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Group Title: FMRI technical reports
Title: Checklists of selected shallow-water marine invertebrates of Florida
CITATION THUMBNAILS PAGE IMAGE ZOOMABLE
Full Citation
STANDARD VIEW MARC VIEW
Permanent Link: http://ufdc.ufl.edu/UF00001489/00001
 Material Information
Title: Checklists of selected shallow-water marine invertebrates of Florida
Series Title: FMRI technical reports
Physical Description: xv, 238 p. : ; 28 cm.
Language: English
Creator: Camp, David K
Lyons, William G
Perkins, Thomas H
Florida Marine Research Institute
Publisher: Florida Dept. of Environmental Protection, Florida Marine Research Institute
Place of Publication: St. Petersburg Fla
Publication Date: 1998
 Subjects
Subject: Marine invertebrates -- Classification -- Florida   ( lcsh )
Genre: government publication (state, provincial, terriorial, dependent)   ( marcgt )
bibliography   ( marcgt )
non-fiction   ( marcgt )
 Notes
Summary: The checklists are a record of all species of mollusks, polychaetous annelids, malacostracan crustaceans, and echinoderms known to occur in Florida's estuarine and coastal marine waters offshore to depths of approximately 37 meters. Compilation of the version ended in November 1997.
Bibliography: Includes bibliographical references and index.
Statement of Responsibility: David K. Camp, William G. Lyons, Thomas H. Perkins.
 Record Information
Bibliographic ID: UF00001489
Volume ID: VID00001
Source Institution: University of Florida
Holding Location: University of Florida
Rights Management: All rights reserved by the source institution and holding location.
Resource Identifier: ltqf - AAA0781
notis - AMF6079
alephbibnum - 002450793
oclc - 40721216

Table of Contents
    Copyright
        Copyright
    Front Cover
        Front cover i
        Front cover ii
    Front Matter
        Page a
        Page b
    Table of Contents
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Full Text












COPYRIGHT NOTICE

[yearof publication] Florida Department of Environmental
Protection [source text]
The FLORDA DEPARTMENT OF ENVIRONMENTAL PROTECTION and FLORA
FISH AND WILDLIFE CONSERVATION COMMISSION holds all lights to the
source text of this electromn resource on behalf of the State of Flonda The
FLORDA DEPARTMENT OF ENVIRONMENTAL PROTECTION and FLORDA FISH
AND WILDLIFE CONSERVATION COMMISSION shall be considered the
copyright holder for the text and images of tus publication
Under the Statutes of Florida (FS 25705, 257105, and 377075), the
FLORDA DEPARTMENT OF ENVIRONMENTAL PROTECTION and FLORDA FISH
AND WILDLIFE CONSERVATION COMMISSION (Tallahassee, FL) publisher of
the Florida Marine Research Institute's Technical Reports, as a division of
state government makes its documents public ( c published) and extends to
the state's official agencies and libraes, including the University of
Florida's Smathers Libary, limited rights of reproduction
The FLORDA DEPARTMENT OF ENVIRONMENTAL PROTECTION and FLORDA
FISH AND WILDLIFE CONS CONSERVATION COMMISSION has made ts publication
available to the University of Florida, on behalf of the State University
System, for purposes of digihzation and Internet distribution


I if




ISSN 1092-194X


A U.S


Checklists of Selected
Shallow-Water Marine Invertebrates
of Florida
David K. Camp, William G. Lyons, and Thomas H. Perkins


Florida Department of "^"
Environmental Protection FLORID ;
FMRI Technical Report TR-3 1998

















Lawton Chiles
Governor of Florida


Florida Department of Environmental Protection

Virginia B. Wetherell
Secretary





FLOR A



The Florida Marine Research Institute (FMRI) is a bureau of the Florida Department of Envi-
ronmental Protection (FDEP).The FDEP's mission is to"protect, conserve, and manage Florida's
environment and natural resources." The FMRI conducts applied research pertinent to manag-
ing marine-fishery resources and marine species of special concern in Florida.
Programs at the FMRI focus on resource-management topics such as managing gamefish and
shellfish populations, restoring depleted fish stocks and the habitats that support them, pro-
tecting coral reefs, preventing and mitigating oil-spill damage, protecting endangered and
threatened species, and managing coastal-resource information.
The FMRI publishes three series: Memoirs of the Hourglass Cruises, Florida Marine Research Publi-
cations, and FMRITechnical Reports. FMRI Technical Reports contain information relevant to imme-
diate resource-management needs.


Kenneth D. Haddad, Chief of Research

Institute Editors
Theresa M. Bert, David K. Camp, Paul R. Carlson, Mark M. Leiby,
William G. Lyons, Anne B. Meylan, Robert G. Muller,
Ruth O. Reese, Carmelo R. Tomas

James F. Quinn, Jr., Science Editor
Judith G. Leiby, Copy Editor
Llyn C. French, Art Editor











Checklists of Selected
Shallow-Water Marine Invertebrates
of Florida







David K. Camp
William G. Lyons
Thomas H. Perkins

Florida Department of Environmental Protection
Florida Marine Research Institute
100 Eighth Avenue Southeast
St. Petersburg, Florida 33701







Florida Department of Environmental Protection
FMRI Technical Report TR-3


1998









Cover Photographs


Clockwise from upper left:
Chloeia viridis, Diadema antillarum, Stenopus hispidus, and Lithopoma tectum.
Reprinted by permission from REEF CREATURE Identification-Florida,
Caribbean, Bahamas, Copyright 1992 by Paul Humann.





Copies of this document may be obtained from

Florida Marine Research Institute
100 Eighth Ave. SE
St. Petersburg, FL 33701-5095
Attn: Librarian





Document Citation

Camp, D. K., W. G. Lyons, and T. H. Perkins. 1998. Checklists of selected shallow-water marine in-
vertebrates of Florida. Florida Marine Research Institute Technical Report TR-3.238 pp.





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The papers used in this publication meet the minimum requirements of the American
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Table of Contents


RATIONALE AND SCOPE OF CHECKLISTS ..................
Acknowledgments .................................
Literature Cited ..................................
HPtrVt Tt TCO n- CL AT T MAT IAT ATECD MA A DTMTT LnT T T TCr A F1 t FO3DATTA


C1


Family Plicatulidae . .


MECK IT %-F a -- . . . . . .
M ethods ...................................................
Acknowledgm ents .............................................
Checklist of the M ollusca ................... ......................
Phylum M ollusca ............................................
Class Aplacophora ........................................
Order Pholidoskepia ......... .. ......................
Family Meiomeniidae .........................
Class Polyplacophora ......................................
Order Neoloricata ...................................
Family Hanleyidae ...........................
Family Ischnochitonidae ........................
Family Callistoplacidae ........................
Family Chaetopleuridae ........................
Family Chitonidae ...........................
Family Tonicellidae ...........................
Family M opaliidae ...........................
Family Acanthochitonidae ......................
Class Scaphopoda ........................................
Order Dentaliida ....................................
Family Dentaliidae ...........................
Family Gadilinidae ...........................
O rder G adilida .....................................
Family Gadilidae ............................
Class Bivalvia ..... .... ... .... ... ..... ... .... ...... ......
Order Solemyoida ...................................
Family Solemyidae ...........................
Order N uculoida ....................................
Family Nuculidae ............................
Family Nuculanidae ..........................
Order M ytiloida ....................................
Family M ytilidae ............................
O rder Arcoida .......................................
Fam ily Arcidae .............................
Family Noetiidae ............................
Family Glycymerididae ........................
O rder Pterioida .....................................
Fam ily Pteriidae ............................
Family Isognomonidae ........................
Family M alleidae ............................
Family Pinnidae .............................
O rder Lim oida .....................................
Fam ily Lim idae .............................
Order Ostreoida ..................................
Family Pectinidae ............................
Family Propeamussidae ........................


. . 1
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..10
. 10


.











Family Spondylidae . .
Family Anomiidae ....
Family Ostreidae ...
Family Gryphaeidae .
OrderVeneroida .............
Family Lucinidae ...
Family Ungulinidae . .
Family Cyrenoididae .
Family Chamidae ...
Family Lasaeidae ...
Family Galeommatidae .
Family Sportellidae . .
Family Carditidae ...
Family Condylocardiidae
Family Astartidae ...
Family Crassatellidae .
Family Cardiidae ....
Family Mactridae ....
Family Solenidae ....
Family Pharidae .....
Family Tellinidae ....
Family Donacidae ...
Family Psammobiidae .
Family Semelidae ...
Family Solecurtidae . .
Family Dreissenidae .
Family Trapezidae ....
Family Corbiculidae .
FamilyVeneridae ....
Family Petricolidae . .
Order Myoida ..............
Family Myidae ......
Family Corbulidae . .
Family Gastrochaenidae
Family Hiatellidae ...
Family Pholadidae . .
Family Teredinidae . .
Order Pholadomyoida .........
Family Lyonsiidae ...
Family Pandoridae . .
Family Thraciidae ...
Family Periplomatidae .
Family Poromyidae . .
FamilyVerticordiidae ..
Family Cuspidariidae ..
Class Gastropoda .................
Order Patellogastropoda ........
Family Lottiidae .....
Order Archaeogastropoda .......
Family Scissurellidae .
Family Fissurellidae ..
Family Turbinidae ....
FamilyTrochidae ....
Family Calliostomatidae


. . . . . 10
. . . . . 10
. . . . .. 10
. . . . . 10
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......... .11
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. . . . . 15
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. . . . . 16
. . . . . 16
. . . . . 16









Family Skeneidae .............................. 17
Order Neritopsina .................. .................... 17
Family Neritidae .............................. 17
Family Phenacolepadidae ......................... 17
Order Neotaenioglossa ...................................... 17
Family Obtortionidae ............................ 17
Family Litiopidae .............................. 17
Family Cerithiidae ............................. 17
Family Batillariidae ............................. 17
Family Potamididae ............................. 17
Family Planaxidae .............................. 17
Family Modulidae ................................. 17
Family Turritellidae ............................. 17
Family Siliquariidae ...................... ........ 18
Family Littorinidae ............................... 18
Family Skeneopsidae ............................ 18
Family Barleeiidae ............................. 18
Family Rissoidae .............................. 18
Family Hydrobiidae ..............................18
Family Truncatellidae .............................18
Family Assimineidae ............................ 19
Family Elachisinidae ............................ 19
FamilyVitrinellidae ................................ 19
Family Caecidae ............................... 19
Family Strombidae ............................. 19
Family Hipponicidae ............................ 20
Family Vanikoridae ..............................20
Family Calyptraeidae ............................ 20
Family Capulidae ...............................20
Family Xenophoridae .............................20
FamilyVermetidae ..............................20
Family Cypraeidae ............................. 20
Family Ovulidae ............................... 20
FamilyTriviidae ................................20
Family Lamellariidae ............................ 21
Family Naticidae .............................. 21
Fam ily Cassidae ............................... 21
FamilyTonnidae .................................. 21
Family Ficidae ................................... 21
Family Ranellidae ................................. 21
Family Personidae .............................. 22
Family Bursidae ............................... 22
Family Cerithiopsidae ........................... 22
Family Triphoridae ............................. 22
Family Epitoniidae ............................. 22
Family Janthinidae ............................. 23
Family Aclididae .............................. 23
Family Eulimidae .............................. 23
Order Neogastropoda ................... ................ 23
Family Muricidae .............................. 23
Family Coralliophilidae .......................... 24
Family Turbinellidae ............................ 24
Family Buccinidae .............................. 24










Family Melongenidae ........................... 24
Family Nassariidae ............................. 24
Family Fasciolariidae ............................ 25
Family Columbellidae ........................... 25
FamilyVolutidae .................................. 25
Family Olividae .................................. 25
Family Harpidae ............................... 26
Family Cystiscidae ............................. 26
Family Marginellidae .............................26
Fam ily M itridae ............................... 26
Family Costellariidae ............................ 26
Family Cancellariidae ........................... 26
Fam ily Drilliidae ............................... 27
Family Terebridae ................................. 27
Family Strictispiridae ............................ 27
FamilyTurridae .................................. 27
Family Conidae .................................. 27
Subclass Heterobranchia ................... .................. 28
Order Heterostropha ................... ................. 28
Family Cornirostridae ........................... 28
Family M athildidae ............................. 28
Family Architectonicidae .......................... 28
Family Rissoellidae ............................. 29
Fam ily Ebalidae ............................... 29
Family Pyramidellidae ........................... 29
Family Amathinidae ............................ 30
Order Uncertain ..................... .................... 30
Family Acteonidae ............................. 30
Subclass Opisthobranchia ................... ................. 30
Order Cephalaspidea ................... ................. 30
Family Ringiculidae ..............................30
Family Aplustridae ............................. 30
Family Cylichnidae ............................. 30
Family Philinidae .............................. 30
Family Aglajidae .............................. 30
Family Gastropteridae ........................... 30
Family Retusidae .............................. 30
Fam ily Bullidae ............................... 31
Family Haminoeidae ............................ 31
Family Runcinidae ............................. 31
Order Acochlidioidea ................... ................. 31
Family Microhedylidae ........................... 31
Order Thecosomata ................... .................. 31
Family Cavoliniidae ..............................31
Order Anaspidea ................... ................... 31
Family Akeridae ............................... 31
Family Aplysiidae .............................. 31
Family Notarchidae ............................. 31
Order Sacoglossa ................... ................... 31
Family Volvatellidae ..............................31
Family Cylindrobullidae .......................... 31
Family Oxynoidae .............................. 31
Fam ily Juliidae ................................ 31
Fam ily Elysiidae ............................... 31










Family Costasiellidae ............................32
Family Stiligeridae ............................. 32
Order Notaspidea ...................... .............. 32
Family Umbraculidae .......................... 32
Family Pleurobranchidae ......................... 32
Order Nudibranchia ....................................... 32
Family Corambidae ................................ 32
Family Goniodorididae ............................32
Family Aegiretidae ..............................32
Family Polyceratidae ............................ 32
Family Chromodorididae ......................... 32
Family Asteronotidae .............................33
Family Dorididae .... .................. .........33
Family Dendrodorididae ...........................33
Family Phyllidiidae ............................. 33
Family Archidorididae ........................... 33
Family Discodorididae ........................... 33
Family Platydorididae ........................... 33
Family Tritoniidae ................................. 33
Family Bornellidae ............................. 33
Family Lomanotidae ............................ 33
Family Scyllaeidae ............................. 33
Family Dotoidae .................................. 34
Family Arminidae .............................. 34
Family Janolidae .................................. 34
Family Flabellinidae ............................ 34
Family Eubranchidae .............................34
Family Tergipedidae ............................ 34
Family Fionidae ............................... 34
Family Facelinidae ............................. 34
Family Aeolidiidae ............................. 34
Family Glaucidae .............................. 34
Subclass Pulmonata ......................................... 34
Order Archaeopulmonata ................................. 34
Family Ellobiidae .............................. 34
Order Basommatophora .................................. 35
Family Siphonariidae .............................35
Family Trimusculidae .............................35
Order Systellommatophora ................................ 35
Family Onchidiidae ................................ 35
Class Cephalopoda. ....................... .................. 35
Order Sepioidea ......................................... 35
Family Spirulidae .............................. 35
Family Sepiolidae .............................. 35
Order Teuthoidea ..................... ................ 35
Family Loliginidae ............................. 35
Order Octopoda ......................................... 35
Family Octopodidae ............................ 35
Family Argonautidae ............................ 35
Pelagic Mollusks ................................................. .. 35
Class Gastropoda .......................... ........... ....... 36
Order Neotaenioglossa ................... ................ 36


Family Polybranchiidae.


. . . . . . . . . 3









Family Atlantidae ...............................36
Family Carinariidae ............................. 36
Family Pterotracheidae ............................36
Order Thecosomata ................... .................. 36
Family Limacinidae ............................. 36
Family Cavoliniidae .............................36
Family Peraclididae ............................. 36
Order Gymnosomata ................... ................. 36
Family Clionidae .................... ......... 36
Family Cliopsidae ................ ............. 37
Family Notobranchaeidae ......................... 37
Family Pneumodermatidae ........................ 37
Family Thliptodontidae ............................37
Order Nudibranchia ................... ................. 37
Family Phylliroidae ............................. 37
References .................... ............................... 37
CHECKLIST OF SHALLOW-WATER MARINE POLYCHAETOUS ANNELIDA OF FLORIDA . 79
Methods ................... .................................. .79
Acknowledgments ...............................................79
Checklist of the Polychaetous Annelida ................... ................ 80
Phylum Annelida ................... .......................... 80
Class Polychaeta .................... ....................... 80
Order Phyllodocida ................... .................. 80
Family Phyllodocidae ............................ 80
Family Lacydoniidae ............................ 81
Family Glyceridae ................................. 81
Family Goniadidae ............................. 81
Family Sphaerodoridae ............................ 81
Family Hesionidae ............................. 81
Family Pilargidae ............................. 82
Fam ily Syllidae ............................... 82
Family Nereididae ............................. 83
Family Nephtyidae ............................. 84
Family Paralacydoniidae .......................... 84
Family Aphroditidae ............................ 84
Family Polynoidae .................... ......... 84
Family Acoetidae ...............................85
Family Eulepethidae ............................ 85
Family Sigalionidae ................................ 85
Family Pholoidae ............................. 85
Family Chrysopetalidae ..... ........... ....... 85
Family Pisionidae ............................. 86
Order Amphinomida ................... .................86
Family Amphinomidae ............................86
Family Euphrosinidae ........................... 86
Order Eunicida .................... ................... 86
Family Onuphidae ............................. 86
Family Eunicidae ............................. 86
Family Lumbrineridae ............................87
Family Oenonidae ............................... 88
Family Dorvilleidae ................................ 88
Order Orbiniida .................... ................... 88
Family Orbiniidae ................................. 88










Order Spionida .............................
Family Spionidae ....................
Family Poecilochaetidae ................
Family Longosomatidae ................
Order Chaetopterida ..........................
Family Chaetopteridae .................
Order M agelonida ...........................
Family Magelonidae ..................
Order Cirratulida ............................
Family Paraonidae ...................
Family Questidae ....................
Family Cirratulidae ...................
Order Ctenodrilida ...........................
Family Ctenodrilidae .. .................
Order Cossurida .............................
Family Cossuridae ...................
Order Flabelligerida ..........................
Family Flabelligeridae .................
Family Acrocirridae ...................
Family Fauveliopsidae .................
Order Opheliida .............................
Family Opheliidae ...................
Family Scalibregmatidae ................
Order Sternaspida ...........................
Family Sternaspidae ..................
Order Capitellida ............................
Family Capitellidae ...................
Family Maldanidae ...................
Family Arenicolidae ..................
Order Oweniida .............................
Family Oweniidae ....................
Order Terebellida ............................
Family Pectinariidae ..................
Family Sabellariidae ..................
Family Ampharetidae .................
Family Trichobranchidae ................
Family Terebellidae ...................
Order Sabellida .............................
Family Sabellidae ....................
Family Serpulidae ....................
Family Spirorbidae ...................
Order Nerillida .............................
Family Nerillidae ....................
Order Polygordiida ...........................
Family Polygordiidae ..................
Order Protodrilida ...........................
Family Saccocirridae ..................
References ......................................... .

CHECKLIST OF SHALLOW-WATER MARINE MALACOSTRACAN CRUSTACEA 4
M ethods . . . . . . . . . . . . . . . . . . . . . .
Acknowledgm ents .....................................
Checklist of the Malacostracan Crustacea .......................
Phylum Arthropoda ...................................


. . . .
. . . .
. . . .
. . . . .

. . . .
. . . .

. . .
. .
. . . .
. . . .

. . . .
. . . .
. . . .
. . . .
. . . .
. . . .








. . . .

. . . .
. . . .










...... .
. . .
,. . .


S88
S88
. 89
S89
S89
S89
S90
. 90
S90
S90
. 90
S90
S91
. 91
S91
S91
S91
S91
S91
S91
S91
S91
S91
S92
S92
S92
S92
S92
. 92
S93
S93
S93
S93
S93
S93
S93
.93
S94
S94
S95
S95
S96
S96
S96
. 96
. 96
S96
S96

123
123
124
124
124









Subphylum Crustacea .................... ....................... 124
Class M alacostraca ................... ...... ................ 124
Subclass Phyllocarida ............................... .. .....124
Order Leptostraca ................... ........ .......... 124
Family Nebaliidae ........................ 124
Subclass Hoplocarida .............................. .. .. . ..... 124
Order Stomatopoda ................... ....... ....... 124
Suborder Unipeltata .................... ... .. .............. 1124
Superfamily Gonodactyloidea . . . . . . .. .. . 124
Family Eurysquillidae ........................... 124
Family Gonodactylidae .......................... 124
Family Odontodactylidae ........................... 124
Family Pseudosquillidae .......................... 125
Superfamily Lysiosquilloidea ........................ 125
Family Lysiosquillidae ................... . .. 125
Family Nannosquillidae .................. .. .... 125
Family Tetrasquillidae ........................... 125
Superfamily Squilloidea ........................... 125
Family Squillidae .................... . .. .. 125
Subclass Eumalacostraca .................... .......... ........ .125
Superorder Peracarida ....................... ............. 125
Order Mysidacea .......................................125
Suborder M ysida .................................... 125
Family Mysidae ................................. 125
Subfamily Siriellinae .................... ....... 125
Subfamily Gastrosaccinae ........................ 125
Subfamily Mysinae ................ ........... 125
Order Amphipoda ..................................... 126
Suborder Ingolfiellidea ................................... 126
Family Ingolfiellidae ............................. 126
Suborder Gammaridea ................................... 126
Family Ampeliscidae .................. ........... 126
Family Amphilochidae ........................... 126
Family Ampithoidae .............................. 126
Family Anamixidae ............................... 127
Family Aoridae .................................. 127
Family Argissidae ................................ 127
Family Aristiidae ................................. 127
Family Bateidae ................................. 127
Family Biancolinidae ................. ............ 127
Family Cheluridae ................................ 127
Family Colomastigidae ........................... 127
Family Corophiidae .............................. 128
Subfamily Corophiinae ........................ 128
Family Cyproideidae ................. ............ 128
Family Dexaminidae ............................. 128
Family Epimeriidae .............................. 128
Family Eusiridae ................................. 128
Family Gammaridae .............................. 128
Family Haustoriidae ...................... ...... 129
Family Hyalellidae ................................ .129
Family Hyalidae ................................ 129
Family Iphimediidae ............................. 129
Family Ischyroceridae ............................. 129





































Subord







Order Iso]
Subord



Subord


Family Leucothoidae .......
Family Liljeborgiidae .......
Family Lysianassidae .......
Subfamily Lysianassinae ....
Subfamily Tryphosinae ....
Family Megaluropidae ......
Family Melphidippidae ......
Family Ochlesidae .........
Family Oedicerotidae .......
Family Phliantidae .........
Family Phoxocephalidae .....
Subfamily Brolginae ......
Subfamily Metharpiniinae ..
Family Platyischnopidae .....
Family Pleustidae .........
Subfamily Parapleustinae ...
Subfamily Stenopleustinae .
Family Podoceridae ........
Family Pontoporeiidae ......
Family Sebidae ...........
Family Stenothoidae ........
Family Synopiidae .........
Family Talitridae ..........
er Caprellidea .............
Family Caprellidae ........
Family Caprellinoididae .....
Family Pariambidae ........
Family Phtisicidae .........
Subfamily Phtisicinae .....
Family Protellidae .........
poda . . . . . . . . . .
er Anthuridea .............
Family Anthuridae .........
Family Hyssuridae .........
Family Paranthuridae .......
er Asellota ...............
Superfamily Gnathostenetroidoidea
Family Gnathostenetroididae .
Superfamily Janiroidea ........
Family Incertae Sedis .......
Family Janiridae ..........
Family Joeropsidae ........
Family Munnidae .........
Family Paramunnidae .......
Family Pleurocopidae .......
Family Santiidae ..........
Superfamily Stenetrioidea ......
Family Stenetriidae ........
erEpicaridea ..............
Family Bopyridae .........
Subfamily Athelginae .....
Subfamily Bopyrinae ......
Subfamily Hemiarthrinae ...


. . . . . . . . . . 130
. . . . . . . . . . 130
. . . . . . . . . . 130
. . . . . . . . . . 130
. . . . . . . . . . 130
. . . . . . . . . . 130
. . . . . . . . . . 130
. . . . . . . . . . 130
. . . . . . . . . . 130
. . . . . . . . . . 130
. . . . . . . . . . 131
. . . . . . . . . . 131
. . . . . . . . . . 131
. . . . . . . . . . 131
. . . . . . . . . . 131
. . . . . . . . . . 131
. . . . . . . . . . 131
. . . . . . . . . . 131
. . . . . . . . . . 131
. . . . . . . . . . 131
. . . . . . . . . . 131
. . . . . . . . . . 131
. . . . . . . . . . 131
. . . . . . . . . . 131
. . . . . . . . . . 132
. . . . . . . . . . 132
. . . . . . . . . . 132
. . . . . . . . . . 132
. . . . . . . . . . 132
. . . . . . . . . . 132
. . . . . . . . . . 132
. . . . . . . . . . 132
. . . . . . . . . . 132
. . . . . . . . . . 132
. . . . . . . . . . 132
. . . . . . . . . . 132
. . . . . . . . . . 132
. . . . . . . . . . 132
. . . . . . . . . . 133
. . . . . . . . . . 133
. . . . . . . . . . 133
. . . . . . . . . . 133
. . . . . . . . . . 133
. . . . . . . . . . 133
. . . . . . . . . . 133
. . . . . . . . . . 133
. . . . . . . . . . 133
. . . . . . . . . . 133
. . . . . . . . . . 133
. . . . . . . . . . 133
. . . . . . . . . . 133
. . . . . . . . . . 133
. . . . . . . . . . 133
. . . . . . . . . . 133
. . . . . . . . . . 134


Subord










Subfamily Ioninae .....
Subfamily Pseudioninae .
Family Entoniscidae ......
Suborder Flabellifera ............
Family Aegidae .........
Family Tridentellidae .....
Family Cirolanidae ......
Subfamily Cirolaninae .. .
Subfamily Conilerinae .. .
Subfamily Eurydicinae .. .
Family Corallanidae ......
Family Cymothoidae .....
Family Limnoriidae ......
Family Serolidae ........
Family Sphaeromatidae ....
Subfamily Ancininae ....
Subfamily Cassidininae ..
Subfamily Dynameninae
Subfamily Sphaeromatinae
Suborder Gnathiidea ............
Family Gnathiidae .......
Suborder Microcerberidea ........
Family Microcerberidae ....
Suborder Oniscidea ............
InfraorderTylomorpha .........
Family Tylidae .........
Infraorder Ligiamorpha ........
Section Diplocheta ..........
Family Ligiidae .........
Section Crinocheta ..........
Superfamily Oniscoidea .....
Family Halophilosciidae ...
Family Philosciidae ......
Family Rhyscotidae ......
Family Scyphacidae ......
SuborderValvifera .............
Family Arcturidae .......
Subfamily Arcturinae ...


Subfamily Edwinjoycinae .
Family Chaetiliidae ......
Family Idoteidae ........
Subfamily Idoteinae ....
Order Tanaidacea ...............
Suborder Tanaidomorpha .........
SuperfamilyTanaoidea ......
Family Tanaidae ........
Subfamily Pancolinae ...
Superfamily Paratanaoidea ..
Family Anarthruridae .....
Family Leptocheliidae .....
Family Nototanaidae .....
Family Pseudotanaidae ....
Subfamily Cryptocopinae
Subfamily Pseudotanainae


. . . . . . . . . . . 134
. . . . . . . . . . . 134
. . . . . . . . . . .. 134
................ ...... 134
. . . . . . . . . . . 134
. . . . . . . . . . . 135
. . . . . . . . . . . 135
. . . . . . . . . . . 135
. . . . . . . . . . . 135
. . . . . . . . . . . 135
. . . . . . . . . . . 135
. . . . . . . . . . . 135
. . . . . . . . . . . 135
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 136
. . . . . . . . . . . 137
. . . . . . . . . . . 137
. . . . . . . . . . . 137
. . . . . . . . . . . 137
. . . . . . . . . . . 137
. . . . . . . . . . . 137
. . . . . . . . . . . 137
. . . . . . . . . . . 137
. . . . . . . . . . . 137
. . . . . . . . . . . 137
. . . . . . . . . . . 137
. . . . . . . . . . . 137
. . . . . . . . . . . 137
. . . . . . . . . . . 137
. . . . . . . . . . . 137
. . . . . . . . . . . 137
. . . . . . . . . . . 137
. . . . . . . . . . . 137


@
O









Suborder Apseudomorpha ..........
Superfamily Apseudoidea ......
Family Apseudidae ........
Family Cirratodactylidae .....
Family Kalliapseudidae ......
Family Pagurapseudidae .....
Family Parapseudidae .......
Order Cumacea ..................
Family Bodotriidae ........
Subfamily Bodotriinae .....
Subfamily Mancocuminae .
Subfamily Vaunthompsoniinae
Family Diastylidae .........
Family Leuconidae .... .....
Family Nannastacidae .......
Superorder Eucarida .................
Order Euphausiacea ................
Family Euphausiidae .......
Order Decapoda ..................
Suborder Dendrobranchiata .........
Superfamily Penaeoidea .......
Family Penaeidae .........
Family Sicyoniidae .........
Family Solenoceridae .......
Superfamily Sergestoidea ......
Family Sergestidae .........
Family Luciferidae .........
Suborder Pleocyemata .............
Infraorder Stenopodidea ..........
Family Spongicolidae .......
Family Stenopodidae .......
Infraorder Caridea .............
Superfamily Pasiphaeoidea .....
Family Pasiphaeidae ........
Superfamily Atyoidea ........
Family Atyidae ...........
Superfamily Bresilioidea .......
Family Bresiliidae .........
Superfamily Nematocarcinoidea .
Family Rhynchocinetidae .....
Superfamily Palaemonoidea .....
Family Anchistioididae ......
Family Gnathophyllidae .....
Family Palaemonidae .......
Subfamily Palaemoninae . .
Subfamily Pontoniinae .....
Superfamily Alpheoidea .......
Family Alpheidae .........
Family Hippolytidae ........
Family Ogyrididae .........
Superfamily Processoidea ......
Family Processidae ........
Superfamily Crangonoidea .....










Family Crangonidae ........ ......... ......... 142
Infraorder Astacidea ................................ 142
Superfamily Enoplometopoidea . . . . . . . ... . 142
Family Enoplometopidae ......................... 142
Infraorder Thalassinidea .............................. 142
Superfamily Callianassoidea ........................ 142
Family Callianassidae ..................... ........ 142
Subfamily Callianassinae ....................... 142
Subfamily Callichirinae ........................ 142
Subfamily Cheraminae ........................ 142
Subfamily Eucalliinae ......................... 142
Family Ctenochelidae ........................... 142
Subfamily Ctenochelinae . . . . ......... . 142
Family Laomediidae ............................ 142
Family Upogebiidae ............................ 142
Superfamily Axioidea ............................ 143
Fam ily Axiidae ............................... 143
Family M icheleidae ............................ 143
Infraorder Palinura ................................. 143
Superfamily Palinuroidea .......................... 143
Fam ily Palinuridae ............................. 143
Fam ily Scyllaridae ............................. 143
Subfamily Ibacinae ...........................143
Subfamily Scyllarinae ......................... 143
Family Synaxidae ...... . . . . ........ 143
Infraorder Anomura . . . . ........... . . . . . ... 143
Superfamily Coenobitoidea . . . . . . . ..... ........ 143
Family Coenobitidae . . . . . . . ..... .......... 143
Family Diogenidae ................... .......... .143
Superfamily Paguroidea . . . . . . . ..... ......... 144
Family Paguridae ........ ..... ..... .... ...... 144
Superfamily Galatheoidea .......... ...... ... ..... 144
Family Galatheidae . . . . . . . ..... ........ . 144
Family Porcellanidae ......... .. .... ...... . . 144
Superfamily Hippoidea ......... ........ ........ 144
Family Albuneidae . . . . . . . ..... . . . ... .. 144
Family Hippidae ........ ......... ........... 145
Superfamily Dromioidea ........................... 145
Family Dromiidae ............................. 145
Infraorder Brachyura ................................ 145
Section Archaeobrachyura ........................... 145
Superfamily Homoloidea .......................... 145
Family Cyclodorippidae ......................... 145
Superfamily Raninoidea ........................... 145
Family Raninidae .................. .......... 145
Subfamily Ranininae .......................... 145
Subfamily Notopodinae ........................ 145
Subfamily Symethinae ......................... 145
Section Oxystomata ........... ..... .............. 145
Superfamily Dorippoidea .......................... 145
Family Dorippidae ............................. 145
Subfamily Ethusinae .......................... 145
Superfamily Leucosioidea .......................... 145
Family Calappidae ............................. 145









Subfamily Calappinae ...........


Subfamily Matutinae .......................... 145
Family Leucosiidae ............................ 145
Subfamily Ebaliinae .......................... 145
Subfam ily Iliinae ............................146
Section Oxyrhyncha ............................... 146
Superfamily M ajoidea ............................ 146
Family Majidae ......... . .... .......... .. 146
Subfamily Inachinae ..........................146
Subfamily Epialtinae .......................... 146
Subfamily Pisinae ........................... 146
Subfamily Majinae ............... .......... 146
Subfamily Tychinae ........................... 147
Superfamily Parthenopoidea ........................ 147
Family Parthenopidae ........................... 147
Subfamily Parthenopinae ....................... 147
Section Brachyrhyncha ............................. 147
Superfamily Portunoidea .......................... 147
Family Portunidae ............................. 147
Subfamily Polybiinae .......................... 147
Subfamily Portuninae ......................... 147
Superfamily Xanthoidea ........................... 147
Family Goneplacidae ........................... 147
Family Xanthidae ............................. 148
Superfamily Grapsidoidea .......................... 149
Family Gecarcinidae ............................ 149
Family Grapsidae ............................. 149
Subfamily Grapsinae .......................... 149
Subfamily Plagusiinae ......................... 149
Subfamily Sesarminae ........ ................... 149
SubfamilyVaruninae ..........................150
Superfamily Pinnotheroidea ........................ 150
Family Pinnotheridae ........................... 150
Subfamily Pinnotherinae ....................... 150
Subfamily Pinnothereliinae ...................... 150
Superfamily Ocypodoidea .......................... 150
Family Ocypodidae ............................ 150
Subfamily Ocypodinae ......................... 150
Fam ily Palicidae ............................ 150
Superfamily Cryptochiroidea . . . . . . ... ........ 150
Family Cryptochiridae .......................... 150
References ................... ............................... .. 151
CHECKLIST OF SHALLOW-WATER ECHINODERMATA OF FLORIDA ............... 191
M methods .................... ............................... 191
Acknowledgm ents ... ...........................................191
Checklist of the Echinodermata ..................... .................... 192
Phylum Echinodermata ................... ....................... 192
Class Crinoidea ................... ....................... 192
Order Comatulida ................... .................. 192
Family Comasteridae ........................... 192
Class Asteroidea ................... ....................... 192
Order Paxillosida ... .................... ........ ...... 192
Family Luidiidae ............................. 192


. . . . 14











Family Astropectinidae ..
OrderValvatida ...............
Family Asterinidae .....
Family Goniasteridae .. .
Family Ophidiasteridae .
Family Mithrodiidae ....
Family Asteropseidae .. .
Family Oreasteridae ....
Order Spinulosida .............
Family Echinasteridae . .
Order Forcipulatida ............
Family Asteriidae .....
Class Ophiuroidea ............. .
Order Phrynophiurida ..........
Family Ophiomyxidae . .
Family Gorgonocephalidae
Order Ophiurida ..............
Family Ophiuridae .....
Family Ophiocomidae .. .
Family Ophionereididae
Family Ophiodermatidae
Family Ophiactidae . .
Family Amphiuridae . .
Family Ophiothricidae .
Class Echinoidea ..................
Order Cidaroida ..............
Family Cidaridae ......
Order Diadematoida ...........
Family Diadematidae . .
Order Arbacioida .............
Family Arbaciidae .....
Order Temnopleuroida ..........
FamilyTemnopleuridae ..
Family Toxopneustidae .
Order Echinoida ..............
Family Echinometridae .
Order Holectypoida ............
Family Echinoneidae ..
Order Clypeasteroida ...........
Family Clypeasteridae .
Family Mellitidae .....
Order Spatangoida ............
Family Schizasteridae .. .
Family Brissidae ......
Class Holothuroidea ................
Order Dendrochirotida ..........
Family Cucumariidae .. .
Family Sclerodactylidae.
Family Phyllophoridae .
Order Aspidochirotida ..........
Family Stichopodidae .. .
Family Holothuriidae .. .
Order Molpadiida .............
Family Caudinidae .....


. . . . . . . . . . . 192
. . . . . . . . . . . 192
. . . . . . . . . . . 192
. . . . . . . . . . . 192
. . . . . . . . . . . 192
. . . . . . . . . . . 192
. . . . . . . . . . . 192
.................... 192
..................... 192
. . . . . . . . . . . 192
. . . . . . . . . . . 192
.................... 192
. . . . . . . . . . . 192
.. .................. 192
. . . . . . . . . . 192
. . . . . . . . . . . 193
. . . . . . . . . . . 193
. . . . . . . . . . . 193
. . . . . . . . . . . 193
. . . . . . . . . .. 193
. . . . . . . . . . . 193
. . . . . . . . . . . 193
. . . . . . . . . . . 194
...... .............. 194
..................... 194
. . . . . . . . . . . 194
. . . . . . . . . . . 194
. . . . . . . . . . . 194
. . . . . . . . . . 194
. . . . . . . . . . . 194
. . . . . . . . . . . 194
. . . . . . . . . . . 194
. . . . . . . . . . 194
. . . . . . . . . . 194
. . . . . . . . . . . 194
. . . . . . . . . . . 194
. . . . . . . . . . . 194
. . . . . . . . . . . 194
. . . . . . . . . . .. 194
. . . . . . . . . . . 194
. . . . . . . . . . . 194
. . . . . . . . . . . 194
. . . . . . . . . . . 194
.................. .. 195
. . . . . . . . . . . 195
. . . . . . . . . . .. 195
. . . . . . . . . . . 195
. . . . . . . . . . . 195
. . . . . . . . . . . 195
. . . . . . . . . . . 195
. . . . . . . . . . . 195
. . . . . . . . . . . 195
. . . . . . . . . . . 195









Order Apodida .......................... ................ 195
Family Synaptidae ........................... .195
Family Chiridotidae .............................. 196
References ........................... .......................... 196
INDEX OF SCIENTIFIC NAMES ........ ......... ..................... 209










Rationale and Scope of Checklists

David K. Camp, William G. Lyons, and Thomas H. Perkins


The Florida Department of Environmental Protection
is preparing to use biological criteria to assess human
impacts on the ambient water quality of Florida's es-
tuaries and nearshore coastal zone.The use of biocri-
teria in environmental regulation and management
is based on the principle that species composition and
biotic abundance can serve as indicators of the envi-
ronmental quality of a community and that these bi-
ological factors can be assessed to detect deviations in
community condition and to evaluate trends in envi-
ronmental health (Karr and Chu, 1997). To be able to
perform such assessments, however, ecologists must
have access to resources that will allow them to make
accurate and consistent identifications of the biologi-
cal components of the community. Accuracy and con-
sistency of identification are integral parts of biologi-
cal quality assurance in regulatory ecology, an enter-
prise whose results must be both scientifically valid and
defensible.The more accurate and consistent the iden-
tifications of the biological components of the marine
community are, the more likely it is that trends can be
detected. If early warning signs of environmental im-
pairment in Florida's marine communities are to be rec-
ognized, identifications must be accurate at the species
level.
Identifying the fauna of Florida's estuarine and
coastal marine communities is a challenging task.
Florida is located in the transition area between two ma-
rine zoogeographic zones, the tropical zone to the
south and the temperate zone to the north, and mul-
titudes of species related to each of these zones con-
tribute to the state's fauna, which also contains a sig-
nificant endemic component. As a consequence, more
species of marine organisms may be found in Florida
waters than in those of any other of the continental
United States.
Most of Florida's benthic marine macroinverte-


brate species belong to four principal groups: mol-
lusks, polychaetous annelids, malacostracan crus-
taceans, and echinoderms (Table 1). Species of these
groups are prominent in epibenthic and infaunal sam-
ples taken in shallow waters, and they generally con-
tribute the greatest numbers of species and individu-
als found in these types of samples. Depending on
the habitat and the type of gear used, it is not unusual
for single samples of Florida marine benthos to con-
tain from 30 to more than 100 species belonging to
these four groups. Because of this great diversity, iden-
tifying organisms to the species level can be difficult
and time-consuming.
Common impediments to the identification process
include scattered or incomplete information on a fauna,
out-of-date literature, and occurrences of"new" (un-
described) or extralimital species. Even with the recent
decline of taxonomy as a research discipline, newly de-
scribed marine invertebrate species are being added
to the list of Florida's fauna every year, and specimens
of other still-undescribed species are present in many
museum collections, including the Florida Marine Re-
search Institute's Marine Specimen Collection at St. Pe-
tersburg. Moreover, the classification of the major ma-
rine invertebrate groups is dynamic, changing as the
understanding of phylogenetic relationships increases,
so even quite familiar names may change from time to
time. All of these factors pose problems of accuracy and
consistency for those charged with conducting long-
term, community-level comparisons of fauna within
and between programs and with maintaining Florida's
marine biodiversity.
The production, distribution, and periodic up-dat-
ing of authoritative identification guides would do
much to improve the accuracy and consistency of iden-
tifications (Carney, 1996). By making these guides avail-
able to those engaged in faunal assessments in many


Table 1. Numbers of families, genera, and species-level taxa of shallow-water (<37-m depth) marine Mollusca, poly-
chaetous Annelida, malacostracan Crustacea, and Echinodermata known from Florida in 1997.

Family Genus Species

Mollusca 221 646 1,407
Polychaetous Annelida 58 328 821
Malacostracan Crustacea 155 532 1,135
Echinodermata 39 77 162
Total 473 1,583 3,525



FMRI Technical Report TR-3 1









Rationale and Scope Macroinvertebrate Checklists Camp eta!. 1998


separate investigations, we can help to ensure that the
same animal will be consistently identified by the same
name and that the name selected will be based upon
the most accurate information. Problems posed by re-
classifications or by the appearances of"new"(unde-
scribed, recently described, or extralimital) species
can be mitigated by periodic and timely updates of the
guides.
A first step in producing identification guides is to
prepare authoritative, up-to-date checklists of all
species of the major marine benthic macroinverte-
brate groups known from shallow Florida waters. Such
checklists will serve two purposes-they will provide
ecologists with up-to-date names for most of Florida's
benthic macroinvertebrates, and they will provide a
basis from which to plan the scope and production of
illustrated guides to the marine fauna of Florida. The
purpose of this paper is to initiate that first step. An
electronic network (perhaps via an Internet website)
should also be established to allow easy access to the
checklists and to inform Florida ecologists of nomen-
clatural changes to the checklists and of associated
identification literature.
The checklists of mollusks, polychaetous annelids,
malacostracan crustaceans, and echinoderms that fol-
low list all species of these groups known to occur in
Florida's estuarine and coastal marine waters offshore
to depths of approximately 37 meters (20 fathoms or
120 feet).The 37-m depth was selected as a deep-water
boundary because this depth has been shown to be an
approximate point of transition between the faunas of
shallow coastal zones and those of intermediate and
deeper shelf zones, at least off western Florida (Lyons,
1980; Lyons and Camp, 1983). Although such points of
transition can differ between animal groups and, cer-
tainly, among locations in a region as large and envi-
ronmentally diverse as Florida, we have found that
many of the species that demonstrate fidelity to par-
ticular depth zones off western Florida maintain that
fidelity in other areas of the state. As a caveat to that
generalization, it is important to recognize the phe-
nomenon of latitudinal submergence-species that
inhabit very shallow waters in the Florida Keys and
along southeastern Florida may be found only in deeper
waters farther north. However, the converse is seldom
true-benthic macroinvertebrate species that cus-
tomarily inhabit Florida's intermediate and deeper
shelf communities are rarely found in inshore waters
of Florida (although a few occur in lesser depths off
New England).
The checklists should not be considered a complete
listing of all species that live in Florida waters, be-
cause undescribed species in museum collections are
usually not included, and many species currently


known only from nearby states and countries (e.g., Ba-
hama Islands, Cuba) could also occur in Florida waters.
These lists are works in progress, and they should be
updated regularly. Compilation of this version ended
in November 1997.
Individual accounts of methods and acknowledg-
ments related to the creation of each list are presented
below, and the list for each group is accompanied by
a separate compilation of pertinent literature.The ref-
erences cited in those compilations include all litera-
ture sources consulted during the preparation of that
list, as well as some other references useful in classi-
fying and identifying organisms of that group. Many
other references have also been included to guide the
user to pertinent information on the communities in
which the organisms live and the forces that may im-
pact those organisms and their communities; how-
ever, the lists of references do not include all papers
that have treated Florida's benthic invertebrates. The
sound management of Florida's marine resources will
depend not only upon the consistent and accurate
identification of organisms but also upon an under-
standing and appreciation of the composition and dy-
namics of the communities in which these organisms
live.

Acknowledgments

We thank the following people, who contributed their
time and effort to improving the quality of this check-
list. Staff of the library at Florida Marine Research In-
stitute (FMRI) (Jan Boyett, Bob Brock, Felice Drouin, and
James Thomas) acquired and provided copies of many
of the publications we consulted. James F. Quinn, Jr.,
Llyn French, and Judy Leiby, of the FMRI Publications
Production Office, edited our copy, assembled our sev-
eral word-processing files into the finished document,
and assisted in improving this work. Funding was pro-
vided by Florida Department of Environmental Pro-
tection, Division of Water Facilities, Bureau of Water Re-
sources Protection from FY 1997 U.S. Environmental
Protection Agency 604(b) grant monies administered
by Mary Paulic. We are grateful to the department for
giving us the opportunity to assemble these lists and
associated references. Finally, we are especially grate-
ful to Paul Humann, marine life author and photog-
rapher, for allowing several of his excellent photographs
to grace the cover of this document.

Literature Cited

CARNEY, R. S. 1996. On the adequacy and improvement
of marine benthic pre-impact surveys. Pp. 295-315 in
Schmitt, R. J., and C. W. Osenberg (eds.), Detecting Eco-


2 FMRI Technical Report TR-3


Rationale and Scope


Macroinvertebrate Checklists


Camp et al. 1998







Macroinvertebrate Checklists


logical Impacts: Concepts and Applications in Coastal
Habitats. Academic Press, NewYork. xx + 401 pp.

KARR, J. R., and E. W. CHU. 1997. Biological monitor-
ing and assessment: Using multimetric indexes effec-
tively. EPA 235-R97-001. University of Washington,
Seattle. vii + 149 pp.

LYONS, W. G. 1980. Molluscan communities of the


West Florida Shelf. Bulletin of the American Malaco-
logical Union, Inc. (1979): 37-40.

LYONS, W. G., and D. K. CAMP. 1983. Zones of faunal
similarity within the Hourglass study area. Pp. 44-46
in Proceedings of the Third Annual Gulf of Mexico
Information Transfer Meeting, December 1982. United
States Department of the Interior, Minerals Manage-
ment Service, New Orleans.


FMRI Technical Report TR-3


Camp et al. 1998


Rationale and Scope













Checklist of Shallow-Water Marine Mollusca of Florida

William G. Lyons


Methods

The initial draft of this list was based, in part, on in-
formation in American Seashells, second edition (Abbott,
1974) and on lists of mollusks prepared by the Coun-
cil of Systematic Malacologists and the American Mala-
cological Union for the American Fisheries Society
(Turgeon et al., 1988; Turgeon et al., 1998). The Florida
list was created by selecting from those larger lists the
estuarine and marine species known from eastern
North America and then by reducing that set of names,
first by deleting the names of species not known from
Florida and then by deleting the names of several hun-
dred species known only from intermediate and deep-
water regions of the continental shelf off Florida. Many
other published and unpublished sources on occur-
rences of mollusks in Florida within the shallow-water
limits set for this list were then examined to confirm
that the names remaining were appropriate for listing.
Regional information sources were consulted for the
Jacksonville area (H. G. Lee, unpublished manuscript);
the Indian River Lagoon (Mikkelsen et al., 1995); the
continental shelf off central and northeastern Florida
(FMRI SEAMAP project, in prep.); the Oculina reefs off
eastern Florida (Reed and Mikkelsen, 1987); shallow
coastal waters off Hutchinson Island (Lyons, 1989);
Peanut Island at Lake Worth Inlet (Lee, 1997b); Biscayne
Bay and nearby environs (Voss and Voss, 1955; Mc-
Nulty, 1961; McNulty et al., 1962; O'Gower and Wacasey,
1967; H. B. Moore et al., 1968; and numerous papers by
Ev. Marcus, Er. Marcus, D. R. Moore, and others); the
Florida Keys National Marine Sanctuary (Lyons and
Quinn, 1995); Florida Bay (Tabb and Manning, 1961; Tur-
ney and Perkins, 1972; FMRI Florida Bay benthic fau-
nal mapping project, in prep.); southwestern Florida
(Perry and Schwengel, 1955); Charlotte Harbor (Es-
tevez, 1986); Tampa Bay and Boca Ciega Bay (Hall and
Saloman, 1975; many other papers in unpublished
database compiled at FMRI by WGL); Crystal River re-
gion (Lyons et al., 1971); Cedar Key region (Dall, 1884;
Lee, 1997a); Florida Middle Ground (Hopkins et al.,
1977; Turgeon and Lyons, 1978; unpublished species list
by Turgeon and Lyons); Apalachee Bay region (Men-
zel, 1971); St. Andrew Bay (Saloman et al., 1982), Pen-
sacola Bay region (Cooley, 1978; Frey and Dirjes, 1988);
shallow coastal waters off western and southeastern
Florida (Godcharles and Jaap, 1971); and the conti-
nental shelf off western Florida (D. R. Moore, 1977;


Bishof, 1980; Lyons, 1980; mollusks of Project Hourglass,
FMRI unpublished). Additional records were con-
sulted in a multitude of other papers.
The names on this list are presented in a phylo-
genetic arrangement of phylum, class, subclass, order,
and family, with genera and species arranged alpha-
betically within families.The sequence of listing follows
that of Turgeon et al. (1998) and is an accurate repre-
sentation of the prevalent classification theory at this
time. The format of the list differs from that of the
larger lists (e.g., Turgeon et al., 1998) by including a
category for subclass and also by providing authorities
(original authors and dates) for higher taxa (phylum,
class, subclass, order, family, and genus). Like the
names themselves, the authorities for higher taxa were
extracted from many sources, among the more promi-
nent of which are Abbott (1974), H. E. Vokes (1980),
Ponder and Waren (1988),Vaught (1989), Millard (1996),
and Coan and Scott (1997). However, it was also nec-
essary to consult many other works to obtain author-
ities for some of the names.
Three species included here are not listed by Tur-
geon et al. (1998) because a criterion for listing in the
latter work requires prior publication of the informa-
tion elsewhere, and there seem to be no published
records of the following species in North America: 1)
Solariorbis semipuncta D. R. Moore, 1965: fresh-dead
shells and fragments among coralline sand in pockets
atop reef structures, depth about 23-24 m, Florida Mid-
dle Ground, northeastern Gulf of Mexico, D. D.Turgeon
and W G. Lyons, unpublished data; 2) Careliopsis bartschi
(Aguayo and Rehder, 1936): Florida Bay, W. G. Lyons,
unpublished data; 3) Bornella calcarata Morch, 1863:
shallow to intermediate shelf waters off northeastern
Florida, SEAMAP benthic community study; also taken
commonly there by a commercial shell dredger dur-
ing the 1970s. Another species, ErycinafloridanaVanatta,
1904, described from Florida, was overlooked in other
recent compilations of taxa (e.g., Abbott, 1974; Turgeon
et al., 1988, 1998).
Undescribed species are not included on this list.
Seemingly unnamed species belonging to the families
Triphoridae, Eulimidae, Buccinidae, Nassariidae,
Columbellidae, Turridae, Conidae (subfamily Man-
geliinae), and Pyramidellidae from shallow waters of
Florida are known to various workers. However, few if
any of these species have been figured or diagnosed
in publications, so nothing would be served by listing


FMRI Technical Report TR-3 5








Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al, 1998


unreferenced names (e.g., Nassarius sp. A). On an en-
couraging note, several of these species are the subjects
of ongoing studies, so names for some of them may be
available soon.
Copies of the draft list were sent to three review-
ers (see acknowledgments), each of whom provided
comments that improved the final document.

References

The accompanying compendium of references pre-
sents citations for nearly 1,100 books, journal articles,
dissertations, theses, contract reports, and popular ar-
ticles containing information on the classification,
identification, distribution, life history, and ecology of
the shallow-water mollusks of Florida. The list is by no
means complete. For example, the classic works in
which many of Florida's most well-known mollusks
were described (e.g., the works of Linnaeus, Gmelin,
Lamarck, Say, and others) have been omitted, although
an attempt has been made to include early faunal sur-
veys that documented and described Florida species
(e.g., Conrad, 1846; Calkins, 1878; Dall, 1884). Many
popular guides to the identification of Florida mol-
lusks have been listed, but some are included princi-
pally for their historical value (e.g., Aldrich and Sny-
der, 1936; Morris, 1947), and some of the less useful pa-
perback guides to seashells have been omitted.
Likewise, most of the popular guides to worldwide
seashells have been omitted, even though some of
them include information on Florida shells. Many pa-
pers concerned only with the mollusks of Florida's
deeper-water shelf and slope have also been omitted.
Papers describing the births, romances, diets, in-
terspecific associates, competitive struggles, deaths,
and similar facets of the natural history of Florida mol-
lusks have been included on the assumption that such
information can lead to a better understanding of the
organisms by those who would study or manage them.
On the other hand, reports of the ordinary data of
fisheries (effort, landings, and economics) have been
excluded, as such information is generally available
elsewhere. Also excluded are papers on the more com-
plex aspects of molluscan biology, such as anatomy, em-
bryology, and biochemistry. Thus, a paper on prey
preferences of a predatory gastropod would be in-
cluded, but a paper describing the cell structure of
that predator's stomach wall would not.
One very useful source of information on Florida's
molluscan fauna is S. D. Kaicher's Card Catalogue of
World-Wide Shells, a series of 60 card packs published
individually between 1973 and 1992. Each pack consists
of approximately 108 3 x 5 cards dedicated to a single
family (or occasionally two related families) of marine


gastropods, and each card presents the name, range,
diagnostic comments, and illustrations for a different
species of that family. Nearly all of the packs contain
cards treating species of the Florida fauna, but such in-
formation is readily available elsewhere for some fam-
ilies (e.g., Strombidae), so those packs are not cited
here. In other cases, however, the information in the
card packs is superior in accuracy and quality of il-
lustration to any other information on those species,
and an effort has been made to list the most pertinent
of those packs. Examples of packs especially useful as
identification aids for Florida species include those
on Olividae (especially Olivella), Nassariidae, andTur-
ridae.
A few articles from shell club newsletters have
been included as examples of this information re-
source. Such articles can provide a wealth of informa-
tion on local faunas, and in many instances they con-
stitute the only record of local occurrence for a given
species. Many useful and informative articles have
been published in Shell-O-Gram, the newsletter of the
Jacksonville Shell Club, in Seafari, the now-defunct
newsletter of the Palm Beach County Shell Club, and
in similar publications. Some of those newsletter ar-
ticles have been later cited in scientific reports, but this
is not peer-reviewed literature, and such articles
should be used with caution. On the other hand, some
of the shell collectors who write these articles are more
knowledgable identifiers of local fauna than are some
of the biologists who identified mollusks for some of
the environmental surveys cited in the present com-
pendium. In fact, there seems to be an increasing ten-
dency towards the publication of incorrect molluscan
identifications in the peer-reviewed scientific literature,
especially in papers reporting general faunal assess-
ments, and some of those papers listed here contain
remarkable errors. The best rule is to accept that in-
formation which seems reasonable and seek verifica-
tion for that which does not, regardless of the source.

Acknowledgments

Dr. Paula M. Mikkelsen, Department of Invertebrates,
American Museum of Natural History, NewYork, and
Dr. James F. Quinn, Jr., Florida Department of Envi-
ronmental Protection, Florida Marine Research Insti-
tute, St. Petersburg, each reviewed the entire draft list,
and Dr. Kerry B. Clark, Department of Biological Sci-
ences, Florida Institute of Technology, Melbourne, re-
viewed sections on aplacophorans and heterobranch
and opisthobranch gastropods. Each of the reviewers
has extensive experience working with the marine
molluscan fauna of Florida, and each contributed valu-
able comments that improved the list.This effort also


6 FMRI Technical Report TR-3


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Mollusca--W. G. Lyons


Macroinvertebrate Checklists








Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


benefited from information provided by many others
who contributed to or reviewed manuscript drafts of
the American Fisheries Society list (Turgeon et al.,
1998). Notable among these are Mr. Kurt Auffenberg,
Florida Museum of Natural History, Gainesville; Dr.
Robert C. Bullock, University of Rhode Island, Kingston;


Mr. EugeneV. Coan, Palo Alto, California; Dr.Terrence
Gosliner, California Academy of Science, San Fran-
cisco; Dr. Harry G. Lee, Jacksonville, Florida; Dr. Gary
McDonald, Long Marine Laboratory, Santa Cruz, Cal-
ifornia; and Dr. Gary Rosenberg, Academy of Natural
Sciences of Philadelphia, Pennsylvania.


Checklist of the Mollusca


Phylum Mollusca
Class Aplacophora von Ihering, 1876
Solenogasters and Glisten-worms

Order Pholidoskepia Salvini-Plawen, 1978
Family Meiomeniidae Salvini-Plawen, 1985

Genus Meiomenia Morse, 1979
Meiomenia arenicola Salvini-Plawen, 1985

Class Polyplacophora J. E. Gray, 1821
Chitons

Order Neoloricata Bergenhayn, 1955
Family Hanleyidae Bergenhayn, 1955

Genus Hanleya J. E. Gray, 1857
Hanleya tropicalis Dall, 1881

Family Ischnochitonidae Dall, 1889

Genus Ischnochiton J. E. Gray, 1847
Ischnochiton dilatosculptus Kaas, 1982
Ischnochiton erythronotus (C. B. Adams, 1845)
Ischnochiton hartmeyeri Thiele, 1910
Ischnochiton niveus Ferreira, 1987
Ischnochiton papillosus (C. B. Adams, 1845)
Ischnochiton pseudovirgatus Kaas, 1972
Ischnochiton striolatus (J. E. Gray, 1828)

Genus Stenoplax Carpenter in Dall, 1879
Stenoplax bahamensis Kaas &Van Belle, 1987
Stenoplax boogii (Haddon, 1886)
Stenoplaxfloridana (Pilsbry, 1892)

Family Callistoplacidae Pilsbry, 1893

Genus Callistochiton Carpenter in Dall, 1879
Callistochiton portobelensis Ferreira, 1976
Callistochiton shuttleworthianus Pilsbry, 1893

Family Chaetopleuridae Plate, 1899

Genus Chaetopleura Shuttleworth, 1853
Chaetopleura apiculata (Say, 1834)


Chaetopleura janeirensis (J. E. Gray, 1828)
Chaetopleura staphylophera Lyons, 1985

Family Chitonidae Rafinesque, 1815

Genus Acanthopleura Guilding, 1829
Acanthopleura granulata (Gmelin, 1791)

Genus Chiton Linnaeus, 1758
Chiton marmoratus Gmelin, 1791
Chiton tuberculatus Linnaeus, 1758
Chiton viridis Spengler, 1797

Genus Tonicia J. E. Gray, 1847
Tonicia schrammi (Shuttleworth, 1856)

Family Tonicellidae Simroth, 1894

Genus Lepidochitona J. E. Gray, 1821
Lepidochitona liozonis (Dall & Simpson, 1901)

Family Mopaliidae Dall, 1889

Genus Ceratozona Dall, 1882
Ceratozona squalida (C. B. Adams, 1845)

Family Acanthochitonidae Pilsbry, 1893

Genus Acanthochitona J. E. Gray, 1821
Acanthochitona balesae Abbott, 1954
Acanthochitona hemphilli (Pilsbry, 1893)
Acanthochitona pygmaea (Pilsbry, 1893)
Acanthochitona roseojugum Lyons, 1988
Acanthochitona zebra Lyons, 1988

Genus Americhiton Watters, 1990
Americhiton andersoni (Watters, 1981)

Genus Cryptoconchus Burrow, 1815
Cryptoconchus floridanus (Dall, 1889)

Class Scaphopoda Bronn, 1862
Tuskshells and Toothshells

Order Dentaliida Da Costa, 1776


FMRI Technical Report TR-3 7


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W. G. Lyons--Mollusca








Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


Family Dentaliidae J. E. Gray, 1834

Genus Antalis H. and A. Adams, 1854
Antalis antillarum (d'Orbigny, 1842)
Antalis pilsbryi (Rehder, 1942)
Antalis taphrium (Dall, 1889)

Genus Dentalium Linnaeus, 1758
Dentalium americanum Chenu, 1843
Dentalium laqueatum A. E.Verrill, 1885

Genus Graptacme Pilsbry & Sharp, 1897
Graptacme calamus (Dall, 1889)
Graptacme eborea (Conrad, 1846)
Graptacme semistriolata (Guilding, 1834)

Family Gadilinidae Chistikov, 1975

Genus Episiphon Pilsbry & Sharp, 1897
Episiphon sowerbyi (Guilding, 1834)

Order Gadilida Starobogatov, 1974
Family Gadilidae Stoliczka, 1868

Genus Gadila J. E. Gray, 1847
Gadila mayor (J. B. Henderson, 1920)

Genus Polyschides Pilsbry & Sharp, 1897
Polyschides agassizii (Dall, 1881)
Polyschides carolinensis (Bush, 1895)
Polyschides quadridentatus (Dall, 1881)
Polyschides tetrodon (Pilsbry & Sharp, 1898)

Class Bivalvia Linnaeus, 1758
Mussels, Scallops, Oysters, Clams, etc.

Order Solemyoida Dall, 1889
Family Solemyidae J. E. Gray, 1840

Genus Solemya Lamarck, 1818
Solemya occidentalis Deshayes, 1857
Solemya velum Say, 1822

Order Nuculoida Dall, 1889
Family Nuculidae J. E. Gray, 1824

Genus Nucula Lamarck, 1799
Nucula calcicola D. R. Moore, 1977
Nucula proxima Say, 1822

Family Nuculanidae H. & A. Adams, 1858

Genus Nuculana Link, 1807


Nuculana acuta (Conrad, 1832)
Nuculana concentrica (Say, 1824)

Order Mytiloida Firussac, 1822
Family Mytilidae Rafinesque, 1815

Genus Amygdalum Miihlfeld, 1811
Amygdalum papyrium (Conrad, 1846)

Genus Botula Morch, 1853
Botulafusca (Gmelin, 1791)

Genus Brachidontes Swainson, 1840
Brachidontes domingensis (Lamarck, 1819)
Brachidontes exustus (Linnaeus, 1758)
Brachidontes modiolus (Linnaeus, 1767)

Genus Crenella Brown, 1827
Crenella decussata (Montagu, 1808)

Genus Dacyridium Torell, 1859
Dacrydium elegantulum hendersoni Salas & Gofas,
1997

Genus Geukensia Poel, 1959
Geukensia demissa (Dillwyn, 1817)
Geukensia granosissima (G. B. Sowerby III, 1914)

Genus Gregariella Monterosato, 1884
Gregariella coralliophaga (Gmelin, 1791)

Genus Ischadium Jukes-Brown, 1905
Ischadium recuroum (Rafinesque, 1820)

Genus Lioberus Dall, 1898
Lioberus castaneus (Say, 1822)

Genus Lithophaga Roding, 1798
Lithophaga antillarum (d'Orbigny, 1842)
Lithophaga aristata (Dillwyn, 1817)
Lithophaga bisulculata (d'Orbigny, 1842)
Lithophaga nigra (d'Orbigny, 1842)

Genus Modiolus Lamarck, 1799
Modiolus americanus (Leach, 1815)
Modiolus modiolus squamosus Beauperthuy, 1967

Genus Musculus R6ding, 1798
Musculus lateralis (Say, 1822)

Genus Mytilus Linnaeus, 1758
Mytilus edulis Linnaeus, 1758


8 FMRI Technical Report TR-3


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Macro! nverte brate Checklists


Camp et al. 1998







Macroinvertebrate Checklists


Order Arcoida Stoliczka, 1871
Family Arcidae Lamarck, 1809

Genus Acar J. E. Gray, 1842
Acar domingensis (Lamarck, 1819)

Genus Anadara J. E. Gray, 1847
Anadara brasiliana (Lamarck, 1819)
Anadarafloridana (Conrad, 1869)
Anadara notabilis (Roding, 1798)
Anadara ovalis (Bruguiere, 1789)
Anadara transversa (Say, 1822)

Genus Arca Linnaeus, 1758
Arca imbricata Bruguiere, 1789
Arca zebra (Swainson, 1833)

Genus Barbatia J. E. Gray, 1842
Barbatia cancellaria (Lamarck, 1819)
Barbatia candida (Helbling, 1779)
Barbatia tenera (C. B. Adams, 1845)

Family Noetiidae Stewart, 1930

Genus Arcopsis von Koenen, 1885
Arcopsis adamsi (Dall, 1886)

Genus Noetia J. E. Gray, 1857
Noetia ponderosa (Say, 1822)

Family Glycymerididae Newton, 1916

Genus Glycymeris da Costa, 1778
Glycymeris americana (DeFrance, 1829)
Glycymeris decussata (Linnaeus, 1758)
Glycymeris pectinata (Gmelin, 1791)
Glycymeris spectralis Nicol, 1952
Glycymeris undata (Linnaeus, 1758)

Order Pterioida Newell, 1965
Family Pteriidae J. E. Gray, 1847

Genus Pinctada Rbding, 1798
Pinctada imbricata Roding, 1798
Pinctada margaritifera (Linnaeus, 1758)

Genus Pteria Scopoli, 1777
Pteria colymbus (R6ding, 1798)
Pteria longisquamosa (Dunker, 1852)

Family Isognomonidae Woodring, 1925

Genus Isognomon Lightfoot, 1786
Isognomon alatus (Gmelin, 1791)


Isognomon bicolor (C. B. Adams, 1845)
Isognomon radiatus (Anton, 1839)

Family Malleidae Lamarck, 1819

Genus Malleus Lamarck, 1799
Malleus candeanus (d'Orbigny, 1842)

Family Pinnidae Leach, 1819

Genus Atrina J. E. Gray, 1842
Atrina rigida (Lightfoot, 1786)
Atrina seminuda (Lamarck, 1819)
Atrina serrata (Sowerby, 1825)

Genus Pinna Linnaeus, 1758
Pinna carnea Gmelin, 1791

Order Limoida Waller, 1978
Family Limidae Rafinesque, 1815

Genus Ctenoides Morch, 1853
Ctenoides scabra (Born, 1778)
Ctenoides tenera (Sowerby, 1843, non Turton, 1822)

Genus Lima Bruguiere, 1797
Lima lima (Linnaeus, 1758)

Genus Limaria Link, 1807
Limaria locklini (McGinty, 1955)
Limaria pellucida (C. B. Adams, 1846)

Genus Limatula S.V. Wood, 1839
Limatula hendersoni Olsson & McGinty, 1958
Limatula subauriculata (Montagu, 1808)

Order Ostreoida Ffrussac, 1822
Family Pectinidae Rafinesque, 1815

Genus Argopecten Monterosato, 1889
Argopecten gibbus (Linnaeus, 1758)
Argopecten irradians (Lamarck, 1819)
Argopecten lineolaris (Lamarck, 1819)
Argopecten nucleus (Born, 1778)

Genus Bractechlamys Iredale, 1939
Bractechlamys antillarum (Recluz, 1853)

Genus Caribachlamys Waller, 1993
Caribachlamys imbricata (Gmelin, 1791)
Caribachlamys mildredae (Bayer, 1941)
Caribachlamys ornata (Lamarck, 1819)
Caribachlamys sentis (Reeve, 1853)


FMRI Technical Report TR-3


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W. G. Lyons--Mollusca








Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


Genus Euvola Dall, 1898
Euvola raveneli (Dall, 1898)
Euvola ziczac (Linnaeus, 1758)

Genus Laevichlamys Waller, 1993
Laevichlamys multisquamata (Dunker, 1864)

Genus Lindapecten Petuch, 1995
Lindapecten exasperatus (G. B. Sowerby II, 1842)
Lindapecten muscosus (W. Wood, 1828)

Genus Nodipecten Dall, 1898
Nodipecten nodosus (Linnaeus, 1758)

Genus Spathochlamys Waller, 1993
Spathochlamys benedicti (A. E.Verrill & Bush,
1897)

Family Propeamussidae Abbott, 1954

Genus Cyclopecten A. E.Verrill, 1897
Cyclopecten nanus A. E.Verrill & Bush, 1897

Family Plicatulidae Watson, 1930

Genus Plicatula Lamarck, 1801
Plicatula gibbosa Lamarck, 1801

Family Spondylidae J. E. Gray, 1826

Genus Spondylus Linnaeus, 1758
Spondylus americanus Hermann, 1781
Spondylus ictericus Reeve, 1856

Family Anomiidae Rafinesque, 1815

Genus Anomia Linnaeus, 1758
Anomia simplex d'Orbigny, 1842

Genus Pododesmus Philippi, 1837
Pododesmus rudis (Broderip, 1834)

Family Ostreidae Rafinesque, 1815

Genus Crassostrea Sacco, 1897
Crassostrea virginica (Gmelin, 1791)

Genus Cryptostrea Harry, 1985
Cryptostrea permollis (G. B. Sowerby II, 1871)

Genus Dendrostrea Sowerby, 1839
Dendostrea frons (Linnaeus, 1758)

Genus Ostrea Linnaeus, 1758


Ostrea equestris Say, 1834

Genus Teskeyostrea Harry, 1985
Teskeyostrea weberi (Olsson, 1951)

Family Gryphaeidae Vyalov, 1936

Genus Parahyotissa Harry, 1985
Parahyotissa mcgintyi Harry, 1985

OrderVeneroida H. & A. Adams, 1856
Family Lucinidae Fleming, 1828

Genus Anodontia Link, 1807
Anodontia alba Link, 1807
Anodontia philippiana (Reeve, 1850)

Genus Codakia Scopoli, 1777
Codakia orbicularis (Linnaeus, 1758)
Codakia orbiculata (Montagu, 1808)
Codakia pectinella (C. B. Adams, 1852)

Genus Divalinga Chavan, 1951
Divalinga quadrisulcata (d'Orbigny, 1842)

Genus Divaricella von Martens, 1880
Divaricella dentata (W. Wood, 1815)

Genus Lucina Bruguiere, 1797
Lucina amianta (Dall, 1901)
Lucinafloridana Conrad, 1833
Lucina keenae Chavan, 1971
Lucina leucocyma Dall, 1886
Lucina pectinata (Gmelin, 1791)
Lucina pensylvanica (Linnaeus, 1758)
Lucina sombrerensis Dall, 1886
Lucina trisulcata Conrad, 1841

Genus Lucinisca Dall, 1901
Lucinisca muricata (Spengler, 1798)
Lucinisca nassula (Conrad, 1846)

Genus Parvilucina Dall, 1901
Parvilucina costata (d'Orbigny, 1842)
Parvilucina multilineata (Tuomey & Holmes, 1857)

Family Ungulinidae H. & A. Adams, 1856

Genus Diplodonta Bronn, 1831
Diplodonta candeana (d'Orbigny, 1842)
Diplodonta notata Dall & Simpson, 1901
Diplodonta nucleiformis (Wagner, 1838)
Diplodonta punctata (Say, 1822)
Diplodonta semiaspera Philippi, 1836


10 FMRI Technical Report TR-3


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Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


Diplodonta soror (C. B. Adams, 1852)
Diplodonta subglobosa (C. B. Adams, 1852)
Diplodonta venezuelensis (Dunker, 1848)

Family Cyrenoididae H. & A. Adams, 1857

Genus Cyrenoida Joannis, 1835
Cyrenoidafloridana (Dall, 1896)

Family Chamidae Lamarck, 1809

Genus Arcinella Schumacher, 1817
Arcinella cornuta Conrad, 1866

Genus Chama Linnaeus, 1758
Chama congregate Conrad, 1833
Chama florida Lamarck, 1819
Chama macerophylla Gmelin, 1791
Chama sarda Reeve, 1847
Chama sinuosa Broderip, 1835

Genus Pseudochama Odhner, 1917
Pseudochama inezae Bayer, 1943
Pseudochama radians (Lamarck, 1819)

Family Lasaeidae J. E. Gray, 1842

Genus Aligena H. C. Lea, 1843
Aligena texasiana Harry, 1969

Genus Erycina Lamarck, 1805
Erycinafloridana Vanatta, 1904

Genus Lasaea Brown, 1827
Lasaea adansoni (Gmelin, 1791)

Genus Mysella Angas, 1877
Mysella planulata (Stimpson, 1851)

Genus Orobitella Dall, 1900
Orobitellafloridana (Dall, 1899)
Orobitella limpida (Dall, 1899)

Genus Parabornia Boss, 1965
Parabornia squillina Boss, 1965

Genus Pythinella Dall, 1899
Pythinella cuneata (A. E.Verrill & Bush, 1898)

Family Galeommatidae J. E. Gray, 1840

Genus Divariscintilla A. W. B. Powell, 1932
Divariscintilla cordiformis P. M. Mikkelsen &
Bieler, 1992


Divariscintilla luteocrinita P. M. Mikkelsen &
Bieler, 1992
Divariscintilla octotentaculata P. M. Mikkelsen &
Bieler, 1992
Divariscintilla troglodytes P. M. Mikkelsen &
Bieler, 1989
Divariscintilla yoyo P. M. Mikkelsen & Bieler, 1989

Family Sportellidae Dall, 1899

Genus Basterotia Hoernes, 1859
Basterotia elliptica (Recluz, 1850)
Basterotia quadrata (Hinds, 1843)

Genus Ensitellops Olsson & Harbison, 1953
Ensitellops protextus (Conrad, 1841)

Family Carditidae Fleming, 1828

Genus Carditamera Conrad, 1838
Carditamerafloridana Conrad, 1838

Genus Glans Miihlfeld, 1811
Glans dominguensis (d'Orbigny, 1842)

Genus Pleuromeris Conrad, 1867
Pleuromeris tridentata (Say, 1826)

Genus Pteromeris Conrad, 1862
Pteromeris perplana (Conrad, 1841)

Family Condylocardiidae Bernard, 1897

Genus Carditopsis E. A. Smith, 1881
Carditopsis smithii (Dall, 1896)

Genus Cuna Hedley, 1902
Cuna dalliVanatta, 1904

Family Astartidae d'Orbigny, 1844

Genus Astarte Sowerby, 1816
Astarte nana Dall, 1886

Family Crassatellidae Ferussac, 1822

Genus Crassinella Guppy, 1874
Crassinella dupliniana (Dall, 1903)
Crassinella lunulata (Conrad, 1834)
Crassinella martinicensis (d'Orbigny, 1842)

Genus Eucrassatella Iredale, 1924
Eucrassatella speciosa (A. Adams, 1852)


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W. G, Lyons--Mollusca








Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


Family Cardiidae Lamarck, 1809

Genus Americardia Stewart, 1930
Americardia guppyi (Thiele, 1910)
Americardia media (Linnaeus, 1758)

Genus Dinocardium Dall, 1900
Dinocardium robustum (Lightfoot, 1786)

Genus Laevicardium Swainson, 1840
Laevicardium laevigatum (Linnaeus, 1758)
Laevicardium mortoni (Conrad, 1831)
Laevicardium pictum (Ravenel, 1861)
Laevicardium sybariticum (Dall, 1886)

Genus Papyridea Swainson, 1840
Papyridea semisulcata (J. E. Gray, 1825)
Papyridea soleniformis (Bruguiere, 1789)

Genus Trachycardium Morch, 1853
Trachycardium egmontianum (Shuttleworth, 1856)
Trachycardium magnum (Linnaeus, 1758)
Trachycardium muricatum (Linnaeus, 1758)

Family Mactridae Lamarck, 1809

Genus Anatina Schumacher, 1817
Anatina anatina (Spengler, 1802)

Genus Mactrotoma Dall, 1894
Mactrotomafragilis (Gmelin, 1791)

Genus Mulinia J. E. Gray, 1837
Mulinia lateralis (Say, 1822)

Genus Raeta J. E. Gray, 1853
Raeta plicatella (Lamarck, 1818)

Genus Rangia Desmoulins, 1832
Rangia cuneata (Sowerby, 1831)

Genus Spisula J. E. Gray, 1837
Spisula raveneli (Conrad, 1831)

Family Solenidae Lamarck, 1809

Genus Solen Linnaeus, 1758
Solen viridis Say, 1822

Family Pharidae H. & A. Adams, 1858

Genus Ensis Schumacher, 1817
Ensis megistus Pilsbry & McGinty, 1943
Ensis minor Dall, 1900


Family Tellinidae de Blainville, 1814

Genus Cymatoica Dall, 1889
Cymatoica orientalis (Dall, 1890)

Genus Leporimetis Iredale, 1930
Leporimetis intastriata (Say, 1826)

Genus Macoma Leach, 1819
Macoma brevifrons (Say, 1834)
Macoma cerina Dall, 1900
Macoma constricta (Bruguiere, 1792)
Macoma limula Dall, 1895
Macoma mitchelli Dall, 1895
Macoma phenax Dall, 1900
Macoma tenta (Say, 1834)

Genus Strigilla Turton, 1822
Strigilla carnaria (Linnaeus, 1758)
Strigilla gabbi Olsson & McGinty, 1958
Strigilla mirabilis (Philippi, 1841)
Strigilla surinamensis Boss, 1972

Genus Tellidora H. & A. Adams, 1856
Tellidora cristata (Rcluz, 1842)

Genus Tellina Linnaeus, 1758
Tellina aequistriata Say, 1824
Tellina alternate Say, 1822
Tellina angulosa Gmelin, 1791
Tellina candeana d'Orbigny, 1842
Tellina consobrina d'Orbigny, 1842
Tellina cristallina Spengler, 1798
Tellinafausta Pulteney, 1799
Tellina gouldii Hanley, 1846
Tellina iris Say, 1822
Tellina laevigata Linnaeus, 1758
Tellina lineata Turton, 1819
Tellina listeri RSding, 1798
Tellina magna Spengler, 1798
Tellina martinicensis d'Orbigny, 1842
Tellina mera Say, 1834
Tellina nitens C. B. Adams, 1845
Tellina paramera Boss, 1964
Tellina radiata Linnaeus, 1758
Tellina similis J. Sowerby, 1806
Tellina squamifera Deshayes, 1855
Tellina sybaritica Dall, 1881
Tellina tampaensis Conrad, 1866
Tellina tenella A. E.Verrill, 1874
Tellina texana Dall, 1900
Tellina versicolor DeKay, 1843

Family Donacidae Fleming, 1828


12 FMRI Technical Report TR-3


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Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


Genus Donax Linnaeus, 1758
Donaxfossor Say, 1822
Donax texasianus Philippi, 1847
Donax variabilis Say, 1822

Genus Iphigenia Schumacher, 1817
Iphigenia brasiliana (Lamarck, 1818)

Family Psammobiidae Fleming, 1828

Genus Asaphis Modeer, 1793
Asaphis deflorata (Linnaeus, 1758)

Genus Heterodonax Morch, 1853
Heterodonax bimaculatus (Linnaeus, 1758)

Genus Sanguinolaria Lamarck, 1799
Sanguinolaria sanguinolenta (Gmelin, 1791)

Family Semelidae Stoliczka, 1870

Genus Abra Lamarck, 1818
Abra aequalis (Say, 1822)
Abra lioica (Dall, 1881)

Genus Cumingia Sowerby, 1833
Cumingia coarctata Sowerby, 1833
Cumingia tellinoides (Conrad, 1831)

Genus Ervilia Turton, 1822
Ervilia concentrica (Holmes, 1860)
Ervilia nitens (Montagu, 1808)

Genus Semele Schumacher, 1817
Semele bellastriata (Conrad, 1837)
Semele proficua (Pulteney, 1799)
Semele purpurascens (Gmelin, 1791)

Genus Semelina Dall, 1900
Semelina nuculoides (Conrad, 1841)

Family Solecurtidae d'Orbigny, 1846

Genus Solecurtus de Blainville, 1824
Solecurtus cumingianus (Dunker, 1861)
Solecurtus sanctaemarthae (d'Orbigny, 1842)

Genus Tagelus J. E. Gray, 1847
Tagelus divisus (Spengler, 1794)
Tagelus plebeius (Lightfoot, 1786)

Family Dreissenidae J. E. Gray, 1840

Genus Mytilopsis Conrad, 1858


Mytilopsis leucophaeata (Conrad, 1831)
Mytilopsis sallei (R6cluz, 1849)

Family Trapezidae Lamy, 1920

Genus Coralliophaga de Blainville, 1824
Coralliophaga coralliophaga (Gmelin, 1791)

Family Corbiculidae J. E. Gray, 1847

Genus Polymesoda Rafinesque, 1820
Polymesoda caroliniana (Bosc, 1801)
Polymesoda maritima (d'Orbigny, 1842)

FamilyVeneridae Rafinesque, 1815

Genus Agriopoma Dall, 1902
Agriopoma texasianum (Dall, 1892)

Genus Anomalocardia Schumacher, 1817
Anomalocardia auberiana (d'Orbigny, 1842)

Genus Callista Poli, 1791
Callista eucymata (Dall, 1890)

Genus Chione Miihlfeld, 1811
Chione cancellata (Linnaeus, 1767)

Genus Cyclinella Dall, 1902
Cyclinella tenuis (R6cluz, 1852)

Genus Dosinia Scopoli, 1777
Dosinia discus (Reeve, 1850)
Dosinia elegans (Conrad, 1843)

Genus Gemma Deshayes, 1853
Gemma gemma (Totten, 1834)

Genus Globivenus Coen, 1934
Globivenus rigida (Dillwyn, 1817)
Globivenus rugatina (Heilprin, 1886)

Genus Gouldia C. B. Adams, 1847
Gouldia cerina (C. B. Adams, 1845)

Genus Lirophora Conrad, 1862
Lirophora latilirata (Conrad, 1841)

Genus Macrocallista Meek, 1876
Macrocallista maculata (Linnaeus, 1758)
Macrocallista nimbosa (Lightfoot, 1786)

Genus Mercenaria Schumacher, 1817
Mercenaria campechiensis (Gmelin, 1791)


FMRI Technical Report TR-3 13


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W. G. Lyons--Mollusca








Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


Mercenaria mercenaria (Linnaeus, 1758)

Genus Parastarte Conrad, 1862
Parastarte triquetra (Conrad, 1846)

Genus Periglypta Jukes-Brown, 1914
Periglypta listeri (J. E. Gray, 1838)

Genus Pitar Rimer, 1857
Pitar cordatus (Schwengel, 1951)
Pitarfiulminatus (Menke, 1828)
Pitar pilula Rehder, 1943
Pitar simpsoni (Dall, 1895)

Genus Puberella Fischer-Piette, 1977
Puberella intapurpurea (Conrad, 1850)
Puberella pubera (Bory Saint-Vincent, 1827)

Genus Timoclea T. Brown, 1827
Timoclea grus (Holmes, 1858)
Timoclea pygmaea (Lamarck, 1818)

Genus Tivela Link, 1807
Tivelafloridana Rehder, 1939

Genus Transennella Dall, 1884
Transennella conradina Dall, 1884
Transennella cubaniana (d'Orbigny, 1842)
Transennella stimpsoni Dall, 1902

Family Petricolidae Deschayes, 1831

Genus Choristodon Jonas, 1844
Choristodon robustum (G. B. Sowerby I, 1834)

Genus Cooperella Carpenter, 1864
Cooperella atlantica Rehder, 1943

Genus Petricola Lamarck, 1801
Petricola lapicida (Gmelin, 1791)

Genus Petricolaria Stoliczka, 1870
Petricolaria pholadiformis (Lamarck, 1818)

Order Myoida Goldfuss, 1820
Family Myidae Lamarck, 1809

Genus Paramya Conrad, 1861
Paramya subovata (Conrad, 1845)

Genus Sphenia Turton, 1822
Sphenia antillensis Dall & Simpson, 1901

Family Corbulidae Lamarck, 1818


Genus Caryocorbula Gardner, 1926
Caryocorbula barrattiana (C. B. Adams, 1852)

Genus Corbula Bruguiere, 1792
Corbula caribaea d'Orbigny, 1842
Corbula chittyana C. B. Adams, 1852
Corbula contract Say, 1822
Corbula dietziana C. B. Adams, 1852
Corbula swiftiana C. B. Adams, 1852

Genus Juliacorbula Olsson & Harbison, 1953
Juliacorbula cubaniana (d'Orbigny, 1842)

Genus Varicorbula Grant & Gale, 1931
Varicorbula operculata (Philippi, 1848)

Family Gastrochaenidae J. E. Gray, 1840

Genus Gastrochaena Spengler, 1783
Gastrochaena hians (Gmelin, 1791)
Gastrochaena ovata G. B. Sowerby I, 1834

Genus Spengleria Tryon, 1862
Spengleria rostrata (Spengler, 1783)

Family Hiatellidae J. E. Gray, 1824

Genus Hiatella Bosc, 1801
Hiatella arctica (Linnaeus, 1767)
Hiatella azaria (Dall, 1881)

Genus Panopea Menard, 1807
Panopea bitruncata (Conrad, 1872)

Family Pholadidae Lamarck, 1809

Genus Bamea Risso, 1826
Barnea truncata (Say, 1822)

Genus Cyrtopleura Tryon, 1862
Cyrtopleura costata (Linnaeus, 1758)

Genus Diplothyra Tryon, 1862
Diplothyra smithii Tryon, 1862

Genus Jouannetia DesMoulins, 1828
Jouannetia quillingi Turner, 1955

Genus Martesia Sowerby, 1824
Martesia cuneiformis (Say, 1822)
Martesiafragilis A. E.Verrill & Bush, 1890
Martesia striata (Linnaeus, 1758)

Genus Pholas Linnaeus, 1758


14 FMRI Technical Report TR-3


Mollusca--W. G. Lyons


Macro! nvertebrate Checklists


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Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


Pholas campechiensis Gmelin, 1791

Family Teredinidae Rafinesque, 1815

Genus Bankia J. E. Gray, 1842
Bankia carinata (J. E. Gray, 1827)
Bankia fimbriatula Moll & Roch, 1931
Bankia gouldi (Bartsch, 1908)

Genus Lyrodus Gould, 1870
Lyrodus bipartitus (Jeffreys, 1860)
Lyrodus floridanus (Bartsch, 1922)
Lyrodus mediolobatus (Edmondson, 1942)
Lyrodus pedicellatus (Quatrefages, 1849)

Genus Nototeredo Bartsch, 1923
Nototeredo knoxi (Bartsch, 1917)

Genus Spathoteredo Roch, 1937
Spathoteredo spatha (Jeffreys, 1860)

Genus Teredo Linnaeus, 1758
Teredo bartschi W. Clapp, 1923
Teredo fulleri W. Clapp, 1924
Teredofurcifera von Martens, 1894
Teredo navalis Linnaeus, 1758
Teredo portoricensis W. Clapp, 1924

Order Pholadomyoida Newell, 1965
Family Lyonsiidae P. Fischer, 1887

Genus Entodesma Philippi, 1845
Entodesma beanum (d'Orbigny, 1842)

Genus Lyonsia Turton, 1822
Lyonsiafloridana Conrad, 1848
Lyonsia hyalina (Conrad, 1831)

Family Pandoridae Rafinesque, 1815

Genus Pandora Bruguiere, 1797
Pandora arenosa Conrad, 1834
Pandora bushiana Dall, 1886
Pandora trilineata Say, 1822

Family Thraciidae Stoliczka, 1870

Genus Asthenothaerus Carpenter, 1864
Asthenothaerus balesi Rehder, 1943
Asthenothaerus hemphillii Dall, 1886

Genus Bushia Dall, 1886
Bushia elegans (Dall, 1886)


Genus Thracia Sowerby, 1823
Thracia morrisoni R. Petit, 1964

Family Periplomatidae Dall, 1895

Genus Periploma Schumacher, 1817
Periploma anguliferum (Philippi, 1847)
Periploma margaritaceum (Lamarck, 1801)
Periploma tenerum P. Fischer, 1882

Family Poromyidae Dall, 1886

Genus Poromya Forbes, 1844
Poromya rostrata Rehder, 1943

Family Verticordiidae Stoliczka, 1871

Genus Haliris Dall, 1886
Haliris fischeriana (Dall, 1881)

Genus Trigonulina d'Orbigny, 1842
Trigonulina ornata d'Orbigny, 1842

Family Cuspidariidae Dall, 1886

Genus Cardiomya A. Adams, 1864
Cardiomya costellata (Deshayes, 1830)
Cardiomya gemma A. E.Verrill & Bush, 1898
Cardiomya ornatissima (d'Orbigny, 1842)

Class Gastropoda Cuvier, 1797
Snails, Limpets, Conchs, Whelks, etc.

Order Patellogastropoda Lindberg, 1986
Family Lottiidae J. E. Gray, 1840

Genus Lottia J. E. Gray, 1833
Lottia leucopleura (Gmelin, 1791)

Genus Patelloida Quoy & Gaimard, 1834
Patelloida pustulata (Helbling, 1779)

Order Archaeogastropoda Thiele, 1925
Family Scissurellidae J. E. Gray, 1847

Genus Scissurella d'Orbigny, 1824
Scissurella cingulata 0. G. Costa, 1861

Family Fissurellidae Fleming, 1822

Genus Diodora J. E. Gray, 1821
Diodora arcuata (G. B. Sowerby II, 1862)
Diodora cayenensis (Lamarck, 1822)
Diodora dysoni (Reeve, 1850)


FMRI Technical Report TR-3 15


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W. G. Lyons--Mollusca








Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


Diodora jaumei Aguayo & Rehder, 1936
Diodora listeri (d'Orbigny, 1842)
Diodora meta (von Ihering, 1927)
Diodora minute (Lamarck, 1822)
Diodora sayi (Dall, 1889)
Diodora variegata G. B. Sowerby II, 1862
Diodora viridula (Lamarck, 1822)

Genus Emarginula Lamarck, 1801
Emarginula dentigera Heilprin, 1889
Emarginula phrixodes Dall, 1927
Emarginula pumila (A. Adams, 1852)

Genus Fissurella Bruguiere, 1789
Fissurella angusta (Gmelin, 1791)
Fissurella barbadensis (Gmelin, 1791)
Fissurella fascicularis Lamarck, 1822
Fissurella nodosa (Born, 1778)
Fissurella rosea (Gmelin, 1791)

Genus Hemitoma Swainson, 1840
Hemitoma emarginata (de Blainville, 1825)
Hemitoma octoradiata (Gmelin, 1791)

Genus Lucapina Sowerby, 1835
Lucapina aegis (Reeve, 1850)
Lucapina philippiana (Finlay, 1930)
Lucapina sowerbii (G. B. Sowerby II, 1835)
Lucapina suffusa (Reeve, 1850)

Genus Lucapinella Pilsbry, 1890
Lucapinella limatula (Reeve, 1850)

Genus Rimula Defrance, 1827
Rimula aequisculpta Dall, 1927
Rimulafrenulata (Dall, 1889)

Family Turbinidae Rafinesque, 1815

Genus Arene H. & A. Adams, 1854
Arene bairdii (Dall, 1889)
Arene briareus (Dall, 1881)
Arene cruentata (Miihlfeld, 1829)
Arene tricarinata (Stearns, 1872)
Arene variabilis (Dall, 1889)
Arene venustula Aguayo & Rehder, 1936

Genus Astralium Link, 1807
Astralium phoebium (Rbding, 1798)

Genus Cyclostrema Marryat, 1818
Cyclostrema cancellatum Marryat, 1818
Cyclostrema tortuganum (Dall, 1927)


Genus Didianema Woodring, 1928
Didianema pauli Pilsbry & McGinty, 1945

Genus Eulithidium Pilsbry, 1898
Eulithidium affine (C. B. Adams, 1850)
Eulithidium bellum (M. Smith, 1937)
Eulithidium pterocladicum (Robertson, 1958)
Eulithidium thalassicola (Robertson, 1958)

Genus Lithopoma J. E. Gray, 1850
Lithopoma americanum (Gmelin, 1791)
Lithopoma caelatum (Gmelin, 1791)
Lithopoma tectum (Lightfoot, 1786)
Lithopoma tuber (Linnaeus, 1767)

Genus Turbo Linnaeus, 1758
Turbo cailletii P. Fischer & Bernardi, 1856
Turbo canaliculatus Hermann, 1781
Turbo castanea Gmelin, 1791

Family Trochidae Rafinesque, 1815

Genus Cittarium Philippi, 1847
Cittarium pica (Linnaeus, 1758)

Genus Euchelus Philippi, 1847
Euchelus guttarosea Dall, 1889

Genus Pseudostomatella Thiele, 1924
Pseudostomatella erythrocoma (Dall, 1889)

Genus Solariella S.V. Wood, 1842
Solariella lacunella (Dall, 1881)

Genus Synaptocochlea Pilsbry, 1890
Synaptocochlea picta (d'Orbigny, 1842)

Genus Tegula Lesson, 1835
Tegula excavata (Lamarck, 1822)
Tegulafasciata (Born, 1778)
Tegula gruneri (Philippi, 1849)
Tegula hotessieriana (d'Orbigny, 1842)
Tegula lividomaculata (C. B. Adams, 1845)

Family Calliostomatidae Thiele, 1921

Genus Calliostoma Swainson, 1840
Calliostoma adelae Schwengel, 1951
Calliostoma barbouri Clench & Aguayo, 1946
Calliostoma euglyptum (A. Adams, 1855)
Calliostomajavanicum (Lamarck, 1822)
Calliostoma jujubinum (Gmelin, 1791)
Calliostoma pulchrum (C. B. Adams, 1850)
Calliostoma roseolum Dall, 1881


16 FMRI Technical Report TR-3


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Camp et al, 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


Calliostoma scalenum Quinn, 1992
Calliostoma tampaense (Conrad, 1846)
Calliostoma yucatecanum Dall, 1881

Family Skeneidae Thiele, 1929

Genus Haplocochlias Carpenter, 1864
Haplocochlias swifti Vanatta, 1913

Genus Parviturbo Pilsbry & McGinty, 1945
Parviturbo calidimaris Pilsbry & McGinty, 1945
Parviturbofrancesae Pilsbry & McGinty, 1945
Parviturbo rehderi Pilsbry & McGinty, 1945
Parviturbo weberi Pilsbry & McGinty, 1945

Order Neritopsina Cox & Knight, 1960
Family Neritidae Rafinesque, 1815

Genus Nerita Linnaeus, 1758
Neritafulgurans Gmelin, 1791
Nerita peloronta Linnaeus, 1758
Nerita tessellata Gmelin, 1791
Nerita versicolor Gmelin, 1791

Genus Neritina Lamarck, 1816
Neritina clenchi Russell, 1940
Neritina usnea (Roding, 1798)
Neritina virginea (Linnaeus, 1758)

Genus Puperita J. E. Gray, 1857
Puperita pupa (Linnaeus, 1767)

Genus Smaragdia Issel, 1869
Smaragdia viridis (Linnaeus, 1758)

Family Phenacolepadidae Pilsbry, 1900

Genus Phenacolepas Pilsbry, 1891
Phenacolepas hamillei (P. Fischer, 1857)

Order Neotaenioglossa Haller, 1882
Family Obtortionidae Thiele, 1925

Genus Finella A. Adams, 1869
Finella adamsi (Dall, 1889)
Finella dubia (d'Orbigny, 1842)

Family Litiopidae H. & A. Adams, 1854

Genus Alaba H. & A. Adams, 1853
Alaba incerta (d'Orbigny, 1842)

Genus Litiopa Rang, 1829
Litiopa melanostoma Rang, 1829


Family Cerithiidae Fleming, 1822

Genus Bittiolum Cossmann, 1906
Bittiolum varium (Pfeiffer, 1840)

Genus Cerithium Bruguiere, 1789
Cerithium atratum (Born, 1778)
Cerithium eburneum Bruguiere, 1792
Cerithium guinaicum Philippi, 1849
Cerithium lindae Petuch, 1987
Cerithium litteratum (Born, 1778)
Cerithium lutosum Menke, 1828
Cerithium muscarum Say, 1822

Family Batillariidae Thiele, 1929

Genus Batillaria Benson, 1842
Batillaria minima (Gmelin, 1791)

Family Potamididae H. & A. Adams, 1854

Genus Cerithidea Swainson, 1840
Cerithidea costata (E. M. da Costa, 1778)
Cerithidea scalariformis (Say, 1825)

Family Planaxidae J. E. Gray, 1850

Genus Fossarus Philippi, 1841
Fossarus bellus Dall, 1889
Fossarus orbignyi P. Fischer, 1864

Genus Hinea J. E. Gray, 1847
Hinea lineata (E. M. da Costa, 1778)

Genus Supplanaxis Thiele, 1929
Supplanaxis nucleus (Bruguiere, 1789)

Family Modulidae P. Fischer, 1884

Genus Modulus J. E. Gray, 1842
Modulus calusa Petuch, 1988
Modulus floridanus Conrad, 1869
Modulus modulus (Linnaeus, 1758)
Modulus pacei Petuch, 1987

Family Turritellidae Lovin, 1847

Genus Turritella Lamarck, 1799
Turritella acropora Dall, 1889
Turritella exoleta (Linnaeus, 1758)

Genus Vermicularia Lamarck, 1799
Vermiculariafargoi Olsson, 1951
Vermicularia knorrii (Deshayes, 1843)


FMRI Technical Report TR-3 17


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Mollusca-W. 0. Lyons Macroinvertebrate Checklists Camp et a!. 1998


Vermicularia spirata (Philippi, 1836)

Family Siliquariidae Anton, 1838

Genus Tenagodus Guettard, 1774
Tenagodus squamatus (de Blainville, 1827)

Family Littorinidae J. E. Gray, 1840

Genus Cenchritis von Martens, 1900
Cenchritis muricatus (Linnaeus, 1758)

Genus Littoraria J. E. Gray, 1834
Littoraria angulifera (Lamarck, 1822)
Littoraria irrorata (Say, 1822)
Littoraria nebulosa (Lamarck, 1822)
Littoraria tessellata (Philippi, 1847)

Genus Nodilittorina von Martens, 1897
Nodilittorina angustior (Morch, 1876)
Nodilittorina dilatata (d'Orbigny, 1842)
Nodilittorina meleagris (Potiez & Michaud, 1838)
Nodilittorina mespillum (Miihlfeld, 1824)
Nodilittorina riisei (Morch, 1876)
Nodilittorina ziczac (Gmelin, 1791)

Genus Tectarius Valenciennes, 1833
Tectarius antonii (Philippi, 1846)

Family Skeneopsidae Iredale, 1915

Genus Skeneopsis Iredale, 1915
Skeneopsis planorbis (Fabricius, 1780)

Family Barleeiidae Thiele, 1925

Genus Amphithalamus Carpenter, 1865
Amphithalamus vallei Aguayo & Jaume, 1947

Genus Barleeia Clark, 1853
Barleeia tincta Guppy, 1895

Family Rissoidae J. E. Gray, 1847

Genus Alvania Risso, 1826
Alvania auberiana (d'Orbigny, 1842)

Genus Folinia Crosse, 1868
Folinia bermudezi (Aguayo & Rehder, 1936)

Genus Microstelma A. Adams, 1863
Microstelma vestale (Rehder, 1943)

Genus Rissoina d'Orbigny, 1840


Rissoina cancellata Philippi, 1847
Rissoina decussata (Montagu, 1803)
Rissoina dyscrita Faber, 1990
Rissoina multicostata (C. B. Adams, 1850)
Rissoina princeps (C. B. Adams, 1850)
Rissoina sagraiana d'Orbigny, 1842
Rissoina striatocostata d'Orbigny, 1842
Rissoina striosa (C. B. Adams, 1850)

Genus Schwartziella Nevill, 1881
Schwartziella bouryi (Desjardin, 1949)
Schwartziella bryerea (Montagu, 1803)
Schwartziella catesbyana (d'Orbigny, 1842)
Schwartziella fischeri Desjardin, 1949

Genus Stosicia Brusina, 1870
Stosicia aberrans (C. B. Adams, 1850)

Genus Zebina H. & A. Adams, 1854
Zebina browniana (d'Orbigny, 1842)
Zebina laevigata (C. B. Adams, 1850)

Family Hydrobiidae Troschel, 1857

Genus Floridiscrobs Pilsbry & McGinty, 1949
Floridiscrobs dysbatus (Pilsbry & McGinty, 1949)

Genus Heleobops F. G. Thompson, 1968
Heleobops docimus F. G. Thompson, 1968

Genus Littoridinops Pilsbry, 1952
Littoridinops monroensis (Frauenfeld, 1863)
Littoridinops palustris F. G. Thompson, 1968
Littoridinops tenuipes (Couper, 1844)

Genus Onobops F. G.Thompson, 1968
Onobops crassus F. G. Thompson, 1968
Onobops jacksoni (Bartsch, 1953)

Genus Probythinella Thiele, 1928
Probythinella protera Pilsbry, 1953

Genus Pyrgophorus Ancey, 1888
Pyrgophorus platyrachis F. G.Thompson, 1968

Genus Spurwinkia Davis, Mazurkiewicz & Mandrac-
chia, 1982
Spurwinkia salsa (Pilsbry, 1905)

Genus Texadina Abbott & Ladd, 1951
Texadina sphinctostoma (Abbott & Ladd, 1951)

Family Truncatellidae J. E. Gray, 1840


18 FMRI Technical Report TR-3


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Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


Genus Truncatella Risso, 1826
Truncatella caribaeensis Reeve, 1842
Truncatella clathrus Lowe, 1932
Truncatella pulchella Pfeiffer, 1839

Family Assimineidae H. & A. Adams, 1856

Genus Assiminea Fleming, 1828
Assiminea succinea (Pfeiffer, 1840)

Family Elachisinidae Ponder, 1985

Genus Elachisina Dall, 1918
Elachisinafloridana (Rehder, 1943)

Family Vitrinellidae Bush, 1897

Genus Anticlimax Pilsbry & McGinty, 1946
Anticlimax athleenae (Pilsbry & McGinty, 1946)
Anticlimax pilsbryi (McGinty, 1945)

Genus Circulus Jeffreys, 1865
Circulus liratus (Verrill, 1882)
Circulus semisculptus (Olsson & McGinty, 1958)
Circulus texanus (Moore, 1965)

Genus Cochliolepis Stimpson, 1858
Cochliolepis holmesii (Dall, 1889)
Cochliolepis parasitica Stimpson, 1858
Cochliolepis striata Dall, 1889

Genus Cyclostremiscus Pilsbry & Olsson, 1945
Cyclostremiscus beauii (P. Fischer, 1857)
Cyclostremiscus cubanus (Pilsbry & Aguayo, 1933)
Cyclostremiscus jeannae Pilsbry & McGinty, 1946
Cyclostremiscus pentagonus (Gabb, 1873)
Cyclostremiscus suppressus (Dall, 1889)

Genus Episcynia Morch, 1875
Episcynia inornata (d'Orbigny, 1842)

Genus Pachystremiscus Olsson & McGinty, 1958
Pachystremiscus ornatus (Olsson & McGinty, 1958)
Pachystremiscus pulchellus (Olsson & McGinty,
1958)

Genus Parviturboides Pilsbry & McGinty, 1950
Parviturboides interruptus (C. B. Adams, 1850)

Genus Pleuromalaxis Pilsbry & McGinty, 1945
Pleuromalaxis balesi (Pilsbry & McGinty, 1945)
Pleuromalaxis pauli (Olsson & McGinty, 1958)

Genus Solariorbis Conrad, 1865


Solariorbis blakei (Rehder, 1944)
Solariorbis infracarinatus (Gabb, 1881)
Solariorbis mooreanus (Vanatta, 1904)
Solariorbis semipunctus D. R. Moore, 1965
Solariorbis terminalis (Pilsbry & McGinty, 1946)

Genus Teinostoma H. & A. Adams, 1854
Teinostoma biscaynense Pilsbry & McGinty, 1945
Teinostoma carinicallus Pilsbry & McGinty, 1946
Teinostoma clavium Pilsbry & McGinty, 1945
Teinostoma goniogyrus Pilsbry & McGinty, 1945
Teinostoma incertum Pilsbry & McGinty, 1945
Teinostoma lerema Pilsbry & McGinty, 1945
Teinostoma lituspalmarum Pilsbry & McGinty,
1945
Teinostoma megastoma (C. B. Adams, 1850)
Teinostoma parvicallum Pilsbry & McGinty, 1945

Genus Vitrinella C. B. Adams, 1850
Vitrinella filifera Pilsbry & McGinty, 1946
Vitrinellafloridana Pilsbry & McGinty, 1946
Vitrinella helicoidea C. B. Adams, 1850
Vitrinella hemphilli Vanatta, 1913

Family Caecidae J. E. Gray, 1850

Genus Caecum Fleming, 1813
Caecum antillarum Carpenter, 1858
Caecum bipartitum de Folin, 1870
Caecum breve de Folin, 1867
Caecum carolinianum Dall, 1892
Caecum clava de Folin, 1867
Caecum cooper S. Smith, 1860
Caecum cycloferum de Folin, 1867
Caecum floridanum Stimpson, 1851
Caecum gurgulio Carpenter, 1858
Caecum heladum Olsson & Harbison, 1953
Caecum imbricatum Carpenter, 1858
Caecum johnsoni Winkley, 1908
Caecum plicatum Carpenter, 1858
Caecum pulchellum Stimpson, 1851
Caecum regular Carpenter, 1858
Caecum strigosum de Folin, 1868
Caecum subvolutum de Folin, 1874
Caecum textile de Folin, 1867
Caecum vestitum de Folin, 1868

Genus Meioceras Carpenter, 1858
Meioceras corucopiae (Carpenter, 1858)
Meioceras cubitatum (de Folin, 1868)
Meioceras nitidum (Stimpson, 1851)

Family Strombidae Rafinesque, 1815


FMRI Technical Report TR-3 19


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Mollusca-W, G, Lyons Macroinvertebrate Checklists Camp et al. 1998


Genus Strombus Linnaeus, 1758
Strombus alatus Gmelin, 1791
Strombus costatus Gmelin, 1791
Strombus gallus Linnaeus, 1758
Strombus gigas Linnaeus, 1758
Strombus pugilis Linnaeus, 1758
Strombus raninus Gmelin, 1791

Family Hipponicidae Troschel, 1861

Genus Hipponix Defrance, 1819
Hipponix antiquatus (Linnaeus, 1767)
Hipponix subrufus (Lamarck, 1819)

Family Vanikoridae J. E. Gray, 1840

Genus Megalomphalus Brusina, 1871
Megalomphalus adamsii (P. Fischer, 1857)
Megalomphalusfloridanus (D. R. Moore, 1965)
Megalomphalus palmalitoris (Pilsbry & McGinty,
1950)

Genus Vanikoro Quoy & Gaimard, 1832
Vanikoro oxychone Morch, 1877

Family Calyptraeidae Lamarck, 1809

Genus Calyptraea Lamarck, 1799
Calyptraea centralis (Conrad, 1841)

Genus Cheilea Modeer, 1793
Cheilea equestris (Linnaeus, 1758)

Genus Crepidula Lamarck, 1799
Crepidula aculeata (Gmelin, 1791)
Crepidula convexa Say, 1822
Crepidulafornicata (Linnaeus, 1758)
Crepidula maculosa Conrad, 1846
Crepidula plana Say, 1822

Genus Crucibulum Schumacher, 1817
Crucibulum auricula (Gmelin, 1791)
Crucibulum striatum (Say, 1826)

Family Capulidae Fleming, 1822

Genus Capulus Montfort, 1810
Capulus incurvus (Gmelin, 1791)

Family Xenophoridae Troschel, 1852

Genus Xenophora G. Fischer, 1807
Xenophora conchyliophora (Born, 1780)


Family Vermetidae Rafinesque, 1815

Genus Dendropoma Morch, 1861
Dendropoma corrodens (d'Orbigny, 1842)
Dendropoma irregulare (d'Orbigny, 1842)

Genus Petaloconchus H. C. Lea, 1843
Petaloconchus erectus (Dall, 1888)
Petaloconchus varians (d'Orbigny, 1841)

Genus Serpulorbis Sassi, 1827
Serpulorbis decussatus (Gmelin, 1791)

Family Cypraeidae Rafinesque, 1815

Genus Erosaria Troschel, 1863
Erosaria acicularis (Gmelin, 1791)

Genus Macrocypraea Schilder, 1930
Macrocypraea cerous (Linnaeus, 1771)
Macrocypraea zebra (Linnaeus, 1758)

Genus Propustularia Schilder, 1927
Propustularia surinamensis (G. Perry, 1811)

Genus Talparia Troschel, 1863
Talparia cinerea (Gmelin, 1791)

Family Ovulidae Fleming, 1822

Genus Cymbovula Cate, 1974
Cymbovula acicularis (Lamarck, 1810)

Genus Cyphoma Riding, 1798
Cyphoma alleneae Cate, 1973
Cyphoma gibbosum (Linnaeus, 1758)
Cyphoma mcgintyi Pilsbry, 1939
Cyphoma rhomba Cate, 1978
Cyphoma sedlaki Cate, 1976
Cyphoma signatum Pilsbry & McGinty, 1939

Genus Neosimnia P. Fischer, 1884
Neosimnia advena (Cate, 1978)

Genus Pseudocyphoma Cate, 1973
Pseudocyphoma gibbulum Cate, 1978

Genus Simnialena Cate, 1973
Simnialena uniplicata (G. B. Sowerby II, 1848)

Family Triviidae Troschel, 1863

Genus Hespererato Schilder, 1932
Hespererato maugeriae (J. E. Gray, 1832)


20 FMRI Technical Report TR-3


Mollusca-WW G. Lyons


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Camp et al. 1998








Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


Genus Niveria Jousseaume, 1884
Niveria atomaria pygmaea (Schilder, 1931)
Niveria candidula (Gaskoin, 1836)
Niveria leucosphaera Schilder, 1931
Niveria pediculus (Linnaeus, 1758)
Niveria quadripunctata (J. E. Gray, 1827)
Niveria suffusa (J. E. Gray, 1827)

Genus Proterato Schilder, 1927
Proterato gemma pygmaea Schilder, 1933

Genus Pusula Jousseaume, 1884
Pusula maltbiana (Schwengel & McGinty, 1942)

Family Lamellariidae d'Orbigny, 1841

Genus Lamellaria Montagu, 1815
Lamellaria koto Schwengel, 1944
Lamellaria leucosphaera Schwengel, 1942
Lamellaria perspicua (Linnaeus, 1758)

Genus Marsenina J. E. Gray, 1850
Marsenina globosa L. M. Perry, 1939

Family Naticidae Forbes, 1838

Genus Haliotinella Souverbie in Souverbie & Mon-
trouzier, 1875
Haliotinella patinaria (Guppy, 1876)

Genus Natica Scopoli, 1777
Natica livida Pfeiffer, 1840
Natica marochiensis (Gmelin, 1791)
Natica tedbayeri Rehder, 1986

Genus Naticarius Dumbril, 1806
Naticarius canrena (Linnaeus, 1758)
Naticarius verae Rehder, 1947

Genus Neverita Risso, 1826
Neverita duplicate (Say, 1822)

Genus Polinices Montfort, 1810
Polinices hepaticus (RSding, 1798)
Polinices lacteus (Guilding, 1834)
Polinices uberinus (d'Orbigny, 1842)

Genus Sigatica Meyer & Aldrich, 1886
Sigatica carolinensis (Dall, 1889)
Sigatica semisulcata (J. E. Gray, 1839)

Genus Sinum Roding, 1798
Sinum maculatum (Say, 1831)
Sinum perspectivum (Say, 1831)


Genus Stigmaulax Morch, 1852
Stigmaulax sulcatus (Born, 1778)

Genus Tectonatica Sacco, 1890
Tectonatica pusilla (Say, 1822)

Family Cassidae Latreille, 1825

Genus Casmaria H. & A. Adams, 1853
Casmaria ponderosa atlantica Clench, 1944

Genus Cassis Scopoli, 1777
Cassisflammea (Linnaeus, 1758)
Cassis madagascariensis Lamarck, 1822
Cassis tuberosa (Linnaeus, 1758)

Genus Cypraecassis Stutchbury, 1837
Cypraecassis testiculus (Linnaeus, 1758)

Genus Phalium Link, 1807
Phalium granulatum (Born, 1778)

Family Tonnidae Suter, 1913

Genus Tonna Briinnich, 1772
Tonna galea (Linnaeus, 1758)
Tonna pennata (Morch, 1852)

Family Ficidae Meek, 1864

Genus Ficus Rbding, 1798
Ficus communis R6ding, 1798

Family Ranellidae J. E. Gray, 1854

Genus Charonia Gistel, 1848
Charonia tritonis variegata (Lamarck, 1816)

Genus Cymatium Roding, 1798
Cymatium aquatile (Reeve, 1844)
Cymatium comptum (A. Adams, 1855)
Cymatium corrugatum krebsii (Morch, 1877)
Cymatium cynocephalum (Lamarck, 1816)
Cymatiumfemorale (Linnaeus, 1758)
Cymatium labiosum (W. Wood, 1828)
Cymatium martinianum (d'Orbigny, 1846)
Cymatium muricinum (RSding, 1798)
Cymatium nicobaricum (R6ding, 1798)
Cymatium occidentale (Morch, 1877)
Cymatium parthenopeum (von Salis, 1793)
Cymatium tenuiliratum (Lischke, 1873)
Cymatium vespaceum (Lamarck, 1822)

Genus Linatella J. E. Gray, 1857


FMRI Technical Report TR-3 21


Camp et a[. 1998


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W. G. Lyons--Mollusca








Mollusca-W. G. Lyons Macroinvertebrote Checklists Camp et ci. 1998


Linatella caudata (Gmelin, 1791)

Family Personidae J. E. Gray, 1854

Genus Distorsio R6ding, 1798
Distorsio clathrata (Lamarck, 1816)
Distorsio constricta mcgintyi Emerson & Puffer,
1953

Family Bursidae Thiele, 1925

Genus Bufonaria Schumacher, 1817
Bufonaria bufo (Bruguiere, 1792)

Genus Bursa Roding, 1798
Bursa corrugata ponderosa (Reeve, 1844)
Bursa granularis cubaniana (d'Orbigny, 1842)
Bursa grayana Dunker, 1862
Bursa rhodostoma thomae (d'Orbigny, 1842)

Family Cerithiopsidae H. & A. Adams, 1854

Genus Cerithiopsis Forbes & Hanley, 1849
Cerithiopsis albovittata (C. B. Adams, 1850)
Cerithiopsis flava (C. B. Adams, 1850)
Cerithiopsisfusiformis (C. B. Adams, 1850)
Cerithiopsis greenii (C. B. Adams, 1839)
Cerithiopsis io Dall & Bartsch, 1911
Cerithiopsis iota (C. B. Adams, 1845)
Cerithiopsis lata (C. B. Adams, 1850)
Cerithiopsis vanhyningi Bartsch, 1918

Genus Horologica Laseron, 1956
Horologica pupa (Dall & Simpson, 1901)

Genus Retilaskeya Marshall, 1978
Retilaskeya bicolor (C. B. Adams, 1845)
Retilaskeya emersonii (C. B. Adams, 1839)

Genus Seila A. Adams, 1861
Seila adamsii (H. C. Lea, 1845)

Family Triphoridae J. E. Gray, 1847

Genus Cosmotriphora Olsson & Harbison, 1953
Cosmotriphora decorate (C. B. Adams, 1850)
Cosmotriphora melanura (C. B. Adams, 1850)
Cosmotriphora olivacea (Dall, 1889)

Genus Iniforis Jousseaume, 1884
Iniforis turristhomae (Holten, 1802)

Genus Marshallora Bouchet, 1985
Marshallora nigrocincta (C. B. Adams, 1839)


Genus Metaxia Monterosato, 1884
Metaxia rugulosa (C. B. Adams, 1850)
Metaxia taeniolata (Dall, 1889)

Genus Similiphora Bouchet, 1985
Similiphora intermedia (C. B. Adams, 1850)

Genus Triphora de Blainville, 1828
Triphora calva Faber & Moolenbeek, 1991
Triphora lilacina (Dall, 1889)
Triphora ornata (Deshayes, 1832)

Family Epitoniidae S. S. Berry, 1910

Genus Alexania Strand, 1928
Alexaniafloridana (Pilsbry, 1945)

Genus Amaea H. & A. Adams, 1854
Amaea retifera (Dall, 1889)

Genus Cirsotrema Morch, 1852
Cirsotrema dalli Rehder, 1945

Genus Depressiscala de Boury, 1909
Depressiscala nautlae (Morch, 1874)

Genus Epitonium R6ding, 1798
Epitonium albidum (d'Orbigny, 1842)
Epitonium angulatum (Say, 1830)
Epitonium apiculatum (Dall, 1889)
Epitonium blainei Clench & Turner, 1953
Epitonium candeanum (d'Orbigny, 1842)
Epitonium champion Clench & Turner, 1952
Epitonium denticulatum (G. B. Sowerby II, 1844)
Epitonium echinaticosta (d'Orbigny, 1842)
Epitoniumfoliaceicosta (d'Orbigny, 1842)
Epitonium humphreysii (Kiener, 1838)
Epitonium krebsii (Morch, 1874)
Epitonium lamellosum (Lamarck, 1822)
Epitonium matthewsae Clench & Turner, 1952
Epitonium multistriatum (Say, 1826)
Epitonium novangliae (Couthouy, 1838)
Epitonium occidentale (Nyst, 1871)
Epitonium phymanthi Robertson, 1994
Epitonium rupicola (Kurtz, 1860)
Epitonium striatissimum (Monterosato, 1878)
Epitonium tollini Bartsch, 1938
Epitonium unifasciatum (G. B. Sowerby II, 1844)

Genus Opalia H. & A. Adams, 1853
Opalia burryi Clench & Turner, 1950
Opalia crenata (Linnaeus, 1758)
Opalia hotessieriana (d'Orbigny, 1842)
Opalia pumilio (Morch, 1874)


22 FMRI Technical Report TR-3


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Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


Genus Opaliopsis Thiele, 1928
Opaliopsis concava (Dall, 1889)

Family Janthinidae Lamarck, 1812

Genus Janthina Rbding, 1798
Janthina exigua Lamarck, 1816
Janthina globosa Swainson, 1822
Janthina janthina (Linnaeus, 1758)
Janthina pallida W. Thompson, 1840

Genus Recluzia Petit de la Saussaye, 1853
Recluzia rollandiana Petit de la Saussaye, 1853

Family Aclididae G. O. Sars, 1878

Genus Graphis Jeffreys, 1867
Graphis underwoodae Bartsch, 1947

Family Eulimidae Troschel, 1853

Genus Ersilia Monterosato, 1872
Ersilia stancyki Waren, 1980

Genus Eulima Risso, 1826
Eulima auricincta (Abbott, 1958)
Eulima bifasciata d'Orbigny, 1842
Eulimafulvocincta C. B. Adams, 1850

Genus Eulimostraca Bartsch, 1917
Eulimostraca subcarinata (d'Orbigny, 1842)

Genus Melanella Bowdich, 1822
Melanella conoidea (Kurtz & Stimpson, 1851)
Melanella eulimoides (C. B. Adams, 1850)
Melanella gracilis (C. B. Adams, 1850)
Melanella jamaicensis (C. B. Adams, 1845)

Genus Microeulima War6n, 1992
Microeulima hemphillii (Dall, 1884)

Genus Nanobalcis Waren in War6n & Mifsud, 1990
Nanobalcis worsfoldi Waren, 1990

Genus Niso Risso, 1826
Niso aeglees Bush, 1885
Niso hendersoni Bartsch, 1953

Genus Oceanida de Folin, 1870
Oceanida graduate de Folin, 1870
Oceanida inglei Lyons, 1978

Genus Pelseneeria Kohler &Vaney, 1908
Pelseneeria stimpsonii (A. E.Verrill, 1872)


Genus Sabinella Monterosato, 1890
Sabinella troglodytes (Thiele, 1925)

Genus Umbilibalcis Bouchet & Waren, 1986
Umbilibalcis lata (Dall, 1889)

Genus Vitreolina Monterosato, 1884
Vitreolina arcuata (C. B. Adams, 1850)
Vitreolina bermudezi (Pilsbry & Aguayo, 1933)

Order Neogastropoda Thiele, 1929
Family Muricidae Rafinesque, 1815

Genus Acanthotrophon Hertlein & Strong, 1951
Acanthotrophon striatoides Vokes, 1980

Genus Aspella Morch, 1877
Aspella senex Dall, 1903

Genus Calotrophon Hertlein & Strong, 1951
Calotrophon ostrearum (Conrad, 1846)

Genus Chicoreus Montfort, 1810
Chicoreus dilectus (A. Adams, 1855)
Chicoreus pomum (Gmelin, 1791)

Genus Dermomurex Monterosato, 1890
Dermomurex elizabethae (McGinty, 1940)
Dermomurex pacei Petuch, 1988
Dermomurex pauperculus (C. B. Adams, 1850)

Genus Eupleura H. & A. Adams, 1853
Eupleura caudata (Say, 1822)
Eupleura sulcidentata Dall, 1890

Genus Favartia Jousseaume, 1880
Favartia alveata (Kiener, 1842)
Favartia cellulosa (Conrad, 1846)
Favartia minirosea (Abbott, 1954)
Favartia pacei Petuch, 1988
Favartia richardbinghami (Petuch, 1987)

Genus Haustellum Schumacher, 1817
Haustellum bellegladeense (Vokes, 1963)
Haustellum cabritii (Bernardi, 1859)
Haustellum rubidum (F. C. Baker, 1897)

Genus Hexaplex G. Perry, 1810
Hexaplex filvescens (G. B. Sowerby II, 1834)

Genus Murexiella Clench & Farfante, 1945
Murexiella glypta (M. Smith, 1938)
Murexiella kalafuti Petuch, 1987
Murexiella levicula (Dall, 1889)
Murexiella macgintyi (M. Smith, 1938)


FMRI Technical Report TR-3 23


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Macroinvertebrate Checklists


W. G, Lyons--Mollusca








Mollusca-W, G. Lyons Macroinvertebrate Checklists Camp et ai. 1998


Genus Muricopsis Bucquoy, Dautzenberg & Dollfus,
1882
Muricopsis caribbaea (Bartsch & Rehder, 1939)
Muricopsis oxytata (M. Smith, 1938)

Genus Plicopurpura Cossmann, 1903
Plicopurpura patula (Linnaeus, 1758)

Genus Pterotyphis Jousseaume, 1880
Pterotyphis triangularis (A. Adams, 1855)

Genus Stramonita Schumacher, 1817
Stramonita haemastoma canaliculata (J. E. Gray,
1839)
Stramonita haemastoma floridana (Conrad, 1837)
Stramonita rustica (Lamarck, 1822)

Genus Thais Roding, 1798
Thais deltoidea (Lamarck, 1822)

Genus Trachypollia Woodring, 1928
Trachypollia nodulosa (C. B. Adams, 1845)
Trachypollia turricula (von Maltzan, 1884)

Genus Urosalpinx Stimpson, 1865
Urosalpinx cinerea (Say, 1822)
Urosalpinx perrugata (Conrad, 1846)
Urosalpinx tampaensis (Conrad, 1846)

Family Coralliophilidae Chenu, 1859

Genus Babelomurex Coen, 1922
Babelomurex mansfieldi (McGinty, 1940)

Genus Coralliophila H. & A. Adams, 1853
Coralliophila aberrans (C. B. Adams, 1850)
Coralliophila caribaea Abbott, 1958
Coralliophila galea (Reeve, 1846)
Coralliophila pacei Petuch, 1987

Genus Quoyula Iredale, 1912
Quoyula kalafuti Petuch, 1987

Family Turbinellidae Swainson, 1840

Genus Vasum Riding, 1798
Vasum muricatum (Born, 1778)

Family Buccinidae Rafinesque, 1815

Genus Antillophos Woodring, 1928
Antillophos candeanus (d'Orbigny, 1842)

Genus Bailya M. Smith, 1944


Bailya intricate (Dall, 1884)
Bailya parva (C. B. Adams, 1850)
Bailya weberi (Watters, 1983)

Genus Cantharus Riding, 1798
Cantharus cancellarius (Conrad, 1846)
Cantharus multangulus (Philippi, 1848)

Genus Colubraria Schumacher, 1817
Colubraria testacea (Morch, 1852)

Genus Engina J. E. Gray, 1839
Engina corinnae Crovo, 1971
Engina turbinella (Kiener, 1835)

Genus Parviphos Sarasua, 1984
Parviphos adelus (Schwengel, 1942)

Genus Pisania Bivona, 1832
Pisania pusio (Linnaeus, 1758)

Genus Pollia J. E. Gray in Sowerby, 1834
Pollia auritula (Link, 1807)
Pollia karinae (Nowell-Usticke, 1959)
Pollia tincta Conrad, 1846

Family Melongenidae Gill, 1867

Genus Busycon Riding, 1798
Busycon carica (Gmelin, 1791)
Busycon sinistrum Hollister, 1958

Genus Busycotypus Wenz, 1943
Busycotypus canaliculatus (Linnaeus, 1758)
Busycotypus spiratus (Lamarck, 1816)

Genus Melongena Schumacher, 1817
Melongena bicolor (Say, 1826)
Melongena corona (Gmelin, 1791)
Melongena sprucecreekensis Tucker, 1994

Family Nassariidae Iredale, 1916

Genus Ilyanassa Stimpson, 1865
Ilyanassa obsoleta (Say, 1822)
Ilyanassa trivittata (Say, 1822)

Genus Nassarius Dumbril, 1806
Nassarius acutus (Say, 1822)
Nassarius albus (Say, 1826)
Nassarius antillarum (d'Orbigny, 1842)
Nassarius consensus (Ravenel, 1861)
Nassarius hotessieri (d'Orbigny, 1845)
Nassarius polygonatus (Lamarck, 1822)


24 FMRI Technical Report TR-3


Mollusca--W, G. Lyons


Macroinvertebrate Checklists


Camp et a]. 1998








Camp et al. 1998 Macrainvertebrate Checklists W. G. Lyons-Mollusca


Nassarius vibex (Say, 1822)

Family Fasciolariidae J. E. Gray, 1853

Genus Fasciolaria Lamarck, 1799
Fasciolaria hunteria (G. Perry, 1811)
Fasciolaria tulipa (Linnaeus, 1758)

Genus Fusinus Rafinesque, 1815
Fusinus eucosmius (Dall, 1889)
Fusinus helenae Bartsch, 1939

Genus Heilprinia Grabau, 1904
Heilprinia timessa (Dall, 1889)

Genus Latirus Montfort, 1810
Latirus angulatus (R6ding, 1798)
Latirus carinifer Lamarck, 1822
Latirus infundibulum (Gmelin, 1791)
Latirus nematus Woodring, 1928

Genus Leucozonia J. E. Gray, 1847
Leucozonia nassa (Gmelin, 1791)
Leucozonia ocellata (Gmelin, 1791)

Genus Pleuroploca P. Fischer, 1884
Pleuroploca gigantea (Kiener, 1840)

Genus Teralatirus Coomans, 1965
Teralatirus cayohuesonicus (G. B. Sowerby III,
1878)

Family Columbellidae Swainson, 1840

Genus Aesopus Gould, 1860
Aesopus stearnsii (Tryon, 1883)

Genus Astyris H. & A. Adams, 1853
Astyris lunata (Say, 1826)
Astyris multilineata (Dall, 1889)
Astyris raveneli (Dall, 1889)

Genus Columbella Lamarck, 1799
Columbella mercatoria (Linnaeus, 1758)
Columbella rusticoides Heilprin, 1886

Genus Columbellopsis Bucquoy, Dautzenberg & Doll-
fus, 1882
Columbellopsis nycteis (Duclos, 1846)

Genus Costoanachis Sacco, 1890
Costoanachis avara (Say, 1822)
Costoanachis catenata (G. B. Sowerby I, 1844)
Costoanachisfloridana (Rehder, 1939)


Costoanachis lafresnayi (P. Fischer & Bernardi,
1857)
Costoanachis semiplicata (Stearns, 1873)
Costoanachis similis (Ravenel, 1861)
Costoanachis sparsa (Reeve, 1859)

Genus Mazatlania Dall, 1913
Mazatlania cosentini (Philippi, 1836)

Genus Mitrella Risso, 1826
Mitrella dichroa (G. B. Sowerby I, 1844)
Mitrella ocellata (Gmelin, 1791)

Genus Nassarina Dall, 1889
Nassarina bushiae (Dall, 1889)
Nassarina glypta (Bush, 1885)

Genus Nitidella Swainson, 1840
Nitidella nitida (Lamarck, 1822)

Genus Parvanachis Radwin, 1968
Parvanachis obesa (C. B. Adams, 1845)
Parvanachis ostreicola (G. B. Sowerby III, 1882)

Genus Rhombinella Radwin, 1968
Rhombinella laevigata (Linnaeus, 1758)

Genus Steironepion Pilsbry & Lowe, 1932
Steironepion minor (C. B. Adams, 1845)
Steironepion moniliferum (G. B. Sowerby I, 1844)

Genus Suturoglypta Radwin, 1968
Suturoglypta iontha (Ravenel, 1861)
Suturoglypta pretrii (Duclos, 1846)

Genus Zafrona Iredale, 1916
Zafrona dicomata (Dall, 1889)
Zafrona idalina (Duclos, 1840)
Zafrona pulchella (de Blainville, 1829)
Zafrona taylorae Petuch, 1987

FamilyVolutidae Rafinesque, 1815

Genus Enaeta H. & A. Adams, 1853
Enaeta cylleniformis (G. B. Sowerby 1, 1844)

Genus Scaphella Swainson, 1832
Scaphella junonia (Lamarck, 1804)

Family Olividae Latreille, 1825

Genus Jaspidella Olsson, 1956
Jaspidella blanesi (Ford, 1898)
Jaspidella jaspidea (Gmelin, 1791)


FMRI Technical Report TR-3 25


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W. G. Lyons--Mollusca








Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


Genus Oliva Bruguiere, 1789
Oliva bollingi Clench, 1937
Oliva sayana Ravenel, 1834

Genus Olivella Swainson, 1831
Olivella bullula (Reeve, 1850)
Olivellafloralia (Duclos, 1853)
Olivella inusta G. B. Sowerby III, 1915
Olivella lactea (Marrat, 1871)
Olivella macgintyi Olsson, 1956
Olivella minute (Link, 1807)
Olivella monilifera (Reeve, 1850)
Olivella mutica (Say, 1822)
Olivella nivea (Gmelin, 1791)
Olivella perplexa Olsson, 1956
Olivella pusilla (Marrat, 1871)

Family Harpidae Bronn, 1849

Genus Morum Roding, 1798
Morum oniscus (Linnaeus, 1767)

Family Cystiscidae Stimpson, 1865

Genus Gibberula Swainson, 1840
Gibberula lavalleeana (d'Orbigny, 1842)

Genus Granulina Jousseaume, 1888
Granulina hadria (Dall, 1889)

Genus Persicula Schumacher, 1817
Persicula catenata (Montagu, 1803)
Persicula pulcherrima (Gaskoin, 1849)

Genus Pugnus Hedley, 1896
Pugnus lachrimula (Gould, 1862)
Pugnus serrei (Bavay, 1911)

Family Marginellidae Fleming, 1828

Genus Dentimargo Cossmann, 1899
Dentimargo aureocinctus (Stearns, 1872)
Dentimargo eburneolus (Conrad, 1834)
Dentimargo idiochila (Schwengel, 1943)
Dentimargo reductus (Bavay, 1922)

Genus Hyalina Schumacher, 1817
Hyalina pallida (Linnaeus, 1758)

Genus Prunum Herrmannsen, 1852
Prunum amabile (Redfield, 1852)
Prunum apicinum (Menke, 1828)
Prunum avenaceum (Deshayes, 1844)
Prunum carneum (Storer, 1837)


Prunum evelynae (Bayer, 1943)
Prunum guttatum (Dillwyn, 1817)
Prunum roscidum (Redfield, 1860)
Prunum succinea (Conrad, 1846)
Prunum virginianum hartleyanum (Schwengel,
1941)
Prunum virginianum virginianum Conrad, 1868

Genus Volvarina Hinds, 1844
Volvarina albolineata (d'Orbigny, 1842)
Volvarina avena (Kiener, 1834)
Volvarina lactea (Kiener, 1841)
Volvarina subtriplicata (d'Orbigny, 1842)

Family Mitridae Swainson, 1831

Genus Mitra Lamarck, 1798
Mitra barbadensis (Gmelin, 1791)
Mitraflorida Gould, 1856
Mitra nodulosa (Gmelin, 1791)
Mitra pallida Nowell-Usticke, 1959
Mitra semiferruginea Reeve, 1845
Mitra straminea A. Adams, 1853

Family Costellariidae MacDonald, 1860

Genus Thala H. & A. Adams, 1853
Thalafloridana (Dall, 1884)

Genus Vexillum R5ding, 1798
Vexillum dermestinum (Lamarck, 1811)
Vexillun epiphaneum (Rehder, 1943)
Vexillum exiguum (C. B. Adams, 1845)
Vexillum gemmatum (G. B. Sowerby II, 1874)
Vexillum hendersoni (Dall, 1927)
Vexillum histrio (Reeve, 1844)
Vexillum puella (Reeve, 1845)
Vexillum pulchellum (Reeve, 1844)
Vexillum sykesi (Melvill, 1925)
Vexillum wandoense (Holmes, 1860)
Vexillum zythochroa (Melvill, 1888)

Family Cancellariidae J. E. Gray, 1853

Genus Agatrix Petit, 1967
Agatrix agassizii (Dall, 1889)

Genus Axelella Petit, 1988
Axelella smithii (Dall, 1888)

Genus Cancellaria Lamarck, 1799
Cancellaria adelae Pilsbry, 1940
Cancellaria reticulata (Linnaeus, 1767)


26 FMRI Technical Report TR-3


Mollusca--W. G, Lyons


Macroinvertebrate Checklists


Camp et al. 1998








Camp eta!. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


Genus Trigonostoma de Blainville, 1827
Trigonostoma rugosum (Lamarck, 1822)
Trigonostoma tenerum (Philippi, 1848)

Genus Tritonoharpa Dall, 1908
Tritonoharpa lanceolata (Menke, 1828)

Family Drilliidae Morrison, 1966

Genus Bellaspira Conrad, 1868
Bellaspira pentagonalis (Dall, 1889)

Genus Cerodrillia Bartsch & Rehder, 1939
Cerodrillia bealiana Schwengel & McGinty, 1942
Cerodrillia clappi Bartsch & Rehder, 1939
Cerodrillia perryae Bartsch & Rehder, 1939
Cerodrillia thea (Dall, 1884)

Genus Drillia J. E. Gray, 1838
Drillia acurugata Dall, 1890
Drillia cydia (Bartsch, 1943)

Genus Lissodrillia Bartsch & Rehder, 1939
Lissodrillia simpsoni (Dall, 1887)

Genus Splendrillia Hedley, 1922
Splendrilliafucata (Reeve, 1845)
Splendrillia halidorema (Schwengel, 1940)
Splendrillia moseri (Dall, 1889)
Splendrillia woodringi (Bartsch, 1934)

Family Terebridae Morch, 1852

Genus Hastula H. & A. Adams, 1853
Hastula cinerea (Born, 1778)
Hastula hastata (Gmelin, 1791)
Hastula salleana (Deshayes, 1859)

Genus Terebra Bruguiere, 1789
Terebra arcas Abbott, 1954
Terebra concava (Say, 1826)
Terebra dislocata (Say, 1822)
Terebra floridana Dall, 1889
Terebra glossema Schwengel, 1942
Terebra protexta (Conrad, 1846)
Terebra rushii Dall, 1889
Terebra taurina (Lightfoot, 1786)
Terebra vinosa Dall, 1889

Family Strictispiridae J. H. McLean, 1971

Genus Strictispira J. H. McLean, 1971
Strictispira solida (C. B. Adams, 1850)


Family Turridae Swainson, 1840

Genus Compsodrillia Woodring, 1928
Compsodrillia eucosmia (Dall, 1889)

Genus Crassispira Swainson, 1840
Crassispira cubana (Melvill, 1923)
Crassispirafuscescens (Reeve, 1845)
Crassispira phasma Schwengel, 1940
Crassispira rhythmica Melvill, 1927
Crassispira tampaensis (Bartsch & Rehder, 1939)

Genus Pilsbryspira Bartsch, 1950
Pilsbryspira albocincta (C. B. Adams, 1845)
Pilsbryspira albomaculata (d'Orbigny, 1842)
Pilsbryspira leucocyma (Dall, 1884)
Pilsbryspira monilis (Bartsch & Rehder, 1939)
Pilsbryspira zebroides (Weinkauff, 1876)

Genus Polystira Woodring, 1928
Polystira albida (G. Perry, 1811)

Genus Pyrgospira J. H. McLean, 1971
Pyrgospira ostrearum (Stearns, 1872)

Genus Viridrillia Bartsch, 1943
Viridrillia williami Bartsch, 1943

Genus Zonulispira Bartsch, 1950
Zonulispira sanibelensis (Bartsch & Rehder, 1939)

Family Conidae Fleming, 1822

Genus Agathotoma Cossmann, 1899
Agathotoma candidissima (C. B. Adams, 1845)

Genus Brachycythara Woodring, 1928
Brachycythara barbarae Lyons, 1972
Brachycythara biconica (C. B. Adams, 1850)

Genus Conus Linnaeus, 1758
Conus amphiurgus Dall, 1889
Conus anabathrum Crosse, 1865
Conus attenuatus Reeve, 1844
Conus daucus Hwass, 1792
Conus delessertii R&cluz, 1843
Conus ermineus Born, 1778
Conus flamingo Petuch, 1980
Conusflavescens G. B. Sowerby II, 1834
Conus floridensis G. B. Sowerby II, 1870
Conus granulatus Linnaeus, 1758
Conus harasewychi Petuch, 1987
Conus mindanus Hwass, 1792
Conus mus Hwass, 1792


FMRI Technical Report TR-3 27


Camp et al. 1998


Macroinvertebrate Checklists


W. G. Lyons--Mollusca








Mollusca-W, G. Lyons Macroinvertebrate Checklists Camp et al. 1998


Conus patae Abbott, 1971
Conus pealii Green, 1830
Conus regius Gmelin, 1791
Conus sennottorum Rehder & Abbott, 1951
Conus spurius Gmelin, 1791
Conus stearnsii Conrad, 1869
Conus stimpsoni Dall, 1902
Conus ziczac Miihlfeld, 1816

Genus Cryoturris Woodring, 1928
Cryoturris cerinella (Dall, 1889)
Cryoturrisfargoi McGinty, 1955
Cryoturris quadrilineata (C. B. Adams, 1850)

Genus Daphnella Hinds, 1844
Daphnella cingulata (Dall, 1890)
Daphnella lymneiformis (Kiener, 1840)
Daphnella margaretae Lyons, 1972

Genus Glyphostoma Gabb, 1872
Glyphostoma dentiferum Gabb, 1872
Glyphostoma gabbii Dall, 1889

Genus Glyphoturris Woodring, 1928
Glyphoturris diminuta (C. B. Adams, 1850)
Glyphoturris eritima (Bush, 1885)
Glyphoturris quadrata (Reeve, 1845)
Glyphoturris rugirima (Dall, 1889)

Genus Glyptaesopus Pilsbry & Olsson, 1941
Glyptaesopus proctorae (M. Smith, 1936)

Genus Granoturris Fargo, 1953
Granoturris presleyi Lyons, 1972

Genus Ithycythara Woodring, 1928
Ithycythara auberiana (d'Orbigny, 1842)
Ithycythara lanceolata (C. B. Adams, 1850)
Ithycythara parkeri Abbott, 1958
Ithycythara pentagonalis (Reeve, 1845)
Ithycythara psila (Bush, 1885)

Genus Kurtziella Dall, 1918
Kurtziella accincta (Montagu, 1808)
Kurtziella atrostyla (Tryon, 1884)
Kurtziella cerina (Kurtz & Stimpson, 1851)
Kurtziella dorvilliae (Reeve, 1845)
Kurtziella limonitella (Dall, 1884)
Kurtziella perryae Bartsch & Rehder, 1939

Genus Lioglyphostoma Woodring, 1928
Lioglyphostoma canna (Dall, 1889)

Genus Mitrolumna Bucquoy, Dautzenberg & Dollfus,
1883


Mitrolumna biplicata (Dall, 1889)

Genus Nannodiella Dall, 1919
Nannodiella oxia (Bush, 1885) A
Nannodiella vespuciana (d'Orbigny, 1842)

Genus Platycythara Woodring, 1928
Platycythara elata (Dall, 1889)

Genus Pyrgocythara Woodring, 1928
Pyrgocythara filosa Rehder, 1943
Pyrgocythara hemphilli Bartsch & Rehder, 1939
Pyrgocythara plicosa (C. B. Adams, 1850)

Genus Rimosodaphnella Cossmann, 1915
Rimosodaphnella morra (Dall, 1881)

Genus Rubellatoma Bartsch & Rehder, 1939
Rubellatoma diomedea Bartsch & Rehder, 1939
Rubellatoma rubella (Kurtz & Stimpson, 1851)

Genus Stellatoma Bartsch & Rehder, 1939
Stellatoma stellata (Stearns, 1872)

Genus Tenaturris Woodring, 1928
Tenaturris bartletti (Dall, 1889)

Genus Thelecythara Woodring, 1928
Thelecythara floridana Fargo, 1953

Genus Vitricythara Fargo, 1953
Vitricythara metria (Dall, 1903)

Subclass Heterobranchia J. E. Gray, 1840
Order Heterostropha P. Fischer, 1885
Family Cornirostridae Ponder, 1990

Genus Tomura Pilsbry & Olsson, 1952
Tomura bicaudata (Pilsbry & McGinty, 1946)

Family Mathildidae Dall, 1889

Genus Mathilda Semper, 1865
Mathilda hendersoni Dall, 1927
Mathilda yucatecana (Dall, 1881)

Family Architectonicidae J. E. Gray, 1850

Genus Architectonica Riding, 1798
Architectonica nobilis Roding, 1798

Genus Heliacus d'Orbigny, 1842
Heliacus bisulcatus (d'Orbigny, 1842)
Heliacus cylindricus (Gmelin, 1791)


28 FMRI Technical Report TR-3


Mollusca-WW G. Lyons


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Camp et al. 1998







Camp et al. 1998 Macroinvertebrate Checklists W. 0. Lyons-Mollusca


Heliacus infundibuliformis perrieri (Rochebrune,
1881)

Genus Psilaxis Woodring, 1928
Psilaxis krebsii (Morch, 1875)

Genus Spirolaxis Monterosato, 1913
Spirolaxis centrifuga (Monterosato, 1890)

Family Rissoellidae J. E. Gray, 1850

Genus Rissoella J. E. Gray, 1847
Rissoella caribaea Rehder, 1943

Family Ebalidae Waren, 1994

Genus Henrya Bartsch, 1947
Henrya morrisoni Bartsch, 1947

Genus Murchisonella Morch, 1875
Murchisonella tampaensis (Bartsch, 1947)

Family Pyramidellidae J. E. Gray, 1840

Genus Boonea Roberston, 1978
Boonea bisuturalis (Say, 1822)
Boonea impressa (Say, 1822)
Boonea seminuda (C. B. Adams, 1839)

Genus Careliopsis Morch, 1875
Careliopsis bartschi (Aguayo & Rehder, 1936)
Careliopsis styliformis (Morch, 1875)

Genus Chrysallida Carpenter, 1857
Chrysallida gemmulosa (C. B. Adams, 1850)
Chrysallida toroensis (Olsson & McGinty, 1958)

Genus Cyclostremella Bush, 1897
Cyclostremella humilis Bush, 1897

Genus Eulimastoma Bartsch, 1904
Eulimastoma engonium (Bush, 1885)
Eulimastoma teres (Bush, 1885)
Eulimastoma weberi (Morrison, 1965))

Genus Fargoa Robertson, 1978
Fargoa bartschi (Winkley, 1909)
Fargoa bushiana (Bartsch, 1909)
Fargoa dianthophila (H. W. Wells & M. J. Wells,
1961)
Fargoa dux (Dall & Bartsch, 1906)

Genus Houbricka Wise, 1996
Houbricka incisa (Bush, 1899)


Genus Ividia Dall & Bartsch, 1904
Ividia abbotti (Olsson & McGinty, 1958)
Ividia havanensis (Pilsbry & Aguayo, 1933)

Genus Kleinella A. Adams, 1860
Kleinella cedrosa (Dall, 1884)

Genus Longchaeus Morch, 1875
Longchaeus candidus (Morch, 1875)
Longchaeus crenulatus (Holmes, 1860)

Genus Odostomia Fleming, 1817
Odostomia acutidens Dall, 1884
Odostomia didyma (A. E.Verrill & Bush, 1900)
Odostomia laevigata (d'Orbigny, 1842)
Odostomia product (C. B. Adams, 1840)
Odostomia somersi A. E.Verrill & Bush, 1900
Odostomia virginica J. B. Henderson & Bartsch,
1914

Genus Orinella Dall & Bartsch, 1904
Orinella vanhyningi Bartsch, 1944

Genus Peristichia Dall, 1889
Peristichia agria Dall, 1889
Peristichia toreta Dall, 1889

Genus Petitilla Wise, 1996
Petitilla crosseana (Dall, 1885)

Genus Pseudoscilla Boettger, 1901
Pseudoscilla babylonia (C. B. Adams, 1845)

Genus Pyramidella Lamarck, 1799
Pyramidella dolabrata (Linnaeus, 1758)
Pyramidella resticula (Dall, 1889)

Genus Sayella Dall, 1885
Sayellafusca (C. B. Adams, 1839)
Sayella hemphillii (Dall, 1884)

Genus Triptychus Morch, 1875
Triptychius niveus Morch, 1875

Genus Turbonilla Risso, 1826
Turbonilla areolata A. E.Verrill, 1873
Turbonilla belotheca Dall, 1889
Turbonilla conradi Bush, 1899
Turbonilla constricta Bush, 1899
Turbonilla coomansiVan Aartsen, 1994
Turbonilla curta Dall, 1889
Turbonilla dalli Bush, 1899
Turbonilla exilis (C. B. Adams, 1850)
Turbonilla hemphilli Bush, 1899


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W, G, Lyons--Mollusca







Macroinvertebrate Checklists


Turbonilla interrupta (Totten, 1835)
Turbonilla levis (C. B. Adams, 1850)
Turbonilla multicostata (C. B. Adams, 1850)
Turbonilla obeliscus (C. B. Adams, 1850)
Turbonilla protracta Dall, 1892
Turbonilla puncta (C. B. Adams, 1850)
Turbonilla punicea Dall, 1884
Turbonilla pupoides d'Orbigny, 1842
Turbonilla reticulata (C. B. Adams, 1850)
Turbonilla riisei Morch, 1875
Turbonilla textilis (Kurtz, 1860)
Turbonilla toyatani J. B. Henderson & Bartsch,
1914
Turbonilla validaVerrill & Bush, 1900
Turbonilla virga Dall, 1884
Turbonilla virgata Dall, 1892
Turbonilla viridaria Dall, 1884
Turbonilla wrightsvillensis Powell, 1981

Family Amathinidae Ponder, 1987

Genus Cyclothyca Stearns, 1891
Cyclothyca pacei Petuch, 1987

Genus Iselica Dall, 1918
Iselica anomala (C. B. Adams, 1850)

Order Uncertain
Family Acteonidae d'Orbigny, 1842

Genus Acteon Montfort, 1810
Acteon candens Rehder, 1939

Genus Rictaxis Dall, 1871
Rictaxis punctostriatus (C. B. Adams, 1840)

Subclass Opisthobranchia Milne-Edwards, 1848
Order Cephalaspidea P. Fischer, 1883
Family Ringiculidae Philippi, 1853

Genus Ringicula Deshayes, 1838
Ringicula semistriata d'Orbigny, 1842

Family Aplustridae J. E. Gray, 1847

Genus Hydatina Schumacher, 1817
Hydatina physis (Linnaeus, 1758)

Genus Micromelo Pilsbry, 1895
Micromelo undatus (Bruguiere, 1792)

Family Cylichnidae A. Adams, 1850

Genus Acteocina J. E. Gray, 1847


Acteocina atrata P. S. Mikkelsen & P. M.
Mikkelsen, 1984
Acteocina bidentata (d'Orbigny, 1841)
Acteocina bullata (Kiener, 1834)
Acteocina canaliculata (Say, 1826)
Acteocina candei (d'Orbigny, 1841)
Acteocina lepta Woodring, 1928
Acteocina recta (d'Orbigny, 1841)

Genus Cylichna Loven, 1846
Cylichna eburnea A. E.Verrill, 1885
Cylichna verrillii Dall, 1889

Genus Scaphander Montfort, 1810
Scaphander pilsbryi McGinty, 1955

Genus Tornatina A. Adams, 1850
Tornatina inconspicua (Olsson & McGinty, 1958)

Family Philinidae J. E. Gray, 1850

Genus Philine Ascanius, 1772
Philine sagra (d'Orbigny, 1841)

Family Aglajidae Pilsbry, 1895

Genus Chelidonura A. Adams, 1850
Chelidonura hirundinina (Quoy & Gaimard, 1833)
Chelidonura sabina Ev. Marcus & Er. Marcus,1970

Genus Navanax Pilsbry, 1895
Navanax aenigmaticus (Bergh, 1893)

Genus Philinopsis Pease, 1860
Philinopsis pusa (Ev. Marcus & Er. Marcus, 1967)

Family Gastropteridae Swainson, 1840

Genus Gastropteron Meckel in Kosse, 1813
Gastropteron chacmol Gosliner, 1989
Gastropteron vespertilium Gosliner & Armes, 1984

Family Retusidae Thiele, 1929

Genus Pyrunculus Pilsbry, 1895
Pyrunculus caelatus (Bush, 1885)

Genus Retusa Brown, 1827
Retusa sulcata (d'Orbigny, 1841)

Genus Volvulella Newton, 1891
Volvulella paupercula (Watson, 1883)
Volvulella persimilis (Morch, 1875)
Volvulella recta (Morch, 1875)


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Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


Family Bullidae Lamarck, 1801

Genus Bulla Linnaeus, 1758
Bulla solida Gmelin, 1791
Bulla striata Bruguiere, 1792

Family Haminoeidae Pilsbry, 1895

Genus Atys Montfort, 1810
Atys caribaeus (d'Orbigny, 1841)
Atys riiseanus Morch, 1875
Atys sandersoni Dall, 1881

Genus Haminoea Turton & Kingston in Carrington,
1830
Haminoea antillarum (d'Orbigny, 1841)
Haminoea elegans (J. E. Gray, 1825)
Haminoea petitii (d'Orbigny, 1841)
Haminoea succinea (Conrad, 1846)

Family Runcinidae H. & A. Adams, 1854

Genus Runcina Forbes & Hanley, 1851
Runcina divae (Ev. Marcus & Er. Marcus, 1963)

Order Acochlidioidea Kiithe, 1935
Family Microhedylidae Odhner, 1938

Genus Unela Er. Marcus, 1953
Unela remanei Er. Marcus, 1953

Order Thecosomata de Blainville, 1824
Family Cavoliniidae H. & A. Adams, 1854

Genus Creseis Rang, 1828
Creseis acicula (Rang, 1828)

Order Anaspidea P. Fischer, 1883
Family Akeridae Pilsbry, 1893

Genus Akera Miiller, 1776
Akera thompsoni Olsson & McGinty, 1951

Family Aplysiidae Lamarck, 1809

Genus Aplysia Linnaeus, 1767
Aplysia brasiliana Rang, 1828
Aplysia cervina (Dall & Simpson, 1901)
Aplysia dactylomela Rang, 1828
Aplysiajuliana Quoy & Gaimard, 1832
Aplysia morio (A. E.Verrill, 1901)
Aplysia parvula Morch, 1863

Family Notarchidae Eales, 1925


Genus Bursatella de Blainville, 1817
Bursatella leachii pleii Rang, 1828

Genus Dolabrifera J. E. Gray, 1847
Dolabrifera dolabrifera (Rang, 1828)

Genus Petalifera J. E. Gray, 1847
Petalifera ramosa Baba, 1959

Genus Phyllaplysia P. Fischer, 1872
Phyllaplysia engeli Er. Marcus, 1955
Phyllaplysia smaragda K. B. Clark, 1977

Genus Stylocheilus Gould, 1852
Stylocheilus citrinus (Rang, 1828)
Stylocheilus longicauda (Quoy & Gaimard, 1825)

Order Sacoglossa von Ihering, 1876
Family Volvatellidae Pilsbry, 1895

Genus Ascobulla Ev. Marcus, 1972
Ascobulla ulla (Er. Marcus & Ev. Marcus, 1970)

Genus Volvatella Pease, 1860
Volvatella bermudae K. B. Clark, 1982

Family Cylindrobullidae Thiele, 1931

Genus Cylindrobulla P. Fischer, 1857
Cylindrobulla beauii P. Fischer, 1857

Family Oxynoidae H. & A. Adams, 1854

Genus Lobiger Krohn, 1847
Lobiger souverbii P. Fischer, 1857

Genus Oxynoe Rafinesque, 1819
Oxynoe antillarum Morch, 1863
Oxynoe azuropunctata K. R. Jensen, 1980

Family Juliidae E. A. Smith, 1885

Genus Berthelinia Crosse, 1875
Berthelinia caribbea Edmunds, 1963

Family Elysiidae H. & A. Adams, 1854

Genus Bosellia Trinchese, 1891
Bosellia corinneae Ev. Marcus, 1973
Bosellia marcusi Ev. Marcus, 1972
Bosellia mimetica Trinchese, 1891

Genus Elysia Risso, 1818
Elysia canguzua Er. Marcus, 1955


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Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


Elysia chlorotica Gould, 1870
Elysia cornigera Nuttall, 1989
Elysia crispata Morch, 1863
Elysia evelinae Er. Marcus, 1957
Elysia ornata (Swainson, 1840)
Elysia papillosa A. E.Verrill, 1901
Elysia patina Ev. Marcus, 1980
Elysia serca Er. Marcus, 1955
Elysia subornata A. E.Verrill, 1901
Elysia tuca Ev. Marcus & Er. Marcus, 1967

Genus Thuridilla Bergh, 1872
Thuridilla picta (A. E.Verrill, 1901)

Family Polybranchiidae H. & A. Adams, 1854

Genus Caliphylla Costa, 1867
Caliphylla mediterranea A. Costa, 1867

Genus Cyerce Bergh, 1871
Cyerce antillensis Engel, 1927
Cyerce cristallina (Trinchese, 1881)

Genus Mourgona Er. Marcus & Ev. Marcus, 1970
Mourgona germaineae Er. Marcus & Ev. Marcus,
1970

Genus Polybranchia Pease, 1860
Polybranchia viridis (Deshayes, 1857)

Family Costasiellidae K. B. Clark, 1984

Genus Costasiella Pruvot-Fol, 1951
Costasiella nonatoi Ev. Marcus & Er. Marcus, 1960
Costasiella ocellifera (Simroth, 1895)

Family Stiligeridae Iredale & O'Donoghue, 1923

Genus Aplysiopsis Deshayes, 1864
Aplysiopsis zebra K. B. Clark, 1982

Genus Ercolania Trinchese, 1872
Ercolania coerulea Trinchese, 1893
Ercolaniafunerea (A. Costa, 1867)
Ercolania fuscata (Gould, 1870)

Genus Hermaea Lov6n, 1844
Hermaea cruciata Gould, 1870
Hermaea kingstoni (T. E. Thompson, 1977)
Hermaea verticilata (Ortea, 1984)

Genus Stiliger Ehrenberg, 1831
Stiligerfuscovittatus Lance, 1962
Stiliger vossi Ev. Marcus & Er. Marcus, 1960


Order Notaspidea P. Fischer, 1883
Family Umbraculidae Dall, 1889

Genus Umbraculum Schumacher, 1817
Umbraculum umbraculum (Lightfoot, 1786)

Family Pleurobranchidae Ferussac, 1822

Genus Berthella de Blainville, 1824
Berthella agassizii (MacFarland, 1909)
Berthella stellata (Risso, 1826)

Genus Pleurobranchaea Meckel in Leue, 1813
Pleurobranchaea inconspicua Bergh, 1897

Genus Pleurobranchus Cuvier, 1804
Pleurobranchus areolatus Morch, 1863
Pleurobranchus reesi White, 1952

Order Nudibranchia de Blainville, 1814
Family Corambidae Bergh, 1869

Genus Corambe Bergh, 1869
Corambe obscura (A. E.Verrill, 1870)

Family Goniodorididae H. & A. Adams, 1854

Genus Okenia Leuckart & Bronn in Menke, 1830
Okenia impexa Er. Marcus, 1957
Okenia zoobotryon (Smallwood, 1910)

Genus Trapania Pruvot-Fol, 1931
Trapania dalva Ev. Marcus, 1972

Family Aegiretidae P. Fischer, 1883

Genus Aegires Lov6n, 1844
Aegires sublaevis Odhner, 1932

Family Polyceratidae Alder & Hancock, 1845

Genus Polycera Cuvier, 1817
Polycera aurisula Er. Marcus, 1957
Polycera hummi Abbott, 1952
Polycera odhneri Er. Marcus, 1955
Polycera rycia Er. Marcus & Ev. Marcus, 1970

Genus Polycerella A. E.Verrill, 1880
Polycerella emertoni A. E.Verrill, 1880

Genus Thecacera Fleming, 1828
Thecacera pennigera (Montagu, 1815)

Family Chromodorididae Bergh, 1891


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Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


Genus Cadlina Bergh, 1878
Cadlina rumia Er. Marcus, 1955

Genus Chromodoris Alder & Hancock, 1855
Chromodoris binza Ev. Marcus & Er. Marcus, 1963
Chromodoris clenchi (Russell, 1935)
Chromodoris kempfi Ev. Marcus, 1970
Chromodoris morchii Bergh, 1879
Chromodoris nyalya Ev. Marcus & Er. Marcus,
1967

Genus Glossodoris Ehrenberg, 1831
Glossodoris sedna (Ev. Marcus & Er. Marcus, 1967)

Genus Hypselodoris Stimpson, 1855
Hypselodoris bayeri (Ev. Marcus & Er. Marcus,
1967)
Hypselodoris sycilla (Bergh, 1890)
Hypselodoris webbi (d'Orbigny, 1839)

Family Asteronotidae Thiele, 1935

Genus Aphelodoris Bergh, 1879
Aphelodoris antillensis (Bergh, 1879)

Family Dorididae Rafinesque, 1815

Genus Doris Linnaeus, 1758
Doris verrucosa Linnaeus, 1758

Genus Siraius Er. Marcus, 1955
Siraius bicolor (Bergh, 1884)
Siraius kyolis Ev. Marcus & Er. Marcus, 1967

Family Dendrodorididae O'Donoghue, 1924

Genus Dendrodoris Ehrenberg, 1831
Dendrodoris krebsii (Morch, 1863)
Dendrodoris warta Ev. Marcus & Gallagher, 1976

Genus Doriopsilla Bergh, 1880
Doriopsilla pharpa Er. Marcus, 1961

Family Phyllidiidae Rafinesque, 1815

Genus Ceratophyllidia Eliot, 1903
Ceratophyllidia papilligera (Bergh, 1890)

Family Archidorididae Bergh, 1892

Genus Atagema J. E. Gray, 1850
Atagema prea (Ev. Marcus & Er. Marcus, 1967)

Family Discodorididae Bergh, 1891


Genus Anisodoris Bergh, 1898
Anisodoris work Ev. Marcus & Er. Marcus, 1967

Genus Discodoris Bergh, 1877
Discodoris alba White, 1952
Discodoris evelinae Er. Marcus, 1955
Discodoris mortenseni Ev. Marcus & Er. Marcus,
1963
Discodoris phoca Ev. Marcus & Er. Marcus, 1967
Discodoris purcina Ev. Marcus & Er. Marcus, 1967

Genus Geitodoris Bergh, 1892
Geitodoris immunda Bergh, 1894
Geitodoris pusae (Er. Marcus, 1955)

Genus Peltodoris Bergh, 1880
Peltodoris greeleyi MacFarland, 1909

Genus Taringa Er. Marcus, 1955
Taringa telopia Er. Marcus, 1955

Family Platydorididae Bergh, 1891

Genus Platydoris Bergh, 1877
Platydoris angustipes (Morch, 1863)

Family Tritoniidae Menke, 1828

Genus Marionia Vayssiere, 1877
Marionia tedi Ev. Marcus, 1983

Genus Tritonia Cuvier, 1798
Tritonia bayeri Ev. Marcus & Er. Marcus, 1967

Genus Tritonidoxa Bergh, 1907
Tritonidoxa wellsi (Er. Marcus, 1961)

Genus Tritoniopsis Eliot, 1905
Tritoniopsisfrydis Er. Marcus & Ev. Marcus, 1970

Family Bornellidae P. Fischer, 1883

Genus Bornella A. Adams & Reeve, 1848
Bornella calcarata Morch, 1863

Family Lomanotidae Bergh, 1892

Genus Lomanotus V6rany, 1844
Lomanotus vermiformis Eliot, 1908

Family Scyllaeidae Rafinesque, 1815

Genus Scyllaea Linnaeus, 1758
Scyllaea pelagica Linnaeus, 1758


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Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et ci. 1998


Family Dotoidae J. E. Gray, 1853

Genus Doto Oken, 1815
Doto caramella Er. Marcus, 1957
Doto chica Ev. Marcus & Er. Marcus, 1960
Doto coronata (Gmelin, 1791)
Doto divae Ev. Marcus & Er. Marcus, 1960
Doto pita Er. Marcus, 1955
Doto uva Er. Marcus, 1955

Genus Miesea Er. Marcus, 1961
Miesea evelinae (Er. Marcus, 1957)

Family Arminidae Pruvot-Fol, 1927

Genus Armina Rafinesque, 1814
Armina wattla Ev. Marcus & Er. Marcus, 1967

Family Janolidae Pruvot-Fol, 1954

Genus Janolus Bergh, 1884
Janolus coins Er. Marcus, 1955
Janolus mucloc (Er. Marcus, 1958)

Family Flabellinidae Bergh, 1889

Genus Flabellina Voigt, 1834
Flabellina engeli Ev. Marcus & Er. Marcus, 1968

Family Eubranchidae Odhner, 1934

Genus Eubranchus Forbes, 1838
Eubranchus coniclus (Er. Marcus, 1958)

Family Tergipedidae Bergh, 1889

Genus Catriona Winckworth, 1941
Catriona maua Ev. Marcus & Er. Marcus, 1960
Catriona oba Ev. Marcus, 1970

Genus Cuthona Alder & Hancock, 1855
Cuthona caerulea (Montagu, 1804)
Cuthona perca (Er. Marcus, 1958)
Cuthona tina (Er. Marcus, 1957)

Genus Tenellia A. Costa, 1866
Tenelliafuscata (Gould, 1870)

Family Fionidae Alder & Hancock, 1855

Genus Fiona Alder & Hancock in Forbes & Hanley,
1851
Fiona pinnata (Eschscholtz, 1831)


Family Facelinidae Bergh in Carus, 1889

Genus Austraeolis Burn, 1962
Austraeolis catina Ev. Marcus & Er. Marcus, 1967

Genus Cratena Bergh, 1864
Cratena pilata (Gould, 1870)

Genus Dondice Er. Marcus, 1958
Dondice occidentalis (Engel, 1925)

Genus Favorinus M. E. Gray, 1850
Favorinus auritulus Er. Marcus, 1955

Genus Godiva Macnae, 1954
Godiva rubrolineata Edmunds, 1964

Genus Learchis Bergh, 1889
Learchis poica Ev. Marcus & Er. Marcus, 1960

Genus Phidiana J. E. Gray, 1850
Phidiana lynceus Bergh, 1867

Family Aeolidiidae d'Orbigny, 1834

Genus Spurilla Bergh, 1864
Spurilla neapolitana (delle Chiaje, 1823)
Spurilla verrucicornis (A. Costa, 1867)

Family Glaucidae Menke, 1828

Genus Glaucus FBrster, 1777
Glaucus atlanticus FSrster, 1777

Subclass Pulmonata Cuvier, 1817
Order Archaeopulmonata Morton, 1955
Family Ellobiidae H. & A. Adams, 1854

Genus Blauneria Shuttleworth, 1854
Blauneria heteroclita (Montagu, 1808)

Genus Creedonia Martins, 1996
Creedonia succinea (Pfeiffer, 1854)

Genus Ellobium Riding, 1798
Ellobium dominicense (Ferussac, 1821)

Genus Laemodonta Philippi, 1846
Laemodonta cubensis (Pfeiffer, 1854)

Genus Leuconopsis Hutton, 1884
Leuconopsis novinundi (Pilsbry & McGinty, 1949)

Genus Melampus Montfort, 1810


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Camp et al. 1998 Macroinvertebrate Checklists W, 0. Lyons-Mollusca


Subgenus Melampus Montfort, 1810
Melampus (Melampus) bidentatus Say, 1822
Melampus (Melampus) coffeus (Linnaeus, 1758)
Subgenus Detracia J. E. Gray, 1840
Melampus (Detracia) bullaoides (Montagu, 1808)
Melampus (Detracia) floridanus Pfeiffer, 1856
Melampus (Detracia) monile (Bruguiere, 1789)
Melampus (Detracia) morrisoni Martins, 1996

Genus Microtralia Dall, 1894
Microtralia occidentalis (Pfeiffer, 1854)

Genus Pedipes Scopoli, 1777
Pedipes mirabilis (Miihlfeld, 1816)
Pedipes ovalis (C. B. Adams, 1849)

Genus Tralia J. E. Gray, 1840
Tralia ovula (Bruguiere, 1789)

Order Basommatophora Schmidt, 1855
Family Siphonariidae J. E. Gray, 1840

Genus Siphonaria Sowerby, 1823
Siphonaria alternate (Say, 1826)
Siphonaria pectinata (Linnaeus, 1758)

Genus Williamia Monterosato, 1884
Williamia krebsii (Morch, 1877)

Family Trimusculidae Zilch, 1959

Genus Trimusculus Schmidt, 1818
Trimusculus carinatus (Dall, 1870)

Order Systellommatophora Pilsbry, 1948
Family Onchidiidae J. E. Gray, 1824

Genus Onchidella M. E. Gray, 1850
Onchidella brattstroemi Ev. Marcus, 1978
Onchidellafloridana (Dall, 1885)
Onchidella wah Ev. Marcus, 1978

Class Cephalopoda Cuvier, 1797
Squids and Octopuses

Order Sepioidea Leach, 1817
Family Spirulidae Rafinesque, 1815

Genus Spirula Lamarck, 1799
Spirula spirula (Linnaeus, 1758)

Family Sepiolidae Leach, 1817

Genus Semirossia Steenstrup, 1881


Semirossia tenera (A. E.Verrill, 1880)

Order Teuthoidea Owen, 1836
Family Loliginidae Lesueur, 1821

Genus Loligo Schneider, 1784
Loligo pealeii Lesueur, 1821
Loligo pleii de Blainville, 1823

Genus Lolliguncula Steenstrup, 1881
Lolliguncula brevis (de Blainville, 1823)

Genus Pickfordiateuthis Voss, 1953
Pickfordiateuthis pulchella Voss, 1953

Genus Sepioteuthis de Blainville, 1824
Sepioteuthis sepioidea (de Blainville, 1823)

Order Octopoda Rafinesque, 1815
Family Octopodidae Rafinesque, 1815

Genus Octopus Lamarck, 1798
Octopus briareus Robson, 1929
Octopus burryi Voss, 1953
Octopus defilippi VWrany, 1851
Octopus filosus Howell, 1868
Octopus joubini Robson, 1929
Octopus macropus Risso, 1826
Octopus vulgaris Lamarck, 1798

Family Argonautidae Rafinesque, 1815

Genus Argonauta Linnaeus, 1758
Argonauta argo Linnaeus, 1758
Argonauta hians Solander, 1786
Argonauta nodosus Solander, 1786

Pelagic Mollusks

Many mollusks that live in the pelagic realm, either
free-swimming or on the alga Sargassum, wash up con-
spicuously on Florida's shores. Most of these species
are evidenced in the nearshore environment principally
by their shells and/or dead or dying animals (e.g., the
gastropods Janthina, Recluzia, Fiona, and Glaucus and the
cephalopod Spirula) and egg cases ("paper nautilus":
cephalopods, genus Argonauta), but some (e.g., the gas-
tropods Litiopa and Scyllaea) may drop off Sargassum
near shore and live for brief periods among the shal-
low-water benthos. All of these species are included in
the general list. That list also includes Creseis acicula, a
pteropod snail with a glassy, needle-like shell that can
become entangled in swimwear, causing a type of
"swimmer's itch."The species sometimes occurs in


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Mollusca-W. G. Lyons Macroinverfebrate Checklists Camp et al. 1998


great densities in coastal waters, and windows of its
shells are occasionally found along Florida beaches.
Consult Lalli and Gilmer (1989) for a comprehensive
summary of these and the following species of pelagic
snails.
The gastropods listed hereafter lead exclusively
pelagic lives in the open ocean. Many of the species
occur throughout the Atlantic Ocean, and some have
world-wide distributions. Although they occur in wa-
ters that may intrude into coastal environments in
areas such as southeastern and northwestern Florida,
representatives of these species are virtually never
collected except deliberately by those sampling plank-
ton and nekton. Pelagic cephalopods, principally
squids, are also encountered occasionally in Florida
coastal waters, but most of these occurrences are ac-
cidental, and the many such deep-water species that
may be involved are not listed here.

Class Gastropoda Cuvier, 1797
Order Neotaenioglossa Hailer, 1882
Family Atlantidae Rang, 1829

Genus Atlanta Lesueur, 1817
Atlanta brunnea J. E. Gray, 1850
Atlanta helicinoides J. E. Gray, 1850
Atlanta inclinata J. E. Gray, 1850
Atlanta lesueurii J. E. Gray, 1850
Atlanta peronii Lesueur, 1817

Genus Oxygyrus Benson, 1835
Oxygyrus keraudrenii (Lesueur, 1817)

Genus Protatlanta Tesch, 1908
Protatlanta souleyeti (E. A. Smith, 1888)

Family Carinariidae de Blainville, 1818

Genus Cardiapoda d'Orbigny, 1835
Cardiapoda placenta (Lesson, 1830)

Genus Carinaria Lamarck, 1801
Carinaria lamarcki P&ron & Lesueur, 1810

Family Pterotracheidae Gray, 1840

Genus Firoloida Lesueur, 1817
Firoloida demarestia Lesueur, 1817

Genus Pterotrachea Forskil, 1775
Pterotrachea hippocampus Philippi, 1836
Pterotrachea scutata Gegenbaur, 1855

Order Thecosomata de Blainville, 1824


Family Limacinidae de Blainville, 1823

Genus Limacina Bosc, 1817
Limacina bulimoides (d'Orbigny, 1836)
Limacina inflata (d'Orbigny, 1836)
Limacina lesueurii (d'Orbigny, 1836)
Limacina trochiformis (d'Orbigny, 1836)

Family Cavoliniidae H. & A. Adams, 1854

Genus Cavolinia Abildgaard, 1791
Cavolinia gibbosa (d'Orbigny, 1836)
Cavolinia inflexa (Lesueur, 1813)
Cavolinia tridentata (Niebuhr, 1775)
Cavolinia uncinata (Rang, 1829)

Genus Clio Linnaeus, 1767
Clio chaptalii J. E. Gray, 1850
Clio cuspidata (Bosc, 1802)
Clio polita (Pelseneer, 1888)
Clio pyramidata Linnaeus, 1767
Clio recurva (Children, 1823)

Genus Creseis Rang, 1828
Creseis virgula (Rang, 1828)

Genus Cuvierina Boas, 1886
Cuvierina columnella (Rang, 1827)

Genus Diacavolinia Van der Spoel, 1987
Diacavolinia deblainvillei van der Spoel, Bleeker
& Kobayashi, 1993

Genus Diacria J. E. Gray, 1842
Diacria trispinosa (de Blainville, 1821)

Genus Hyalocylis Fol, 1875
Hyalocylis striata (Rang, 1828)

Genus Styliola Lesueur, 1825
Styliola subula (Quoy & Gaimard, 1827)

Family Peraclididae C. W. Johnson, 1915

Genus Peracle Forbes, 1844
Peracle bispinosa Pelseneer, 1888
Peracle diverse Monterosato, 1875
Peracle reticulata (d'Orbigny, 1836)
Peracle triacantha P. Fischer, 1882

Order Gymnosomata de Blainville, 1824
Family Clionidae Rafinesque, 1815

Genus Paraclione Tesch, 1903


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Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


Paraclione longicaudata (Souleyet, 1852)

Family Cliopsidae O. G. Costa, 1873

Genus Cliopsis Troschel, 1854
Cliopsis krohnii Troschel, 1854

Family Notobranchaeidae Pelseneer, 1886

Genus Notobranchaea Pelseneer, 1886
Notobranchaea macdonaldi Pelseneer, 1886

Genus Prionoglossa Tesch, 1950
Prionoglossa tetrabranchiata (Bonnevie, 1913)

Family Pneumodermatidae Latreille, 1825

Genus Pneumoderma Cuvier, 1805
Pneumoderma mediterraneum (van Beneden, 1838)


Pneumoderma violaceum violaceum (d'Orbigny,
1836)

Genus Pneumodermopsis Keferstein, 1862
Pneumodermopsis macrochira (Meisenheimer,
1905)
Pneumodermopsis paucidens (Boas, 1886)

Family Thliptodontidae Kwietniewski, 1910

Genus Thliptodon Boas, 1886
Thliptodon diaphanus (Meisenheimer, 1903)

Order Nudibranchia Cuvier, 1817
Family Phylliroidae Menke, 1830

Genus Phylliroe P&ron & Lesueur, 1810
Phylliroe atlantica Bergh, 1871
Phylliroe bucephalum Peron & Lesueur, 1810


References


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FMRI Technical Report TR-3 37


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Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


ABBOTT, R.T. 1986. Seashells of North America Guide
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38 FMRI Technical Report TR-3


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Camp et 01. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


BAKER, H. B. 1957. Fasciolaria distans Lamarck. Minutes
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BALES, B. R. 1942. Albinos of Muricidea ostrearum Con-
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western Atlantic. Johnsonia 1(5): 1-11.


FMRI Technical Report TR-3 39


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Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


BEQUAERT,J. C. 1943.The genus Littorina in the west-
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BERGH, L. S. R. 1894. Reports on the dredging opera-
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BERTSCH, H. 1975. Distributional and anatomical ob-
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BERTSCH, H. 1988. Anatomy and zoogeography of
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BEU, A. G. 1970. The Mollusca of the genus Charonia
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BEU, A. G. 1970. The Mollusca of the subgenus Mono-
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BEU, A. G. 1971. Genera of the bivalve family Mesodes-
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BEU, A. G. 1985. A classification and catalogue of liv-
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BEU, A. G. 1986. Taxonomy of gastropods of the fami-
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BEU, A. G. 1988. Taxonomy of gastropods of the fami-
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Early history of the families, with four new genera
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BEU, A. G., and W. O. CERNOHORSKY. 1986. Taxon-
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BEU, A. G., and E. A. KAY. 1988. Taxonomy of gas-
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BEU, A. G., and P. A. MAXWELL. 1987. A revision of the
fossil and living gastropods related to Plesiotriton Fis-
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BIELER, R. 1984. Die gattungen der Architectonicidae
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BIELER, R. 1985. Die gattungen der Architectonicidae
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BIELER, R. 1985. Die gattungen der Architectonicidae
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BIELER, R. 1987. Die gattungen der Architectonicidae
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203-215.

BIELER, R. 1988. Phylogenetic relationships in the gas-


40 FMRI Technical Report TR-3


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Camp et al. 1998







Macroinvertebrate Checklists


tropod family Architectonicidae, with notes on the
family Mathildidae (Allogastropoda). Malacological
Review, Supplement 4: 205-240.

BIELER, R. 1992. Tenagodus or Siliquaria? Unraveling tax-
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BIELER, R. 1993. Architectonicidae of the Indo-Pacific
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BIELER, R., and M. G. HADFIELD. 1990. Reproductive
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BIELER, R., and P. M. MIKKELSEN. 1988. Anatomy
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1-29.

BINGHAM, F O. 1969. Littorina nebulosa in Florida.The
Nautilus 82(4): 146, 147.

BINGHAM, F. O. 1972. First recorded occurrence of
Littorina tessellata Philippi, 1847, from the shores of
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BINGHAM, F 0. 1972. Shell growth in the gastropod
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BINGHAM, F. O. 1972. Several aspects of the repro-
ductive biology of Littorina irrorata (Gastropoda). The
Nautilus 86(1): 8-10.

BINGHAM, F. O. 1972. The influence of environmen-
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toral gastropod Littorina irrorata. Bulletin of Marine
Science 22(2): 309-335.

BINGHAM, F. O.1972. The mucus holdfast of Littorina
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BISHOF, D. E. 1980.The Ecology of Molluscan Infauna
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BLAKE, N. J., L. J. DOYLE, andT. E. PYLE. 1976.The mac-
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BLOOM, S. A., J. L. SIMON, andV. D. HUNTER. 1972.
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BONSDORFF, E., and W. G. NELSON. 1992. The ecol-
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The Veliger 35(4): 358-365.

BORKOWSKI, T.V. 1974. Growth, mortality, and pro-
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BORKOWSKI,T.V. 1975.Variability among Caribbean
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BORKOWSKI,T.V., and M. R. BORKOWSKI. 1969.The
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BOSS, K. J. 1966. The subfamily Tellininae in the west-
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BOSS, K. J. 1968. The subfamily Tellininae in the west-
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Johnsonia 4(46): 273-344.

BOSS, K. J. 1969. The subfamily Tellininae in the west-
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BOSS, K. J. 1972. The genus Semele in the western At-
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BOSS, K.J. 1982. Mollusca. Pp. 945-1166 in S. P. Parker.
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BOSS, K. J. 1982. Classification of Mollusca. Pp.
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BOSS, K. J., N. K. EBBS, and W. C. STEWART. 1967. Mon-
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BOSS, K. J., and A. S. MERRILL. 1965.The family Pan-
doridae in the western Atlantic. Johnsonia 4(44):
181-215.


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BOSS, K. J., and D. R. MOORE. 1967. Notes on Malleus
(Paramalleus) candeanus (d'Orbigny) (Mollusca: Bi-
valvia). Bulletin of Marine Science 17(1): 85-94.

BOSS, K. J., and M. L. WASS. 1970. Northward range
extension of Cyclinella tenuis Recluz.The Nautilus 83(3):
112,113.

BOUCHET, P. 1984. Les Triphoridae de Mediterrande
et du proche Atlantique (Mollusca, Gastropoda). La-
vori della Societa Italiana di Malacologia 2: 15-58.

BOUCHET, P., and J. ORTEA. 1980. Quelques Chro-
modorididae bleus (Mollusca, Gastropoda, Nudi-
branchiata) de l'Atlantique oriental. Annales de l'In-
stitut Oceanographique 56(2): 117-125.

BOUCHET, P., and A. WAREN. 1985. Revision of the
northeast Atlantic bathyal and abyssal Neogastropoda
excluding Turridae (Mollusca, Gastropoda). Bollettino
Malacologica, Supplemento 1: 193-296.

BOUCHET, P., and A. WAREN. 1986. Revision of the
northeast Atlantic bathyal and abyssal Aclididae, Eu-
limidae, Epitoniidae (Mollusca, Gastropoda). Bollettino
Malacologico, Supplemento 2: 297-576.

BOUCHET, P., and A. WAREN. 1993. Revision of the
northeast Atlantic bathyal and abyssal Mesogastropoda.
Bollettino Malacologico, Supplemento 3: 577-840.

BOWER, W. J. 1945. Egg laying process of Strombus
pugilis alatus Gmelin. The Nautilus 59(1): 35.

BOWLING, C. 1994. Habitat and size of the Florida
crown conch (Melongena corona Gmelin): Why big snails
hang out at bars. Journal of Experimental Marine Bi-
ology and Ecology 175: 181-195.

BRAKONIECKI, T. F. 1996, A revision of the genus Pick-
fordiateuthisVoss, 1953 (Cephalopoda; Myopsida). Bul-
letin of Marine Science 58(1): 9-28.

BRATCHER,T., andW. O. CERNOHORSKY. 1987. Liv-
ing Terebras of the World; A Monograph of the Recent
Terebridae of the World. American Malacologists, Inc.,
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BRETSKY, S. S. 1969. Phenetic and Phylogenetic Clas-
sifications of the Lucinidae (Mollusca: Bivalvia). Ph. D.
Dissertation,Yale University, New Haven. 485 pp., 22 pls.

BRETSKY, S. S. 1971. Evaluation of the efficacy of nu-
merical taxonomic methods: An example from the bi-


valve mollusks. Systematic Zoology 20(2): 204-222.

BRETSKY, S. S. 1976. Evolution and classification of the
Lucinidae (Mollusca: Bivalvia). Palaeontographica
Americana 8(50): 1-337, 36 pls.

BRITTON, J. C., JR. 1970. The Lucinidae (Mollusca: Bi-
valvia) of the Western Atlantic Ocean. Ph.D. Disserta-
tion, The George Washington University, Washington,
D. C. 566 pp., 23 pls.

BRITTON, J. C., JR. 1972. Two new species and a new
subgenus of Lucinidae (Mollusca: Bivalvia), with notes
on certain aspects of lucinid phylogeny. Smithsonian
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BRITTON, J. C., JR., and B. MORTON. 1989. Shore
Ecology of the Gulf of Mexico. University of Texas
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BROOK, I. M. 1978. Comparative macrofaunal abun-
dance in turtlegrass (Thalassia testudinum) communi-
ties in south Florida characterized by high blade den-
sity. Bulletin of Marine Science 28(1): 212-217.

BROOKS, J. 1970. Sebastian Inlet in April. Seafari 12(6): 6.

BROOKS, J. 1970. Diodora from Sebastian Inlet. Seafari
12(8-9): 10.

BROOKS, J. 1971. Bear Cut in April. Seafari 13(4): 6.

BROWNELL, W. N., and J. M. STEVELY. 1981. The bi-
ology, fisheries, and management of the queen conch,
Strombus gigas. Marine Fisheries Review 43(7): 1-12.

BRUNCKHORST, D. J. 1990. Description of a new
species of Phyllidiopsis Bergh (Nudibranchia: Dori-
doidea: Phyllidiidae) from the tropical western Pacific,
with comments on the Atlantic species. Journal of Mol-
luscan Studies 56(4): 577-584.

BULLIS, H. R., JR., and J. R. THOMPSON. 1965. Col-
lections by the exploratory fishing vessels Oregon, Sil-
ver Bay, Combat, and Pelican made during 1956-1960 in
the southwestern North Atlantic. United States Fish and
Wildlife Service, Special Scientific Report-Fisheries
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BULLIS, H. R., JR. 1956. The genus Scaphander in the
Gulf of Mexico and notes on the western Atlantic
species. Bulletin of Marine Science of the Gulf and
Caribbean 6(1): 1-17.


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Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


BULLIS, H. R., JR., and R. CUMMINS, JR. 1961. An in-
terim report of the Cape Canaveral calico scallop bed.
Commercial Fisheries Review 23(10): 1-8.

BULLIS, H. R., JR., and R. M. INGLE. 1959. A new fish-
ery for scallops in western Florida. Proceedings of the
Gulf and Caribbean Fisheries Institute, Eleventh An-
nual Session: 75-78.

BULLOCK, R. C. 1974. A contribution to the systematics
of some West Indian Latirus (Gastropoda: Fasciolari-
idae). The Nautilus 88(3): 69-79.

BULLOCK, R. C. 1976. On the identity of two sup-
posed fasciolariid gastropods, Fusus mexicanus Reeve,
1848, and Peristernia nodulosa A. Adams, 1855. Tulane
Studies in Geology and Paleontology 12: 133-136.

BULLOCK, R. C. 1985. The Stenoplax limaciformis
(Sowerby, 1832) species complex in the New World
(Mollusca: Polyplacophora: Ischnochitonidae). The
Veliger 27(3): 291-307.

BULLOCK, R. C. 1988. The genus Chiton in the New
World (Polyplacophora: Chitonidae).TheVeliger 31(3/4):
141-191.

BURGESS, C. M. 1985. Cowries of the World. Gordon
Verhoef Seacomber Publications, Cape Town. 289 pp.

BURGGRAF, P. 1970.Turridae and beach drift. Seafari
12(7): 14, 15.

BURGGRAF, P. 1970. Pompano Beach dredging. Seafari
12(8-9): 11, 12.

BURGHARDT, G. 1971. A Mopaliella from Florida. Of
Sea and Shore 2(3): 147.

BUSH, K. J. 1899. Descriptions of new species of Tur-
bonilla of the western Atlantic fauna, with notes on
those previously known. Proceedings of the Academy
of Natural Sciences of Philadelphia (1899): 145-177,
pl. 8.

BUSH, K. J. 1909. Notes on the family Pyramidellidae.
American Journal of Science 27: 475-484.

BUTTON, F. L. 1901. Trivia paucilirata Sowerby. The
Nautilus 14(12): 144.

CAIRNS, S. 1992. Oligohaline areas in Tampa Bay trib-
utaries: Spatial extent and species list.Tampa Bay Na-
tional Estuary Program Technical Report 04-92.25 pp.


CALDWELL, D. K. 1959. Notes on the crown conch
Melongena corona. The Nautilus 72(4): 117-122.

CALKINS, W. W. 1878. Catalogue of the marine shells
of Florida, with notes and descriptions of several new
species. Proceedings of the Davenport Academy of
Natural Sciences 2: 232-252.

CARPENTER, J. S. 1967. History of scallop and clam ex-
plorations in the Gulf of Mexico. Commercial Fish-
eries Review 29(1): 47-53.

CARTER, J. G. 1978. Ecology and evolution of the Gas-
trochaenacea (Mollusca, Bivalvia) with notes on the evo-
lution of the endolithic habitat. Peabody Museum of
Natural History,Yale University, Bulletin 41: 1-92.

CATE, C. N. 1973. A systematic revision of the Recent
cypraeid family Ovulidae (Mollusca: Gastropoda).The
Veliger 15, Supplement: i-iv, 1-116,51 pls.

CATE, C. N. 1976. Five new species of Ovulidae (Mol-
lusca: Gastropoda). The Veliger 19(2): 159-162, 1 pl.

CATE, C. N. 1978. New species of Ovulidae and rein-
statement of Margovula pyrulina (A. Adams, 1854) (Gas-
tropoda). The Nautilus 92(4): 160-167.

CATE, C. N. 1979. A review of the Triviidae (Mollusca:
Gastropoda). San Diego Society of Natural History
Memoir 10: 1-126.

CATHER, J. N., and M. E. CROVO. 1972. The spawn,
early development and larvae of Cyphoma gibbosum
(Cypraeacea). The Nautilus 85(4): 111-114.

CERNOHORSKY, W. O.1970. Systematics of the fam-
ilies Mitridae &Volutomitridae. Bulletin of the Auck-
land Institute and Museum 8: i-iv, 1-190.

CERNOHORSKY, W. 0. 1975. The taxonomy of some
west American and Atlantic Nassariidae based on
their type-specimens. Records of the Auckland Insti-
tute and Museum 12: 121-173.

CERNOHORSKY, W. O. 1976. The Mitridae of the
World. Part I. The subfamily Mitrinae. Indo-Pacific
Mollusca 3(17): 273-528.

CERNOHORSKY, W. 0. 1977. The taxonomy of some
Indo-Pacific Mollusca. Part 5. With descriptions of new
taxa and remarks on Nassarius albus (Say). Records of
the Auckland Institute and Museum 14: 121-132.


FMRI Technical Report TR-3 43


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W. G. Lyons--Mollusca







Macroinvertebrate Checklists


CERNOHORSKY, W. 0.1978. The date of publication
of Anton's "Verzeichniss der Conchylien."TheVeliger
20(3): 229.

CERNOHORSKY, W. 0. 1978. The taxonomy of
Caribbean-Atlantic Costellariidae (Mollusca: Gas-
tropoda). Records of the Auckland Institute and Mu-
seum 15: 87-109.

CERNOHORSKY, W. 0.1984. Systematics of the fam-
ily Nassariidae (Mollusca: Gastropoda). Bulletin of the
Auckland Institute and Museum 14: i-iv, 1-356.

CERNOHORSKY, W. 0. 1991. The Mitridae of the
World. Part 2. The subfamily Mitrinae concluded and
subfamilies Imbricariinae and Cylindromitrinae.
Monographs of Marine Mollusca 4: 1-164.

CHANLEY, P. 1969. Donaxfossor: A summer range ex-
tension of Donax variabilis. The Nautilus 83(1): 1-14.

CHOI, D. R., and R. N. GINSBURG. 1983. Distribution
of coelobites (cavity-dwellers) in coral rubble across the
Florida reef tract. Coral Reefs 2: 165-172.

CLAMPIT, L. 1993. Shelling in the Florida Panhandle.
Texas Conchologist 30(1): 8, 9.

CLARK, K. B. 1977. Phyllaplysia smaragda (Opistho-
branchia: Notarchidae), a new anaspidean from Florida.
Bulletin of Marine Science 27(4): 651-657.

CLARK, K. B. 1982. A new Aplysiopsis (Opisthobranchia:
Hermaeidae) from central Florida, with a brief sum-
mary of the ceratiform families of the order Ascoglossa
(= Sacoglossa). Bulletin of Marine Science 32(1):
213-219.

CLARK, K. B., and D. DeFREESE. 1987. Population
ecology of Caribbean Ascoglossa (Mollusca: Opistho-
branchia): A study of specialized algal herbivores.
American Malacological Bulletin 5(2): 259-208.

CLARK, K. B., and A. GOETZFRIED. 1976. Lomanotus
stauberi, a new dendronotacean nudibranch from cen-
tral Florida (Mollusca: Opisthobranchia). Bulletin of
Marine Science 26(4): 474-478.

CLARKE, A. H., JR. 1965. The scallop superspecies Ae-
quipecten irradians (Lamarck). Malacologia 2: 161-188.

CLENCH, W. J. 1923. The marine shells of Sanibel,
Florida. The Nautilus 37(2): 52-56.


CLENCH, W. J. 1925. Additions to the list of marine
shells of Sanibel, Florida. The Nautilus 38(3): 93-95.

CLENCH, W. J. 1934. A new subspecies of Oliva retic-
ularis from southern Florida.The Nautilus 47(4): 142,143,
pl. 7.

CLENCH, W. J. 1937. Onchidium (Onchidella) floridanum
Dall. The Nautilus 50(3): 85, 86.

CLENCH, WJ. J1939. Additional notes on Terebraflam-
mea Lamarck. The Nautilus 53(1): 7, 8.

CLENCH, W.J. 1942.The genus Ficus in the western At-
lantic. Johnsonia 1(2): 1, 2.

CLENCH, W. J. 1942.The genera Dosinia, Macrocallista
and Amiantis in the western Atlantic. Johnsonia 1(3): 1-8.

CLENCH, W. J. 1942. The genus Conus in the western
Atlantic. Johnsonia 1(6): 1-40.

CLENCH, W. J. 1944. The genera Casmaria, Galeodea,
Phalium and Cassis in the western Atlantic. Johnsonia
1(16): 1-16.

CLENCH, W. J. 1945. The West Indian fauna in south-
ern Florida. The Nautilus 59(1): 32, 33.

CLENCH, W.J. 1945.The genus Ficus in the western At-
lantic. Johnsonia 1(18): 1-3.

CLENCH, W. J. 1945. The genus Conus in the western
Atlantic. Johnsonia 1(18): 4.

CLENCH, W. J. 1946. The genera Bathyaurinia, Rehder-
ia and Scaphella in the western Atlantic. Johnsonia
2(22): 41-60.

CLENCH, W. J. 1947. The genera Purpura and Thais in
the western Atlantic. Johnsonia 2(23): 61-91.

CLENCH, W. J. 1953. The genus Murex in the western
Atlantic. Johnsonia 2(32): 359-361.

CLENCH, W. J. 1953. The genus Conus in the western
Atlantic. Johnsonia 2(32): 363-376.

CLENCH, W. J. 1953. The genera Scaphella and Aurin-
iopsis in the western Atlantic. Johnsonia 2(32): 376-380.

CLENCH, W. J. 1954. The occurrence of dines in mol-
luscan populations. Systematic Zoology 3(3): 122-125.


44 FMRI Technical Report TR-3


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Macroinvertebrate Checklists


CLENCH, W. J. 1959. The genus Murex in the western
Atlantic. Johnsonia 3(39): 331-334.

CLENCH, W. J. 1964. The genera Pedipes and Laemod-
onta in the western Atlantic. Johnsonia 4(42): 117-128.

CLENCH, W.J. 1966. Extension of range for Dosinia dis-
cus. The Nautilus 80(2): 70.

CLENCH, W. J. 1967. Ficus carolae and F.floridensis.The
Nautilus 81(4): 107.

CLENCH, W. J., and R. T. ABBOTT. 1941. The genus
Strombus in the western Atlantic. Johnsonia 1(1): 1-15.

CLENCH, W. J., and R.T. ABBOTT. 1942.The genera Tec-
tarius and Echininus in the western Atlantic. Johnsonia
1(4): 1-4.

CLENCH, W. J., and R. T. ABBOTT. 1943. The genera
Cypraecassis, Morum, Sconsia and Dalium in the west-
ern Atlantic. Johnsonia 1(9): 1-8.

CLENCH, W. J., and R. T. ABBOTT. 1943. The genera
Gaza and Livona in the western Atlantic. Johnsonia
1(12): 1-9.

CLENCH, W. J., and R. T. ABBOTT. 1945. The genus
Strombus in the western Atlantic. Johnsonia 1(18): 1.

CLENCH, W. J., and C. G. AGUAYO. 1943.The genera
Xenophora and Tugurium in the western Atlantic. John-
sonia 1(8): 1-6.

CLENCH, W. J., and R. C. BULLOCK. 1970.The genus
Conus in the western Atlantic. Johnsonia 4(48): 372-379.

CLENCH, W. J., and I. PtREZ-FARFANTE. 1945. The
genus Murex in the western Atlantic. Johnsonia 1(17):
1-58.

CLENCH, W. J., and L. C. SMITH. 1944. The family
Cardiidae in the western Atlantic. Johnsonia 1(13):
1-32.

CLENCH, W. J., and R. D. TURNER. 1946. The genus
Bankia in the western Atlantic. Johnsonia 2(19): 1-28.

CLENCH, W. J., and R. D. TURNER. 1948. A catalogue
of the family Truncatellidae with notes and descriptions
of new species. Occasional Papers on Mollusks, Har-
vard University 1(13): 157-212.


CLENCH, W. J., and R. D. TURNER. 1948. The genus
Truncatella in the western Atlantic. Johnsonia 2(25):
149-164.

CLENCH, W. J., and R. D. TURNER. 1950. The genera
Sthenorytis, Cirsotrema, Acirsa, Opalia and Amaea in the
western Atlantic. Johnsonia 2(29): 221-246.

CLENCH, W. J., and R. D.TURNER. 1951.The genus Epi-
tonium in the western Atlantic: Part I. Johnsonia 2(30):
249-288.

CLENCH, W. J., and R. D. TURNER. 1952. The genera
Epitonium (Part II), Depressiscala, Cylindriscala, Nystiella
and Solutiscala in the western Atlantic. Johnsonia 2(31):
289-356.

CLENCH, W. J., and R. D. TURNER. 1953. The genera
Epitonium, Opalia, and Cylindriscala in the western At-
lantic. Johnsonia 2(32): 361-363.

CLENCH, W. J., and R. D. TURNER. 1956. The family
Melongenidae in the western Atlantic. Johnsonia 3(35):
161-188.

CLENCH, W. J., and R. D.TURNER. 1957.The family Cy-
matiidae in the western Atlantic. Johnsonia 3(36):
189-244.

CLENCH, W. J., and R. D.TURNER. 1960.The genus Cal-
liostoma in the western Atlantic. Johnsonia 4(40): 1-80.

COAN, E.V. 1997. Recent species of the genus Petricola
in the eastern Pacific (Bivalvia:Veneroida).TheVeliger
40(4): 298-340.

COAN, E.V., and P. H. SCOTT. 1997. Checklist of the ma-
rine bivalves of the northeastern Pacific Ocean. Santa
Barbara Museum of Natural History, Contributions
in Science No. 1: 1-28.

COLTON, H. S. 1905. Sexual dimorphism in Strombus
pugilis Linnaeus. The Nautilus 18(12): 138-140.

COLTON, H. S. 1905. Some notes on living Strombus
pugilis. The Nautilus 19(8): 85-88, pl. 3.

CONRAD, T. A. 1846. Observations on the geology of
a part of east Florida, with a catalogue of Recent shells
of the coast. American Journal of Science, Series 2,
2(4): 36-45.

CONRAD, T. A1846. Descriptions of new species of fos-


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Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


sil and Recent shells and corals. Proceedings of the
Academy of Natural Sciences of Philadelphia 3(1):
19-27, pls. 1, 2.

CONRAD, T. A. 1846. Catalogue of shells inhabiting
Tampa Bay and other parts of the Florida coast. Amer-
ican Journal of Science, Series 2, 2(6): 393-398.

CONRAD, T. A. 1848. Descriptions of two new genera
and new species of Recent shells, etc. Proceedings of
the Academy of Natural Sciences of Philadelphia 4(6):
121.

CONRAD, T. A. 1849. Descriptions of new fossil and Re-
cent shells of the United States. Journal of the Acad-
emy of Natural Sciences of Philadelphia, Series 2,1(4):
207-209.

CONRAD, T. A. 1862. Descriptions of new genera, sub-
genera and species of Tertiary and Recent shells. Pro-
ceedings of the Academy of Natural Sciences of
Philadelphia (1862): 284-291.

CONRAD, T. A. 1866. Descriptions of new marine bi-
valve Mollusca. American Journal of Conchology 2:
280, 281, pl. 15.

CONRAD,T. A. 1869. Notes on Recent Mollusca. Amer-
ican Journal of Conchology 5: 104-108, pls. 12, 13.

COOGAN, A. H. 1968. Bahamian and Floridian biofa-
cies. Pp. 141-154 in H. G. Multer, ed. Field Guide to some
Carbonate Rock Environments, Florida Keys and West-
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COOLEY, N. R. 1978. An inventory of the estuarine
fauna in the vicinity of Pensacola, Florida. Florida Ma-
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COOMANS, H. E. 1965. Teralatirus, a new genus in the
Fasciolariidae. Basteria 29(1-4): 10-14, pl. 1.

COOVERT, G. A. 1987. Proper usage of the generic
names Haloginella, Hyalina, and Volvarina with a diag-
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COOVERT, G. A. 1987. A literature review and sum-
mary of marginellid anatomy. Marginella Marginalia
3(2/3): 8-25.

COOVERT, G. A. 1987. Notes on the genus Granulina
and its type species. Marginella Marginalia 3(4): 26-32.


COOVERT, G. A. 1988. Marginellidae of Florida, Part
1: Granulina hadria. Marginella Marginalia 4(1): 1-7.

COOVERT, G. A. 1988. Marginellidae of Florida, Part
2: Prunum succinea with a discussion of Prunum and
Volvarina. Marginella Marginalia 4(5): 35-42.

COOVERT, G. A. 1989. A literature review and sum-
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COOVERT, G. A., and H. K. COOVERT. 1990. A study
of marginellid radulae. Part 1: type 6 radula,'Prunum/
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1-68.

COOVERT, G. A., and H. K. COOVERT. 1995. Revi-
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CORGAN, J. X. 1971. Review of Parodostomia, Telloda, Go-
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COSSIGNANI,T. 1994. Bursidae of the World. Mostra
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COSTA, F. H. A. 1994. On the Conusjaspideus complex
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COURTNEY, C. M. 1974. The marine macroinverte-
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COURTNEY, C. M. 1976. Mangrove and seawall oys-
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American Malacological Union, Inc. (1975): 29-32.

COURTNEY, C. M., andT. D. EDMOND. 1975.The ma-
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Deltona Corporation, Miami, Florida.

COURTNEY, C. M.,J. M. HATCHER, and P. G. GENOA.
1978.The occurrence of Tagelus plebeius in a manmade
brackish water lake with comments on its associates.
Bulletin of the American Malacological Union, Inc.
(1977): 68-75.

CRAIG, A. K., S. DOBKIN, R. B. GRIMM, and J. B.


46 FMRI Technical Report TR-3


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Mocroinvertebrate Checklists


Camp et a]. 1998








Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


DAVIDSON. 1969. The gastropod Siphonaria pectinata:
A factor in destruction of beach rock. American Zool-
ogist 9(3): 895-901.

CROVO, M. E. 1970. Gastrochaena hians Gmelin. Seafari
12(11): 6, 7.

CROVO, M. E. 1971. Cypraea cervus and Cypraea zebra
in Florida-One species or two? The Veliger 13(3):
292-295, 1 pl.

CROVO, M. E. 1971. A new Engina in Florida.TheVeliger
14(1): 30, 1 pl.

CROVO, M. E. 1974. Further notes and corrections con-
cerning the spawn of Florida Cyphoma (Ovulidae).The
Nautilus 88(2): 53-55.

CULTER, J. K. 1986. Manual for identification of marine
invertebrates. A guide to some common estuarine
macroinvertebrates of the Big Bend Region, Tampa
Bay, Florida. Environmental Monitoring and Support
Laboratory, U. S. EPA/600/4-86/002.

DALL, W. H. 1884. On a collection of shells sent from
Florida by Mr. Henry Hemphill. Proceedings of the
United States National Museum 6(384): 318-342, pl. 10.

DALL, W. H. 1885. Notes on some Floridian land and
fresh water shells with a revision of the Auriculacea of
the eastern United States. Proceedings of the United
States National Museum 8(17): 255-289, pls. 17, 18.

DALL, W. H. 1886. Reports on the results of dredging,
under supervision of Alexander Agassiz, in the Gulf of
Mexico (1877-78) and the Caribbean Sea (1879-80), by
the U. S. Coast Survey steamer "Blake", Lieut.-Com-
mander C. D. Sigsbee, U. S. N., and Commander J. R.
Bartlett, U. S. N., commanding. XXIX. Report on the
Mollusca. Part I. Brachiopoda and Pelecypoda. Bul-
letin of the Museum of Comparative Zoology at Har-
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DALL, W. H. 1889. Reports on the results of dredging,
under supervision of Alexander Agassiz, in the Gulf of
Mexico (1877-78) and the Caribbean Sea (1879-80), by
the U. S. Coast Survey steamer"Blake", Lieut.-Com-
mander C. D. Sigsbee, U. S. N., and Commander J. R.
Bartlett, U. S. N., commanding. XXIX. Report on the
Mollusca. Part II. Gastropoda and Scaphopoda. Bulletin
of the Museum of Comparative Zoology at Harvard
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DALL, W. H. 1889. A preliminary catalogue of the shell-


bearing marine mollusks and brachiopods of the south-
eastern coast of the United States, with illustrations of
many of the species. Bulletin of the United States Na-
tional Museum 37: 1-221, pls. 1-74.

DALL, W. H. 1895. An undescribed Meretrix from
Florida. The Nautilus 9(1): 10, 11.

DALL, W. H. 1895. Three new species of Macoma from
the Gulf of Mexico. The Nautilus 9(3): 32-34.

DALL, W. H. 1896. On the American species of Ervilia.
The Nautilus 10(3): 25-27.

DALL, W. H. 1896. On the American species of
Cyrenoidea.The Nautilus 10(5): 51, 52.

DALL, W. H. 1896.The mollusks and brachiopods of the
Bahama Expedition of the State University of Iowa. Bul-
letin of the Laboratory of Natural History, State Uni-
versity of Iowa 4(1): 12-27, pl. 1.

DALL, W. H. 1897. Synopsis of the Pinnidae of the
United States and West Indies.The Nautilus 11(3): 25,26.

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Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


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48 FMRI Technical Report TR-3


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Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


nean Society of London 44: 731-739.

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56-64.


FMRI Technical Report TR-3 49


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Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


FERREIRA, A. J. 1985. Chiton (Mollusca: Polypla-
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commercial hydraulic clam dredge on benthic com-


50 FMRI Technical Report TR-3


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Camp et al. 1998








Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


munities in estuarine areas. Florida Department of
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FMRI Technical Report TR-3 51


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Macroinvertebrate Checklists


W. G. Lyons--Mollusca







Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


onymy, Aphelodoris antillensis equals Chromodoris bis-
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52 FMRI Technical Report TR-3


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Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


vertebrates of the northeastern Gulf of Mexico. Mis-
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HENDERSON, J. B. 1920. A monograph of the east
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HENNIG, T., and J. HEMMEN. 1993. Ranellidae & Per-
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HENRY, C. A. 1976.The commensal clam, Paramya sub-
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HENRY, L. M. 1952. Observations on the sea hare Bur-
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HOAGLAND, K. E. 1979. The behavior of three sym-
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HOAGLAND, K. E. 1983. Ecology and larval develop-
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14-27; 88(2): 68.


FMRI Technical Report TR-3 53


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Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


HOUBRICK, R. S. 1980. Observations on the anatomy
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HOUBRICK, R. S. 1987. Anatomy of Alaba and Litiopa
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Florida State Board of Conservation Marine Laboratory
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INGHAM, R. E., and J. A. ZISCHKE. 1977. Prey pref-
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INTERNATIONAL COMMISSION ON ZOOLOGI-
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JENNER, C. E., and A. B. McCRARY. 1970. Paramya
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JENSEN, K. R. 1980. Oxynoe azuropunctata, n. sp., a new
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JENSEN, K. R., and K. B. CLARK. 1983. Annotated
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JOHNSON, C. W. 1911. Spondylus echinatus Martyn.
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JOHNSON, C. W. 1912. Notes on the variation of Strom-
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JOHNSON, C. W. 1913. Murexfulvescens Sowerby.The
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JOHNSON, C. W. 1919. An old collecting ground re-
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JOHNSON, C. W. 1919. Notes on the species of Fasci-
olaria of the southeastern United States. The Nautilus
33(2): 44-48.


54 FMRI Technical Report TR-3


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Camp et al. 1998 Macroinvertebrate Checklists W. 0. Lyons-Mollusca


JOHNSON, C. W. 1929.The egg-capsules of Fasciolaria
gigantea Kiener. The Nautilus 43(2): 63, pl. 4.

JOHNSON, C. W. 1931. Lima inflata and its nest. The
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JOHNSON, C. W. 1934. List of marine Mollusca of the
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JOHNSON, C. W., and H. A. PILSBRY. 1895. A re-
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JOHNSON, M. C. 1956. A living specimen of the east
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JOLLEY, J. W., JR. 1972. Exploratory fishing for the sun-
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JONES, D. S., L R. QUITMYER, W. S. ARNOLD, and D.
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KAAS, P. 1982. Ischnochiton dilatosculptus, a new species
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KAAS, P., and R. A.VAN BELLE. 1985. Monograph of
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KAAS, P., and R. A.VAN BELLE. 1985. Monograph of
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KAAS, P., and R. A.VAN BELLE. 1987. Monograph of
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KAAS, P., and R. A.VAN BELLE. 1990. Monograph of
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KAAS, P., and R. A.VAN BELLE. 1994. Monograph of
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KAICHER, S. D. 1982. Nassariidae, Part I. Card Cata-
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KAICHER, S. D. 1983. Nassariidae, Part II. Card Cata-
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KAICHER, S. D. 1984. Columbellidae, Part I. Card Cat-
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KAICHER, S. D. 1984. Columbellidae, Part II. Card Cat-
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KAICHER, S. D. 1984.Turridae, Part I. Card Catalogue
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KAICHER, S. D. 1985. Nassariidae, Part III. Card Cat-
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KAICHER, S. D. 1987. Olividae, Part II. Card Catalogue
of World-Wide Shells, No. 49: cards 4942-5046.


FMRI Technical Report TR-3 55


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W. G. Lyons--Mollusca







Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


KAICHER, S. D. 1988. Olividae, Part III. Card Cata-
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KAICHER, S. D. 1989. Olividae, Part IV. Card Cata-
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KRAKAUER, J. M. 1969.The Ecology ofAplysia willcoxi
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LENDERKING, R. E. 1954. Some recent observations
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LERMOND, N.W. 1927. Collecting in Florida.The Nau-
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LEWIS, F. G., III, and A. W. STONER. 1981. An exam-
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LIGHTFOOT, J. 1992. Caecidae of the western Atlantic.


56 FMRI Technical Report TR-3


Mollusca--W. G. Lyons


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Camp et al. 1998








Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


Of Sea and Shore 14(4): 171-185.

LIGHTFOOT, J. 1992. Caecidae of the western Atlantic,
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LIPE, R. 1984. Tampa Bay shells. American Concholo-
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LYMAN, F. B. 1936. Discovery and observation of liv-
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LYMAN, F. B. 1937. Conus mus Hwass in Lake Worth,
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LYMAN, F. B. 1938. Hydatina physis (L.) in Lake Worth,
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LYMAN, F. [B.] 1942. Dredging at Destin. Shell Notes
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LYONS, W. G. 1972. New Turridae (Gastropoda: Tox-
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LYONS, W. G. 1976. Distribution of Cerithium litteratum


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LYONS, W. G. 1977. Comments on three Jamaican
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LYONS, W. G. 1977. Notes on occurrence of Eupleura sul-
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LYONS, W. G. 1978. Status of Eulima subcarinata Or-
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LYONS, W. G. 1978. Status of the genus Oceanida De-
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LYONS, W. G. 1980. Molluscan communities of the
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LYONS, W. G. 1981. Polyplacophora of Dry Tortugas,
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LYONS, W. G. 1985. Chaetopleura staphylophera (Poly-
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LYONS, W. G. 1988. A review of Caribbean Acantho-
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cata (Conrad). Pp. 48-51 in M. Deyrup and R. Franz, eds.


FMRI Technical Report TR-3 57


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Mollusca-W. G. Lyons Macroinverlebrate Checklists Camp et al. 1998


Rare and Endangered Biota of Florida,Vol. IV. Inver-
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LYONS, W. G., S. P. COBB, D. K. CAMP, J. A. MOUN-
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LYONS, W. G., and S. B. COLLARD. 1974. Benthic in-
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LYONS, W. G., and J. F. QUINN, JR. 1995. Appendix J.
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MARCUS, EV. Du B.-R. 1972. On some opisthobranchs
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MARCUS, EV. Du B.-R. 1972. On the Anaspidea (Gas-
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MARCUS, EV. Du B.-R. 1973. On the genus Bosellia
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MARCUS, EV. Du B.-R. 1974. On some Cephalaspidea
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MARCUS, EV. Du B.-R. 1976. Opisthobranchia von
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MARCUS, EV. Du B.-R. 1977. On the genus Tornatina
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MARCUS, EV. Du B.-R. 1977. An annoted checklist of
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MARCUS, EV. Du B.-R. 1978. The western Atlantic
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MARCUS, EV. Du B.-R. 1979. The Atlantic species of
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MARCUS, EV. Du B.-R. 1980. Review of the western At-
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MARCUS, EV. Du B.-R. 1982. Systematics of the gen-
era of the order Ascoglossa (Gastropoda). Journal of


58 FMRI Technical Report TR-3


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Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


Molluscan Studies, Supplement 10: 1-31.

MARCUS, EV. Du B.-R. 1983.The western Atlantic Tri-
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MARCUS, EV. Du B.-R. 1984. The western Atlantic
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MARCUS, EV. Du B.-R. 1985. Subclass Opisthobranchia.
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MARCUS, EV. Du B.-R. 1985. The western Atlantic
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MARCUS, EV. Du B.-R., and S. B. GALLAGHER. 1976.
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MARCUS, EV. Du B.-R., andT. M. GOSLINER. 1984. Re-
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MARCUS, EV. Du B.-R., and ER. MARCUS. 1960.
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MARCUS, EV. Du B.-R., and ER. MARCUS. 1962.
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MARCUS, EV. Du B.-R., and ER. MARCUS. 1963.
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MARCUS, EV. Du B.-R., and ER. MARCUS. 1964. On
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MARCUS, EV. Du B.-R., and ER. MARCUS. 1967. Amer-
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MARCUS, EV. Du B.-R., and ER. MARCUS. 1967.
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MARCUS, EV. Du B.-R., and ER. MARCUS. 1967. Some
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MARELLI, D. C., and S. GRAY. 1983. Conchological
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MARELLI, D. C., and S. GRAY. 1985. Comments on
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MARELLI, D. C, W. G. LYONS, W. S. ARNOLD, and M.
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MASON, W. T., JR., and S. A. ZENGEL. 1996. Foods of


FMRI Technical Report TR-3 59


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W. G. Lyons--Mollusca








Mollusca-W. 0. Lyons Macroinvertebrate Checklists Camp et al. 1998


juvenile spotted seatrout in seagrasses at Seahorse
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McGINTY, T. L. 1937. A double-lipped Strombus. The
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McGINTY, T. L. 1937.Acanthochitona pygmaea (Pilsbry).
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McGINTY, T. L1939. Terebraflammea, a new record for
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McGINTY, T. L. 1939. Collecting on a coral reef in
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McGINTY,T. L. 1939. Littorina minima (Wood).The Nau-
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McGINTY, T. L. 1940. A new naticoid snail from Florida.
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McGINTY, T. L. 1941. Solarium bisulcatum Orbigny,


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McGINTY, T. L. 1946. A new Florida strombus, Strom-
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McLEAN, R. A. 1941.The oysters of the western Atlantic.
Notulae Naturae No. 67: 1-14.


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Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


McLEAN, R. A., and J. S. SCHWENGEL. 1944. A new
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Nautilus 97(3): 93-98.


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Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


MOOK, D. 1977. Studies on fouling invertebrates in the
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MOORE, D. R. 1956. Observations of predation on
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MOORE, D. R. 1957. A note on Cuna dalli.The Nautilus
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MORRISON, J. P. E. 1951. Two new western Atlantic
species of pulmonate mollusks of the genera Detracia


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Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


and two old ones (family Ellobiidae). Journal of the
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MORRISON, J. P. E. 1967. Notes on American Hastula.
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guide to the larger marine gastropods of Florida, the


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W. G. Lyons--Mollusca







Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


Gulf of Mexico, and the Caribbean region. University
of Miami Sea Grant Field Guide Series No. 5: 1-55.

ORTEA, J., A. A. LUQUE, and J. TEMPLADO. 1988.
Elysia picta Verrill, 1901, and Geitodoris pusae (Marcus,
1955), two amphiatlantic opisthobranch gastropods.
Journal of Molluscan Studies 54(2): 243-247.

ORTEA, J., A. A. LUQUE, and J. TEMPLADO. 1990.
Contributions to the knowledge of the genus Aegires
Lov6n, 1844 (Opisthobranchia: Doridoidea: Aegiretidae)
in the north Atlantic, with descriptions of two new
species. Journal of Molluscan Studies 56(3): 333-337.

ORTEA, J., and J. TEMPLADO. 1989. A new species of
Cyerce Bergh, 1871 (Opisthobranchia: Ascoglossa) from
Cuba. Iberus 8: 11-14.

ORTEA, J., and J. TEMPLADO. 1990. A new species of
the genus Cyerce Bergh, 1871, from the CapeVerde Is-
lands (Opisthobranchia: Ascoglossa).TheVeliger 33(2):
202-205.

ORTEA, J., A.VALDES, and J. ESPINOSA. 1994. North
Atlantic nudibranchs of the Chromodoris clenchi colour
group (Opisthobranchia: Chromodorididae). Journal of
Molluscan Studies 60(3): 237-248.

ORTIZ TOUZET, M., and R. LALANA RUEDA. 1984.
Primer hallazgo de la clase Aplacophora (Mollusca) en
aguas someras cubanas. Revista de Investigaciones
Marinas 5(3): 25-28.

PAIGE, J. A. 1973. The Nudibranchs of Cedar Key,
Florida, With Special Reference to the Life History
and Ecology of Hypselodoris edenticulata (White, 1952).
M. Sc.Thesis, University of Florida, Gainesville. 119 pp.

PAINE, R.T. 1962. Ecological diversification in sympatric
gastropods of the genus Busycon. Evolution 16(4):
515-523.

PAINE, R.T. 1962. Reproduction of Olivella mutica.The
Nautilus 75(4): 139-142.

PAINE, R.T. 1963.Trophic relationships of 8 sympatric
predatory gastropods. Ecology 44(1): 63-73.

PAINE, R. T. 1963. Feeding rate of a predaceous gas-
tropod, Pleuroploca gigantea. Ecology 44(2): 402, 403.

PALMER, K.V. W. 1947. Notes on Costacallista eucymata
(Dall). The Nautilus 61(2): 44-47, pl. 4.


PENZIAS, L. P. 1969. Tellina martinicensis (Mollusca:
Bivalvia): Biology and productivity. Bulletin of Marine
Science 19(3): 568-579.

PtREZ FARFANTE, I. 1943. The genera Fissurella, Lu-
capina and Lucapinella in the western Atlantic. Johnsonia
1(10): 1-20.

PEREZ FARFANTE, I. 1943. The genus Diodora in the
western Atlantic. Johnsonia 1(11): 1-20.

PtREZ FARFANTE, I. 1945. The genera Lucapina and
Diodora in the western Atlantic. Johnsonia 1(18): 4-6.

PtREZ FARFANTE, I. 1947.The genera Zeidora, Nesta,
Emarginula, Rimula and Puncturella in the western At-
lantic. Johnsonia 2(24): 93-148.

PERRY, L. M. 1936. A marine tenement. Science
84(2172): 156, 157.

PERRY, L. M. 1939. Conus melvilli Sowerby, and a new
species of the Lamellariidae.The Nautilus 53(2): 40-42,
pl. 8.

PERRY, L. M. 1940. Marine shells of the southwest
coast of Florida. Bulletins of American Paleontology
26(95): 1-260, pls. 1-40.

PERRY, L. M. 1940. A new tellin of the subgenus Angulus,
a new Lamellaria and a new subspecies of Crassispira,
from southwest Florida. The Nautilus 53(3): 79-81.

PERRY, L. M., and J. S. SCHWENGEL. 1955. Marine
Shells of the Western Coast of Florida. Paleontological
Research Institution, Ithaca, NewYork. 318 pp., 55 pls.

PERRY, L. M., J. S. SCHWENGEL, and T. DRANGA.
1938. Note on unreported marine molluscs from Sani-
bel, Florida. The Nautilus 52(1): 27-29.

PETERSON, R. 1971. Collecting at Palm Beach Inlet.
Seafari 13(6): 10.

PETIT, R. E., and M. G. HARASEWYCH. 1990. Cata-
logue of the superfamily Cancellarioidea Forbes and
Hanley, 1851 (Gastropoda: Prosobranchia). The Nau-
tilus 103(supplement 1): 1-69.

PETIT, R. E., and D. WILSON. 1986. Publication dates
of Heilprin's explorations on the west coast of Florida.
Tulane Studies in Geology and Paleontology 19(2): 95,
96.


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Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


PETUCH, E. J. 1980. A new species of Conus from south-
eastern Florida (Mollusca: Gastropoda). Proceedings
of the Biological Society of Washington 93(2): 299-302.

PETUCH, E. J. 1982. Geographical heterochrony: Con-
temporaneous coexistence of Neogene and Recent
molluscan faunas in the Americas. Palaeogeography,
Palaeoclimatology, Palaeoecology 37: 277-312.

PETUCH, E.J. 1982. Paraprovincialism: Remnants of pa-
leoprovincial boundaries in Recent marine molluscan
provinces. Proceedings of the Biological Society of
Washington 95(4): 774-780.

PETUCH, E. J. 1987. New Caribbean Molluscan Faunas.
Coastal Education and Research Foundation, Char-
lottesville, Virginia. 154 pp. + 4 appendix pp.

PETUCH, E. J. 1988. Neogene History of Tropical Amer-
ican Mollusks: Biogeography and Evolutionary Pat-
terns of Tropical Western Atlantic Mollusca. Coastal Ed-
ucation and Research Foundation, Charlottesville,Vir-
ginia. 217 pp.

PETUCH, E. J. 1995. Molluscan discoveries from the
tropical western Atlantic region. La Conchiglia 27(275):
36-41.

PETUCH, E.J., and D. M. SARGENT. 1986. Atlas of the
Living Olive Shells of the World. Coastal Education and
Research Foundation, Charlottesville,Virginia. 253 pp.

PIECH, B. J. 1993. New records for ranellid gastropods
in the western Atlantic (Ranellidae: Cymatiinae). The
Veliger 36(1): 88-91.

PIEL, W. H. 1991. Pycnogonid predation on nudibranchs
and ceratal autotomy. The Veliger 34(4): 366, 367.

PILSBRY, H. A. 1888. Monographs of the Turbinidae and
Trochidae. Manual of Conchology 10(2): 161-323, pls.
31-69.

PILSBRY, H. A. 1889. FamilyTrochidae. Manual of Con-
chology 11: 1-519, pls. 1-67.

PILSBRY, H. A. 1890. Monographs of the families Sto-
matellidae, Scissurellidae, Haliotidae, etc. Manual of
Conchology 12: 1-323, pls. 1-65.

PILSBRY, H. A. 1891. Monographs of the Acmaeidae,
Lepetidae, Patellidae andTitiscaniidae. Manual of Con-
chology 13: 1-195, pls. 1-74.


PILSBRY, H. A. 1891. Acmaea candeana vs. Acmaea an-
tillarum. The Nautilus 5(8): 85, 86.

PILSBRY, H. A. 1892. Polyplacophora (Chitons). Lepi-
dopleuridae, Ischnochitonidae, Chitonidae, Mopali-
idae. Manual of Conchology 14: i-xxxiv, 1-128, pls.
1-30.

PILSBRY, H. A. 1893. Polyplacophora (Chitons). Lepi-
dopleuridae, Ischnochitonidae, Chitonidae, Mopali-
idae. Manual of Conchology 14: 129-350, pls. 31-68.

PILSBRY, H. A. 1893. Order Opisthobranchiata. Man-
ual of Conchology 15: 134-436.

PILSBRY, H. A. 1893. Notes on the Acanthochitidae
with descriptions of new American species. The Nau-
tilus 7(3): 31,32.

PILSBRY, H. A. 1893. Polyplacophora (Chitons). Acan-
thochitidae, Cryptoplacidae, and Appendix. Manual of
Conchology 15:1-64, pls. 1-10.

PILSBRY, H. A. 1894. Polyplacophora (Chitons). Acan-
thochitidae, Cryptoplacidae, and Appendix. Manual of
Conchology 15: 65-132, pls. 11-17.

PILSBRY, H. A. 1895. Philinidae, Gastropteridae, Aglaj-
idae, Aplysiidae, Oxynoeidae, Runcinidae, Umbrac-
ulidae, Pleurobranchidae. Manual of Conchology 16:
i-vii, 1-262, pls. 1-74.

PILSBRY, H. A. 1896. Descriptions of new species of
mollusks. Proceedings of the Academy of Natural Sci-
ences of Philadelphia 48: 21-24.

PILSBRY, H. A. 1898. Order Aplacophora. Manual of
Conchology 17: 281-310, 346-48, pls. 40-48.

PILSBRY, H. A. 1900. A new Calliostoma from Florida.
The Nautilus 13(11): 128, 129.

PILSBRY, H. A. 1902. Fasciolaria gigantea, subsp. reevei.
Proceedings of the Academy of Natural Sciences of
Philadelphia (1902): 552.

PILSBRY, H. A. 1905. Land shells of the Florida Keys.
The Nautilus 19(4): 37-41.

PILSBRY, H. A. 1922. Gemma gemma purpurea (Lea).
The Nautilus 36(1): 32.

PILSBRY, H. A. 1936. Melongena corona (Gmelin). The
Nautilus 50(1): 29.


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Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


PILSBRY, H. A. 1938. Dimensions of Florida Chamas.
The Nautilus 51(4): 143.

PILSBRY, H. A. 1939. A triad of umbilicate Latirus, Re-
cent and Pliocene. The Nautilus 52(3): 84-86, pl. 5.

PILSBRY, H. A. 1939. Cyphoma mcgintyi, new species.The
Nautilus 52(3): 108.

PILSBRY, H. A. 1939. A new Floridan Cyclostrema.The
Nautilus 53(2): 53, pl. 8.

PILSBRY, H. A. 1940. A new race of Cancellaria from
Floridd. The Nautilus 54(2): 54, pl. 3.

PILSBRY, H. A. 1943. Florida species of Rimula. The
Nautilus 57(2): 37-40, pl. 7.

PILSBRY, H. A. 1945. Stenacmefloridana, an American
member of the Amphibolacea. The Nautilus 58(4):
112-116, pl. 5.

PILSBRY, H. A. 1945. New Floridian marine mollusks.
The Nautilus 59(2): 59, 60, pl. 6.

PILSBRY, H. A. 1946. The type specimens of C. B.
Adams's Jamaican species of Vitrinella. Notulae Natu-
rae No. 162: 1-5.

PILSBRY, H. A. 1949. A West Indian Hydatina.The Nau-
tilus 63(1): 15-17, pl. 1.

PILSBRY, H. A. 1949. New Cerithiidae from Florida. The
Nautilus 63(2): 65, 66, pl. 1.

PILSBRY, H. A. 1951. Aplysia badistes, a peculiar Flori-
dan sea-hare. Notulae Naturae No. 240: 1-6.

PILSBRY, H. A. 1955. Another Florida Conus.The Nau-
tilus 69(2): 47,48, pl. 3.

PILSBRY, H. A. 1956. A gastropod domiciliary in sea
urchin spines. The Nautilus 69(4): 109, 110, pl. 6.

PILSBRY, H. A., and C. W. JOHNSON. 1892. Catalogue
of Fissurellidae of the United States.The Nautilus 5(9):
102-107.

PILSBRY, H. A., and T. L. McGINTY. 1938. Review of
Florida Chamidae. The Nautilus 51(3): 73-79, pl. 7.

PILSBRY, H. A., and T. L. McGINTY. 1939. Small Cas-
sididae of Florida and the West Indies. The Nautilus
52(3): 73-77, pl. 5.


PILSBRY, H. A., and T. L. McGINTY. 1939. The genus
Cyphoma in Florida. The Nautilus 53(1): 1-4, pl. 1.

PILSBRY, H. A., and T. L. McGINTY. 1943. Ensis minor
megistus n. subsp., a west Florida razor clam. The Nau-
tilus 57(1): 33,34.

PILSBRY, H. A., and T. L. McGINTY. 1945. Cy-
clostrematidae and Vitrinellidae of Florida-I. The
Nautilus 59(1): 1-13, pls. 1, 2.

PILSBRY, H. A., and T. L. McGINTY. 1945. "Cy-
clostrematidae" and Vitrinellidae of Florida-II. The
Nautilus 59(2): 52-59, 72 (addendum), pl. 6.

PILSBRY, H. A., and T. L. McGINTY. 1946. "Cy-
clostrematidae"andVitrinellidae of Florida, Part III.The
Nautilus 59(3): 77-83, pl. 8.

PILSBRY, H. A., andT. L. McGINTY. 1946.Vitrinellidae
of Florida, Part 4. The Nautilus 60(1): 12-18, pls. 1, 2.

PILSBRY, H. A., andT. L. McGINTY. 1946. Note on Vit-
rinella (Tomura) bicaudata. The Nautilus 60(1): 36.

PILSBRY, H. A., andT. L. McGINTY. 1949. New marine
mollusks of Florida and the Bahamas. The Nautilus
63(1): 9-15, pl. 1.

PILSBRY, H. A., andT. L. McGINTY. 1950.Vitrinellidae
of Florida, Part 5. The Nautilus 63(3): 85-87, pl. 5.

PILSBRY, H. A., andT. L. McGINTY. 1952. Notes on Nesta
(Laevinesta) atlantica, a Floridan fissurellid mollusk.
The Nautilus 66(1): 1-3.

PILSBRY, H. A., and R. A. McLEAN. 1939. A new Arca
from the West Indian region. Notulae Naturae No. 39:
1,2.

PILSBRY, H. A., and A. A. OLSSON. 1946. Condylocar-
dia in Florida and Middle America.The Nautilus 60(1):
6, 7, pl. 1.

PILSBRY, H. A., and A. A. OLSSON. 1950. Review of An-
ticlimax, with new Tertiary species (Gastropoda, Vit-
rinellidae). Bulletins of American Paleontology 33(135):
103-124, pls. 17-20.

PILSBRY, H. A., and A. A. OLSSON. 1953. Materials for
a revision of east coast and Floridian volutes.The Nau-
tilus 67(1): 1-13, pls. 1-3.

PILSBRY, H. A., and B. SHARP. 1897-98. Class Scapho-


66 FMRI Technical Report TR-3


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Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


poda. Manual of Conchology 17: v-xxxii, 1-280,310,311,
337-346, pls. 1-39.

PILSBRY, H. A., and E. G.VANATTA. 1934. Melongena
corona and its races.The Nautilus 47(4): 117-121, pl. 12.

PLOCKELMAN, C. 1965. Encounter between Murex
pomum Gmelin and Terebra taurina Solander. Seafari
7(10): 6.

PLOCKELMAN, C. 1965. Sequel to attack ofM. pomum
on T. taurinus. Seafari 7(11): 8.

PLOCKELMAN, C. 1970. Shells of Palm Beach County,
#23:Seafari 12(6): 2-4.

PLOCKELMAN, C. 1970.The wind vs... Bear Cut. Sea-
fari 12(7): 13, 14.

PLOCKELMAN, C. 1970. Finger Channels luck. Seafari
12(10): 1, 2.

PLOCKELMAN, C. 1970. Spondylus: Home, nursery,
trash can. Seafari 12(11):1-5.

PLOCKELMAN, C. 1970. Mullet Key revisited. Seafari
12(12): 14, 15.

PLOCKELMAN, C. 1971. Estero Island field trip. Sea-
fari 13(1): 3, 4.

PLOCKELMAN, C. 1971. January-Bear Cut. Seafari
13(2): 9.

PONDER, W. F. 1973. The origin and evolution of the
Neogastropoda. Malacologia 12: 295-338.

PONDER, W. F. 1983. Review of the genera of the Bar-
leeidae (Mollusca: Gastropoda: Rissoacea). Records of
the Australian Museum 35: 231-281.

PONDER, W. F. 1983. Reclassification of some Ameri-
can species assigned to the Rissoidae (sensu lato). The
Nautilus 97(3): 90-91.

PONDER, W. F. 1985. The anatomy and relationships
of Elachisina Dall (Gastropoda: Rissoacea). Journal of
Molluscan Studies 51(1): 23-34.

PONDER, W. F. 1985. A review of the genera of the Ris-
soidae (Mollusca: Mesogastropoda: Rissoacea). Records
of the Australian Museum, Supplement 4: 1-221.

PONDER, W. F. 1987. The anatomy and relationships


of the pyramidellacean limpet Amathina tricarinata
(Mollusca: Gastropoda). Asian Marine Biology 4:1-34.

PONDER, W. F. 1990. The anatomy and relationships
of the Orbitestellidae (Gastropoda: Heterobranchia).
Journal of Molluscan Studies 56(4): 515-532.

PONDER, W. F. 1990. The anatomy and relationships
of a marine valvatoidean (Gastropoda: Heterobranchia).
Journal of Molluscan Studies 56(4): 533-555.

PONDER, W. F., and E. H.VOKES. 1988. A revision of
the Indo-West Pacific fossil and Recent species of
Murex s.s. and Haustellum (Mollusca: Gastropoda: Mu-
ricidae). Records of the Australian Museum, Supple-
ment 8: 1-160.

PONDER, W. F., and A. WAREN. 1988. Classification of
the Caenogastropoda and Heterostropha-a list of the
family-group names and higher taxa. Malacological Re-
view, Supplement 4: 288-328.

POPE, V. A. 1955. The geoduck clam in Florida. Quar-
terly Journal of the Florida Academy of Sciences (1954)
17(4): 252.

POPPE, G.T., andY. GOTO. 1992.Volutes. Mostra Mon-
diale Malacologia, Ancona, Italy. 348 pp.

POST, E. J. 1899. Collecting on the Gulf coast of Florida.
The Nautilus 13(5): 52-54.

POWELL, E. N. 1981.Three Turbonilla (Pyramidellidae,
Gastropoda) of North Carolina, with comments on
pyramidellid systematics. The Journal of the Elisha
Mitchell Scientific Society 97(1): 37-54.

PRESBREY, E.W. 1913. Concerning Cypraea exanthema,
cervus and cervinetta. The Nautilus 27(1): 8-11.

PREZANT, R. S. 1981. Taxonomic re-evaluation of the
bivalve family Lyonsiidae. The Nautilus 95(2): 58-72.

PUFFER, E. L., and W. K. EMERSON. 1954. Catalogue
and notes on the gastropod genus Busycon. Proceed-
ings of the Biological Society of Washington 67(13):
115-150.

PULLEY,T. E. 1959. Busycon perversum (Linn6) and some
related species. The Rice Institute Pamphlet 46(1):
70-89.

QUINN, J. F., JR. 1979. Biological results of the University
of Miami deep-sea expeditions. 130. The systematics


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Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


and zoogeography of the gastropod family Trochidae
collected in the Straits of Florida and its approaches.
Malacologia 19(1): 1-62.

QUINN, J. F., JR. 1992. New species of Calliostoma
Swainson, 1840 (Gastropoda: Trochidae).The Nautilus
106(3): 77-114.

RADWIN, G. E. 1968.The systematic position of Glyptae-
sopus.The Nautilus 82(1): 18, 19.

RADWIN, G. E. 1977.The family Columbellidae in the
western Atlantic. The Veliger 19(4): 403-417, 2 pls.

RADWIN, G. E. 1977.The family Columbellidae in the
western Atlantic. Part IIa.-The Pyreninae.TheVeliger
20(2): 119-133, 4 pls.

RADWIN, G. E. 1978.The family Columbellidae in the
western Atlantic. Part IIb.-The Pyreninae (contin-
ued). The Veliger 20(4): 328-344, 4 pls.

RADWIN, G. E., and A. D'ATTILIO. 1976. Murex Shells
of the World: An Illustrated Guide to the Muricidae.
Stanford University Press, Stanford, California. 284
pp., 32 pls.

RADWIN, G. E., and H. W. WELLS. 1968. Comparative
radular morphology and feeding habits of muricid
gastropods from the Gulf of Mexico. Bulletin of Marine
Science 18(1): 72-85.

RAEIHLE, D. 1967. Notes on captive Leucozonia nassa
Gmelin, Chaetopleura apiculata Say, and Ischnochiton
floridanus Pilsbry. American Malacological Union, Inc.,
Annual Report (1967): 13, 14.

RAEIHLE, D. 1975. Remarks on western Atlantic
Anachis. Bulletin of the American Malacological Union,
Inc. (1974): 40,41.

RAKOCINSKI, C. F., R. W. HEARD, S. E. LECROY,J. A.
McLELLAND, andT. SIMONS. 1993. Seaward change
and zonation of the sandy-shore macrofauna at Perdido
Key, Florida, U.S.A. Estuarine, Coastal and Shelf Sci-
ence 36: 81-104.

RAKOCINSKI, C. F., R. W. HEARD, S. E. LECROY, J. A.
McLELLAND, and T. SIMONS. 1996. Responses by
macrobenthic assemblages to extensive beach restora-
tion at Perdido Key, Florida, U.S.A. Journal of Coastal
Research 12(1): 326-353.


REED, J. K., and P. M. MIKKELSEN. 1987.The mollus-
can community associated with the scleractinian coral
Oculina varicosa. Bulletin of Marine Science 40(1):
99-131.

REHDER, H. A. 1935. New Caribbean marine shells.The
Nautilus 48(4): 127-130, pl. 7.

REHDER, H. A. 1939. New marine mollusks from the
west Atlantic. The Nautilus 53(1): 16-21, pl. 6.

REHDER, H. A. 1940. Notes on Thericium auricoma
Schwengel and stantoni Dall.The Nautilus 54(2): 72,73.

REHDER, H. A. 1943. New marine mollusks from the
Antillean Region. Proceedings of the United States
National Museum 93(3161): 187-203, pls. 19, 20.

REHDER, H. A. 1943. Corrections and ecological notes
on some recently described Florida marine shells.The
Nautilus 57(1): 32, 33.

REHDER, H. A. 1944. A new subspecies of Conus ver-
rucosus Hwass from Florida.The Nautilus 57(3): 105,106.

REHDER, H. A. 1945. Two new species of Cirsostrema
(Epitoniidae) from Florida. Proceedings of the Biolog-
ical Society of Washington 58(31): 127-129.

REHDER, H. A. 1947. A new species of Naticarius from
Florida. The Nautilus 61(1): 19, 20, pl. 1.

REHDER, H. A. 1954. Mollusks. Pp. 469-474 in P. S.
Galtsoff, ed. Gulf of Mexico, its origin, waters, and ma-
rine life. United States Department of the Interior,
Fishery Bulletin 89.

REHDER, H. A. 1981.The Audubon Society Field Guide
to North American Seashells. Alfred A. Knopf, Inc.,
NewYork. 894 pp.

REHDER, H. A. 1986. Natica (Glyphepithema) bayeri,
new name. The Nautilus 100(1): 38.

REHDER, H. A. 1986. Natica (Glyphepithema) tedbayeri,
a replacement name.The Nautilus 100(3): 112.

REHDER, H. A., and R.T. ABBOTT. 1951. Some new and
interesting mollusks from the deeper waters of the
Gulf of Mexico. Revista de la Sociedad Malacologica
"Carlos de la Torre"8(2): 53-68, pls. 8.

REID, D. G. 1986.The Littorinid Molluscs of Mangrove


68 FMRI Technical Report TR-3


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Camp et al, 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


Forests in the Indo-Pacific Region; the Genus Littoraria.
British Museum (Natural History), London. xv + 228 pp.,
1 pl.

RICE, S. A., G. W. PATTON, and S. MAHADEVAN.
1981. An ecological study of the effects of offshore
dredge material disposal with special reference to
hard-bottom habitats in the eastern Gulf of Mexico. A
technical report to Manatee County Chamber of Com-
merce, Bradenton, Florida. Mote Marine Laboratory,
Sarasota, Florida. 45 pp.

RICHARDS, B. R., and W. F. CLAPP. 1944. A prelimi-
nary report on the fouling characteristics of Ponce de
Leon Tidal Inlet, Daytona Beach, Florida. Journal of
Marine Research 5(3): 189-195.

RICHARDS, H. G. 1933. New varieties of Melongena and
Gyrineum.The Nautilus 47(2): 57, pl. 6.

ROBERTSON, R. 1957. A study of Cantharus multangu-
lus (Philippi), with notes on Cantharus and Pseudonep-
tunea (Gastropoda: Buccinidae). Notulae Naturae No.
300: 1-10.

ROBERTSON, R. 1958.The family Phasianellidae in the
western Atlantic. Johnsonia 3(37): 245-283.

ROBERTSON, R. 1959.The family Phasianellidae in the
western Atlantic. Johnsonia 3(39): 344-346.

ROBERTSON, R. 1961. A second western Atlantic Ris-
soella and a list of the species in the Rissoellidae. The
Nautilus 74(4): 131-136, pl. 9; 75(1): 21-26.

ROBERTSON, R. 1963. Bathymetric and geographic
distribution of Panopea bitruncata. The Nautilus 76(3):
75-82.

ROBERTSON, R. 1973. Cyclostremella: A planispiral
pyramidellid. The Nautilus 87(3): 88.

ROBERTSON, R. 1978. Spermatophores of six eastern
North American pyramidellid gastropods and their
systematic significance (with the new genus Boonea).
Biological Bulletin 155(2): 360-382.

ROBERTSON, R. 1985. Archaeogastropod biology and
the systematics of the genus Tricolia (Trochacea: Tri-
coliidae) in the Indo-West-Pacific. Monographs of Ma-
rine Mollusca 3: 1-103.

ROBERTSON, R. 1994. Two new tropical western At-
lantic species of Epitonium, with notes on similar global


species and natural history.The Nautilus 107(3): 81-93.

ROBERTSON, R. 1996. Fargoa bartschi (Winkley, 1909):
A little-known Atlantic and Gulf coast American odos-
tomian (Pyramidellidae) and its generic relationships.
American Malacological Bulletin 13(1/2): 11-21.

ROBERTSON, R., andT. HABE. 1965.Alexania replaces
Habea (Epitoniidae). The Nautilus 78(4): 140, 141.

ROBERTSON, R, andT. MAU-LASTOVICKA. 1979.The
ectoparasitism of Boonea and Fargoa (Gastropoda: Pyra-
midellidae). Biological Bulletin 157(2): 320-333.

ROBERTSON, R., and K. OYAMA. 1958. The family
Stenacmidae. The Nautilus 72(2): 68, 69.

ROLAN, E. 1992.The family Omalogyridae G. O. Sars,
1878 (Mollusca, Gastropoda) in Cuba with description
of eight new species. Apex 7(2): 35-46.

ROLAN, E., and J. ESPINOSA. 1992. La familiar
Cerithiopsidae H. y A. Adams, 1853 (Mollusca, Gas-
tropoda), en la Isla de Cuba. 1. El genero Retilaskeya
Marshall, 1978. Publicacoes Ocasionis da Sociedad
Portuguesa de Malacologia 16: 39-43, 1 pl.

ROLAN, E., and J. ESPINOSA. 1992. La familiar
Cerithiopsidae H. y A. Adams, 1853 (Mollusca, Gas-
tropoda), en la Isla de Cuba. 2. El genero Horologica
Laseron 1956. Publicacoes Ocasionis da Sociedad Por-
tuguesa de Malacologia 16: 45-49,1 pl.

ROLAN, E., and J. ESPINOSA. 1994.The familyTriphor-
idae (Mollusca, Gastropoda) in Cuba. 3.The genus Iso-
triphora. Basteria 58(1-2): 63-68.

ROLAN, E., and R. FERNANDEZ-GARCES. 1993. La fa-
milia Triphoridae en la isla de Cuba. 1. El genero
Metaxia Monterosato, 1884. Bollettino Malacologico
28(6-12): 169-176.

ROLAN, E., and R. FERNANDEZ-GARCtS. 1993.The
familyTriphoridae (Mollusca, Gastropoda) in Cuba. 2.
The genus Iniforis Jousseaume, 1884. Apex 8(3): 95-106.

ROLAN, E., and R. FERNANDEZ-GARCES. 1994.The
familyTriphoridae (Mollusca, Gastropoda) in Cuba. 4.
The genera Monophorus, Nototriphora, Cosmotriphora
and Cheirodonta with the description of three new
species. Apex 9(1): 17-27.

ROLAN, E., and R. FERNANDEZ-GARCtS. 1995.The
family Triphoridae (Mollusca, Gastropoda) in Cuba. 5.


FMRI Technical Report TR-3 69


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W. G. Lyons--Mollusca








Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


The genera Marshallora, Mesophora, Similiphora, Eu-
triphora, Latitriphora, Aclophora and other species with-
out generic affiliation. Apex 10(1): 9-24.

ROOPNARINE, P. D. 1996. Systematics, biogeography
and extinction of chionine bivalves (Bivalvia:Veneridae)
in tropical America early Oligocene-Recent. Mala-
cologia 38(1-2): 103-142.

ROSENBERG, G. 1992.The Encyclopedia of Seashells.
Michael Friedman Publishing Group, Inc., NewYork.
224 pp.

ROSENBERG, G. 1996. Independent evolution of ter-
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ROSEWATER, J. 1978. The zoogeography of West
African Littorinidae. Bulletin of the American Mala-
cological Union, Inc. (1977): 31-34.

ROSEWATER, J. 1980. Subspecies of the gastropod Lit-
torina scabra. The Nautilus 94(4): 158-162.

RUPPERT, E. E., and R. S. FOX. 1988. Seashore Animals
of the Southeast: A Guide to Common Shallow-Water
Invertebrates of the Southeastern Atlantic Coast. Uni-
versity of South Carolina Press, Columbia, South Car-
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RUSSELL, H. D. 1941.The Recent mollusks of the fam-
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RUSSELL, H. D. 1971. Index Nudibranchia; a Cata-
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SALOMAN, C. H. 1976. The benthic fauna and sedi-
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SALOMAN, C. H., S. P. NAUGHTON, and J. L.TAYLOR.
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SALOMAN, C. H., S. P. NAUGHTON, and J. L.TAYLOR.
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SALOMAN, C. H., and J. L. TAYLOR. 1969. Age and
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SALVINI-PLAWEN, L. v. 1985. New interstitial
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SAVERCOOL, D. M., and R. R. LEWIS, III. 1994. Hard
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SCHELTEMA, A. H. 1968. Redescriptions of Anachis
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SCHWENGEL, J. S. 1940. Two new Floridan marine
shells. The Nautilus 53(3): pl. 12; 53(4): 109, 110.

SCHWENGEL, J. S. 1940. New Mollusca from Florida.
The Nautilus 54(2): 49-52, pl. 3.

SCHWENGEL, J. S. 1941. A genus and family of ma-
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55(2): 37-40, pl. 3.

SCHWENGEL, J. S. 1941. Marginella hartleyana, new
species. The Nautilus 55(2): 65, pl. 3.

SCHWENGEL, J. S. 1942. Living Mitraflorida. The Nau-
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SCHWENGEL, J. S. 1942. New Floridan marine mol-
lusks. The Nautilus 56(2): 62-66, pl. 3, 6.

SCHWENGEL, J. S. 1943. New marine shells from
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SCHWENGEL, J. S. 1943. [Note on Diadora jaumei].
The Nautilus 57(1): 32.

SCHWENGEL, J. S. 1944. Smaragdia viridis viridemaris
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SCHWENGEL, J. S. 1944. A new Floridian Lamellaria.
The Nautilus 58(1): 17, 18, pl. 1.

SCHWENGEL, J. S.1949. Fusinus spectrum (Adams and
Reeve), a new record for the Gulf of Mexico.The Nau-
tilus 62(3): 101, 102.

SCHWENGEL, J. S. 1951. New marine mollusks from
British West Indies and Florida Keys. The Nautilus
64(4): 116-119, pl. 8.


70 FMRI Technical Report TR-3


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Macroinvertebrate Checklists


Forests in the Indo-Pacific Region; the Genus Littoraria.
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RICHARDS, B. R., and W. F. CLAPP. 1944. A prelimi-
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RICHARDS, H. G. 1933. New varieties of Melongena and
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ROBERTSON, R. 1957. A study of Cantharus multangu-
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ROBERTSON, R. 1958.The family Phasianellidae in the
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ROBERTSON, R. 1959.The family Phasianellidae in the
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ROBERTSON, R. 1961. A second western Atlantic Ris-
soella and a list of the species in the Rissoellidae. The
Nautilus 74(4): 131-136, pl. 9; 75(1): 21-26.

ROBERTSON, R. 1963. Bathymetric and geographic
distribution of Panopea bitruncata. The Nautilus 76(3):
75-82.

ROBERTSON, R. 1973. Cyclostremella: A planispiral
pyramidellid. The Nautilus 87(3): 88.

ROBERTSON, R. 1978. Spermatophores of six eastern
North American pyramidellid gastropods and their
systematic significance (with the new genus Boonea).
Biological Bulletin 155(2): 360-382.

ROBERTSON, R. 1985. Archaeogastropod biology and
the systematics of the genus Tricolia (Trochacea: Tri-
coliidae) in the Indo-West-Pacific. Monographs of Ma-
rine Mollusca 3: 1-103.

ROBERTSON, R. 1994. Two new tropical western At-
lantic species of Epitonium, with notes on similar global


species and natural history.The Nautilus 107(3): 81-93.

ROBERTSON, R. 1996. Fargoa bartschi (Winkley, 1909):
A little-known Atlantic and Gulf coast American odos-
tomian (Pyramidellidae) and its generic relationships.
American Malacological Bulletin 13(1/2): 11-21.

ROBERTSON, R., andT. HABE. 1965.Alexania replaces
Habea (Epitoniidae). The Nautilus 78(4): 140, 141.

ROBERTSON, R, andT. MAU-LASTOVICKA. 1979.The
ectoparasitism of Boonea and Fargoa (Gastropoda: Pyra-
midellidae). Biological Bulletin 157(2): 320-333.

ROBERTSON, R., and K. OYAMA. 1958. The family
Stenacmidae. The Nautilus 72(2): 68, 69.

ROLAN, E. 1992.The family Omalogyridae G. O. Sars,
1878 (Mollusca, Gastropoda) in Cuba with description
of eight new species. Apex 7(2): 35-46.

ROLAN, E., and J. ESPINOSA. 1992. La familiar
Cerithiopsidae H. y A. Adams, 1853 (Mollusca, Gas-
tropoda), en la Isla de Cuba. 1. El genero Retilaskeya
Marshall, 1978. Publicacoes Ocasionis da Sociedad
Portuguesa de Malacologia 16: 39-43, 1 pl.

ROLAN, E., and J. ESPINOSA. 1992. La familiar
Cerithiopsidae H. y A. Adams, 1853 (Mollusca, Gas-
tropoda), en la Isla de Cuba. 2. El genero Horologica
Laseron 1956. Publicacoes Ocasionis da Sociedad Por-
tuguesa de Malacologia 16: 45-49, 1 pl.

ROLAN, E., and J. ESPINOSA. 1994.The familyTriphor-
idae (Mollusca, Gastropoda) in Cuba. 3.The genus Iso-
triphora. Basteria 58(1-2): 63-68.

ROLAN, E., and R. FERNANDEZ-GARCtS. 1993. La fa-
milia Triphoridae en la isla de Cuba. 1. El g6nero
Metaxia Monterosato, 1884. Bollettino Malacologico
28(6-12): 169-176.

ROLAN, E., and R. FERNANDEZ-GARCES. 1993.The
family Triphoridae (Mollusca, Gastropoda) in Cuba. 2.
The genus Iniforis Jousseaume, 1884. Apex 8(3): 95-106.

ROLAN, E., and R. FERNANDEZ-GARCES. 1994.The
familyTriphoridae (Mollusca, Gastropoda) in Cuba. 4.
The genera Monophorus, Nototriphora, Cosmotriphora
and Cheirodonta with the description of three new
species. Apex 9(1): 17-27.

ROLAN, E., and R. FERNANDEZ-GARCtS. 1995.The
family Triphoridae (Mollusca, Gastropoda) in Cuba. 5.


FMRI Technical Report TR-3


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W. G. Lyons--Mollusca








Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


SCHWENGEL, J. S., and T. L. McGINTY. 1942. Some
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FMRI Technical Report TR-3 71


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W. G. Lyons--Mollusca







Macroinvertebrate Checklists


new species of Aegires Loven, 1844 (Opisthobranchia:
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FMRI Technical Report TR-3


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W. G, Lyons--Mollusca







Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


scription ofMelongena sprucecreekensis n. sp. Bulletin of
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TURGEON, D. D., A. E. BOGAN, E. V. COAN, W. K.
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VERRILL, A. H. 1947. Additional Strombus samba
(Clench) from Florida. The Nautilus 61(1): 32.


74 FMRI Technical Report TR-3


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Camp et al. 1998 Macroinvertebrate Checklists W. G. Lyons-Mollusca


VERRILL, A. H. 1947. Additions to the Mollusca of
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VERRILL, A. H. 1947. Ensis directs in Lake Worth,
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VERRILL, A. H. 1947. Unrecorded habit of Cymatium
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VOKES, E. H., and A. D'ATTILIO. 1982. Review of the
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FMRI Technical Report TR-3 75


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Macroinvertebrate Checklists


VOKES, E. H., and R. HOUART. 1986. An evaluation of
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Prosobranchia, Eulimidae). Zoologica Scripta 9:187-210.


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WAREN, A. 1980. Descriptions of new taxa of Eulim-
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new species. The Nautilus 74(4): 149-157.


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Mollusca-W. G. Lyons Macroinvertebrate Checklists Camp et al. 1998


WELLS, H. W., and M. J. WELLS. 1962.The distinction
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and Sons, NewYork. 132 pp.


78 FMRI Technical Report TR-3


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Macroinvertebrate Checklists


Camp et al. 1998










Checklist of Shallow-Water Marine Polychaetous
Annelida of Florida

Thomas H. Perkins


Methods

This checklist of Florida shallow-water polychaetes,
which current research is showing to be a weakly sup-
ported monophyletic group (Rouse and Fauchald,
1997), was derived from a variety of sources. These
sources include 1) the extensive systematic literature
on Florida polychaetes, which begins primarily with the
publication on Florida annelids, in German, of Ehlers
(1887), and which has not been systematically synthe-
sized to allow for rapid identification; 2) the published
results of ecological surveys; 3) data from specimen col-
lections; 4) unpublished data and specimens examined
from recently completed estuarine environmental-
assessment programs (EMAP) conducted in Florida by
the U.S. Environmental Protection Agency and the
National Oceanic and Atmospheric Administration
(NOAA) Coastal Oceans Program; and 5) other pub-
lished and unpublished information from surveys con-
ducted by the State of Florida and by qualified con-
sultants for the state, the federal government, and pri-
vate organizations. As a result, reliable polychaete
identification data are available from almost all areas
of the state.
The primary specimen repository from which
species names were extracted for inclusion on this list
is the Marine Specimen Collection housed at the
Florida Marine Research Institute, St. Petersburg. Data
from the Harbor Branch Oceanographic Museum, Fort
Pierce, Florida, were also used. Other specimens from
Florida, some of which have been examined but data
for which were not generally used in compiling the list,
are housed at the Florida Department of Environ-
mental Protection's central laboratory, Tallahassee,
and district offices; in the Texas A&M Oceanographic
Collection, College Station, Texas; at Gulf Coast Re-
search Laboratory, Ocean Springs, Mississippi; at Barry
Vittor & Associates, Inc., Mobile, Alabama; in collec-
tions of the National Museum of Natural History,
Washington, D.C.; and in collections of other national
and international museums.
Citations for most of the literature listed as help-
ful in identifying Florida polychaetes (see References)
were extracted from the Ward and Fauchald bibliog-
raphy (Ward and Fauchald, 1997) available on the in-
ternet. Additional, more recent citations and local ref-
erences not included in that bibliography are also


listed among the polychaete references, resulting in
about 500 literature citations that are pertinent to the
identification of Florida polychaetes.
Criteria used to determine which species names to
include in the list were various combinations of the fol-
lowing: 1) location in estuarine and near-coastal wa-
ters of the state in depths of about 37 m or less; 2) a
moderately skeptical determination by me of the reli-
ability of an identification, continued availability of
vouchers that will allow specimens to be reliably iden-
tified, or both; and 3) for published "lettered" or un-
named species, the availability of a description and
voucher specimens that, with locality data, will allow
species to subsequently be identified or described. A
few taxa listed as "Genus name" cf. "species name,"
based on specimens in the FMRI Marine Specimen Col-
lection that possibly are that species and may not de-
serve the"cf."designation have been included on the
list.
Considering the state of knowledge of Florida
polychaetes, this list is a work in progress.There is no
doubt that there are names on the list that will prove
to be incorrect, and some species may be listed under
two or more names. There are specimens available of
many undescribed species in Florida waters that are
not listed, just as there are species that remain undis-
covered.
The classification of the Polychaeta is currently
unstable, especially at categories higher than family.
The system in which the species are reported here is
based on the classical system of Pettibone (1982). An-
other commonly used but different system, that of
Fauchald (1977a), was used by the authors of the Tax-
onomic Guide to the Polychaetes of the Northern Gulf of
Mexico (Uebelacker and Johnson, 1984). A new, well-
documented classification system for family groups
and higher categories has been proposed (Rouse and
Fauchald, 1997). Rouse and Fauchald state that it is
unlikely that any new classification will have any
longevity, but their system should provide the basis for
a stable and phylogenetically correct classification in
the future.

Acknowledgments

Many biologists have provided specimens over the
years. In particular, I thank Robert Ernest, Ecological


FMRI Technical Report TR-3 79








Polychaeta-T. H. Perkins Macroinvertebrate Checklists Camp et al. 1998


Associates (formerly Applied Biology, Inc.), Jensen
Beach, Florida; Harvey Rudolph, Palm Beach County
Government, formerly Florida Department of Envi-
ronmental Protection (FDEP); Steve Kent, Florida De-
partment of Environmental Protection, Orlando; David
Karlen, Environmental Protection Commission of Hills-
borough County; Vicki Whiting, FDEP central labora-
tory, Tallahassee; Jerry McLelland, Gulf Coast Research
Laboratory; Dr. Gary Gaston and Carol Cleveland,
University-of Mississippi; and Dr. Marianne Whitehurst
andTimThibault, BarryVittor & Assoc., Inc. Bob Ernest
also provided data on specimen identifications from
many studies conducted by Applied Biology, Inc. Har-
vey Rudolph provided data on specimens he and oth-
ers identified from Biscayne Bay. Jerry McLelland and
Sarah LeCroy, Gulf Coast Research Laboratory, pro-
vided data for their specimens collected at Perdido
Key, Florida, during studies for the National Park Ser-
vice; David Karlen provided data for specimens from
Tampa Bay collected for the Tampa Bay National Es-
tuary Program. Tom Dix, Florida Marine Research In-
stitute (FMRI), provided data for polychaetes identified


from FMRI studies in Florida Bay. Marianne Whitehurst
and Tim Thibault provided data from studies con-
ducted by BarryVittor & Assoc., Inc., for NOAA in Bis-
cayne Bay and Florida Bay and provided updates to the
nomenclature of species reported in the Taxonomic
Guide to the Polychaetes of the Northern Gulf ofMexico (Ue-
belacker and Johnson, 1984). Gary Gaston and Carol
Cleveland also provided information from their stud-
ies for NOAA in Florida Bay and Biscayne Bay and
from their Florida-West Indian Province EMAP stud-
ies for the U.S. Environmental Protection Agency (EPA)
laboratory at Gulf Breeze, Florida. Dr. John Macauley,
EPA, Gulf Breeze, Florida, and Linda Harwell, Johnson
Controls, Inc., Gulf Breeze, Florida, the EPA data con-
sultant, provided detailed data for specimens collected
in the Florida area of the Louisianan Province estuar-
ine EMAP program. Dr. Kristian Fauchald, National
Museum of Natural History, Washington, D.C., Tim
Thibault, Marianne Whitehurst, Gary Gaston, and
Carol Cleveland reviewed draft copies of the species
list and corrected several of my mistakes. All are grate-
fully acknowledged.


Checklist of the Polychaetous Annelida


Phylum Annelida
Class Polychaeta
Order Phyllodocida
Family Phyllodocidae Orsted, 1843

Genus Eulalia Savigny, 1822
Eulalia hutchinsonensis (Perkins, 1984)
Eulalia myriacyclum (Schmarda, 1861)

Genus Eumida Malmgren, 1865
Eumida sanguinea (Orsted, 1843)

Genus Eteone Savigny, 1822
Subgenus Hypereteone BergstrBm, 1914
Eteone (Hypereteone) alba Webster, 1879
Eteone (Hypereteone) foliosa Quatrefages, 1866
Eteone (Hypereteone) heteropoda Hartman, 1951

Genus Hesionura Hartmann-Schr6der, 1958
Hesionura coineaui (Laubier, 1962)
Hesionura laubieri ? (Hartmann-Schroder, 1963)

Genus Mystides Th6el, 1879
Mystides borealis Th6el, 1879
(= M. nr. borealis,fide Eibye-Jacobsen, 1992)

Genus Nereiphylla Blainville, 1828
Nereiphylla castanea (Marenzeller, 1879)


Nereiphyllafragilis (Webster, 1879)
Nereiphylla nr. paretti of Hartman, 1951
Nereiphylla sp. A of Gathof, 1984
(as Genetyllis sp. A)

Genus Paranaitis Southern, 1914
Paranaitis capensis (Day, 1960)
Paranaitis gardineri Perkins, 1984
Paranaitis papillosa (Ehlers, 1887)
Paranaitis speciosa (Webster, 1879)

Genus Phyllodoce Lamarck, 1818
Subgenus Anaitides Czerniavsky, 1882
Phyllodoce (Anaitides) erythrophylla (Schmarda,
1861)
Phyllodoce (Anaitides) groenlandica (Orsted, 1843)
Phyllodoce (Anaitides) longipes (Kinberg, 1866)
Phyllodoce (Anaitides) maderiensis Langerhans,
1880
Phyllodoce (Anaitides) mucosa (Orsted, 1843)
Subgenus Phyllodoce Lamarck, 1818
Phyllodoce (Phyllodoce) arenae Webster, 1879
Phyllodoce (Phyllodoce) tortugaeTreadwell, 1917

Genus Pirakia Bergstrim, 1914
Pirakia punctifera (Grube, 1860)

Genus Protomystides Czerniavsky, 1882


FMRI Technical Report TR-3


Polychaeta--T. H. Perkins


Macroinvertebrate Checklists


Camp et al. 1998




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