The National Parks of Haiti
The Butterflies (Lepidoptera:Rhophalocera)
of Morne La Visite
and Pic Macaya
Miami Springs, Fla
Albert Schwartz, PhD
Miami Dade Community College
Prepared for USAID/Haiti under contract Number
A4;th 'B,-i riliea.5
HY~;> *"'~a^^l j
THE BUTTERFLIES (LEPIDOPTERA: RHOPALOCERA) OF
MORNE LA VISIT AND PIC MACAYA, HAITI
156 S. Melrose Dr.,
Miami Springs, FL 33166
Miami-Dade Community College, North Campus
Miami, FL 33167
ABSTRACT. Twenty five species of rhopaloceran
Lepidoptera are known from the Morne la Visite and Pic
Macaya areas. Those species that are most common are
confirmed high elevation Hispaniolan butterflies, but there
are single or a very few specimens of low to moderate
elevation species. The lack of specimens of papilionids,
libytheids, and ithomiids may be due to the elevation or the
ecology of these upland areas.
The high uplands of the Massif de la Selle and Massif
de la Hotte on the Hispaniolan south island have been rarely
sampled as far as their lepidopteran faunas are concerned.
No one has made a serious effort to collect butterflies on
the Massif de la Hotte, and in fact the biology of that
range has been very poorly known, due primarily to its
inaccessibility. Biologists have visited the Massif de la
Selle with some frequency, but their itineraries did not
carry them into the area now included within the national
park boundary. Major exceptions are P.J. Darlington and M.
Bates, who, on 16-22.ix.1934, collected carabid beetles in
the high uplands of the Massif de la Selle on Morne la
Visite (Darlington, 1935). They collected, in addition to
beetles, some lepidopteran material. Darlington alone
collected in the Massif de la Hotte area (10-22.x.1934).
All specimens were deposited in the Museum of Comparative
Zoology at Harvard University, and the collections served as
sources of 5 holotypes of new taxa.
S.R. Yocom made collections at both Morne la Visite
(12.ii.1984-16.ii.1984) and in the vicinity of Pic Macaya
(23.i.1984-9.ii.1984). Gali likewise visited both areas and
collected specimens at Morne la Visite (l.ix.1984-5.ix.1984)
and at Pic Macaya and vicinity (9.ix.1984-13.ix.1984).
Chris Fastie and Koren Zimmerman collected in the La Visite
area (10.i.1985-16.i.1985) and in the Pic Macaya region
(21.i.1985-28.i.-1985). All collected in other areas
peripheral to the proposed national parks, but that material
is not included in the present report; we have included
here only those species that have been taken within the park
The high upland lepidopteran fauna of Hispaniola is
sparse. Although all collections were made during winter
and fall, not the prime times for butterfly activity on
Hispaniola, it is not likely that any collectors would have
encountered many additional species if they had collected in
the summer, the time of greatest abundance of many species.
The notably poor representation of some families
(Nymphalidae, for instance) is probably not an artifact of
the season but rather the reflection of the fact that there
are very few nymphalids that occur at these high elevations,
even during the summer. The scarcity of lycaenids and
hesperiids likewise is due to the high elevations involved.
Lycaenids are, on Hispaniola, primarily butterflies of
lowland xeric areas, and skippers have their greatest
species diversity from low to moderate elevations. The
exception to this statement is Paratrytone batesi, an upland
skipper described from the Massif de la Selle and common not
only there but in the Massif de la Hotte, as well as on the
Cordillera Central and the Sierra de Neiba in the Republica
Dominicana. Areas wherein all collectors worked are
primarily pine forest, a habitat that is not rich in
Of the 7 families of butterflies collected, the
outstanding exception is the Satyridae. The sole Antillean
representative of this family, Calisto, is noteworthy for
its explosive radiation on Hispaniola, whence there are now
28 species known, in contrast to a maximum elsewhere of 4
species on Cuba. Jamaica and Puerto Rico have 1 species
each, and the Bahama Islands have 2 species, derived from
Cuba. Many Hispaniolan Calisto are high upland butterflies,
often restricted to 1 or a very few mountain massifs. These
upland species do not shun pine forest and often have their
maximum abundance there. It is not surprising that more
kinds of satyrids were collected than representatives of any
Since many localities on both Morne la Visite and in
the Pic Macaya region are not shown on any map, the senior
author compiled the following gazeteer for his localities.
This listing includes directions for reachingthe various
sites and gives a brief annotation of the habitat.
l'Ouest: Scierie (old sawmill), Morne la Visite. The
following localities are approximately 2 hours by vehicle
south of Port-au-Prince. On the road to Kenscoff, one
passes the towns of Petionville, Fermate, and Furcy. In
Furcy, the main road forks into a lower and an upper road.
The upper road takes one directly above Kenscoff, whereas
the lower road continues along the same elevational band and
then abruptly ascends toward a higher elevation. This road
is narrow and contains numerous mudholes. After a rainfall,
the road can be as smooth and slick as ice and warrants
extreme caution. After approximately 10 tedious km, one
comes to a flat stretch which is more or less the entrance
to the park area. The road continues through open mixed
pine-deciduous forest for approximately 3 km and eventually
descends to a creek almost immediately followed by a small
stream. This small stream feeds into the Riviere Blanche a
few 100 m below the road. Beyond the stream, less than 1 km
ahead, one arrives at Scierie.
5 km WNW Scierie. When traveling up the main road from
Furcy heading toward Scierie, there is a field with large
"cut up" boulders and/or rocks on the left with an
inconspicuous path. This path ascends and continues along
some open pineland with a tall grass understory. The path
follows a "strict" elevational band and eventually descends
toward Scierie. There are several open meadows, corn
fields, and side paths. It is very important to remain on
the main path which follows the same elevation.
East side of Pic Cabaio, ca. 2.4 km from Scierie. From
Scierie, one travels south and ascends Pic Cabaio, and then
descends to the south side of the Pic into fields of corn.
Once past the corn fields, one enters semi-open pinewoods.
1 km W Roche Cabrit. This locality can be approached
from the road that passes by Scierie. It is approximately 2
km from Scierie.
1.2 km S Scierie. When travelling on the road toward
Furcy from Scierie, at about 0.4 km down the road, there is
a path on the left-hand side, before the small stream.
Ascending this path and keeping to the right, one first
passes corn fields on the right and then through the middle
of a corn field. Eventually, one reaches some pinewoods
along the Riviere Blanche, which is below. The path is now
situated at the bottom of a shrubby hillslope with pines,
and below the path are many tall and short ferns mixed with
a few pines.
Sud: Massif de la Hotte and vicinity.
The high uplands of the Massif de la Hotte are
approached most easily from Camp Perrin near the southern
coast of the Tiburon Peninsula. Biologists have found Camp
Perrin and nearby Les Cayes (on the southern coast) to be
convenient locations to set up for field work on the distal
portion of the Tiburon Peninsula, due primarily to their
close proximity to the Massif de la Hotte and their
accessibility from Port-au-Prince.
From Camp Perrin, one travels on the road
south-southwest toward, and then west, across the Riviere du
Sud. From the west, one heads north across the Plaine des
Cayes to the Riviere 1'Acul and parallels the l'Acul on its
east side in a northeast direction. The 1'Acul switches
back, and one crosses to the west side to continue along the
1'Acul which switches back once again, and one travels
across to the west side to continue in a northeast
A gradual ascent is made up the Massif de la Hotte on a
tortuous road that can be quite an adventure for one
unfamiliar with Haitian high elevation roads. When one
arrives at Les Platons, travel by foot or mule is the only
mode of transportation available. From Les Platons, one
travels north across a fairly flat stretch along a path
through open scrub and rock. At a fairly brisk pace, one
can cross this flat stretch in 1.5-2 hr. Several small
native communities lie along the way where bananas and other
refreshments can be purchased for a minimal fee.
Subsequently, one arrives on the Plaine de Formon, at the
base of Pic Formon; this is the home of Madame Robert a
native hostess, where one can camp for the evening (base
camp #1). From the Plaine de Formon, one has a choice of
two directions in which to cross the Pic Formon range. The
route taken by Gali is given below.
When traveling from Plain de Formon one ascends the
southern slope of the Formon range. The slope is moderately
inclined, and at a fairly good pace one can arrive at base
camp #2 near the top of the Formon range in 2 hr. The
incline becomes rather steep at this elevation for several
hundred meters until it finally becomes more or less level.
At this locality, one is near Pic Formon; the aspect of the
range becomes more dense and wet. The situation typical of
high elevation collecting (sunny in the morning, overcast
and raining by midday) is usual. Although one is on level
ground, when traveling along, movement is quite difficult
due to the mud and slippery conditions. On the other side,
one begins a fairly steep and tortuous descent in which the
path courses in a zigzag pattern. Without question, the
stamina of one's quadriceps muscles is tested on this path!
The path courses downward into a large ravine between the
Formon and Macaya ranges. The location in the ravine is at
the head of the Riviere du Sud and is a fine campsite
because of the accessibility of water sources.
Ravine locality. The ravine is fairly wide and
continues for several kilometers in the direction of Les
Cayes on one hand and Pic Macaya on the other. The river
courses through the ravine and is outlined with boulders and
rocks of various sizes with paths of varying lengths at the
riversides. Landslides are known to be common and warrant
precaution in obvious situations. Vegetation along the
paths in the ravine is sparse, with tall shrubs and some
herbaceous plants along the paths.
1400 m. When travelling along the ravine in the
direction of Les Cayes, one continues for approximately 1
km; on the left hand there is a path that courses up the
From the ravine, one travels up-river along the Riviere
du Sud as far as its source. There one ascends along a
mixed dirt and gravel path, and after approximately 2 hr, he
reaches the saddle between the Formon and Macaya ranges.
This saddle is approximately 400 m below Pic Macaya. The
incline is dangerously steep until Pic Macaya. One travels
through mixed overgrown pine and deciduous forest with
shrub-fern understory. A magnificent view is to the right
of the path, with a 300 m drop. Butterflies were collected
as they perched on the shrubs and ferns. Some Calisto were
seen; however, they regularly escaped into the forest,
using the shrubs and ferns as havens. The exceptionally
steep slope made collecting quite difficult. When one
arrives in the vicinity of Pic Macaya, the aspect of the
range became more dark and dense. A canopy is formed above
and allows for minimal sunlight. We encountered a small
patch of open area where previous biologists had camped. No
butterflies were seen or collected on Pic Macaya.
Urbanus proteus domingo Scudder, 1872. Sud: ravine
between Pic Formon and Macaya, 1050 m, 1 (ll.ix, 0900-1300
h; 24-29 C).
Wallengrenia druryi Latreille, 1824. 1'Ouest: 2 km NW
Scierie, 1785-1810 m, 1 (3.ix, 0930-1100 h, 19-21 C); Sud:
ravine between Pic Formon and Macaya, 1050 m, 1 (ll.ix,
0900-1300 h, 24-29 C); Macaya, 1485 m, 1 (6.ii); o.5 km SE
Pic Macaya, 1732 m, 1 (28.i, 1345 h).
Paratrytone batesi Bell, 1935. l'Ouest: 200 m N
Scierie, 1891 m, 2 (3.ix, 1230-1330 h, 25 C); 1.2 km S
Scierie, 1950 m, 1 (5.ix, 1230-1330 h, 24 C); 1 km SE
Scierie, 1891 m, 1 (l.ix, 1230-1330 h, 24 C); 2 km NW
Scierie, 1785-1810 m, 8 (3.ix. 0930-1100 h, 19-21 C); 4.8 km
NW Scierie, 2187 m, 1 (2.ix, 1230-1330 h, 19 C); 5 km WNW
Scierie, 1891 m, 24 (4.ix, 0930-1030 h, 21 C); 1-4 km WNW
Scierie, 1830-1891 m, 2 (4.ix, 1030-1230 h, 21 C); 1 km W
Roche Cabrit, 2000 m, 1 (5.ix, 1000-1030 h, 24 C); Sud:
Pic Formon, near top, 580 m, 3 (9.ix); 2 (12.ix, 1200-1300
h, 28 C); Pic Macaya, near top, 2300 m, 2 (10.ix, 1000-1200
h, 25 C); Morne Formon, 1750 m, 1 (2.ii). Yocom took the
last-listed specimen feeding on Ageratum (Asteraceae).
Panoquina sylvicola woodruffi Watson, 1937. l'Ouest:
2 km NW Scierie, 1785-1810 m, 1 (3.ix, 0930-1100 h, 19-21
Panoquina nero Fabricius, 1798. 1'Ouest: 5 km WNW
Scierie, 1891 m, 1 (4.ix, 0930-1030, 21 C).
Eurema pyro Godart, 1819. Sud: between Pic Formon and
Macaya, 1050 m, 1 (11.ix, 0900-1300 h, 24-29 C).
Nathalis iole Boisduval, 1836. l'Ouest: 200 m NW
Scierie, 1952 m, 4 (2-3.ix, 0900-1330 h, 22-24 C); 0.2 km
NE Scierie, 1484 m, 2 (11.i, 0900, 24 C); 1.8 km SE
Scierie, 1865 m, 4 (12.i, 0920, 21 C); Morne La Visite,
2100 m, 5 (13.ii); 2 km E Morne La Visite, 2121 m, 1 (15.i,
1445 h, 14 C).
Dismorphia spio Godart, 1819. l'Ouest: 1 km SE
Scierie, 1891 m, 1 (l.ix, 1230-1330 h, 24 C).
Strymon columella cybirus Hewitson, 1874. l'Ouest: 5
km WNW Scierie, 1891 m, 1 (4.ix, 0930-1030 h, 21 C).
Hemiargus hanno watsoni Comstock and Huntington, 1943.
1'Ouest: 5 km WNW Scierie, 1891 m, 1 (4.ix, 0930-1030 h, 21
Heliconius charitonius church Comstock and Brown,
1950. l'Ouest: Morne La Visite, 1430-1880 m, 3 (12, 16.ii).
Dryas iulia hispaniola Hall, 1925. l'Ouest: Scierie,
1685 m, 1 (10.i, 1140 h, 22 C).
Junonia evarete zonalis Felder and Felder, 1867.
l'Ouest: Scierie, 1936 m, 3 (12.i, 1020 h, 20 C); 2 km NW
Scierie, 1788-1810 m, 1 (3.ix, 0930-1100 h, 19-21 C); 1.2
km SW Scierie, 1806 m, 1 (10.i, 1230 h, 17 C); 1.4 km SW
Scierie, 1794 m, 1 (10.i, 1245 h, 15 C); 0.2 km NE Scierie,
1948 m, 5 (11.i, 0900-1520 h, 17-24 C); Morne La Visite,
2100 m, 8 (13.ii).
Vanessa cardui cardui Linnaeus, 1758. l'Ouest:
Scierie, 1891 m, 1 (28.ix, 1730 h, 23 C); 1685 m, 2 (10.i,
0900-1055 h, 19-21 C); 1685 m, 1 (10.i, 1600 h, 17 C); 5
km WNW Scierie, 1891 m, 1 (4.ix, 0930-1030 h, 21 C); 1.4 km
SSW Scierie, 1794 m, 2 (10.i, 1245 h, 15 C); 0.2 km NE
Scierie, 1948 m, 1 (11.i, 0906 h, 17 C); Morne La Visite,
2100 m, 1 (13.ii); Morne La Visite, 2165 m, 1 (16.i, 1130
h, 19 C); 0.5 km S Morne La Visite, 1988 m, 1 (16.1, 1430
h, 18 C).
Vanessa virginiensis Drury, 1770. l'Ouest: ca.
Scierie, 2030 m, 1 (11.i, 1500 h, 18 C); 4 km NW Scierie,
2000 m, 1 (2.ix, 1100-1145 h, 21 C); 2 km NW Scierie,
1785-1810 m, 1 (3.ix, 0930-1100 h, 19-21 C); 5 km WNW
Scierie, 1891 m, 1 (4.ix, 0930-1030 h, 21 C); Morne La
Visite, 2100 m, 3 (13.ii); Morne La Visite, S slope, 2100 m,
1 (14.i, 0845 h, 16 C); Morne La Visite, 2200 m, 1 (14.i,
1545 h, 18 C); Roche Cabrit, 2200 m, 1 (15.ii); Sud: Morne
Formon, 1635-1800 m, 5 (27.i-4.ii); Pic Macaya, 2345 m, 3
(25.1, 1400 h).
Calisto archebates Menetries, 1832. l'Ouest: 1 km SE
Scierie, 1891 m, 23 (l.ix, 1230-1330 h, 24 C); 2 km NW
Scierie, 1785-1810 m, 1 (3.ix, 0930-1100 h, 19-21 C); 200 m
N Scierie, 1891 m, 1 (3.ix, 1230-1330 h, 24 C); 200 m W
Scierie, 1675 m, 4 (10.i, 1105 h, 21 C); 0.6 km W Scierie,
1820 m, 1 (10.i, 1210 h, 22 C); 5 km WNW Scierie, 1891 m, 2
(4.ix, 0930-1030 h, 21 C); Morne La Visite, S slope, 2100
m, 1 (14.i, 1115 h, 23 C).
Calisto loxias Bates, 1935. Sud: Pic Formon, nr. top,
1900-1910 m, 7 (9, 12.ix, 1200-1300 h, 28 C).
Calisto chrysaoros Bates, 1935. l'Ouest: 1.2 km S
Scierie, 1950 m, 12 (5.ix, 1100-1230 h, 29 C); Sud: Pic
Formon, nr. top, 1900-1910 m, 31 (9-13.ix, 0830-1330 h,
21-28 C); Morne Formon, 1830 m, 3 (2.ii).
Calisto tragia Bates, 1935. 1'Ouest: 4 km NW Scierie,
610 m, 2 (2.ix, 1100-1145 h, 21 C); 5 km WNW Scierie, 1891
m, 9 (4.ix, 0930-1030 h, 21 C); 1-4 km WNW Scierie,
1830-1891 m, 19 (4.ix, 1030-1230 h, 21 C); 1 km SE Scierie,
1891 m, 1 (l.ix, 1230-1330 h, 24 C); 1.2 km S Scierie, 1950
m, 2 (5.ix, 1100-1230 h, 29 C); 1 km W Roche Cabrit, 2000
m, 3 (5.ix, 1000-1030 h, 24 C); east side, Pic Cabaio, 2100
m, 2 (5.ix, 0900-0950, 22 C); Morne La Visite, S slope,
2085 m, 1 (14.i, 1020 h, 19 C).
Calisto hysia Godart, 1819. Sud: Morne Formon, 1000 m,
2 (l.ii); Pic Formon, nr. top, 1900-1901 m, 1 (12.ix,
1200-1300 h, 28 C); western end of Pic Macaya ridge, 1400
m, 8 (11.ix, 1300-1340 h). Bates (1935) reported this
species from "La Visite", but no recent collectors have
taken it there.
Calisto clenchi Schwartz and Gali, 1984. l'Ouest: 1.2
km S Scierie, 1950 m, 2 (5.ix, 1100-1230 h, 29 C).
Danaus cleophile Godart, 1819. Sud: between Pic
Formon and Macaya, 1050 m, 1 (11.ix, 0900-1300 h, 24-29 C).
Danaus plexippus megalippe Hubner, 1819. 1'Ouest:
Scierie, 1936 m, 1 (12.i, 1020 h, 21 C); 1 km W Scierie,
1827 m, 1 (10.i, 1215 h, 21 C); 2 km NW Scierie, 1785-1810
m, 1 (3.ix, 0930-1100 h, 21 C); Morne La Visite, 1430-1880
m, 3 (12, 16.ii); Sud: Morne Formon, 1000-1200 m, 3
Anetia briarea briarea Godart, 1819. l'Ouest: 5 km
WNW Scierie, 1891 m, 1 (4.ix, 0930-1030 m, 21 C); Morne La
Visite, S slope, 2085-2100 m, 7 (13-14.i, 1030-1245 h, 18-24
C). Fastie and Zimmerman observed as many as 12 individuals
of this species feeding on the flowers of Eupatorium
(Asteraceae) on 14.i at 1245 h and 24 C.
Anetia jaegeri Menetries, 1832. l'Ouest: Scierie, 1685
m, 3 (10.i, 0900-1045 h, 19-20 C); 200 m NW Scierie, 1952
m, 2 (2.ix, 0900-0930, 22 C); 0.4 km S Scierie, 580 m, 3
(5.ix, 1045-1050 h, 25 C); 0.5 km S Scierie, 1845 m, 3
(12.i, 0920 h, 16 C); ca. 4.8 km NW Scierie, 2187 m, 1
(2.ix, 1230-1330 h, 19 C); Morne La Visite, 2058-2066 m, 5
(14-15.i, 0945-1530 h, 15-24 C); Morne La Visite 1880 m, 6
(12.ii); Sud: Pic Formon, nr. top, 1910 m, 2 (9.ix); Pic
Macaya, nr. top, 2300 m, 1 (10.ix, 1000-1200 h, 25 C); Pic
Macaya, 2320 m, 3 (27.i, 1045-1115 h); Morne Formon,
1800-1880 m, 3 (29-30 i).
Twenty five species of butterflies, representative of
seven families, are herein reported from the Morne la Visite
and Pic Macaya areas. These are: 5 skippers, 3 pierids, 2
lycaenids, 2 heliconiids, 3 nymphalids, 6 satyrids, and 5
danaids. The two areas have six species in common: W.
druryi, P. batesi, V. virginiensis, C. chrysaoros, D.
plexippus, and A. jaegeri. Although the remaining 19
species are generally widespread on Hispaniola, some (U. p.
domingo, P. s. woodruffi, E. pyro, N. iole, D. spio, S. c.
cybirus, H. h. watsoni, H. ch. church, D. i. hispaniola, J.
e. zonalis) have their elevations of maximum abundance in
the lowlands or reach to only moderate elevations in the
mountain ranges. On the other hand, some species (P.
batesi, V. c. cardui, V. virginiensis, C. archebates, C.
loxias, C. chrysaoros, C. tragia, C. clenchi, D. cleophile,
A. b. briarea, A. jaegeri) are predominantly high elevation
butterflies, rarely descending to moderate or low
elevations. Of this group, C. archebates is restricted to
the Massif de la Selle (in which range Morne La Visite lies)
and C. loxias has been known only from the Massif de la
Hotte (of which Pic Macaya is the culminating peak).
Although it is perhaps foolhardy to make
generalizations based on collections which total only 63
man/days, all in fall and winter, still there are certain
peculiarities that bear noting.
The lack of records for J. e. zonalis in the Macaya
region is peculiar; this species is abundant at Morne La
Visite. This nymphalid is a butterfly of open areas
-fields, roadsides, pastures, and other disturbed areas.
Vanessa c. cardui is unknown from the Macaya region,
yet it appears to be common at Morne La Visite. This
species too is a butterfly of more or less open areas.
There are no records of C. hysia from Morne La Visite
(although it has been reported from that area by Bates
), but this satyrid is not uncommon in the Macaya
area. The species is altitudinaly and geographically
widespread on the Hispaniolan south island, and its absence
from Morne La Visite is inexplicable.
Likewise, A. b. briarea is an upland butterfly
throughout Hispaniola; there are no Macaya records for this
species, although it is modestly abundant at La Visite. Its
congener, Anetia pantherata Martyn 1797, has not been taken
in either area; this species likewise is a common butterfly
of Hispaniolan high elevations.
The apparent rarity of D. cleophile, known from only
one Macaya specimen and unknown from Morne La Visite, is
peculiar, since this species is a moderate to high elevation
The absence of 3 families of butterflies whose
representatives occur on Hispaniola (Papilionidae,
Libytheidae, Ithomiidae) from the park areas deserves
comment. The single Hispaniolan libytheid (Libytheana
terena Godart 1819) does occur in the uplands but is
distinctly uncommon there and is more abundant in lowland
forests, either mesic or xeric. It is rarely encountered in
pine forest. The single Hispaniolan ithomiid (Greta
diaphana Drury 1773) is widely distributed on the Dominican
portion of the north island and is known in the Republica
Dominicana from the Sierra de Baoruco on the south island.
The species has not been taken in the Dominican portion of
the Massif de la Selle (between Los Arroyos and El Aguacate)
and may well be absent from the Massif de la Selle as well
as the Massif de la Hotte. This clearing is a butterfly of
dappled sun-and-shade situations that occur along streams or
forest margins. Schwartz has never seen this "proper"
habitat in Haiti, but Gali found what appeared to be
suitable habitat near Morne La Visite (a deciduous wooded
stream near Scierie; Fig.2) but did not encounter G.
diaphana. It may be that (1) the species does not occur in
Haiti (there are no Haitian records) or (2) that the
requisite habitat is very uncommon and local.
The swallowtails (Papilionidae) on Hispaniola are
almost without exception butterflies of low to moderate
elevations. It is rare indeed that one encounters such
species as Battus polydamas Linnaeus 1758, Heraclides
aristodemus Esper 1794, H. machaonides Esper 1796, or
Priamides pelaus Fabricius 1775 in the uplands, and even
more so if the forest cover is pine. Other papilionids
(Heraclides aristor Godart 1819, Eurytides zonarius Butler
1869) are distinctly xeric lowland species, although
occasional individuals may wonder to moderate elevations.
The single upland papilionid is Battus zetides Munroe 1971.
This species is locally very abundant but occurs only in
deciduous upland forest. Although there are no recent
Haitian records (due to habitat destruction?), the species
is expected in both park areas, either as a vagrant in pine
forest or in streamside deciduous forest. It is most
abundant in the summer (June-August), but individuals may be
seen or taken at other times of the year.
Bates, M. 1935. The satyrid genus Calisto. Occ. Papers
Boston Soc. Nat. Hist., 8:229-248.
Darlington, P.J. 1935. West Indian Carabidae II. Itinerary
of 1934; forests of Haiti; new species; and a new key
to Colpodes. Psyche, 42:167-215.
Butterflies of the National Parks of Haiti
Parc La Visite Pare Macaya
Urbanus proteus domingo 0 +
Wallengrenia druryi + +
Paratrytone batesi + +
Panoquina sylvicola woodruffi + 0
Panoquina nero + 0
Eurema pyro 0 +
Nathalis iole + 0
Dismorphia spio + 0
Strymon columella cybirus + 0
Hemiarqus hanna watsoni + 0
Heliconius charitonius church + 0
Dryas iulia hispaniola + 0
Junonia evarete zonalis + 0
Vanessa cardui cardui + 0
Vanessa virginiensis + +
Calisto archebates + 0
Calisto loxias 0 +
Calisto chrysaoros + +
Calisto tragia + 0
Calisto hysia ? +
Calisto clenchi + 0
Danaus cleophile 0 +
Danaus plexippus megalippe + +
Anetia briarea briarea + 0
Anatia jaegeri + +
Total number of families 7
Total number of species 25