture on the neck more irregular and in part
stronger; neck rounded; last half of body whorl detached
from penultimate whorl, aperture much protruding, round-
ish-oval, yellowish inside, peristome not expanded; su-
perior lamella continuous with spiral lamella by a more
or less weak curve, lamellae at the connection lowered;
inferior lamella high in position (near to superior and spi-
ral lamella), ascending with a spiral, relatively high in-
wards, not continued to the peristome; subcolumellar
lamella descending to the lunellar, + connected with the
lunella, in oblique view visible far inwards; lunellar dor-
sal-dorsolateral in position, principal plica ending
dorsolaterally, in part more laterally; upper palatal plica
short, upper part of lunella and upper palatal plica
weakened or nearly missing; clausilium plate in oblique
view fully visible, occasionally leaving a gap at the pala-
tal side, relatively broad, distally pointed, with an indis-
tinct outer corer. Spiral lamella penetrating inwards to
a nearly dorsolateral position, inferior lamella to a nearly
ventral one (one specimen examined).
Type Material.- Peru: Depto. Huanuco, 14.9 km
NE of Tingo Maria, 800 m alt., F. G. Thompson leg.,
17.03.1969 (FGT-1105). Holotype: UF 268775.
Paratypes: UF 298610 (15), SMF 323441/3.


Measurements.- Holotype: H = 15.0 mm; D = 3.1
mm; AH = 2.34 mm; AD = 2.16; W = 10 (less decol-
lated, from decollation closure W = 7). Paratypes (in mm,
n 18): H = 11.5 14.2 (- 15.0, if less decollated);
D = 2.9 3.2; W (n = 14): 6 7/2 (- 10, if less decol-
lated); R, (n = 18): 10- 18, mean 13.5.
Etymology.- The subspecies is called amoena
from Latin amoena (= pleasant, delightful), because of
the nice appearance of the shell.
Note.- The new subspecies was compared with
the holotype and voucher specimens of P h. huallagana
from the type locality (SMF 62674/14, 89493/5, 199114/
1). The investigated populations of the nominal subspe-
cies display no noteworthy variation in shell characteris-
tics. They are recorded from areas south and west of
Tingo Maria, while the new subspecies comes from the
northeast of Tingo Maria.

Genus Temesa H. & A. Adams, 1855
1855 Balea (Temesa) H. & A. Adams, The genera of
recent Mollusca, 2: 175.
Note.- The genus Temesa was redefined and dis-
cussed by Nordsieck (1999: 177).

Temesa clausilioides (Reeve, 1849)
(Figs. 24-25)
1849 Bulimus clausilioides Reeve, Conchologia Iconica,
5: pl. 73, sp. 523 [Peru, Andes of Caxamarca].
Note.- The type species of Temesa has not been
found again since its description and is only known by
three syntypes in NHM. Therefore, the sample collected
by Thompson is described; one syntype (fig. 24) and a
specimen from this sample (fig. 25) are figured.
Description.- Shell conical, with conical apical
part, suture of upper whorls deep, whorls in part shoul-
dered by a blunt edge, aperture in part shifted to the left;
greyish-brown or bluish-grey, protoconch brown;
teleoconch whorls densely rib-striated, on the neck in
part more coarsely, riblets white, especially at the su-
ture, greater part of the shell covered with bluish-grey
surface layer, with a spiral sculpture, especially on the
neck; neck nearly rounded, basal edge of body whorl
running to the peristome like an indistinct dorsal keel,
occasionally with sutural bulge, umbilicus open; aper-
ture + protruding, in part adnate, oval to triangular-piri-
form, brown inside, peristome somewhat expanded,
lighter in colour; superior lamella reduced to a weak callus
near to the peristome or missing, inferior lamella weak,
steeply ascending on the columellar edge; other lamel-
lae and plicae and clausilium missing.
Material. UF 268783, SMF 323444/10: Depto.
Cajamarca, 15 km SW of Encafiada, 3110 m alt., FGT-







1807.
Measurements.- FGT-1807 (in mm, n = 104):
H= 11.3 15.9; D = 3.0 4.0; W (n = 18): 7- 8/2.

Temesa gradata n. sp.
(Fig. 23)
Diagnosis.- A Temesa species with reduced
clausilial apparatus with white surface layer, shouldered
whorls and distinct rib-striation. In comparison with T.
incarum Pilsbry, 1926 and T. bicolor Pilsbry, 1949 shell
less conical; whorls more rounded (shouldered); sculp-
ture stronger; aperture mostly detached.
Description.- Shell + conical, with broad conical
apical part, suture deep, whorls much rounded (shoul-
dered), especially upper ones; bluish greyish-brown,
protoconch brown; teleoconch whorls densely ribbed,
lower ones and on the neck more densely and irregu-
larly, ribs white, in part white surface layer also between
the ribs, thus intervals bluish-white to brown, with a spi-
ral sculpture; neck nearly rounded; aperture detached,
in part adnate, oval, brown within, peristome not ex-
panded; superior lamella reduced to a diffuse callus near
to the peristome or missing; inferior lamella weak, steeply
ascending on the columellar edge or indistinct; other
lamellae and plicae and clausilium missing.
Type material.- Peru: Depto. Cajamarca, 30 km
NE of Encafiada, 3550 m alt., F. G. Thompson leg.,
22.04.1972 (FGT-1818). Holotype: UF 268790.
Paratypes: UF 298625 (6), SMF 311960/3.
Measurements.- Holotype: H = 12.45 mm;
D = 3.1 mm; AH = 3.1 mm; AD = 2.45 mm; W = 72.
Paratypes (in mm, n = 9): H = 11.5 13.2; D = 3.2 -
3.7; W = 7- 8; R, (n = 10): 9/2- 12, mean 10.9.
Etymology.- This species is called gradata from
Latin gradatus (= stepped) for the stepped whorls of its
shell.

Temesa pilsbryi Weyrauch, 1956
1956 Temesapilsbryi Weyrauch, Archiv fur Mollusken-
kunde, 85 (4/6): 146, pl. 11, figs. 1-4.

Temesa pilsbryi perfectecostata n. ssp.
(Fig. 22)
Diagnosis.- In comparison with T.p. pilsbryi (fig.
21) whorls more rounded; sculpture stronger.
Description (of type series).- Shell conical-fusi-
form, apical part slender conical, suture deep, especially
upper whorls much rounded; yellowish- to greyish-
brown, protoconch brown; teleoconch whorls coarsely
ribbed, on the neck more irregularly, ribs white, inter-
vals finely striated; neck nearly rounded, umbilicus
open; aperture detached and protruding, obliquely oval,


BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 45(2)

yellowish inside, peristome not expanded; superior
lamella a diffuse callus near to the peristome or missing;
inferior lamella on the columellar edge indistinct or miss-
ing; other lamellae and plicae and clausilium missing.
Type Material.- Peru: Depto. Ancash, 5 km SW
of Chavin de Huantar, 3300 m alt., F. G. Thompson leg.,
02.04.1972 (FGT-1771). Holotype: UF 268781.
Paratypes: UF 298617 (22), SMF 311961/4, NEUB (2).
Additional material.- UF 268782: 5 km SW of Chavin
de Huantar, 3150 m alt., FGT-1782.
Measurements.- Holotype: H = 13.3 mm;
D = 2.85 mm; AH = 2.75 mm; AD = 2.05 mm;
W = 10/4. Paratypes (in mm, n = 16): H = 11.6 13.7;
D = 2.7 3.2; W = 9/4 11; R, (n = 20): 3'/ 8, mean
5.9. 5 km SW of Chavin de Huantar, 3150 m alt. (in
mm, n= 25): H = 10.5 13.3; D = 2.6 3.0; W (n = 12):
8/4 -10; R, (n = 20): 4 9'/, mean 6.4.
Etymology.- This subspecies is called
perfectecostata from Latin perfectus (= perfect) and
costa (= rib) because of the even and straight ribs of its
shell.
Note.- The new subspecies was compared with
the holotype, paratypes and additional voucher speci-
mens of T.p. pilsbryi from the type locality (El Infiernillo,
right side of Rio Rimac, 3360 3370 m alt.) (Holotype
T p. pilsbryi SMF 155296; paratypes SMF 155297/2,
155298/1,155299/7, 156280/50, 162116/100; additional
specimens SMF 199139/11,209347/14,264391/10).

Temesa parcecostata (Polinski, 1922)
1922 Neniaparcecostata Polinski, Bulletin de l'Acad-
mie Polonaise des Sciences et des Lettres Cracovie,
(B) 1921: 130 [Peru]
Material.- UF 268773: Depto. Huanuco, 9.2 km
S of Tingo Maria, 800 m alt., FGT-1102.

Genus Incaglaia Pilsbry, 1949
1949 Nenia (Incaglaia) Pilsbry, Proceedings of the Aca-
demy of Natural Sciences Philadelphia, 101: 215.
Note.- This genus was defined and delimited by
Nordsieck (1999: 178).

Incaglaia huanucensis (Pilsbry, 1949)
1949 Nenia [Neniatracta] angrandi huanucensis Pil-
sbry, Proceedings of the Academy of Natural Scien-
ces Philadelphia, 101: 228, pl. 21, figs. 4-4c, 5. [Peru,
Huanuco, 1900 m alt.].
Material.- UF 268757: Depto. Huanuco, 13.1 km
N of Huanuco, 6100 ft. alt., FGT-1038; UF 268756: 13
km N of Huanuco, 6200 ft. alt., FGT-1037; UF 268760:
9.7 km NE of Huanuco, 1900 m alt., FGT-1076; UF
268755, SMF 323445/10:9.5 km N of Huanuco, 6300 ft.





NEUBERT & NORDSIECK: New South American Clausiliidae

alt., FGT-1034; UF 268754, SMF 323446/10: 6.9 km N
of Huanuco, 6200 ft. alt., FGT-1029; UF 268753: 2.3 km
N of Huanuco, 6200 ft. alt., FGT-1028; UF 268761: 3.1
km W of Huanuco, 1900 malt., FGT-1082; UF 268758:
0.4 km N ofAmbo, 6800 ft. alt., FGT-1039; UF 268759:
7.4 km S ofAmbo, 7200 ft. alt., FGT-1040; UF 268780:
7.7 km S ofAmbo, 2300 m alt., FGT-1114; UF 268750,
SMF 323447/8: 6.1 km S of Vichaycoto, 6300 ft. alt.,
FGT-1024; UF 268751: 2.5 kmN ofTomayquichua, 6500
ft. alt., FGT-1026; UF 268752: 1.7 km S of
Tomayquichua, 6500 ft. alt., FGT-1027; UF 268762: 4.4
km NW of Tambillo, 1900 m alt., FGT-1089.

Incaglaia leucostoma n. sp.
(Fig. 19)
Diagnosis. Differs from the other species of the
genus by the following character states: shell with pro-
nounced dorsal keel; inferior lamella high in position, very
low; rudimentary anterior lower palatal plica present.
Description. Shell with conical or somewhat at-
tenuated apical part; yellowish-brown; teleoconch whorls
densely rib-striated, lower ones in part somewhat more
densely, on the neck much more coarsely, rib-striation
between the keels and in the neck depression fading,
riblets largely white, at the suture patches of white riblets
alternating with non-white weakly sculptured ones; neck
with rounded basal keel and more prominent dorsal keel,
the latter long and diverging from the basal keel behind,
with distinct neck depression and sutural bulge, umbili-
cus open; aperture much protruding, oblique broadly
oval to roundish, yellowish inside, peristome expanded
and somewhat thickened, white; superior lamella high,
continuous with the spiral lamella by a marked curve, in
part indistinctly elongated by its side, lamellae at the con-
nection much lowered, spiral lamella low, principal plica
close to spiral lamella, in front elongated into the sinulus;
inferior lamella high in position, ascending with a spiral,
very low inwards, far inwards very close to the spiral
lamella, in front more or less indistinct, mostly bent and
continued with two branches to the peristome;
subcolumellar lamella descending to the lunellar, ap-
proaching the lunella, in oblique view visible far inwards;
lunellar nearly dorsally or dorsally situated, principal plica
ending nearly dorsolaterally to nearly laterally; upper
palatal plica continuous with lunella by a curve, lunella
thick below, ending with a spur-like anterior lower platal
plica, posterior lower palatal plica short to reduced;
clausilium plate in oblique view fully visible, relatively
broad, distally narrower, palatal edge distally bent up,
with a differently pronounced outer corner. Spiral lamella
low, penetrating inwards to a dorsal position, inferior
lamella to a lateral or ventrolateral one (two specimens


examined).
Type Material.- Peru: Depto. Cajamarca, 25 km
E of Celendin, 2510 m alt., F. G. Thompson leg.,
23.04.1972 (FGT-1824). Holotype: UF 268797.
Paratypes: UF 298624 (35 + some fragments), SMF
311959/6, NEUB (3).
Measurements.- Holotype: H = 15.75 mm;
D = 3.7 mm; AH = 3.45 mm; AD = 3.3 mm; W = 10.
Paratypes (in mm, n = 39): H = 13.2 17.5; D = 3.3
4.0; W (n = 20): 82 10; R (n = 20): 11/2 16, mean
13.6.
Etymology.- This species is called leucostoma
from Greek leukos (= white) and stoma (= mouth).

Genus Cvlindronenia Ehrmann in Pilsbry, 1949
1949 Nenia (Cvlindronenia) Ehrmann in Pilsbry, The
Nautilus, 62 (3): 104.
Note.- Cvlindronenia is frequently cited with
the authorship of Ehrnann in Zilch, 1949, Archiv fir
Molluskenkunde. 78 (1/3): 95 (15th June 1949). This
publication of Zilch was based on unpublished manu-
scripts of Ehrrnann, and Zilch remarks in a footnote that
he received the printed copy of The Nautilus, 62 (3)
already on 19th April 1949. In fact, this volume was
issued in January 1949 and predates Zilch's work by
half a year. The note on Cvlindronenia in The Nautilus
is authored by "P. Ehrmann, Ms", and it is very likely (as
Zilch also points out) that Pilsbry took the opportunity to
publish Ehrmann's description posthum. Thus we con-
sider the correct citation of Cvlindronenia as Ehrmann
in Pilsbry, 1949.
The relationship of Cvlindronenia and Steeriana
was discussed by Nordsieck (1999: 179), the classifica-
tion of Nenia cumulloana Pilsbry, 1949 with
Cilindronenia by Nordsieck (1999: 182).

Cvlindronenia cicatricosa (Loosjes & Loosjes-van
Bemmel, 1989)
1989 Steeriana (Cvlindronenia) cicatricosa Loosjes
and Loosjes-van Bemmel, Basteria, 53 (4/6): 84,
fig. 3 [Peru, dept. Amazonas, between Leimebam-
ba and Balsas].

Cvlindronenia cicatricosa cicatricosa
Material.- UF 268815: Depto. Amazonas, 21 km
ENE of Balsas, 1850 m alt., FGT-1856.

Cylindronenia cicatricosa leimebambensis H.
Nordsieck, 1999
1999 Cylindronenia cicatricosa leimebambensis H.
Nordsieck, Basteria, 63 (4/6): 180, figs. 13-14 [Peru,
dept. Amazonas, Leimebamba].







Material.- UF 268807, UF 268808, SMF 323438/
8: Depto. Amazonas, 11 km SSW of Leimebamba, 2810
m alt., FGT-1836.

Cylindronenia cumulloana (Pilsbry, 1949)
1949 Nenia [Cylindronenia] cumulloana Pilsbry, Pro-
ceedings of the Academy of Natural Sciences Phila-
delphia, 101: 224, pl. 20, figs. 3-3f [Road from Caja-
marca to Celendin at kilometer 73, in the Cordillera
de Cumulloa].
Material.- UF 268789, 268791, SMF 323439/5:
Depto. Cajamarca, 30 km NE of Encafiada, 3550 m alt.,
FGT- 1818; UF 268792:25 km SW of Celendin, 3300 m
alt., FGT-1819; UF 268812: 3 km E of Celendin, 2800 m
alt., FGT-1820; UF 268794:3 km ESE of Celendin, 2800
malt., FGT-1821; UF 268796, SMF 311965/3: 13 km by
road E of Celendin, 2980 m alt., FGT-1823; UF 268798,
298611, SMF 311966/8: 25 km E of Celendin, 2510 m
alt., FGT-1825.

Cylindronenia violacea n. sp.
(Fig. 16)
Diagnosis.- A small Cylindronenia species with
rib-folds on the whole teleoconch and protruding aper-
ture. In comparison with C. cumulloana (Pilsbry, 1949)
shell with strong rib-folds on all whorls; neck nearly
rounded; aperture more protruding; inferior lamella in
part nearer to superior and spiral lamella.
Description.- Shell decollated, suture shallow; yel-
lowish reddish-brown; teleoconch whorls densely rib-
striated, lower ones in part more densely, riblets partly
white, at the suture indistinct patches of white riblets
alternating with non-white weakly sculptured ones, giv-
ing rise to oblique protractive rib-folds on all whorls, more
distinct on the neck forming nodes on the dorsal keel;
body whorl compressed, neck nearly rounded (dorsal
keel indistinct, with sutural bulge behind); aperture de-
tached and far protruding, obliquely piriform or rhombic
oval, violet-brown inside, peristome expanded, light in
colour; superior lamella high, continuous with the spiral
lamella by a curve, occasionally elongated by its side;
inferior lamella straightly ascending with a weak spiral,
moderately high inwards, in front ending or bent on the
columellar edge with a thickening, occasionally contin-
ued to the peristome; subcolumellar lamella descending
to the lunellar, not or indistinctly connected with it; lunellar
dorsolateral, occasionally dorsolateral-lateral in position,
principal plica ending laterally or more ventrolaterally;
upper palatal plica divergent, in part long, continuous to
the lunella by an angle, occasionally posterior lower pala-
tal plica present; clausilium plate in oblique view fully
visible, leaving a gap at the palatal side, relatively broad,


BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 45(2)

with an indistinct outer corer. Spiral lamella penetrat-
ing inwards to a ventrolateral position, inferior lamella
nearly to the beginning of the penultimate whorl (one
specimen examined).
Type Material.- Peru: Depto. Huanuco, 10.8 km
W of Huancapallac, 2950 m alt., F. G. Thompson leg.,
21.03.1969 (FGT-1111). Holotype: UF 268779.
Paratypes: UF 298612 (7 + 1 fragment), SMF 311962/1.
Measurements.- Holotype: H = 15.95 mm;
D =3.75 mm; AH =4.15 mm; AD = 3.5 mm; W = 6/2
decollatee). Paratypes (in mm, n = 7): H = 14.2 16.7;
D =3.7-4.1; W=6 -7/2; R, (n = 9): 12 18, mean
14.5.
Etymology.- This species is called violacea from
Latin violaceus (= violet) for the remarkably coloured
aperture of its shell.

Genus Andinia Polinski, 1922
1922 Nenia (Andinia) Polinski, Bulletin de l'Acad6mie
Polonaise des Sciences et des Lettres Cracovie,
(B) 1921: 124.

Andinia taczanowskii (Lubomirski, 1879)
1879 Nenia taczanowskii Lubomirski, Proceedings of
the Zoological Society London, 47: 726, pl. 56, figs. 3,
4 [Peru, Bambamarca].
Material.- UF 210197 210200, SMF 323436/
10: Depto. Cajamarca, 7 km by road SW of
Bambamarca, 2920 m alt., FGT-1804.

Genus Steeriana Jousseaume, 1900
1900 Steeriana Jousseaume, Bulletin de la Soci6t6 Philo-
matique Paris, (9) 2: 34.

Steeriana malleolata (Philippi, 1867)
1867 Clausilia malleolata Philippi, Malakozoologische
Bl3itter, 14 : 194, Taf. 2, Fig. 3-4 [Peru, between Con-
tumaza and Cajamarca].
Material.- UF 268785,268786, SMF 323442/10:
Depto. Cajamarca, 2 km SW of Encafiada, 3040 m alt.,
FGT- 1809.

Steeriana cajamarcana Weyrauch & Zilch, 1954
1954 Steeriana (S.) cajamarcana Weyrauch & Zilch,
Archiv fur Molluskenkunde, 83 (1/3): 73, pl. 5, fig. 8,
text-fig. 6.

Steeriana cajamarcana sororminor n. ssp.
(Fig. 13)
Diagnosis.- In comparison with S. c.
cajamarcana (fig. 11) shell smaller and more slender;
rib-striation stronger; peristome detached; inferior lamella





NEUBERT & NORDSIECK: New South American Clausiliidae

high, simply arched.
Description.- Shell decollated, suture shallow; yel-
lowish-white; teleoconch whorls densely rib-striated,
lower ones more densely, on the neck more coarsely,
riblets white, at the suture patches of white riblets alter-
nating with non-white weakly sculptured ones; neck
nearly rounded, both keels weakly developed, umbilicus
open; aperture detached and protruding, obliquely piri-
form, whitish inside; superior lamella moderately high,
widely separated from the short spiral lamella; inferior
lamella straightly ascending, arched, high inwards, in front
ending on the columellar edge; subcolumellar lamella de-
scending to the lunellar, relatively well visible; lunellar
nearly dorsal to dorsal in position, principal plica ending
nearly dorsolaterally, in front far from the peristome;
upper palatal plica continuous to the lunella by an angle,
both strongly developed, without posterior lower palatal
plica; clausilium plate in oblique view fully visible, not
filling the lumen of the body whorl, more or less narrow
and pointed.
Type Material.- Peru: Depto. Cajamarca, 2 km
SW of Encafiada, 3040 m alt., F. G. Thompson leg.,
21.04.1972 (FGT-1809). Holotype: UF 268784.
Paratypes: UF 298623 (2).
Measurements.- Holotype: H = 9.8 mm; D = 3.2
mm; W = 4 (decollated). Paratypes (in mm, n = 2):
H = 9.4, (9.0); D = 2.9, 2.9; W = 4'4.
Etymology. The subspecies is called sororminor
from Latin soror (= sister) and minor (= smaller), as it
looks like the smaller sister of S. c. cajamarcana.
Note.- The new subspecies was compared with
the holotype, paratypes and additional voucher speci-
mens of S. c. cajamarcana from the type locality (Ha-
cienda Cochambul, 18 km from Cajamarca, at the road
to San Marcos, 2850 m alt.) holotypee S. c.
cajamarcana SMF 135516; paratypes SMF 69818/14,
139782/1, 157914/1; additional specimens SMF 199131 /
4,264382/10).

Steeriana sorormajor n. sp.
(Fig. 12)
Diagnosis.- A large Steeriana species with weak
dorsal keel and clausilium plate filling nearly the lumen
of the body whorl. In comparison with S. c.
cajamarcana Weyrauch & Zilch, 1954 shell bigger;
sculpture somewhat stronger, sutural patches distinct;
dorsal keel somewhat stronger; aperture detached; su-
perior lamella widely separated from spiral lamella; in-
ferior lamella higher; clausilium plate nearly filling the
lumen of body whorl. S. malleolata (Philippi, 1867) has
a more ventricose shell, with distinct rib-folds and pro-
nounced keels; inferior lamella lowered in front;


clausilium plate narrower.
Description. Shell decollated, suture shallow, that
of upper whorls deeper; greyish-white; teleoconch whorls
densely rib-striated, lower ones hardly more densely, less
undulate, on the neck somewhat coarser, more undu-
late, riblets white with interruptions, at the suture patches
of white riblets alternating with reddish-brown weakly
sculptured ones, on the upper whorls indistinct protrac-
tive rib folds; neck nearly rounded (dorsal keel dis-
tinct, filiform, basal keel swollen), umbilicus open; ap-
erture detached and protruding, rhombic-oval, brown
inside, peristome somewhat expanded, white; superior
lamella of different height, short, widely separated from
the spiral lamella; inferior lamella straightly ascending,
arched, high inwards, in front ending on columellar edge
with distinct thickening; subcolumellar lamella descend-
ing to the lunellar; lunellar dorsal to dorsal-dorsolateral
in position, principal plica ending laterally or more vent-
rolaterally, in front far from the peristome; upper palatal
plica continuous to the lunella by a curve or an angle,
lunella strongly developed, posterior lower palatal plica
a basal plica-like broadening of the lunella; clausilium
plate in oblique view nearly fully visible, mostly leaving a
gap at the palatal side, moderately broad, distally pointed,
forming an outer corner. Spiral lamella penetrating in-
wards to a lateral position, inferior lamella to a more
ventrolateral one (one specimen examined).
Type Material.- Peru: Depto. Cajamarca, 2 km
NE of Encafiada, 3200 m alt., F. G. Thompson leg.,
21.04.1972 (FGT-1811). Holotype UF 268787. Paratypes
UF 298607 (10), SMF 311963/3, NEUB (1).
Measurements. Holotype: H = 18.4 mm;
D = 5.65 mm; AH = 5.05 mm; AD = 4.05 mm; W = 5
(decollated). Paratypes (in mm, n = 11): H = 17.3 19.6;
D = 5.5 6.2; W (n = 13): 44 6; R, (n = 14): 12 18,
mean 15.4.
Etymology.- This species is called sorormajor
from Latin soror (= sister) and major (= bigger), as it
looks like the bigger sister of S. cajamarcana.

Steeriana celendinensis Weyrauch & Zilch, 1954
1954 Steeriana (S.) celendinensis Weyrauch & Zilch
in Zilch, Archiv fir Molluskenkunde, 83 (1/3): 70, Taf.
5, Fig. 6, Abb. 3 [Peru, 3 km SE Celendin, 2700 m
alt.].
Material.- UF 268793, 268813, SMF 323443/10:
Depto. Cajamarca, 3 km ESE of Celendin, 2800 m alt.,
FGT- 1821, 1852; UF 268814:4 km SE of Celendin, 2800
malt., FGT-1854.

Brevinenia n. gen.
Diagnosis.- Shell decollate; neck nearly rounded;







streaked; superior lamella continuous with or separated
from spiral lamella; subcolumellar lamella connected with
the lunella; lower palatal plicae missing. Differs from
Ehrmanniella Zilch, 1949 and Andiniella Weyrauch,
1958 by the fusion of the anterior end of the subcolumellar
lamella with the lunella (cf. Nordsieck, 1999: 182).
The genital organs of one specimen of the type
species (Balsas 65 km in direction to Leimebamba, 3350
m alt., leg. Hemmen) could be examined. It turned out
that the new genus is more closely related to
Ehrmaniella Zilch, 1949 than to Andiniella Weyrauch
1958. But the comparison with that genus is difficult,
because the descriptions and figures of the male copu-
latory organs (Loosjes & Loosjes van Bemmel 1966: 6-
8, figs. 2-3; 1984: 6, fig. 1) are insufficient (the demar-
cation of penis and epiphallus is unclear). The examined
genitalia are characterized as follows: Female genitalia
like in Ehrmaniella; male copulatory organs much dif-
ferent if compared to the cited figures: epiphallus con-
sisting of parts of differing width, most distal one run-
ning into penis with a marked narrowing, penis consist-
ing of two parts, proximal penis (seen from the gonad)
forming a loop held together by muscular strands.
Type Species.- Brevinenia richardsi (Grego &
Szekeres, 2004).

Brevinenia richardsi (Grego & Szekeres, 2004)
(Fig. 20)
2004 Andiniella richardsi Grego & Szekeres, Schriften
zur Malakozoologie, 21: 6, fig. 2.
Description.- Shell decollated, suture moderately
deep; yellowish reddish-brown to greyish-brown;
teleoconch whorls strongly rib-striated, lower ones more
densely, on the neck more coarsely, riblets more or less
white, especially on the upper whorls and the neck, at
the suture patches of white riblets alternating with brown
weakly sculptured ones; neck rounded, inflated, with
weak sutural bulge behind; aperture detached and pro-
truding, obliquely roundish-oval or oval, + brown inside,
peristome somewhat expanded, lighter in colour; supe-
rior lamella continuous with the spiral lamella by a more
or less marked curve, occasionally elongated by its side,
lamellae at the connection lowered; inferior lamella
high in position (near to superior and spiral lamella),
straightly ascending with a weak spiral, mostly high in-
wards, in front bent on the columellar edge with a
thickening; subcolumellar lamella descending to the
lunellar, anterior end fused with the lunella, in oblique
view farly visible inwards; lunellar in dorsal or more dor-
solateral position, principal plica ending nearly
dorsolaterally or dorsolaterally, in front elongated into
the sinulus; upper palatal plica continuous with lunella


BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 45(2)

by an angle or a curve, lunella strongly developed, ba-
sally continuous with the subcolumellar lamella;
clausilium plate in oblique view fully visible, in part leav-
ing a gap at the palatal side, moderately broad, distally
rounded or somewhat pointed, occasionally with an in-
distinct outer comer. Spiral lamella penetrating inwards
to a nearly dorsolateral or nearly lateral position, inferior
lamella to a nearly ventral or ventral one (two speci-
mens examined).
Material.- UF 268800,298619, SMF 311970/10,
NEUB (4): Depto. Amazonas, 24 km SSW of
Leimebamba, 3370 m alt., FGT-1834. UF 268806: 11
km SSW of Leimebamba, 2810 m alt., FGT-1836.
Measurements.- 24 km SSW of Leimebamba (in
mm, n = 57): H= 14.0 17.0; D = 3.7 4.3; W (n = 24):
44 63/4; RI (n = 20): 9 13, mean 11.2. 11 km SSW
of Leimebamba (in mm, n = 3): H = 15.1 15.8;
D 3.9 4.1; W= 54-6.
Note.- In comparison with B. dedicate
(Weyrauch & Zilch, 1954) shell more slender; sculpture
stronger, sutural patches more distinct; superior lamella
constantly connected with spiral lamella; inferior lamella
higher in position (near to superior and spiral lamella);
principal plica running farther to the peristome.

Brevinenia dedicate (Weyrauch & Zilch, 1954)
1954 Andinia (Ehrmaniella) dedicate Weyrauch &
Zilch in Zilch, Archiv fiLr Molluskenkunde, 83 (1/3):
68, Taf. 5, Fig. 5, Abb. 5 [Peru, valley of Rio Payan-
chan, at the road from Cerro de Pasco to Huanuco,
3800 m alt.].
Material.- UF 268770, SMF 323437/5: Depto.
Huanuco, 9.2 km S of Tingo Maria, 800 m alt., FGT-
1102; UF 268777, SMF 311969/7, NEUB (3): 27.1 km
SE of Aucayacu, 650 m alt., FGT-1107.

Genus Zilchiella Weyrauch, 1957
1957 Zilchiella Weyrauch, Archiv ftir Molluskenkunde,
86 (1/3): 9.

Zilchiella scala n. sp.
(Fig. 5)
Diagnosis.- A Zilchiella species with shouldered
whorls, weak sculpture and sharp dorsal keel. In com-
parison with Z. grandiportus (Weyrauch, 1957) shell
occasionally more slender, constantly decollated, sutural
edge of the whorls strong; sculpture weaker; dorsal keel
more pronounced (mainly behind).
Description. Shell decollated, suture shallow,
whorls with a blunt sutural edge = shouldered, becoming
weaker on the lower whorls, ending on the body whorl
as sutural bulge, aperture more or less shifted to the left;





NEUBERT & NORDSIECK: New South American Clausiliidae

yellowish reddish-brown to greyish-violet; teleoconch
whorls with very densely spaced weak rib-striae, more
distinct on the upper whorls, above the sutural edge and
on the neck, rib-striae white, mainly at suture and base
of the whorls, on the neck also on dorsal keel and
periomphalum, therefore whorls with diffuse white sur-
face layer, forming a white band at the base continued
on the dorsal keel; neck with sharp dorsal keel continu-
ing the pronounced basal edge of the body whorl, basal
keel indistinct, umbilicus open or closed; aperture de-
tached and far protruding, rectangular, with a chan-
nel formed by the dorsal keel, brown inside, peristome
expanded, lighter in colour; superior lamella a differently
weak oblique callus near to the peristome, occasionally
plica-like elongated; inferior lamella strong, not steeply
ascending, of different height, in front ending far from
the peristome; other parts of clausilial apparatus miss-
ing. Inferior lamella penetrating inwards to a lateral or
nearly ventral position (two specimens examined).
Type Material. Peru: Depto. Cajamarca, 20 km
ESE of Magdalena, 1830 m alt., F. G. Thompson leg.,
05.05.1972 (FGT-1859). Holotype: UF 268816.
Paratypes: UF 298613 (54 + 2 fragments), SMF 311967/
10, NEUB (5).
Measurements. Holotype: H = 21.9 mm; D = 7.1
mm; AH = 6.35 mm; AD = 4.35 mm; W = 6 decollatee).
Paratypes (in mm, n = 45): H = 16.1 -24.6; D= 5.5-
7.2; W (n = 18): 4/2 7/4.
Etymology. This species is named from Latin
scala (= stair, staircase) for the characteristic form of
its shell.

Zilchiella palatalis n. sp.
(Fig. 4)
Diagnosis.- A Zilchiella species with character-
istic sculpture, sharp dorsal keel and with subcolumellar
lamella and palatal plica. In comparison with Z.
grandiportus (Weyrauch, 1957) shell more slender,
constantly decollated, lighter in colour; sculpture coarser,
rib-striae in part elevated to hollow ribs; dorsal keel more
pronounced (mainly behind); inferior lamella less steeply
ascending; subcolumellar lamella and long palatal plica
present.
Description.- Shell decollated, suture shallow, ap-
erture in part shifted to the left; yellowish-brown;
teleoconch whorls densely rib-striated, rib-striae partly
white, some elevated mainly at the suture and base of
the whorls to hollow ribs, prominent on the dorsal keel,
upper whorls with rib-folds; neck with sharp dorsal keel,
continuing the pronounced basal edge of the body whorl,
basal keel indistinct, umbilicus open; aperture detached
and far protruding, rectangular, with a channel formed


by the dorsal keel, brownish inside, peristome expanded,
lighter in colour; superior lamella a weakly developed
node near to the peristome or missing; inferior lamella
not steeply ascending, high, in front ending far from the
peristome; subcolumellar lamella present, steeply ascend-
ing, only visible in oblique view; palatal plica in the middle
of the palatal wall present, running inwards from a dor-
sal position of the body whorl to the penultimate whorl;
other lamellae and plicae and clausilium missing. Infe-
rior lamella penetrating inwards to the penultimate whorl
(one specimen examined).
Type material.- Peru: Depto. Cajamarca, 2 km E
of Magdalena, 1280 m alt., F. G. Thompson leg.,
05.05.1972 (FGT-1861). Holotype: UF 268817.
Paratypes: UF 298608 (3), SMF 311968/1.
Measurements.- Holotype: H = 22.8 mm; D = 5.5
mm; AH = 4.9 mm; AD = 4.0 mm; W = 7% decollatee).
Paratypes (in mm, n = 2): H = 22.2, 21.5; D = 5.5, 5.0;
W (n = 4): 7 72.
Etymology.- This species is called palatalis for
the unusual presence of a palatal plica in a Zilchiella
species.

ECOLOGICAL REMARKS
As in many groups of snail species worldwide, precise
information on basic environmental factors influencing
the habitats as well as autecological observations are
missing. For the species of Clausiliidae mentioned here
we are able to present some of these data, because F.
Thompson provided us with his notes taken in the field.
Although not overall complete they offer a basic under-
standing of the habitats inhabited by these species.
Browsing the field notes for information on the
substrata it becomes obvious that the majority of the
species live on limestone or marbles, a fact, which
matches the observations in many parts of the world. A
remarkable exception is Incaglaia huanucensis, which
is exclusively recorded from non-limestome areas (10
places). For this species, substrata like gneisses and
schists are characteristic in combination with a xeric
environment. The vegetation of these habitats mainly
consists of Agavacea, Bromeliacea (Tillandsia) and
Cactacea (Opuntia), and the specimens were recorded
from under leaves, grasses, stones and ground slabs.
The altitudes of these habitats range from 750 to 2300
m. Another species recorded from non-limestone sub-
stratum is Hemicena polinskiana ancashensis from
Llupash, Depto. Ancash. The substratum consisted of
grants and basalts in an altitude of 3310 m with some
trees in ravine, but a mostly herbaceous vegetation.
An interesting combination of species can be found
in lowland habitats around Tingo Maria (Depto.





BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 45(2)


Table 1. Sympatric occurences of species of Neniinae.

CC 00 '- r1 in
Co Cl Cl C C
00 00 o W )-
oo oo oo oo
Species/Locality 0 0 0 0 -4 -


Columbinia bryantwalkeri x x
Incania platystoma x
Incania p. imbecilla x
Cyclonenia gibber x
Bequaertinenia delicate x
Peruinia tingamariae x x
Pseudogracilinenia h. huallagana x
Temesa gradata x
Temesa parcecostata x
Incaglaia leucostoma x
Cylindronenia c. leimebambensis x
Cylindronenia cumulloana x x x x
Steeriana malleolata x
Steeriana c. sororminor x
Steeriana celendinensis x
Brevinenia dedicate x
Brevinenia richardsi x


Huanuco). The area is characterized by wet tropical
forests or often with second growth trees and bushes.
The key species for this area seems to be Peruinia
tingamariae, which was recorded from 7 out of 10
places. It was found to live in sympatry with Columbinia
bryantwalkeri (FGT-1094, FGT-1102), and
Pseudogracilinenia huallagana huallagana (FGT-
1094). At FGT-I 102 (9.2 km S of Tingo Maria, 800 m
alt.), it was found together with Columbinia
bryantwalkeri, Temesa parcecostata, and Brevinenia
dedicate (Tab. 1).
Most other species originate from mesic to
submesic limestone habitats in considerably high altitudes
ranging from 2500 to 3600 m. The potential vegetation
seems to consist of low shrubs with small interspersed
tufts of trees, the latter under more favourable condi-
tions in ravines etc. and grassy hillslopes. Many of these
areas visited by Thompson are more or less intensively
cultivated and thus, their vegetation has changed to a
type of vegetation dominated by grass. Still, many spe-
cies of Clausiliidae persist under this environmental pres-
sure, and in several cases even sympatric occurence
can be found (Tab. 1, FGT-1809, 1820/1821, 1824/1825,
1835/1836). The Zilchiella species both occur in lower


altitudes, Zilchiella scala from a limestone cliff at 1830
m, and Zilchiella palatalis from slabs and sandstones
above road cut at 1280 m.
No other observations concerning autecological
topics were recorded. Temesa pilsbryi perfectecostata
and Cyclonenia albosuturalis were found actively
crawling on wet mosses and limestone rubble. Others
aestivated under dead leaves or were found attached to
bare rock faces. All species found by Thompson seem
to belong to the ecotype of rock-dwelling clausiliids. Tree
living species or species living in decaying wood were
not recorded yet.

ACKNOWLEDGEMENTS
The authors are grateful to F. G. Thompson for the pos-
sibility to work on the material he collected during his
trips to South America. We are much indebted to the
curators of all institutions holding type material of
Neniinae, whose fast response and deliberate actions
made it possible to conduct this study (in alphabetical
order): P. Bouchet (Paris), E. Gittenberger (Leiden), R.
Hershler (Washington), R. Janssen (Frankfurt), P.
Mordan (London), A. Riedel (Warszawa), G. Rosenberg
(Philadelphia). Special thanks go to R. Janssen,





NEUBERT & NORDSIECK: New South American Clausiliidae

Forschungsinstitut Senckenberg, for giving us the chance
to use the museum's facilities such as the library, collec-
tion, and technical equipment.

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Handbuch der Paldozoologie, 6(2)(1/4): XII +
835 pp.





BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 45(2)


~.- .j


Figures. 1-7 (all figures frontal view x4, dorsal view x6, phot.: E. Neubert). Shells of Neniinae. Fig. 1: Columbinia
(Steatonenia) pachygastris n. sp., holotype (UF 268818), H = 27.45 mm. Fig. 2: Cyclonenia gibber n. sp., holotype
(UF 268799), H = 25.90 mm. Fig. 3: Cyclonenia sanmarcos Grego & Szekeres, 2004, H = 26.10 mm. Fig. 4: Zilchiella
palatalis n. sp., holotype (UF 268817), H = 22.8 mm. Fig. 5: Zilchiella scala n. sp., holotype (UF 268816), H = 21.90
mm. Fig. 6: Hemicena polinskiana polinskiana (Pilsbry, 1949), holotype (ANSP 185113a), H = 21.40 mm. Fig. 7:
Hemicena polinskiana ancashensis n. ssp., holotype (UF 210212), H = 18.10 mm.






NEUBERT & NORDSIECK: New South American Clausiliidae


Figures. 8-16 (all figures frontal view x5, dorsal view x7, phot.: E. Neubert). Shells of Neniinae. Fig. 8: Incania
papillosa papillosa n. sp., holotype (UF 268801), H = 18.70 mm. Fig. 9: Incania papillosa imbecilla n. ssp.,
holotype (UF 298626), H = 13.20 mm. Fig. 10: Incaniaplatystoma n. sp., holotype (UF 268805), H =13.15 mm. Fig.
11: Steeriana cajamarcana cajamarcana Weyrauch & Zilch, 1954, holotype (SMF 135516), H =12.00 mm. Fig.
12: Steeriana sorormajor n. sp., holotype (UF 268787), H = 18.40 mm. Fig. 13: Steeriana cajamarcana sororminor
n. ssp., holotype (UF 268784), H = 9.80 mm. Fig. 14: Bequaertinenia delicate n. sp., holotype (UF 268795), H
= 17.0 mm. Fig. 15: Cyclonenia albosuturalis n. sp., holotype (UF 268788), H = 13.55 mm. Fig. 16: Cylindronenia
violacea n. sp., holotype (UF 268779), H = 15.95 mm.





BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 45(2)


Figures. 17-25 (all figures frontal view x5, dorsal view x7, phot.: E. Neubert). Shells of Neniinae. Fig. 17:
Pseudogracilinenia huallagana huallagana (Pilsbry, 1949), holotype (ANSP 185174a), H= 11.00 mm. Fig. 18:
Pseudogracilinenia huallagana amoena n. ssp., holotype (UF 268775), H = 15.00 mm. Fig. 19: Incaglaia leucostoma
n. sp., holotype (UF 268797), H = 15.75 mm. Fig. 20: Brevinenia richardsi (Grego & Szekeres, 2004), UF 268800,
H = 15.40 mm. Fig. 21: Temesa (Temesa)pilsbryipilsbryi, holotype (SMF 155296), H = 12.05 mm. Fig. 22: Temesa
(Temesa) pilsbryi perfectecostata n. ssp.,holotype (UF 268781), H = 13.30 mm. Fig. 23: Temesa (Temesa) gradata
n. sp., holotype (UF 268790), H = 12.45 mm. Fig. 24: Temesa (Temesa) clausilioides (Reeve, 1849), syntype NHM
1969273, H = 12.90 mm. Fig. 25: Temesa clausilioides (Reeve, 1849), UF 268783, H = 14.0 mm.







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RECENT PUBLICATIONS OF THE BULLETIN


Neubert, E., & H. Nordsieck. 2005. New South American Clausiliidae from the collections of the Florida Museum
of Natural History (Gastropoda, Clausiliidae, Neniidae). Bull. Florida Mus. Nat. Hist. 45(2): 45-62. Price $5.00

Dilcher, D.L. & T.A. Lott. 2005. A Middle Eocene fossil plant assemblage (Powers Clay Pit) from western
Tennessee. Bull. Florida Museum Nat. Hist. 45(1): 1-43. Price $7.00

King, F. W. and C. M. Porter, (Editors). 2003. Zooarchaeology: Papers to honor Elizabeth S. Wing. Volume 44, No. 1,
pp.1-208. Price $20.00

MacFadden, B. J. and O Carranza-Castaneda. 2002. Cranium of Dinohippus mexicanus (Mammalia: Equidae)
from the early Pliocene (Latest Hemphillian) of central Mexico, and the origin of Equus. Volume 43, No.5, pp. 163-
185. Price $5.00

Kratter, A. W., T. Webber, T. Taylor, and D. W. Steadman. 2002. New specimen-based records of Florida birds.
Volume 43, No.4, pp.111-161. Price $5.50

MacFadden, B. J. 2001. Three-toed browsing horse AIl hilth.,'iJm clarencei from the early Miocene (Hemingfordian)
Thomas Farm, Florida. Volume 43, No.3, pp. 79-109. Price $5.50

F. G. Thompson and G. P. Brewer. 2000. Land snails of the genus Humboldtiana from northern Mexico (Gastropoda,
Pulmonata, Helicoidea, Humboldtianidae). Volume 43, No. 2, pp. 49-77. Price $5.00

Hayes, F. G. 2000. The Brooksville 2 local fauna (Arikareean, latest Oligocene): Hernando County, Florida. Volume
43, No.1, pp. 1-47. Price $6.50


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