An annotated checklist and bibliography of the land and freshwater snails of México and Central America

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Title:
An annotated checklist and bibliography of the land and freshwater snails of México and Central America
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Bulletin of the Florida Museum of Natural History ;
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299 p. : ; 28 cm.
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English
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Thompson, Fred G ( Fred Gilbert ), 1934-
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Florida Museum of Natural History, University of Florida
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Gainesville, Fla
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Gastropoda -- Mexico   ( lcsh )
Gastropoda -- Central America   ( lcsh )
Gastropoda -- Bibliography   ( lcsh )
Pulmonata -- Mexico   ( lcsh )
Pulmonata -- Central America   ( lcsh )
Pulmonata -- Bibliography   ( lcsh )
Freshwater snails -- Mexico   ( lcsh )
Freshwater snails -- Central America   ( lcsh )
Freshwater snails -- Bibliography   ( lcsh )
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bibliography   ( marcgt )
non-fiction   ( marcgt )

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Includes bibliographical references and index.
Statement of Responsibility:
Fred G. Thompson.

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University of Florida
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BULLETINAN ANNOTATED CHECKLIST AND BIBLIOGRAPHY OF THE LAND AND FRESHWATER SNAILS OF MXICO AND CENTRAL AMERICA Fred G. ThompsonVol. 50, No. 1, pp. 1 2011 UNIVERSITY OF FLORIDA GAINESVILLE

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The FLORIDA MUSEUM OF NATURAL HISTORY is Floridas state museum of natural history, dedicated to understanding, preserving, and interpreting biological diversity and cultural heritage. The BULLETIN OF THE FLORIDA MUSEUM OF NATURAL HISTORY is a peer-reviewed journal that publishes results of original research in zoology, botany, paleontology, archaeology, and museum science. The Bulletin is published at irregular intervals, and volumes are not necessarily completed in any one year. Volumes contain between 150 and 300 pages, sometimes more. The number of papers contained in each volume varies, depending upon the number of pages in each paper, but four numbers is the current standard. Multi-author issues of related papers have been published together, and inquiries about putting together such issues are welcomed. Address all inquiries to the Managing Editor of the Bulletin. Richard C. Hulbert Jr., Editor Bulletin Committee Ann S. Cordell Richard C. Hulbert Jr. Jacqueline Miller Larry M. Page Roger W. Portell, Treasurer Irvy R. Quitmyer David W. Steadman, ISSN: 0071-6154 Copyright 2011 by the Florida Museum of Natural History, University of Florida. All rights reserved. Text, commercial use or republication by printed or electronic media is strictly prohibited without written permission of the museum. Publication Date: September 30, 2011 Price: $45.00 This issue completes Volume 50 Send communications concerning puchase or exchange of this publication and manuscript queries to: Managing Editor of the Bulletin Florida Museum of Natural History University of Florida P.O. Box 117800 Gainesville, FL 32611-7800 USA FAX: 352-846-0287

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ABSTRACTThe checklist of the non-marine gastropod fauna summarizes the known species and subspecies that are recognized from Mxico and Central America. It is an annotated list of 1800 terminal taxa presented in a hierarchical framework. These include 1502 native species plus 278 native subspecies, and 20 introduced species. The native species include 175 aquatic operculates, 84 aquatic pulmonates, 130 terrestrial operculates, and 1112 terrestrial pulmonates. In most cases in order to be as objective as possible the list uncritically records the most recent assignment of terminal taxa. In a few instances some changes are deemed necessary. For higher-level changes competing schemes are treated equally. In cases of terminals and higher taxa, readers are directed to the systematic works that discuss relevant taxonomy. It is anticipated that the annotated list will be a useful resource for everyone interested in non-marine gastropods and their nomenclature. In addition to clarifying some issues or points of confusion, this list should also provide an impetus for future work aimed at clarifying and resolving areas of taxonomic disagreement and/or uncertainty, and to make better known the non-marine molluscan fauna of Mxico and Central America.RESUMEN de Mxico y Centroamrica. Esta es una lista anotada de 1800 taxa terminales presentes en el sistema jerrquico. Estas incluyen 1502 especies nativas, 278 subespecies nativas y 20 especies introducidas. Estas especies nativas incluyen 175 especies de operculados, 84 pulmonados acuticos, 130 de operculados terrestres y 1112 de pulmonados terrestres. Para ser lo ms objetivo posible, en la mayora de los casos algunos cambios se consideraron necesarios. Para cambios a un nivel jerarquico ms alto, los esquemas contradictorios propuestos fueron tratados equitativamente. En los casos de taxa terminales y superiores, los lectores son dirigidos a los trabajos sistemticos que discuten la taxonoma relevante. Se prev que la como para hacer ms conocida la fauna de moluscos no marina de Mxico y Centroamrica. Key Words: Gastropoda; biogeography; Mxico; Central AmericaAN ANNOTATED CHECKLIST AND BIBLIOGRAPHY OF THE LAND AND FRESHWATER SNAILS OF MXICO AND CENTRAL AMERICA Fred G. Thompson1Thompson, F. G. 2011. An annotated checklist and bibliography of the land and freshwater snails of Mxico and Central America. Bulletin of the Florida Museum of Natural History 50(1):1.1

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2 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)TABLE OF CONTENTSIntroduction ..........................................................................3 Subclass Vetigastropoda .......................................................4 Superfamily Helicinoidea ..........................................4 Superfamily Ampullarioidea ...................................21 Superfamily Viviparoidea ........................................24 Superfamily Cyclophoroidea ...................................25 Superfamily Cerithioidea ........................................34 Superfamily Littorinoidea .......................................43 Superfamily Rissooidea ...........................................47 Superfamily Valvatoidea .........................................61 Subclass Pulmonata ...........................................................62 Superfamily Lymnaeoidea .......................................62 Superfamily Planorboidea .......................................63 Superfamily Ellobioidea ..........................................79 Superfamily Veronicelloidea ...................................80 Superfamily Succineoidea .......................................82 Superfamily Achatinelloidea ...................................87 Superfamily Cochlicopoidea ...................................87 Superfamily Pupilloidea ..........................................87 Superfamily Orthalicoidea ....................................100 Superfamily Urocoptoidea ....................................130 Superfamily Achatinoidea .....................................157 Superfamily Testacelloidea ....................................167 Superfamily Streptaxoidea ....................................201 Superfamily Rhytidoidea .......................................202 Superfamily Acavoidea .........................................203 Superfamily Punctoidea ........................................204 Superfamily Sagdoidea ..........................................209 Superfamily Helicarionoidea .................................210 Superfamily Gastrodontoidea ................................215 Superfamily Zonitoidea .........................................216 Superfamily Limacoidea .......................................222 Superfamily Arionoidea ........................................223 Superfamily Helicoidea .........................................224 Bibliography of the Land and Freshwater Snails of Mxico and Central America ...........................260 Appendix. Faunal lists for Central American countries and Mxican states ................................................278 Taxonomic Index (for Families, Subfamilies, Genera, and Subgenera only) ..............................................297

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 3INTRODUCTION An annotated checklist and bibliography of the land and freshwater snails of Mxico and Central malacology/mexico-central_america_snail_ many idividuals who have offered corrections and comments that have made this a better contribution to neotropical biology. Since then many taxonomic changes and editions to the fauna require an updated revision. The objective of this study is to list all species and subspecies of land and freshwater gastropods reported from Mxico and Central America. The list includes 1502 native species plus an additional 238 native subspecies, and 20 introduced species. The list includes published distribution records within this area for all terminal taxa. In only a few instances are unpublished distribution records included. The following geographic terms are used. North America refers to the North American Continent, which extends south to the Polochic Valley in Guatemala. Central America refers to the countries of Panam, Costa Rica, Nicaragua, Honduras, El Salvador, Guatemala, and Belize, to these countries. Middle America refers to Central America and the West Indies. Records for New Providence Island, Colombia, are included because America, not Colombia. The geographic area of this study is vast, and it contains many biomes, different geological structures, complex physiographic features and a myriad of ecological settings. Local surveys have been published for only a small portion of this area. It is not possible on the basis of published information to identify biotic hotspots, because the available biogeographic information does not allow for objective comparisons of one region with another. A tabulation of the faunal diversity for each Central American country and each Mexican state is provided in the appendix. A disproportionate large number of 293 species and subspecies have been recorded from Veracruz State, Mxico compared to of investigators who have worked there. Even their investigations were localized within smaller subregions of central Veracruz. The localized regional data are not comparable because of historical socio-political circumstances effecting access, and the emphasis different investigators placed on certain taxa. The extensive limestone terrains of southern and western Mxico promise to be as species diverse as central Veracruz, but their faunas are greatly under-reported. The number of known land and freshwater gastropods from Mxico and Central America is large. Nearly all of the know species are based on sound taxonomic studies and reviews dating back to Shuttleworth, Menke, and Pfeiffer. I estimate that the number of recorded taxa is about 35% of the actual fauna. The high number of estimated undescribed species is based on the fact that most of Panam, much of Costa Rica, most of Nicaragua, nearly all of Honduras, most of Guatemala, nearly all of Belize, and 85% of Mxico have not been explored, or have been poorly explored for mollusks, especially for small and minute species. I have collected extensively in these areas, and reported faunas of better known regions. This of the extensive specimens deposited in the Florida additional new species. I suspect the estimate that only 35% of the actual Mexican-Central American fauna is known is conservative. Systematic biology is made unnecessarily species or to resurrect them from synonymy responsibility is on authors to provide evidence supporting their actions. To do otherwise is poor science. Merely to state that a name is a synonym does not constitute a valid action. In accordance with ICZN Article 45.6.4, and for purposes of this

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4 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)work, varieties and forms of species published prior is made concerning the validity of such subspecies. and subfamilies used here follows Bouchet and Rocroi (2005:240). That reference should be consulted in cases where the type genus, synonymy names include the publication in which the name Fischer and Crosse (1970), Strebel (1873 1882), Von Martens (1890), Bequaert and they are general works, and no other publications MUSEUM ACRONYMS Type specimens are cited only when this information is readily available in publication. The following acronyms refer to museum collections where cited specimens are housed: ANSP, Academy of Natural Sciences, Philadelphia, Pennsylvania; BMNH, The Natural History Museum, London, UK; CAS, California Academy of Sciences, San Francisco, California; FMNH, Field Museum of Natural History, Chicago, Illinois; INBio, Museo de Zoologa, Universidad de Costa Rica, San Jos, Costa Rica; MCZ, Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts; MHNN, Museum dHistoire Naturelle Neuchtel; MNHN, Musum National dHistoire Naturelle, Paris, France; NMBE, Natural History Museum, Bern; RMNH, Nationaal Natuurhistorisch Museum Naturalis, Leiden, the Netherlands; SMF, Natur-Museum Senckenberg, Frankfurt am Main, Germany; UF, Florida Museum of Natural History, Gainesville, Florida; UMMZ, Museum of Zoology, University of Michigan, Ann Arbor, Michigan; USNM, National Museum of Natural History, Washington, DC; ZMA, Zologisch Museum, Amsterdam, the Netherlands; ZMB, Museum fr Naturkunde, Humboldt Universitt, Berlin, Germany; and ZMH, Zoologisches Museum der Universitt Hamburg, Hamburg, Germany.Class GASTROPODA Cuvier 1795 Subclass VETIGASTROPODA Salvini-Plawen 1989 Superfamily HELICINOIDEA Frussac 1822 Family CERESIDAE Thiele 1925 Ceresinae Thiele 1925; Handbuck der Zoologie 5:78. Ceresidae Thompson 1980; Malacologia 20:13.Distribution.Mxico, Venezuela, Colombia, Equador, Peru, and Brazil. Taxonomy.Six genera are recognized. Three occur in the study area. Genus Ceres Gray 1856Ceres Gray 1856; Proc. Zool. Soc. Lond. 24:100.Thompson 1980; Malacologia 20:17.Type Species.Carocolla eolina Ducluz 1834. Distribution.Eastern Mxico in the states of Veracruz, San Lus Potos and Tamaulipas. Taxonomy.Three species are recognized. Ceres eolina (Duclos 1834)Carocolla eolina Duclos 1834; in Gurins Mag. Zool.: pl. 30. Odontostoma eolina (Duclos). Pfeiffer 1848; Monog. Helic. Viv. 1:11. Proserpina eolina (Duclos). Pfeiffer 1853; Monogr. Helic. Viv. 3:290. Ceres eolina (Duclos). Gray 1856; Proc. Zool. Soc. Lond. 24:102.Fischer & Crosse 1892; Miss. Sci. Mex. II:382; pl. 54, Proserpina (Ceres) eolina (Duclos). Pfeiffer, 1856; Proc. Zool. 1890; Biol. Cent. Amer.:44.Von Martens 1901; Biol. Cent. Amer.:609.Type Locality.Cerro de Plumas [Cerro de Palmas] Sierra de Matlaquihahuitl, Veracruz, Mxico. Distribution.Known only from the type locality. Ceres nelsoni Dall 1898Ceres nelsoni Dall 1898; Nautilus 12:27.Fischer & Crosse 1, 3, 5, 8 (shell).Solem 1954; Nautilus 64:7.Thompson 1980; Correa-Sandoval, Garca-Cubas & Reguero 1998; Acta Zool. Mex. (73):13.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):235. Proserpina (Ceres) nelsoni (Dall). Von Martens 1901; Biol. Cent. Amer.:609.Type Locality.Pilitla [Xilitla], San Lus Potos, Mxico. Holotype USNM 107823. Distribution.SAN LUS POTOS: numerous localities

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 5 (Correa-Sandoval et al. 1998). TAMAULIPAS: Aserradero del Paraiso, 15 km NNW of Chamal (Solem 1957); numerous localities in southern Tamaulipas (Correa-Sandoval & Rodriguez 2002). Ceres salleana Gray 1856Ceres salleana Gray 1856; Proc. Zool. Soc. Lond. 24:100, 102.Pfeiffer 1876; Monogr. Pneumo. Viv. 4:295.Fischer & Malacologia 24:6, 17. Proserpina (Ceres) salleana (Gray). Pfeiffer 1856; Proc. Zool. Soc. Acad. Nat. Sci. Phila. 43:332.Von Martens 1890:45.Von Martens 1901; Biol. Cent. Amer.:609. Distribution.VERACRUZ: Orizaba (Pilsbry 1891); Huatusco, Barranca de Dos Puentes; Cerro de Palma, Sierra de Matlaquihuahuitl, near Toxpan (Von Martens 1901). Genus Linidiella Jousseaume 1889Linidiella Jousseaume 1889; Mem. Zool. Soc. France 2:256.H. B. Baker 1923; Nautilus 36:84.Thompson 1980; Malacologia 24:19. Chersodespoena Sykes 1900; Proc. Malac. Soc. London 4:136.Type Species.Linidiella : Proserpina swifti Bland 1863 (H. B. Baker 1923). Chersodespoena: Despoena (Chersodespoena) cinnanomea Sykes 1900 (monotypy). Distribution.Ecuador, Venezuela, eastern Mxico. Taxonomy.Four species are recognized. Two occur in Mxico. Linidiella citrina Thompson 1987Linidiella citrina Thompson 1987; Arch. fr Mollusk. 117:160; Type Locality. km NE of Comolapa, Veracruz, Distribution.VERACRUZ: known only from the immediate vicinity of Comolapa. Linidiella sulfureous Thompson 1967Linidiella sulfureous Thompson 1967; Proc. Biol. Soc. Wash. 80:61; Type Locality..2 mi. S of Solusuchiapa, Chiapas, Mxico; 1600 ft. alt. Holotype UF 19025. Distribution.Known only from the type locality. Genus Proserpinella Bland 1865Proserpinella Bland 1865:157.Thompson 1980; Malacologia 24:23.Type Species.Proserpinella berendti Bland 1865 (monotypy). Distribution.Mexican states of Veracruz, Jalisco and Nayarit. Taxonomy.Three species are recognized. Proserpinella berendti (Bland 1865)Proserpinella berendti Bland 1865; Ann. Lyceum Nat. Hist. Thompson 1980; Malacologia 24:24. Proserina (Proserpinella ) berendti (Bland). Von Martens 1890; Biol. Cent. Amer.:45.Von Martens 1901; Biol. Cent. Amer.:609.Type Locality.Mirador, Veracruz, Mxico. Distribution.Known only from the type locality. Proserpinella edentula Naranjo-Garcia 1994Proserpinella edentula Naranjo-Garcia 1994; Arch. fr Mollusk. reproductive system).Type Locality.Municipio La Huerta, Estacion de W), Jalisco, Mxico. Holotype Universidad Nacional Autonoma de Mxico, Instituto de Biologia 260. Distribution.Known only from the type locality. Proserpinella hannae Dall 1926Proserpinella hannae Dall 1926; Proc. Calif. Acad. Sci., ser. Malacologia 24:24.Type Locality.Isla Mara Madre, Islas Maras, Nayarit, Mxico. Holotype CAS 2211. Distribution.Known only from the type locality. Family HELICINIDAE Frussac 1822 Islands, Australia, Indian Ocean islands and southeast Asia. Taxonomy.Numerous genera and subgenera. There is little consensus on the status and validity of many of these. The family Helicinidae is poorly studied, and new investigations will require many changes. A study by Richling (2004) is a major step in addressing many taxonomic problems in Central America and Mxico. Three subfamilies are recognized in the Neotropical region: Helicininae, Vianinae and Stoastominae. and coastal Venezuela. Subfamily HELICININAE Frussac 1822 Distribution.New World, ranging from Argentina north to the southeastern United States and throughout the West Indies. Taxonomy.Four genera of Helicininae occur in Mxico and Central America. Fifty-seven species and twelve subspecies are recognized in the area covered in this study. Careful studies of morphology, biogeography and ecology by H. B. Baker (1922, 1926, 1928) and Richling (2004) demonstrated that many forms that were previously considered subspecies or varieties are valid species. This summary relies heavily upon the work of Richling (2004). Genus Ceochasma Thompson 1968Ceochasma Thompson 1968, Proc. Biol. Soc. Wash. 81:45.

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6 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Helicina (Ceochasma) (Thompson). Richling 2004; Malacologia 45:426, 427.Type Species.By monotypy Ceochasma phrixina Thompson 1968. Distribution.Colima, Mxico. Taxonomy.The systematic position of this genus remains uncertain (Richling 2004). A single species is recognized. Ceochasma phrixina Thompson 1968Ceochasma phrixina Thompson 1968; Proc. Biol. Soc. Wash. (reproductive anatomy).Type Locality.A collapsed limestone ridge 0.3 mile east of Tamala, Colima, Mxico; 500 ft alt. [Tamala is a small village ca. 2 km NE and ca. 6 km E of Tacolapa]. Holotype UF 20138. Distribution.Known only from the type locality. Genus Helicina Lamarck 1799Helicina Lamarck 1799:76.Type Species.Helicina neritella Lamarck 1799. Distribution.The West Indies, Mxico, Central America and South America. Taxonomy.Nine subgenera are recognized in this study. Helicina s. s., Oxyrhombus, Tristramia, Pseudolygyra Cinctella Oligyra, Gemma, Analcadia and Sericea The superfamily Helicinoidea is a nearly morphostatic group of organisms in which the soft anatomy shows little differentiation between and within genera. In this case shell characters prove useful for grouping species, even though in some instances the distinctions between subgenera approach being arbitrary. H. B. Baker (1922a, 1926, 1928), Thompson (1968, 1980, 1982) and Richling (2004) provided fauna. Richling greatly advanced our knowledge of the Costa fewer subgenera. Integrating the rest of the Mxico and proposed requires recognition of additional subgenera. This by H. B. Baker (1928). Subgenus Oligyra Say 1818Olygyra Say 1818. Oligyra Say 1819 (amended for Olygyra). Orbiculata Wagner 1905; Denksch. Kaiserli. Akad...Type Species.Oligyra orbiculata Say 1818 (type by monotypy). Distribution.Southeastern United States, Mxico and Central America. Taxonomy.Three species and one subspecies occur within the study area. Helicina (Oligyra) orbiculata orbiculata (Say 1818)Oligyra orbiculata Say 1818; Jour. Acad. Nat. Sci. Phila. pt. 2:283. Oligyra (Oligyra) orbiculata orbiculata Say. H. B. Baker 1922; (radula). Helicina orbiculata Wagner 1910; in Martini & Chemnitz, Syst. Helicina (Oligyra) orbiculata orbiculata Say. H. B. Baker 1926; anatomy). Helicina hanleyana Pfeiffer 1849; Proc. Zool. Soc. Lond. Type Locality.Oyster shell hammocks near the mouth of the St. Johns River, Florida. Holotype ANSP 10573. Distribution.In the United States the species is widely distributed in Florida, Georgia, Alabama, Tennessee, Mississippi, Louisiana, Arkansas and Oklahoma. Numerous localities in southern TAMAULIPAS and eastern SAN LUS POTOS (Correa-Sandoval et al. 1998; Correa-Sandoval & Rodriguez 2002). Helicina (Oligyra) orbiculata tropica Pfeiffer 1852Helicina tropica 9, 10 (shell).Strenth & Littleton 2000; Texas Jour. Sci. 52: 25. Helicina orbiculata tropica Pfeiffer. Pilsbry 1896:534.Pilsbry Oligyra (Oligyra) orbiculata tropica (Pfeiffer). H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:44. Helicina (Oligyra) orbiculata tropica Pfeiffer. H. B. Baker 1926; Type Locality.Not given. Distribution.COAHUILA (Pilsbry 1948). NUEVO LEON: Santiago (Correa-Sandoval 1993); Cerro de la Silla (Correa-Sandoval & Salazar 2005). TAMAULIPAS (Pilsbry 1948). Helicina (Oligyra) borealis Von Martens 1890Helicina borealis 15 (shell).Fischer & Crosse 1893:435.Von Martens 1901; Biol. Cent. Amer.:606.Wagner 1910; in Martini & Chemnitz, (shell). Oligyra (Oligyra) borealis (Von Martens). H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:44.Type Locality.Villa Lerdo, Durango, Mxico. Distribution.Known only from the type locality. Helicina (Oligyra) cordillerae Pfeiffer 1857Helicina cordillerae Pfeiffer 1857; Proc. Zool. Soc. Lond. (1886):323.Reeve 1874; Conch. Icon., Helicina 149 (shell).Von Martens 1890; Biol. Cent. Amer.:32.Von Martens 1901; Biol. Cent. Amer.:603.Wagner 1910; in Martini & Chemnitz, Syst. Conch. Cab., Helicinidae:300; pl. Oligyra (Oliyra) codillerae (Pfeiffer). H. B. Baker. 1922; Proc. Acad. Nat. Sci. Phila. 74:44.Type Locality.Volcn Orizaba, Rancheria de Jacale, Veracruz, Mxico; 3500 m alt. (Fischer & Crosse 1893).

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 7 Distribution.Known only from the type locality. Subgenus Oxyrhombus Crosse & Fischer 1893Oxyrhombus Crosse & Fischer 1893:401. Concentrica Wagner 1905; Denksch. Kaiserli. Akad. :282. Punctisulcata Wagner 1905; Denksch. Kaiserli. Akad. :294. Cinctella Wagner 1910; in Martini Chemnitz, Syst. Conch. Cab., Helicinidae:298.Type Species.Oxyrhombus: Helicina amoena (selected by H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:84). Concentrica: Helicina concentrica Pfeiffer 1849 (type by tautonomy). Punctisulcata : Helicina punctisulcata Von Martens 1900 (type by tautonomy). Cinctella : Helicina cinctella Shuttleworth 1852 (type by tautonomy). Distribution.Tamaulipas and Guerrero, Mxico south to Venezuela. Taxonomy.Eleven species are recognized within the study area. Additional species occur in northern South America. Helicina (Oxyrhombus) amoena Pfeiffer 1849Helicina amoena Pfeiffer 1845:119.Von Martens 1890; Biol. Cent. 4.Von Martens 1901; Biol. Cent. Amer.:602.Fluck 1906; Nautilus 20:1.Wagner 1910; in Martini & Chemnitz, Syst. Bequaert 1957; Bull. Mus. Comp. Zool. 115:206.Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):7. Helicina (Oxyrhombus) amoena Pfeiffer. H. B. Baker 1922; Proc. Morelet 1849; Test. Noviss. I:19. Helicina amoena var. minor Von Martens 1890; Biol. Cent. Amer.:28. Helicina amoena var. depressa Von Martens 1890; Biol. Cent. Amer.:28.Type Localities.Helicina amoena: Honduras. Helicina amoena var. minor: Nicaragua. Helicina amoena var. depressa: Bugaba, west of David, Panam. Helicina : Petn, Guatemala. Distribution.PANAM. NICARAGUA: near Rio Wounta (Rio Kukallaya), NW of Kukallaya (Fluck 1906). GUATEMALA, Dept. Alta Verapaz: Cobn (Von Martens 1890). Dept. Baja Verapaz: San Juan; Sabo, 2800 ft. alt.; Chacoj; Panzos; Teleman (Von Martens 1890). Dept. Petn: Tikal (Basch 1959). CAMPECHE: (Von Martens 1890). CHIAPAS: Laguna Ocotal, 950 m alt.; El Censo to Laguna Ocotal, 1000 m alt.; Ocosingo, 850 m alt.; El Real, 600 m alt.; Monte Libano, 600 m alt. (Bequaert 1957). VERACRUZ (Wagner 1910). Helicina (Oxyrhombus) bocourti Crosse & Fischer 1869Helicina bocourti Crosse & Fischer 1869; Jour. de Conchyl. 17:251. Helicina dysoni bocourti Crosse & Fischer. Fischer & Crosse Helicina dysoni Pfeiffer. Simpson 1897; Nautilus 11:13. Oligyra (Analcadia ) dysoni bocourti Crosse & Fischer. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:47. Helicina (Tristramia) bocourti Crosse & Fischer. Rehder 1966; 18 (radula).Richling 2004; Malacologia 45:425. Helicina dysoni var. jansoni Von Martens 1890; Biol. Cent. Alcadia (Analcadia ) dysoni bocourti (Crosse & Fischer). Wagner Type Locality.Helicina bocourti: Isla de tila, Honduras. Helicina dysoni var. jansoni: Bonacca Island [Isla Guanaja], Honduras. Distribution.HONDURAS, Dept. Islas de la Baha: Isla tila; Isla Roatn; Isla Guanaja. BELIZE: along the coast from Rocky Point to Puerto Honduras, near the Guatemala (Rehder 1966). Helicina (Oxyrhombus) cinctella cinctella Shuttleworth 1852Helicina cinctella Shuttleworth 1852; in Bern Mittheil.:304.Von Martens 1890; Biol. Cent. Amer.:29.Pilsbry 1891; Proc. Acad. Nat. Sci. Phila. 43:332.Fischer & Crosse 1893:403; pl. 55, in Martini & Chemnitz, Helicina (Oxyrhombus) cinctella cinctella Shuttleworth. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:48.H. B. Baker Syntypes: NMBE 15275/3. Distribution.JALISCO: Tepc (Von Martens 1890). (Von Martens 1890); Sumidero (H. B. Baker 1928). Helicina (Oxyrhombus) cinctella bautistae Wagner 1910Helicina (Oxyrhombus) cinctella bautistae Wagner 1910; in Martini 18 (shell). Helicina (Oxyrhombus) cinctella bautista Wagner. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:48.Type Locality.San Juan Bautista, Tabasco, Mxico. Holotype ZMB 103323. Distribution.Known only from the type locality. Helicina (Oxyrhombus) ghiesbreghti Pfeiffer 1856Helicina ghiesbreghti Pfeiffer 1856:381.Von Martens 1890; Biol. Cent. Amer.:28.Pilsbry 1892:339.Fischer & Crosse Cent. Amer.:202.Thompson 1957; Nautilus 70:100. Helicina (Oxyrhombus) ghiesbreghti Pfeiffer. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:47 Helicina ghiesbreghti Pfeiffer. Reeve 1874; Conch. Icon., Helicina: Helicina ghiesbreghti Pfeiffer. Wagner 1910; in Martini & Chemnitz, Syst. Conch. Cab., Helicinidae:296; pl. 59, Type Locality.Chiapas, Mxico. Distribution.GUATEMALA: El Reposo, 8000 ft. alt.; San Francisco Miramar; Costa Cuca (Von Martens 1890).

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8 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) CHIAPAS (Von Martens 1890). TABASCO: mountains of Poan (Pilsbry 1892); 0.5.0 mi. E of Teapa (Thompson 1957). Helicina (Oxyrhombus) heighwayana Dall 1909Helicina heighwayana Dall 1909; Smiths. Misc. Coll. 52:392.7 (shell).Type Locality.Sierra Darin, near the Rio Atrato, Panam. Holotype USNM 111074. Distribution.PANAM: Rio Puerco (Pilsbry 1926). Helicina (Oxyrhombus) isthmica Pilsbry 1926Helicina isthmica Pilsbry 1926; Acad. Nat. Sci. Phila. 78:68; textType Locality.Rio Puerco, Panam. Holotype ANSP 140713. Distribution.Known only from the type locality. Helicina ( Oxyrhombus) oxyrhyncha Crosse & Debeaux 1863Helicina oxyrhyncha (shell).Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:69; Type Locality.not given. Distribution.PANAM: Lowlands of Rio Tucuti, Panam (Pilsbry 1926). Helicina (Oxyrhombus) ptychophora Sykes 1902Helicina ptychophora Sykes 1902; Proc. Malac. Soc. London 5:20; 78:69.Type Locality.Guatemala. Distribution.GUATEMALA: exact localities are unknown. Helicina (Oxyrhombus) sanguinea Pfeiffer 1849Helicina sanguinea Pfeiffer 1849; Proc. Zool. Soc. Lond. (1848):124.Sowerby 1866; Thesaurus Conch. 3:292; pl. 275, Helicina : pl. Oligyra (Analcadia) sanguinea (Pfeiffer). H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:47. Type Locality.Honduras. Distribution.HONDURAS. Unknown. Helicina (Oxyrhombus) sowerbyana Pfeiffer 1849Helicina sowerbyana Pfeiffer 1849; Proc. Zool. Soc. Lond. (1848):124.Von Martens 1890; Biol. Cent. Amer.:28.Fischer & Crosse 1893:444.Von Martens 1901; Biol. Cent. Amer.:602.Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:782; Reguero 1998; Acta Zool. Mex. (73):13.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):235: Helicina (Oxyrhombus) sowerbyana Pfeiffer. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:48.Type Locality.Guatemala. Distribution.NUEVO LEON: Santiago (CorreaSandoval 1993). SAN LUS POTOS: Las Cascadas, (Correa-Sandoval et al. 1998). TAMAULIPAS: canyon 4 mi. (Correa-Sandoval & Rodriguez 2002, 2005). Subgenus Succincta Wagner 1905Succincta Wagner 1905; Denksch. Kaiserli. Akad. :315. Gemma Wagner 1905; Denksch. Kaiserli. Akad. : (not Gemma Deshayes 1853).Type Species.Succincta : Helicina succincta Von Martens 1890. Gemma: Helicina gemma Preston 1903. Distribution.Mxico and Central America. Taxonomy.Six species and six subspecies. Helicina (Succincta) arenicola arenicola Morelet 1849Helicina arenicola Morelet 1849; Test. Noviss. I:21.Von Martens 1890; Biol. Cent. Amer.:36.Pilsbry 1891; Proc. Acad. Nat. Sci. Phila. 43:332.Fischer & Crosse 1893:424; pl. 56, Wagner 1910; in Martini & Chemnitz, Syst. Conch. Cab., Proc. Amer. Philos. Soc. 77:257.Harry 1950; Occ. Pap. Mus. Zool. Univ. Mich. (524):29.Rehder 1966; Proc. Biol. Soc. Wash. 79:276. Oligyra (Succincta) arenicola (Morelet). H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:45.Thompson 1967; Bull. Fla. St. Mus. 11:229.Type Locality.Sisal, Yucatn, Mxico; in sandy places. Distribution.CAMPECHE: numerous localities (Thompson 1967). QUINTANA ROO: Chancanab; Laguna Chichankanab, near Esmeralda; Tulm; San Miguel, Isla Cozumel; San Gerbacio (Rehder 1966); 7.1 mi. NNE of Xiatil; 2.3 mi. SSE of Xiatil (Thompson 1967). YUCATN: Silam; Ticul; Tabi; Uxmal; Tunkas; Santa Ana (Pilsbry 1891); Chichen Itza; Progreso; 8 mi. S of Progreso; 8 mi. NW of Chichen Itza (Harry 1950); 3.2 mi. S of Progreso; 1.0 mi. SSE of Puerto Telchac (Thompson 1967). Helicina (Succincta) arenicola raresulcata Pfeiffer 1861Helicina raresulcata Pfeiffer 1861; Malak. Bltt. 8:173.Strebel Biol. Cent. Amer.:36.Wagner 1910; in Martini & Chemnitz, Oligyra (Succincta) arenicola raresulcata (Pfeiffer). H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:45.Type Locality.Veracruz, [Veracruz], Mxico Distribution.Known only from the type locality. Helicina (Succincta) Menke 1828 (shell).Von Martens 1901; Biol. Cent. Amer.:606.Hinkley 1907; Nautilus 21:71.Richling 2004; Malacologia 45:386 387.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 9 (Menke). H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:45.Bequaert 1957; Bull. Mus. Comp. Zool. 115:206.Thompson 1967; Bull. Fla. St. Mus. 11:229. Helicina (Succincta) Menke. H. B. Baker 1928; (reproductive anatomy). Type Locality.Jamaica. Dept. Copn: Copn (Von Martens 1901). GUATEMALA, Dept. Alta Verapaz: Cobn (Von Martens 1890). Dept. Baja Verapaz: Senahu (Von Martens 1890). CAMPECHE: 10.2 mi. E of Escarcega; (Thompson 1967). CHIAPAS: Laguna Ocotal, 950 m alt. (Bequaert 1957; Bull. Mus. Comp. Zool. 115); Palenque (Thompson 1967). PUEBLA: Necaxa (H. B. Baker 1928). QUINTANA ROO:4.0 mi. E of Xpujil [Campeche] (Thompson 1967). SAN LUS POTOS: 1998). TABASCO: Teapa; San Jan Bautista (Von Martens 1890). TAMAULIPAS: Tampico (Hinkley 1907);El Cielo Biosphere Reserve, Gomez Farias (Correa-Sandoval & Rodriguez 2002, 2005). VERACRUZ: Papantla; Veracruz; Misantla; Mirador (Von Martens 1890); San Rafael, Jicaltepec (Pilsbry 1896); numerous localities in northern Veracruz (Correa-Sandoval 2000). Helicina (Succincta) Pfeiffer 1857 Pfeiffer 1857; Proc. Zool. Soc. Lond. (1856):380.Wagner 1910; in Martini & Chemnitz, Syst. Oligyra (Succincta ) (Pfeiffer). H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:45.Type Locality.Chiapas, Mxico. Distribution.CHIAPAS (Wagner 1910). TABASCO: Teapa; San Juan Bautista (Wagner 1910). VERACRUZ: Helicina (Succincta) Wagner 1905Helicina brevilabris incommoda Wagner 1905; Denksch. Kaiserli. in Martini & (shell).Type Locality.Guatemala. Distribution.GUATEMALA: exact locality unknown. Helicina (Succincta) Pfeiffer 1861Helicina strebeli Pfeiffer 1861; Malak. Bltt. 8:173.Strebel Oligyra (Succincta) (Pfeiffer). H. B. Baker 1922; (radula). Helicina (Succincta) Pfeiffer. H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):28.Type Locality.Mirador, Veracruz, Mxico. Distribution.PUEBLA: Necaxa, 2215 ft. alt. (H. Helicina (Succincta) oaxacana Pilsbry 1920Helicina oaxacana Oligyra (Succincta) oaxacana (Pilsbry). H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:45.Type Locality.Puerto Angel, Oaxaca [Mxico]. Distribution.Known only from the type locality. Helicina (Succincta) oweniana oweniana Pfeiffer 1849Helicina oweniana Pfeiffer 1849:123.Pfeiffer 1850:40; pl. 7, Von Martens 1901; Biol. Cent. Amer.:605.Thompson 1957:100. Alcadia (Leialcadia ) oweniana (Pfeiffer). Wagner 1908:81; pl. 12, Oligyra (Succincta ) oweniana oweniana (Pfeiffer). H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:45. Helicina (Succincta ) oweniana oweniana Pfeiffer. Richling 2004; Type Locality.Chiapas, Mxico. Lectotype BMNH 20010751 (Richling 2004). Distribution.GUATEMALA, Dept. Petn: Tikal National Park (Basch 1959). MXICO. TABASCO: Teapa; 0.5.0 mi. E of Teapa (Thompson 1957). CHIAPAS: (Wagner 1908). Helicina ( Succincta) oweniana anozona Von Martens 1875Helicina anozona Von Martens 1875:649.Von Martens 1876:261; Helicina oweniana anozona Von Martens. Fischer & Crosse Alcadia (Leialcadia ) oweniana anozona (Von Martens). Wagner Oligyra oweniana var. anozona (Von Martens). Bequaert 1957; Bull. Mus. Comp. Zool. 115:206.Type Locality.Vicinity of Cobn, Alta Verapaz, Guatemala. Lectotype ZMB 25604a (Richling 2004:389). Distribution.GUATEMALA, Dept. Alta Verapaz: Cubilquitz. Dept. Baja Verapaz: Teleman (Von Martens 1890). CHIAPAS: Monte Libano, 600 m alt.; El Real, 600 m alt. (Bequaert 1957). Helicina ( Succincta) oweniana coccinostoma Morelet 1849Helicina coccinostoma Morelet 1849; Test. Noviss. I:19. Helicina oweniana coccinostoma Morelet. Von Martens 1890; Biol. 5, 5a (shell).Von Martens 1901; Biol. Cent. Amer.:605.Alcadia (Leialcadia) oweniana coccinostoma (Morelet). Wagner

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10 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Oligyra (Succincta ) oweniana coccinostoma (Morelet). H. B. Baker Bequaert 1957; Bull. Mus. Comp. Zool. 115:206.Type Locality. [Dept. Petn], Guatemala. Lectotype: BMNH 1893.2.4.1605 (Richling 2004:388). Distribution.GUATEMALA, Dept. Petn: Betalulen (Wagner 1908). CHIAPAS: Laguna Ocotal, 950 m alt. (Bequaert 1957). Helicina (Succincta) succincta Von Martens 1890Helicina succincta Von Martens 1890; Biol. Cent. Amer.:36; pl. 1, 55:782.Wagner 1910; in Martini & Chemnitz, Syst. Conch. Oligyra (Succincta ) arenicola succincta (Von Martens). H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:45. Helicina (Succincta) succincta Von Martens. H. B. Baker 1928; Mxico. Distribution.PUEBLA: Necaxa (H. B. Baker 1928). VERACRUZ: Texolo (Pilsbry 1903); Tlacolula; Cuesta de Misantla (Von Martens 1890). Subgenus Gemma Wagner 1905Gemma Wagner 1907:81.Richling 2004; Malacologia 45: 427 (not Gemma Deshayes 1853).Type Species.Helicina gemma Preston 1903. Distribution.Eastern Mxico and Central America. Taxonomy.We continue the use of the preoccupied subgeneric name Gemma, following Richling (2004), because of the uncertain taxonomic status of closely related subgenera. It may be decided that Gemma is synonymous with Succincta where H. B. Baker (1922) had placed the name. Twelve species and three subspecies are recognized in Gemma. Helicina (Gemma) beatrix beatrix Angas 1879Helicina beatrix Angas 1879; Proc. Zool. Soc. Lond. 47:484; pl. 43:332.Fischer & Crosse. 1893; Miss. Sci. Mex.:435. Pilsbry 1926b:127.Monge-Najera: 1997:113. var. Von Martens 1890; Biol. Cent. Amer.:39. var. beatrix Angas. Von Martens 1900; Biol. Cent. Amer.:606. Alcadia (Leialcadia) beatrix (Angas). Wagner 1908; in Martini & Oligyra (Succincta) beatrix beatrix (Angas). H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:45. Helicina (Oligyra) beatrix (shell).Pilsbry 1926b:127. Helicina (Gemma) beatrix bearix Angas. Richling 2004; Type Locality.Costa Rica, only on hills up to 2500 ft. alt. Lectotype BMNH 1879.7.22.79 (Richling 2004). Distribution.COSTA RICA; Caribbean mountain slopes of the Cordierra Central and the Cordierra de Talamanca Helicina (Gemma) beatrix confusa (Wagner 1908)Alcadia (Liealcadia) beatrix confusa 25 (shell). Oligyra (Succincta) beatrix confusa (Wagner). H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:45. Helicina (Gemma) beatrix confusa (Wagner). Richling 2004; 130 (map). Lectotype MIZ 8409 (Richling 2004). Distribution.COSTA RICA, the SE area of Prov. Helicina (Gemma) beatrix nicaraguae (Wagner 1908)Alcadia (Leiacadia) nicaraguae (shell). Helicina (Gemma) beatrix nicaraguae (Wagner). Richling 2004; Type Locality.Nicaragua. Lectotype MIZ 8408 (Richling 2004). Distribution.NICARAGUA: exact locality unknown Helicina (Gemma) beatrix riopejensis Richling 2004Helicina (Gemma) beatrix riopejensis Richling 2004; Malacologia Type Locality.SW of Liverpool, ca. 24 km W of Puerto from the immediate vicinity of the type locality. Helicina (Gemma) chiquitica (Richling 2001)Oligyra chiquitica Richling 2001; Schriften zur Malak.; Schriften Helicina (Gemma) chiquitica (Richling). Richling 2004; Type Locality.Approx. 9 km W of Matina, a little stream on the Rio Barbilla from the crossing of the road Holotype: UCR-INBio 3404977. Distribution.COSTA RICA, Prov. Cartago: Parque

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 11 W); Zona Protectora Rio Pacuare, Sector de La Estacion Heredia: S of Puerto Viejo de Sarapiqui, Zona Protectora La of Siquierres, along footpath stream up to Rio Pacuarito, 110 Helicina (Gemma) diaphana Pfeiffer 1852Helicina diaphana Pfeiffer 1852:98. Helicina (Gemma) diaphana Pfeiffer. Richling 2004; Malacologia Oligyra (Analcoadia) dysoni diaphana (Pfeiffer). H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:47.Type Locality.Honduras. Possible syntypes BMNH 196282 (Richling 2004). Distribution.HONDURAS: no other locality information given. Helicina (Gemma) escondida Richling 2004Helicina (Gemma) escondida Richling 2004; Malacologia 45; (distribution). 9 km W of Matina, a little upstream on the Rio Barbilla from W). Holotype UCR-INBio 354263. Distribution .COSTA RICA: known only from Cartago Turrialba, International American Agricultural Institute, 600 W); road along S bank of Rio Banano, opposite La Bomba Helicina (Gemma) fragilis fragilis Morelet 1851Helicina fragilis Morelet 1851; Test. Noviss. II:17.Von Martens 1890; Biol. Cent. Amer.:35.Fischer & Crosse 1893:420; pl. Amer.:605.Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:782. Helicina (Gemma) fragilis Morelet. Richling 2004; Malacologia Alcadia (Leialcadia) fragilis 9 (shell). Oligyra (Succincta) fragilis fragilis (Morelet). H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:45.Type Locality.Sylvas Petenenses, Dept. Petn, Guatemala. Lectotype BMNH 1893.2.4.809 (Richling 2004). Distribution.GUATEMALA, Dept. Alta Verapaz: Cobn (Wagner 1908). Dept. Baja Verapaz: Teleman, in the valley of the Rio Polochic; Purula, 4000 ft. alt. (Von Martens 1890). GUERRERO: Omilteme, 8000 ft. alt. (Von Martens 1890). VERACRUZ; Orizaba (Von Martens 1901). Helicina (Gemma) fragilis elata Shuttleworth 1852Helicina elata Shuttleworth 1852; in Bern Mittheil.:304. Helicina (Gemma) elata Shuttleworth. Richling 2004; Malacologia Helicina fragilis Morelet (in part). Von Martens 1890; Biol. Cent. Amer.:35. Oligyra (Succincta ) fragilis elata (Shuttleworth). H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:45. Helicina (Succincta) fragilis elata Shuttleworth. H. B. Baker 1928; (female reproductive anatomy). Distribution.VERACRUZ: Atoyac to Sumidero (H. B. Baker 1928). Helicina (Gemma ) gemma Preston 1903Helicina gemma Alcadia (Leialcadia) gemma 17 (shell). Oligyra (Succincta) gemma (Preston). H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:45. Helicina (Gemma) gemma Preston. Richling 2004; Malacologia (Richling 2004). Distribution.COSTA RICA; generally distributed throughout the Caribbean slope of the northeastern sector of the country below 1200 m alt. NICARAGUA. Dept. Zelaya Norte: Cerro Salaya, Bosawas (Richling 2004). Helicina (Gemma) merdigera Pfeiffer 1855Helicina merdigera Pfeiffer 1855:102. Helicina (Gemma) merdigera Pfeiffer. Richling 2004; Malacologia Helicina fragilis Morelet (in part). Von Martens 1890; Biol. Cent. Amer.:35. Oligyra (Succincta ) fragilis merdigera (Pfeiffer). H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:45.Type Locality.Veracruz [City], [Veracruz], Mxico. Lectotype BMNH 20010752.1 (Richling 2004). Distribution.Known only from the type locality. Helicin a (Gemma) mohriana Pfeiffer 1861.Helicina mohriana Pfeiffer 1861; Malak. Bltt. 8:172. Helicina (Gemma) mohriana Pfeiffer. Richling 2004; Malacologia 45:334. Alcadia (Leialcadia ) mohriana (Pfeiffer). Wagner 1908:85; pl. 14, Type Locality.Orizaba, Mxico. Type specimens lost (Richling 2004). 1908).

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12 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Helicina (Gemma) Richling 2004Helicina (Gemma) Richling 2004; Malacologia 209 (distribution).Type Locality.Cloud forest, Sendero Bosque Nuboso Monteverde, Reserva Biologica Bosque Nebosos, Cordillera Tilarn near Monterverde, Prov. Puntarenas, Costa Rica. Distribution.COSTA RICA: numerous localities in Alajuela, Guanecaste, and Puntarenas Provinces in the immediate vicinity of the type locality. Prov. Guanacaste:6 mi. NNE of Tilarn (ca. 10 N, 84 W); Prov. San Jos: Helicina (Gemma) talamancensis (Richling 2001)Oligyra talamancensis Richling 2001; Schriften zur Malak.:3; Helicina (Gemma) talamancensis (Richling). Richling 2004; Type Locality.Primary forest bordered by secondary growth, Fila Costea, north of Bajo Bonta, north of Rio Distribution.COSTA RICA: Prov. Puntarenas and Prov. San Jos Know from numerous localities in the southcentral region of the Cordillera Talamanca (Richling 2004). Helicina (Gemma) terryae Rehder 1940Helicina terryae Rehder 1940:350; Jour. Wash. Acad. Sci. 32:350; Helicina (Gemma) terryae Rehder. Richling 2004; Malacologia Type Locality .Prov. Chiriqu, Panam. Holotype USNM 536026 (Richling 2004). Distribution.Known only from the type locality. Subgenus Tristramia Crosse 1863Tristramia Crosse & Fischer 1863. Caloplisma Crosse & Fischer in Fischer & Crosse 1893:405. Retorquata Wagner 1905; Denksch. Kaiserli. Akad. 78:230. Turbinata Wagner 1905; Denksch. Kaiserli. Akad. 42:218, 306. Rostrata Wagner 1905; Denksch. Kaiserli. Akad. 42:218.Type Species.Tristramia: type by monotypy Helicina Tristram 1861 (= Helicina rostrata Morelet 1849. Caloplisma: type by monotypy Helicina rostrata Morelet 1849. Retorquata: Helicina zephyrina species). Turbinata: type by tautonomy Helicina turbinata Pfeiffer 1846. Rostrata: type by tautonomy Helicina rostrata Morelet 1849. Distribution.Mxico and south to Colombia. Taxonomy.Numerous species. Eighteen species and seven subspecies are recognized in Mxico and Central America. Helicina ( Tristramia) chrysocheila chrysocheila Binney 1851Helicina chrysocheila Martens 1890; Biol. Cent. Amer.:33.Pilsbry 1948:1081 Trop. 41:685.Correa-Sandoval, Garca-Cubas & Reguero 1998; Acta Zool. Mex. (73):13.Correa-Sandoval 2000; Acta Zool. Mex. (79):8.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):235.Correa-Sandoval & Salazar 2005; Acta Zool. Mex. (21) 59. Helicina (Tristramia) chrysocheila chrysocheila Binney. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:51.Type Locality.No locality given. Distribution.NUEVO LEON: numerous localities in southern Nuevo Leon (Correa-Sandoval 2005). SAN LUS POTOS: numerous localities in southeastern part of state (Correa-Sandoval et al. 1998); 0.1 km W of Ejido Buenavista (Correa-Sandoval et al. 1998). TAMAULIPAS: numerous localities (Correa-Sandoval & Rodriguez 2002, 2005)). VERACRUZ: numerous localities in the northern part of state (Correa-Sandoval 2000). Helicina (Tristramia) chrysocheila schuttleworthi Von Martens 1890Helicina chrysocheila Shuttleworth 1852; in Bern Mittheil.:303.(Shell). Helicina chrysocheila var. shuttleworthi Von Martens 1890; Biol. Helicina (Tristramia) chrysocheila schuttleworthi Von Martens. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:51. Type Locality.Cordova, Veracruz, Mxico. Holotype NMBE 15267. Distribution.Known only from the type locality. Taxonomy.There seems to be little basis for recognizing shuttleworthi as distinct from H. chrysocheila. Helicina (Tristramia) delicatula Shuttleworth 1852Helicina delicatula Shuttleworth 1852; in Bern Mittheil.:303.Von Martens 1890; Biol. Cent. Amer.:37.Wagner 1910; in Martini & Chemnitz, Syst. Conch. Cab., Helicinidae:3101; pl. 62, Helicina (Tenuis) delicatula delicatula Shuttleworth. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:50. Helicina (Oligyra) delicatula Shuttleworth. H. B. Baker 1928; Occ. Helicina heloisiae Pfeiffer 1856:322.Sowerby 1866; Thes. Conch. Holotype: NMBE 19074. Martens 1890).

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 13 Helicina (Tristramia) durangoana durangoana Mousson 1883Helicina durangoana Von Martens 1890; Biol. Cent. Amer.:37.Fischer & Crosse in Martini 4 (shell). Helicina (Tenuis) durangoana durangoana Mousson. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:50.Type Locality.Sierra Madre, Durango, Mxico. Distribution.DURANGO: Ventanas (Von Martens 1890). Helicina ( Tristramia) durangoana sagulensis Wagner 1910Helicina durangoana sagulensis Wagner 1910; in Martini & (shell). Helicina (Tenuis) durangoana sagulensis Wagner. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:50.Type Locality.Sagula, Jalisco, Mxico. Distribution.Known only from the type locality. Helicina (Tristramia) echandiensis Richling 2004Helicina echandiensis Richling 2004; Malacologia 45:271; (female reproductive anatomy).Type Locality.Parque Nacional de Amistad, Sector Las Alturas, southern Cordillera de Talamanca, S of Cerro INBio 3542520. Distribution.Known only from the type locality. Helicina (Tristramia) elatior Von Martens 1890Helicina turbinata (shell). Helicina zephyrina var. elatior Von Martens 1890; Biol. Cent. Amer.:31. Helicina elatior Von Martens. Dall 1905; Smiths. Misc. Coll. 48:48:192.Type Locality.Not stated. Distribution.SAN LUIS POTOS: Sierra Alvarz; San Dieguito (Dall 1905). VERACRUZ: Papantla; Misantla; Helicina (Tristramia) Pilsbry 1930Helicina (Tristramia) Pilsbry 1930:248, Type Locality.Isla de Providencia, ridge north of High Peak and valley running westward. Holotype ANSP 150717. Distribution.Known only from the type locality. Helicina (Tristramia) funcki Pfeiffer 1849Helicina funcki Pfeiffer 1849; Proceedings of the Zoological Society of London, 16 (1848):121.Pfeiffer 1850:33; pl. 9, Helicina : pl. 17, Ancey 1897:87.Von Martens 1900; Biol. Cent. Amer.:603 604.Wagner 1910; in Martini & Chemnitz, Syst. Conch. 1920a:3.Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:59, 69, 78:127.Pres 1994:746.Monge-Njero 1997:113. Helicina (Retorquata) funcki Pfeiffer. Wagner 1905; Denksch. Kaiserli. Akad. :232. Helicina (Retorquata) funcki costaricensis Wagner 1905; Denksch. Wagner 1910; in Martini & Chemnitz, Syst. Conch. Cab., Proc. Acad. Nat. Sci. Phila. 74:51. Pfeiffer. Pilsbry 1920a:3. Helicina (Tristramia) Pilsbry. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:51. Helicina (Tristramia) funcki funcki Pfeiffer. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:51.H. B. Baker 1926; Proc. Acad. male reproductive systems). Helicina (Tristramia) funcki Pfeiffer. Richling 2004; Malacologia (Costa Rica, distribution).T ype Localities. Helicina funcki : San Yago [Santiago], Prov. Veraguas, Panam; lectotype: BMNH 20010497.1 (Richling 2004). Helicina funcki costariceneis : Sta. Clara Costa Rica; 250 m alt.; lectotype MIZ 8989a (Richling 2004). : Juan Vias, Prov. Cartago, holotype ANSP 105286. Distribution.COSTA RICA: numerous localities up to about 1600 m in the Provs. Alajuela, Cartago, Guanacaste, NICARAGUA, Dept. Rio San Juan: Greytown (Ancey 1897). PANAM: numerous localities in Depts. Bocas del Santiago (Von Martens 1890). Helicina (Tristemia ) hondurana Richards 1938Helicina hondurana Richards 1938; Proc. Amer. Philos. Soc. Type Locality.West End, Isla de Roatn, Dept. Islas de la Baha, Honduras.Holotype ANSP 17019. Distribution.HONDURAS: known only from Isla de Roatn. Helicina (Trisramia) notata Pfeiffer 1856Helicina delicatula notata Pfeiffer 1856:323.Von Martens 1890; 10b, 111a (shell).Wagner 1910; in Martini & Chemnitz, Helicina (Tenuis) tenuis notata Pfeiffer. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:50. Helicina (Tristramia) notata Pfeiffer. H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):27.Typ Distribution.VERACRUZ: Atoyac (Von Martens 1890).

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14 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Helicina (Tristramia) pitalensis Wagner 1910Helicina pitalensis Wagner 1910; in Martini & Chemnitz, Syst. Helicina (Tristramia) pitalensis Wagner. Richling 2004; Malacologia (distribution). Helicina funcki Pfeiffer. Von Martens 1900; Biol. Cent. Amer.:33 (in part). Helicina (Tristramia) funcki pitalensis Wagner. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:51 Helicina amoena Pfeiffer. Monge Njero 1997:113. 103240. Distribution.COSTA RICA, numerous localities in Alajuela and Puntarena provinces (Richling 2004). Helicina (Tristramia) punctisulcata punctisulcata Von Martens 1890Helicina punctisulcata (shell).Wagner 1910; in Martini & Chemnitz, Syst. Conch. 2004; Malacologia 45:277; pl. 87 (shell). Helicina (Oxyrhomba) punctisulcata punctisulcata Von Martens. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:48.Type Locality.Omilteme, Sierra Madre del Sur, Guerrero, Mxico. Lectotype ZMB 103326a (Richling 2004). Distribution.Known only from the type locality. Helicina (Tristramia) punctisulcata cuericensis Richling 2004Helicina punctisulcata cuericensis Richling 2004; Malacologia Type Locality.Estacion Cuerici, 4.5 km E of De Villa Mills, Sendero El Mirador, Cordillera de Talamanca, Prov. UCR-INBio 3542622. Distribution.COSTA RICA: known only from the immediate vicinity of the type locality. Helicina (Tristramia) punctisulcata zunilensis Wagner 1910Helicina punctisulcata zunilensis Wagner 1910; in Martini & Helicina (Oxyrhomba) punctisulcata zunilensis Wagner. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:48.Type Locality.Volcn Zunl on the border of Quetzaltenango and Solol departments, Guatemala. Holotype ZMB 103324 (Richling 2004). Distribution.Known only from the type locality. Helicina (Tristramia) rostrata rostrata Morelet 1849Helicina rostrata Morelet 1849; Test. Noviss. I:17.Tate 1870; Amer. Jour. Conch. 5:159.Sowerby 1866; Thes. Conch. Biol. Cent. Amer.:30.Fischer & Crosse 1893:465; pl. 55, Wagner 1910; in Martini & Chemnitz, Syst. Conch. Cab., Lopz 2002: 45. Helicina (Tristramia) rostrata rostrata Morelet. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:51. Tristram 1861; Proc. Zool. Soc. Lond. 29:233; pl. Helicina rostrata var. simplex Fischer & Crosse 1893:465; pl. 55, Type Localities. Helicina rostrata: San Agustin Lanquin, Alta Vera Paz, Guatemala. : Guatemala. Helicina rostrata var. simplex: not stated. Distribution.GUATEMALA, Dept. Alta Verapaz: Cobn NICARAGUA, Dept. Chontales: Acoyapa (Von Martens 1890). Dept. Managua: San Diego (Tate 1870); Managua (Von Martens 1901). Reg. Autonono Atlantico Sur: Isla del Maz (Richards 1939). Depts. Chontales, Boaco and Helicina (Tristramia) rostrata denticulata Pfeiffer 1855Helicina denticulata Pfeiffer 1855; Proc. Zool. Soc. Lond. 23:103.Tate 1870; Amer. Jour. Conch. 5:159.Sowerby 1866; Thes. Biol. Cent. Amer.:30.Wagner 1910; in Martini & Chemnitz, Helicina (Tristramia) rostrata denticulata Pfeiffer. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:51.Type Locality.San Pedro [San Pedro Sula], Honduras. Distribution.Known only from the type locality. NICARAGUA: forests of Chontales (Von Martens 1890). Helicina (Tristramia) rostrata matagalpensis Wagner 1910Helicina rostrata matagalpensis Wagner 1910; in Martini & 19 (shell). Helicina (Tristramia) rostrata matagalpensis Wagner. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:51.Type Locality.Matagalpa, Nicaragua. Distribution.Known only from the type locality. Helicina (Tristramia) senachuensis Wagner 1905 Wagner 1905; Denksch. Kaiserli. Helicina senachuensis 7 (shell). Helicina (Tenuis) senachuensis Wagner. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:50. Type Locality.Senachu, Guatemala, Distribution.Known only from the type locality. I have been unable to locate Senachu. There is a locality in Dept. Alta Verapaz named Senahu (15.4 N, 89.8 W). Helicina (Tristramia) sinuosa Pfeiffer 1850Helicina sinuosa Pfeiffer 1850; Zeitschr. Malak.:191.Von Martens

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 151890; Biol. Cent. Amer.:32.Wagner 1910; in Martini & Chemnitz, Syst. Conch. Cab., Helicinidae:297; pl. 59, Type Locality.Mxico. Distribution.Unknown. Remarks.Status uncertain; this may be synonymous with Helicina zephyrina Duclos 1833. Helicina (Tristramia) tenuis Pfeiffer 1849Helicina tenuis Pfeiffer 1849; Proc. Zool. Soc. Lond. (1848):124 125.Von Martens 1890; Biol. Cent. Amer.:34.Fischer Von Martens 1901; Biol. Cent. Amer.: 604.Wagner 1910; in Martini & Chemnitz, Syst. Conch. Cab., Helicinidae:302; Mus. Zool. Univ. Mich. (186):35.Pilsbry 1926a:339.Von Martens 1901; Biol. Cent. Amer.:605.Goodrich & Van der Schalie 1937; Misc. Pub. Mus. Zool. Univ. Mich. (34):12, 15, 32.Van der Schalie 1940; Occ. Pap. Mus. Zool. Univ. Mich. (413):6, 9, 10.Haas 1949; Nautilus 62:137, 138.Thompson 1957; Nautilus 70:100.Bequaert 1957; Bull. Mus. Comp. Zool. 115:207. Helicina (Tenuis) tenuis tenuis Pfeiffer. H. B. Baker 1922; Proc. Helicina tenuis tenuis Pfeiffer. Thompson 1967; Bull. Fla. St. Mus. 11:228. Morelet 1849; Test. Noviss. I:20.Sowerby 1866; Conch. Icon., Helicina (shell). Helicina chiapensis Pfeiffer 1856.Sowerby 1866; Thes. Conch. Helicina Helicina tenuis chiapensis Pfeiffer. Pilsbry 1892:339. ?Helicina lindeni Tristram 1862:5.Sowerby 1866; Thes. Conch. Pub. Carnegie Inst. Wash. (431):543. Helicina (Oligyra) lindeni Tristram. Fischer & Crosse 1893:416 Helicina tenuis lindeni Pfeiffer. Hinkley 1920; Nautilus 34:49, 52. Wagner 1905; Denksch. Kaiserli. Helicina teunis pitieri Wagner 1910; in Martini & Chemnitz, Syst. Helicina (Tristramia) tenuis Pfeiffer. Richling 2004; Malacologia Type Localities. Helicina tenuis: Yucatn; lectotype BMNH 20010496 (Richling 2004). : Dept. Petn, Guatemala; lectotype BMNH 1893.2.4.1991 (Fischer & Crosse 1893). Helicina chiapensis: Chiapas, Mxico; syntypes ZMB 65624. Helicina lindeni : Mxico. Helicina tenuis pittieri : Rio de los Plataneros S of : Verapaz, Guatemala. Distribution.Panam north to Guerrero, Mxico on Most commonly encountered at lower elevations below 1500 m alt. PANAM: Isla Barro Colorado (Pilsbry 1930). COSTA RICA: numerous localities. HONDURAS, Dept. Dept. Olancho: vicinity of Magua Cave, 15 km SSW of Ahuachapn: 6 km W of Atiquizaya, on road to Ahuachapn. GUATEMALA: numerous localities. CAMPECHE: 16.4 km E of Escarciga. CHIAPAS: 15 1. km W of San Cristobal, 2669 m alt.; 18.3 km N of Tuxtla Gutierrez, 1372 m alt.; 12.9 km N of Tuxtla Gutierrez, 1158 m alt.; 4.8 km SSE of Tuxtla Gutierrez, 822 m alt.; 7.5 km NNE of Huixtla 183 m alt.; 21.3 km NW of Huixtla, 300 m alt.; 44.4 km NW of Ocozocoautla, 610 m alt.; 24.5 km NW of Ocozocoautla, 823 m alt.; 34.1 km E, 16.4 km S of Comitan, 1524 m alt.; Ruins of Palenque. GUERRERO: 1 km E of Petaquillas, 1158 m alt.; 2.2 km NNE of Mazatln, 1475 m alt.; limestone hill 1 W). OAXACA: Lagunas, 259 m alt.; limestone ridge 4 km VERACRUZ: 5 km ENE of Cuautemoc, Oaxaca, 75 m alt. 350 m alt.; Laguna Encantada; limestone knoll 2 km SW of The above records are from Richling (2004). The following records are given by other authors: GUATEMALA, Dept. Alta Verapaz: Cobn; Cubilguitz (Von Martens 1890). Dept. Baja Verapaz: Panzos; Chacoj; San Juan; Purula (Von Martens 1890). Dept. Quetzaltenango; Cerro Zunil; San Isidro; El Reposa; Zapote (Von Martens 1890). Dept. Totonicapan: Totonicapam Mountains (Von Martens 1890). CHIAPAS: El Real, 600 m alt.; Monte Libano, 600 m alt. (Bequaert 1957). TABASCO: San Juan Bautista; Teapa (Von Martens 1890); Poan (Von Martens 1901); 0.5.0 mi. E of Teapa (Thompson 1957). VERACRUZ: Soledad (Von Martens 1890). Helicina (Tristramia) trossula Morelet 1849Helicina trossula Morelet 1849; Test. Noviss. I:19.Wagner 1910; in Martini & Chemnitz, Syst. Conch. Cab., Helicinidae:311; (in part). Von Martens 1890; Biol. Cent. Amer.:39.Type Locality.Petn, Guatemala. Distribution.Known only from the type locality. Helicina (Tristramia) turbinata Pfeiffer 1848Helicina turbinata Pfeiffer 1848; Zeitschri. Malak. 5:87.Wagner 1910; in Martini & Chemnitz, Syst. Conch. Cab., Proc. Acad. Nat. Sci. Phila. 55:781. Helicina zephyrina Menke. Von Martens 1890; Biol. Cent. Amer.:30 (in part). Helicina zephyrina var. Von Martens 1890; Biol.

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16 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Cent. Amer.:31.Type Localities. Helicina turbinata: Jalapa, Veracruz, Mxico. Helicina zephyrina var. : Veracruz, Mxico. Distribution.VERACRUZ: Antigua; Misantla; 1910). Helicina (Tristramia) Pilsbry 1910 Pilsbry 1910; Proc. Acad. Nat. Sci. Phila. 61 Helicina (Tristramia) chysocheila Pilsbry. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:51.Correa-Sandoval, Garca-Cubas & Reguero 1998; Acta Zool. Mex. 73:13.Correa-Sandoval 2000; Acta Zool. Mex. 79:8.CorreaSandoval & Rodriguez 2002; Acta Zool. Mex. (86):235. Type Locality.On the sides of the canyon below Los Canoas, San Lus Potos. Holotype ANSP 99025. Distribution.SAN LUS POTOS: eastern part of state; sides of the highest peak S of Mecos Falls; Cascadas Micos Cueva El Salitre, Xilitla, numerous loclities in the region; Sandoval et al.. 1998). Numerous localities in southern TAMAULIPAS (Correa-Sandoval & Rodriguez 2002 2005). VERACRUZ: San Juan Cuajinampa, La Ordea, Papantla (Correa-Sandoval 2000).. Helicina (Tristamia) zephyrina zephyrina Duclos 1833Helicina zephyrina Duclos 1833; in (shell).Von Martens 1890; Biol. Cent. Amer.:30 (in part).Martens 1901; Biol. Cent. Amer.:603.Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:782.Correa-Sandoval, Garca-Cubas & Reguero 1998; Acta Zool. Mex. (73):13.Correa-Sandoval 2000; Acta Zool. Mex. (79):8.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):235. Helicina (Tristamia) zephyrina zephyrina Duclos. H. B. Baker 13 (radula).H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):32; pl. 6 (female reproductive anatomy). Helicina sandozi Shuttleworth 1852; in Bern Mittheil.:303.Neubert since its original description; a nomen oblitum). Helicina behrendti 14 (shell).Type Localities. Helicina zephyrina: Vera Cruz. Helicina sandozi: Mxico. Syntypes: NMBE 19039/1. Helicina behrendti : not stated. Distribution.AXACA: Tustepec [Tuxtepec] (Von Martens 1901). SAN LUS POTOS: numerous localities in eastern part of state (Correa-Sandoval et al. 1998); 3 km E of W); Barrio San Rafael, Ro Moctezuma, Tamazunchale numerous localities in southern part of state (Correa-Sandoval & Rodriguez 2002); El Cielo Biosphere Reserve (CorrreaSandoval & Rodriguez 2005); Tampico (Hinkley 1907). VERACRUZ: numerous localities in northern part of state Huatusco (Von Martens 1890); Texolo (Pilsbry 1903). Helicina ( Tristramia) zephyrina deppeana Von Martens 1863Helicina deppeana Von Martens 1863; Monatsb. Knig. Preuss. Akad. Wiss. Berlin:540.Sowerby 1866; Thes. Conch. 3:298; in Martini & Chemnitz, Syst. Conch. Cab., Helicinidae:309; pl. 62, Helicina (Tristramia) zephyrina deppeana Von Martens. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:51.Type Locality.Mxico. Distribution.OAXACA: Yalalag, south of Villa Alta (Von Martens 1890). VERACRUZ: Jalapa; Soledad (Von Martens 1890). Helicina (Tristramia) zephyrina dientensis Pilsbry 1903Helicina zephyrina dientensis Pilsbry 1904; Proc. Acad. Nat. Sci. Helicina zephyrina dientensis Pilsbry. H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:51. Mxico. Holotype ANSP 77238. Distribution.Known only from the type locality. Genus Alcadia Gray 1840 Type Species.Helicina major Gray 1824. Distribution.The West Indies and Central America. Taxonomy.Richling (2003) recognizes three subgenera. Alcadia s. s. and Idesa H. Adams. & A. Adams Microalcadia is endemic to Central America. Subgenus Microalcadia Richling 2004Microalcadia Richling 2004; Malacologia 45:370.Type Species.Helicina hojarasca Richling 2004. Distribution.Costa Rica. Taxonomy.Three species are referred to the subgenus.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 17 Alcadia (Microalcadia ) boeckeleri (Richling 2001)Helicina boeckeleri (operculum), 101, 12B (shell). Alcadia (Microlcadia) boeckeleri (Richling). Richling 2004; Type Locality.Primary forest, Parqu Nacional Guanacaste, about 10 km S of Santa Cecilia, Volcn Oros, 3404980. Distribution.COSTA RICA: known only from the immediate vicinity of the type locality. Alcadia (Miscroalcadia) hojarasca (Richling 2001)Helicina hojarasca (operculum), 7, 11A (shell). Alcadia (Microalcadia) hojarasca (Richling). Richling 2004; Type Locality.Cordillera de Tilarn, about 9 km N of Santa Elena, near Mirador Gerardo, Guanacaste Province, UCR-INBio 3404389. Distribution.COSTA RICA: in addition to the type locality the species has been recorded from one other locality. Puntarens Prov.: Zona Protectora Arenal-Monteverde, 1500 Alcadia (Microalcadia ) exigua (Pfeiffer 1849)Helicina exigua Pfeiffer 1849; Proc. Zool. Soc. Lond. (1848):121. ?Alcadia (Micoalcadia ) exigua (Pfeiffer). Richling 2004; Malacologia 45:374.Type Locality.Honduras. Distribution.Central America. In 1849 Honduras included most of the Caribbean coast of Central America from the Yucatn Peninsula south to Costa Rica and Panam. Genus Lucidella Swainson 1840Lucidella Swainson 1840, Richling 2004; Malacologia 45:194, 330.Type Species.Helix aureola Frussac 1822. Distribution.South Florida, the West Indies, and the Caribbean region of the South American mainland from French Guyana north to Veracruz, Mxico. Taxonomy.Four subgenera are recognized. Lucidella s.s. and Poeniella Indies region. Lidsleya Chitty 1857 is found in Jamaica and on the South American mainland (H. B. Baker 1922b). Poenia H. & A. Adams 1856, is found on the mainland from French Guyana north to Mxico and on some near-by mainland islands in the Caribbean region. Subgenus Poenia H. Adams & A. Adams 1856Poenia H. Adams & A. Adams 1856:304 (type species: Helicina unidentata Pfeiffer 1847). Perenna Guppy 1867:260 (type species: Helicina lamellosa Guppy 1867).Distribution.French Guyana and coastal islands north to Veracruz, Mxico. Taxonomy.Five species are recognized. One is widely distributed on the mainland, one occurs on Isla de Roatn, one occurs on Islas del Cisne (Lucidella pilsbryi Clapp 1914), one occurs on Cuba (Lucidella granum Pfeiffer 1856), and one occurs on Grand Cayman Island (Lucidella caymanensis Pilsbry 1930). Lucidella (Poenia) lirata (Pfeiffer 1847)Helicina lirata Pfeiffer 1847; Zeitschr. Malak., 4:150.Pfeiffer 1850; in Martini & Chemnitz, Syst. Conch. Cab., Helicina :14; pl. Helicina (Poenia ) lirata Pfeiffer. Fischer & Crosse 1893:397. Lucidella lirata (Pfeiffer). Wagner 1911; in Martini & Chemniz, Syst. Conch. Cab., Lucidella Hinkley 1920; Nautilus 34:41, 49. 52.Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:59, 71.Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:127.Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. 82:339.Goodrich & Van der Schalie 1937; Misc. Pub. Mus. Zool. Univ. Mich. (34):12, 14, 33.Richards 1938; Proc. Amer. Philos. Soc. 79:176.Bequaert 1957; Bull. Mus. Comp. Zool. 115:208.Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):8.Haas & Solem 1960; Nautilus 73:130.Branson & McCoy 1963; Nautilus 76:102.Thompson Costa Rica). Lucidella (Poenia ) lirata (Pfeiffer). H. B. Baker 1922; Proc. Acad. B. Baker 1922b; Occ. Pap. Mus. Zool. Univ. Mich. (186):36.H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):33 Clench 1933; Pub. Carnegie Inst. Wash. (431):543. Helicina unidentata Pfeiffer 1849:125.Sowerby 1866; Thes. Helicina lirata var unidentata Pfeiffer. Von Martens 1891 41, 607. Helicina (Perena) lamellosa Lucidella lirata lamellosa Wagner 1911; in Martini & Chemnitz, Syst. Conch. Cab., Lucidella Helicina rusticella Morelet 1849; Test. Noviss. I:21. Helicina lirata var. rusticella Morelet. Von Martens 1891:41.Fischer & Crosse 1893:397.Von Martens 1900:607. Helicina semistriata Sowerby 1866; Thes. Conch. 3:281; pl. 268, Helicina lirata var. semistriata Sowerby. Von Martens 1891:41.Type Localities. Helicina lirata: Yucatn, Mxico; primary type destroyed (Boss & Jacobson 1974). Helicina unidentata : Honduras; type probably lost (Boss & Jacobson 1974:37). Helicina rusticella: Isla de Carmen, Mxico; syntype BMNH 1893.2.4. Helicina semistriata: not given; types destroyed (Bosse & Jacobson 1974). Helicina lamellosa: Gulf of Paria, Trinidad; synty pe BMNH 1874.10.30.12.

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18 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Distribution.Widely distributed along the Caribbean Coast from French Guiana to central Veracruz, Mxico, and on coastal islands. Central American and Mexican records follow. PANAM: Tabernillo, Canal Zone; near Darin; Juan Minas; Cd. Panam; Isla Taboga (Pilsbry 1926a); Bocas del Toro (Von Martens 1891). COSTA RICA: numerous localities 2004). NICARAGUA, Depts. Boaco, Estel, Matagalpa, Rivas and Rio San Juan Matagalpa: 4.5 km S of Matagalpa, 1200 m alt. (Richling 2004). HONDURAS: Isla de Roatn (Richards 1938); Isla de tila (Pilsby 1891). GUATEMALA, Dept. Izabal: Jocola (Hinkley 1920). Dept. Alta Verapaz: Cham (Hinkley 1920). Dept. Petn: Tikal (Basch 1959). BELIZE, Cayo District: Rio Frio Cave (Haas & Solem 1960). QUINTANA ROO: 4 mi. E of Xpujil; 7.1 m. NNW of Xiatil (Thompson 1967). YUCATN: Chichen Itza (Bequaert & Clench 1933); Labna (Pilsbry 1901). CAMPECHE: Ciudad del Carmen (Branson & McCoy 1963); numerous localities (Thompson 1967). TABASCO: Teapa; San Juan Bautista (Von Martens 1891); 0.5.0 mi. E of Teapa (Thompson 1957). CHIAPAS: Selva Lacondona, Laguna Ocotal, 950 m alt.; Laguna Ocotal to El Censo, 1000 m alt. (Bequaert 1957). YUCATN. Lucidella (Poenia) midyetti Richards 1938Lucidella midyetti Richards 1938; Proc. Amer. Philos. Soc. 79:175 Type Locality.Between Coxen Hole and French Harbor, Isla de Roatn, Dept. Islas de la Baha, Honduras. Holotype ANSP 170021. Distribution.HONDURAS: known only from Isla de Roatn. Lucidella (Poenia) pilsbryi pilsbryi Clapp 1914Lucidella pilsbryi (shell).Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. 82:243; pl. Type Locality.Islas del Cisne, Dept. Islas de la Baha, Honduras. Holotype MCZ 22890. Distribution.Known only from the type locality. Lucidella (Poennia) pilsbryi indecora Pilsbry 1930Lucidella (Poennia) pilsbryi indecora Pilsbry 1930; Proc. Acad. Type Locality.Isla del Cisne Pequeo, at the western third a short distance from the north shore, Dept. Islas de la Baha, Honduras. Holotype ANSP 150881. Distribution.Known only from the type locality. Genus Schasicheila Shuttleworth 1852Schasicheila Shuttleworth 1852; in Bern Mittheil.:301.Von Martens 1890; Biol. Cent. Amer.:43, 608.Wagner 1911; in Martini & Chemnitz, Syst. Conch. Cab., Schasicheila :351.H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:56.H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):36.Richling 2002:408.Type Species.Helicina nicoleti Shuttleworth 1852. Distribution.Eastern Mxico from Tamaulipas south to Guatemala. Taxonomy.The genus is divided into three subgenera and includes ten species and two subspecies. Subgenus Atoyac H. B. Baker 1928Atoyac H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):36.Type Species.Helicina alata Pfeiffer 1849. southern Tamaulipas, southeastern San Luis Potosi and Veracruz. Taxonomy.Three species are recognized. Schasicheila (Atoyac ) alata (Pfeiffer 1849)Helicina alata Pfeiffer 1849; Proc. Zool. Soc. Lond. 17:87. Schazichila alata Schasicheila alata (Pfeiffer). Von Martens 1890; Biol. Cent. Amer.:43.Von Martens 1901; Biol. Cent. Amer.:608.Wagner 1911; in Martini & Chemnitz, Syst. Conch. Cab., Schasicheila 1922; Proc. Acad. Nat. Sci. Phila. 74:57.Richling 2004; (female reproductive anatomy). Schasicheila (Atoyac) alata Pfeiffer. H. B. Baker 1928; Occ. anatomy).Type Locality.Cord Distribution.CHIAPAS: Monte Libano, 600 m alt. (Bequaert 1957; Bull. Mus. Comp. Zool. 115). VERACRUZ: Mizantla (Strebel 1873); Mirador; San Cristobal; Atoyac; Jalapa (Von Martens 1890); Orizaba (Von Martens 1901). Schasicheila (Atoyac) fragilis Pilsbry 1899Schasicheila fragilis Pilsbry 1899; Proc. Acad. Nat. Sci. Phila. 51:391.Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:782 Chemnitz, Syst. Conch. Cab., Schasicheila 20 (shell).H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:57.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):235.Correea-Sandovl & Salazar 2005; Acta Zool. Mex. (21):59. Schasicheila (Atoyac) fragilis Pilsbry. H. B. Baker 1928; Occ. Type Locality.Diente, Mxico. Holotype ANSP 77237. Distribution.NUEVO LEN: Laguna de Sanchez (25 Salazar 2005). TAMAULIPAS: Victoria-Gomez Farias

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 19 Schasicheila (Atoyac ) xanthia Pilsbry 1909Schazicheila xanthia Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:57.CorreaSandoval, Garca-Cubas & Reguero 1998; Acta Zool. Mex. (73):13. Schasicheila (Atoyac) xanthia Pilsbry. H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):36.Type Locality.Canyon below Los Canoas, San Lus Potos. Holotype ANSP 99024. Distribution.SAN LUS POTOS: Cueva El Salitre, 6 (Correa-Sandoval et al. 1998). Subgenus Necaxia H. B. Baker 1928Necaxia H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):36.Type Species.Helicina minuscula Pfeiffer 1859. Distribution.Guatemala north to San Lus Potos. Taxonomy.A single species is recognized. Schasicheila (Necaxia) minuscula (Pfeiffer 1859)Helicina minuscula Pfeiffer 1859; Proc. Zool. Soc. Lond. 27:29. Schasicheila minuscula (Pfeiffer). Fischer & Crosse 1893:452; pl. Sci. Phila. 74:57.Correa-Sandoval, Garca-Cubas & Reguero 1998; Acta Zool. Mex. (73):13.Correa-Sandoval 2000; Acta Zool. Mex. (79):8. Schasichila minuscula (Pfeiffer). Von Martens 1901; Biol. Cent. Amer.:608. Schasicheila (Necaxia) minuscula (Pfeiffer). H. B. Baker 1928; Occ. Schasicheila minima (Pfeiffer). Strebel 1880; Beitrag. Mex. Type Locality.Misantla, Veracruz, Mxico (H. B. Baker 1928). Distribution.GUATEMALA, Dept. Alta Verapaz: Chama (H. B. Baker 1928). SAN LUS POTOS: 4925 m alt. (H. B. Baker 1928); numerous localities in northern Veracruz (Correa-Sandoval 2000). Subgenus Schasicheila Shuttleworth 1852 Distribution.Eastern Mxico from Tamaulipas south to Alta Verapaz, Guatemala. Taxonomy.Nine species and two subspecies are recognized. Schasicheila (Schasicheila) hidalgoana Dall 1897Schasicheila hidalgoana Dall 1897; Nautilus 11:62.Pilsbry 1904; (shell).Hinkley 1907; Nautilus 21:71.H. B. Baker 1922; H. B. Baker 1926; Proc. Acad. Nat. Sci. Phila. 76:49; pl. 8, Cubas & Reguero 1998; Acta Zool. Mex. (73):13.CorreaSandoval 2000; Acta Zool. Mex. (79):8.Correa-Sandoval & Rodriguez 2002; Acta Zol. Mex., (86):235. Schasicheila (Schasicheila ) misantlensis Fischer & Crosse. H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):41.Type Locality.Encarnacion, Hidalgo, Mxico. Holotype in the USNM. Distribution.SAN LUS POTOS: numerous localities in eastern region of state; (Correa-Sandoval et al. 1998). NUEVO LEN: Don Soltero, Hwy. Laguna Sanchezcanyon 4 mi. W of Cd. Victoria (Pilsbry 1903); Ejido Filipe 2002); El Cielo Biosphere Reserve, Gomez Farias (CorreaSandoval & Rodriguez. 2005). VERACRUZ: Rio Cazones, Schasicheila (Schasicheila) hinkleyi Pilsbry 1919Schasicheila hinkleyi Pilsbry 1919; Proc. Acad. Nat. Sci. Phila. Acad. Nat. Sci. Phila. 74:57. Schasicheila (Schasicheila ) hinkleyi Pilsbry. H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):37.Type Locality.Chama, [Dept. Alta Verapaz], Guatemala. Holotype in the ANSP. Distribution.Known only from the type locality. Schasicheila (Schasicheila) misantlensis Fischer & Crosse 1893Schasichila pannucea misantlensis Fischer & Crosse 1893:449; pl. Phila. 74:57. Schasicheila (Schasicheila ) misantlensis Fischer & Crosse. H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):41.Type Locality.Misantla, [Veracruz], Mxico. Distribution.VERACRUZ: Necaxa, 2215 ft. alt. (H. B. Baker 1918). Schasicheila (Schasicheila) nicoleti Shuttleworth 1852Shasicheila nicoleti Shuttleworth 1852; in Bern Mittheil.:302.H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:57. CorreaSandoval & Rodriguez 2002; Acta Zool. Mex. (86):235.Helicina nicoleti (Shuttleworth). Reeve 1874; Conch. Icon., Helicina Schasichila nicoleti (Shuttleworth). Von Martens 1890; Biol. Cent. Amer.:43. Schasicheila (Schasicheila ) nicoleti Shuttleworth. H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):40. Distribution.TAMAULIPAS: Los Angeles, Sierra

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20 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Rodriguez 2002). VERACRUZ: Cerro de Plumas, near alt. (H. B. Baker 1928). Schasicheila (Schasicheila) palmeri Dall 1905Schasicheila palmeri Dall 1905; Smiths. Misc. Coll. 48:19, pl. Phila. 74:57. Schasicheila (Schasicheila ) misantlensis Fischer & Crosse. H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):41.Type Locality.Sierra Alvarz, San Lus Potos, Mxico; 7900 ft. alt. Holotype USNM 110397. Distribution.SAN LUS POTOS: San Dieguito (Dall 1905). Schasicheila (Schasicheila) pannucea (Morelet 1849)Helicina pannucea Morelet 1849; Test. Noviss. I:21. Schasicheila pannucea (Morelet). Sowerby 1866; Thes. Conch. Chemnitz, Syst. Conch. Cab., Schasicheila 1 (shell).H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:57.Bequaert 1957; Bull. Mus. Comp. Zool. 115:207. Schasicheila (Schasicheila ) pannucea (Morelet). H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):42. Schasichila pannucea (Morelet). Von Martens 1890; Biol. Cent. (shell). Helicina alata (Menke). Reeve 1874; Conch. Icon., Helicina: pl. 13, Type Locality.San Lus, Dept. Petn, Guatemala. Distribution.GUATEMALA, Dept. Alta Verapaz: Cobn. Dept. Baja Verapaz: Senahu, N of Polochic Valley. Dept. Petn: San Lus (Von Martens 1890). CHIAPAS: Monte Libano, 600 m alt. Schasicheila (Schasicheila) pilsbryi Wagner 1911Schasicheila pilsbryi Wagner 1911; in Martini & Chemnitz, Syst. Conch. Cab., Schasicheila Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:57. Schasicheila (Schasicheila ) pilsbryi Wagner. H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):37.Type Locality.Mxico or Guatemala. Distribution.Unknown. Schazicheila (Schasicheila) Pilsbry 1899 Pilsbry 1899; Proc. Acad. Nat. Sci. Phila. 51:391.Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. Acad. Nat. Sci. Phila. 74:57. Schasicheila (Schasicheila ) Pilsbry. H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):36.CorreaSandoval & Salazar 2005; Acta Zool. Mex. (21):59. Mxico. Holotype in the ANSP. Distribution.NUEVO LEN: Santiago (CorreaSandoval 1993); numerous localities in southern region of state (Correa-Sandoval & Salazar 2005). Schasicheila ( Schasicheila) Pilsbry 1903 Pilsbry 1904; Proc. Acad. Nat. Sci. Acad. Nat. Sci. Phila. 74:57. Schasicheila (Schasicheila ) Pilsbry. H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):37.Type Locality.Canyon 4 miles west of [Ciudad] Victoria, Tamaulipas, Mxico. Holotype ANSP 85809. Distribution.Known only from the type locality. Schasicheila (Schasicheila) walkeri Hinkley 1920Schasicheila walkeri Type Locality.A bluff on the mountain north of Chama, Dept. Alta Verapaz, Guatemala. Holotype in the ANSP. Distribution.Known only from the type locality. Subfamily VIANINAE Baker 1922 The subfamily Vianinae is distributed primarily in the West Indies. A single genus occurs in the study area. Genus Pyrgodomus Crosse & Fischer 1893Pyrgodomus Crosse & Fischer, in Fischer & Crosse 1893:440.H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):43.Type Species.Helicina chryseis Tristram 1861. Distribution. Pyrgodomus is widely distributed at lower and intermittent elevations on calcareous substrata from Tamaulipas and Guerrero south to Prov. Puntarenas, Costa Rica. Taxonomy.Three species and one subspecies are recognized. The small size of the species and the paucity Pyrgodomus to be under-recorded. The genus is in critical need of review. Pilsbry 1930 Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. Type Locality.Islas del Cisne, Dept. Islas de la Baha, Honduras. Holotype ANSP 150868. Distribution.Known only from the type locality. Pyrgodomus microdinus microdinus (Morelet 1851)Helicina microdina Morelet 1851; Test. Noviss. II:18.Von Martens 1890; Biol. Cent. Amer.:39. Helicina (Idesa) microdina Morelet. Fischer & Crosse 1893:438 Eutrochatella (Artecallossa ) microdina (Morelet). Wagner Eutrochatella (Pyrgodomus) microdinus microdinus (Morelet). H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:61. Pyrgodomus microdinus microdinus (Morelet). H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):45.Goodrich & Van der Schalie 1937; Misc. Pub. Mus. Zool. Univ. Mich.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 21 Helicina chryseis Tristram 1862; Proc. Zool. Soc. Lond. (1861):5.Von Martens 1900; Biol. Cent. Amer.:606. Helicina (Pyrgodomus) chryseis Tristram. Fischer & Crosse Eutrochatella microdina chryseis (Tristram). Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 72:197.Hinkley 1920; Nautilus 34:52.H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:61. Pyrgodomus microdinus chryseis (Tristram). H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):45.Type Localities. Helicina microdina: Vera-paz [Dept. Alta Verapaz], Guatemala. Lectotype BMNH 1893.2.3.1986. (Richling 2004). Helicina chryseis: Guatemala, mountain forests of Vera Paz; location of the primary types are unknown. Distribution.Widely distributed at lower and intermittent elevations on calcareous substrata from Tamaulipas and Guerrero, Mxico south to Prov. Puntarenas, Costa Rica. Numerous specimen lots of Pyrgodomus from Costa Rica, Nicaragua, Honduras, Guatemala, Belize, and Mxico are in the UF collection. GUATEMALA, Dept. Alta Verapaz: Chama (Pilsbry 1920). MXICO. Numerous specimen lots of Pyrgodomus in the UF are from the states of Yucatn, Quintana Roo, Campeche, Tabasco, Chiapas, Oaxaca, Veracruz, San Lus Potos, and Tamaulipas. Pyrgodomus microdinus abditus H. B. Baker 1928Pyrgodomus microdina abditus H. B. Baker 1928; Occ. Pap. Mus. Mxico; 1300 ft. alt. Holotype in the UMMZ. Sumidero (H. B. Baker 1928). Pyrgodomus simpsoni (Ancey 1886)Trochatella simpsoni Ancey 1886; Bull. Soc. Malac. France 2:235. Pyrgodomus simpsoni (Ancey). Pilsbry 1904; Proc. Acad. Nat. Sci. Eutrochatela (Pyrgodomus) simpsoni (Ancey). H. B. Baker 1922; Proc. Acad. Nat. Sci. Phila. 74:61.Type Locality.Isla de tila, Dept. Islas de la Baha, Honduras. Distribution.Known only from the type locality. Superfamily AMPULLARIOIDEA Gray 1824 Family AMPULLARIIDAE Gray 1824Ampullariidae Gray 1824.Cowie 1997; Bulletin of Zoological Nomenclature 54 (2).Cowie & Thiengo 2003; Malacologia 45. Pilidae Preston 1915.Type Genus.Ampullaria Lamarck 1799. Ampullaria is a junior synonym of Pila Rding 1798. The family name Pilidae was based on the genus Pila The International Commission on Zoological Nomenclature [ICZN] 1999a, Opinion 1913 ruled that Ampullariidae is the valid name for this family. Distribution.Freshwater in Tropical America, Africa and southeast Asia east to the Philippine Islands. Taxonomy.The taxonomy of this family is subjective and unstable. There is little agreement on the number of genera or subgenera, and the species level taxonomy is chaotic. Nearly all species were described on the basis of only the shell. In most instances only a single specimen was included in the original description. Frequently the locality of origin of type specimens was unknown or only vaguely known. The result of this history is that there is very little agreement concerning how many valid species exist among the hundreds of names that have been proposed. Cowie and Thiengo (2003) recognized 140 Neotropical species and Their work represented a major effort to bring some order to this chaos. The following arrangement departs very little from their scheme. Genus Marisa Gray 1824Marisa Gray 1824; Philos. Mag. & Jour. 63:276.Cowie & Thiengo 2003; Malacologia 45:49. Ceratodes Guilding 1828; Zool. Jour. 3:537. Type Species.Marisa; Helix cornuarietis Linneaus 1758 by subsequent designation of Gray 1847. Ceratodes: Helix cornuarietis Linnaeus 1758 by original designation. Distribution.South America, Central America. Taxonomy.Eight species have been described. Three are currently recognized. Marisa cornuaurietis (Linnaeus) is found in northern South America; M. planogyra Pilsbry is found in the Rio Parana system, and M. rotula (Mousson) is known from Colombia, Panam and Costa Rica. Marisa rotula (Mousson 1869)Ampullaria rotula Mousson 1869; Malac. Bltt. 16:183.Von Ceratodes rotula (Mousson). Mousson 1873; Malac. Bltt. 16:19. Marisa rotula (Mousson). Pilsbry 1933; Proc. Acad. Nat. Sci. Phila. 85:71. Marisa cornuarietis (Linnaeus). Cowie & Thiengo 2003; Malacologia 45:51.Type Locality.Colombia, lower part of the Rio Magdalena. Syntypes ZMZ 525321. Distribution.Colombia, Panam, and Costa Rica. COSTA RICA (Von Martens 1899). Genus Pomacea Perry 1812Pomacea Perry 1812; Arcana: unnumbered plate.Cowie & Thiengo 2003; Malacologia 45:51. Conchylium Cuvier 1816:426. Limnopomus Dall 1904; Jour. of Conch. 11:52. Type Species.Pomacea: Pomacea maculata Perry 1812, by original designation. Conchylium : Nerita urceus Mller 1774. Limnopomus: Ampullaria columellaris Gould 1848. Distribution.South America north to Tamaulipas along coast; the Windward Island of the Lesser Antilles; Jamaica;

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22 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Cuba; the southeastern United States. Taxonomy.Cowie & Thiengo (2003) recognize three subgenera. Only one occurs in Central America and Mxico. Subgenus Pomacea Perry 1812 Distribution.Same as for the genus. Taxonomy.Cowie and Thiengo (2003) listed 97 species in the subgenus Pomacea. Fourteen occur in the study area. Pomacea auriformis (Reeve 1856)Ampullaria auriformis Reeve 1856; Conchologica Iconica 10: pl. Pomacea auriformis (Reeve). Pain 1964; Jour. of Conch. 25:225.Cowie & Thiengo 2003; Malacologia 45:57.Type Locality.Honduras. Syntype BMNH 20020646. Distribution.Unknown. At the time that this species was described, Honduras included nearly the entire Caribbean Coast of Central America. Pomacea belizensis (Crosse & Fischer 1888)Ampullaria belizensis Crosse & Fischer in Fischer & Crosse 1888: Martens 1899; Biol. Cent. Amer.:417. (Say). Pain 1964; Jour. of Conch. 25:227.Cowie & Thiengo 2003; Malacologia 45:58.Type Locality.No locality given by Crosse and Fischer in 1888. Belize (Fischer & Crosse 1890). Syntypes in MNHN. Distribution.Known only from the type locality. Pomacea catemacensis (H. B. Baker 1922)Ampullaria patula catemacensis H. B. Baker 1922; Occ. Pap. Mus. Pomacea patula (H. B. Baker). Naranjo-Garcia & Garca-Cubas 1986; Ann. Inst. Biol. 56:603.Espinoza-Chvez & MartnezCowie & Thiengo 2003; Malacologia 45:59.Diupotex-Chong, Cazzaniga, Hernndez-Santoyo & Betancourt-Rule 2004; Biocell. 28:279 (karyotype).Type Locality.Lago de Catemaco, Veracruz, Mxico. Holotype: UMMZ 31850. Distribution.Known only from the type locality. Remarks.Pomacea catemacensis was described as a subspecies of Ampullaria patula Reeve 1856, from New Grenada (= Colombia) and Panam (see Cowie & Thiengo 2003:71). The name Ampullaria patula Reeve is a primary junior homonym of Ampullaria patula Lamarck 1804 (Naticidae), and is not available for use. The identity of Reeves species from New Grenada is unresolved, but it has no bearing on the status of P. catemacensis. Pomacea cerasum (Hanley 1854)Ampullaria cerasum Hanley 1854:unnumbered page.Von Martens Pomacea cerasum (Hanley). Cowie & Thiengo 2003; Malacologia 45:59.Type Locality.Not given. Distribution.TABASCO: Teapa (Von Martens 1899:421). Pomacea conoidea (Von Martens 1899)Ampullaria conoidea Von Martens 1899; Biol. Cent. Amer.:423; pl. Pomacea conoidea (Von Martens). Cowie & Thiengo 2003:60.Type Locality.Costa Rica. Distribution.Known only from the type locality. Pomacea costaricana (Von Martens 1899)Ampullaria costaricana Von Martens 1899; Biol. Cent. Amer.:418 Pomacea costaricana (Von Martens). Cowie & Thiengo 2003:60.Type Locality.Not given. No type designated. Distribution.Von Martens (1899) gives the following locality records. NICARAGUA: Lago de Nicaragua. COSTA RICA, Prov. Puntarenas: Rio Saveyre, at Boca Culebra; Palmar, south of the Rio Grande de Terraba. PANAM, Prov. Chiriqu: Chiriqu. Pomacea cumingi cumingi (King & Broderip 1831)Ampullaria cumingii King & Broderip 1831; Zool. Jour. 5:332. Pomacea cumingi (King & Broderip). Morrison 1946; Smithson. Misc. Coll. 106:6.Morrison 1952; Nautilus 65:105.Cowie & Thiengo 2003:61.Type Locality.Panam, Archipilago de las Perlas: Isla Saboga (Morrison 1952:105). Distribution.Known only from the type locality. Morrison (1946, 1952) recorded this species from Isla Pacheco, but later he discounted this locality. Pomacea cumingi sanjoseensis Morrison 1946Pomacea cumingii sanjoseensis Morrison 1946; Smithson. Malacologia 45:74.Type Locality.Three small streams at the west end of Isla San Jos, Archipilago de las Perlas, Panam. Holotype USNM 542136. Distribution.Known only from the type locality. Pomacea dacostae (Sowerby 1909)Ampullaria dacostae G. B. Sowerby 1909; Proc. Malac. Soc. Pomacea dacostae (G. B. Sowerby). Cowie & Thiengo 2003; Malacologia 45:61.Type Locality.Costa Rica. Syntype BMNH 10.19.35. Distribution.Known only from the type locality. (Say 1827) B. Baker 1922; Occ. Pap. Mus. Zool. Univ. Mich. (106):37. (Say 1827). Pain 1964; Jour. of Conch. 25:226 Univ. Nac. Auton. Mxico 58:21; pl. 1 (shell, operculum); pl. 2 (radula); pls. 3 (anatomy).Cowie & Thiengo 2003;

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 23Malacologia 45:64. Pain (1964) and Cowie & Thiengo (2003) listed the following names as synonyms: Valenciennes 1833. (type locality not given). Ampullaria ochracea Jay 1836. Ampullaria malleata Jonas 1844. (type locality: Tabasco). Ampullaria labiosa (Koch) Philippi 1851. Ampullaria lamarckii Philippi 1851. Ampullaria hondurasensis Reeve 1856. Reeve 1856. (type locality: Tabasco). Ampullaria fumata Reeve 1856. Ampullaria columbiensis (Sowerby) Reeve 1856. Ampullaria Von Martensiana Neville 1884. Ampullaria strebeli Fischer & Crosse 1890. (type locality: Misantla, Veacruz, in a small stream called the Brazo Seco). Ampullaria strebeli var. prasina Fischer & Crosse 1890. Ampullaria belizensis Fischer & Crosse 1890 [recognized as a distinct species here]. var. esculpta Fischer & Crosse 1890. (type locality not given). Ampullaria malleata var. arata Fischer & Crosse 1890. (type locality: not given). Ampullaria occlusa Fischer & Crosse 1890. (type locality: freshwater mouth of the Rio Guacalata). Ampullaria monachus Fischer & Crosse 1890. (type locality: Santa Ampullaria yucatanensis Fischer & Crosse 1890. Ampullaria eumicra Fischer & Crosse 1890. (type locality: State of Oaxaca) Ampullaria lemniscata Fischer & Crosse 1890. Ampullaria innexa Fischer & Crosse 1890. (type locality: Monte de Mistam, near Coapam, Oaxaca). Ampullaria malleata var. prasina Fischer & Crosse 1890. Ampullaria malleata var. oajacensis Fischer & Crosse 1890. (type locality: Monte de Mistam, near Coapam, Oaxaca). Ampullaria yucatanensis var. izabelensis Von Martens 1899.Type Locality.For only. A short distance below [Ciudad] Veracruz, Mxico. Lectotype ANSP 50645 (Pilsbry 1991a:325; H. B. Baker 1964:68). Distribution.From Tamaulipas south along eastern Mxico to Dept. Petn, Guatemala and Belize, and along and Thiengo (2003:64) Pain 1964 stated that this species is distributed from central Mxico to the Rio Magdalena Basin, Colombia. Some records are as follow. COSTA RICA, Prov. Guanacaste: Rio de Caas north of Santa Cruz (Pilsbry 1920:9). GUATEMALA, Dept. Petn: Tikal National Park (Basch 1959) MXICO. CAMPECHE: 51 mi. E of Campeche (Branson & McCoy 1965). TABASCO: Villa Hermosa (Thompson 1967). VERACRUZ: Los Tuxtlas, Laguna Escondida, Laguna Zacatl and Laguna Sontecomapn (Rangel-Ruiz 1987). Widely introduced in the Taxonomy.The systematic status of the many names associated with this widely distributed species is unsatisfactory. All of the names are based on shell characters that are variable and unreliable. However, to treat them as synonyms without careful morphological study or genetic analysis is as arbitrary as was their original descriptions. Very few locality citations from the literature are repeated here, because of the long lists of synonyms that may or may not apply to and its subspecies. The reader is referred to Cowie and Thiengo (2003) for these locality citations. (Hanley 1854)Ampullaria dysoni Martini & Chemnitz, Syst. Conch. Cab., Ampullaria:110; pl. (Hanley). Pain 1964; Jour. of Conch. Type Locality.Honduras. Syntype BMNH 1907 11.21.65. Distribution.Known only from the type locality. (Crosse & Fischer 1890)Ampullaria erogata Crosse & Fischer 1890; Jour. de Conchyl. Kobelt 1912, in Martini & Chemnitz, Syst. Conch. Cab., Ampullaria (Crosse & Fischer). H. B. Baker 1922; Occ. Pap. Mus. Zool. Univ. Mich. (106):38.Pain 1964; Jour. of Type Locality.Not given. Distribution.Same as for (Pain 1964). (Von Martens 1899) var. guatemalensis Von Martens 1899; Biol. (Say). Cowie & Thiengo 2003; Malacologia 45:65.Type Locality.Paso Antonio, in the lower part of the Distribution.Known only from the type locality. (Reeve 1856) Reeve. Kobelt 1912; in Martini & Chemnitz, Syst. Conch. Cab., Ampullaria (Reeve). Pain 1964; Jour. of Conch. Ampullaria ghiesbreghtii Ampullaria ghiesbreghti Reeve.Fischer & Crosse 1890:233; pl. 47, Pomacea ghiesbreghti (Reeve). Pain 1953; Proc. Malac. Soc. London 29:222. (Reeve). Bequaert 1957:210. Pomus giganteus Tristram 1864:414 (type locality: Lago de Petn, Guatemala). Ampullaria chiapasensis Crosse & Fischer 1890; Jour. de Conchyl. 38:235. (type locality: Las Playas, Chiapas, in marshes). (Tristram). Von Martens 1899; Biol.

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24 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Ampullaria tristrami Crosse & Fischer 1890; Jour. de Conchyl. 38:234. (type locality Lago de Petn, Guatemala). var. tristami Von Martens 1899; Biol. Cent. Hinkley 1920; Nautilus 34:45, 47, 49. Type Locality.For only. Lake Petn, Dept. Petn, Guatemala. Syntype BMNH 1986214. Distribution.Chiapas, Tabasco, and the Dept. Petn, Guatemala (Pain 1964). BELIZE: (UF collections). TABASCO: Rio Grijalva, 4 mi. S of Villlahermosa; Rio Grijalva 14 mi. N of Teapa (Thompson 1957). GUATEMALA, Dept. Izabal: Plantera, Rio Dulces near Livingston; Jocola, north side of Lago de Izabal. Dept. Alta Verapaz: Panzos, Rio Polochic (Hinkley 1920). Pomacea lattrei lattrei (Reeve 1856)Ampullaria lattrei Reeve 1856; Conchologica Iconographia, 10 Ampullaria delattrei Reeve. Fischer & Crosse 1890:246 (emend.); Reeve. Hinkley 1920; Nautilus 34:47. Pomacea lattrei (Reeve), Cowie & Thiengo 2003:67.Type Locality.Cobn, Guatemala. Lectotype BMNH 2020665 (Cowie & Thiengo 2003). Distribution.Northern and eastern Guatemala. Hinkley (1920) lists this species from rocks near Castillo San Filipe, Lago de Izabal, [Dept. Izabal], Guatemala. Remarks.The species was named by Reeve (1856) as Ampullaria lattrei for the French naturalist M. A. Delattre, because Reeve took the discoverers name to be de Lattre, a titular designation of nobility. The ICZN code Appendix D, para. 21 (d) prohibits the use of a particle denoting nobility the name to delattrei, conforming to its correct orthography. However, ICZN Article 32 requires that the original spelling must be maintained, except as provided by para. 32 (c) (ii). If a name is misspelled due to a typographic error or a lapsus calami, and this is clear in the publication without reference to other sources, then the name can be emended. Reeves original name must be retained, regardless of its incorrect etymology. Pomacea lattrei chamana (Hinkley 1920)Ampullaria lattrei chamana Hinkley 1920; Nautilus 34:53. Pomacea lattrei chamana (Hinkley). Cowie & Thiengo 2003:59.Type Locality.Chama, near the Rio Tsalbha, Dept. Alta Verapaz, Guatemala. Lectotype ANSP 46321 (H. B. Baker 1964:168). Distribution.Known only from the type locality. Pomacea miltocheila (Reeve 1856)Ampullarius miltocheilus Reeve 1856. Conchologica Iconica 10: pl. Pain 1953; Proc. Malac. Soc. London 29:122. Ampullaria miltochilus Reeve. Fischer & Crosse 1890:247. Pomacea cumings (King & Broderip 1831). Cowie & Thiengo 2003; Malacologia 45:69.Type Locality.Province of Chiapes [= Chiapas], Mxico. Lectotype BMNH 20020668. Distribution.VERACRUZ: Cazcajal, on the upper part of the Rio Uzpanapa (Pain 1953) Remarks.Cowie and Thiengo (2003) gave no Pomacea cumingi (King & Broderip 1831), a species from Panam. Pomacea pealiana (Lea 1834)Ampullaria pealiana Ampullaria pealeana Lea. Von Martens 1899; Biol. Cent. Amer.:423; Pomacea pealiana (Lea). Cowie & Thiengo 2003; Malacologia 45:71.Type Locality.Turbaco, Colombia. Holotype ANSP 192933 (Abbott 1955). Distribution.Colombia, Ecuador, Venezuela, and Panam (Cowie & Thiengo 2003:71). PANAM, Prov. Veraguas: Veraguas (Von Martens 1899). Pomacea picta (Reeve 1856)Ampullaria picta Pomacea picta (Reeve). Cowie & Thiengo 2003; Malacologia 45:72.Type Locality.Mazatln. Here restricted to Mazatln, Guerrero, which is on the road south from Mxico City to Acapulco. This is the locality from where many 19th Century species originated, very few of which occur at Mazatln, Sinaloa, Mxico. Syntypes BMNH 1907.11.21.91. Distribution.Known only from the type locality. Pomacea zeteki Morrison 1946Ampullaria cumingi King & Broderip. Von Martens 1899; Biol. Cent. Amer.:422 (in part). Pomacea zeteki Morrison 1946; Smithson. Misc. Coll. 106:8.Cowie & Theingo 2003; Malacologia 45:77.Type Locality.A shallow margin of the Rio Chagres, near Gatuncilla, Panam. Holotype USNM 542137. Distribution.Known only from the type locality. Superfamily VIVIPAROIDEA Gray 1847 Family VIVIPARIDAE Gray 1847 A single species of Viviparidae of questionable status has been recorded from Mxico. Subfamily VIVIPARINAE Gray 1847 Genus Montfort 1810 (Binney 1865) 1 (shell).Binney 1865; Smiths. Misc. Coll. (144):11314; Paludina inornata (Binney). Fischer & Crosse 1891:291. (Lea). Clench & Fuller 1965; Occ. Pap. on

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 25 Type Locality.Chopatilo. Von Martens (1899) suggested that the orthography of Chopatilo probably is incorrect. He gives three localities with different spellings that may have been the place of origin of the types, one in Michoacn, is here selected as the most probable: Hacienda Chapotito, Lectotype MCZ 234704 (Clench & Fuller 1965:397). Distribution.Known only from the type locality. Remarks.Clench and Fuller (1965:398) suggested that may be a synonym of georgianus (Lea 1834) because of its resemblance to atypically green and unbanded specimens from northern Florida. This synonymy seems unlikely because the weakly impressed suture that characterizes inornatus differs strikingly from the much more deeply impressed suture of Superfamily CYCLOPHOROIDEA Gray 1847 Family NEOCYCLOTIDAE Kobelt & Moellendorff 1897 Type Genus.Neocyclotus Fischer & Crosse 1886. Taxonomy.Twenty-eight genera of Neocyclotidae are recognized currently in the neotropical region. Another six taxonomy of the cyclophorids is very unsettled. There is little of families, subfamilies and genera. Three subfamilies of Neocyclotidae are recognized in the Neotropics. The Megalomastomatinae is found in Cuba, Hispaniola, Puerto Rico, and the Virgin Islands, and in eastern Mxico, Guatemala and Belize. The Crocidopomatinae is found in Cuba, Hispaniola and the Lesser Antilles (Thompson 1967). The Neocyclotinae is found in the Greater and Lesser Antilles, and the mainland from Mxico south to Bolivia. Subfamily MEGALOMASTOMATINAE Blanford 1864Megalomastominae Torre & Bartsch 1942; Bull. U. S. Nat. Mus. 181:2.Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. 181:142. Thompson 1969; Zoologica 54:37.Type Genus.Megalomastoma Swainson 1840. Distribution.Greater Antilles, Eastern Mxico, Guatemala, and Belize. Taxonomy.Five genera are recognized, two in the Mxico-Central American Region. Genus Aperostoma Troschel 1847Aperostoma Troschel 1847:44.Heremanssen 1852:10.H. B. Baker 1932; Nautilus 35:14.H. B. Baker 1943; Nautilus 56:137.Morrison 1955; Jour. Wash. Acad. Sci. 45:152.Solem 1956; Proc. Acad. Nat. Sci. Phila. 108:46.Thompson 1969; Zoologica 54:37. Cyrtotoma Mrch 1852; Catalogus Conchyliorum 1:40.Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. 181:169. Habropoma Crosse & Fischer 1880:8.Fischer & Crosse 1890:127 129.Type Species. Aperostoma: Cyclostoma mexicanum Menke 1830. Cyrtotoma: Cyclostoma mexicanum Menke 1830.Habropoma: Cyclostoma mexicanum Menke 1830. Distribution.Eastern Mxico from eastern San Lus Potos south to Oaxaca and Veracruz. Taxonomy.Three species and one subspecies are recognized currently. Bartsch and Morrison (1942) recognized eight, most of which Solem (1956a) placed in synonymy. Aperostoma mexicanum mexicanum (Menke 1830)Cyclostoma mexicanum Menke 1830; Symopsis method. mollusc.:133. Aperostoma mexicanum (Menke). Troschel 1847; Zeitschr. Malak., 4:44.Solem 1956; Proc. Acad. Nat. Sci. Phila. 108:48; pl. 5, Cyrtotoma mexicanum (Menke). Mrch 1852; Catalogus concyliorum, 1:40.Bartsch & Morrison 1942; Bull. U. S. Nat. Cyrtotoma mexicanum mexicanum (Menke). H. B. Baker 1922; Occ. Pap. Mus. Zool. Univ. Mich. (106):43. Habropoma mexicanum (Menke). Fischer & Crosse 1890:130; Cyclophorus mexicanus Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. Cyrtotoma ignotum Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. Type Localities. Cyclostoma mexicanum: Papantla, Veracruz, Mxico. : Motzorongo, Veracruz, Mxico; holotype USNM 128285. Cyrtotoma ignotum: Mxico; holotype USNM 523515. : Hacienda Coaotolapam, Veracruz, Mxico; holotype USNM 515789. Distribution.OAXACA: Playa Vicente; Tustepe (Von Martens 1890). PUEBLA: Multatoyuca (Bartsch & Morrison 1942). VERACRUZ: Papantla; Misantla; Cuesta de Papantla; Jalapa; Cerro de Plumas (Von Martens 1890); Coatotolapam; (Solem 1956a). Taxonomy.The synonymy used here follows Solem museum collections. The taxonomy of the genus needs to be reviewed. Until then the distribution records given above should be regarded as tentative. Aperostoma mexicanum salleanum (Von Martens 1865)Cyclophorus salleanus Von Martens 1865:151. Habropoma salleana (Martens). Fischer & Crosse 1880:133; Cyclophorus (Cyrtotoma) salleanus Von Martens. Von Martens 1890; Biol. Cent. Amer.:7. Cyrtotoma salleanum (Von Martens). Kobelt & Mllendorff 1907; Nach. Bltt. Deut. Ges., 29:140.Bartsch & Morrison 1942; Cyrtotoma mexicanum salleanum (Von Martens). H. B. Baker 1922; (shell).

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26 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Aperostoma mexicanum salleanum (Von Martens). H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):51.Thompson system). Aperostoma mexicanum mexicanum (Menke) (in part). Solem 1956a:48. Distribution.VERACRUZ: Barranca de Santa Maria, near Mirador; Orizaba; Misantla; Quilate, near Misantla; Atoyac; Coatepec; Jalapa (Von Martens 1890); 2.9 mi. E of mi. SW, 0.8 mi. N of Fortn, 2900 ft. alt. (Thompson 1969). Aperostoma palmeri (Bartsch & Morrison 1942)Cyrtotoma palmeri Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. Aperostoma mexicanum palmeri (Batsch & Morrison). Solem 1956; Reguero 1998; Acta Zol. Mex., n. s. (73):13.Correa-Sandoval & Rodriguez 2002; cta Zool. Mex., (86):235. Aperostoma palmeri (Bartsch & Morrison). Thompson 1969; Type Locality.Gomez Faras, Tamaulipas, Mxico. Holotype USNM 198079. Distribution.HIDALGO: Chapulhuacan (Solem 1956a). SAN LUS POTOS: 11.4 mi. E of Xilitla, 1100 ft. alt.; 7.5 mi. E of Xilitla (Thompson 1969); 6 km SE of Sandoval et al. 1998). TAMAULIPAS: El Cielo Biosphere Reserve (Correa-Sandoval & Rodriguez 2005); numerous localties in the southern region of the state (Correa-Sandoval & Rodriguez 2002).. Aperostoma walkeri H. B. Baker 1928Aperostoma walkeri H. B. Baker 1928; Occ. Pap. Mus. Zool. 39 (operculum).Solem 1956; Proc. Acad. Nat. Sci. Phila. Zool. Mex. n. s. (79):8. Cyrtotoma walkeri (H. B. Baker). Bartsch & Morrison 1942; Bull. Cyrtotoma goldmani Bartsch & Morrison 1942; Bull. U. S. Nat. Type Localities .Aperostoma walkeri : forests on south side of Vaso de Tenango, Nacaxa, Puebla, Mxico; holotype in the UMMZ. Cyrtotoma goldmani: Multatoyuca [Metlaltoyuca], Puebla, Mxico; holotype USNM 53516. Distribution.PUEBLA: Metlaltoyuca; Necaxa. VERACRUZ: Coyutla; Misantla (Solem 1956a); El Cedral, 2000). Genus Tomocyclus Crosse & Fischer 1872Tomocyclus Crosse & Fischer 1872; Jour. de Conchyl. 20:76.Fischer & Crosse 1888:1131.Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. 181:142.Thompson 1963; Breviora (181):1.Type Species.Tomocyclus gealei Crosse & Fischer 1872. Distribution.Belize, eastern Guatemala and southeast Mxico in the states of Chiapas and Veracruz. Taxonomy.Five species are recognized. Tomocyclus gealei Crosse & Fischer 1872Tomocyclus gealei Crosse & Fischer 1872:77.Fischer & Crosse Thompson 1963:2. Megalomastoma (Tomocyclus) gealei (Crosse & Fischer). Von Martens 1890; Biol. Cent. Amer.:10.Type Locality.Chiapas, Mxico. Distribution.GUATEMALA, Dept. Baja Verapaz: woods between Tactic and Tamahu; Polochic Valley above Panzos and Senahu (Von Martens 1890). Thompson 1963 Thompson 1963; Breviora (181):3; pl. 1, Type Locality.High rainforest at Valentine Camp, 50 mi. southwest of Cayo, British Honduras [Belize]. Holotype UMMZ 194095. Distribution.BELIZE or GUATEMALA. The type locality, if correct, would place the species in the Dept. Petn, Guatemala. Tomocyclus guatemalensis (Pfeiffer 1851)Cyclostoma guatemalensis Pfeiffer 1851:245.Pfeiffer 1853:267; Tomocyclus guatemalensis (Pfeiffer). Crosse & Fischer 1872:76.(shell).Thompson 1963; Breviora (181):8Type Locality.Vera Paz, Guatemala. Distribution.Known only from the type locality. Tomocyclus lunai Bartsch 1945Tomocyclus guatemalensis (in part) Fischer & Crosse 1886:124; pl. Tomocyclus lunai Bartsch 1945; Proc. Biol. Soc. Wash. 58:63.Thompson 1963; Breviora (181):8.Type Locality.Santecomopan, Veracruz, Mxico. Holotype USNM 573547. Distribution.VERACRUZ: Santecomopan; San Martin Tuxtla; south slope of Volcn San Martin, 1040 ft. alt.; Volcn San Martin (Thompson 1963). Tomocyclus simulacrum (Morelet 1849)Cyclostoma simulacrum Morelet 1849; Test. Noviss. I:22. Cyslostoma copanense Sowerby 1850; Thesaurus Conchyl., 1 Tomocyclus simulacrum (Morelet). Crosse & Fischer 1872; Jour. de

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 27Thompson 1963; Breviora (181):5. Megalomastoma simulacrum var. gracilius Von Martens 1890; Biol. Cent. Amer.:10. Tomocyclus siphonis Bartsch & Morrison 1942; Bull. U. S. Nat. Tomocyclus constrictus Bartsch & Morrison 1942; Bull. U. S. Nat. Type Localities. Cyclostoma simulacrum: Dept. Petn, Guatemala. Cyclostoma copanensis: Cobn, Guatemala (?). Megalomastoma simulacrum var. gracilius: Between Tactic and Tamahu, Dept. Baja Verapaz, Guatemala. Tomocyclus siphonis: Alta Verapaz, Guatemala. Tomocyclus constrictus: Cobn, Guatemala. Distribution.GUATEMALA. Dept. Alta Verapaz: Cobn; Chama; Finca de Providensia; near Arroyo Yalchatila, 4 km SW of Seiba; 1 km N of Finca Samac; 55 km NE of Cobn; Chinaja (Thompson 1963). Dept. El Quich:4 km NW of Hacienda Pacala (Thompson 1963). Subfamily NEOCYCLOTINAE Kobelt & Mllendorff 1897 Distribution.Widely distributed throughout the neotropical realm. Taxonomy.About 25 genera and numerous subgenera are recognized. Thirty-seven species occur in the study area. Genus Neocyclotus Fischer & Crosse 1886Neocyclotus Fischer & Crosse 1886:148.Morrison 1955; Jour. Wash. Acad. Sci. 45:156.Solem 1956; Proc. Acad. Nat. Sci. Phila. 108:52.Thompson 1969; Zoologica 54:65. Aperostoma (Neocyclotus ) Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. 181:203. Austrocyclotus Bartsch 1942:132.Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. 181:195.Morrison 1955; Jour. Wash. Acad. Sci. 45:156.Type Species.Neocyclotus : Cyclostoma dysoni Pfeiffer 1851 (by subsequent designation; Pilsbry 1910:533). Austrocyclotus : Cyclostoma stramineum Reeve 1843 (by original designation). Distribution.Veracruz south to Ecuador, Venezuela and the Lesser Antilles. Taxonomy.About 25 species are recognized. Six occur in the study area. Neocyclotus dysoni is widely distributed from Panam north to Veracruz. Bartsch & Morrison (1842) divided the species into 13 subspecies. These are listed below, although the validity of some are in doubt (Solem 1956a). Many earlier distribution records are omitted from this report for Neocyclotus dysoni because their identities are nearly Neocyclotus bisinuatus (Von Martens 1864)Cyclotus bisinuatus Von Martens 1864; Malak. Bltt. 11:113; pl. 3, Cyclotus (Aperostoma) bisinuatus (Von Martens). Von Martens 1890; Biol. Cent. Amer.:3. Neocyclotus (Neocyclotus ) bisinuatusm (Von Martens). Kobelt & Mllendorff 1897; Nachrb. Deut. Malak. Ges. 29:137. Aperostoma (Aperostoma) bisinuatum (Martens). Bartsch & 6 (shell). Neocyclotus bisinuatus (Von Martens). Thompson 1969; Zoologica system).Type Locality.Elevated plains of Costa Rica. Distribution.COSTA RICA: Prov. Cartago: Tapant, 4300 ft. alt.; 5 mi. E of Villa Neily; Inter American Agricultural Institute, Turrialba, 2000 ft. alt. (Thompson 1969). GUATEMALA, Dept. Retalhuleu: Costa Cuco above Retalhuleu; San Francisco Miramar, 2500 ft. alt. Dept. Huehuetenango: on the slope of Volcn de Santa Maria, 3000 ft. alt. (Von Martens 1890). Neocyclotus capscelius Thompson 1969Neocyclotus capscelius Thompson 1969; Zoologica 54:73; pl. V, Type Locality.A wooded ravine on Cerro de la Muerte, 10.5 km N of San Isidro El General, Prov. San Jos, Costa Rica. Holotype UF 20148. Distribution.Known only from the type locality. Neocyclotus chrysacme (Bartsch & Morrison 1942)Aperostoma (Neocyclotus ) chrysacme Bartsch & Morrison 1942; Type Locality.Wani, Nicaragua. Holotype USNM 186112. Distribution.Known only from the type locality. Neocyclotus dysoni dysoni (Pfeiffer 1851)Cyclostoma dysoni Pfeiffer 1851; Proc. Zool. Soc. Lond. 19:243.Pfeiffer 1853; in Martini & Chemnitz Syst. Conch. Cab., 1:259; Neocyclotus dysoni (Pfeiffer) in part. Fischer & Crosse 1888:164. Cyclotus (Aperostoma) dysoni (Pfeiffer) in part. Von Martens 1890; Biol. Cent. Amer.:3. Poteria (Neocyclotus) dysoni (Pfeiffer). H. B. Baker 1923; Occ. Pap. Mus. Zool. Univ. Mich. (137):33. Aperostoma (Neocyclotus ) dysoni dysoni (Pfeiffer). Bartsch & 28.Thompson 1969; Zoologica 54:67.Type Locality.Honduras. Distribution.HONDURAS, Dept. Comayagua:4 km SSW of Comayagua; 15.1 km N of Comayagua (Thompson 1969). (Von Martens 1890)Cyclotus (Aperostoma) Von Martens 1890; Biol. Cent. Neocyclotus (Neocyclotus ) (Von Martens). Kobelt & Mllendorff 1897; Nachrb. Deut. Malak. Ges. 29:137. Poteria (Neocyclotus) (Von Martens). H. B. Baker 1923; Occ. Pap. Mus. Zool. Univ. Mich. (137):34. Aperostoma (Neocyclotus ) (Von Martens). Bartsch & 22 (shell).Type Locality.Bugaba, Panam.

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28 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Distribution.Known only from the type locality. Neocyclotus dysoni ambiguus (Von Martens 1890)Cyclotus (Aperostoma) dysoni ambiguus Von Martens 1890; Biol. Cent. Amer.:4. Neocyclotus (Neocyclotus ) dysoni ambiguus (Von Martens). Kobelt & Mllendorff 1897; Nachrb. Deut. Malak. Ges. 29:137. Poteria (Neocyclotus) dysoni ambiguus (Von Martens 1890). H. B. Baker 1923; Occ. Pap. Mus. Zool. Univ. Mich. (137):34. Aperostoma (Neocyclotus ) dysoni ambiguum (Von Martens). Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. 181:211; pl. Neocyclotus dysoni ambiguum (Von Martens). Solem 1956; Proc. Acad. Nat. Sci. Phila. 108:53.Thompson 1969; Zoologica 13, A (female reproductive system). Veracruz, Mxico. Distribution.CHIAPAS: Ixtapa; Tecpatn (Solem 1956); 8.0 mi. N of Tuxtla Gutierrez, 3100 ft. alt.; 8.6 mi. E of Chiapa de Corze, 3100 ft alt.; 14.9 mi. E of Chiapa de Corzo, 4400 ft. alt. (Thompson 1969). OAXACA: Rio Grande (Solem 1956a). TABASCO:2.4 mi. E of Teapa; 2.6 mi. E of Teapa; 4.0 mi. W of Teapa (Thompson 1969). VERACRUZ: Jalapa; Octopan (Bartsch & Morrison 1942); Laguna Encontada, near San Andres Tuxtla (Thompson 1969). Neocyclotus dysoni aureus (Bartsch & Morrison 1942)Aperostoma (Neocyclotus ) dysoni aureus Bartsch & Morrison 1942; Neocyclotus dysoni aureum (Bartsch & Morrison). Solem 1956; Proc. Acad. Nat. Sci. Phila. 108:53.Branson & McCoy 1963; Nautilus 76:103.Type Locality.Panistlahuaca, Oaxaca, Mxico. Holotype USNM 523970. Distribution.CAMPECHE: 51 mi. E of Campeche (Branson & McCoy 1963). OAXACA: Gamboa; Panistlahuaca (Solem 1956a). EL SALVADOR, Dept. Santa Ana: Lago de Coatepeque. GUATEMALA: Lago de Amatitlan (Solem 1956a). Neocyclotus dysoni berendti (Pfeiffer 1861)Cyclotus berendti Pfeiffer 1861; Malak. Bltt. 8:171. Neocyclotus berendti 4 (shell).Fischer & Crosse 1888:167.Branson & McCoy 1963; Nautilus 76:103. Cyclotus (Aperotoma) dysoni berendti (Pfeiffer). Von Martens 1890; Biol. Cent. Amer.:5. Cyclotus dysoni form multilineatus Pilsbry 1891; Proc. Acad. Nat. Sci. Phila. 43:325. Poteria (Neocyclotus) berendti (Pfeiffer). H. B. Baker 1923; Occ. Pap. Mus. Zool. Univ. Mich. (137) 33. Bequaert & Clench 1933; Pub. Carnegie Inst. Wash. (431):350. Aperostoma (Neocyclotus ) dysoni berendti (Pfeiffer). Bartsch & (shell). Aperostoma dysoni berendti (Pfeiffer). Harry 1950; Occ. Pap. Mus. Zool. Univ. Mich. (524):25. Neocyclotus dysoni berendti (Pfeiffer). Solem 1956; Proc. Acad. Nat. Sci. Phila. 108:53.Rehder 1966; Proc. Biol. Soc. Wash. 79:278.Thompson 1967; Bull. Fl. St. Mus., 11:228.Type Locality.Dead shells found on the shore, Veracruz, Veracruz (Von Martens doubts this locality). Distribution.CAMPECHE: 5.1 mi. NNW of Dzibalchn (Thompson 1967). QUINTANA ROO: 12 km from Icaiche (Rehder 1966; 7.1 mi. NNW of Xiatil; 4.0 mi. E of Xpujil (Thompson 1967). YUCATN: 8 mi. S of Progreso; near Chichen Itza (Harry 1950); Chichen Itza (Bequaert & Clench 1933); 0.5 mi. E of Becanchn; 10.0 mi. SE of Becanchn (Thompson 1967); 1.5 mi. S off Libre Union (Branson & McCoy 1963). Neocyclotus dysoni cookei (Bartsch & Morrison 1942)Aperostoma (Neocyclotus ) dysoni cookei Bartsch & Morrison 1942; Neocyclotus dysoni cookei (Bartsch & Morrison 1942). Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):6.Rehder 1966; Proc. Biol. Soc. Wash. 79:278.Thompson 1967; Bull. Fla. St. Mus. 11:228.Type Locality.Uaxactun, Dept, Petn, Guatemala. Holotype USNM 524006. Distribution.BELIZE: near Punta Gorda; foot of a limestone hill, left bank of the Rio Belize, 1.5 mi. S of Cayo; W of Gales Point (Bartsch & Morrison 1942). GUATEMALA, Dept. Petn: Magalango; 2 km S of Flores; 2 km S of Puebla Nueva (Bartsch & Morrison 1942); Tikal National Park (Basch 1959). CAMPECHE: 7.2 mi. S of Pixtun; 19.2 mi. E of Silvituc; 10.2 mi. E of Escarcega (Thompson 1967). QUINTANA ROO: 12 km from Icaiche (Rehder 1966). Neocyclotus dysoni dyeri (Bartsch & Morrison 1942)Aperostoma (Neocyclotus ) dysoni dyeri Bartsch & Morrison 1942; Neocyclotus dysoni dyeri (Bartsch & Morrison). Thompson 1969; Zoologica 54:68.Type Locality.La Ceiba, Honduras. Holotype USNM 215592. (Thompson 1969). Neocyclotus dysoni hinkleyi (Bartsch & Morrison 1942)Aperostoma (Neocyclotus ) dysoni hinkleyi Bartsch & Morrison (shell). Type Locality.Guatemala. Holotype USNM 523968. Distribution.GUATEMALA, Dept. Alta Verapaz: Cacao Finca, Tres Aguas, near Senahu; Chama; Secanquim, 500 m alt. Dept. Izabal: Maya Farms, Quiriqua (Bartsch & Morrison 1942). Neocyclotus dysoni nicaraguensis (Bartsch & Morrison 1942)Aperostoma (Neocyclotus ) dysoni nicaraguensis Bartsch & 16 (shell). Neocyclotus dysoni nicaraguense (Bartsch & Morrison). Thompson

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 29 524005. Distribution.COSTA RICA, Prov. Guanacaste:3.8 mi. S of Nicoya; 2.2 mi. SE of Nicoya, 500 ft. alt.; 1.2 mi. E of Caimital (Thompson 1969). NICARAGUA, Dept. Grenada, Dirio [Diria ?] (Bartsch & Morrison 1942). Numerous localities in the Dept. Estel, Dept. Matagalpa, Dept. Managua, Dept. Leon Neocyclotus dysoni ruatanensus (Bartsch & Morrison 1942)Aperostoma (Neocyclotus ) dysoni ruatanensus Bartsch & Morrison Neocyclotus dysoni var. ruatanensus (Bartsch & Morrison). Solem 1956; Proc. Acad. Nat. Sci. Phila. 108:54.Type Locality.Ruatan [Roatn] Island, Dept. Islas de la Baha, Honduras.Holotype USNM 364702. Distribution.Known only from the type locality. Neocyclotus dysoni sallei (Bartsch & Morrison 1942)Cyclotus (Aperostoma) dysoni var. minor Von Martens 1890: pl. 1, Cyclotus corrugatior minor Chitty 1857). Poteria (Neocyclotus) dysoni minor (Von Martens). H. B. Baker 1923; Occ. Pap. Mus. Zool. Univ. Mich. (137) 33. Aperostoma (Neocyclotus ) dysoni sallei Bartsch & Morrison 1942; Neocyclotus dysoni sallei (Bartsch & Morrison 1942). Solem 1956; Proc. Acad. Nat. Sci. Phila. 108:53.Type Locality.Yucatn, Mxico. Distribution.Known only from the type locality. Neocyclotus dysoni sumichrasti (Bartsch & Morrison 1942)Aperostoma (Neocyclotus ) dysoni sumichrasti Bartsch & Morrison (shell).Type Locality.Chontales forest, Nicaragua. Holotype USNM 523674. Distribution.Known only from the type locality. (Bartsch & Morrison 1942)Aperostoma (Neocyclotus ) Bartsch & Morrison (shell).Type Locality.Cervantes, [Prov. Cartago], Costa Rica. Holotype USNM 524003. Distribution.Known only from the type locality. Neocyclotus panamensis Da Costa 1904Neocyclotus panamensis Da Costa 1904; Proc. Malac. Soc. London Poteria (Neocyclotus) panamensis (Da Costa). H. B. Baker 1923; Occ. Pap. Mus. Zool. Univ. Mich. (137) 37. Aperostoma (Austrocyclotus ) panamense (Da Costa). Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. 181:197; pl. 27, Type Locality.Chiriqu, Panam. Distribution.Known only from the type locality. Neocyclotus simplicostus Thompson 1969Neocyclotus simplicostus Thompson 1969; Zoologica 54:68; system).Type Locality.A ravine 4.2 mi. northwest of Escuintla, Chiapas; 300 ft. alt. Holotype UF 20167. Distribution.CHIAPAS: 21.3 mi. NW of Huixtla, 300 ft. alt.; 32.5 mi. NW of Huixtla, 200 ft. alt. (Thompson 1969). Genus Incidostoma Bartsch & Morrison 1942Aperostoma (Incidostoma) Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. 181:187. Incidostoma Morrison 1955; Jour. Wash. Acad. Sci. 45:155. Neocyclotus (Incidostoma), Thompson 1969; Zoologica 54:73.Type Species. Aperostoma (Incidostoma) malleatum Bartsch & Morrison 1942. Distribution.Costa Rica south to Bolivia and Brazil. Taxonomy.Aperostoma Troschel was used by Bartsch and Morrison (1942:187) for a large group of mainland species. The genus name is not available for this group because Herrmannsen (1852) designated Cyclostoma mexicanum as the type species, which belongs in a different subfamily, the Megalomastomatinae. Of the eight subgenera of Aperostoma recognized by Bartsch (1942), Bartsch and Morrison (1942), and Morrison (1955), Neocyclotus is the oldest available name, and Incidostoma is the next oldest available name. Incidostoma and Neocyclotus as separate genera, following Morrison (1955). Bartsch (1942) and Bartsch and Morrison (1942) listed 72 species in Aperostoma (= Incidostoma as used here). Ten species occur in the study area. Incidostoma brujense (Bartsch & Morrison 1942)Aperostoma (Aperostoma) brujense Bartsch & Morrison 1942; Bull. Type Locality.Cerro Bruja, Panam; 500 ft. alt. Holotype USNM 251418. Distribution.PANAM: mountains around Gaspasalana, High Mamoni (Bartsch & Morrison 1942). Incidostoma carmioli (Bartsch & Morrison 1942)Aperostoma (Incidostoma) carmioli Bartsch & Morrison 1942; Bull. Neocyclotus (Incidostoma) carmioli (Bartsch & Morrison). Thompson 1969; Zoologica 54:74.Type Locality.Chitaria, Costa Rica. Holotype USNM 405227. Distribution.COSTA RICA, Prov. Cartago: Rio Chitaria, 2 km. N of Jabillas, 3000 ft. alt. (Thompson 1969). Incidostoma confusum (Sykes 1901)Cyclostoma giganteum Reeve. Pfeiffer 1848; in Martini & Chemnitz Aperostoma confusum Sykes 1901; Jour. of Malac., 8:106; pl. 10, Poteria (Neocycolus) confusum (Sykes). H. B. Baker 1923; Occ.

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30 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Pap. Mus. Zool. Univ. Mich. (137) 42. Aperostoma (Aperostoma) confusum (Sykes). Bartsch & Morrison Type Locality.Panam. Distribution.COLOMBIA, Dept. Choco: Acand, a small village in the Gulf of Atrato, on the boundary between Panam and Colombia (Bartsch & Morrison 1942). Incidostoma costaricense (Von Martens 1876)Cyclotus quitesis costaricensis Von Martens 1876:257. Aperostoma (Incidosoma) costaricense (Von Martens). Bartsch & 16 (shell).Type Locality.Costa Rica. Distribution.COSTA RICA (?): unknown. Incidostoma exiguum (Bartsch & Morrison 1942)Aperostoma (Incidostoma) exiguum Bartsch & Morrison 1942; Bull. Type Locality.Zorquin Valley, Talamanca, Costa Rica. Holotype USNM 190281. Distribution.Known only from the type locality. Incidostoma gigantea (Reeve 1842)Cyclostoma giganteum Reeve 1842; Conchologica systematica, 2: Neocyclotus giganteus (Reeve). Kobelt & Mllendorff 1897; Nachrb. Deut. Malak. Ges. 29:137. Aperostoma giganteum (shell). Poteria (Neocyclotus) gigantea (Reeve). H. B. Baker 1923; Occ. Pap. Mus. Zool. Univ. Mich. (137) 40, 42. Aperostoma (Incidostoma) giganteum (Reeve). Bartsch & Morrison (shell). Incidostoma gigantea (Reeve). Morrison 1955; Jour. Wash. Acad. Sci. 45:157.Type Locality.In woods near Panam, Panam. Distribution.PANAM: Cerro de Garagara, 20 m alt. (Bartsch & Morrison 1942). Incidostoma impressum (Thompson 1969)Neocyclotus (Incidostoma) impressus Thompson 1969; Zoologica (female reproductive system).Type Locality.Along road immediaely east of Nueva 20151. (Thompson 1969). Incidostoma irregulare (Pfeiffer 1855)Cyclostoma (Cyclotus) irregulare Pfeiffer 1855; Proc. Zool. Soc. Lond. 23:117. Cyclotus irregularis (Pfeiffer). Reeve 1864; Conch. Iconica, 14: pl. Cyclotus (Aperostoma) irregularis (Pfeiffer). Von Martens 1890; Biol. Cent. Amer.:3.Von Martens 1900; Biol. Cent. Amer.:596. Poteria (Neocyclotus) irregulare (Pfeiffer). H. B. Baker 1923; Occ. Pap. Mus. Zool. Univ. Mich. (137):43. Aperostoma (Aperostoma) irregulare (Pfeiffer). Bartsch & Morrison Neocyclotus (Incidostoma) irregulare (Pfeiffer). Thompson 1969; Zoologica 54:74.Type Locality.Costa Rica. Distribution.COSTA RICA: Cache; Talamanca (Von Martens 1890); San Mateo, 250 m alt.; Ravine of Vijaqual in the valley of San Savenque; El Pital, in the alley of the Rio Naranjo; Alto de Mano Tigre, in the valley of the Rio Grande de Terraba; Quebrada de Tocori, in the valley of the (Thompson 1969). Incidostoma pittieri (Von Martens 1900)Cyclotus (Aperostoma) irregularis pittieri Von Martens 1890; Biol. Cent. Amer.:597. Poteria (Noecyclotus ) irregularis pittieri (Von Martens). H. B. Baker 1923; Occ. Pap. Mus. Zool. Univ. Mich. (137) 41. Aperostoma (Aperostoma) pittieri (Von Martens). Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. 181:236.Type Locality.Salinas Bay northwestern Costa Rica. Distribution.Known only from the type locality. Incidostoma portobellense (Bartsch & Morrison 1942)Aperostoma (Aperostoma) portobellense Bartsch & Morrison 1942; Type Locality.Porto [Puerto] Bello, Panam. Holotype USNM 251434. Distribution.Known only from the type locality. Genus Dicrista Thompson 1969Dicrista Thompson 1969; Zoologica 54:42.Type Species.Dicrista liobasis Thompson 1969 (by original designation). Distribution.West coast of Mxico from Sinaloa south to Oaxaca, Costa Rica (?). Taxonomy.Seven species are recognized. Dicrista cooperi (Tryon 1863)Cyclotus cooperi Tryon 1863; Proc. Acad. Nat. Sci. Phila.:281; pl. Mexcyclotus cooperi (Tryon). Bartsch & Morrison 1842:180; pl. 24, Mexcyclotus lutescens (Pfeiffer). Bartsch & Morrison 1942; Bull. U. Dicrista cooperi (Tryon). Thompson 1969; Zoologica 54:43; pl. Type Locality.Near Mazatln, Sinaloa, Mxico. Holotype ANSP 13019. Distribution.SINALOA: Rosario. JALISCO: Hacienda de Istapa; San Sebastian. COLIMA: 7.6 mi. NNE of Manzanilla, 200 ft. alt. GUERRERO: Zihuatenajo.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 31 locality. (All records from Thompson 1969.) Dicrista damianensis (Solem 1956)Mexcyclotus damianensis Solem 1956; Proc. Acad. Nat. Sci. Phila. Dicrista damianensis (Solem). Thompson 1969; Zoologica (operculum).Type Locality. mile north of San Pedro Damian Holotype UMMZ 184837. Distribution.Known only from the type locality. Thompson 1969 Thompson 1969; Zoologica 54:49; pl. II, Type Locality..8 miles south of Mazatln, Guerrero, Mxico. 3500 ft. alt. Holotype UF 20197. Distribution.Known only from the type locality. Dicrista indentata Thompson 1969Dicrista indentata Type Locality..0 miles southeast of San Vicente, Michoacn, Mxico. 200 ft. alt. Holotype UF 20191. Distribution.Known only from the type locality. Dicrista liobasis Thompson 1969Dicrista liobasis Thompson 1969; Zoologica 54:48; pl. III, reproductive system).Type Locality.A limestone sink 6.0 miles southwest and 6.6 miles east of Pihuamo, Jalisco, Mxico. [ca. 19 N, 103 W]; 2000 ft. alt. Holotype UF 20194. Distribution.Known only from the type locality. Dicrista petersi (Solem 1956)Mexcyclotus petersi Solem 1956; Proc. Acad. Nat. Sci. Phila. Dicrista petersi (Solem). Thompson 1969; Zoologica 54:52; p. IV, Type Locality.La Placita [Sulatillo], Michoacn,, Distribution.Known only from the type locality. Dicrista rugosa Thompson 1969Dicrista rugosa C-D (male reproductive system).Type Locality.A small limestone knoll 1.4 miles east of Colima, Mxico.; 1800 ft. alt. Holotype UF 20199. Distribution.Known only from the type locality. Genus Xenocyclus Thompson 1969Xenocyclus Thompson 1969; Zoologica 54:55.Type Species.Xenocyclus patulus Thompson 1969 (by original designation). Distribution.Colima, Mxico. Taxonomy.A single species is recognized. Xenocyclus patulus Thompson 1969Xenocyclus patulus Thompson 1969; Zoologica 54:54: pl. 5, reproductive system).Type Locality.A limestone ridge 0.3 miles southeast of Tamala, Mxico. 500 ft. alt. Holotype UF 20185. Distribution.Known only from the type locality. Genus Amphicyclotus Crosse & Fischer 1879Amphicyclotus Crosse & Fischer 1879; Jour. de Conchyl. 27:46.Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. 181:183.Morrison 1955; Jour. Wash. Acad. Sci. 45:160.Solem 1956; Proc. Acad. Nat. Sci. Phila. 108:43.Thompson 1969; Zoologica 54:56. Megacyclotus Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. 181:181.Type Species.Amphicyclotus : Cyclostoma (Cyclophorus) boucardi Pfeiffer 1856, by original designation. Megacyclotus : Cyclostoma ponderosum Pfeiffer 1851. Distribution.Southeast Mxico, Guatemala, Belize, El Salvador and Honduras. Taxonomy.Eight species and two subspecies are recognized. Amphicyclotus boucardi (Pfeiffer 1856)Cyclostoma (Cyclophorus) boucardi Pfeiffer 1856; Proc. Zool. Soc. Cyclophorus boucardi (Pfeiffer). Pfeiffer 1858; Monogr. pnemon. viv., 2:65. Amphicyclotus boucardi (Pfeiffer). Fischer & Crosse 1880:pl. 4 (shell). Cyclophorus (Amphicyclotus ) boucardi (Pfeiffer). Von Martens 1890; Biol. Cent. Amer.:6. Distribution.Known only from the type locality. Amphicyclotus maleri Crosse & Fischer 1883Amphicyclotus maleri Crosse & Fischer 1883:102.Fischer & 1888:145.Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. Cyclophorus (Amphicyclotus) maleri (Crosse & Fischer). Von Martens 1890; Biol. Cent. Amer.:6.Type Locality.Tabasco, Mxico. (Von Martens 1890). TABASCO: Sierra Poan (Solem 1956a). Amphicyclotus megaplanus Morrison 1955Amphicyclotus megaplanus Morrison 1955; Jour. Wash. Acad. Sci.

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32 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Type Locality.Forests of Ocote, near Ocozocoantla [Ocozocoautla], Chiapas, Mxico; 600 m alt. Holotype USNM 618777. Distribution.CHIAPAS: 15.8 mi. NW of Ocozocoautla, 2700 ft. alt. (Thompson 1969). Amphicyclotus palenquensis (Pilsbry 1935)Aperostoma (Amphicyclotus ) palenquense Pilsbry 1935; Proc. Acad. Megacyclotus palenquensis (Pilsbry). Bartsch & Morrison 1942; Amphicyclotus (A.) palenquensis (Pilsbry). Solem 1956; Proc. Acad. Nat. Sci. Phila. 108:44.Type Locality.District of Palenque, Chiapas, Mxico. Holotype ANSP 106344. Distribution.CHIAPAS: Palenque. VERACRUZ: Motzorongo (Solem 1956a). Thompson 1963 system).Type Locality.Hacienda Monte Cristo, Metapn, Dept. Santa Ana, El Salvador; 2200 m alt. Holotype UMMZ 195882. Distribution.Known only from the type locality. Amphicyclotus paulsonorum Thompson 1969Amphicyclotus paulsonorum Thompson 1969; Zoologica 54:61; Type Locality.A ravine 4.2 miles northwest of Escuintla, Chiapas, Mxico; 300 ft. alt. Holotype UF 20180. Distribution.CHIAPAS:21.3 mi. NW of Huixtla, 300 ft. alt.; 32.5 mi. NW of Huixtla, 200 ft. alt.; 10 mi. NW of Pijijiapan, 100 ft. alt. (Thompson 1969). Amphicyclotus ponderosus (Pfeiffer 1851)Cyclostoma ponderosum Pfeiffer 1851; Proc. Zool. Soc. Lond. 19:283. Cyclophorus ponderosus (Pfeiffer). Pfeiffer 1852; Monogr. pneumon. viv. 1:97. Amphicyclotus ponderosus (Pfeiffer). Fischer & Crosse 1880: pl. 35, Acad. Nat. Sci. Phila. 108:46. Cyclophorus (Amphicyclotus ) ponderosus (Pfeiffer). Von Martens 1890; Biol. Cent. Amer.:5.Von Martens 1900:597. Megacyclotus ponderosus (Pfeiffer). Bartsch & Morrison 1942; Type Locality.Not given. Distribution.GUATEMALA, Dept. Alta Verapaz: Cobn; Senahu (Von Martens 1890); San Juan (Von Martens 1900); Chama; Secanquin, 550 m alt. (Bartsch & Morrison 1942). Amphicyclotus texturatus texturatus (Sowerby 1850)Cyclostoma texturatum Sowerby 1850; Thesaurus conchyliorum Amphicyclotus texturatus (Sowerby). Fischer & Crosse 1880: pl. 1 (shell). Amphicyclotus texturatus texturatus (Sowerby). Solem 1956; Proc. Acad. Nat. Sci. Phila. 108:45.Type Locality.Guatemala. Distribution.GUATEMALA, Dept. Alta Verapaz: Cobn; Finca Providencia; Finca Caluchen; Finca Samac (Solem 1956a). CHIAPAS: Chiquihuite; Soconusco (Solem 1956a). Amphicyclotus texturatus goldfussi (Boettger 1892)Aperostoma (Amphicyclotus ) goldfussi Boettger 1892; Nach. Bltt. Deut. Malak. Ges., 24:203. Amphicyclotus goldfussi (Boettger). Kobelt & Mllendorff 1907; Nach. Bltt. Deut. Malak. Ges., 29:139.Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. 181:184. Cyclophorus (Amphicyclotus ) texturatus goldfussi (Boettger). Von Martens 1900; Biol. Cent. Amer.:598. Amphicyclotus texturatus goldfussi (Boettger). Solem 1956; Proc. Acad. Nat. Sci. Phila. 108:145.Type Locality.San Pedro Sula, Honduras. (Solem 1956a). Amphicyclotus texturatus spiralis Thompson 1969Amphicyclotus (A.) texturatus texturatus (Sowerby). Solem 1956; Proc. Acad. Nat. Sci. Phila. 108:45 ( in part). Amphicyclotus texturatus spiralis Thompson 1969; Zoologica system).Type Locality.Coffee grove 4.7 mi. north-northeast of Huixtla, Chiapas, Mxico; 600 ft. alt. Holotype UF 20176. Distribution.CHIAPAS: Escuintla (Solem 1956a); 12.7 mi. NNE of Huixtla, 1500 ft. alt.; 14.6 mi. NNE of Huixtla, 2100 ft. alt. (Thompson 1969). Genus Barbacyclus Bartsch & Morrison 1942Barbacyclus Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. 181:175.Thompson 1969; Zoologica 54:64.Type Species.Cyclophorus underwoodi Da Costa 1900, by original designation. Distribution.Costa Rica. Taxonomy.Three species are recognized. Barbacyclus boucardi (Angas 1878)Cyclotus boucardi Angas 1878; Proc. Zool. Soc. Lond. 46:73; pl. 5, Neocyclotus (Neocyclotus ) boucardi (Angas). Kobelt & Mllendorff 1907; Nach. Bltt. Deut. Malak. Ges., 29:137. Aperostoma princeps angasianum Pilsbry 1935:3. Barbacyclus boucardi (Angas). Bartsch & Morrison 1942; Bull. U.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 33 Type Locality.San Carlos, Costa Rica. near Guapiles (Pilsbry 1935). Barbacyclus princeps (Pilsbry 1935)Cyclotus boucardi Angas 1879; Proc. Zool. Soc. Lond. 47:483 (not Cyclotus boucardi Angas 1878:73). Aperostoma (Amphicyclotus ) princeps 1b (shell). Barbacyclus princeps (Pilsbry). Bartsch & Morrison 1942; Bull. U. system).Type Locality.Mouth of the Rio Banana, 5 miles from (Thompson 1969). Barbacyclus underwoodi (Da Costa 1900)Cyclophorus underwoodi Da Costa 1900; Proc. Malac. Soc. London Cyclophorus (Amphicyclotus ) underwoodi Da Costa. Von Martens 1900; Biol. Cent. Amer.:598. Amphicyclotus underwoodi (Da Costa). Kobelt 1902; Das Tierreich, Cyclophoridae:257.Kobelt 1912; in Martini & Chemnitz Babacyclus underwoodi (Da Costa). Bartsch & Morrison 1942; Type Locality.Carillo, Costa Rica. Distribution.Known only from the type locality. Genus Calacyclotus Bartsch & Morrison 1942Calacyclotus Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. 181:178.Type Species.Amphicyclotus olssoni Pilsbry 1926 by original designation. Distribution.Panam and immediately adjacent Colombia. Taxonomy.Two species are known. Calacyclotus atratensis Bartsch & Morrison 1942Calacyclotus atratensis Bartsch & Morrison 1942; Bull. U. S. Nat. Type Locality.Mountains near the mouth of the Rio Atrato in Colombia. Holotype USNM 206291. Distribution.PANAM-COLOMBIA: known only from the type locality. The Rio Atrato lies on the border between Colombia and Panam. Calacyclotus olssoni (Pilsbry 1926)Amphicyclotus olssoni Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. Calacyclotus olssoni (Pilsbry). Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. 181:178. Holotype ANSP 47354. Distribution.COLOMBIA, Dept. Choco: Acand, Gulf of Atrato (Bartsch & Morrison 1942). Neocyclotidae of doubtful occurrence in Mxico and Central America Mexcyclotus lutescens (Pfeiffer 1851)Cyclosytoma lutescens Pfeiffer 1851; Proc. Zool. Soc. Lond. 19: 250. Mexcyclotus lutescens (Pfeiffer). Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. 181:179.Solem 1956; Proc. Acad. Nat. Sci. Phila. 108:55.Thompson 1969; Zoologica 54:61.Taxonomy.The species name is a nomen dubium (Thompson 1969; Zoologica 54:41). As a consequence the genus name is also a nomen dubium described from Brazil, but was later credited to Mxico. The syntypes lack opercula, and the identity of the species cannot be determined with certainty. This, coupled with the doubtful untenable. Family DIPLOMMATINIDAE Pfeiffer 1856 Distribution.The family is widely distributed on mainland and in the Neotropical Region. The family includes numerous genera and species. A single genus occurs in the Neotropical Region. Genus Adelopoma Doering 1884Adelopoma Ancey 1899; Jour. de Conchyl. 47:194.Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. 181:148.Type Species.Adelopoma tucma Doering 1884. Distribution.Eastern Mxico south to Argentina. Taxonomy.Eight species are recognized. Two occur in the study area. Adelopoma costaricense Bartsch & Morrison 1942Adelopoma costaricense Bartsch & Morrison 1942; Bull. U. S. Malac. Rev. 33/34:92.Type Locality.Santa Mara, Costa Rica; 1550 m alt. Holotype USNM 516034. Distribution.Known only from the type locality. NICARAGUA, Dept. Jinotega: Apans; Cartuja; Selva Negra Adelopoma stolli (Von Martens 1890)Diplommatina stolli Von Martens 1890; Biol. Cent. Amer.:20; Palaina (Eupalaina) stolli (Von Martens). Kobelt & Moellendorff 1898:133. Adelopoma Stolli (Von Martens). Ancey 1899; Jour. de Conchyl. 47:197.Kobelt 1902:480.Hinkley 1907; Nautilus 21:78.Bartsch & Morrison 1942; Bull. U. S. Nat. Mus. 181:149; 2001; Malac. Rev. 33/34:92.Type Locality.NW Guatemala: in the District

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34 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Cholhuitz, on the slope of the Volcn de Santa Mara, at the plantation Helvitia. I have not located this place with certainty. The Global Gazetteer Version 2.1. Jan 26, 2006, lists Cholhuitz, Dept. Huehuetenango (15.87 N, 91.55 W). Distribution.Reported from northeastern Mxico, Guatemala and Nicaragua. GUATEMALA: only from the type locality. NICARAGUA, Dept. Jinotega: Puene Pra; Las TAMAULIPAS: Tampico, in river drift (Hinkley 1907); El Cielo Biosphere Reserve (Correa-Sadoval & Rodriguez 2005). Superfamily CERITHIOIDEA Flemming 1822 Family PLEUROCERIDAE Fischer 1885 Distribution.North and South America, east Asia, and Africa. The geographic restrictions of the family are poorly understood because of the unsatisfactory taxonomy and phylogeny of the many genera placed here. Taxonomy.Ten genera are recognized in North America. One occurs in Mxico. Genus Lithasiopsis Pilsbry 1910Lithasiopsis Pilsbry 1910; Proc. Malac. Soc. London 9:47.Thompson 1959; Occ. Pap. Univ. Mich. Mus. Zool., (600):1. Pachychilus (Oxymelania). Morrison 1954; Proc. U. S. Nat. Mus. 103:365.Type Species.Litasiopsis hinkleyi Pilsbry 1910. the Rio Guayalejo system of northeastern Mxico. Taxonomy.Four species are recognized. Lithasiopsis crassus Thompson 1959Lithasiopsis crassa Thompson 1959; Occ. Pap. Univ. Mich. Mus. Type Locality.Rio Sabinas [Rio Guayalejo system], Pano Ayuctle, 5 miles NE of Gomez, Farias, Tamaulipas, Mxico. Holotype UMMZ 195130. Distribution.TAMAULIPAS: known only from the Rio Sabinas (Thompson 1959). Lithasiopsis darnelli Thompson 1959Lithasiopsis darnelli Thompson 1959; Occ. Pap. Univ. Mich. Mus. Type Locality.Rio Sabinas [Rio Guayalejo system], 195098. Distribution.TAMAULIPAS: known only from the Rio Sabinas (Thompson 1959). Lithasiopis hinkleyi Pilsbry 1910Lithasiopis hinkleyi Pilsbry 1910; Proc. Malac. Soc. London 9:48; Type Locality.Rio Coy [Rio Panuco system], San Lus Potos, Mxico. Holotype in the ANSP. Distribution.Known only from the type locality. Lithasiopsis mexicanus Pilsbry 1910Lithasiopsis mexicanus Pilsbry 1910; Proc. Malac. Soc. London Type Locality.Rio Montezuma [= Moctezuma] [Rio Panuco system], at the ford, a short distance above Distribution.Known only from the type locality. Family PACHYCHILIDAE Troschel 1857 Distribution.Mxico south to northern South America, southeastern Asia, Africa. Taxonomy.As with the Pleuroceridae, the geographic, morphological, taxonomic, and phylogenetic parameters relating to this family remain poorly understood. Two genera are recognized in Mxico and Central America. Genus Amnipila Pilsbry 1956Amnipila Pilsbry 1956, Proc. Acad. Nat. Sci. Phila. 108:38. Type Species.Pachychilus pila Pilsbry & Hinkley 1910. Distribution.Rio Tamasopo, San Lus Potos, Mxico. Taxonomy.The genus is monotypic. Amnipila pila (Pilsbry & Hinkley 1910)Pachychilus pila Pilsbry & Hinkley 1910, Proc. Acad. Nat. Sci. Amnipila pila (Pilsbry & Hinkley). Pilsbry 1956, Proc. Acad. Nat. Type Locality.Rio Tamosopo above and below the natural Bridge, near Verastuagu, San Lus Potos, Mxico. Holotype ANSP 99559 (Pilsbry 1956). Distribution.SAN LUS POTOS: known only from the immediate vicinity of the type locality. Genus Pachychilus I. Lea and H. C. Lea 1850Pachychilus I. and H. C. Lea 1850; Proc. Zool. Soc. Lond. 18:179.Brot 1874; in Martini & Chemnitz, Syst. Conch. Cab. (Melaniaceen):19.Fischer & Crosse 1892; Mission Sci. Mex., II:320.Von Martens 1899; Biol. Cent. Amer.:435 464.Morrison 1954; Proc. U. S. Nat. Mus. 103:364. Cercimelania Crosse & Fischer 1892; Miss. Sci. Mex. II:327, 328, 340. Sphaeromelania Rovereto 1899; Atti. Soc. Ligust. Sci. Nat. Geogr., 10:109 (substitute name for Pachychilus ).Distribution.Eastern Mxico from Tamaulipas south to the Isthmus of Tehuantepc and from there southeast along east and west coast drainages at lower elevations to Nicaragua. From there the range is disjunct to the area near Puerto Cabello, Venezuela. Three species occur in Cuba. Taxonomy.Pachychilus are large, handsome aquatic snails. Generally they are abundant, and they are the most conspicuous invertebrates present in streams and lakes where they occur. The snails have very limited dispersal ability.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 35 Populations are limited genetically and geographically, and numerous forms occur in nature. Because of their abundance and conspicuous appearance specimens found their way into the laboratories of early European and American naturalists. account for the observed variation. Whether these represent valid taxa or not remains to be determined. For the most part the taxonomic arrangement used herein follows Von Martens (1899), with subsequently described species inserted where it seems appropriate. Fischer and Crosse (1892) provided an exhaustive synonymy of the species known to then. Von Martens (1899) summarized subsequent literature. The reader is referred to the monographs by Reeve (1861), Brot (1874), Fischer & Crosse (1892), and Von Martens (1899) for aid in identifying the earlier known species. Subgenus Pachychilus I. Lea & H. C. Lea 1850Pachychilus I. Lea & H. Lea 1850; Proc. Zool. Soc. Lond. 18:179. Cercimelania Fischer & Crosse 1892; Miss. Sci. Mex. II:327, 328, 340.Type Species.Melania Sowerby 1824 Cercimelania : Melania liebmanni Philippi 1848. Distribution.Eastern Mxico from Veracruz south to Nicaragua. One species, occurs in Venezuela. Three species occur in Cuba. been proposed for members of this subgenus. Pachychilus (Pachychilus) apis (Lea 1850)Melania apis Lea 1850; Proc. Zool. Soc. Lond. 18:90.Reeve 1861; in Martini Pachychilus apis (Lea). Fischer & Crosse 1892; Miss. Sci. Mex. II:356.Von Martens 1899; Biol. Cent. Amer.:455.Type Locality.Vera Cruz, in marshy places. Distribution.Known only from the type locality. Pachychilus (Pachychilus) chrysalis chrysalis (Brot 1872)Melania chrysalis Brot 1872; Mtereiaux fam. Melaniens, 3:39: pl. Pachychilus chrysalis (Brot). Fischer & Crosse 1892; Miss. Sci. Sci. Phila. 44:340.Von Martens 1899; Biol. Cent. Amer.:457; Biol. Cent. Amer.:646. Pachychilus chrysalis var. Fischer & Crosse 1892; Miss. Fischer & Crosse. Pilsbry 1900; Nautilus 13:139. Brot 1874; in Martini & Chemnitz, Syst. & Crosse 1892; Miss. Sci. Mex. II:344.Type Localities. Melania chrysalis: not given. Pachychilus chrysalis var. : San Pedro Gineta, Isthmus of Tehuantepec, Oaxaca, Mxico. Pachychilus : San Pedro Gineta, Isthmus of Tehuantepec, Oaxaca, Mxico. Distribution.CHIAPAS: San Pedro Jineta; Ixtacomitan. TABASCO: Teapa; Rio Puyacatengo, near Teapa (Pilsbry 1900). NICARAGUA: Lago de Managua (Von Martens 1901). Pachychilus (Pachychilus) chrysalis nympha Von Martens 1899Pachychilus chrysalis var. nympha Von Martens 1899; Biol. Cent. Type Locality.Rio Sucio, El Salvador. Distribution.Known only from the type locality. Pachychilus ( Pachychilus) (Morelet 1849) Morelet 1849; Test. Noviss. I:26.Reeve 1861; in Martini (Morelet).H. & A. Adams 1854; Gen. (Morelet). Fischer & Crosse 1892; Miss. Sci. Sci. Phila. 44:340.Goodrich & Van der Schalie 1937; Misc. Pub. Mus. Zool. Univ. Mich. (34):41. Melania cinerea Morelet 1849; Test. Noviss. I:26.Reeve 1861; in Martini Pachychilus cinereus (Morelet). Fischer & Crosse 1892; Miss. Sci. Cent. Amer.:459. Melania panucula Morelet 1851; Test. Noviss. II:23.Brot 1874; in Martini & Chemnitz, Syst. Conch. Cab. (Melaniaceen):27; pl. Pachychilus panucula (Morelet). Fischer & Crosse 1892; Miss. Sci. Amer.:458. Melania tumidus Tristram 1863; Proc. Zool. Soc. Lond. 31:413. Melania tumida Tristram. Brot 1874; in Martini & Chemnitz, Syst. Conch. Cab. (Melaniaceen):23. Pachychilus tumidus (Tristram). Fischer & Crosse 1892; Miss. Sci. Mex. II:357. Pachychilus panucula var. tumidus (Tristram). Von Martens 1899; Melania mexicana Reeve 1861; Conch. Icon. (Melania 129.Brot 1874; in Martini & Chemnitz, Syst. Conch. Cab. Pachychilus mexicanus (Reeve). Fischer & Crosse 1892; Miss. Sci. Mex. II:340. var. lutescens Fischer & Crosse 1892; Miss. Type Localities. : small streams of the Department of Verapaz, Guatemala. Melania cinerea: in the rivers of Cobn, Alta Vera Paz, Guatemala. Melania panucula: rivers of the Department of Petn, Guatemala. Melania tumida: lake of Petn, Guatemala. Melania mexicana : Mxico. var. lutescens: Cobn, [Dept. Alta Verapaz], Guatemala. Distribution.TABASCO: Sierra Poan. GUATEMALA, Dept. Alta Verapaz: Chejel; Chama; Tactc; Cobn; between Chama and Cobn (Hinkley 1920); Rio de la Pasion

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36 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) system (Goodrich & Van der Schalie 1937). Dept. Baja Verapaz: Stream of Santa Rosa. Dept. Izabal: Rio Cavech; Rio Dulce near Livingston (Hinkley 1920). NICARAGUA: Rio Tungla, near Quiquina (Fluck 1905). Pachychilus (Pachychilus) Crosse & Fischer 1891Pachychilus indifferens Crosse & Fischer 1891; Jour. de Chonchyl., 39:25.Fischer & Crosse 1892; Miss. Sci. Mex. II:337; pl. 50, Crosse & Fischer. Goodrich & Van der Schalie 1937; Misc. Pub. Univ. Mich. Mus. Zool. (34):41.Type Locality.Rio Motagua, Guatemala. Distribution.GUATEMALA, Dept. Baja Verapaz: Rio Motagua. Dept. Petn: Lago de Petn; Lago de Eckibix (Goodrich & Van der Schalie 1937). Pachychilus ( Pachychilus) explicatus Fischer & Crosse 1892Pachychilus indifferens var. explicata Fischer & Crosse 1892; Miss. Pachychilus explicatus Fischer & Crosse. Von Martens 1899; Biol. Cent. Amer.:459.Type Locality.Rio Machaquila, Dept. Petn, Guatemala. Distribution.Known only from the type locality. Pachychilus (Pachychilus) hinkleyi (Marshall 1921)Sphaeromelania hinkleyi Marshall 1921; Proc. U. S. Nat. Mus. Type Locality.Rio Tsalbha, Cham, Dept. Alta Verapaz, Guatemala. Holotype USNM 336412. Distribution.GUATEMALA, Dept. Alta Verapaz: Chama (Hinkley 1920). Pachychilus (Pachychilus) indiorum (Morelet 1849)Melania indiorum Morelet 1849; Test. Noviss. I:25. Sowerby. Brot 1874; in Martini & Chemnitz, (Sowerby). H. & A. Adams 1854:299; pl. (Sowerby). Fischer & Crosse, 338. Pachycheilus indiorum (Morelet). H. & A. Adams 1854:299.Von Martens 1899; Biol. Cent. Amer.:455.Goodrich & Van der Schalie 1937:42. var. costato-plicata Brot 1874; in Martini & var. costatoplicatus (Brot). Fischer & Pachychilus indiorus var. costatoplicatus (Brot). Von Martens 1899; Biol. Cent. Amer.:456. var. Fischer & Crosse 1892; Pachychilus indiorum var. Fischer & Crosse. Von Martens 1899; Biol. Cent. Amer.:456. Melania sallei Melania radix Brot 1874; in Martini & Chemnitz, Syst. Conch. Cab. Type Localities. Melania indiorum: Near the ruins, Palenque, Chiapas, Mxico. var. costatoplicata : Palenque, Chiapas, Mxico. var.: Palenque, Chiapas, Mxico. Melania sallei : not given. Melania radix : not given. Distribution.GUATEMALA, Dept. Alta Verapaz: La Ceiba (Goodrich & Van der Schalie 1937); Panzos; Chejel (Hinkley 1920). Dept. Guatemala: Lago de Amatitln (Hinkley 1920). Dept. Izabal: Rio Cavech; Rio Dulce, near Livingston; Jocola; Panzos (Hinkley 1920). MXICO, CHIAPAS: Palenque. OAXACA: Isthmus of Tehuantepec. VERACRUZ: San Andres Tuxtla. Pachychilus (Pachychilus) liebmanni liebmanni (Philippi 1848)Melania liebmanni Conch. Icon. (Melania Pachychilus liebmanni (Philippi). Fischer & Crosse 1892; Miss. Sci. Mex. II:347.Von Martens 1899; Biol. Cent. Amer.:453; pl. 27, Type Locality.Mxico. Distribution.VERACRUZ: Playa Vicente (Von Martens 1899). Pachychilus ( Pachychilus) liebmanni gassiesi (Reeve 1861)Melania gassiesi 236. Pachychilus liebmanni var. gassiesi (Reeve). Von Martens 1899; Biol. Cent. Amer.:454.Type Locality.Central America. Distribution.Unknown. Pachychilus (Pachychilus) liebmanni gracilior Von Martens 1899Pachychilus liebmanni var. gracilior Von Martens 1899; Biol. Cent. Amer.:453. Melania gassiesi (Reeve) Gassies 1863:93.Brot 1874; in Martini & Pachychilus gassiesi (Reeve). Fischer & Crosse 1892; Miss. Sci. Type Locality.Rio Teotalcingo, Oaxaca, Mxico. Distribution.Known only from the type locality. Pachychilus (Pachychilus) oerstedi oerstedi (Mrch 1860)Pachychilus rstedi Mrch 1860; Malak. Bltt. 7:79.Von Martens Melania rstedi (Mrch). Brot 1874; in Martini & Chemnitz, Syst. Pachychilus jansoni H. Adams 1870. Proc. Biol. Soc. London:795.Brot 1874; in Martini & Chemnitz, Syst. Conch. Cab. (Melaniaceen):40.Type Locality.Pachychilus oerstedi: Segovia, Nicaragua. Pachychilus jansoni : Chontales, Nicaragua. Distribution.HONDURAS, Dept. Copn: Lancetilla (Pilsbry 1931:84). NICARAGUA, Dept. Chontales: Chontales. Dept. Matagalpa: Arroyo Alasan, Matagalpa. Remarks.Spelling of species and subspecies name rstedi is corrected to oerstedi following ICZN Article 32.5.2.1.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 37 Pachychilus (Pachychilus) oerstedi planensis (Lea 1858)Melania planensis Lea 1858; Proc. Acad. Nat. Sci. Phila.:118.Lea in Martini & Chemnitz, Syst. Conch. Cab. (Melaniaceen):33; pl. Pachychilus rstedi var. planensis (Lea). Von Martens 1899; Biol. Cent. Amer.:459. Pachychilus planensis (Lea). Goodrich & Van der Schalie 1937; Misc. Pub. Mus. Zool. Univ. Mich. (34):42.Type Locality.Plan and Omoa, valley of the Rio Ula, Honduras. Distribution.GUATEMALA, Dept. Izabal: Puerto Barrios (Goodrich & Van der Schalie 1937). HONDURAS: only from the type locality. Remarks.Spelling of species name rstedi is corrected to oerstedi following ICZN Article 32.5.2.1. Pachychilus (Pachychilus) pottsianus Hinkley 1920Pachychilus pottsianus Hinkley 1920; Nautilus 33:54.Type Locality.A small stream near Jocola, Dept. Izabal, Guatemala. Type specimen not designated; presumably in Hinkley Collection, Illinois Museum of Natural History, University of Illinois. Distribution.GUATEMALA, Dept. Izabal: known only from the immediate vicinity of the type locality. Jocola was a plantation on the north side of the Lago de Izabel. Pachychila pottsianus was collected from two small rills on a hillside behind Jocola. Pachychilus (Pachychil us) schumoi Pilsbry 1931Pachychilus schumoi (shell).Type Locality.Rio Negro [Rio Salinas], Cham, Dept. Alta Verapaz, Guatemala. Holotype ANSP 76231. Distribution.Known only from the type locality. Pachychilus (Pachychilus) turati (Villa 1854)Melania turati Villa 1854; in Strobel, Giornale di Malacologia 2:113.Brot 1874; in Martini & Chemnitz, Syst. Conch. Cab. Pachychilus turatii (Villa). Fischer & Crosse 1892; Miss. Sci. Melania gassiesi Type Locality.Not given. Distribution.VERACRUZ: Rio Atoyac; Rio Jamapa; Subgenus Pilsbrychilus Morrison 1952Pilsbrychilus Morrison 1952; American Malacological Union News Bulletin and Annual Report (1948):7.Morrison 1954; Proc. U. S. Nat. Mus. 103:366. Type Species.Pachyc heilus dalli Pilsbry 1896. Distribution.Southern Mxico. Taxonomy.A single species is recognized. Pachychilus (Pilsbrychilus) dalli Pilsbry 1896Pachycheilus dalli Pilsbry 1896; Science, 3:608.Pilsbry 1896; Proc. Acad. Nat. Sci. Phila. 48:269. Pachychilus dalli Pilsbry. Von Martens 1899; Biol. Cent. Amer.:456; Type Locality.Tehuantepec, Oaxaca, Mxico. Distribution.Known only from the type locality. The type locality most likely is the Isthmus of Tehuantepec, and not the city. Numerous searches have been made in the area Subgenus Glyptomelania Crosse & Fischer 1892Glyptomelania Fischer & Crosse 1892; Miss. Sci. Mex. II:328, 351.Von Martens 1899; Biol. Cent. Amer.:438.Morrison 1954; Proc. U. S. Nat. Mus. 103:366.Type Species.Melania glaphyra Morelet 1849. Distribution.Nicaragua, Honduras, El Salvador, and Guatemala north to Chiapas and Tabasco, Mxico. Taxonomy.Five species and twelve subspecies are recognized. Pachychilus (Glyptomelania ) glaphyrus glaphyrus (Morelet 1849)Melania glaphyra Morelet 1849; Test. Noviss. I:24.Reeve 1861; Conch. Icon. (Melania Pachychilus glaphyrus (Morelet). Fischer & Crosse 1892; Miss. Sci. Amer.:444.Goodrich & Van der Schalie 1937; Misc. Pub. Mus. Zool. Univ. Mich. (34):39. Type Locality.Not given. Distribution.GUATEMALA. Dept. Izabal: Jocolo (Hinkley 1920). Dept. Alta Verapaz: Arroyo Yalchactil, Rio de la Passion, ca. 4 miles southwest of La Ceiba (Goodrich & Van der Schalie 1939). Dept. Petn: Springs of the [Rio] Usumacinta (Fischer & Crosse 1892). Pachychilus (Glyptomelania ) glaphyrus bicarinatus Von Martens 1901Pachychilus glaphyrus var. Pilsbry 1893; Proc. Acad. Nat. Sci. Pachychilus glaphyrus var. bicarinatus Von Martens 1901; Biol. Cent. Amer.:445.Type Locality.Tabasco, Mxico. Distribution.Known only from the type locality. Pachychilus (Glyptomelania ) glaphyrus glaphyroides Fischer & Crosse 1892.Melania immanis var. Pachychilus glaphyrus glaphyroides Fischer & Crosse 1892; Miss. Sci. Mex. II:352.Von Martens 1899; Biol. Cent. Amer.:444.Type Locality.Springs of the Rio Usumacinta, Guatemala. Distribution.Known only from the type locality. Pachychilus (Glyptomelania ) glaphyrus immanis (Morelet 1851)Melania immanis Morelet 1851; Test. Noviss. II:22.Reeve 1861;

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38 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Pachychilus glaphyrus immanis (Morelet). Fischer & Crosse 1892; Miss. Sci. Mex. II:351; pl. 53: Figs. 1, 1a.Von Martens 1899; Biol. Cent. Amer.:438.Type Locality.Small streams in the Province of Petn, Guatemala. Distribution.GUATEMALA, Dept. Alta Verapaz: 1899:444). Pachychilus (Glyptomelania ) glaphyrus polygonotus (Lea 1850)Melania polygonotus Lea 1850:195.Reeve 1861; Conch. Icon. Pachychilus glaphyrus polygonotus (Lea). Fischer & Crosse 1892; Miss. Sci. Mex. II:351.Von Martens 1899; Biol. Cent. Amer.:444. Pilsbry 1892; Proc. Acad. Nat. Sci. Type Localities. Melania polygonotus: Copan. Syntypes in the USNM. (It is not clear whether Lea intended Copn, Dept. Santa Brbara, Honduras, or Cobn, Dept. Alta Verapaz, Guatemala, as suggested by Von Martens. If Copn is the type locality it seems highly unlikely that could be a synonym of polygonotus as suggested by Von Martens 1899:444.) Mxico. Holotype ANSP. Distribution.GUATEMALA, Dept. Alta Verapaz: Cobn; Dolores (Von Martens 1899). TABASCO: western 1893b). Pachychilus (Glyptomelania ) glaphyrus pyramidalis (Morelet 1851)Melania pyramidalis Morelet 1849; Test. Noviss. I:25. Melania opiparis Morelet. Reeve 1861; Conch. Icon. (Melania): pl. Melania godmani Tristram 1863:113. Pachychilus godmani (Tristram). Fischer & Crosse 1892; Miss. Sci. Mex. II:363. Pachychilus glaphyrus Pachychilus glaphyrus pyramidalis (Morelet). Von Martens 1899; Type Localities. Melania pyramidalis: Small streams of the interior of Tabasco, Mxico. Melania godmani: Lake Petn, Guatemala. Distribution.GUATEMALA, Dept. Alta Verapaz: Panzos (Hinkley 1920). Dept. Izabal: Jocolo (Hinkley 1920); Esmeralda (Hinkley 1920); Livingston (Hinkley 1920); across the Rio Dulce from Livingston (Hinkley 1920). Dept. Petn: San Lus; Lake Petn (Von Martens 1899). Pachychilus (Glyptomelania ) glaphyrus scamnatus Fischer & Crosse 1892Pachychilus glaphyrus scamnatus Fischer & Crosse 1892; Cent. Amer.:444.Type Locality.Cobn, Alta Verapaz, Guatemala. Distribution.Known only from the type locality. Pachychilus (Glyptomelania ) glaphyrus opilaris (Morelet 1851)Melania opiparis Morelet 1851:23. Pachychilus immanis Pachychilus glaphyrus var Fischer & Crosse 1892; Biol. Cent. Amer.:445. Pachychilus glaphyrus opiparis Fischer & Crosse 1892; Miss. Amer.:445.Type Localities. Melania opilaris: Dolores, Dept. Petn, Guatemala. Pachychilus glaphyrus var : Dolores, Dept. Petn, Guatemala. Distribution.Known only from the type locality. Pachychilus (Glyptomelania ) lacustris lacustris (Morelet 1849)Melania lacustris Morelet 1849; Test. Noviss. I:25.Brot 1874; in Martini & Chemnitz, Syst. Conch. Cab. (Melaniaceen):25. Pachychilus lacustris (Morelet). Fischer & Crosse 1892; Miss. Sci. 1899; Biol. Cent. Amer.:447. Pachychilus glaphyrus lacustris (Morelet). Goodrich & Van der Schalie 1937; Misc. Pub. Mus. Zool. Univ. Mich. (34):40.Type Locality.Lake near Izabal, Guatemala. Distribution.GUATEMALA, Dept. Guatemala: Lago de Amatitln (Hinkley 1920). Dept. Izabal: Jocolo (Hinkley); Lago de Izabal (Von Martens 1899). Pachychilus (Glyptomelania ) lacustris amphibolus Fischer & Crosse 1892Pachychila lacustris var. amphibolus Fischer & Crosse 1892; Miss. Amer.:447. Pachychilus lacustris var. pumila Fischer & Crosse 1892; Miss. Sci. Pachychilus lacustris var. pumilus Fischer & Crosse. Von Martens 1899; Biol. Cent. Amer.:448. Pachychilus lacustris var. major Fischer & Crosse 1892; Miss. Sci. Pachychilus lacustris var. terebralis Fischer & Crosse 1892; Miss. Amer.:448. Pachychilus lacustris var. eliminatus Fischer & Crosse 1892; Miss. Sci. Mex. II:360.Von Martens 1899; Biol. Cent. Amer.:448. Pachychilus lacustris var. conradi Von Martens 1899; Biol. Cent. Pachychlus glaphyrus lacustris (Morelet). Goodrich & Van der Schalie 1937; Misc. Pub. Mus. Zool. Univ. Mich. (34):40.Type Localities.All of these varieties were described from Lago de Izabal, Dept. Izabal, Guatemala. Distribution.GUATEMALA, Dept. Izabal: Lago de Izabal. Dept. Petn: Arroyo Subn; Rio de la Pasion (Goodrich & Van der Schalie 1937).

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 39 Pachychilus (Glyptomelania ) largillierti largillierti (Philippi 1843)Melania largillierti in Martini 1a-c. Pachychilus largillierti (Philippi).Fischer & Crosse 1892; Miss. der Schalie 1937:40. Melania intermedia Von dem Busch, in Melania rusticula Von dem Busch 1858:36. Melania rubicunda 206. Tristram 1863:413. Melania salwini Tristram. Brot 1874; in Martini & Chemnitz, Syst. Conch. Cab. (Melaniaceen):27 (amended name). Pachychilus subexaratus Crosse & Fischer 1891:219.Fischer & Pachychilus largillierti var. nodulosus Von Martens 1899; Biol. Type Localities. Melania largillierti : not given. Melania intermedia: Lago de Nicaragua. Melania rusticula: not given. Melania rubicunda: not given. : Rio de La Pason, Dept. Alta Verapaz, Guatemala. Pachychilus subexaratus: a small stream near Lago de Izabal, Guatemala. Pachychilus largillierti var. nodulosus; Paso Antonio, Guatemala. Distribution.GUATEMALA, Dept. Escuintla: Mirandilla. Dept. Guatemala: Lago de Amatitlan; Rio Michatoya, issuing from the lake; Rio Maria Linda; (Morelet); Paso Antonio, near Torola. Dept. Izabal: Rio Cavech; Plantera (Hinkley 1920). Dept. Petn: Rio de la Pasion. Dept. Zacapa: Lagarto, near Zacapa. NICARAGUA: Lago de Nicaragua. EL SALVADOR, Dept. San Miguel: Joya. Pachychilus (Glyptomelania ) largillierti stolli Von Martens 1899Pachychilus largillierti var. stolli Von Martens 1899; Biol. Cent. Type Locality.Environs of Retalhuleu, Dept. Retalhuleu, Guatemala. Distribution.Known only from the type locality. Pachychilus (Glyptomelania ) obeliscus obeliscus (Reeve 1861)Melania obeliscus Pachychilus obeliscus (Reeve). Fischer & Crosse 1892; Miss. Sci. Mex. II:358.Von Martens 1899; Biol. Cent. Amer.:446.Type Locality.Honduras. Distribution.HONDURAS: unknown. Pachychilus (Glyptomelania ) obeliscus examartus Fischer & Crosse 1892Melania obeliscus var. Brot 1874; in Martini & Chemnitz, Syst. Fischer & Crosse 1892; Miss. Sci. Mex. II:358.Von Martens 1899; Biol. Cent. Amer.:447.Type Locality.Not given. Distribution.Unknown. Pachychilus (Glyptomelania ) obeliscus pyrgiscus Fischer & Crosse 1892Pachychilus obeliscus var. pyrgiscus Fischer & Crosse 1892; Miss. Type Locality.Not given. Distribution.GUATEMALA, Dept. Petn: Lago de Petn. Pachychilus (Glyptomelania ) potomarchus Pilsbry 1892Pachychilus potomarchus Pilsbry 1893; Proc. Acad. Nat. Sci. Phila. Pachychilus glaphyrus potomarchus Pilsbry. Von Martens 1899; Biol. Cent. Amer.:446.Type Locality.Tabasco, Mxico. Distribution.Known only from the type locality. Pachychilus (Glyptomelania ) subnodosus (Philippi 1847)Melania subnodosa Martini & Chemnitz, Syst. Conch. Cab. (Melaniaceen):29; pl. Pachychilus subnodosus (Philippi). Fischer & Crosse 1892; Miss. Sci. Mex. II:366.Von Martens 1899; Biol. Cent. Amer.:450; Type Locality.Not given. Distribution.NICARAGUA, Dept. Managua: Managua (Von Martens 1899). Subgenus Oxymelania Fischer & Crosse 1892Oxymelania Fischer & Crosse 1892; Miss. Sci. Mex. II:328, 366.Von Martens 1899; Biol. Cent. Amer.:461. Potamanax Pilsbry 1893; Proc. Acad. Nat. Sci. Phila. 44:304.Type Species. Oxymelania : Melania schiedeanus Philippi 1843. Potamanax: Pachychilus (Potamanax) Pilsbry 1893. Distribution.Streams from the Rio Guayalejo system in northeastern Mxico south to the Rio Usumacinta system in Guatemala. Taxonomy.Eighteen species and four subspecies are recognized. Pachychilus (Oxymelania ) apheles Thompson 1967Pachychilus apheles 1.Type Locality.A spring run 14 miles west northwest of Ciudad Valles, on the road to Rio Verde, San Lus Potos, Mxico. Holotype UF 19756. Distribution.Known only from the type locality. Pachychilus ( Oxymelania) atratus Pilsbry & Hinkley 1910Pachychilus atratus Pilsbry & Hinkley 1910; Proc. Acad. Nat. Sci.

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40 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Pachychilus atratus ganinas 19, 20.Pilsbry 1956, Proc. Acad. Nat. Sci. Phila. 108:36. Pachychilus atratus suprastriatus Pilsbry & Hinkley 1910:526.Pilsbry 1956, Proc. Acad. Nat. Sci. Phila. 108:36 (as multistriatus). Type Localities. Pachychilus atratus: Rio Tamasopo near Verastagu, San Lus Potos, Mxico; holotype and Pachychilus atratus ganinas: Rio Ganina [Gallina] 3 miles SW of San Dieguito, San Lus Potos; holotype ANSP 99577. Pachychilus atratus suprastriatus: Rio Valles at Mecos Falls, and some rapids 2 miles west of Mecos, San Lus Potos; holotype ANSP 99575. Distribution.SAN LUS POTOS: known from the the type localities cited above. Pachychilus (Oxymelania ) corpulentus Thompson 1967Pachychilus corpulentus Thompson 1967; Nautilus 81:28; pl. 28, Type Locality.Nacimiento de Rio Mante, about 5 miles west of Ciudad Mante, Tamaulipas, Mxico [Rio Guayalejo system]. Holotype UF 19754. Distribution.Known only from the type locality. Pachychilus (Oxymelania ) graphium (Morelet 1849)Melania graphium Morelet 1849; Test. Noviss. I:26.Reeve 1861; Pachycheilus graphium (Morelet). H. & A. Adams 1854:298. Pachychilus graphium (Morelet). Fischer & Crosse 1892; Miss. Cent. Amer.:461.Von Martens 1901; Biol. Cent. Amer.:646.Goodrich & Van der Schalie 1937; Misc. Pub. Mus. Zool. Univ. Pachychilus cumingi Lea 1850. Proc. Zool. Soc. Lond. 18:179. Brot 1862; Matriaux fam. Mlaniens, 1:43.Brot 1874; in Martini & Chemnitz, Syst. Conch. Cab. Melania gracilis Tristram 1863; Proc. Zool. Soc. Lond. 31:413. Pachychilus tristrami Fischer & Crosse 1892; Miss. Sci. Mex. II:369. Pachychilus graphium var. reducta Fischer & Crosse 1892; Miss. Sci. Mex. II:333. Pachychilus graphium var. transcendens Fischer & Crosse 1892; Type Localities. Melania graphium: streams of Verapaz. Pachychilus cumingi: Cobn, in large rivers. : not given. Melania gracilis: Lago de Petn. Pachychilus tristrami: Lago de Petn. Pachychilus graphium var. reducta: Verapaz. Pachychilus graphium var. transcendens Uspatan, Dept. El Quich, Guatemala. Distribution.GUATEMALA, Dept. Alta Verapaz: on road between Cobn and Cham (Hinkley 1920). Dept. Izabal: Esmeralda (Hinkley 1920). Dept. El Quich. Dept. Petn: Lago de Petn. YUCATN: (Von Martens 1901). Pachychilus (Oxymelania ) humerosus Pilsbry & Hinkley 1910Pachychilus humerosus Pilsbry & Hinkley 1910; Proc. Acad. Nat. Sci. Phila. 61:526Type Locality.Rio Tamasopo near Verastagua, above and below the Natural Bridge, San Lus Potos, Mxico. Holotype and paratype ANSP 99579. Distribution.Known only from the type locality. Pachychilus (Oxymelania ) moctezumensis Pilsbry & Hinkley 1910Pachychilus moctezumensis Pilsbry & Hinkley 1910; Proc. Acad. Type Locality.Rio Moctezuma at the ford south of Distribution.Known only from the type locality. Pachychilus (Oxymelania ) monachus Pilsbry & Hinkley 1910Pachychilus monachus Pilsbry & Hinkley 1910; Proc. Acad. Nat. Type Locality.Rio Coy [Choy?], 3 miles south of Los paratypes ANSP 99581. Distribution.Known only from the type locality. Pachychilus (Oxymelania ) pleurostriatus pleurostriatus (Say 1831)Melania pleurostriata Say 1831; New Harmony Disseminator of Useful Knowledge, December 1831.Brot 1874; in Martini & Chemnitz, Syst. Conch. Cab. (Melaniaceen):44. Pachychilus pleurostriatus (Say). Fischer & Crosse 1892; Miss. Sci. Mex. II:370.Von Martens 1899; Biol. Cent. Amer.:449; pl. 26, Melania rubida Lea 1856; Proc. Acad. Nat. Sci. Phila.:145.Lea Martini & Chemnitz, Syst. Conch. Cab. (Melaniaceen):341; pl. Type Localities. Melania pleurostriata: not given. Melania rubida : Mxico. Distribution.SAN LUS POTOS: Rio Coy. A record from Laguna de Chapala, Jalisco is discounted here (Von Martens 1899). Pachychilus (Oxymelania ) pleurostriatus longus Pilsbry & Hinkley 1910Pachychilus pleurostriatus longus Pilsbry & Hinkley 1910; Proc. Proc. Acad. Nat. Sci. Phila. 108:33.Type Locality.Rio Ganina [Gallina], three miles southwest of San Dieguito, San Lus Potos, Mxico.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 41 Distribution.Known only from the type locality. Pachychilus (Oxymelania ) pleurostriatus tamasopensis Pilsbry & Hinkley 1910Pachychilus pleurostriatus tamasopensis Pilsbry & Hinkley 1910; 1956, Proc. Acad. Nat. Sci. Phila. 108:33.Type Locality.A small stream north of the mill south of the Tamasopo Sugar Co., San Lus Potos, Mxico. Holotype Distribution.Known only from the type locality. Pachychilus (Oxymelania ) pleaurotoma Pilsbry & Hinkley 1910Pachychilus pleaurotoma Pilsbry & Hinkley 1910; Proc. Acad. Nat. Acad. Nat. Sci. Phila. 108:37. to the Rio Moctezuma, San Lus Potos, Mxico. Holotype Distribution.SAN LUS POTOS: small streams near Pachychilus (Oxymelania ) schiedeanus schiedeanus (Philippi 1843)Melania schiedeanus Conch. Icon. (Melania in Martini & Chemnitz, Syst. Conch. Pachychilus schiedeanus (Philippi). Fischer & Crosse 1892; Miss. Cent. Amer.:462.Type Locality.Small brooks and ditches around the City of Mxico. This occurrence has not been corroborated by subsequent investigations. Distribution.VERACRUZ: Misantla; Rio de Misantla; Pachychilus (Oxymelania ) schiedeanus strebelianus Fischer & Crosse 1892Melania saussurei (Brot?). Strebel 1873; Beitrag. Mex. Landund Sssw.-Conch. I:36; 43, 43a. Pachychilus schiedeanus var. strebelianus Fischer & Crosse 1892; Miss. Sci. Mex. II:396. Pachychilus schiedeanus var. strebelianus Fischer & Crosse. Von Martens 1899; Biol. Cent. Amer.: 462.Type Locality.Streams of Palpoala, close to Misantla, Veracruz; Mxico. Distribution.VERACRUZ: known only from the vicinity of the type locality. Pachychilus (Oxymelania ) saussurei ( Brot 1874)Melania saussurei Brot; 1860; in 11.Brot 1874; in Martini & Chemnitz, Syst. Conch. Cab. Pachychilus saussurei (Brot). Fischer & Crosse 1892; Miss. Sci. Mex. II:368.Von Martens 1899; Biol. Cent. Amer.:462.Type Locality.Swamps in the woods bordering the Rio Grande, between Tampico and Mxico. This Rio Grande runs into the Lagoon of Meztitln. The road from Tampico to Mxico City crosses the river between Zacualtipn and Atotonilco, Hidalgo (Von Martens 1899). Distribution.Known only from the type locality. Apparently this species is extinct. Attempts by the author to locate populations in 2000 were unsuccessful. Pachychilus ( Oxymelania) suturalis Pilsbry & Hinkley 1910Pachychilus suturalis Pilsbry & Hinkley 1910; Proc. Acad. Nat. Sci. Nat. Sci. Phila. 108:37. 99585. Distribution.SAN LUS POTOS: streams in the Huichihuayn; Tamazunchale (Pilsbry 1956). Pachychilus (Oxymelania ) tristis Pilsbry & Hinkley 1910Pachychilus tristis Pilsbry & Hinkley 1910; Proc. Acad. Nat. Sci. Nat. Sci. Phila. 108:33.Type Locality.Rio Tamasopo at the natural bridge, near Verastagu, San Lus Potos, Mxico. Holotype and 3 Distribution.SAN LUS POTOS: known only from the immediate vicinity of the type locality. Pachychilus (Oxymelania ) Hinkley 1907 1, 5.Pilsbry & Hinkley 1910; Proc. Acad. Nat. Sci. Phila. 61:528.Pilsbry 1956; Proc. Acad. Nat. Sci. Phila. 108:37.Type Locality.Rio Valles, Valles, San Lus Potos, Mxico. Holotype ANSP 96592. Distribution.TAMAULIPAS, Rio Guayalejo system: Rio Frio, Gomez Farias; Rio Temes, below the bridge at Magiscatzin (Pilsbry 1956). SAN LUS POTOS, Rio Panuco Rio Panuco, Pujal (Pilsbry & Hinkley 1910). Pachychilus (Oxymelania )Pilsbry & Hinkley 1910 8, 9. Pilsbry & Hinkley 1910; Proc. Acad. Nat. Sci. Phila. 61:528.Type Locality.Chaimai Creek, about halfway between Valles and Pujal, San Lus Potos, Mxico. Holotype and Distribution.SAN LUS POTOS: in addition to the type locality this subspecies is known from the Rio Casas Viejas, coming from the north of Valles and emptying into the

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42 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Rio Valles some miles east of Mecos Falls (Pilsbry & Hinkley 1910). Subgenus Potamanax Pilsbry 1892Potamanax Pilsbry 1892:431.Pilsbry 1893b:63. Pachychilus (Oxymelania) Fischer & Crosse, in part. Morrison 1954; Proc. U. S. Nat. Mus. 103:365.Type Species.Potamanax rovirosai Pilsbry 1892. Distribution.Three species are recognized from the Rio Usumacinta system of Guatemala and adjacent Mxico. Pachychilus (Potamanax) pasionensis Pilsbry 1956Pachychilus pilsbryi (Von Martens). Goodrich & Van der Schalie 1937; Misc. Pub. Univ. Mich. Mus. Zool., (34):42. Pachychilus pasionensis Pilsbry 1956; Proc. Acad. Nat. Sci. Phila. Type Locality.A small arroyo tributary to the Rio Holotype UMMZ 65510. Distribution.GUATEMALA: known only from the (Goodrich & Van der Schalie 1937). Pachychilus (Potamanax) pilsbryi Von Martens 1899.P. [otamanax] Pilsbry 1893; Proc. Acad. Nat. Sci. Phila. Pachychilus (Potamanax) (Pilsbry). Fischer & Crosse Pachychilus pilsbryi Von Martens 1899; Biol. Cent. Amer.:463; pl. P. [Potamanax] Pilsbry 1892, non Pachychilus glaphyrus var. Pilsbry 1892.Von Type Locality.Sierra Poan, Tabasco, Mxico. Cotypes (2) ANSP 63386. Distribution.TABASCO: Rio Usumacinta, Santa Gertruda (Von Martens 1899); Santa Gertrudis, Rio Usumacinta (Von Martens 1901). Remarks.The name pilsbryi was proposed by Von Martens, because he considered Potamanax not to be distinct from Pachychilus, subgen. Oxymelania. Thus, Potamanax Pilsbry 1892, p. 341 is a primary homonym of Pilsbry 1892, p. 153. However, Pilsbry (1893) compared Potamanax to the Cuban species (Orbigny) (Thiaridae) and not to Pachychilus remain to be resolved. Pilsbry did not address these questions in 1956 (Proc. Acad. Nat. Sci. Phila. 109). Pachychilus (Potamanax) sargi (Crosse & Fischer 1875)Melania sargi Crosse & Fischer 1875; Jour. de Conchyl. 23:225, in Martini & Chemnitz, Syst. Conch. Pachychilus sargi (Crosse & Fischer). Fischer & Crosse 1892; Miss. Cent. Amer.:463.Type Locality.Guatemala. Distribution.GUATEMALA, Dept. Alta Verapaz: Family THIARIDAE Gill 1871 Genus Aylacostoma Spix 1827Aylacostoma Braziliam, Munich:15; pl. 8.Morrison 1954; Proc. U. S. Nat. Mus. 103:375, 376.Type Species.Melania glabra Spix 1827. Distribution.Central America south to southern Brazil. Taxonomy.The genus contains four subgenera, two of which occur in Central America (Morrison 1954). Subgenus Aylacostoma Spix 1827 Distribution.Guatemala, Colombia and Venezuela south to southeastern Brazil. Taxonomy.One species occurs in Guatemala. Numerous species are found in South America. Aylacostoma (Aylacostoma ) rubiginosum (Morelet 1849)Melania rubiginosa Morelet 1849; Testacea Noviss. I:25. Hemisinus rubiginosus (Morelet). H. & A. Adams 1854; Gen. Recent in Martini & Chemnitz, Syst. Conch. Cab. (Melanidae):394; pl. Semisinus rubiginosus (Morelet). Fischer & Crosse 1892:318; pl. Aylacostoma rubiginosum (Morelet). Morrison 1954; Proc. U. S. Nat. Mus. 103:377. Melania petenensis Tristram 1862; Proceedings of the Zoological Society:441. Hemisinus petenensis (Tristram). Brot 1862; Materiaux fam. Melaniens, 2:51. Hemisinus zoster Brot 1862; Materiaux fam. Melaniens, 2:62.Type Localities. Melania ruiginosa: Lago de Izabal, [Dept. Izabal], Guatemala. Melania petenensis : Lake of Petn, Guatemala. Hemisenus zoster: not given. Distribution.GUATEMALA, Dept. Izabal: Jocala, on the north shore of Lago de Izabal (Hinkley 1920). Dept. Petn: Lake of Petn (Von Martens 1899). Subgenus Hemisinus Swainson 1840Hemisinus Swainson 1840; Treatise on Malacology:200, 341.Brot 1874; in Martini & Chemnitz, Syst. Conch. Cab., (Melanidae):369.Morrison 1954; Proc. U. S. Nat. Mus. 103:367 Gomez, Strong & Glaubrecht, 2011; Malacologia 53:229-250. Semisinus P. Fischer 1885; Manuel de Conchyl., 701.Fischer & Crosse 1892:313.Von Martens 1899; Biol. Cent. Amer.:464.Type Species.Melania lineolata Gray 1828. Distribution.Jamaica, Central America south to Ecuador and to Peru. Aylacostoma (Hemisinus) maculatum ( Lea 1832)Malania maculata Lea 1832; Trans. Amer. Philos. Soc. 5:194; pl.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 43Semisinus maculatus (Lea).Von Martens 1901; Biol. Cent. Melania osculatii Ant. Villa 1855; Notizie interno al genera Melania in Atti dell Accad. Fisico-Medio-statistica di Milano, 2:8. Hemisinus osculatii in Martini & Chemnitz, Syst. Conch. Cab., (Melanidae):379; Type Localities. Melania maculate : Peru. Melania osculatii : Quito, Ecuador. Shei and the Rio Tararia (Von Martens 1901). COLOMBIA. ECUADOR. Genus Melanoides Olivier 1804 Type Species.Turba tuberculata O. F. Mller, 1774. Distribution.Africa east through southern and eastern Taxonomy.Numerous species have been described. The number of valid taxa remains undetermined. Melanoides tuberculata (O. F. Mller 1774) Islands. Introduced widely into Neotropical and Neotemparate regions. The species is spreading rapidly throughout Mxico and Central America, and by now it is assumed to be established in nearly all drainage systems in these countries. Taxonomy.Brandt (1974:164) listed 37 synonyms for Melanoides tuberculatus. Melanoides are parthenogenetic uni-parental species, in which each individual is reproductively independent from all other individuals. Biparental species evolutionary concepts do not apply to such organisms. Melanoides turriculus (Lea 1850) is a name that is commonly encountered in the Neotropical literature. It is generally regarded as a synonym of M. tuberculatus. Superfamily LITTORINOIDEA Children 1834 Family ANNULARIIDAE Henderson & Bartsch 1920 Distribution.West Indies, south Florida, Mxico south to Panam, Colombia and Venezuela. Taxonomy.Numerous genera, subgenera and species. and their sister group, the Eur-Asian-Ethiopean Pomatiasidae, has been based on structure of the operculum and upon breathing devises associated with the aperture. Henderson of the Annulariidae based on these features. The opercula of many of the Mxico-Central American species were not new material I have examined. Solem (1961) adapted subgeneric terminology for the mainland species proposed by H. B. Baker (1928). Watters (2006) reviewed the family and by previous workers. He de-emphasisised the importance that other authors placed upon opercular and apertural features. because they do not adequately discriminate between some genera, even in different subfamilies. It is beyond the scope of this study to try to address the generic nomenclature proposed by Watters (2006), because most of the names apply to West Indian taxa and have little bearing on the fauna of Mxico utilized by Solem (1961), because it depicts phylogenetic and biogeographic relationships more clearly than later attempts to address these problems. Watters (2006) provided a wealth of information on the species. He listed over 1500 species and subspecies names. Henderson & Bartsch (1920), and it was a monumental achievement by itself. Fifteen species are known from Mxico and Central America. Subfamily CHONDROPOMATINAE Henderson & Bartsch 1920 Genus Chondropoma Pfeiffer 1847Chondropoma Pfeiffer 1847; Zeit. Malak. 4:109.Henderson 1914; Nautilus 27:138.Henderson & Bartsch 1920; Proc. U. S. Nat. Mus. 58.Torre & Bartsch 1942; Proc. U. S. Nat. Mus. 89:349.Bartsch 1946; Bull. U. S. Nat. Mus. 192:5.Solem 1961; Arch. Moll. 90:204.Watters 2006:25. Gouldipoma Watters 2006:71. Type Species.Chondropoma: Cyclostoma sagra Orbigny 1842; by original designation. Gouldipoma: Chondropoma callipeplum Solem 1961a, by original designation. Distribution.Widely distributed in the West Indies and the circum-Caribbean region of the mainland from Florida and Mxico south tto Venezuela. Taxonomy.Many species and subspecies. Watters (2006) distinguished Gouldipoma from Chondropoma on the basis of shell sculpture. He transfered Parachondria Dall 1905 from the Subfamily Rhytodopominae to subgeneric status with Chondropoma of both and included many disparate species. He synonymized ten other subgenera and genera with Parachondria, further complicating a taxonomic dilemma. For purposes of this study I follow Solem (1961). The species tentatively are referred to the subgenus Chondropomium. Five species of Chondropoma are recognized from Mxico and Central America. Subgenus Chondropomium Henderson & Bartsch 1920Chondropomium Henderson & Bartsch 1920; Proc. U. S. Nat. Mus. 58:60.Watters 2006:62.Type Species.Chondropoma weinlandi Pfeiffer 1862. Distribution.Hispaniola, Jamaica, Mxico, Central America and Venezuela. Taxonomy.About 20 species are included in the subgenus. Chondropoma (Chondropomium) callipeplum Solem 1961Chondropoma callipeplum Solem 1961; Arch. Moll. 90:207; pl. 11,

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44 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Gouldipoma callipeplum (Solem). Watters 2006:71 183.Type Locality.Wani, Nicaragua. Holotype USNM 186093. Distribution.Known only from the type locality. Chondropoma ( Chondropomium) Pfeiffer 1856 Pfeiffer 1856; Proc. Zool. Soc. Lond. Biol. Cent. Amer.:15, 17; Solem 1961; Arch. Moll. 90:204 Chondropoma (Chondropomium) Pfeiffer. H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):50. (Pfeiffer). Watters 2006:73, 216. Syntypes in the BMNH, ZMB and the MNHN. Distribution.VERACRUZ: Central Veracruz, near Chondropoma (Chondropomium) gruneri (Pfeiffer 1846)Cyclostoma gruneri Pfeiffer 1846; Zeit. Malak. 3:47.Pfeiffer 1847: Cistula gruneri (Pfeiffer). Gray 1850; Cyclophoridae:59.Pfeiffer 1851; Zeit. Malak. 8:170. Chondropoma gruneri (Pfeiffer). Reeve 1863; Conch. Icon., Chondropoma Ctenopoma gruneri (Pfeiffer). Fischer & Crosse 1890:188, 191. Choanopoma (Choanopomops) gruneri (Pfeiffer). Solem 1961; Arch. Moll. 90:203. Halotudora gruneri (Pfeiffer). Watters 2006:73, 283.Type Locality.Honduras; here restricted to Belize. Distribution.BELIZE: numerous specimen samples in the Florida Museum of Natural History. Chondropoma (Chondropomium) kuesteri (Pfeiffer 1851)Cyclostoma ksteri Pfeiffer. Pfeiffer 1851; Proc. Zool. Soc. Lond. Cistula ksteri Pfeiffer 1852; Monogr. pneumo. vivent., 1:265. Chondropoma ksteri (Pfeiffer). Reeve 1863; Conch. Icon., Chondropoma Ctenopoma ksteri (Pfeiffer). Fischer & Crosse 1890:194; pl. Choanopoma (Choanopomops) ksteri (Pfeiffer). Solem 1961; Arch. Moll. 90:203. Halotudora ksteri (Pfeiffer). Watters 2006:73, 321.Type Locality.Honduras; here restricted to Belize. Distribution.BELIZE: numerous specimen samples in the Florida Museum of Natural History. Chondropoma (Chondropomium) rubicundum (Morelet 1849)Cyclostoma rubicundum Morelet 1849; Test. Noviss. I:22.Pfeiffer 1854; in Martini & Chemnitz Syst. Conch. Cab., Cyclostoma Chondropoma rubicundum (Morelet). Pfeiffer 1851; Zeit. Malak. 8:173. Reeve 1863; Conch. Icon., Chondropoma Von Martens 1890:16.Hinkley 1920; Nautilus 34:42, 44, 52.Goodrich & Van der Schalie 1937; Misc. Pub. Mus. Zool. Univ. Mich. (34):31.Bequaert 1957:212.Solem 1961; Arch. (shell). Chondropoma (Chondropomium) rubicundum (Morelet). Henderson & Bartsch 1920; Proc. U.S. Nat. Mus. 58:60.H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):50. (Pfeiffer). Watters 2006:72, 448. Cyclostoma acerbulum Morelet 1851; Test. Noviss. II:19. Chondropoma acerbulum (Morelet). Fischer & Crosse 190:210 Cistula subangulatum Von Martens 1886; Sitz.-berg. Gesell. naturf. Freud. berlin, (10):162. Chondropoma subangulatum (Von Martens). Von Martens 1890; Type Localities. Cyclostoma rubicundum: Petn et Verapaz, Guatemala. Cyclostoma acerbulum: Verapaz, Guatemala. Chondropoma subangulatum: Teleman, Guatemala. Distribution.BELIZE: Cayo Dist. Succolz (Solem 1961a). HONDURAS: extreme northwest of the country in the Dept. Corts (UF). GUATEMALA: numerous localities in the Depts. Petn, Alta Verapaz, and Baja Verapaz (Goodrich & van der Schalie 1937; Hinkley 1920; Solem 1961). CHIAPAS: Laguna Ocotal, 950 m alt. (Bequaert 1957). TABASCO (Solem 1961). Subfamily ANNULARIINAE Henderson & Bartsch 1920 Taxonomy.Two genera occur in the Mxico-Central American region. Genus Annularia Schumacher 1817Annularia Schumacher 1817:60, 196.Henderson & Bartsch 1920; Proc. U. S. Nat. Mus. 58:71, 72.Watters 2006:85 (in part). Type Species.Turbo lincina Linnaeus 1758. Distribution.The Greater Antilles, Mxico. Taxonomy.A single species occurs in the study area. Numerous species occur in the West Indies. Annularia sumichrasti Crosse & Fischer 1874Choanopoma sumichrasti Crosse & Fischer 1874; Jour. de Conchyl. (shell).Solem 1961; Arch. Moll. 90:196 (in part).Halotudora sumichrasti (Crosse & Fischer). Watters 2006: 73.Type Locality.Isthmus of Tehuantepac, [Oaxaca], Mxico. Distribution.OAXACA: reported only from the type locality. Common in a small area on the south side of the Isthmus of Tehuantepec (UF). Taxonomy.Solem (1961) confused this species with Choanopoma chiapensis Crosse & Fischer 1877. Opercular differences indicate that they belong in different genera. Genus Tudora Gray 1850Tudora Gray 1850:48.Henderson & Bartsch 1920; Proc. U. S. Nat. Mus. 58:75, 76.Type Species.Cyclostoma similis Gray 1843 [= Cyclostoma megacheilos Potiez & Michaud 1838].

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 45 Distribution.Jamaica, Caribbean coastal areas of the mainland; Dutch West Islands. Taxonomy.Henderson and Bartsch (1920) listed seven subgenera. H. B. Baker proposed the subgenus Tudorata to include mainland species found in Central America and South America. Subgenus Tudorata H. B. Baker 1924Tudorata H. B. Baker 1924; Occ. Pap. Mus. Zool. Univ. Mich. (152):152:44.Solem 1960; Arch. Moll. 90:428. Tudora Gray 1850: Watters 2006:77.Type Species.Tudora muskusi H. B. Baker 1924. Distribution.Coastal areas of the mainland from Venezuela north to Panam; Dutch West Indies. Taxonomy.Six species are recognized. One occurs in Central America. Tudora (Tudorata) thomasi Solem 1961Tudora (Tudorata) thomasi Solem 1961; Arch. Moll. 90:208; pl. 11, Gouldipoma thomasi (Solem). Watters 2006:72, 508.Type Locality.Natural Bridge of the Rio Puerto near the shore of Madden Lake, Canal Zone, Panam. Holotype FMNH 73390. Distribution.Known only from the type locality. Subfamily RHYTIDOPOMINAE Henderson & Bartsch 1920 Genus Choanopoma Pfeiffer 1847Choanopoma Pfeiffer 1847; Zeit. Malak. 3:47.H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):47. Type Species.Turbo lincina Linnaeus 1758. Distribution.West Indies, Caribbean coastal areas of the mainland. Taxonomy.Much controversy centers about the use of the name Choanopoma. Henderson & Bartsch (1920) and Waters (2006) treated Choanopoma as a synonym of Annularia. H. B. Baker (1928) recognized Choanopoma as a valid genus. Solem (1961) followed H. B. Baker but pointed out the unstable taxonomy surrounding the name Choanopoma. A single subgenus, Choanopomops, is recognized in Mxico and Central America. Subgenus Choanopomops H. B. Baker 1928Choanopomops H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):48.Solem 1961; Arch. Moll. 90:194.Watters 2006:58 (in part). Halotudora Watters 2006:73.Type Species.Chondropomops: Cyclostoma largillieti Pfeiffer 1846. Halotudora: Cyclostoma radiosum Morelet 1849. Distribution.Caribbean coastal areas of the mainland, Jamaica and Cuba (H. B. Baker 1928). Taxonomy.Nine species are recognized in Mxico and Central America. Additional species occur in northern South America and Jamaica. Choanopoma (Choanopomops) andrewsae (Ancey 1886)Cyclostoma andrewsae Ancey 1886; Ann. de malac. 2:251. Colobostylus andrewsae (Ancey). Fischer & Crosse 1890:218. Choanopoma andrewsae (Ancey). Von Martens 1890; Biol. Cent. Philos. Soc. 77:256.Richards 1938; Proc. Amer. Philos. Soc. 79:176, 178.Thompson 1967; Bull. Fla. St. Mus. 11:222, Chondropoma andrewsae (Ancey).Henderson & Bartsch 1920; Proc. U. S. Nat. Mus. 58:60. Choanopoma (Choanopomops) andrewsae andrewsae (Ancey). Choanopoma cozumelense Richards 1937; Proc. Amer. Philos. Soc. Choanopoma andrewsae roatanense Richards 1938; Proc. Amer. Annularia (Choanopomops) cozumelensis (Richards). Rehder 1966; Proc. Biol. Soc. Wash. 79:279. Tudorisca andrewsae andrewsae (Ancey). Watters 2006:79, 140. Tudorisca andrewsae roatanense (Richards). Watters 2006:79, 140. Tudorisca cozumelense (Richards). Watters 2006:79, 218.Type Localities. Cyclostoma andrewsae: Isla de tila, Dept. Islas de la Baha, Honduras. Choanopoma cozumelense: San Gerbacio, Isla Cozumel, Quintana Roo, Mxico; holotype ANSP 167749. Choanopoma andrewsae roatanense: West End, Isla de Roatn, Honduras; holotype ANSP 170018. Distribution.HONDURAS: Dept. Islas de la Baha: Isla de Roatn; Isla de tila. CAMPECHE: 5.1 mi. NNW of Dzilbachn; 3.6 mi. S of Hopelchn; 3.4 mi. S of Cayl. QUINTANA ROO: Tulm (Rehder 1966); 4.0 mi. E of Xpujil, Campeche; 7.1 mi. NNW Xiatil; 2.3 mi. SSE Xiatil; San Sebastian, Isla Cozumel (Thompson 1967c). YUCATN: 0.8 mi. NE Becanchn; 10.0 mi. NE Becanchn; Chichen Itza (Thompson 1967c). Choanopoma (Choanopomops) chiapasense Crosse & Fischer 1877Choanopoma chiapasense Crosse & Fischer 1877; Jour. de Conchyl. 8a (shell).Bequaert & Clench 1931; Occ. Pap. Boston Soc. Nat. Hist. 5:426.Bequaert & Clench 1933; Pub. Carnegie Inst. Wash. (431):542.Bequaert 1957:225. Tudora (Tudorisca) chiapensis [sic] (Crosse & Fischer 1877). Henderson & Bartsch 1920; Proc. U. S. Nat. Mus. 58:76. Choanopoma (Choanopomops) sumichrasti var. chiapasense Choanopoma (Choanopomops) chiapensis [sic] Crosse & Fischer. Halotudora chiapasense (Crosse & Fischer). Watters 2006:73, 201.Type Locality.Chiapas, Mxico. Distribution.CHIAPAS: Santo Domingo; Chivela; Lagunas (Solem 1961a); 8 km. E of Chiapa de Corzo, 3100 ft. alt; 14.9 mi. E of Chiapa de Corzo, 4400 ft. alt.; 8 mi. km of Tuxtla Gutierrez, 380 ft. alt. (Thompson 1966).

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46 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Choanopoma (Choanopomops) gaigei Bequaert & Clench 1931Choanopoma gaigei Bequaert & Clench 1931; Occ. Pap. Boston Soc. Nat. Hist. 5:425.Bequeart & Clench 1933; Pub. 81 (shell).Bequaert & Clench 1936; Pub. Carnegie Inst. Wash. (457):72.Goodrich & van der Schalie 1937; Misc. Pub. Mus. Zool. Univ. Mich. (34):12, 15, 22, 32.Harry 1950; Occ. Pap. Mus. Zool. Univ. Mich. (524):26.Solem 1961; Arch. Branson & McCoy 1963; Nautilus 76:104.Thompson 1966; Nautilus 80:27.Thompson 1967; Bull. Fla. St. Mus. 11:225, 227. Choanopoma gagei [sic] Bequaert & Clench.Richards 1937; Proc. Amer. Philos. Soc. 77:257. Halotudora gaigei (Bequaert & Clench). Watters 2006:73, 269.Type Locality.Chichen Itza, Yucatn, Mxico. Holotype UMMZ 49190. Distribution.GUATEMALA, Dept. Petn: 1 mi. NW of Paso Caballo; 4 mi. N of Paso Caballo (Goodrich & van der Schalie 1937). CAMPECHE: numerous localities (Thompson 1967c). QUINTANA ROO: 4.0 mi. E of Xpujil (Campeche); 7.1 mi. NNE of Polyuc; 2.3 mi. SSE Polyuc (Thompson 1966). YUCATN: vicinity of Chichen Itza (Bequaert & Clench 1931, 1933, 1936; Harry 1950); 0.8 mi. NE Becanchen (Thompson 1966). Choanopoma (Choanopomops) largillierti (Pfeiffer 1846)Cyclostoma largillierti Pfeiffer 1846; Zeit. Malak. 3:46.Pfeiffer Cyclostoma grataloupi Pfeiffer 1853 [for 1851]; Proc. Zool. Soc. Ctenopoma largillierti (Pfeiffer). Fischer & Crosse 1890:196; Ctenopoma grateloupi (Pfeiffer). Fischer & Crosse 1890:199; Chondropoma largillierti (Pfeiffer). Von Martens 1890; Biol. Cent. Amer.:19. Licina (Choanopomops) largillierti (Pfeiffer). H. B. Baker 1928; Licina (Choanopomops) grateloupi (Pfeiffer). H. B. Baker 1928; (radula). Choanopoma largillierti (Pfeiffer). Bequaert & Clench 1933; Pub. Carnegie Inst. Wash. (431):540.Richards 1937; Proc. Amer. Philos. Soc. 77:257.Bequaert & Clench 1938; Pub. Carnegie Inst. Wash. (491):259.Harry 1950; Occ. Pap. Mus. Zool. 1963; Nautilus 76:104.Thompson 1966; Nautilus 80:27.Thompson 1967; Bull. Fla. St. Museum, 11:224.Watters 2006:58, 326. Choanopoma (Choanopomops) largillierti (Pfeiffer). Bequaert & Clench 1936; Pub. Carnegie Inst. Wash. (457):71.Haas & Solem 1960; Nautilus 73:130.Solem 1961; Arch. Moll. (sculpture). Choanopomops grateloupi (Pfeiffer). Watters 2006:58, 279. Choanopomops largillierti (Pfeiffer). Watters 2006:58, 326.Type Locality.Cyclostoma largillierti : Yucatn, Mxico. Cyclostoma grateloupi : Yucatn, Mxico. Distribution.CAMPECHE: numerous localities (Thompson 1967c). YUCATN: numerous localities (Bequaert & Clench 1938; Branson & McCoy 1963; Thompson 1966, 1967). QUINTANA ROO: Tulm (Rehder 1966); numerous localities (Thompson 1966). BELIZE: Chetumal (Solem 1961). Choanopoma ( Choanopomops) martensianum (Pilsbry 1900)Chondropoma martensianum Pilsbry 1900; Nautilus 13:140.4, 4a (shell).Thompson 1957; Nautilus 70:100.Solem 1961; (sculpture).Type Locality.Sierra Poan, Tabasco, Mxico. Holotype ANSP 63417. Distribution.TABASCO: known only from the type locality and from near-by areas near Teapa (Thompson 1957). Choanopoma (Choanopomops) osberti (Tristram 1861)Adamsilla osberti Tristram 1861; Proc. Zool. Soc. Lond. 29:232.Licina osberti (Tristram). H. B. Baker 1928; Occ. Pap. Mus. Zool. Univ. Mich. (193):48. Choanopoma osberti (Tristram). Von Martens 1890; Biol. Cent. Annularia rigidula (Morelet). Haas 1949; Nautilus 62:136. Parachondrops osberti (Tristram). Watters 2006:47, 386.Type Locality.Guatemala. Distribution.GUATEMALA, Dept. Sacatepquez: Sacatepquez (Solem 1961). Choanopoma (Choanopomops) radiosum (Morelet 1849)Cyclostoma radiosum Morelet 1849; Test. Noviss. I:22. Cistula radiosa.(Morelet). Pfeiffer 1851; Zeit. Malak. 8:169.Crosse & Fischer 1883:103.Hinkley 1920; Nautilus 34:44. Chondropoma radiosum (Morelet). Reeve 1963; Conch. Icon., Chonropoma Cent. Amer.:18.Pilsbry 1900; Nautilus 13:140. Ctenopoma sargi Crosse & Fischer 1883; Jour. de Conchyl. 31:103. Ctenopoma radiosum (Morelet). Fischer & Crosse 1890:188; Ctenopoma sargi Crosse & Fischer. Fischer & Crosse 1890:191 Choanopoma radiosum (Morelet). Goodrich & van der Schalie 1937; Misc. Pub. Mus. Zool. Univ. Mich. (34):20. 31.Thompson 1966; Nautilus 80:28. Licina radiosa (Morelet). Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):7. Choanopoma (Colobostylus) radiosum (Morelet). Haas & Solem Choanopoma (Choanopomops) radiosum (Morelet). Solem 1961; Chondropoma sargi (Crosse & Fischer). Von Martens 1890:18. Halotudora radiosua (Morelet). Watters 2006:73, 430.Type Locality.Cyclostoma radiosum: Petn, Guatemala. Cistula sargi: San Miguel de Tucuru, Dept. Alta

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 47 Verapaz, Guatemala (Solem 1961). Distribution.BELIZE: Punta Gorda; Gales Point; Bienque Viejo; Augustino (Solem 1961). GUATEMALA: widely distributed in the Depts. Petn, Alta Verapaz, and Izabal (Solem 1961). Choanopoma (Choanopomops) rigidulum (Morelet 1851)Cyclostoma rigidulum Morelet 1851; Test. Noviss. II:18. Adamsiella rigidula (Morelet). Fischer & Crosse 1890:176; pl. Choanopoma rigidulum (Morelet). Von Martens 1890; Biol. Cent. Choanopoma (Choanopoma) rigidulum (Morelet). Haas & Solem 1960; Nautilus 73:130. Choanopoma (Choanopomops) rigidulum (Morelet). Solem. 1961; (sculpture). Licina (Choanopomops) rigidulum (Morelet). H. B. Baker 1928; Occ. Pap. Mus. Zol. Univ. Mich. (193):48. Parachondrops rigidulus (Morelet). Watters 2006:47, 442.Type Locality.San Lus, Dept. Petn, Guatemala [Watters 2006]. Holotype BMNH 93.2.4.704. Distribution.GUATEMALA: reported only from the type locality. BELIZE: Cayo District: Rio Frio Cave, 2 mi. from Augustine (Haas & Solem 1960). Choanopoma (Choanopomops) terecostatum Thompson 1966Choanopoma terecostatum Thompson 1966; Nautilus 80:24; Halotudora terecostata (Thompson). Watters 1906:73, 506.Type Locality.Limestone hillside 15.8 miles [25 km] northwest of Ocozocoautla, Chiapas, 2700 ft. [830 m] alt. Holotype UMMZ 216554. Distribution.Known only from the type locality. Annulariidae of doubtful occurrence in Mxico and Central America Chondropoma turritum Pfeiffer 1853Chondropoma turritum Pfeiffer 1853 [for 1851]; Proc. Zool. Soc. Martens 1890; Biol. Cent. Amer.:19.Solem 1962; Arch. Gouldipoma turritum (Pfeiffer). Watters 2006:72, 527.Type Locality.Honduras. Distribution.Unknown. Remarks.Chondropoma turritum is very similar to Chondropoma canescens (Pfeiffer 1852) from Cuba. It may be a West Indian species incorrectly attributed to Honduras. Morelet 1851C Morelet 1851; Test. Noviss. II:19. (Morelet). Fischer & Crosse 1890:211 Torre & Bartsch 1938; Proc. U. S. Nat. Mus. 85:341.Solem 1961; Arch. Moll. 90:209.Watters 2006:534.Remarks.This species was erroniusly recorded by Morelet (1851) as coming from Palenque, Chiapas, Mxico. Isle of Pines, Cuba. Cistula pleurophorum Pfeiffer 1852Cistula pleurophorum Pfeiffer 1853 [for 1851]; Proc. Zool. Soc. Lond. 19:169.Fischer & Crosse 1890:218. Chondropoma pleurophorum (Pfeiffer). Von Martens 1890:16. Choanopoma (Choanopomops) pleurophorum (Pfeiffer). Solem Type Locality.Honduras. Distribution.Unknown. This species has not been found since its original description. It may be a West Indian species (Solem 1961). Cistula trochlearis Pfeiffer 1852Cistula trochlea Pheiffer 1851; Zeitsch. Malakozoologie 8 (11):171. (Non Benson 1851). Cistula trochlearis Pfeiffer 1852; Mono. Pneum. Viv.:275.Solem 1961; Arch. Moll. 90:2091.Type Locality.Guadeloupe. Distribution.Unknown. Solem (1961) and Watters (2006) regarded this specioes as a nomen dubium. Superfamily RISSOOIDEA Gray 1847 Family HYDROBIIDAE Troschel 1857 Taxonomy.This is a very large, cosmopolitan family of fresh and brackish water snails. The family includes several hundred genera. One hundred and thirteen (113) species occur in the study area. Subfamily COCHLIOPINAE Tryon 1866Cochliopinae Tryon 1866.Taylor 1966.Hershler & Thompson 1992. Littoridininae Thiele 1928.Taylor 1966 (in part). Mexithauminae Taylor 1966 (in part). Paludiscalinae Taylor 1966 (in part). Semiscalinae Giusti & Pezolli 1980.Type Genus.Cochliopa Stimpson 1865. Distribution.North America, throughout South America, the West Indies, central Africa and the Mediterranean region. Taxonomy.About thirty-nie genera are recognized in the study area. Genus Aroapyrgus Baker 1931Aroa Baker 1930; Occ. Pap. Mus. Zool. Univ. Mich. (210):33. (Type species by original designation: Baker 1930.) Non Walker 1855, Lepidoptera. Aroapyrgus, Baker 1931; Nautilus 44:124. Hershler & Thompson 1992; Malac. Rev. Suppl. 5:24. (Replacement name for Aroa H. B. Baker 1930.) Sioliella Haas 1949; Anal. Inst. Biol. Mex.,20:307 (type species by original designation: Sioliella effusa Haas 1949 (subjective synonym).Distribution.North and South America from San Lus Potos, Mxico, south to Colombia and east to Par State, Brazil. Taxonomy.Twenty-nine species are recognized

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48 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) (Hershler & Thompson 1992). Nineteen occur in the study area. Aroapyrgus alleei Morrison 1946Aroapyrgus alleei Morrison 1946; Smiths. Misc. Coll. 106:14; pl. 2, Suppl. 5:26.Type Locality.Allee Stream, Barrow Colorado Island, Gutun Lake, Canal Zone, Panam. Holotype USNM 542142. Distribution.Known only from the type locality. Aroapyrgus chagresensis Morrison 1946Aroapyrgus chagresensis Morrison 1946; Smiths. Misc. Coll. 1992; Malac. Rev. Suppl. 5:26.Type Locality.Ro Chagres, Gatuncilla, Panam. Holotype USNM 542144. Distribution.PANAM: recorded only from the Ro Chagres at the type locality and Madronal. Aroapyrgus cisterninus (Walker 1919)Amnicola cisternina Walker 1919; Occ. Pap. Mus. Zool. Univ. Mich. Aroapyrgus cisternina (Walker). Thompson 1968; Fla. Hydrob.:150.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:26Type Locality.Reservoir north of Guatemala City, Dept. Guatemala, Guatemala. Holotype UMMZ 47161. Distribution.Known only from the type locality. Aroapyrgus clenchi (Goodrich & Van der Schalie 1937)Somatogyrus clenchi Goodrich & Van der Schalie 1937; Misc. Pub. Aroapyrgus clenchi (Goodrich & Van der Schalie). Hershler & Thompson 1992; Malac. Rev. Suppl. 5:26. Petn, Guatemala. Holotype in the UMMZ. Distribution.GUATEMALA, Dept. Petn: Ro de La tributaries. Aroapyrgus conchensensis (Walker 1919)Amnicola conchensensis Walker 1919; Occ. Pap. Mus. Zool. Univ. Aroapyrgus conchenensis (Walker). Thompson 1968; Fla. Hydrob.:150.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:27.Type Locality.Conchens River, Guatemala. UMMZ 38508. Distribution.Known only from the type locality. Aroapyrgus costaricenis (Mrch 1860)Hydrobia costaricensis Mrch 1860:67.Frauenfeld 1864:594. Amnicola costaricensis (Mrch). Von Martens 1899; Biol. Cent. Aroapyrgus costaricensis (Mrch). Thompson 1968; Fla. Hydrob.:150.Malek, E. A., R. Brenes & G. Rojas 1975; Jour. parasit., 61:355.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:27.Type Locality.Ro Torres, Prov. San Jos, Costa Rica. Location of syntypes unknown. Distribution.COSTA RICA: Streams at intermediate elevations of San Jos, Cartago, and Puntarenas provinces. Aroapyrgus guatemalensis (Fischer & Crosse 1891)Amnicola guatemalensis Fischer & Crosse 1891; Miss. Sci. Mex. II: Amnicola hyalina (Fischer & Crosse) (in part). Von Martens 1899; Biol. Cent. Amer.:432. Aroapyrgus guatemalensis (Fischer & Crosse). Thompson 1968; Fla. Hydrob.:150.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:27.Type Locality.Lago de Amatitln, Dept. Guatemala, Guatemala. Distribution.Known only from the type locality. A record from Lago de Catemaco, VERACRUZ, Mxico Aroapyrgus hinkleyi (Walker 1919)Amnicola hinkleyi Walker 1919; Occ. Pap. Mus. Zool. Univ. Mich. Aroapyrgus hinkleyi (Walker). Thompson 1968; Fla. Hydrob.:27.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:27.Type Locality.Reservoir north of Guatemala City, Dept. Guatemala, Guatemala. Holotype UMMZ 47162. Distribution.Known only from the type locality. Aroapyrgus joseana Morrison 1946Aroapyrgus josana Morrison 1946; Smiths. Misc. Coll. 106:15; Rev. Suppl. 5:27.Type Locality.A northern tributary of the Ro Mata Puerco Estuary, Isla San Jos, Archipelago de Las Perlas, Panam. Holotype USNM 542147. Distribution.PANAM: known only from small streams on Isla San Jos. Aroapyrgus mexicanus (Pilsbry 1910)Somatogyrus mexicanus 3. Aroapyrgus mexicanus (Pislbry). Hershler & Thompson 1992; Malacological Rev., Suppl. 5:28.Type Locality.Ro Coy, on the road to Tampamoln, San Lus Potos, Mxico. Lectotype ANSP 99023 (Baker 1964). Distribution.Known only from the type locality. Aroapyrgus orizaben sis (Crosse & Fischer 1891)Amnicola orizabensis Crosse & Fischer 1891; Jour. de Conchyl. 39: 4b.Von Martens 1899; Biol. Cent. Amer.:432. Aroapyrgus orizabensis (Crosse & Fischer). Hershler & Thompson 1992; Malac. Rev. Suppl. 5:28.Type Loca lity.Near Orizaba, Veracruz State, Mxico. Distributio n.VERACRUZ: known only from the immediate vicinity of the type locality.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 49 Aroapyrgus panamensis (Tryon 1863)Amnicola panamensis Tryon 1963; Proc. Acad. Nat. Sci. Phila. Aroapyrgus panamensis (Tryon). Thompson 1968; Fla. Hydrob.:150.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:27.Type Locality.Panam. Holotype ANSP 58067. Distribution.Known only from the type locality. A record for Javal, Dept. Chontales, Nicaragua (Tate 1870:153) refers to A. tryoni Aroapyrgus panzosensis (Walker 1919)Amnicola panzosensis Walker 1919; Occ. Pap. Mus. Zool. Univ. Aroapyrgus panzosensis (Walker). Thompson 1968; Fla. Hydrob.:150.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:28.Type Locality.Panzos, Ro Polochc Valley, Dept. Alta Verapaz, Guatemala. Holotype UMMZ 37389. Distribution.Known only from the type locality. Aroapyrgus pasionensis (Goodrich & Van der Schalie 1937)Amnicola pasionensis Goodrich & Van der Schalie 1937; Misc. Pub. Aroapyrgus pasionensis (Goodrich & Van der Schalie). Thompson 1968; Fla. Hydrob.:150.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:28.Type Locality.Arroyo Subn, a tributary of the Ro Guatemala. Holotype UMMZ. Distribution.GUATEMALA: known from the Ro Usumacinta and tributaries, TABASCO, Mxico. Aroapyrgus petenensis (Morelet 1851)Paludina petenensis Morelet 1851; Test. Noviss. II:21. Amnicola petenensis (Morelet). Frauenfeld 1864:635. Fischer & Martens 1899; Biol. Cent. Amer.:430.Goodrich, and Van der Schalie 1937; Misc. Pub. Mus. Zool. Univ. Mich. (34):36. Aroapyrgus petenensis (Morelet). Thompson 1968; Fla. Hydrob.:150.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:28. Type Locality.Lago de Itza, Dept. Petn, Guatemala. Distribution.GUATEMALA: known from various lakes in the Dept. Petn Aroapyrgus rhegoides (Morelet 1851)Paludina rhegoides Morelet 1851; Test. Noviss. II:22. Amnicola rhegoides (Morelet). Von Martens 1899; Biol. Cent. Aroapyrgus rhegoides (Morelet). Hershler & Thompson 1992; Malac. Rev. Suppl. 5:28.Type Locality.Lago de Coatepeque, Dept. Santa Ana, El Salvador. Distribution.Known only from the type locality. Aroapyrgus stolli (Von Martens 1901)Amnicola stolli Von Martens 1901; Biol. Cent. Amer.:645; pl. 46, Aroapyrgus stolli (Von Martens). Thompson 1968; Fla. Hydrob.:150.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:28.Type Locality.Lago de Atitln, Dept. Solol, Guatemala. Distribution.Known only from the type locality. Aroapyrgus subangulatus (Von Martens 1899)Amnicola subangulata Von Martens 1899; Biol. Cent. Amer.:435; Aroapyrgus subangulata (Von Martens). Thompson 1968; Fla. Hydrob.:150.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:28.Type Locality.Ro Torres, in the environs of San Jos, Prov. San Jos, Costa Rica; 1100 m alt. from the type locality and from a small stream between the Ro Madre de Dios and the Ro Hondo, 80 m alt. Aroapyrgus tryoni (Pilsbry 1904)Amnicola tryoni Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:781; Aroapyrgus tryoni (Pilsbry). Thompson 1968; Fla. Hydrob.:150.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:29.Type Locality.Roots of plants in a swampy pool near Javal, Dept. Chontales, Nicaragua. Holotype ANSP 58066a. Distribution.Known only from the type locality. mi SW of Cartago, 1200 m (Pilsbry 1920). A record from Tampico, Mxico almost certainly is in error (Hinkley 1907:79). Genus Chorrobius Hershler, Liu & Landye 2011Chorrobius Hershler, Liu & Landye, 2011; J. Molluscan Studies 77:16.Type Species.Chorrobia crassilabrum Hershler, Liu & Landye 2011. Distribution.COAHUILA. Taxonomy.A single species is known. Corrobius crassilabrum Hershler, Liu & Landye 2011Corrobius crassilabrum Hershler, Liu & Landye 2011; J. Molluscan 7A-D (radula), 8A-C (reproductive anatomy)Type Locality.An unnamed spring in a private recreation area on the road to Los Lirios (south-southeast of Distribution.COAHUILA: known from four small springs in the immediate vicinity of the type locality.Genus Coahuilix Taylor 1966Coahuilix Taylor 1966; Veliger 9:180.Hershler 1985; Malacologia 26:53.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:31.

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50 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Type Species.By original designation: Coahuilix hubbsi Taylor 1966. Distribution.Cuatro Cinegas Basin, Coahuila, Mxico. Taxonomy.Two species are recognized (Hershler & Thompson 1992). Coahuilix hubbsi Taylor 1966Coahuilix hubbsi 17. Type Locality.Pozo de Becerra, Cuatro Cinegas Basin, Coahuila, Mxico. Holotype UMMZ 220180. Distribution.COAHUILA: endemic to springs in the Cuatro Cinegas Basin. Coahuilix landyei Hershler 1985Coahuilix landyei G, I, J, K.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:33.Type Locality.A small spring 1.12 km south of the north tip of the Sierra de San Marcos, Cuatro Cinegas Basin, Coahuila, Mxico. Holotype ANSP A9894n. Distribution.COAHUILA: endemic to springs in the Cuatro Cinegas Basin. Genus Cochliopa Stimpson 1865Cochliopa Stimpson 1865; Amer. Jour. Conch. 1:52. Morrison 1946; Smiths. Misc. Coll. 106:27.Taylor 1966; Veliger 9:175.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:33. Type Species.Amnicola rowelli Tryon 1863, by original designation. Costa Rica. Habitat.Brackish water and freshwater just above the brackish zone. Taxonomy.Four species are recognized. Cochliopa diazensis Morrison 1946Cochliopa diazensis Morrison 1946; Smiths. Misc. Coll. 106:27; Rev. Suppl. 5:33.Type Locality.Ro Juan Diaz, just below Las Sabanas Road bridge east of Cd. Panam, Prov. Panam, Panam. USNM 542170. Distribution.Known only from the type locality. Cochliopa josana Morrison 1946.Cochiopa josana Morrison 1946; Smiths. Misc. Coll. 106: 28; Type Locality.Ro Mata Puerco, which opens into the Ensenada de Bodega, southwest part of Isla San Jos, Archipelago de Las Perlas, Panam. Holotype USNM 542173. Distribution.PANAM: known only from small streams on Isla San Jos. Cochliopa perforata Thompson & Hershler 1991Cochliopa perforata Thompson & Hershler 1991; Proc. Biol. Soc. Bay, Prov. Puntarenas, Costa Rica. Holotype UF 175911. Distribution.COSTA RICA, Prov. Puntarenas: known Cochliopa rowelli (Tryon 1863)Amnicola rowelli Tryon 1863; Proc. Acad. Nat. Sci. Phila. 15:147; Cochliopa rowelli (Tryon). Stimpson 1865; Amer. Jour. Conch. 1:52.Pilsbry 1905; Nautilus 19:91.Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 72:198.Morrison 1946; Smiths. Misc. Hershler & Thompson 1992; Malac. Rev. Suppl. 5:34.Type Locality.Restricted to Ro Matasnillo, Cd. Panam, Panam (Morrison 1946). Lectotype ANSP 58282a (Baker 1946). Distribution.PANAM: known only from the type locality, where it is now extinct. Genus Cochliopina Morrison 1946Cochliopina Morrison 1946; Smiths. Misc. Coll. 106:18.Taylor 1966; Veliger 9: 176.Hershler 1985; Malacologia 26:65.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:36. Type Species.By original designation: Cochliopa riograndensis Pilsbry and Ferriss 1906. Distribution.North America, Central America, and South America from Texas south to the Ro Santiago, Ecuador. Taxonomy.Nineteen species are recognized, all occuring the study area. Cochliopina australis Morrison 1946Cochliopina australis Morrison 1946; Smiths. Misc. Coll. 106:23 Hershler & Thompson 1992; Malac. Rev. Suppl. 5:37.Type Locality.Musselshell Creek just above the lagoon on the southeast end of Isla San Jos, Archipelago de Las Perlas, Panam. Holotype USNM 542165. Distribution.Known only from the type locality. Cochliopina compacta (Pilsbry 1910)Cochliopa compacta Pilsbry 1920; Proc. Acad. nat. Sci. Phila. 72:197. Cochliopina compacta (Pilsbry). Morrison 1946; Smiths. Misc. Coll. 106:19.Taylor 1966; Veliger 9:176.Hershler & Thompson Type Locality.Ro Choy, at the cave, south of Las Palmas, San Lus Potos, Mxico. Holotype ANSP 99007a. Distribution.Known only from the type locality. Cochliopina dulcensis (Marshall 1920)Cochliopa dulcensis Marshall 1920; Proc. U. S. Nat. Mus. 58:302; Cochliopina dulcensis (Marshall). Morrison 1946; Smiths. Misc.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 51Coll. 106:19.Taylor 1966; Veliger 9:176.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:38.Type Locality.Ro Dulce, Dept. Izabal, Guatemala. Holotype USNM 336413. Distribution.Known only from the type locality. Cochliopina extremis Morrison 1946Cochliopina extremis Morrison 1946; Smiths. Misc. Coll. 106:22 Hershler & Thompson 1992; Malac. Rev. Suppl. 5:38.Type Locality.Small stream with a 100-yard lagoon just east of the southern tip of Isla San Jos, Archipelago de Las Perlas, Panam. Holotype USNM 542163. Distribution.Known only from the type locality. Cochliopina francesae (Goodrich & Van der Schalie 1937)Cochiopa francesae Goodrich & Van der Schalie 1937; Misc. Pub. Cochliopina francesae (Goodrich & Van der Schalie). Morrison 1946; Smiths. Misc. Coll. 106:19.Taylor 1966; Veliger 9:176.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:38.Type Locality.Ro de La Pasion, at the mouth of the Arroyo Chajchinic, northwest of Porvenir, Dept. Alta Verapaz, Guatemala. Holotype in the UMMZ. GUATEMALA, and the Ro San Pedro, TABASCO, Mxico. Cochliopina fratercula Morrison 1946Cochliopina fratercula Morrison 1946; Smiths. Misc. Coll. 106:21 Hershler & Thompson 1992; Malac. Rev. Suppl. 5:38.Type Locality.Ro Mata Puerco, Isla San Jos, Archipelago de Las Perlas, Panam. Holotype USNM 542157. Distribution.Known only from the type locality. Cochliopina guatemalensis (Morelet 1851) Morelet 1851; Test. Noviss. II:22.Fischer & Cochliopa guatemalensis (Morelet). Von Martens 1899; Biol. Cent. Amer.:428.Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 72:198. Cochliopina guatemalensis (Morelet). Morrison 1946; Smiths. Misc. Coll. 106:19.Taylor 1966; Veliger 9:176.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:38.Type Locality.Ro Michatoya, near the port of Istapa [Puerto Ixstapa], Dept. Escuintla, Guatemala. Distribution.GUATEMALA: known for certain only from the type locality. Cochiopina hinkleyi (Pilsbry 1920)Cochliopa hinkleyi Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. Cochliopina hinkleyi (Pilsbry). Morrison 1946; Smiths. Misc. Coll. 106:19.Taylor 1966; Veliger 9:176.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:39.Type Locality.Lago de Izabal near Jocolo, Dept. Izabal, Guatemala. Holotype ANSP 46246a. Distribution.Known only from the type locality. Cochliopina infundibulum (Von Martens 1899)Cochliopa infundibulum Von Martens 1899; Biol. Cent. Amer.:429; Cochliopina infundibulum (Von Martens). Morrison 1946; Smiths. Misc. Coll. 106:19.Taylor 1966; Veliger 9:176.Hershler & Type Locality.Guatemala. Distribution.Ro Usumacinta System of TABASCO and CHIAPAS, Mxico. GUATEMALA: Dept. Petn Cochliopina izabal (Pilsbry 1920)Cochliopa izabal Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 72:200 Cochliopa izabal perstriata Pilsbry 1920; Proc. Acad. Nat. Sci. Coichliopina izabal (Pilsbry). Morrison. 1946:19.Taylor 1966; Veliger 9:176.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:39.Type Locality.Lago de Izabal, near Jocolo, Dept. Izabal, Guatemala. Holotype ANSP 45638a. Distribution.Known only from the type locality. Cochliopina juradoi Morrison 1946Cochliopina juradoi Morrison 1946; Smiths. Misc. Coll. 106:20; & Thompson 1992; Malac. Rev. Suppl. 5:39.Type Locality.Stream leading to the northwest mangrove marsh on Isla San Jos, Archipelago de las Perlas, Panam. Holotype USNM 542154. Distribution.Known only from the type locality. Cochliopina milleri Taylor 1966Cochliopina milleri h, 24.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:39.Type Locality.Ro Mesquites at the main road 9 km southwest of Quatro Cinegas, Coahuila, Mxico. Holotype UMMZ 220182. Distribution.COAHUILA: known only from the immediate vicinity of the type locality. Cochliopina minor (Pilsbry 1920)Cochliopa minor Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 72:199 Cochliopina minor (Pilsbry). Morrison 1946; Smiths. Misc. Coll. 106:19.Taylor 1966; Veliger 9:177.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:39. Holotype ANSP 58286. Distribution.Known only from the type locality. Morrison 1946 Morrison 1946; Smiths. Misc. Coll. 106:22; pl.

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52 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)& Thompson 1992; Malac. Rev. Suppl. 5:39. southwestern bay, Isla San Jos, Archipelago de Les Perlas, Panam. Holotype USNM 542160. Distribution.Known only from the type locality. Cochliopina picta (Pilsbry 1910)Cochliopa picta Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 72:197. Cochliopina picta (Pilsbry). Morrison 1946; Smiths. Misc. Coll. 106:19.Taylor 1966; Veliger 9:177.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:39.Type Locality.Ro Coy, near the ford on the road to Tampamolon, San Lus Potos, Mxico. Holotype ANSP 99013a. Distribution.SAN LUS POTOS: known only from the vicinity of the type locality, and the Ro Ganina near Cochliopina riograndensis (Pilsbry & Ferriss 1906)Cochliopa riograndensis 10.Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 72:197. Cochliopina riograndensis (Pilsbry & Ferriss). Morrison 1946; Smiths. Misc. Coll. 106:18.Taylor 1966; Veliger 9:177.Thompson 1992; Malac. Rev. Suppl. 5:93.Type Locality.Ro San Filipe, near the Ro Grande, Val Verde County, Texas. Holotype ANSP 91324a. Distribution.Streams in Val Verde County, Texas. COAHUILA: Quatro Cinegas Basin. SAN LUS POTOS: Ro Valles. TAMAULIPAS: Ro Panuco and Ro Sabinas systems. Cochliopina tryoniana (Pilsbry 1890)Cochliopa tryoniana Pilsbry 1890; Nautilus 4:52.Pilsbry 1891; Pilsbry 1920:198. Cochliopina tryoniana (Pilsbry). Morrison 1946; Smiths. Misc. Coll. 106:18.Taylor 1966; Veliger 9:177.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:40. Nicaragua. ANSP 58285a. south to the Ro Terraba. COSTA RICA. Cochliopina wetmorei Morrison 1946Cochliopina wetmorei Morrison 1946; Smiths. Misc. Coll. 106:24 Hershler & Thompson 1992; Malac. Rev. Suppl. 5:40.Type Locality.Ro Marina, east side of Isla San Jos, Archipelago de Las Perlas, Panam. Holotype USNM 542167. Distribution.Known only from the type locality. Cochliopina zeteki Morrison 1946Cochliopina zeteki Morrison 1946; Smiths. Misc. Coll. 106:19; Hershler & Thompson 1992; Malac. Rev. Suppl. 5:40.Type Locality.Dead leaves along the east margin of the Ro Juan Diz, just below the Las Sabanas Road bridge, east of Cd. Panam, Prov. Panam, Panam. Holotype USNM 542152. Distribution.Known only from the type locality. Genus Emmericiella Pilsbry 1909Emmericiella Pilsbry 1909; Nautilus 23:46.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:42.Type Species.By original designation: Ememericia (Emmericiella )Pilsbry 1909. Distribution.Subterranean headwaters of the Ro Choy, San Lus Potos, Mxico. Taxonomy.The genus contains two species. Emmericiella longa (Pilsbry 1909)Emmericia (Emmericiella ) longa Pilsbry 1909; Nautilus 23:46; pl. Emmericiella longa (Pilsbry). Hershler & Thompson 1992; Malac. Rev. Suppl. 5:44.Type Locality.Ro Choy, Luis Potos, Mxico. Holotype ANSP 99022. Distribution.Known only from the type locality. (Pilsbry 1909)Emmericia (Emmericiella ) Pilsbry 1909; Nautilus 23:46; (Pilsbry). Hershler & Thompson 1992; Type Locality.Banks of the Ro Choy near the cave, San Luis Potos, Mxico. Holotype ANSP 99021a. Distribution.Known only from the type locality. Genus Eremopyrgus Hershler 1999Eremopyrgus Hershler 1999; Veliger 42:306.Type Species.Eremopyrgus eganensis Hershler, 1999. Distribution.Nevada State, U.S.A. and Chihuahua State, Mxico. Taxonomy.Two species are recognized. One occurs in Mxico. Eremopyrgus elegans Hershler, Liu & Lande 2002Eremopyrgus elegans Hershler, Liu & Lande 2002; J. Molluscan Studies 68:81; Fig. 1 (shell); Fig. 2 (operculum); Fig. 3 (radula); Fig. 4 (reproductive anatomy); Fig. 5 (map) .Type Locality.Ojo Vareleo, 2.4 km NW of Casas Holotype USNM 892119. Distribution.Known only from the type locality. Genus Juturnia Hershler, Liu and Stockwell 2002Juturnia Hershler, Liu and Stockwell 2002; Proc. Biol. Soc. Wash. 115:172. Type Species.Durangonella coahiilae Taylor 1966.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 53 Distribution.Lower Rio Grande region of North America. Taxonomy.Three species, one of which occurs in Mxico. Juturnia coahuilae (Taylor 1966)Durangonella coahuilae Taylor 1966; Veliger 9:184; pl. 14, 24b-d, 25. Juturnia coahuilae (Taylor).Hershler, Liu and Stockwell 2002; Proc. Biol. Soc. Wash. 115:175. Type Locality.Laguna Grande within about 300 m of the mouth of the Ro Churince, and about 17 km southwest of Cuatro Cinegas, Coahuila, Mxico. Holotype UMMZ 220159. Distribution.COAHUILA: endemic to the Cuatro Cinegas Basin. Genus Lithococcus Pilsbry 1911Lithococcus Pilsbry 1911; Princeton Exp. Patagonia, 3:602.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:61.Type Species.By original designation: Lithoglyphus multicarinatus Miller 1878. Costa Rica south to the Rio Santiago system in northwest Ecuador. Taxonomy.Three species are recognized, one of which occurs in Costa Rica. Lithococcus aletes Thompson & Hershler 1991Lithococcus aletes Hershler & Thompson 1992; Malac. Rev. Suppl. 5:63.Type Locality.Ro Bar, 0.5 km above mouth of river, Prov. Puntarenas, Costa Rica. Holotype UF 36378. Distribution.Known only from the type locality. Genus Littoridina Souleyet 1852Littoridina Souleyet 1852:563.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:63. Type Species.By original designation Littoridina guadichaudii Souleyet 1952. Distribution.North America, Central America and South America: coastal streams and estuaries from Texas and Colima, Mxico south to Ecuador. Habitat.Brackish marshes and freshwater streams. Taxonomy.Four species. Many tropical American hydrobioids have been described as species of Littoridina Most of these now are placed in Heleobia (Hershler & Thompson 1992). Littoridina crosseana (Pilsbry 1910)Amnicola crosseana Littoridina crosseana (Pilsbry). Hershler & Thompson 1992; Malac. Type Locality.Ponds at La Barra, Tampico, Tamaulipas, Mxico. Holotype ANSP 99502a. Distribution.Common in brackish marshes in the immediate area of Corpus Cristi, Texas. TAMAULIPAS: Tampico. TABASCO. Littoridina microcona Thompson & Hershler 1991Littoridina microcona Thompson & Hershler 1991; Proc. Biol. Soc. Type Locality.Marshy pool 1.0 km north of Boca de Barranca, Prov. Puntarenas, Costa Rica. Holotype UF 175912. streams and marshes in Prov. Puntarenas Littoridina orcutti (Pilsbry 1928)Amnicola orcutti Pilsbry 1928; Proc. Acad. Nat. Sci. Phila. 80:115 Littoridina orcutti (Plsbry). Hershler & Thompson 1992; Malac. Type Locality.Manzanillo, Colima, Mxico. Holotype ANSP 144801. Distribution.NAYARIT: brackish marshes 2 km northeast of San Blas, southeast to the Laguna de Mitla, GUERRERO. Genus Littoridinops Pilsbry 1952Littoridinops Pilsbry 1952; Nautilus 66:51.Thompson 1968:56.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:68.Type Species.By original designation Amnicola tenuipes Couper 1844. Distribution.Atlantic coast of North America from New Jersey south to Campeche, Mxico; Bahama Islands, Grand Cayman Island, Cuba. Habitat.Brackish and freshwater. Taxonomy.Four species, one of which occurs in Mxico. Littoridinops tampicoensis (Pilsbry & Hinkley 1907)Paludestrina tampicoensis Pilsbry & Hinkley 1907; Nautilus 21:39; Littoridinops tampicoensis (Pilsbry & Hinkley). Taylor 1966; Veliger 9:187.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:70.Type Locality.In river drift, Tampico, Mxico. ANSP 93813a. Distribution.TAMAULIPAS-VERACRUZ: known only the immediate vicinity of the type locality. Genus Mesobia Thompson & Hershler 1991Mesobia Thompson & Hershler 1991; Proc. Biol. Soc. Wash. 104:679.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:74. Type Species.Mesobia pristina Thompson and Hershler 1991. Distribution.Known only from Lago de Yajoa,

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54 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Honduras. Habitat.Freshwater. Taxonomy.One species. Mesobia pristina Thompson & Hershler 1991Mesobia pristina Thompson & Hershler 1991; Proc. Biol. Soc. Type Locality.Lago de Yajoa, at the north side of an island near Agua Azul, Dept. Corts, Honduras. Holotype UF 40421. Distribution.Known only from the type locality. Genus Mexipyrgus Taylor 1966Mexipyrgus Taylor 1966; Veliger 9:188.Hershler 1985; Malacologia 26:87.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:74.Type Species.Mexipyrgus carranzae Taylor 1966. Distribution.COAHUILA: endemic to the Cuatro Cinegas Basin. Habitat.Freshwater. Taxonomy.The genus is monotypic. Mexipyrgus carranzae Taylor 1966Mexipyrgus carranzae Mexipyrgus churinceanus Taylor 1966; Veliger 9:190; pl. 16, Mexipyrgus escobedae 23. Mexipyrgus lugoi 45. Mexipyrgus mojarralis 46; 51. Mexipyrgus multilineatus Taylor 1966; Veliger 9:193; pl. 18, Holotypes.Mexipyrgus carranzae: UMMZ 220211. Mexipyrgus churinceanus: UMMZ 220150. Mexipyrgus escobedae: UMMZ 220202. Mexipyrgus lugoi: UMMZ 220185. Mexipyrgus mojarralis: UMMZ 220192. Mexipyrgus multilineatus : UMMZ 220197. Type Localities. Mexipyrgus carranzae Laguna Tio Candido, 14 km south of Cuatro Cinegas. Mexipyrgus churinceanus: Laguna Churince, 16 km southwest of Cuatro Cinegas. Mexipyrgus escobedae: Laguna Escobeda, 12 km south of Cuatro Cinegas. Mexipyrgus lugoi: Ro Mesquites at the main road 9 km southwest of Cuatro Cinegas. Mexipyrgus mojarralis: West Laguna in El Mojarral, 1.7 km ENE of the northern tip of Sierra San Marcos. Mexipyrgus multilineatus : East Laguna in El Mojarral, 1.9 km ENE of the northern tip of Sierra San Marcos. All localities are in the Cuatro Cinegas Basin, Coahuila, Mxico. Distribution.COAHUILA: endemic to the Cuatro Cinegas Basin. Genus Mexithauma Taylor 1966Mexithauma Taylor 1966; Veliger 9:205.Hershler 1985; Malacologia 26:72.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:78.Type Species.Mexithauma quadripaludium Taylor 1966. Distribution.Mxico, endemic to the Cuatro Cinegas basin, Coahuila. Habitat.Freshwater. Taxonomy.Monotypic. Mexithauma quadripaludium Taylor 1966Mexithauma quadripaludium Taylor 1966; Veliger 9:205; Rev. Suppl. 5:78; 79.Type Locality.Laguna Tio Candido, 9.3 km south of the northern tip of the Sierra de San Marcos, Cuatro Cinegas Basin, Coahuila (Hershler 1985). Holotype UMMZ 220214. Distribution.COAHUILA: endemic to the Cuatro Cinegas Basin (Hershler 1985). Genus Minkleyella Hershler, Liu & Landye 2011Minkleyella Hershler, Liu & Landye 2011; J. Molluscan Studies 77:20.Type Species.Minkleyella balnearis Hershler, Liu, & Landye 2011. Distribution.Known from a single locality in Chihuahua. Taxonomy.A single species is recognized. Minkleyella balnearis Hershler, Liu & Landye 2011Minkleyella balnearis Hershler, Liu & Landye 2011; J. Molluscan 7E-H (radula), 8D-F (reproductive anatomy). end of parking lot) south-southwest of Ciudad Jimenez, 874037. Distribution.Known only from the type locality. Genus Onobops Thompson 1968Onobops Thompson 1968; Fla. Hydrob.:28.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:83.Type Species.Onobops crassa Thompson 1968. Distribution.Atlantic coast of the United States from Maryland south to Campeche, Mxico. Habitat.Brackish marshes. Taxonomy.Two species occur in the southeastern present in the Florida Museum of Natural History from the Gulf coastal region of the United States and Mxico. Genus Paludiscala Taylor 1966Paludiscala Taylor 1966; Veliger 9:207.Hershler 1985; Malacologia 26:58.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:85. Type Species.Paludiscala caramba Taylor 1966. Distribution.Endemic to the Cuatro Cinegas Basin, Coahuila, Mxico.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 55 Habitat.Freshwater. Taxonomy.The genus is monotypic. Paludiscala caramba Taylor 1966Paludiscala caramba Type Locality.Stream draining marsh due east of Majorral West Laguna, about 10 km south of Cuatro Cinegas, Coahuila, Mxico. Holotype UMMZ 220164. Distribution.COAHUILA: springs in the Cuatro Cinegas Basin (Hershler 1985). Genus Phreatoceras Hershler & Longley 1987Hadoceras Hershler & Longley 1986; Proc. Biol. Soc. Wash. 99:122. (Not Hadoceras Strand 1934, Cephalopoda.) Phreatoceras Hershler & Longley 1987; Proc. Biol. Soc. Wash. 100:402.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:87. Type Species.Hadoceras taylori Hershler & Longley 1986. Distribution.North America; Springs in Real County, Texas, and in the Cuatro Cinegas Basin, Coahuila, Mxico. Taxonomy.Monotypic. Phreatoceras taylori (Hershler & Longley 1986)Hadoceras taylori Hershler & Longley 1986; Proc. Biol. Soc. Phreatoceras taylori (Hershler & Longley). Hershler & Longley 1987:402.Hershler & Thompson 1992; Malac. Rev. Suppl. Type Locality.Roaring Springs, Real County, Texas. Holotype USNM 849000. Distribution.Known from springs in Real Co., and Williamson Co., Texas, and the Cuatro Cinegas Basin, COAHUILA. Genus Pseudotryponia Hershler 2001Pseudotryonia Hershler 2001; Smithonian Contr. Zool., 612:15 16.Hershler, Liu & Landye 2011; J. Molluscan Studies 77:10.Type Species.Tryonia alamosae Taylor 1987. Distribution.Southeastern United States, Texas, New Mexico, Chihuahua and Durango, Mxico. Taxonomy.The genus includes six specie. Two occur in Mxico. Pseudotryonia mica Hershler, Liu & Landye 2011Pseudotryonia mica Hershler, Liu & Landye 2011; J. Molluscan 3A-E (radula), 4 A-C (rproductive anatomy).Type Locality.Ojo de Dolores, mani spring pool, Distribution.CHIHUAHUA: known only from the immediate vicinity of the type locality. Tryonia pasajae Hershler, Liu & Landye 2011Tryonia pasajae Hershler, Liu & Landye 2011; J. Molluscan Studies (radula), 4D,-E (reproductive aatomy).Type Locality.An unnamed spring at El Tanque (second pool below spring source) wet-northwest of Cuencame de Ceniceros, Durango, Mxico. Holotype USNM 1133662. Distribution.Known only from the type locality. Genus Pyrgophorus Ancey 1888Pyrgophorus Ancey 1888; Bull. Soc. Malac. France 5:188.Taylor 1966; Veliger 9:194.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:89.Type Species.Pyrgulopsis spinosus Pilsbry & Call 1886. Distribution.West Indies, North America, Central America and South America; south Florida, the West Indies excluding the Bahama Islands and the Cayman Islands; coastal Mississippi, Louisiana, Texas, south though the coastal lowlands of east Mxico to the Isthmus of Tehuantepec; lowland areas from Chiapas and Tabasco south the Caribbean-Atlantic drainage systems east and south to Par State, Brazil. Habitat.Primarily freshwater; occasionally in brackish water. Taxonomy.The species-level systematics of Pyrgophorus subspecies have been described on the basis of shell characters. The shells frequently bear ornate sculpture which varies greatly within and between populations. Similar ornate or smooth shell forms appear throughout the range of the genus. The assignment of names to specimens is highly subjective because most species and subspecies were not adequately diagnosed, illustrated or critically compared to related forms. Taylor (1966) and Hershler and Thompson (1992) reviewed the taxonomic history of the nominate forms. It is uncertain if more than just a few of these are valid. The oldest species name for the genus is Guilding 1828, described from the Windward Islands, West Indies. Most Mxican and Central America populations are referred by authors to Pyrgophorus coronatus (Pfeiffer 1840). Names applicable to most other Mxican and Central American forms are listed in the synonymy given for coronatus. I am comfortable in recognizing only two species in Mxico and Central America, Pyrgophorus coronatus (Pfeiffer 1840) and Pyrgophorus exiguus (Morelet 1851), although other nominate species are also retained in this list. Extralimital taxa are listed in Hershler and Thompson (1992:91). Pyrgophorus coronatus (Pfeiffer 1840)Paludina coronata Pfeiffer 1840:253.Kster 1852; in Martini & Chemnitz, Syst. Conch. Cab., ed. 2 (Paludina 11. Hydrobia coronata (Pfeiffer). Von Martens 1858; in Wiegmanns Archis f. Naturg. 24:192.

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56 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Amnicola coronata (Pfeiffer). Fischer 1860; Jour. de Conchyl. 8:363. Pyrgulopsis coronatus (Pfeiffer). Ancey 1888; Bull. Soc. Malac. France 5:197. Potamopyrgus coronatus (Pfeiffer). Pilsbry 1891; Proc. Acad. Nat. Sci. Phila. 34:328. Pyrgophorus coronatus (Pfeiffer). Taylor 1966; Veliger 9:194. Paludina crystallina Pfeiffer 1840:253. (Matanzas, Matanzas Province, Cuba [Aguayo and Jaume 1947].) Melania spinifera C. B. Adams 1845:17. (Jamaica.) Paludina jamaicensis C. B. Adams 1849. (Jamaica.) Paludina cisternicola Morelet 1851; Test. Noviss. II:21. (Campeche, Campeche, Mxico.) Paludina ornata Morelet 1851; Test. Noviss. II:21. (Lago de Coatepeque, El Salvador.) Paludina cisternina Campeche, Mxico.) Amnicola candeana inermis Fischer 1860; Jour. de Conchyl. 8:363. (Type locality not stated.) Amnicola candeana gibbosa Fischer 1860; Jour. de Conchyl. 8:364. (Bay of Campeche, Mxico.) Frauenfeld 1863:526; pl. 8. Pyrgulopsis nicaraguanus Ancey 1888; Bull. Soc. Malac. France 5:194. (Nicaragua.) Pyrgula nicaraguensis Newcomb, MS; in Ancey 1888; Bull. Soc. Malac. France 5:194. Pyrgulopsis nicaraguanus costuliferus Ancey 1888; Bull. Soc. Malac. France 5:195. (Nicaragua.) Purgulopsis nicaraguanus duplicatus Ancey 1888; Bull. Soc. Malac. France 5:195. (Nicaragua.) Pyrgulopsis newcombianus Ancey 1888; Bull. Soc. Malac. France 5:196. (Nicaragua.) Pyrgulopsis productus Ancey 1888; Bull. Soc. Malac. France 5:197. (Nicaragua.) Purgulopsis wrighti Ancey 1888; Bull. Soc. Malac. France 5:199. (Lago de Coatepeque, El Salvador.) Pyrgulopsis wrighti plicosus Ancey 1888; Bull. Soc. Malac. France 5:199. (Lago de Coatepeque, El Salvador). Pyrgulopsis wrighti transitans Ancey 1888; Bull. Soc. Malac. France 5:200. (Lago de Coatepeque, El Salvador). Pyrgulopsis wrighti oblongus Ancey 1888; Bull. Soc. Malac. France 5:200. (Lago de Coatepeque, El Salvador). Pyrgulopsis wrighti minimus Ancey 1888; Bull. Soc. Malac. France 5:201. (Lago de Coatepeque, El Salvador). Pyrgulopsis wrighti obesus Ancey 1888; Bull. Soc. Malac. France 5:201. (Lago de Coatepeque, El Salvador). Amnicola coronata unicarinata Von Martens 1899; Biol. Cent. Amer.:433. Type locality not stated. Here restricted to Veracruz, Veracruz, Mxico. Earlier, Von Martens (1873) recorded this form from Vieque. Taylor (1966) and Hershler & Thompson (1992) listed unicarinatus as a nomum nudum, but the form is diagnosed by Von Martens (1899:431), and he cited 34a), thus validating the use of the name. Pyrgulopsis coronatus Ancey 1888; Bull. Soc. Malac. France 5:198 (non Paludina coronata Pfeiffer 1840). (Veracruz, Mxico).Type Locality.Matanzas, Matanzas Province, Cuba (Aguayo and Jaume 1947). Distribution.Coastal and lowland regions of Mxico from Tamaulipas and Oaxaca south to Venezuela; West Indies in general. Pyrgophorus chagresensis (Morrison 1946)Lyrodes chagresensis Morrison 1946; Smiths. Misc. Coll. 106:16 Pyrgophorus chagresensis (Morrison). Taylor 1966; Veliger 9:194.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:92.Type Locality.Ro Chagres near Gatuncillo, Prov. Panam, Panam. Holotype USNM 542149. Distribution.Known only from the type locality. Pyrgophorus conoideus (Ancey 1888)Pyrgulopsis conoideus Ancey 1888; Bull. Soc. Malac. France 5:196. Pyrgophorus conoideus (Ancey).Taylor 1966; Veliger 9:194.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:92.Type Locality.Nicaragua. Distribution.NICARAGUA: uncertain distribution. Pyrgophorus hydrobioides (Ancey 1888)Pyrgulopsis hydrobioides Ancey 1888; Bull. Soc. Malac. France 5:201. Amnicola (?) hydrobioides (Ancey). Von Martens 1899; Biol. Cent. Amer.:435. Pyrgophorus hydrobioides (Ancey). Taylor 1966; Veliger 9:194.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:93.Type Locality.Lago Coatepeque, Dept. Santa Ana, El Salvador. Distribution.Known only from the type locality. Remarks.Von Martens (1899) suggested that the type is a mislabeled specimen of from the coast of France. Pyrgophorus spinosus (Call & Pilsbry 1886)Pyrgulopsis spinosus Call & Pilsbry 1886; Proc. Davenport Acad. Potamopyrgus spinosus (Call & Pilsbry).Pilsbry 1891; Proc. Acad. Nat. Sci. Phila. 43:327.Walker 1918; Misc. Pub. Mus. Zool. Univ. Mich. (6):140. Hydrobia texana Pilsbry 1887; Proc. Davenport Acad. Nat. Sci., Braunfels, Comal County, Texas.) Ancey 1888; Bull. Soc. Malac. France 5:193. (Comal Creek, New Braunfels, Comal, County, Texas.) Pyrgophorus spinosus (Call & Pilsbry). Taylor 1966; Veliger 9:195.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:93.Type Locality.Comal Creek, New Braunfels, Comal County, Texas. Syntypes of Potamopyrgus spinosus: not located. Lectotype of Hydrobia texanus: ANSP 68908a (H. B. Baker 1964) Distribution.South Central Texas, including the Rio Bravo del Norte (Rio Grand), COAHUILA and CHIHUAHUA. Remarks.Pilsbry (1891) found little reason for retaining spinosus as a distinct species from coronatus. (Von Martens 1901) von Martens 1901; Biol. Cent.-Ameri.:645; pl.44,

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 57 (von Martens), Thompson 1968; Fla. Hyrdrobiidae:150.Type Locality.Lago de Atitlan, Dept. Solol, Guatemala. Distribution.Known only from the type locality. Pyrgophorus zeteki (Morrison 1946)Lyrodes zeteki Morrison 1946; Smiths. Misc. Coll. 106:17; pl. 2, Pyrgophorus zeteki (Morrison). Taylor 1966; Veliger 9:195.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:94.Type Locality.Pedro Miguel, Canal Zone, Panam. Holotype USNM 432871. Distribution.Known only from the type locality. Genus Subcochliopa Morrison 1946Subcochliopa Morrison 1946; Smiths. Misc. Coll. 106:25.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:99.Type Species. Subcochliopa trochus Morrison 1946. and Costa Rica. Habitat.Brackish water and freshwater zone immediately above brackish zone. Taxonomy.Three species are recognized. Subcochliopa colabrensis Morrison 1946Subcochliopa colabrensis Morrison 1946; Smiths. Misc. Coll. 9:179.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:101.Type Locality.Ro Colabre [Ro Culebra], Ro Bayamo drainage, Prov. Panam, Panam. Holotype USNM 542169. Distribution.Known only from the type locality. Subcochliopa trochulus (Von Martens 1899)Cochliopa trochulus Von Martens 1899; Biol. Cent. Amer.:429; pl. Subcochliopa trochulus (Von Martens). Morrison 1946; Smiths. Misc. Coll. 106:25.Taylor 1966; Veliger 9:197.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:101.Type Locality.Here restricted to El Pozo, Ro Grande de Terraba, Puntarenas Province, Costa Rica. Distribution.Southwestern COSTA RICA. In addition to the type locality the species has been recorded from marshes at Sierpe. Subcochliopa trochus Morrison 1946Subcochliopa trochus Morrison 1946; Smiths. Misc. Coll. 106:25 26.Taylor 1966; Veliger 9:179.Hershler & Thompson (operculum), 65e (radula), 66 (reproductive system).Type Locality.Sona, Ro Tribique, Prov. Veraguas, Panam. Holotype USNM 542168. Distribution.PANAM. occurring in smaller streams draining into the Ro San Pablo, Prov. Veraguas: Quebrada Tolerique, 3 km SE of Sona; Rio San Juan, 2.5 km SE of Soledad. Genus Tepalcatia Thompson & Hershler 2002Tepalcatia Thompson & Hershler 2002; Proc. Biol. Soc. Wash. 115:196.Type Species.By original designation Tepalcatia tela Thompson & Hershler 2002. Distribution.The Rio Balsas basin of southern Mxico. Habitat.Freshwater springs and streams. Taxonomy.Three species are recognized. Tepalcatia bakeri (Pilsbry 1891)Potamopyrgus ? bakeri Pilsbry 1891; Nautilus 5:9Pilsbry 1891; Fischer & Crosse 1891; Miss. Sci. Mex. II:277.Baker 1964; Proc. Acad. Nat. Sci. Phila. 116:171 (lectotype selection). Amnicola bakeri (Pilsbry 1891). Von Martens 1899; Biol. Cent. Amer.:435. Tryonia bakeria (Pilsbry 1891). Taylor 1966; Veliger 9:196. Tepalcatia bakeri (Pilsbry 1891). Thompson & Hershler 2002; Proc. Type Locality.Specimens were dug from the bank of a stream east of Yauntepec, Morelos. Lectotype ANSP 61578 (Thompson & Hershler). Distribution.Known only from the type locality. Tepalcatia polia (Thompson & Hershler 1991)Aroapyrgus polius Thompson & Hershler 1991; Proc. Biol. Soc. Malac. Rev. Suppl. 5:28. Tepalcatia polia (Thompson & Hershler). Thompson & Hershler Type Locality.A small spring-fed pool 3.5 km northeast of Tamazulapn del Progreso, Oaxaca, Mxico. Holotype UF 175028. Distribution.Known only from the type locality. Tepalcatia tela Thompson & Hershler 2002Tepalcatia tela Thompson & Hershler 2002; Proc. Biol. Soc. Wash. Type Locality.A stream (Los Ultimos) 3.0 km north Distribution.MICHOACN: in addition to the type locality this snail is known from a spring-fed lagoon, La Mojada, 7.5 km NW of Apatzingan. Genus Texadina Abbott & Ladd 1951Texadina Abbott & Ladd 1951; Jour. Wash. Acad. Sci. 41:335.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:102. Type Species.Littoridina (Texadina) sphinctostoma Abbott & Ladd 1951. Distribution. Coast of the Gulf of Mexico from Pensacola Bay, Florida, west and south to the Laguna de Terminos, Campeche, Mxico; Jamaica. Habitat.Brackish marshes. Taxonomy.Two species are recognized. One occurs in Mxico.

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58 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Texadina sphinctostoma (Abbott & Ladd 1951)Littoridina (Texadina ) sphinctostoma Abbott & Ladd 1951; Jour. Nautilus 74:158. Texadina sphinctostoma (Abbott & Ladd). Taylor 1966; Veliger 9:196.Garca-Cubas 1963; Bol. Inst. Geol. Univ. Nac. Aut. Mx., 67:1.Hershler & Thompson 1992; Malac. Rev. (radula), 68 (reproductive anatomy).Type Locality..5 miles north of Webb Point, northwest side of San Antonia Bay, 27 miles northeast of Rockport, Texas. Holotype USNM 596722. Distribution.Marshes along the Gulf of Mexico coast from Pensacola Bay, Florida, west and south to Campeche. The only record in Mxico is the Laguna de Terminos, CAMPECHE (Garca-Cubas 1963). Genus Tryonia Stimpson 1865Tryonia Stimpson 1865; Smiths. Misc. Coll. (201):54.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:107. (Type species: Tryonia clathrata Stimpson 1865.).Hershler 2001; Smiths. Contrib. Zool., (612):3. Calipyrgula Pilsbry 1934; Nautilus 48:15. (Type species: Calipyrgula carinifera Pilsbry 1934.) Durangonella Morrison 1945:20.Taylor 1966; Veliger 9:184.Hershler 1985; Malacologia 26:78.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:40. (Type species: Hydrobia seemani Frauenfeld 1863.) Hyalopyrgus Thompson 1968:43. (Type species: Bythinella aequicostata Pilsbry 1889.) Paupertryonia Taylor 1987; Bull. New Mexico Bur. Mines and Min. Res. 116:41. (Type species: Potamopyrgus cheatumi Pilsbry 1935.) Distribution.Northern Sonora and Chihuahua, Mxico; southwestern desert region of California, Nevada, Utah, Arizona, New Mexico and western Texas, and in the southeast in the central part of the Florida Peninsula. Habitat.Freshwater. Taxonomy.The genus includes about 16 extant species (Hershler 2001). Two occur in Mxico. Hershler, Liu and Stockwell (2002) demonstrated that the type species of Durangonella is a Tryonia. Three other species that are known only from shells were described as species of Durangonella, and they are assigned to Tryonia tentatively. Tryonia dugesiana (Morrison 1945)Durangonella dugesiana 3.Taylor 1966; Veliger 9:184.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:42.Type Locality.Andocutira, Michoacn (amended to Andocutin, Guanajuato by Taylor 1966). Holotype USNM 433473. Distribution.Known only from the type locality. Tryonia exiguus (Morelet 1851)Melania exigua Morelet 1851; Test. Noviss. II:23. Tryonia exigua (Morelet). Fischer & Crosse 1891; Miss. Sci. Mex. Amer.:436.Taylor 1966; Veliger 9:197.Hershler 2001; Smiths. Contrib. Zool., (612):9. Paludestrina exigua (Morelet). Goodrich and Van der Schalie 1937; Misc. Pub. Mus. Zool. Univ. Mich. (34):36. Pyrgophorus exigua (Morelet). Hershler & Thompson 1992; Malac. Rev. Suppl. 5:93, 94. Melania minuta Brot 1862:42.Type Locality.Lago de Petn Itza, Dept. Petn, Guatemala. Distribution.Known only from the type locality. Tryonia hertleini (Drake 1956)Lyrodes hertleini Drake 1956; Bull. South. Calif. Acad. Sci. 55:44 46; pl. 15. Tryonia hertleini (Drake). Hershler & Thompson 1992; Malac. Rev. Suppl. 5:110.Hershler 2001; Smiths. Contrib. Zool., (612):10.Type Locality.Springs, Las Palomas, Chihuahua, Mxico. Holotype CAS 9982. Distribution.Known only from the type locality. Possibly extinct. Tryonia mariae (Morrison 1945)Durangonella mariae 2.Taylor 1966; Veliger 9:184.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:42.Type Locality.One meter below the bed of the presently dry Lago de Tlahuac, 20 km east of Xochimilco, Distrito Federal, Mxico. Holotype USNM 433399. Distribution.Known only from the type locality. Tryonia pilsbryi (Morrison 1945)Durangonella pilsbryi 4.Taylor 1966; Veliger 9:184.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:42.Type Locality.Paso del Ro, Colima, Mxico. Holotype USNM 362551. Distribution.Known only from the type locality. Tryonia seemani (Frauenfeld 1863)Hydrobia seemani Frauenfeld 1863:1025.Frauenfeld 1865:1; pl. 8, Amnicola (?) seemanni (Frauenfeld). Von Martens 1899; Biol. Cent. Amer.:435. Durangonella seemani (Frauenfeld). Morrison 1945:20.Taylor 1966; Veliger 9:184.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:42. Tryonia seemani (Frauenfeld). Hershler, Liu and Stockwell 2002; Proc. Biol. Soc. Wash. 115.Type Locality.Durango (it is not clear whether Frauenfeld intended this to mean the city or the state). Cotypes in the Vienna Naturhistorisches Museum. Distribution.DURANGO: known only from a spring at Ciudad Durango, which was a domestic water source for the city many years earlier. Possibly extinct. Genus Zetekina Morrison 1947Zetekella Morrison 1946; Smiths. Misc. Coll. 106:11 [non Zetekella Drake 1944, Hemiptera].

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 59Zetekina Morrison 1947; Nautilus 60:102. Hershler & Thompson 1992; Malac. Rev. Suppl. 5:11214.Type Species.Littoridina frenata Pilsbry 1935. Nicaragua, Costa Rica, and Panam. Habitat.Brackish water and freshwater just above the brackish zone. Taxonomy.Seven species are recognized. Zetekina frenata (Pilsbry 1935)Littoridina frenata Zetekella frenata (Pilsbry). Morrison 1946; Smiths. Misc. Coll. 106:12. Zetekina frenata (Pilsbry). Morrison 1947; Nautilus 60:102.Taylor 1966; Veliger 9:198.Hershler & Thompson 1992; Malac. Rev. Type Locality.Ro Juan Diaz, Cd. Panam, Prov. Panam, Panam. Holotype ANSP 161776. Distribution.Known only from the type locality. Zetekina kompi (Morrison 1946)Zetekella kompi Morrison 1946; Smiths. Misc. Coll. 106:13; pl. 2, Zetekina kompi (Morrison). Taylor 1966; Veliger 9:198.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:114.Type Locality.Ro Mata Puerco, west side of Isla San Jos, Archipelago de Las Perlas, Panam. Holotype USNM 542140. Distribution.Known only from the type locality. Zetekina martensi (Pilsbry 1935)Littoridina martensi Pilsbry 1935; Proc. Acad. Nat. Sci. Phila. 87:5; Zetekella martensi (Pilsbry). Morrison 1946; Smiths. Misc. Coll. 106:11. Zetekina martensi (Pilsbry). Taylor 1966; Veliger 9:198.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:114. Holotype ANSP 27227. Distribution.Known only from the type locality. Zetekina melanioides (Von Martens 1899)Amnicola melanioides Von Martens 1899; Biol. Cent. Amer.:439; Littoridina melanioides (Von Martens). Pilsbry 1935; Proc. Acad. Nat. Sci. Phila. 87:5. Zetekella melanioides (Von Martens). Morrison 1946; Smiths. Misc. Coll. 106:11. Zetekina melanioides (Von Martens). Taylor 1966; Veliger 9:198.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:115.Type Locality.Golfo Dulce, Ro de los Platanales, Prov. Puntarenas, Costa Rica. Distribution.Known only from the type locality. Zetekina panamensis (Bartsch 1920)Syncera panamensis Bartsch 1920; Bull. U. S. Nat. Mus. 58:164; Zetekella panamensis (Bartsch). Morrison 1946; Smiths. Misc. Coll. 106:11. Zetekina panamensis (Bartsch). Taylor 1966; Veliger 9:198.Type Locality.Ro Matasnillo, Panam. Holotype USNM 150870. Distribution.Known only from the type locality. Apparently extinct. Zetekina tenuis (Von Martens 1899)Amnicola melanioides tenuis Von Martens 1899; Biol. Cent. Littoridina tenuis (Von Martens). Pilsbry 1935; Proc. Acad. Nat. Sci. Phila. 87:5. Zetekella tenuis (Von Martens). Morrison 1946; Smiths. Misc. Coll. 106:11. Zetekina tenuis (Von Martens). Taylor 1966; Veliger 9:198.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:112.Type Locality.In a small stream running into the Ro Boto, Golfo Dulce, Prov. Puntarenas, Costa Rica. Distribution.COSTA RICA: known only from the type locality and from marshes at Seripe, Prov. Puntarenas (Von Martens 1899). (Morrison 1946) Morrison 1946; Smiths. Misc. Coll. 106:12, (Morrison). Taylor 1966; Veliger 9:198.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:115.Type Locality.Ro Tribique, Sona, Prov. Veraguas, Panam. Holotype USNM 542139. Distribution.PANAM: streams draining into the Ro San Pablo, Prov. Veraguas Subfamily LITHOGLYPHINAE Tryon 1866 Genus Pterides Pilsbry 1909Pterides Pilsbry 1909; Nautilus 23:48.Herhsler & Thompson 1992; Malac. Rev. Supp. 5:130. Type Species.Pterides pterostoma Pilsbry 1909. Distribution.Ro Choy and Ro Ganina in eastern San Lus Potos, Mxico. Taxonomy.Three species are recognized. Pterides bisinulabris Pilsbry 1909Pterides bisinulabris 7.Type Locality.Ro Ganina, three miles southwest of San Dieguito, San Lus Potos, Mxico. Holotype ANSP 99015. Distribution.Known only from the type locality. Pterides pterostoma Pilsbry 1909Pterides pterostoma 5.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:130.Type Locality.Not given. Holotype ANSP 99016a. Distribution.SAN LUS POTOS (?): unknown. Pterides rhabdus Pilsbry 1909Pterides rhabdus

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60 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Type Locality.Ro Choy near the cave, San Lus Potos, Mxico. Holotype ANSP 99107. Distribution.Known only from the type locality. Subfamily NYMPHOPHILINAE Taylor 1966 Genus Pyrgulopsis Call & Pilsbry 1886Pyrgulopsis Call & Pilsbry 1886; Proc. Davenport Acad. Sci. 5:9.Hershler & Thompson 1987; Nautilus 101:28.Hershler 1994; Smiths. Contrib. Zool. (554):5.Liu & Hershler 2005; Molecular Phylogeny and Evolution 34:284. (Type species by original designation: Stearns 1883.) Fontelicella Gregg & Taylor 1965:103. (Type species by original designation: Fontelicella californiensis Gregg & Taylor 1965.) Natricola Gregg & Taylor 1965:108. (Type species by original designation: Pomatiopsis robusta Walker 1908.) Microamnicola Gregg & Taylor 1965:109. (Type species by original designation: Amnicola micrococcus Pilsbry 1893.) Nymphophilus Taylor 1966. (Type species by original designation: Nymphophilus minkleyi Taylor 1966.) Taylor 1966a:130. (Type species by original designation: Paludestrina nana Chamberlain & Berry 1933.) Mexistiobia Hershler 1985; Malacologia 26:46. (Type species by original designation: Mexistiobia manantiali Hershler 1985.) Apachecoccus Taylor 1987:32. (Type species by original designation: Apachecoccus arizonae Taylor 1987.) Yaquicoccus Taylor 1987:34. (Type species by original designation: Yaquicoccus bernardinus Taylor 1987.) Distribution.Western North America, including the Snake River basin, California coastal drainages, Baja California, Colorado River basin, Rio Grande basin, Pecos River basin and internal drainages of northern Mxico. Taxonomy.About 125 species are currently recognized, seven in Mxico. Pyrgulopsis acarinatus (Hershler 1985)Nymphophilus acarinatus (shell). Pyrgulopsis acarinata (Hershler). Liu & Hershler 2005; Molecular Phylogeny and Evolution 34:296.Type Locality.A large spring along a dirt road, 10.7 km south of the tip of the Sierra de San Marcos, Cuatro Cinegas Basin, Coahuila, Mxico. Holotype ANSP 355255. Distribution.COAHUILA: known only from the type locality and Santa Tecla Laguna, 22 km south of the tip of the Sierra de San Marcos. Pyrgulopsis brandi (Drake 1953)Amnicola brandi 1.Taylor 1975; Pap. Paleo. Mus. Paleo. Univ. Mich. 10:44. Pyrgulopsis brandi (Drake). Hershler 1994; Smiths. Contrib. (operculum), 33b (radula), 44b (penis).Type Locality.Springs at Las Palomas, Chihuahua, Mxico. Holotype USNM 601494. Distribution.Known only from the type locality. Presumed to be extinct due to drying up of springs at the type locality. Pyrgulopsis cedrosensis (Pilsbry 1927)Paludestrina cedroensis Fontelicella cedroensis (Pilsbry). Gregg & Taylor 1965:108.Taylor 1976:55. Pyrgulopsis cedroensis (Pilsbry). Hershler 1994; Smiths. Contrib. (radula).Type Locality.Bernsteins Spring on east side of Isla de Cedros, Baja California Norte, Mxico. Holotype ANSP 141408. Distribution.BAJA CALIFORNIA NORTE: apparently endemic to springs on Isla de Cedros. Not found in recent years. Pyrgulopis chihuahua (Pilsbry 1928)Fluminicola chihuahua 1964:171. Cochliopa chihuahua (Pilsbry). Pilsbry 1935:92.Taylor 1966; Veliger 9:179.Taylor 1967:153.Taylor 1975:56. Pyrgulopsis chihuahua (Pilsbry). Hershler 1994; Smiths. Contrib. (radula), 45a (penis).Type Locality.Ojo Caliente de Santa Rosa, northcentral Chihuahua (Taylor 1967). Holotype ANSP 141266 (Baker 1964:171). Distribution.CHIHUAHUA: known from the type locality, and from Ojo Caliente de Rosetilla, east of Delicias. Pyrgulopsis manantiali (Hershler 1985)Mexistiobia manantiali Hershler 1985; Malacologia 26:47; (female reproductive system). Pyrgulopsis manantiali (Hershler). Hershler & Thompson 1987:29, Type Locality.A small spring 100 m south of Rio Mesquites at Tierra Blanca, Cuatro Cinegas basin, Coahuila, Mxico. Holotype ANSP 355205. Distribution.COAHUILA: known only from the Cuatro Cinegas basin. Pyrgulopsis minkleyi (Taylor 1966)Nymphophilus minkleyi (nervous system), 10 (radula), 11 (reproductive anatomy).Pyrgulopsis minkleyi (Taylor). Liu & Hershler 2005; Molecular Phylogeny and Evolution 34:296. Type Locality.Ro Mesquites at the main road 9 km southwest of Cuatro Cinegas, Coahuila, Mxico. Holotype UMMZ 220188. Distribution.COAHUILA: endemic to the Cuatro Cinegas basin. Pyrgulopsis palomasensis (Pilsbry 1895)Bythinella palomasensis Pilsbry 1895; Nautilus 9:68.Dall 1897;

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 611916; Nautilus 29:111.Drake 1953; Jour. Wash. Acad. Sci. 43:27.Drake 1956; Bull. South. Calif. Acad. Sci. 55:46.Taylor 1967; Veliger 10:156. Amnicola palomasensis (Pilsbry). Von Martens 1899; Biol. Cent. Amer.:134. Fontelicella palomasensis (Pilsbry). Taylor 1975; Pap. Paleo. Mus. Paleo. Univ. Mich. 10:141. Pyrgulopsis palomasensis (Pilsbry). Hershler 1994; Smiths. Contrib. Type Locality.Laguna de Palomas, 1.6 km south of International Boundary Survey Monument Number 21, Mimbres Valley, Chihuahua (Taylor 1967). Lectotype USNM 130016 (Hershler 1994). Distribution.Known only from the type locality. Pyrgulopsis patzcuarensis Pilsbry 1891Pyrgulopsis patzcuarensis Pilsbry 1891; Nautilus 5:9.Pilsbry 1891; Tryonia patzcuarensis (Pilsbry). Taylor 1966; Veliger 9:197.Type Locality.Lake Patzcuaro, Michoacn, Mxico. Holotype ANSP 61588. Distribution.Known only from the type locality. Possibly extinct. Subfamily status uncertain Genus Rachipteron Thompson 1964Rachipteron Thompson 1964; Rev. Biol. Trop. 12:97.Taylor 1966; Veliger 9:171.Hershler & Thompson 1992; Malac. Rev. Suppl. 5:4. (Type species by monotypy: Rachipteron philopelum Thompson 1964.) of Costa Rica. Taxonomy.The family relationships of the genus remain in question. Taylor (1966) suggested a relationship with the Stenothyridae, an East Asian family of brackish water snails. Higher group relationships cannot be resolved until the female reproductive anatomy is examined. The genus is monotypic. Rachipteron philopelum Thompson 1964Rachipteron philopelum Thompson 1964; Rev. Biol. Trop. 12:98 (penis), 1f (radula), 2 (shell).Type Locality.Seepage pools along the banks of the Rio Grande de Trcoles, about 1/4 mile above the mouth of the river, Costa Rica (09 216478. Distribution.COSTA RICA: known only from brackish to the type locality the species is recorded from Quebrada Bomba Vieja at Boca de Barranca, Prov. Puntarenas Family ASSIMINEIDAE H. Adams & A. Adams 1856 Genus Assiminea Fleming 1828 Type Species.Assiminea grayana Fleming 1828. Distribution.Tropical and sub-tropical regions of much of the world. have been recorded from the New World. Two are brackish water inhabitants. Three occur in freshwater (Hershler, Liu & Lang 2007), of which a single species is found in Mxico. Assiminea cienegensis Hershler, Liu & Lang 2007Assiminea sp. Taylor 1966; Veliger 9:208. Assiminea pecos Taylor 1987; Bull. New Mexico Bur. Mines and Mineral Res. 116:8 (in part). Assiminea cienegensis Hershler, Liu & Lang 2007; Hydrobiologia Type Locality.Southern portion of a spring-marsh complex just W of Mex. Hwy. 30, ca. 2.5 km N of Poza de la Becerra, Cuatro Cienegas Basin, Coahuila, Mxico (ca. Distribution.COAHUILA: known only from the Cuatro Cienegas Basin. Superfamily VALVATOIDEA Gray 1840 Family VALVATIDAE Gray 1840 Distribution.Holarctic. Taxonomy.A single genus is recognized generally. Genus Mller 1774 O. F. Mller 1774:198.Fischer & Crosse 1891:293.Von Martens 1899; Biol. Cent. Amer.:426. (Type species by original designation: Mller 1774.Distribution.Holarctic in distribution. Taxonomy.Thirteen species occur in North America. Two species are known from Mxico. Contreras-Arquiet 1993 Contreras-Arquiet 1993; Publ. Biol., Univ. Auto. Type Locality.Manantiel Charco Azul, San Jos de 860587. Distribution.NUEVO LEN: known only from the type locality, and from Manantiel La Prensa, 2 km N of the type locality. Say 1829 Say 1829; New Harmony Disseminator of Useful (shell).Pilsbry 1903; Proc. Acad. Nat. Sci. Phila. 55:778; pl. 52, fgs. 12, 12a (shell).Hinkley 1907; Nautilus 21:79. Say (humerosa Say). 42 (shell). Fischer & Crosse 1891:304. Fischer & Crosse.Von Martens 1899; Biol. Centr. Amer., 427.Type Locality.Lakes around Mxico City, Distrito Federal, Mxico. Syntypes ANSP 1904 (H. B. Baker 1965). Distribution.MXICO, DISTRITO FEDERAL: Mxico City; Lago de Xochimilco (Pilsbry 1903).

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62 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) humeralis humeralis is found from Montana south to Colorado and west to British Columbia and California (Burch 1989). Henderson (1929) contended that western U.S. records are based on Pilsbry 1908, which most authors subsequent to 1908 recognize as a valid subspecies. Burch (1989) did not list V. h. californica In Mxico, humeralis is known only from the type locality near Ciudad Mxico, and from one locality in SAN LUIS POTOS: Rio Valles near Valles (Hinkley 1907). Von Martens 1899 Say. Pilsbry 1891; Proc. Acad. Nat. Sci. Phila. 43:326. var. pilsbryi Von Martens 1899; Biol. Cent. Amer.:427. var. patzcuaroensis Pilsbry 1899; Proc. Acad. Nat. Sci. Phila. 51:392.Type Locality.Lago de Patzcuaro, Michoacn, Mxico. Holotype ANSP 77192a [H. B. Baker 1965]. Distribution.Known only from the type locality. Subclass PULMONATA Cuvier 1814 Genus Fossaria Westerlund 1885Fossaria Westerlund 1885; Fauna der in der palartischen Region lebenden Binnenconchylien, V:49.Type Species.Buccinum truncatula Mller 1774 Distribution.Europe, Asia, Africa, North America, the West Indies, Central America and South America. Taxonomy.Numerous species. Subgenus Fossaria Westerlund 1885 Distribution.The Holarctic Realm. Taxonomy.Numerous species. One species and two subspecies occurs in the study area. Fossaria obrussa obrussa (Say 1825)Lymnaeus obrussus Say 1825; Jour. Phila. Acad. 5:123. Galba obrussa (Say). F. C. Baker 1911; Chicago Acad. Sci. Spec. Fossaria obrussa (Say). Taylor 1966; Veliger 9:209.Burch 1980; Lymnaea decidiosa (Gould). Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 54:777.Type Locality.Harrowgate, Philadelphia Co., Pennsylvania. Two syntypes ANSP 58700. Distribution.Widely distributed in the Canada and the United States south to Texas and Arizona. A singe record exists for Mxico. COAHUILA: Saltillo (Pilsbry 1904) Fossaria (Fossarria) obrussa modicella (Say 1825)Lymnaeus modicellus Say 1825; Jour. Acad. Nat. Sci. Phila. 5:122. Galba humilis modicella (Say). F. C. Baker 1911; Chicago Acad. 1 (shell). Fossaria (Fossarria) modicella (Say). Bequaert & Miller 1973:190 191. Fossarria obrussa modicella (Say). Burch 1980; Walkerana 1:172; Type Locality.Susquehanna River at Oswego, Tioga Co., New Youk. Syntypes ANSP 58790 (2). Distribution.Widespread in North America south to the Mexican border. BAJA CALIFORNIA NORTE: Sierra Laguna (F. C. Baker 1911). CHIHUAHUA: Laguna Toronto, (Bequaert & Miller 1973). SONORA: 10 mi. N of Imuris, Subgenus Bakerlymnaea Weyrauch 1964Nasonia F. C. Baker 1928; Bull. Wisconsin Geol. Nat. Hist. Survey 70 (1):264 (non Nasonia Ashmead 1904, Hymenoptera). Bakerlymnaea Weyrauch 1964; Arch. fr Moll. 93:169. Type Species.Lymnaea cubensis Pfeiffer 1839. Distribution.Southern United States and the West Indies south to Patagonia. Taxonomy.Numerous species and subspecies are recognized. Three occur in the study area. Fossarria (Bakerlymnaea ) bulimoides techella (Haldeman 1867)Limnaea techella Haldeman 1867; Amer. Jour. Conch. 3:194; pl. 6, Galba bulimoides techella (Haldeman). F. C. Baker 1911; Chicago Stagnicola (Bakerlymnaea ) bulimoides techella (Haldeman). Bequaert & Miller 1971:195. Fossarria (Bakerlymnaea) bulimoides techella (Haldeman). Burch Type Locality.Texas. Syntypes ANSP 59604 (4). Distribution.BAJA CALIFORNIA NORTE: near San Diego Co., California (F. C. Baker 1911). SONORA: pond on hwy. 15, 55 mi. S of Nogales; drift of Rio Sonoyta, Sonoyta; estuary of Rio Mayo at Huatobampito; Rio Yaqui, 4 mi. N of Cd. Obregon (Bequaert & Miller 1973). TAMAULIPAS; (Bequaert & Miller 1973). Fossaria (Bakerlymnaea ) cubensis (Pfeiffer 1839)Limnaea cubensis Pfeiffer 1839; in Wiegm. Arch. 1:354.Strebel 1891; Proc. Acad. Nat. Sci. Phila. 55:320.Von Martens 1899; 379. Lymnaea cubensis (Pfeiffer). Hinkley 1907; Nautilus 21:78. Galba cubensis (Pfeiffer). F. C. Baker 1911; Chicago Acad. Sci. Sp. Fossaria (Bakerlymnaea ) cubensis 387 (shell).Type Locality.Cuba. Distribution.Florida and Texas south to Venezuela; the West Indies. GUATEMALA, Dept. Sacatepquez: Antigua. BAJA CALIFORNIA NORTE: Sanzal, Bahia Todos Santo (F. C. Baker 1911). SAN LUIS POTOS: El Abra; Cd. Victoria

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 63 (Hinkley 1907). VERACRUZ: Cd. Veracruz; Rio Tenoya; Orizaba (Von Martens 1899). Fossaria (Bakerlymnaea ) (Orbigny 1835) Orbigny 1835; Magazin de Zoologie 1835:24. Orbigny. Pilsbry 1911; Princeton Univ. Exped. to Hubendick 1951.Knglica Svenka Vetenskapsakademiens Orbigny 1835; Voyage dans LAmrique Limnaeus Orbigny). (Orbigny). Paraense 1982; Memorias do Instituto Oswaldo Cruz, 77:181. (Orbigny). Rangel-Ruiz 1994; Walkerana 7:29 28:71.Type Locality.Restricted by Pilsbry to banks of the Rio Negro, 7 or 8 leagues above the mouth, 41 S lat., Argentina. Distribution.Mxico and Cuba south to Chile and Argentina (Paraense 1982). The identity of Mexican CAMPECHE: Ciudad de Carmen. JALISCO: Lago de Chapala. TABASCO: Tecotlapa; Teapa. VERACRUZ: Jalapa; Teoloyucn. Genus Pseudosuccinea F. C. Baker 1908Pseudosuccinea F. C. Baker 1908; Science, n. s., 27:943.Type Species.Lymnaea columella Say 1817. Distribution.North America south to Panam, and the West Indies. Taxonomy.A single species, Pseudosuccinea columlla is recognized in North and Central America. Pseudosuccinea fransisca (Poey 1858) is a Cuban species. Pseudosuccinea columella (Say 1817)Lymnaea columella Say 1917; Jour. Acad. Nat. Sci., 1:44. Pseudosuccinea columella (Say 1817). F. C. Baker 1911; Sp. Pub. Type Locality.Not stated. Distribution.The typical form is found from Nova Scotia west to Manitoba, south to Florida and Texas. It has been widely introduced elsewhere (Hubendick 1951:52). Taxonomy.F. C. Baker (1911) listed four subspecies. He reported only Pseudosuccinea columella championi from the study area. Pseudosuccinea columella championi (Von Martens 1899)Limnaea columella var. championi Von Martens 1899; Biol. Cent. Pseudosuccinea columella championi (Von Martens). F. C. Baker (shell).Type Locality.Bigabo, Panam. Distribution.Central Mxico south to Panam. MXICO: Cd. Mxico. MICHOACN: lake near Uruapan. NAYARIT: Tepc. Remarks.The taxonomic status and geographic distribution of this subspecies need to be investigated. Genus Stagnicola Leach 1830Stagnicola Leach, in Jeffreys, 1830; Trans. Linn. Soc. 16:370.Type Species.Buccinum paluistris Mller 1774. Distribution.Circumboreal; North America south to central Mxico. Taxonomy.Two subgenera are recognized in the Nearctic Region. One occurs in the study area. The number of species in Stagnicola is debatable. Stagnicola elodes is the only recognized species in the study area. Lymnaeus attenuatus is considered a synonym, but further study may reveal it to be a distinct species. Stagnicola elodes (Say 1821)Lymnaeus elodes Say 1821; Jour. Acad. Nat. Sci. Phila. 2:169. Galba elodes (Say). F. C. Baker 1911; Chicago Acad. Sci. Sp. Pub. (shell). Stagnicola elodes 606, 611 (shell). Lymnaea elodes (Say). Paraense 1994; Malacological Review (digestive system), 12 (pallial organs), 13 (radula). Lymnaeus decidiosus (Say). Pilsbry 1904: Proc. Acad. Nat. Sci. Phila. 55:777. Lymnaeus attenuatus Say 1829; New Harmony Dis., II:244. Lymnaea attenuata Fischer & Crosse 1880:49.Von Martens 1899:375.Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:776. Galba attenuata (Say). F. C. Baker 1911; Chicago Acad. Sci. Spec. PuType Locality.Lymnaeus attenuatus : ditches and ponds in the vicinity of Mxico City, Mxico. Syntypes in the ANSP. Distribution.Interior basin of Canada south to new England, California and New Mexico with outlying populations in central Mxico. COAHUILA: Saltillo (Pilsbry 1904). HIDALGO: Zimapan (Von Martens 1899). MXICO: Tlalpam (Pilsbry 1904). Lago de Chalco (Von Martens 1899). MORELOS: Laguna de Quila, near Zempoala (Paraense 1994). Subfamily FERRISSIINAE Walker 1917 Genus Ferrissia Walker 1908Ferrissia Walker 1908; Nautilus 17:15.Basch 1963; Bull. Mus. Comp. Zool. 129:126.Type Species. Say 1817. Distribution.North America and the West Indies south to Panam. Taxonomy.Numerous species have been described. Basch (1963) placed many of these in synonymy. The taxonomic status of the Middle American species remains to

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64 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) be determined. Two species are listed for the study area. Ferrissia bayacalifornica Walker 1924Ferrissia bayacalifornica Walker 1924; Jour. Wash. Acad. Sci. Ferrissia walkeri (Pilsbry & Ferriss). Basch 1963; Bull. Mus. Comp. Zool. 129:433.Type Locality.A small pool 2.5 miles inland from San Jos del Cabo, Baja California Sur, Mxico. Holotype in the UMMZ. Distribution .Known only from the type locality. Ferrissia occidentalis Walker 1924Ferrissia occidentalis Walker 1924; Jour. Wash. Acad. Sci. 14:431; Ferrissia walkeri (Pilsbry & Ferriss). Basch 1963; Bull. Mus. Comp. Zool. 129:433.Type Locality.A small pool 2.5 miles inland from San Jos del Cabo, Baja California Sur, Mxico. Holotype in the UMMZ. Distribution .Known only from the type locality. Genus Gundlachia Pfeiffer 1849Gundlachia Pfeiffer 1849; Zeitsch. fr Malak. 6:98.Wurtz 1951; Nautilus 64:126.Type Species.Gundlachia ancyliformis Pfeiffer 1849. Distribution.North America south to Guatemala and Honduras. Taxonomy.Wurtz (1951) recognized seven species. Gundlachia hinkleyi Walker 1917Gundlachia hinkleyi 34:38, 40.Wurtz 1951; Nautilus 64:126.Type Locality.Maya Farms, Quirigua, Dept. Izabal, Guatemala. Holotype UMMZ 43455. Distribution.GUATEMALA, Dept. Guatemala: Lago de Amatitlan (Hinkley 1920). Dept. Izabal: Quirigua. Gundlachia hjalmarsoni Pfeiffer 1858Gundlachia hjalmarsoni Pfeiffer 1859; Malak. Bltt. 5:197.Von Martens 1899:403.Wurtz 1951; Nautilus 64:126.Type Locality.Santa Rosa, Honduras. Santa Rosa is the name of a municipality in both the Honduran departments Distribution.Known only from the type locality. Genus Hebetancylus Pilsbry 1914Hebetancylus Pilsbry 1914; Proc. Acad. Nat. Sci. Phila. 65:671.Wurtz 1951; Nautilus 64:126.Basch 1963; Bull. Mus. Comp. Zool. 129:422.Type Species.Ancylus moricandi Orbigny 1837. Distribution.Cuba and Hispaniola in the West Indies; Florida and Texas south to Brazil and Paraguay. Taxonomy.Wurtz lists eight species in the genus. Two occur in the study area. Hebetancylus excentricus (Morelet 1851)Ancylus excentricus Morelet 1851; Test. Noviss. II:17.Fischer & Proc. Acad. Nat. Sci. Phila. 43:3.Von Martens 1899:402 403.Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:777. Velletia excentrica (Morelet). Tate 1970; Amer. Jour. Conch. 5:158. (Morelet). Hinkley 1920; Nautilus 34:39, 40.Walker 1923; The Ancylidae of South Africa:10.Bequaert & Clench 1936; Pub. Carnegie Inst. Wash. (457):70. Ferrissia excentrica (Morelet). Wurtz 1951; Nautilus 64:125.Solem 1954; Nautilus 68:6. Gundlachia excentrica (Morelet). Taylor 1966; Veliger 9:206. Hebetancylus excentricus (Morelet). Basch 1963; Bull. Mus. Comp. Ancylus excentricus var. biolleyi Von Martens 1899; Biol. Cent. Amer.:402. Ancylus excentricus var. pittieri Von Martens 1899; Biol. Cent. Amer.:402.Type Locality.Lago de Itza [Petn], Dept. Petn, Guatemala. Distribution.Florida and Texas south to Costa Rica. COSTA RICA, Prov. San Jos: Rio Torres; San Jos (Von Martens 1899). Prov. Puntarenas: Rio de Las Platanales (Von Martens 1899). NICARAGUA, Dept. Chontales: San Nicolas, San Agustin (Tate 1870). GUATEMALA, Dept. Guatemala: Lago de Amatitlan (Hinkley 1920). Dept. Izabal: Quirigua (Hinkley 1920). Dept. Petn: Lago de Itza. COAHUILA: Rio Salado de los Nadadores, at El Cario de la Montaa, 20 mi. E of Cuatro Cienegas (Taylor 1966). DURANGO: Laguna del Progreso (Solem 1944). MICHOACN: pressa near Uruapan (Pilsbry 1903). YUCATN: Shkolak (Pilsbry 1891); Xanaba Cenote Grande, 8 mi. SW of Chichen Itza; Chotch Cenote 2 mi. NE of Pista (Bequaret & Clench 1936). Wurtz 1951 Wurtz 1951; Nautilus 64:127; pl. Colombia. Holotype ANSP 186643a. Distribution.Known only from the from type locality. Genus Walker 1908 Walker 1908; Nautilus 1908, 17:15.Wurtz 1951; Nautilus 64:125.Basch 1963; Bull. Mus. Comp. Zool. 129:418.Type Species.Ancylus fuscus C. B. Adams 1841. Distribution.North America and the West Indies south to Panam. Taxonomy.Approximately ten species are recognized in the literature. Four occur in the study area. (Goodrich & van der Schalie 1937)Ferrissia () aguadae Goodrich & van der Schalie 1937; 7a (shell). Libertad, Dept. Petn, Guatemala. Holotype in the UMMZ.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 65 Distribution.GUATEMALA: Goodrich & van der localities and the type locality. (Morrison 1946)Ferrissia () joseana Morrison 1946; Smiths. Misc. Coll. Type Locality.Flood-plain pools along the middle reaches of a stream opening into the northwest mangrove swamp on Isla San Jos, Archipelago de Las Perlas, Panam. Holotype USNM 542179. Distribution.Known only from the type locality. (Von Martens 1899)Ancylus papillaris Von Martens 1899; Biol. Cent. Amer.:402; pl. 21, Sci. Phila. 55:777.Wurtz 1951; Nautilus 64:130.Type Locality.Rio Ameca, Jalisco, Mxico. Distribution.JALISCO: Rio Ameca. MXICO: Tlalpam. MICHOACN: Uruapan. NUEVO LEN: Monterey (Pilsbry 1904). (Bourguignat 1857)Ancylus sallei Bourguignat 1857; Amnits Malac., ii:32.Bourguignat 1857; Rev. et Mag. Zool. 1857:16.Strebel Veracruz, Mxico. Distribution.VERACRUZ: Veracruz (Strebel 1873). Genus Uncancylus Pilsbry 1914Uncancylus Pilsbry 1914; Proc. Acad. Nat. Sci. Phila. 65:671.Wurtz 1951; Nautilus 64:128.Type Species.Ancylus barilensis Moricand 1845. Distribution.Guadeloupe, Jamaica, Costa Rica south to Brazil and Argentina. Taxonomy.Wurtz (1951) lists seven species and one subspecies. Two occur in the study area. Uncancylus ameliae Pilsbry 1920Uncancylus ameliae Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 72:9; ANSP 105260. Distribution.Known only from the type locality. Pilsbry 1920 Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 72:7 Vias, Prov. Cartago, Costa Rica; 2500 ft. alt. Holotype ANSP 105277. Distribution.Known only from the type locality. Undetermined species of Ancylidae Ancylus haldemani Boettger 1853Ancylus depressus Boettger 1844; Monogr.:4 (non Ancylus depressus Deshayes 1824). Ancylus haldemani Boettger 1853; Jour. de Conchyl. 4:180.Binney Wurtz 1951; Nautilus 64:130.Type Locality.Mxico. Distribution.Unknown. Ancylus sallei Boettger 1857Ancylus sallei Boettger 1857; Rev. Mag. Zool., 9:16.Wurtz 1951; Nautilus 64:131.Type Locality.Mxico and Cuba. Distribution.Unknown. Genus Gyraulus Charpentier 1837Gyraulus Charpentier 1837; Mem. Soc. Helv. De Sci. Natur., 1:21.Type Species.Planorbis hispidus Draparnaud (= Planorbis albus O. F. Mller 1774). Distribution.Europe, Africa, Asia, the Philippines, Australia, Tasmania, New Caledonia, Fiji and North America. Taxonomy.Four subgenera are recognized (Zilch 1959). One occurs in the study area. Subgenus Torquis Dall 1905Torquis Dall 1905; Harriman Alaska Exp., XIII:83, 86.Type Species. Say 1817. Distribution.North America, northern Asia, Europe. Taxonomy.Three species occur in North America (Burch 1980). A single species occurs in the study area. A second species is placed here provisionally. Gyraulus (Torquis)(Say 1817) 5 (shell).Pilsbry 1891; Proc. Acad. Nat. Sci. Phila. 43:322. Planorbis (Gyraulus) Say. Von Martens 1899:394. (Say). Branson, McCoy & Sisk 1964; Southwestern Natur. 9:103.Bequaert & Miller 1973:204.Burch Type Locality.Delaware River, eastern North America. Distribution.North America from Alaska and northern Canada to the Greater Antilles and south in Mxico at least to Morelos. MORELOS: Yautepec (Pilsbry 1891). SONORA: Rio Nacuzari, 7 mi. S of Nacuzari (Branson et al. 1964). Gyraulus percarinatus Paraense 2000Gyraulus percarinatus Type Locality.Grounds of the Faculty of Agronomy, Panam University of Tocumen, Prov. Panam, Panam. No type specimens designated. Distribution.Known only from the type locality.

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66 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Taxonomy.The name Gyraulus percarinatus Paraense 2000 is invalid, because the description does not include the designation of a holotype as required by ICZN Articles 16.4 and 62.4. Subfamilly DREPANOTREMINAE Zilch 1959 Distribution.American subtropics and tropics from Texas and the Greater Antilles south to Argentina. Taxonomy.Three genera are recognized, Antillorbis, Drepanotrema, and a Brazilian genus, Acrorbis Ohdner 1937. Genus Antillorbis Harry & Hubendick 1964Antillorbis Harry & Hubendick 1964; Geborgs Kungl. Vetenskapsoch Vitterhets-samhlles handlingar, ser. 6, ser. B:29.Type Species.Planorbis circumlineatus Shuttleworth 1854 (= Planorbis aerugnosus Morelet 1851). Distribution.The west Indies, Mxico and Central America. Taxonomy.A single species is recognized. Antillorbis aeruginosus (Morelet 1851)Planorbis aeruginosus Morelet 1851; Test. Noviss. II:15. Gyraulus aeruginosus (Morelet). Goodrich & van der Schalie 1937; Misc. Pub. Mus. Zool. Univ. Mich. (34):33. Antillorbis feruginosus (Morelet). Harry & Hubendick 1964; Geborgs Kungl. Vetenskaps-och Vitterhets-samhlles Drepanotrema (Antillorbis ) aeruguinosus (Morelet). Bequaert & 710 (shell). Planorbis salleanus Dunker 1853; Proc. Zool. Soc. Lond. 54. Planorbis circumlineatus Shuttleworth 1854; Ann. Lyc. Nat. Hist. N.Y. 6:72.Shuttleworth 1854; Mill. Naturf. Ges. Bern 1854:96. Planorbaria circumlineatus (Shuttleworth). Neubert & Gosteli Planorbis lineatus sonorensis Cooper 1893; Proc. Calif. Acad. Sci. Pilsbry & Ferriss 1906; Proc. Acad. Nat. Sci. Planorbis von Sandberger 1875). Planorbis arizonensis Pilsbry & Ferriss 1915; Proc. Acad. Nat. Sci. Phila. 67:390 (replacement name for P. & F.). Gyraulus arizonensis (Pilsbry & Ferriss). Richards 1937; Trans. Amer. Philos. Soc. 77:255. Planorbis santacruzensis Germain 1932; Rec. Indian Museum Type Localities. Planorbis aeruginosis: Lago de Izabal, Guatemala. Planorbis salleans: Santo Domingo (= Hispaniola). Planorbis circumlineatus: Humacoa, Puerto Rico. Syntypes Naturhistorische Museum Bern 18952/16 (Neubert & Gosteli 2003:17). Planorbis lineatus sonorensis: San Miguel, Sonora, Mxico. : drift of San Pedro River, Benson, Cochis Co., Arizona. Planorbis santacruzensis: St. Croix. Distribution.West Indies, southern Texas and Arizona south to Panam. GUATEMALA, Dept. Guatemala: outlet of Lago Amatitlan, near Cd. Guatemala (Goodrich & van der Schalie 1937). Dept. Izabal: Lago de Izabal (Morelet 1851). QUINTANA ROO: Isla Cozumel, swamp near San Miguel; swamp near San Gerbacio (Richards 1937). SONORA: San Miguel (Cooper 1893). Genus Drepanotrema Fischer & Crosse 1880Drepanotrema Fischer & Crosse 1880:59, 75.H. B. Baker 1930; Occ. Pap. Mus. Zool. Univ. Mich. (210):48.Pilsbry 1934; Proc. Acad. Nat. Sci. Phila. 86:58.Harry & Hubendick 1964; Meddel. Gotenb. Mus. Zool. Avdel. 136:26.Type Species.Planorbis yzabelensis Crosse & Fischer 1879 (= Planorbis anatinus Orbigny 1835). Distribution.Tropical America from Texas and the west Indies south to Argentina. Taxonomy.Two subgenera are recognized. Subgenus Drepanotrema Fischer & Crosse 1880 Distribution.Same as for the genus. Taxonomy.F. C. Baker (1945:118) listed eight species. Two occur in the study area. Drepanotrema (Drepanotrema) anatinum (Orbigny 1835)Planorbis anatinus Orbigny 1835; Mag. de Zool. 5 (62):28. Drepanotrema anatinum (Orbigny). Aguayo 1933; Nautilus 47:64 Hubendick 1964; Meddel. Gotenb. Mus. Zool. Avdel. 136:21, Planorbis haldemani C. B. Adams 1849; Contrib. to Conch.:43. Planorbis caracasnsis Gundlach 1857; Malak Bltt. 4:179. Planorbis esperanzensis Tryon 1866; Amer. Jour. Conch. 2:10; pl. Planorbis yzabalensis Crosse & Fischer 1879; Jour. de Conchyl. Planorbis (Hippeutus) yzabalensis (Crosse & Fischer). Von Martens 1899; Biol. Cent. Amer.:397. Anisus (Gyraulus) lenzi Benthem Jutting 1943; Archiv fr Hydrobiologie 39:460.Type Locality.Planorbis anatinus: Rio Parana, Argentina; syntype in the BMNH (Aguayo 1933). Planorbis haldemani: Jamaica. Planorbis esperanzensis: Esperanza, Cuba. Planorbis aracasnsis: Trinidad, Cuba. Planorbis yzabalensis: Lago de Izabal, Guatemala. Anisus (Gyraulus) lenzi: Rio So Francisco, San Pedro Dias-Buchr, Pernambuco, Brazil. Distribution.Widely distributed throughout the West Indies and from Texas south to Argentina and Brazil. PANAM, Canal Zone: Gamboa (Aguayo 1933). Prov. Panam: ditch at the National University, Tocumn (Paraense 2000). COSTA RICA, Prov. Guanacaste: Catalina (Paraense 2000). NICARAGUA, Dept. Rivas: Rivas (Paraense 2000). GUATEMALA, Dept. Izabal: Lago de Izabal, Mariscos; El Prado (Paraense 2000). Dept. Santa Rosa: Rio Helado,

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 67 Taxisco. BELIZE: Belize Dist.: Boston Village (Paraense 2000). TABASCO: Rio Usumacinta; Balancan (Von Martens 1899). Drepanotrema (Drepanotrema) lucidum (Pfeiffer 1839)Planorbis lucidus Pfeiffer 1839; in Weigmanns, Arch. fr Naturgesch.:354. Planorbis (Dreopanotrema) lucidus Pfeiffer. Bequaert & Clench 1936; Pub. Canegie Inst. Wash. (457):67. Drepanotrema lucidum (Pfeiffer). H. B. Baker 1930; Occ. Pap. 1 (shell).Harry & Hubendick 1964; Meddel. Gotenb. Mus. Type Locality.Cuba. Distribution.The West Indies and Mxico south to Brazil and Peru. NICARAGUA, Dept. Grenada: Grenada. Dept. Rivas: Rivas (Paraense 2000). GUATEMALA, Dept. Izabal: Mariscos (Paraense 2000). BELIZE: Belize Dist.: pond by road to International Airport (Paraense 2000). YUCATN: Aguado Halal near Merida; aguado 14 km N, 2 km E of Merida; Aguado Daadz, 6.5 mi. SW of Chichen Itza (Bequaert & Clench 1936). Subgenus Fossulorbis Pilsbry 1934Fossulorbis Pilsbry 1934; Proc. Acad. Nat. Sci. Phila. 86:59.F. C. Baker 1945; Moll. Fam. Planorbidae:118.Type Species.Planorbis cultratus Orbigny 1841. Distribution.The West Indies and southern Texas south to Argentina. Taxonomy.Six species and four subspecies occur in the study area. Drepanotrema (Fossulorbis) cimex (Moricand 1839)Planorbis cimex Moricand 1839; Mm. Soc. Phys. Hist. Nat., Drepanotrema (Fossulorbis) cimex (Moricand). F. C. Baker 1945; Type Locality.Bahia, Brazil. Distribution.Known from southern Texas, Mxico, Central America, the West Indies, Venezuela and Brazil. Drepanotrema (Fossulorbis) cultratum cultratum (Orbigny 1841)Planorbis cultratus Orbigny 1841; in Sagra, Hist. Fis. Polit. y nat. Cent. Amer.:395.Hinkley 1920; Nautilus 34:38. Planorbis (Spiralina) cultratus Orbigny. Bequaert & Clench 1936; Pub. Canegie Inst. Wash. (457):67. Drepanotrema cultratum (Orbigny). H. B. Baker 1930; Occ. Pap. (shell).Richards 1937; Trans. Amer. Philos. Soc. 77:255.F. C. Baker 1945; Moll. Fam. Planorbidae:118, 388, 392; pl. 76, Type Locality.Martinique (Pilsbry 1934:60). Distribution.The Lesser Antilles, Venezuela and Colombia. NICARAGUA, Dept. Chontales: San Agustin, near Acoyapa (Von Martens 1988). GUATEMALA, Dept. Guatemala: ditch along railroad at Laguna, Lago Amatitlan (Hinkley 1920). QUINTANA ROO: swamp a few miles N of San Miguel (Richards 1937). YUCATN: Xanaba Cenote Grande, 8 km SW of Chichen Itza (Bequaert & Clench 1936). Drepanotrema (Fossulorbis) cultratum anitense (Cooper 1893)Planorbis (Anisus) anitensis Cooper 1893; Proc. Calif. Acad. Sci. Drepanotrema cultratum anitense (Cooper). F. C. Baker 1945; Moll. Type Locality.Santa Anita, Baja California Sur, Mxico. Distribution.BAJA CALIFORNIA SUR: lagoon, Santa Rita (F. C. Bater 1945). Drepanotrema (Fossulorbis) cultratum duenasianum (Tristram 1861)Planorbis dueasensis Tristram 1861; Proc. Zool. Soc. Lond. 29:232. Drepanotrema cultratum dueasianum (Tristram). Pilsbry 1934; 9 (shell).Type Locality.Lago de Dueas, Dept. Sacatepquez, Guatemala. Distribution.GUATEMALA, Dept. Guatemala: pools along railroad near Lago de Amatitlan (Pilsbry 1934). Dept. Sacatepquez: Lago de Dueas. Remarks.Spelling of the name of the subspecies dueasianum is corrected to duenasianum as mandated by Article 35.2.1 of the ICZN. Drepanotrema ( Fossulorbis) cultratum labrosum Pilsbry 1934Drepanotrema cultratum labrosum Pilsbry 1934; Proc. Acad. Nat. Type Locality.Brownsvile, Texas. Holotype ANSP 143416. Distribution.Southern Texas south to southern Tamaulipas. TAMAULIPAS: drift debris of the Rio Panuco (Pilsbry 1934). Drepanotrema ( Fossulorbis) cultratum panuco Pilsbry 1934Drepanotrema cultratum panuco Pilsbry 1934; Proc. Acad. Nat. Sci.

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68 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Type Locality.Pasture west of San Dieguito, San Luis Potosi, Mxico. Holotype ANSP 107619. Distribution.SAN LUIS POTOS: San Dieguito. TAMAULIPAS: Tampico (Pilsbry 1934). Drepanotrema ( Fossulorbis) depressissimum (Moricand 1839)Planorbis depressissimus Moricand 1839; Mm. Soc. Phys. Hist. Drepanotrema depressissimum (Moricand). F. C. Baker 1945; Moll. (reproductive anatomy).Type Locality.Bahia, Brazil. Distribution.Widely distributed in South America, Central America and the West Indies (Paraense 2000). COSTA RICA, Prov. Guanacaste: Catalina (Paraense 2000). NICARAGUA, Dept. Chontales: Acoyapa. Dept. Rivas: Rivas (Paraense 2000). Drepanotrema (Fossulorbis) kermatoides (Orbigny 1835)Planorbis kermatoides Orbigny 1835; Mag. de Zool., (62):27. Drepanotrema (Fossulorbis) kermatoides (Orbigny). F. C. Baker Harry & Hubendick 1964; Meddel. Gotenb. Mus. Zool. Avdel. Soc. Wash. 79:280.Type Locality.Providence of Lima, Peru. Distribution.Lesser Antilles, Texas, Mxico, Central America, Venezuela, Peru and Brazil (Harry & Hubendick 1964). QUINTANA ROO: Isla Cozumel (Rehder 1966). Taxonomy.Paraense (2000) suggested that records for Drepanotrema kermaoides from Mxico and Central America may pertain to D. depressissimum. Drepanotrema (Fossulorbis) sumichrasti (Crosse & Fischer 1879)Planorbis sumichrasti Crosse & Fischer 1879; Jour. de Conchyl. Von Martens 1899; Biol. Cent. Amer.:396. Drepanotrema sumichrasti (Crosse & Fischer). F. C. Baker 1945; Type Locality.Cacoprieto, Isthmus of Tehuantepec, Oaxaca, Mxico. Distribution.PANAM, Canal Zone: Isla Barro Colorado (F. C. Baker 1945). OAXACA: Cacoprieto. Drepanotrema (Fossulorbis) surinamense (Clessin 1884)Planorbis surinamensis Clessin 1884; in Martini & Chemnitz; System. Conch. Cab., Ed. 2, Planorbis Drepanotrema surinamense (Clessin). Paraense 2000; Malac. Rev. Type Locality.Surinam. Distribution.PANAM, Prov. Panam: Juan Diaz, ca. 6 km NE of Ciudad Panam; Las Cumbres; Hippodrome RICA: Prov. Guanacaste: Catalina (Paraense 2000). Subfamily BIOMPHALARIINAE H. Watson 1954 Genus Biomphalaria Preston 1910Biomphalaria Preston 1910; Ann. Mag. Nat. Hist. ser. 8, 6:533.Paraense & Deslandes 1958; Rev. Brasil. Biol. 18:65.IZCN Opinion 735 1965. Tropicorbis Pilsbry & Brown 1914; Proc. Acad. Nat. Sci. Phila. 66:212. Platytaphius Pilsbry 1924; Proc. Acad. Nat. Sci. Phila. 78: Australorbis Pilsbry 1934; Proc. Acad. Nat. Sci. Phila. 86:55. Obstructio Haas 1939; Field Mus. Nat. Hist. Zool. Ser., 24:100. Lateorbis F. C. Baker 1945; Moll. Fam. Planorbidae:85.Type Species.Biomphalaria: Biomphalaria smithi Preston 1910. Platytaphius : Planorbis hertopleurus Pilsbry & Vanatta 1896. Tropicorbis: Planorbis liebmanni Philippi 1850 (= Planorbis maya Morelet 1849; = Planorbis Pfeiffer 1839, Paraense & Deslandes 1958). Australorbis: Planorbis glabratus Say 1818. Obstructio: Planorbis janeirensis Clesson 1884. Lateorbis: Planorbis pallidus C. B. Adams 1846. Distribution.Africa, Tropical America. Taxonomy.The systematics of this genus is very unsettled at present. There is a large number of species in Africa and tropical America. The genus has received a great deal of study because of its importance as intermediate hosts for human schistosome parasites. However, many species not directly implicated as schistosome hosts remain poorly studied. Division of Biomphalaria into subgenera is not advisable at present. Biomphalaria albicans (Pfeiffer 1848)Planorbis albicans Pfeiffer 1839; Weigmanns Arch fr Naturgesch.:354. Taphius albicans (Pfeiffer). Harry & Hubendick 1964; Geborgs Kungl. Vetenskaps-och Vitterhets-samhlles handlingar, ser. 6, Biomphalaria albicans (Pfeiffer). Abdel-Malek 1969; Malacologia 18 (shell).Type Locality.Cuba. Distribution.Reported from the Greater Antilles, some Lesser Antilles, Louisiana, Texas, Mxico and Central America (Harry & Hubendick 1964). COSTA RICA, Prov. Cartago: near Turrialba (Abdel-Malek 1969). BELIZE: Belize Dist.: Central Farms (Abdel-Malek 1969). Taxonomy.Paraense (1996a) treated this species as a synonym of Biomphalaria helophila (Orbigny 1835). Biomphalaria belizensis (Crosse & Fischer 1878)Planorbis belizensis Crosse & Fischer 1879; Jour. de Conchyl. (shell).Von Martens 1899; Biol. Cent. Amer.:390. Biomphalaria belizensis (Crosse & Fischer 1878).Type Locality.Belize. Distribution.GUATEMALA, Dept. Alta Verapaz: Cobn; San Miguel Tucar (Fischer & Crosse 1880).

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 69 BELIZE. TABASCO (Von Martens 1899). Biomphalaria boucardianus (Preston 1907)Planorbis boucardianus Preston 1907; Ann. & Mag. Nat. Hist. ser. Tropicorbis boucardianus (Preston). F. C. Baker 1945; Moll. Fam. Type Locality.Mxico. Distribution.NARARIT: Manzanillo (F. C. Baker 1945). (Tryon 1863) Tryon 1863; Proc. Acad. Nat. Sci. Phila. 15:281; pl. Planorbis (Gyraulus) Tryon. Von Martens 1899; Biol. Cent. (Tryon). F. C. Baker 1945; Moll. Fam. (Tryon). Abdel-Malek 1969; Malacologia Type Locality.Panam. Distribution.PANAM, Canal Zone: Gatn (F. C. Baker 1945; Abdel-Malek 1969). NICARAGUA, Dept. Grenada: Lago de Nicaragua, Grenada (Tate 1870). Biomphalaria gracilenta (Gould 1855)Planorbis gracilentus Gould 1855; Proc. Boston Soc. Nat. Hist. 5:129. Tropicorbis (Tropicorbis) gracilentus (Gould). F. C. Bakeer 1945; Type Locality.Great Colorado Desert lowlands, United States. Holotype USNM 26477 (F. C. Baker 1945:496). Distribution.BAJA CALIFORNIA SUR: 30 mi. SW of Santiago, Mxico (F. C. Baker 1945). (Pfeiffer 1839) Pfeiffer 1839; Arch. Naturgesch., 5:354. (Pfeiffer). F. C. Baker 1945; Moll. Fam. (Pfeffer). Paraense & Deslandes 1958; Rev. (Pfeiffer). Bequaert & Miller 1973:205 155 (shell).Yung, Pointier & Parera 1997; Malac. Rev. 3 (shell). Planorbis dicipiens C. B. Adams 1849; Contr. Conch. 3:43. Planorbis maya Morelet 1849; Test. Noviss. I:16.Pilsbry 1891; Proc. Acad. Nat. Sci. Phila. 43:322.Bequaert & Clench 1933; Pub. Carnegie Inst. Wash. (431):538. Planorbis (Gyrorbis) maya Morelet. Fischer & Crosse 1880:72; Planorbis (Gyraulus) maya Morelet. Von Martens 1899; Biol. Cent. Planorbis (Tropicorbis) maya (Morelet). Bequaert & Clench 1936; Pub. Canegie Inst. Wash. (457):66. Tropicorbis (Tropicorbis) maya (Morelet). F. C. Baker 1945; Moll. Taphius maya (Morelet). Paraense & Deslandes 1957a; Jour. de Planorbis liebmanni Philippi 1850; in Dunker, Martini & Chemnitz, 1891; Proc. Acad. Nat. Sci. Phila. 43:322.Pilsbry 1903; Proc. Acad. Nat. Sci. Phila. 55:777.Pilsbry 1928; Proc. Acad. Nat. Sci. Phila. 80:117. Tropicorbis liebmanni (Philippi). Paraense & Deslandes 1957b: Rev. Brasil. Biol. 17:427.Paraense 1958; Rev. Brasil. Type Localities. : Havana, Cuba (Yung et al. 1997). Planorbis decipiens : Jamaica. Planorbis maya: in cisterns, Ciudad Campeche, Campeche. Planorbis liebmanni : Veracruz, Mxico. Distribution.West Indies, Mxico and Central America; South American distribution remains to be determined. EL SALVADOR, Dept. San Salvador: Cd. San Salvador (Paraense 2000). GUATEMALA, Dept. Sacatepquez: Antigua (Von Martens 1899); Lago de Dueas (Paraense 2000). BELIZE: Belize Dist.: Boston Village (Paraense 2000). CAMPECHE: Isla de Carmen; Campeche; lake at km 49, road from Cd. Campeche to Merida (Paraense & Delandes 1957b). NUEVO LEN: Topo Chico, near Monterey (Pilsbry 1903). OAXACA: Isthmus of Tehuantepec. TAMAULIPAS: Mesa de Solis, near La Lajilla, between Padilla and Jimenez (Pilsbry 1928). VERACRUZ. YUCATN: Merida (Pilsbry 1891); Tabi (Von Martens 1899); cave at Actun Has; hacienda Yocat; Ticul District (Bequaert & Clench 1933); aguado 14 km N, 2 km E of Merida; Aguado Halal, near Merida; pool 3 km S of Progreso (Bequaert & Clench 1936). Taxonomy.Paraense and Deslandes (1957b) synonymized Planorbis liebmanni (Philippi 1850) with Planorbis maya (Morelet 1849), and in turn (1958) they synonymized Planorbis maya with (Pfeiffer 1839). Records of from Florida, Missisippi, Louisiana and Texas (Abdel-Malek 1969) require Biomphalaria helophila (Orbigny 1835)Planorbis helophilus Orbigny 1835; Mag. de Zool., (62):27. Biomphalaria helophila (Orbigny).Ruz 1987; Universidad y Ciencias, 4:33; map 1; pl. 1 (shell); pl. 2 (radula); pl. 3 (reproductive anatomy).Paraense 2000; Malac. Rev.; 33:4; Australorbis albicans (Pfeiffer 1839). Paraense and Deslandes anatomy). Planorbis dentiens Morelet 1849; Test. Noviss. I:18.Fischer & Planorbis dentiens var. edentula Fischer & Crosse 180:81; pl. 34, Tate 1870: Amer. Jour. of Conch. 5:159 (not Braun 1848).

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70 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Tropicorbis tatei Baker 1941; Nautilus 54:96 (replacement name for Tate 1870). Tropicorbis shimeki F. C. Baker 1945; Moll. Fam. Type Localities. Planorbis helophila : Lima, Peru. Planorbis cannarum: Belize. Planorbis dentiens : Belize. Tropicorbis shimeki: Ometope [Isla de Ometepe], Nicaragua; holotype USNM 534290. Distribution.EL SALVADOR, Dept. Santa Ana: Lago de Coatepeque (Paraense 2000). GUATEMALA, Dept. Izabal: El Prodo (Paraense 2000). NICARAGUA, Isla de Ometepe (F. C. Baker 1945). Dept. Chontales : Acoyapa (Paraense 2000). VERACRUZ: Coatzacoalcos (F. C. Baker 1945). Los Tuxtlas, Laguna Escondida and Laguna Zacatal Biomphalaria hondurasensis (Clesson 1878)Planorbis hondurasensis Clesson 1878; in Martini & Chemnitz, (shell).Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 72:6. Planorbis (Gyraulus) hondurasensis Clesson. Von Martens 1899; Biol. Cent. Amer.:393.Type Locality.Santa Rosa, Honduras. Distribution.HONDURAS: Santa Rosa. COSTA RICA: Rio Torres (Von Martens 1899). Prov. San Jos: ditch along road from San Jos to La Verbena. Biomphalaria kuhniana (Clessin 1885)Planorbis kuhnerianus Clessen 1883; in Martini & Chemnitz, Syst. Conch. Cab., ed. 2, Planorbis Planorbis kuhniana Clessen 1885; in Martini & Chemnitz, Syst. Conch. Cab., ed. 2, Planorbis:413 (substitute name for Planorbis kuhnerianus Clessen 1883). Tropicorbis kuhnianus (Clessen). F. C. Baker 1945; Moll. Fam. Biomphalaria kuhniana (Clessin). Paraense 1988; Mem. do Inst. Ozwaldo Cruz, 83:1.Paraense 2000; Malac. Rev. 33:5; Planorbis isthmicus Tropicorbis (lateorbis) isthmicus (Pilsbry). F. C. Baker 1945; Moll. Type Locality.Planorbis kuhniana: Paramaribo, Surinam. Planorbis isthmicus: Chinese wells at Cd. Panam, Panam. Distribution.PANAM, Prov. Panam: Cd. Panam; Cd. Panam; National University, Tucuman; Lago Isleta, environs of Cd. Panam (Paraense 2000) Biomphalaria nicaraguana (Morelet 1851)Planorbis nicaraguana Morelet 1851; Test. Noviss. II:14.Von Biomphalaria nicaraguana (Morelet). Paraense 2000; Malac. anatomy).Type Locality.Lago de Nicaragua. Type BMNH 1993.2.4.594). Distribution.NICARAGUA, Dept. Chontales: Acoyapa, near Lago de Nicaragua (Paraense 2000). Dept. Rio San Juan: Rio San Juan (Von Martens 1899). Biomphalaria obstructa (Morelet 1849)Planorbis obsructus Morelet 1849; Test. Noviss. I:16. Planorbula obstructa (Morelet). Fischer & Crosse 1880:78; pl. 33, Proc. Acad. Nat. Sci. Phila. 43:322.Hinkley 1920; Nautilus 34:38.Bequaert & Clench 1936; Pub. Canegie Inst. Wash. (457):68,Richards 1937; Trans. Amer. Philos. Soc. 77:255. Planorbis (Planorbula) obstructus (Morelet). Von Martens 1899; Biol. Cent. Amer.:398. Tropicorbis (Obstructio) obstrutus (Morelet). Goodrich & van der Schalie 1937; Misc. Pub. Mus. Zool. Univ. Mich. (34):33.F. C. Baker 1945; Moll. Fam. Planorbidae:85, 128, 262, 336, Harry 1950; Occ. Pap. Mus. Zool. Univ. Mich. (524):7, 24.Bequaert 1957; Bull. Mus. Comp. Zool. 116:214. Biomphalaria obstructa (Morelet). Abdel-Malek 1969; Malacologia do Inst. Ozwaldo Cruz, 85:391.Paraense 2000; Malac. Planorbis cannarum Morelet 1849; Test. Noviss. I:16. Segmentina donbilli Tristram 1861; Proc. Zool. Soc. Lond. 29:232. Tropicorbis obstructs donbilli (Tristram). F. C. Baker 1945; Moll. (shell). Planorbis berendti Planorbula obstructa anodonta Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 71:219 (not illustrated). Tropicorbis obstructus anodontus (Pilsbry). F. C. Baker 1945: Moll. Type Localities. Planorbis obstructus: Isla de Carmen, Campeche, Mxico. Planorbis cannarum: Belize. Segmentina donbilli : Lago de Dueas, Guatemala. Planorbis berendti : Orizaba, Veracruz, Mxico. Planorbula obstructa anodonta: reservoir 4 miles north of Guatemala City, Guatemala. Distribution.Gulf coastal regions of the United States south to Belize, Guatemala, and El Salvador. EL SALVADOR, Dept. San Salvador: Cd. San Salvador (Paraense 2000). GUATEMALA, Dept. Escuintla: Escuintla (Von Martens 1899). Dept. Guatemala: Lago Amatitlan; reservoir N of Cd. Guatemala (F. C. Baker 1945). Dept. Izabal: Lago de Izabal, Marisco. Dept. Sacatepquez: Lago de Dueas, San Miguel Dueas (Paraense 2000). Dept. Petn: Lago de Petn; aguado NW of San Benito (Goodrich & van der Schalie 1937). BELIZE: Belize Dist.: Boston Village (Paraense 2000). CAMPECHE: Isla de Carmen; Aguado Yalic, Chapoton (Beqaert & Clench 1936). CHIAPAS: Laguna Ocotal to El Censo (Bequaert 1957). OAXACA: San Carlos, east of Oaxaca; Isthmus of Tehuantepec (Fischer & Crosse 1880). QUINTANA ROO: swamp a few miles N of San Miguel, Isla Cozumel (Richards 1937); near San Miguel, Isla Cozumel

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 71 (F. C. Baker 1945). SAN LUIS POTOS: Catamas Lake, El Abra; falls of the Rio Valles, Cd. Valles; Tamosopo (F. C. Baker 1945). TAMAULIPAs: Tampico (F. C. Baker 1945). VERACRUZ: Quilate, near Misantla; Orizaba; Veracruz; Rio Tenoya, Veracruz (Strebel 1873). YUCATN: Ticul (Pilsbry 1891); aguado 14 km N, 2 km E of Merida; Aguado Yalal, near Merida; Piste (Bequaert & Clench 1936); Piste (F. C. Baker 1945); 0.5 mi. W of Progreso; irrigation cistern, 3 mi. N of merida (Harry 1950). Biomphalaria orbicula (Morelet 1849)Planorbis orbiculus Morelet 1849; Test. Noviss. I:17.Fischer & Acad. Nat. Sci. Phila. 43:322.Hinkley 1920; Nautilus 34:38.Richards 1937; Proc. Amer. Philos. Soc. 77:255. Planorbis (Menetus) orbiculus Morelet. Von Martens 1899; Biol. Cent. Amer.:390. Planorbis (Tropicorbis) orbiculus (Morelet). Bequaert & Clench 1936; Pub. Canegie Inst. Wash. (457):66. Tropicorbis (Tropicorbis) orbiculus (Morelet). F. C. Baker 1945; Tropicorbis orbiculus dunkeri F. C. Baker 1945; Moll. Fam. Malacologia 1:45 (a nomen nudum, holotype not designated).Type Locality.Planorbis oriculus: La Palizada, Yucatn, on the frontier of Tabasco, Mxico. Tropicorbis orbiculus dunkeri : not stated. Distribution.Guatemala and eastern Mxico from Quintana Roo north to San Luis Potosi and Tamaulipas. GUATEMALA, Dept. Guatemala: ditch along railroad at Laguna, Lago Amatitlan (Hinkley 1920). CAMPECHE: Cd. Carmen; Laguna de Terminos. OAXACA: San Carlos (Von Martens 1899). QUINTANA ROO: swamp near San Gerbacio, Isla Cozumel (Richards 1937); near San Miguel, Isla Cozumel (F. C. Baker 1945). SAN LUIS POTOS: drift at falls, Cd. Valles; Chamiai Creek, below Cd. Valles; Catamas Lake, El Abra (F. C. Baker 1945). TABASCO: Balacan, near San Juan Bautista (Von Martens 1899). TAMAULIPAS: pool near Tampico (F. C. Baker 1945). VERACRUZ: Veracruz; San Juan (Pilsbry 1891). YUCATN: La Palizada (Von Martens 1899). DISTRITO FEDERAL: near Ciudad Mxico. Biomphalaria panamensis Dunker 1848Planorbis panamensis Dunker 1848; Proc. Zool. Soc. Lond. 16:41.Sowerby, in Reeve; Conch. Icon., 22, Planorbis Planorbis (Gyraulus) panamensis Dunker 1848. Von Martens 1899; Biol. Cent. Amer.:395.Type Locality.West Panam, in a small stream. Distribution.Known only from the type locality. Biomphalaria petenensis (Morelet 1851)Planorbis petenensis Morelet 1851; Test. Noviss. II:15. Planorbis (Gyrorbis) petenensis Morelet. Fischer & Crosse 1880; Planorbis (Gyraulus) petenensis Morelet. Von Martens 1899; Biol. Cent. Amer.:393. Tropicorbis (Lateorbis) petenensis (Morelet). F. C. Baker 1945; Type Locality.Lago de Itza Petn, Dept. Petn, Guatemala. Distribution.COSTA RICA, Prov. San Jos: Rio Torres, near San Jos. Prov. Puntarens: Laguna between Lagarto and Boruca (Von Martens 1899). GUATEMALA, Dept. Petn: Lago Petn Itza; Lago de Petn, Remate (F. C. Baker 1945). OAXACA: Cacoprieto, isthmus of Tehuantepec (Von Martens 1899); Tehuantepec (F. C. Baker 1945). Biomphalaria retusus (Morelet 1849)Planorbis retusus Morelet 1849; Test. Noviss. I:17.Pilsbry 1891; Proc. Acad. Nat. Sci. Phila. 43:322. Planorbis (Gyrorbis) retusus Morelet. Fischer & Crosse 1880:73; Planorbis (Gyraulus) retusus Morelet. Von Martens 1899:393.Type Locality.Isla de Carmen, Campeche, Mxico. Distribution.CAMPECHE: Isla de Carmen. YUCATN: Shkolac, between Merida and Valladolid (Pilsbry 1891). Biomphalaria straminea (Dunker 1848)Planorbis stramineus Dunker 1848; Proc. Zool. Soc. Lond. 16:42.Dunker 1850; in Martini & Chemnitz, Syst. Conch. Cab., Ed. 2, 1, Planorbis Biomphalaria straminea (Dunker). Paraense 2000; Malac. Rev. 33:12.Type Locality.South America. Distribution.COSTA RICA, Prov. Cartago: Rio Coris (Paraense 2000). Biomphalaria subprona (Von Martens 1899)Planorbis (Taphius) subpronus Von Martens 1899; Biol. Cent. Taphius sbpronus (Von Martens). F. C. Baker 1945; Moll. Fam. 1957; Bull. Mus. Comp. Zool. 116:214. Biomphalaria subprona (Von Martens). Paraense 1996; Mem. do Inst. Ozwaldo Cruz, 91:187.Paraense 2000; Malac. Rev. 33:12.Type Locality.Amatitlan, Tabasco [Guatemala]. Distribution.COSTA RICA: Turrialba (F. C. Baker 1945). EL SALVADOR, Dept. Santa Ana: Lago de Coatepeque (Bequaert 1957). GUATEMALA, Dept. Guatemala: Lago de Amatitlan (Paraense 2000). CHIAPAS: Laguna Ocotal, 950 m alt. (Bequaert 1957). Biomphalaria temascalensis Rangel-Ruiz 1987Biomphalaria temascalensis Rangel-Ruiz 1987; Universidad (radula); pls. 3 (reproductive system).Type Locality.Estacion de Acuacultura Tropical Holotype not designated. Distribution.Known only from the type locality.

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72 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Taxonomy.The name Biomphalaria temascalensis Rangel Ruz 1987 is invalid, because the description does not include the designation of a holotype as required by ICZN Articles 16.4 and 62.4. Biomphalaria tepicensis (Von Martens 1899)Planorbis (Gyraulus) tepicensis Von Martens 1899; Biol. Cent. Tropicobis tepicensis (Von Martens). F. C. Baker 1945; Moll. fam. Type Locality.Tepc, Jalisco [Nayarit], Mxico. Distribution.NAYARIT: Tepc. OAXACA: Tehuantepec (F. C. Baker 1945). Subfamily HELISOMATINAE F. C. Baker 1928 Genus Helisoma Swainson 1840Helisoma Swainson 1840; Treatise on Malacology, VIII:337.F. C. Baker 1945; Moll. Fam. Planorbidae:123.Type Species.Planorbis bicarinatus Say 1817 (= Planorbis anceps Menke 1830). Distribution.Throughout North America south to northern Mxico. Taxonomy.Two subgenera are recognized (Burch 1980:198). Subgenus Helisoma Swainson 1840 Distribution.As for the genus. Taxonomy.A single species is recognized. Numerous subspecies and varieties have been proposed (Burch 1880:198, 274). Two subspecies are recognized currently. One occurs in Mxico. Helisoma anceps anceps (Menke 1830)Planorbis bicarinatus Say 1817; Nicholes Encycl., 1st ed.: species Planorbis bicarinatus Lamarck 1804). Planorbis anceps Menke 1830; Synopsis Methodica Molluscorum. 2nd Ed. 242 Planorbis antrosa Conrad 1834; Amer. Jour. Sci., 25:343. Helisoma anceps anceps (Menke 1830). F. C. Baker 1945; Moll. Fam. Planorbidae:402.Taylor 1966; Veliger 9:209.Type Locality.Delaware River, Virginia. Distribution.Rio de Los Natadores, at El Cario, 20 mi. E. of Cuatro Cienegas (Taylor 1966). SONORA: mouth of the Rio Yaqui (F. C. Baker 1945), Genus Planorbella Haldeman 1842Planorbella Haldeman 1842; Monogr. NA freshwater Moll., Physidae:14.Burch 1980; Walkerana 1:202. Type Species.Planorbis campanulatus Say 1821. Distribution.North America and the west Indies south to Peru. Taxonomy.Three subgenera are recognized. Two occur in the study area. Subgenus Pierosoma Dall 1905Pierosoma Dall 1905; Smithsonian Inst. Harriman Alaska Expedition 1899, Alaska Ser. 13:81, 85. Type Species. Say 1818. Distribution.North America south to Panam and the West Indies. Taxonomy.Numerous species. Seven occur in the study area. Planorbella (Pierosoma) caloderma (Pilsbry 1923)Planorbis caloderma Pilsbry. Hinkley 1920; Nautilus 34:46, 48 (nomen nudum). Planorbis caloderma Pilsbry 1923; Nautilus 36:143.Helisoma calodermum (Pilsbry). F. C. Baker 1945; Moll. Fam. Type Locality.Esmeralda, Dept. Jutiapa, Guatemala. Holotype and paratypes ANSP 45662. Distribution.GUATEMALA, Dept. Izabal: Jocolo (Hinkley 1920). Dept. Jutiapa: Esmeralda. Planorbella (Pierosoma) (Menke 1830) Menke 1930; Symp. Method. Mollusc.:37. (Menke). F. C. Baker 1945; Moll. Fam. 12 (shell).Harry & Hubendick 1964; Geborgs Kungl. Vetenskaps-och Vitterhets-samhlles handlingar, ser. 6, ser. Planorbis caribaeus Orbigny 1841, in Sagra, Hist. Fis. Polit. y nat. 1891; Proc. Acad. Nat. Sci. Phila. 43:322.Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 72:6.Hinkley 1920; Nautilus 34:38, 48. Planorbis (Helisoma) caribaeus (Orbigny). Bequaert & Clench 1936; Pub. Canegie Inst. Wash. (457):66. Helisoma caribaeum (Orbigny). van der Schalie 1940; Occ. Pap. Mus. Zool. Univ. Mich. (413):6. Helisoma (Pieronoma) caribaeum caribaeum (Orbigny). F. C. Baker 1945; Moll. Fam. Planorbidae:149, 466, 434, 436, 438; (shell).Bequaert 1957; Bull. Mus. Comp. Zool. 116:214. Planorbis tumidus Dunker 1850; in Martini & Chemnitz, Syst. Conch. Cab., Ed. 2, Planorbis Dall 1896; Proc. U. S. Nat. Mus. 19:369.Pilsbry 1891; Proc. Acad. Nat. Sci. Phila. 43:322. Planorbis ancylostomus Crosse & Fisher 1879; Jour. de Conchyl. (shell). Planorbis ancylostoma var. chiapnesis Fischer & Crosse 1886:63; Planorbis (Helisoma) caribaeus chiapensis Fischer & Crosse. Von Martens 1893; Biol. Cent. Amer.:389.Type Localities. : Jamaica (Pilsbry 1934). Planorbis caribeaus: Cuba. Planorbis dunkeri: not Planorbis ancylostomus: Veracruz, Veracruz, Mxico. Planorbis ancylostoma var. chiapensis: Chiapas, Mxico. Distribution.COSTA RICA, Prov. Cartago: E of Cartago; 4 mi. SW of Cartago; San Isidro del Tejar (Pilsbry 1920). GUATEMALA, Naranjo (F. C. Baker 1945). Dept. Alta Verapaz: a small aguada ca. 8 km N of Samanzana; near Samac (van der Schalie 1940). Dept. Izabal: Jocolo (Hinkley 1920). Dept. Guatemala: Lago Amatitlan. CAMPECHE:

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 73 Aguado Yalic, near Chompoton (Bequaert & Clench 1936). CHIAPAS: Laguna Ocotal, 950 m alt.; Rio Amarillo at the Sumidero, near Las Casas (Bequaert 1957). SONORA: Rio San Bernardino; Rio Sonoyta (Dall 1896). VERACRUZ: Orizaba (Pilsbry 1891); Veracruz (F. C. Baker 1945). YUCATN: Shkolak (Pilsbry 1891). Taxonomy.The following subspecies has been proposed from the study area. Planorbella ( Pierosoma) (Clessin 1889)Planorbis guatemalenis Clessin 1889; in Martini & Chemnitz, Syst. Conch. Cab., Ed. 2, Planorbis Helisoma (Pierosoma) caribaeum guatemalense (Clessin). F. C. 32 (shell).Type Locality.Guatemala. Distribution.NICARAGUA (F. C. Baker 1945). GUATEMALA. Planorbella (Pierosoma) costaricense (Preston 1907)Planorbis costaricensis Preston 1907; Ann. Mag. Nat. Hist. Ser. 7, Helisoma (Pierosoma) costaricense (Preston). F. C. Baker 1945; T ype Locality.Catalina, Prov. Guanacaste, Costa Rica. Distribution.COSTA RICA, Prov. Guanacaste: Catalina (F. C. Baker 1945). Planorbella (Pierosoma) contrerasi (Pilsbry 1920)Planorbis contrerasi Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. Helisoma (Pierosoma) contrerasi (Pilsbry). F. C. Baker 1945; Moll. Type Locality.Lago de Chapala, Jalisco, Mxico. Holotype ANSP 46193. Distribution.Known only from the type locality. Planorbella (Pierosoma) tenue (Dunker 1850)Planorbis tenuis Dunker 1850; Syst. Conch. Cab., Ed. 2, 1904; Proc. Acad. Nat. Sci. Phila. 55:777.Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 72:6. Helisoma tenuis (Dunker). Goodrich & van der Schalie 1937; Misc. Pub. Mus. Zool. Univ. Mich. (34):33. Helisoma (Pierosoma) tenue tenue (Dunker). F. C. Baker 1945; Bequaert & Miller 1987:208. Planorbella (Pierosoma) tenue (Dunker). J. B. Burch 1980; Planorbis mexicanus Dunker 1850; in Martini & Chemnitz, Syst. Conch.-Cab., Ed. 2, Planorbis:45. Planorbis tumens Carpenter 1857; Cat. Mazatlan shells:81.Fischer in Martini & Chemnitz, Syst. Conch. Cab., ed. 2; Limnaeiden:88; ser. 4 12:516. Planorbis (Helisoma) tumens Carpenter. Von Martens 1899:387. Helisoma tumens (Carpenter). F. C. Baker 1945; Moll. Fam. Planorbis tenuis var. Von Martens 1899; Biol. Cent. Planorbis tenuis var. uhdei Von Martens 1899; Biol. Cent. Type Localities. Planorbis tenuis: ditches in the neighborhood of Ciudad Mxico. Planorbis mexicanus: Mxico. Planorbis tumens: Mazatln, Sinaloa, Mxico. Planorbis tenuis var. : City of Mxico. Planorbis tenuis var. uhdei: central mexico. Distribution.GUATEMALA, Dept. Petn: Arroyo Subn, near Sanat Teresa (Goodrich & van der Schalie 1937). BAJA CALIFORNIA SUR: Mulege (Hanna 1923). DURANGO. DISTRITO FEDERAL: Ciudad Mxico; Lago de Chalco (Pilsbry 1920). GUANAJUATO: Jaral (Von Martens 1899); Acambaro (Pilsbry 1903). JALISCO; Lago de Chapala (Von Martens 1899). MXICO: Tlapam (Pilsbry 1903); Toluca; sources of the Rio Lerma, near Toluca (F. C. Baker 1945). SINALOA: Mazatln (Von Martens 1899). Taxonomy.F. C. Baker (1945) recognized the following subspecies from the study area. Planorbella (Pierosoma) tenue boucardi (Fischer & Crosse 1880)Planorbis tenuis var. boucardi Fischer & Crosse 1880:61; pl. 32, Helisoma tenue boucardi (Fischer & Crosse). F. C. Baker 1945; Type Locality.Environs of Ciudad Mxico. Distribution.DISTRITO FEDERAL: known only from the environs of Ciudad Mxico. Planorbella (Pierosoma) tenue chapalensis (Pilsbry 1920)Planorbis chapalensis Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. Helisoma tenue chapalensis (Pilsbry). F. C. Baker 1945; Moll. Fam. Type Locality.Lago de Chapala, Jalisco, Mxico. Holotype ANSP 46194. Distribution.Known only from the type locality. Planorbella (Pierosoma) tenue exaggeratum (Von Martens 1899)Planorbis tenuis Philippi. Pilsbry 1891; Proc. Acad. Nat. Sci. Phila. Planorbis tenuis var. exaggeratus Von Martens 1899; Biol. Cent. Amer.:385.Pilsbry 1903; Proc. Acad. Nat. Sci. Phila. 55:777. Helisoma tenue exaggeratum (Von Martens). F. C. Baker 1945; Type Locality.Lago de Patzcuaro, Michoacn, Mxico. Distribution.Known only from the type locality. Planorbella ( Pierosoma) tenue pertenue (F. C. Baker 1940)Planorbis tenuis var. applanatus Von Martens 1899:384; pl.

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74 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Planorbis applanatus Thomae 1845). Helisoma tenuis pertenuis F. C. Baker 1940; Nautilus 54:97 (replacement name for Planorbis tenuis applanatus Von Martens). Helisoma tenue applanatum (Von Martens). F. C. Baker 1945; Moll. Type Locality.Plateau of Mxico. Distribution.CHIHUAHUA: Rio Lanos (F. C. Baker 1945). SONORA: Rio Yaqui, near Ciudad Obregon (F. C. Baker 1945). VERACRUZ: Orizaba (F. C. Baker 1945). BAJA CALIFORNIA NORTE: Vallecitos (F. C. Baker 1945). Planorbella (Pierosoma) tenue strebelianum (Fischer & Crosse 1880)Planorbis tenuis var. strebialunus Fischer & Crosse 1880:63. Helisoma tenue strebelianum (Fischer & Crosse). F. C. Baker 1945; (shell).Type Locality.Laguna de Los Cocos, Rio Tenoya and ditches in environs of Veracruz, Mxico. Distribution.SAN LUIS POTOS: Penasco, 5 mi. N of San Luis Potosi; Rio Coy (F. C. Baker 1945). Tamaulipas: Alta Mira (F. C. Baker 1945). VERACRUZ: environs of Veracruz. Planorbella (Pierosoma) (Say 1817) Say 1817; Nicholsons Encyclopedia, American Planorbella (Pierosoma) (Say). J. B. Burch 1980; (Say). Paraense 2000; Malac. Rev. 33:2; textType Locality.Lake Erie, North America. Distribution.Widely distributed in North America from the Atlantic coast westward to Missouri and northward to the Arctic Circle and Alaska. Records from Mxico and Central America seem questionable. COSTA RICA, Prov. Cartago: Rio Coris (Paraense 2000). NICARAGUA, Dept. Grenada: Lake Nicaragua, Grenada (Paraense 2000). GUATEMALA, Dept. Alta Verapaz: Lago Petencito, near Cobn (Paraense 200o). BELIZE: Belize Dist.: Boston Village (Paraense 2000). Planorbella (Pierosoma) wyldi (Tristram 1861)Planorbis tumens 4, 4a-c (shell) Planorbis wyldi Tristram 1861; Proc. Zool. Soc. Lond. 29:232. Planorbis tenuis var. wyldi Tristram. Fischer & Crosse 1880; 62; pl. Planorbis (Helisoma) wyldi Tristram. Von Martens 1899; Biol. Helisoma (Pierosoma) wyldi (Tristram). F. C. Baker 1945; Moll. Clessin 1884; in Martini & Chemnitz, Syst. Conch. Cab., Ed. 2, Planorbis (Clessin). F. C. Baker 1945; Moll. Fam. Type Locality.Planorbis wyldi : Lago de Dueas, Dept. Sacatepquez, Guatemala. : Guatemala. Distribution.Known in Central America from Costa Prov. Puntarenas: along Rio Diquis, below Terraba (Von Martens 1899). NICARAGUA, Dept. Grenada: near Volcan Monbacho (Von Martens 1899); San Carlos (F. C. Baker 1945). EL SALVADOR, Dept. San Miguel: Lago de Olomega (F. C. Baker 1945). Dept. Santa Ana: Lago de Guija; Lajo de Metapan (F. C. Baker 1945). GUATEMALA, Dept. Alta Verapaz: Cobn; Dept. Guatemala: Aceituno; Naranjo. Dept. Sacatepquez: Lago de Dueas; Santiago Zamora (Von Martens 1899). Subgenus Seminolina Pilsbry 1934Seminolina Pilsbry 1934; Proc. Acad. Nat. Sci. Phila. 86:31.F. C. Baker 1945; Moll. Fam. Planorbidae:129.Type Species.Planorbis scalare Jay 1839. peninsular Florida. It has been introduced widely in other parts of the world. Taxonomy.Two species are recognized. Planorbella (Seminolina) duryi (Wetherby 1879)Planorbis duryi Wetherby 1879; Jour. Cincinnati Soc. Nat. Hist. Helisoma (Seminolina) duryi duryi (Wetherby). Pilsbry 1934; Proc. 1 (shell). Planorbella duryi 718, 719 (shell).Thompson 1984; Freshwater snail of Fla. Malac. Rev. 33:2, 4.Type Locality.The Florida Everglades, Florida. distribution. Introduced widely elsewhere. COSTA RICA, Prov. Cartago: Rio Coris (Paraense 2000). Taxonomy.Pilsbry (1934) and F. C. Baker (1945) recognized six subspecies. Their status has not been investigated since then. Genus Micromenetus F. C. Baker 1945Micromenetus F. C. Baker 1945; Moll. Fam. Planorbidae:197.Type Species.Planorbis dilatatus Gould 1841. Distribution.Eastern North America; Bermuda. Taxonomy.Eight species are recognized. One occurs in the study area. (Pilsbry 1905)Planorbis alabamensis var. Pilsbry 1905; Nautilus 19:34. Promenetus (Micromenetus) Pilsbry. F. C. Baker (Pilsbry). Thompson 1983; Nautilus 97:68.Thompson 1984; Freshwater Snails Fla., Man. for

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 75 Promenetus minutus Taylor 1954; Rev. Soc. Malac. Carlos de la Torre, 6:37.Type Localities.Pliocene Caloosahatchee Formation, Florida. Distribution.Known from South Florida, Jamaica, Haiti, and Panam. PANAM, Canal Zone: Allee Stream, Isla Barro Colorado (Taylor 1954). Family PHYSIDAE Fischer & Crosse 1886 Distribution.Europe, temperate Asia, North America, Meso-America, South America. Taxonomy.The family contains two subfamilies and 23 by shell characters (Taylor 2003). The locality records listed below are taken from Taylor. Previous published records are omitted because of the chaotic taxonomic history of the family in Mxico and Central America. Subfamily APLEXINAE Starobogatov 1967 Taxonomy.Taylor (2003) lists twelve genera. Distribution.Holarctic in distribution and extending into the tropical realm in South America. Genus Austrinauta Taylor 2003.Austrinauta Taylor 2003; Rev. Biol. Trop. 51 (suppl.):43.Type Species.Physa elata Gould 1853. Distribution.Known only from Nayarit State, Mxico. Taxonomy.One species. Austrinauta elatus (Gould 1853)Physa elata Gould 1853; Jour. Boston Soc. Nat. Hist. 6:379; pl. 14, Aplecta elata (Gould). Fischer & Crosse 1870; 2:92; pl. 32, Austrinauta elatus (Gould). Taylor 2003; Rev. Biol. Trop. 51 Type Locality.Lower California. Lectotype MCZ 169130 (Johnson 1964:71). Distribution.Known only from a single locality. NAYARIT: freshwater marsh beside Matachn-Santa Cruz W). Genus Amecanauta Taylor 2003Amecanauta Taylor 2003; Rev. Bio. Trop., 51 (suppl.):72.Type Species.Amecanauta jaliscoensis Taylor 2003. Distribution.Nayarit and Jalisco, Mxico. Taxonomy.A single species. Amecanauta jaliscoensis Taylor 2003Amecanauta jaliscoensis Taylor 2003; Rev. Biol. Trop. 51 Type Locality.Jalisco: roadside ditch on W side of hwy. 200, opposite entrance to Modulo de Abasto de Distribution.Known only from the type locality. Genus Mexinauta Taylor 2003Mexinauta Taylor 2003; Rev. Biol. Trop. 51 (suppl.):74.Type Species.Physa nitens Philippi 1841. Distribution.Texas, Mxico, Guatemala, Nicaragua, Ecuador, Peru. Taxonomy.Eight species, seven of which occur in Mxico and Central America. Mexinauta aurantia (Carpenter 1857)Aplexa aurantia Carpenter 1857; Cat. Mazatlan shells:179. Bulinus aurantius (Carpenter). Binney 1865; Smiths. Misc. Coll. Stenophysa aurantia (Carpenter). Taylor 1966; Veliger 9:111. Mexinauta aurantia (Carpenter). Taylor 2003; Rev. Biol. Trop. 51 Type Locality.Mazatln, Sinaloa, Mxico. Syntype BMNH 815819. Mxico and northwestern Costa Rica. JALISCO: marsh at km NE Puerto Marques. COSTA RICA: Prov. Guanacaste; Parque Nacional Palo Verde, edge of marsh 100 m E of W immediate vicinity. Mexinauta gracilentus (Fischer & Crosse 1886)Aplecta aurantia var. gracilenta Fischer & Crosse 1886; 2:87; pl. Mexinauta gracilentus (Fischer & Crosse). Taylor 2003; Rev. Biol. Trop. 51 (suppl.):76.Type Locality.Cobn, Dept. Alta Verapaz, Guatemala. Holotype in MHNP. Distribution.Known only from the type locality. (Morelet 1849) Morelet 1849; Test. Noviss. I:18. (Morelet). Taylor 2003; Rev. Biol. Trop. 51 9, 11 (shell).Type Locality.Ditches in Guatemala City, Guatemala. Syntypes in the BMNH (3 uncatalogued specimens). Distribution.CHIAPAS: pond at Pueblo Nuevo, Solistahuacn; San Cristobal de las Casas, 2200 m alt.; km 65, S of Tuxtla Gutierrez along road to Nuena Concordia, 2700 ft. alt.; 15 km S of Ocozocuautla, along road to Villa Flores, 2800 ft. alt. GUATEMALA, Dept. Chimaltenango: Yepocapa, in fountain; Parque Nacional Los Aposentos, 2.5 km S of Chimaltenango; ditch near Parque Chimaltenango. Dept. Guatemala: Laguna El Naranjo, Finca El Naranjo,

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76 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) ca. 5 km NW of Ciudad Guatemala, 1560 m alt.; environs of Guatemala City; Guatemala City. Dept. Sacatepquez: Antigua; Laguneta Quilisimate, between Santiago Zamora and Santa Catarina Barahona, 1460 m alt.; small stream entering Lago de Dueas, above Santiago Zamora. Dept. Santa Rosa: Rio Taxisco, Taxisco. Dept. Zacapa; Zacapa. Mexinauta laetus (Von Martens 1898) var. laeta Von Martens 1898; Biol. Cent. Mexinauta laetus (Von Martens). Taylor 2003; Rev. Biol. Trop. 51 (suppl.):76.Type Locality.Vera Paz, Guatemala. Lectotype ZMB 51234a (Kilias 1961). Distribution.Known only from the type locality. Mexinauta nicaraguana (Morelet 1851)Physa nicaraguana Morelet 1851; 2:16.Von Martens 1898; 366; Mexinautan nicaraguana (Morelet). Taylor 2003; Rev. Biol. Trop. 51 (suppl.):76.Type Locality.Lake Nicaragua, Nicaragua. Syntype BMNH 1893.2.4.610. Distribution.Known only from the type locality. Mexinauta nitens (Philippi 1841)Physa nitens Philippi 1841, in 1. Aplecta nitens 1b. Mexinauta nitens (Philippi). Taylor 203; Rev. Biol. Trop. 51 5 (shell). Bulinus berlandierianus Binney 1865; Amer. Jour. Conch. 1:51; pl. Aplecta nitens var. acutalis Fischer & Crosse 1886; 2:88; pl. 39, Aplecta nitens var. gigantea Fischer & Crosse 1886; 2:89; pl. 39, Type Locality.Physa nitens: Mexiko. Bulinus berlandierianus: Texas, in the vicinity of Matamoros, Tamaulipas, Mxico. Distribution.Coastal lowlands along the Gulf of Mexico from the vicinity of Brownsville, Texas south to Campeche. TAMAULIPAS: Hacienda Acua, 808 m alt. VERACRUZ: Charco de Buenos Aires; Misantla; Veracruz; la Llave and Piedras Negras; Cosamaloapan. TABASCO. Pasture pond at km 21.1, NW of Taxco, on road Nacajuco to Villahermosa; pond beside Mex. Hwy. 180, 1 km N of road to Ignacio Allende; 2.3 km NE of Playa Miramar; marsh about 1 km N of junction Tacotalpa-Tapijulpa road with road to Teapa; pond beside Jalapa-Tacolapa road 2 km S of road to Guanal, N of Rancho Nuevo. CAMPECHE. Ditch beside Mex. Hwy. 180, 0.5 km E of Nuevo Progreso. Mexinauta princeps (Phillipi 1846)Physa princeps Phillipi 1846; in Kster, Martini & Chemnits, Syst. Aplexa princeps (Phillipi). Bequaert & Clench 1933; Pub. Carnegie Inst. Wash. (431):539. Stenophysa princeps (Phillipi). Rehder 1966; Proc. Biol. Soc. Wash. 79:280. Mexinauta princeps (Phillipi). Taylor 2003; Rev. Biol. Trop. 51 Type Locality.Yucatn. Holotype ANSP 21184a (H. B. Baker 1964:155). Distribution.Yucatn Peninsula, from Yucatn and Quintana Roo, Mxico through northern Guatemala and Belize (Taylor 2003). QUINTANA ROO: Marsh 0.5 km W of Puerto Morelos; Laguna de Cob, Cob, Rio Honda. YUCATN: 4 km S of Purto Progresso; Dzadz Cenote, ca. 1 km SW of Chichn Itz. GUATEMALA, Dept. Petn: Aguada 0.5 km from La Libertad road to San Francisco; small dry aguada just SW of San Benito (opposite Flores on mainland). BELIZE: Corazal Dist.: Louisville Forest. Orange Walk Dist.: Orange Walk. Genus Mayabina Taylor 2003Mayabina Taylor 2003; Rev. Biol. Trop. 51 (suppl.):88.Type Species.Physa spiculata Morelet 1849. Distribution.From Oaxaca and Veracruz, Mxico, south to Costa Rica; Ecuador to northernmost Chile (Taylor 2003). Taxonomy.Taylor (2003) lists eleven species, ten of which occur in Mxico and Central America. Mayabina bullula (Crosse & Fischer 1882)Aplecta bullula Crosse & Fisher 1882; Jour. de Conchyl. 30:334.Mayabina bullula (Crosse & Fischer). Taylor 2003; Rev. Biol. Trop. Type Locality.Tuxpan, Veracruz, Mxico. Holotype lost (Taylor 2003). Distribution.Known only from the vicinity of the type locality. VERACRUZ: Tuxpan; pool on W side of Mex. Hwy. Mayabina petenensis Taylor 2003Mayabina petenensis Taylor 2003; Rev. Biol. Trop. 51 (suppl.):96; Type Locality.Guatemala, Dept. Petn, aguada at NE Distribution.Known only from the Dept. Petn, Guatemala. Taylor (2003) listed numerous localities from there. Mayabina pliculosa (Von Martens 1898)Physa fuliginea var. pliculosa Von Martens; 1898; Biol. Cent. Stenophysa pliculosa (Von Martens). Tonn et al. 1964; Rev. Biol. Trop. 12:60. Mayabina pliculosa (Von Martens). Taylor 2003; Rev. Biol. Trop.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 77 Cartago, Costa Rica. Distribution.Western Costa Rica from the Central Valley northwestward to the southern tip of the Nicoya Peninsula (Taylor 2003). COSTA RICA, Alajuela Sur Prov.: Sachi Sar. Prov. Cartago: Rio Turialba, Turialba; Turialba; Alto de Ochomogo; Aguas Calientes, near Cartago; Quebrada Barahona, 1380 m alt.; Juan Vis. Prov. Guanacaste: ditch 3 km toward Hacienda La Taboga, from Caas-Bebedero Pavas; Uruca; San Jos; Rio Torres, San Jos; Rio Torres; Ocloro. Mayabina polita Taylor 2003Aplecta cisternina var. Fischer & Crosse 1886; 2:95; pl. Aplecta cisternina var. gracilis Fischer & Crosse 1886, 2:95; pl. 30, Aplexa spiculata var. gracilis (Fischer & Crosse). Bequaert & 5 (shell). Aplecta cisternina var. minor Fischer & Crosse 1886; 2:95; pl. 30, Mayabina polita Taylor 2003; Rev. Biol. Trop. 51 (suppl.):99; Type Localities. : Mrida, Yucatn; holotype lost. Aplecta gracilis: vicinity of Mrida, Yucatn; holotype lost. Mayabina polita : pasture pool 50 m W of Rio Tulija, 1.5 km S of Mex. Hwy. 186 toward Zopo Norte Distribution.Southeastern Mxico, in Tabasco and northern Chiapas through the Yucatn Peninsula in eastern Campeche, Yucatn and Quintana Roo. Taylor (2003) provided numerous locality records. Mayabina sanctijohannis Taylor 2003Mayabina sanctijohannis Taylor 2003; Rev. Biol. Trop. 51 Colorado; marsh on left bank of Rio Colorado. Mayabina spiculata (Morelet 1849)Physa spiculata Morelet 1849, 1:18.Von Martens 1898; Biol. Cent. Amer.:366. Aplecta spiculata 13 (shell). Aplexa spiculata (Morelet). Bequaert & Clench 1936; Pub. Canegie Mus. Zool. Univ. Mich. (524):23. Physa (Aplexa ) spiculata Morelet. Branson & McCoy 1965; Univ. Colorado Stu., Ser. Biology, 13:12. Stenophysa spiculata (Morelet). Rehder 1966; Proc. Biol. Soc. Wash. 79:280. Aplexa nitens var. spiculata (Morelet). Pilsbry 1891; Proc. Acad. Nat. Sci. Phila. 43:325. Physa cisternina Morelet 1851, 2:15. Aplecta cisternina (Morelet). Fischer & Crosse; 1886, 2:94; pl. 30, Mayabina spiculata (Morelet). Taylor 2003; Rev. Biol. Trop. 51 Type Locality.Cd. Campeche, Campeche Types lost. Distribution.Yucatn Peninsula in eastern Campeche, Yucatn and Quintana Roo (Taylor 2003). CAMPECHE: Cd. 93 km E of Silvituc. QUINTANA ROO: San Gerbacio, Isla Cozumel; Ruinas Kuhunlich; potreros at Toms Garrido. YUCATN: cienaga 2 km SW of Progreso, Cerro Isla; south side of cienaga near Progreso; irrigation cistern ca. 5 km N of Mrida; charcos at Pemex plant, 4 km S of Puerto Progreo; Izamal; Cenote Shkolak, and Tekanto; Hatal Aguada, near Mrida; Yunku Aguada, near Yunku; Chichen-Itz; aguada S Mayabina tapanensis (Crosse & Fischer 1882)Aplecta tapanensis Crosse & Fischer 1882; Jour. de Conchyl. 29:334. Aplecta spiculata var. tapanensis Fischer & Crosse 1886, 2:93: pl. Physa tapanensis (Crosse & Fischer). Von Martens 1898:367. Aplexa tapanensis guatemalensis (Crosse & Fischer). Hinkley 1920; Nautilus 34:38. Mayabina tapanensis (Crosse & Fischer). Taylor 2003; Rev. Biol. Trop. 51 (suppl.):107.Type Locality.Near Tapana [San Pedro Tapanatepec], 2003). Distribution.Southeastern Oaxaca to southern Guatemala (Taylor 2003). OAXACA: San Pedro Tapanatepec, in northern part of town, in a marsh 100 m E of km post 90 on Mex. Hwy. 190. CHIAPAS: 65 km S of Tuxtla Gutierrez, along road to Nueva Concordia. GUATEMALA, Dept. Guatemala: pools along railroad across Laguna Amatitln from Laguna Station. Dept.Solol: San Buenaventura, 1 km NW of Panajachel, 1565 m alt.; Lago de Atitln at mouth of Rio Catarina, 1.5 km NW of Panahachel. Mayabina tempisquensis Taylor 2003Mayabina tempisquensis Taylor 2003; Rev. Biol. Trop. Type Locality.Parque Nacional Palo Verde, edge of marsh 100 m east of west end of airstrip, Prov. Guanacaste, 146092. Distribution.COSTA RICA, Prov. Guanacaste: edge of marsh 100 m E of W end of airstrip, Parque Nacional Palo Verde; ornamental pool inside Hotel Las Espuelas, 2 km SE of La Libertad; Rio Caas, 9 km S of Cartagena.

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78 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Genus Tropinauta Taylor 2003Tropinauta Taylor 2003; Rev. Biol. Trop. 51 (suppl.):11011.Type Species.Tropinauta sinusdulcensis Taylor 2003. Distribution.Costa Rica. Taxonomy.A single species is currently recognized. Tropinauta sinusdulcensis Taylor 2003Tropinauta sinusdulcensis Taylor 2003; Rev. Biol. Trop. 51 Type Locality.A small stream in pasture 3 km SE of W). Holotype CAS 146095. Distribution.Known only from the type locality. Subfamily PHYSINAE Fischer & Crosse 1886 Taxonomy.Taylor (2003) listed eleven genera. Distribution.Holarctic in distribution, extending into the neotropical realm in Middle America. Genus Chiapaphysa Taylor 2003Chiapaphysa Taylor 2003; Rev. Biol. Trop. 51 (suppl.):167.Type Species. Taylor 2003. Distribution.Chiapas, Mxico, and Costa Rica. Taxonomy.Two species are included in the genus. Taylor 2003 Taylor 2003; Rev. Biol. Trop. 51 Type Locality.Rio Suchiapa, 2 km SE of Suchiapa, Ocean and the Gulf of Mexico. CHIAPAS: Rio La Venta, Las Taylor 2003 Taylor 2003; Rev. Biol. Trop. 51 8 (shell). Holotype CA 114784. Distribution.Known only from Costa Rica. COSTA RICA, Prov. Guanacaste: Rio Salina, 1 km NW of Soley Genus Haitia Clench & Aguayo 1932Haitia Clench & Aguayo 1932; Proc. New England Zoological Club 13:37.Taylor 2003; Rev. Biol. Trop. 51 (suppl.):128. Type Species.Physa elegans Clench & Aguayo 1932. Distribution.Tropical and temperate North America, Central America, the West Indies, South America in Chile (Taylor 2003). Haitia lacustris (Clessin 1886)Physa lacustris Clessin 1886; Syst. Conch. Cab., ed. 2, 338:334; pl. Haitia lacustris (Clessin). Taylor 2003; Rev. Biol. Trop. 51 (suppl.):133.Type Locality.Lago de Coatepeque, Dept. Sanata Ana, El Salvador. Holotype BMNH 93.2.4.1370. Distribution.Known only from the type locality. Taxonomy.Possibly only an ecotype of Haitia mexicana (Taylor 2003). Haitia mexicana (Philippi 1841)Physa mexicana Philippi 1841; in Kster, Martini & Chemnitz, Syst. Haitia mexicana (Philippi). Taylor 2003; Rev. Biol. Trop. 51 1 (shell). Taylor listed the following forty-two names as synonyms: Physa solida Philippi 1841; in Kster, Martini & Chemnitz, Syst. Conch. Cab., ed. 2, (Limnaeaceen):6. Physa squalida Morelet 1851; Test. Noviss. II:16. Physa humerosa Gould 1855; Proc. Boston Soc. Nat. Hist. 5:128. Gould 1855; Proc. Boston Soc. Nat. Hist. 5:128. Physa fosheyi Lea 1864; Proc. Acad. Nat. Sci. Phila. 16:114. Lea 1864; Proc. Acad. Nat. Sci. Phila. 16:114. Physa halei Lea 1864; Proc. Acad. Nat. Sci. Phila. 16:114. Physa tenuissima Lea 1864; Proc. Acad. Nat. Sci. Phila. 16:114. Lea 1864; Proc. Acad. Nat. Sci. Phila. 16:115. Physa striata Lea 1864; Proc. Acad. Nat. Sci. Phila. 16:115. Physa traski Lea 1884; Proc. Acad. Nat. Sci. Phila. 16:115. Physa mexicana var. minor Von Martens 1865; Malak. Bltt. 12:57. Physa mexicana var. Von Martens 1865; Malak. Bltt. 12:58. Physa mexicana var. Von Martens 1865; Malak. Bltt. 12:58.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 79Physa mexicana var. minima Von Martens 1865; Malak. Bltt. 12:58. Physa sparsestriata Tryon 1865; Amer. Jour. Conch. 1:224. Physa distinguenda Tryon 1865; Amer. Jour. Conch. 1:225. Physa politissima Tryon 1865; Amer. Jour. Conch. 1:226. Physa dorbigniana Lea 1867; Jour. Acad. Nat. Sci. Phila. 6:166. Lymnaea ambigua Pease 1870; Amer. Jour. Conch. 6:6. Lymnaea compacta Pease 1870; Amer. Jour. Conch. 6:6. Physa mexicana minor Strebel 1874:51. Physa berendti Danker Strebel 1874:55. Physa boucardi Crosse & Fischer 1882; Jour. de Conchyl. 29:334. Physa strebeli Crosse & Fischer 1882; Jour. de. Conchyl., 29:335. Physa tehuantepecensis Crosse & Fischer 1882; Jour. de Conchyl. 29:335. Physa mexicana var. acutissima Fischer & Crosse 1886:100. Physa mexicana var. conoidea Fischer & Crosse 1886:100. Physa mexicana var. plicata Fischer & Crosse 1886:100. Physa tolucensis Fischer & Crosse 1886:100. Physa intermedia Fischer & Crosse 1886:104. Physa polakowskii Clessin 1886; in Martini & Chemnitz, Syst. Conch. Cab., Physa, 338:352. Physa cupreonitens Cockerelll 1889; Jour. Conchol., 6:63. Physa penicillata Hemphill 1890; Cat. N. Amer. Shells:19. Physa osculans rhyssa Pilsbry 1899; Proc. Acad. Nat. Sci. Phila. 51:401. Physa rhomboidea Crandall 1901; Nautilus 15:44. Cockerell 1902; Jour. Malac., 9:138. Physa cradalli F. C. Baker 1906; Trans. Acad. Sci. St. Louis, 16:8. Physa balteata Preston 1907; Ann. & Mag. Nat. Hist. 20:497. Physa bottimeri Clench 1924; Nautilus 38:12. Physa marci F. C. Baker 1924; Nautilus 38:15. Physa humerosa interioris Pilsbry 1932; Nautilus 45:139.Type Locality.Physa mexicana : Mexiko. Holotype: destroyed in SMF during 1939 war. Distribution.Continuous from the Central Valley of California southward over the Mexican Plateau to Oaxaca and Tabasco, and south through Central America at least to Colombia; eastward through southern Nevada, central and southern Utah, Arizona and New Mexico across the southern Great Plains from Colorado to Illinois, Missouri and Louisiana. Patchy occurrences in Washington, Oregon and Idaho (Taylor 2003). In Mxico, recorded by Taylor (2003) from the states of Campeche, Colima, Distrito Federal, Guerrero, Michoacn, Oaxaca, and Tabasco. In Costa Rica, recorded from the Providencias Cartago, Heredia, Puntarenas, and San Jos. Haitia moreleti Taylor 2003Haitia moreleti Taylor 2003; Rev. Biol. Trop. 51 (suppl.):146; Type Locality.Marshy border of Lago de Petn-Itz, Santa Elena, Dept. Petn, Guatemala. Holotype CAS 114821. Distribution.GUATEMALA: known only from the immediate vicinity of the type locality (Taylor 2003). Haitia patzcuarensis (Pilsbry 1891)Physa osculans var. patzcuarensis Pilsbry 1891a:9.Pilsbry Haitia patzcuarensis (Pilsbry). Taylor 2003; Rev. Biol. Trop. 51 Type Locality.Lago de Patzcuaro, Michoacn, Mxico. Holotype ANSP 61629. Distribution.MICHOACAN: known only from Lago de Patzcuaro. Haitia solidissima (Pilsbry 1920)Physa solidissima Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 71:219; Haitia solidissima (Pilsbry). Taylor 2003; Rev. Biol. Trop. 51 (suppl.):135.Type Locality.Laguna de Chapala, Jalisco, Mxico. Holotype ANSP 107482. Distribution.JALISCO: known only from Laguna de Chapala. Genus Ultraphysella Taylor 2003Ultraphysella Taylor 2003; Rev. Biol. Trop. 51 (suppl.):191.Type Species.Ultraphysella sinaloae Taylor 2003. Distribution.Known from Sinaloa and Nayarit, northwestern Mxico. Taxonomy.One species is known for the genus. Ultraphysella sinaloae Taylor 2003Ultraphysella sinaloae Taylor 2003; Rev. Biol. Trop. 51 W). Holotype CAS 146096. Distribution.NAYARIT: Rio de Palillo, El Palillo km E of exit to San Pedro Lagunillas, 0.1 km E of km post in culvert to Siqueiro, 4.2 km from Mex. Superfamily ELLOBIOIDEA L. Pfeiffer 1854 [1822] Family CARYCHIIDAE Jeffreys 1829 Genus Carychium Mller 1774CarychiumType Species.Carychium minimum Mller 1774. Distribution.Holarctic realm in general. Extending south to Java and the Philippine Islands in the Old World, and to Costa Rica and Jamaica in the New World. Taxonomy.Eleven species occur in North America. Two occur in the study area. Carychium costaricensis Von Martens 1898Carychium costaricensis Von Martens 1898; Biol. Cent. Amer.:253; Type Locality.San Jos, Costa Rica.

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80 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Distribution.COSTA RICA: type locality. GUATEMALA, Dept. Alta Verapaz: Chama (Hinkley 1920). Carychium mexicanum Pilsbry 1891Carychium exiguum mexicanum Pilsbry 1891; Proc. Acad. Nat. Sci. 1930a:2.Goodrich & van der Schalie 1937; Misc. Pub. Mus. Zool. Univ. Mich. (34):30.Pilsbry 1948; Land Moll. N. Amer., Univ. Mich. (612):12.Correa-Sandoval 1993; Rev. Biol. Trop. 41:675.Correa-Sandoval, Garca-Cubas & Reguero 1998:13.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):235.Correa-Sandoval & Rodriguez 2005:59. Carychium mexicanum Pilsbry. Hubricht 1985:6; map 14. Type Locality.Hills around Orizaba, Veracruz, 500 ft. above the town. Holotype ANSP 61628a (Baker 1964:193). Distribution.Widely distributed across the southern United States from Georgia and Florida west to Texas, and south to Guatemala. GUATEMALA (Goodrich & van der Scahlie 1937). Dept. Escuintla: Costa Cuca (Von Martens 1898). Dept. Huehuetenango: Hacienda Helvetia, in upper Cholhuitz. Dept. Izabal: Chama. Dept. Petn: Tikal (Basch 1959). NUEVO LEN: Diente, near Monterrey (Pilsbry 1948); Santiago (Correa-Sandoval 1993); near Laguna de N) (Correa-Sandoval & Rodriguez 2005). PUEBLA: near Necaxa, 2625 ft. alt. (Baker 1930a). SAN LUS POTOS: carr. Jalpan-Xilitla, 6 km SW of Xilitla, 830 m alt. TAMAULIPAS: Tampico; Ejida Santa Juana, W of Nuevo 1930a); Texolo (Pilsbry 1948). Genus Coelostele Benson 1864Coelostele Benson 1864; Ann. Mag. Nat. Hist. (3), 13:136.Type Species.Coelostele scalaris Benson 1864. Distribution.India, southern Arabia, Egypt, Syria and southern Spain; northeastern Mxico. Taxonomy.Many species. The single Mexican species may be introduced. Coelostele tampicoensis (Pilsbry 1907)Spiraxis tampicoensis 1907; Nautilus 21:77. Coelostele tampicoensis (Pilsbry). Pilsbry 1908; Man. Conch. 19:346.Correa-Sandoval, Gutierrez & Raza 1998:15.Type Locality.Tampico, Tamaulipas, Mxico; in river drift. Holotype ANSP 93786a (Baker 1964:193). Distribution.SAN LUS POTOS: numerous localities (Correa-Sandoval et al. 1998). TAMAULIPAS: only from the type locality. SYSTELOMMATOPHORA Superfamily VERONICELLOIDEA Gray 1840 Family VERONICELLIDAE Gray 1840 Distribution.Pantropical. Taxonomy.Approximately 300 species have been proposed, but many of these probably are synonyms (Thom, Santos & Perdott 1997). Some veronicellid slugs are readily distributed by human agency. They have been introduced into many places in tropical and subtropical regions, often with dire agricultural consequences because they feed upon newly emergent seedlings. Four species are native to the study area. Five others have been introduced. Genus Belocaulus Hoffmann 1925Belocaulus Hoffmann 1925; Jenaische Zeit, fr Naturw. 61:198, 245.Thom 1975; Iheringia, Zool., 48; :11.Type Species.Vaginula angustipes Heynemann 1885. Distribution.Brazil, Paraguay and Argentina. Introduced elsewhere. Taxonomy.The genus includes two species. Belocaulus angustipes (Heynemann 1885)Vaginulus angustipes Heynemann 1885; Jahrb. Deut, Malakozool. Gesell. 12:275.Simroth 1914; Mem. Soc. Neucht. Sci. (reproductive anatomy). Belocaulus angustipes (Heynemann).Hoffmann 1925; Jenaische Zeit, fr Naturw. 61:198.Lopez-Pitone and Thom 1981; Rev. Bras. Biol. 41:586.Thom 1993; Biocincias, 1:70.Thom, Santos & Pedott 1997; Proc. Biol. Soc. Wash. 110:521.Type Locality.Taquera, Rio Grande do Sul, Brazil. Lectotype SMF 194290 (Thom 1969:334). Distribution.Argentina, Brazil, Paraguay, Colombia. Introduced into Honduras, Florida, Alabama, Louisiana and Genus Diploselenodes Thom 1975Diploselenodes Thom 1975; Ihringia, Zoologie (48):13. Type Species.Vaginula bielanbergi Semper 1885. Distribution.Chile, Brazil, Ecuador, Venezuela, Guayana, Lesser Antilles, Puerto Rico; Hispaniola, Central America. Introduced elsewhere. Taxonomy.Six species are recognized (Thom 1975). Diplosolenodes occidentalis (Guilding 1825)Onchidium occidentalis Guilding 1825, Trans. Linn. Soc. 14:323: Vaginulus occidentalis (Guilding). Fischer 1871; Nouv. Arch. Mus. Hist. Nat. 7:164. Vaginulus (Latipes) occidentalis (Guilding). H. B. Baker 1925; (reproductive anatomy). Cylindrocaulus occidentalis Hoffmann 1925; Jenaische Zeit. fr Naturw. 61:144, 234.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 81Diplosolenodes occidentalis (Guilding).Thom 1975; Ihringia, Zoologie (48):14.Thom 1985; Rev. Bras. Zool. 2 (6):411 417.Thom 1993:71.Thom, Santos, & Pedott 1997; Proc. map.Type Locality.Puerto Rico. Distribution.Guayana, Venezuela, Colombia, Panam, Costa Rica, El Salvador, Nicaragua, Honduras and Dominica. Introduced into Hawaii and Bahamas (Thom, Santos & Pedott 1997:522). COSTA RICA, Prov. Puntarenas: Rio et al. 1997). PANAM, Prov. Panam: Punta de Pia; Cd. Panam. Canal Zone: Isla Flamenco (Thom et al. 1997). EL SALVADOR (Thom et al. 1997). (Stearns 1871) Stearns 1871; Conch. Mem. 8:1. Vaginulus (latipes ) (Stearns). H. B. Baker 1925; Proc. (reproductive anatomy). (Stearns). Hoffmann 1925;Jenaische (Stearns).Thom 1975; Ihringia, Zoologie (48):14.Thom & Lopez-Pitoni 1976; Rev. Bras. Biol. 36:710. Distribution.Known only from the type locality. Taxonomy.This may be a synonym of Diplosoleodes occidentalis (Baker 1925c). Genus Leidyula H. B. Baker 1925Leidyula H. B. Baker 1925; Proc. Acad. Nat. Sci. Phila. 77:158. Hoffmann 1927:211.Pilsbry 1948; Land Moll. N. Amer. 2:1063.Forcart 1967; Ann. Natal Mus. 18:251.Thom 1975; Ihringia, Zoologie (48):17.Type Species.Vaginula moreleti Fischer 1871. Distribution.Guatemala, Mxico, Jamaica, Puerto Rico, Cuba, southeastern United States. Taxonomy.About twelve species are recognized. Leidy 1851 Leidy 1851; in Binney, Terrestrial air-breathing mollusks of the United States and the adjacent territories of North America:198, 251; pl. 4. (Leidy). Binney 1878; Terr. Moll. N. Amer. (reproductive anatomy).Pilsbry 1948; Land Moll. N. Amer. anatomy). (Leidy). Naranjo-Garcia, Thom & Castillejo 2007; Rev. Mex. Biol. 78:42.Type Locality.Punta Rasa, Florida. Restricted to MetaLee-Chee Key, Charlotte Harbour, Florida (Thom 1989). Holotype USNM 180860. Distribution.CHIAPAS: at Rio Coatancito, Tapachula. NUEVO LEN: Hualahuises; Presa Los Magueyes, km 118, Cd. Victoria-Linares Hwy.; Cerro de la Silla, Guadalupe; Presa la Boca, Santiago; Las Adjuntas, Santiago. VERACRUZ: Poza Rica; Tlapacoyan (Naranjo-Garcia, Thom & Castillejo 2007). Leidyula moreleti (Fischer 1871)Vaginula moreleti Fischer 1871; Arch. Mus. Hist. Nat. 7:168, de Conchyl. 20:59.Fischer & Crosse 1878; Miss. Sci. Mex. Clench 1936; Publ. Carnegie Inst. Wash., (457):66. Veronicella (Leidyula) moreleti (Crosse & Fischer). Pilsbry 1919; Proc. Acad. Nat. Sci. Phila. 71:219.H. B. Baker 1925; Proc. Leidyula moreleti (Fischer). Thom 1971; Iheringia, Zool. (40):32 34.Thom, Santos & Pedott 1997; Proc. Biol. Soc. Wash. 110:529.Thom 1993:71.Correa-Sandoval, Garca-Cubas & Reguero 1998; Acta Zool. Mex. (73):13.Correa-Sandoval 2000; Acta Zool. Mex. (97):8.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):235.Naranjo-Garcia, Thom & Catillejo 2007; Rev. Mex. Biod. 78:42. Vaginulus mexicanus Strebel & Pfeffer 1882:130, 140, 23, 46 (reproductive anatomy). Veronicella mexicana betheli Cockerell 1913; Nautilus 27:1.Type Localities. Vaginula moreleti : Palenque, Chiapas, in Fischer 1871 (Thom 1971). Vaginulus mexicanus: Mxico; lectotype SMF 194283 (Thom 1969) Distribution.Colombia, Nicaragua, Honduras, Guatemala and Mxico. Introduced into Florida and the West Indies (Thom, Santos & Pedott 1997:529). GUATEMALA, Dept. Izabal: Jacola; Livingston (Pilsbry 1919; Baker 1925); Puerto Barios (Cockerell 1913). HONDURAS (Cockerell CAMPECHE: Isla de Carmen; Cd. Campeche (Thompson 1967). CHIAPAS: Palenque (type locality). NAYARIT: Tepc (Baker 1925); Cacaprieto. OAXACA: Mecos; Tehuantepec, Cd. Valles (Baker 1925). SAN LUIS POTOS: Cd. Valles (Hinkley 1907); Huichihuayn; Mecos; Tamazunchale (Thom et al. 1997); km 5 on road Tamazunchale-El Taiman TABASCO (Baker 1925). TAMAULIPAS: various localities in SE part of the state (Correa-Sandoval & Rodriguez 2002, 2005). VERACRUZ: Cuatotolapam (Baker 1925c); Las Tuxtlas Biological Station (Naranjo-Garcia et al. 2007); 14 km N and 2 km E of Mrida; 1.6 km N of Mrida (Bequaert & Clench 1936); Mrida; Xmakuil (Naranjo-Garcia et al. 2007). Genus Phyllocaulis Colosi 1922Phyllocaulis Colosi 1922; Ann. Mus. Nac. Hist. Nat. Buenos Aires,

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82 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)31:486.Thom 1975; Ihringia, Zool., (48):21.Thom 1976; Iheringia, Zool., 49:67.Type Species.By subsequent designation: Phylocaulus borellini Colosi 1921 (H. B. Baker 1925) (= Vaginulus soleiformis Orbigny 1835). Distribution.Brazil, Uruguay, Argentina, Chile and Bolivia. Introduced elsewhere. Taxonomy.About twenty species are recognized (Thom 1975, 1976). One had been introduced into western Mxico, but it has not been found there since it was reported in 1925. Phyllocaulis gayi (Fischer 1871)Vaginulus gayi Fischer 1871; Arch. Mus. Hist. Nat. 7:172. Vaginulus (Phyllocolis) gayi (Fischer). H. B. Baker 1925; Nautilus 39:15. Phyllocolis gayi (Fischer). Thom 1971; Iheringia, Zool., (4):27 anatomy).Naranjo-Garcia, Thom & Castillejo 2007; Rev. Mex. Biod. 78:44.Type Locality.Valdivia, Chile. Lectotype in the Museum dHistorie Naturelle, Paris (Forcart 1952). Distribution.Southern Chile (Naranjo-Garcia, Thom & Castillejo 2007). SINALOA: Mazatln (Hoffmann 1925). Genus Sarasinula Grimpe & Hoffmann 1924Sarasinula Grimpe & Hoffmann 1924; Zool. Anz., 68:177.Thom 1975; Ihringia, Zool., (48):25.Type Species.By original designation: Vaginulus plebeius Fischer 1868. Brazil, Peru, Ecuador, Colombia, Venezuela, Lesser Antilles. Introduced elsewhere. Taxonomy.Twelve species are recorded from the neotropical region (Thom 1975). Additional species occur into Mxico and Central America. Sarasinula dubia (Semper 1885)Vaginua dubia Semper 1885; Landmollusken 7:296.Thom 1972:252. Sarasinula dubia (Semper).Thom 1993; Biocincias 1:71.Thom, Santos & Pedott 1997; Proc. Biol. Soc. Wash. 110:530.Naranjo-Garcia, Thom & Castillejo 2007; Rev. Mex. Biod. 78:46. Type Locality.St. Thomas, Virgin Islands. Lectotype ZMB 39057a (Naranjo-Garcia, Thom & Casillejo 2007). Distribution.Brazil, Venezuela, Colombia, Honduras and Mxico. Introduced into Texas, Florida, the West Indies distribution in Central America and Mxico is questionable. Dept. Corts: La Lima (Thom et al. 1997). EL SALVADOR, Dept. La Libertad: Santa Telca. CHIAPAS: ca. 1 km E of Finca Custepec, E side of Rio Custec; 5 km E of Caada W). JALISCO: El Grullo; Pueblo Careyes, km 54 on Fed. Hwy. 200, Barra de NavidadPuerto Vallara; Puerto Vallara. MORELOS: Tetecalita, 16.5 km S, 65. km E of Cuernavaca; 25 km W of Huamantla. OAXACA: 4 km NW of Tepantepec SAN LUS POTOS: Cascada de Tamasopo, 4 km E of Tamasopo. SINALOA: Mazatln. VERACRUZ: Cuitlhuac (all Mexican records from Naranjo-Garcia et al. 2007). Sarasinula plebeia (Fischer 1868)Vaginulus prebeius Fischer 1868; Jour. de Conchyl. 16:145. Sarasinula plebeia (Fischer).Andrews & Dundee 1987; Ceiba 28:163.Thom 1993; Biocincias, 1:71.Thom, Santos & Pedott 1997; Proc. Biol. Soc. Wash. 110:530.NaranjoGarcia, Thom & Catillejo 2007; Rev. Mex. Biod. 78:45.Type Locality.Nouma, New Caledonia (Thom 1971). Distribution.Native to Chile, Brazil, Venezuela, Colombia, Costa Rica, El Salvador, Honduras, Guatemala, and Mxico. Introduced into Florida, Texas, the West Indies, Australia, Asia, and Africa (Thom, Santos & Pedott SALVADOR, Dept. La Libertad: Santa Telca. HONDURAS: MXICO, VERACRUZ: Los Tuxtlas Biological Station (Naranjo-Garcia et al. 2007). YUCATN: Merida (Andrews & Dundee 1987). Genus Veronicella Blainville 1817Veronicella Blainville 1817; Journal de Physique, de Chimie, et dHistoire Naturelle..., 85 (12):437.H. B. Baker 1925:158.Thom 1975; Ihringia, Zoologie (48):32.Type Species. Blainville 1817, by monotypy. Distribution.Native to Jamaica, Hispaniola, Cuba, Ecuador, and Chile. Introduced elsewhere. Taxonomy.Seven species are recognized (Thom 1975, 1988). Veronicella sloanei (Cuvier 1816)Onchdium sloanei Cuvier 1816; Reg. An., II:411. Veronicella sloanei (Cuvier).H. B. Baker 1925; Proc. Acad. Nat. Zool. (67):13.Thom 1993; Biocincias, 1:70.Thom, Santos & Pedott 1997; Proc. Biol. Soc. Wash. 110:532. Leidyula sloanei (Cuvier). Thom 1975; Ihringia, Zool., (48):18.Type Locality.Jamaica. Distribution.Jamaica, Cayman Islands. Introduced into Barbados, Bahama Islands, Bermuda, Colombia (Isla de Providencia), Nicaragua, and Honduras (Thom, Santos & NICARAGUA: Machuca. STYLOMMATOPHORA Superfamily SUCCINEOIDEA Beck 1837 Family SUCCINEIDAE Beck 1837 Genus Catinella Pease 1870Catinella Pease 1870; Jour. de Conchyl. 18:97.Type Species.Catinella rubida Peace 1870.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 83 Distribution.Northern Europe, North America, Hawaii, and Samoa. Taxonomy.Three subgenera are recognized. One occurs in the study area. Subgenus Mediappendix Pilsbry 1948Mediappendix Pilsbry 1948; Land Moll. N. Amer. 2:843.Type Species. Pilsbry 1900. Distribution.North America, Europe, and temperate Asia.. Taxonomy.Nine species are recognized. Two occur in Mxico. Catinella (Mediappendix) (Say 1824) Say 1824; Major Longs Second Expedition 1930; Occ. Pap. Mus. Zool. Univ. Mich. (220):4.Bequaert & Clench 1936; Pub. Canegie Inst. Wash. (457):65.Pilsbry (Say). Burch 1962:67.Hubricht 1985:16; map 134. Say 1829; New Harmony Disseminator, 2:230. (Say). Hubricht 1958; Nautilus 72:60. (Say). Grimm 1960; Nautilus 74:12.Hubricht 1985:16. Catinella texana Hubricht 1961; Nautilus 75:61.Type Locality.Northwest territory. Holotype ANSP 59542. Distribution.Widespread across North America. CAMPECHE: Pantel Aguada, near Champoton (Bequaert & Clench 1936). MXICO: San Juan Teotihuacan (H. B. Baker 1930a). YUCATN: San Ignacio (H. B. Baker 1930a). Catinella (Mediappendix) rehderi Pilsbry 1948Quickella rehderi Pilsbry 1948; Land Mol. N. Amer., 2:845; anatomy). Catinella rehderi (Pilsbry). Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:128.Type Locality. miles west of Davenport, Lincoln Co., Washington. Holotype ANSP 147757a. Distribution.Washington and Montana south to Baja California Sur. BAJA CALIFORNIA NORTE: Isla Coronados; Tecate Valley, near U.S. border; 6.8 km S of San road ca. 15 km N of San Toms; 0.8 km N San Vicente, 90 m alt.; San Telmo Mesa; 40 km N of El Rosario; Millers Landing; 14 km E of Millers Landing; 13 km S of Millers Landing; mesa just S of Rancho Mezquital (Smith et al. 1990). BAJA CALIFORNIA SUR: Bajia San Bartolom; 19 km SW of San Miguel Comond; Bajia San Pedrito, 6.3 km SE of Todos Santos; 0.3 km SSE of San Jos del Cabo, on road to La Playa (Smith et al. 1990). Genus Oxyloma Westerlund 1885Oxyloma Westerlund 1885; Malak. Bltter. 3:48.Pilsbry 1948; Land Moll. N. Amer. 2 (2) 775.Type Species.Succinea duneri Pfeiffer 1865. Distribution.Northern continents and South Africa. Taxonomy.Three subgenera are recognized. One occurs in Mxico. Subgenus Neoxyloma Pilsbry 1948Neoxyloma Pilsbry 1948; Land Moll. N. Amer. 2:775.Type Species.Succinea effuse Pfeifer 1853. Distribution.North America. Taxonomy.About a dozen species are recognized. Two species and one subspecies occur in Mxico. Oxyloma (Neoxyloma ) nuttalliana Lea 1841Succinea nuttalliana Lea 1841; Proc. Amer. Philos. Soc. 2:32. Oxyloma nuttalliana (Lea).Pilsbry 1948; Land Moll. N. Amer. 2:794.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:129.Type Locality.Oregon. Distribution.BAJA CALIFORNIA SUR:19 km SW of San Miguel Comand (Smith et al. 1990). Oxyloma ( Neoxyloma) tlalpamensis tlalpamensis Pilsbry 1899Succinea tlalpamensis Pilsbry 1899; Proc. Acad. Nat. Sci. Phila. 51:401.Von Martens 1901; Biol. Cent. Amer.:647.Pilsbry (shell). Type Locality.Near Tlalpam, State of Mxico. Lectotype ANSP 77207a (H. B. Baker 1963:215. Distribution.Known only from the type locality. Oxyloma ( Neoxyloma) tlalpamensis cuitseana Pilsbry 1899Succinea tlalpamensis cuitseana Pilsbry 1899; Proc. Acad. Nat. Sci. Phila. 51:401.Von Martens 1901; Biol. Cent. Amer.:647.Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:776Type Locality.Lago de Cuitseo, near Huango, Michoacn, Mxico. Lectotype ANSP 77208a (H. B. Baker 1963:215. Distribution.Known only from the type locality. Genus Succinea Draparnaud 1801Succinea Draparnaud 1801; Tableau des Mollques terrestres et Type Species.Helix putris Linnaeus 1756. Distribution.The northern Hemisphere, North Africa, Taxonomy.Five subgenera are recognized. Subgeneric assignment is based on anatomical data, which is lacking for the vast majority of Mexican and Central American species. Two species of known anatomy are so assigned. The rest are left provisionally in the subgenus Succinea Subgenus Calcisuccinea Pilsbry 1948.Calcisuccinea Pilsbry 1948; Land Moll. N. Amer. 2:820.Type Species.Succinea campestris Say 1817. Distribution.North America. Taxonomy.About a dozen species are recognized.

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84 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Two species and three subspecies that occur in Mxico are assigned to this subgenus. Succinea (Calcisuccinea ) campestris Say 1817Succinea campestris Say 1817; Jour. Acad. Nat. Sci. Phila. 1:281.Pilsbry 1891; Proc. Acad. Nat. Sci. Phila. 43:318. Succinea (Calcisuccinea ) campestris Say. Pilsbry 1948; Land Moll. anatomy).Type Locality.Sea Islands of Georgia and Cumberland Island; Amelia Island, Florida. Distribution.Coastal areas of North Carolina, South Carolina, Georgia and Florida. MXICO, Distrito Federal: lago de Texcoco (Pilsbry 1901). Remarks.This species has not been recorded from Mxico subsequent to Pilsbry (1891). Succinea (Calcisuccinea ) luteola luteola Gould 1848Succinea luteola Gould 1848; Boston Soc. Nat. Hist. 3:37.Fischer (shell).Dall 1896; Proc. U. S. Nat. Mus. 19:365.Von Martens 1898; Biol. Centr. Amer.:331.Bequaert & Clench 1936; Pub. Canegie Inst. Wash. (457):65.Goodrich & van der Schalie 1937:31.Pilsbry 1948; Land Moll. N. Amer. 1:828; Branson, McCoy & Sisk 1964; Southwestern Natur. 9:104.Correa-Sandoval, Garca-Cubas & Reguero 1898:14.CorreaSandoval 2000; Acta Zool. Mex. (79):8.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):236.Correa-Sandoval & Rodriguez 2005:59. Succinea texasiana Pfeiffer 1848; Mon. helic. Vivent., 2:526.Type Locality.Galveston, Texas (Pilsbry 1948:830). Distribution.Widely distributed from Louisiana west to Arizona and south through much of Mxico. GUATEMALA (Goodrich & van der Schalie 1937). CHIHUAHUA: Lago de Palomas (Dall 1896). GUERRERO; Venta de Zopilote (Von Martens 1898). NUEVO LEN: km 8 on road from LUS POTOS: numerous localities (Correa-Sandoval et al. 1998). SONORA: Rio Bavispa, 21 mi. S of Agua Prieta (Branson et al. 1964). TAMAULIPAS: numerous localities in southern part of state (Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):236). VERACRUZ: Rancho El Sol, 2000). YUCATN: Progreso (Pilsbry 1891); Cerro Isla; Cienega, near Progreso (Bequaert & Clench 1936). Succinea (Calcisuccinea ) luteola rudiuscula Von Martens 1898Succinea luteola rudiuscula Von Martens 1898; Biol. Cent. Type Locality.Tehuacn, Puebla, Mxico. Distribution.Known only from the type locality. Succinea (Calcisuccinea ) luteola sonoreneis Fischer & Crosse 1878Sucinea lineata var. sonoreneis Fischer & Crosse 1878; Miss. Sci. Succinea luteola sonorensis Fischer & Crosse. Bequaert & Miller 1973:155.Type Locality.Vicinity of Rio Yaqui, Sonora, Mxico. Distribution.SONORA: 33 mi. W of Santa Ana; 1 mi. N of Santa Ana; 2 mi. N of Granados, on Rio Bavispe (Bequaert & Miller 1973). Succinea (Calcisuccinea ) luteola subtilis Von Martens 1898Succinea luteola subtilis Von Martens 1898; Biol. Centr. Amer.:331; Type Locality.Vera Cruz. Distribution.SAN LUIS POTOS: Valles (Hinkley 1907). VERACRUZ: type locality. Subgenus Succinea Draparnaud 1801 Taxonomy.The following species are left in this subgenus, pending further study of their anatomies. Twentytwo species and three subspecies occur in the study area. Sucinea (Succinea) ampulacea Von Martens 1898Succinea ampulacea Von Martens 1898; Biol. Centr. Amer.:340; pl. Type Locality.Ameca, Jalisco, Mxico. Distribution.Known only from the type locality. Succinea (Succinea) Dunker 1850 Dunker 1850; in Pfeiffer, Malak. Bltt. 7:84.14a, 14b (shell) Von Martens 1898; Biol. Centr. Amer.:339; 116:225.Type Locality.Zimapan, Hidalgo, Mxico. Distribution.CHIAPAS: El Real, 600 m alt. (Bequaert 1957). HIDALGO: Zimapan. Succinea (Succinea) californica Crosse & Fischer 1890Succinea californica Crosse & Fischer 1878; Jour. de Conchyl. 457d (shell).Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:128.Type Locality.San Toms, Baja California Norte, Mxico. Distribution.BAJA CALIFORNIA NORTE: Santo Toms; Santo Toms Valley (Smith et al. 1990).

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 85 Succinea (Succinea) carmenensis Fischer & Crosse 1878Succinea undulate var. carmenensis Fischer & Crosse 1878; Miss. 1898; Biol. Centr. Amer.:341. Succinea carmenensis Fischer & Crosse. Bequaert & Clench 1933; Pub. Carnegie Inst. Wash. (431):537.Richards 1937; Proc. der Schalie 1937:31.Rehder 1966; Proc. Biol. Soc. Wash. 79:283.Type Locality.Isla de Carmen, Campeche, Mxico. Distribution.GUATEMALA, Dept. Petn (Goodrich & van der Schalie 1937). CAMPECHE: Isla de Carmen. QUINTANA ROO: arid swamp a few mi. N of San Miguel, Isla Cozumel (Richards 1937); Tulm (Rehder 1966). YUCATN: Santa Ana, near Calcehtok; Chichen Itza (Bequaert & Clench 1933). Succinea (Succinea) clarionensis Dall 1926Succinea clarionensis Dall 1926; Proc. Calif. Acad. Sci. (4) 15:485 alt. Holotype CAS 2209. Distribution.Known only from the type locality. Succinea (Succinea) colorata Fischer & Crosse 1878Succinea colorata Fischer & Crosse 1878; Miss. Sci. Mex. 1:657; Amer.:336.Type Locality.Tabasco, Mxico. Distribution.TABASCO: San Juan Bautista (Von Martens 1898). Succinea (Succinea) Fischer & Crosse 1878Succinea undulate var. Fischer & Crosse 1878; Miss. Succinea salleana var. Von Martens 1898; Biol. Cent. Amer.:339. Distribution.Known only from the type locality. Succinea (Succinea) costaricensis Von Martens 1898Succinea costaricensis Von Martens 1898; Biol. Centr. Amer.:338; Barrientos & J. Franco 1995; Rev. Biol. Trop. 43:181. Distribution.COSTA RICA, Prov. San Jos: San Jos and at La Palma, 1161 m alt. (Von Martens 1898). Prov. Puntarenas: Savana de Guacimo, in the valley of the Rio Brus; thermal springs of Djiri Durunia, valley of the Rio Diquis, 900 m alt. (Von Martens 1898). Succinea (Succinea) globispira Von Martens 1898Succinea globispira Von Martens 1898; Sitzungs-Berichte der Gesllschaft naturforschenden Freude zu Berlin:158.Von Dall 1900; Proc. Acad. Nat. Sci. Phila. 52:97.Type Locality.Isla del Coco. Distribution.COSTA RICA: Isla del Coco (von Martens 1901). Succinea (Succinea) guadelupensis Dall 1900Succinea (rusticana Gould var.) guadelupensis Dall 1900:102; Succinea guadelupensis Dall.Pilsbry 1927; Proc. Calif. Acad. Sci. 455: I (shell).Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:128.Type Locality.Isla Guadalupe, Baja California Norte, Mxico. Holotype in the USNM. Distribution.BAJA CALIFORNIA NORTE: Known only from Isla Guadalupe (Smith et al. 1990). Succinea (Succinea) guatemalensis Morelet 1849Succinea guatemalensis Morelet 1849; Test. Noviss. I:16.Fischer Type Locality.On banks of small streams, Cd. Guatemala, Guatemala. Distribution.COSTA RICA, Prov. Cartago: Turrialba (Von Martens 1898); Juan Vias (Pilsbry 1920). Prov. Puntarenas: Turubares; Rio de las Cascadas, at the foot of the Great Fall, valley of the Rio Diquis (Von Martens 1898). 2002). GUATEMALA, Dept. Alta Verapaz: Panzos (Von Martens 1898). Dept. Sacatepquez: Antigua (Von Martens 1898). Succinea (Succinea) haustrellum Rehder 1942Succinea haustrellum Rehder 1942; Jour. Wash. Acad. Sci. 33:350; Type Locality.Pedernal, Prov. Guanacaste, Costa Rica; 200 m. alt. Holotype USNM 536013. Distribution.COSTA RICA. Prov. San Jos: San Jos (Rehder 1942). Succinea (Succinea) hortulana Morelet 1851Succinea hortulana Morelet 1851; Test. Noviss. II:14.Fischer (shell).Von Martens 1898; Biol. Centr. Amer.:334.Type Locality.Adhering to wall in gardens, Suburbs of Cd. Guatemala, Guatemala. Distribution.GUATEMALA, Dept. Sacatepquez: Antigua. Succinea (Succinea) lutosa Pilsbry 1926Succinea lutosa Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:102 Type Locality.Alhajuela, Canal Zone, Panam. Holotype ANSP 141010a (H. B. Baker 1963:215). Distribution.PANAM, Isla Taboga; Prov. Panam: Cd. Panam (Pilsbry 1926).

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86 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Succinea (Succinea) mcgregori Pilsbry 1898Succinea mcgregori Pilsbry 1898; Proc. Acad. Nat. Sci. Phila. Sci. Phila. 115:215. Nayarit, Mxico. Lectotype 72770a (H. B. Baker 1963:215). Distribution.NAYARIT: known only from the type locality/ Succinea (Succinea) panamensis Pilsbry 1920Succinea panamensis Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. Acad. Nat. Sci. Phila. 82:351.Type Locality.Las Cascades, Canal Zone, Panam. Holotype ANSP 48522a (H. B. Baker 1963:215) Distribution.PAMAMA, Canal Zone: Las Cascades (Pilsbry 1926). Succinea (Succinea) panucoensis Pilsbry 1910Succinea panucoensis Pilsbry 1910; Proc. Acad. Nat. Sci. Phila. Sci. Phila. 115:215.Type Locality.Alligator Lake, San Luis Potosi, Mxico. Lectotype ANSP 99504a (H. B. Baker 1963:215). Distribution.SAN LUIS POTOS: Pujal; Lago de Catamas (Pilsbry 1910). TAMAULIPAS: Tampico (Pilsbry 1910). Succinea (Succinea) pueblensis Fischer & Crosse 1878Succinea pueblensis Fischer & Crosse 1878; Miss. Sci. Mex.:669; Type Locality.Neighborhood of Puebla [City], Puebla, Mxico. Distribution.Known only from the type locality. Succinea (Succinea) recisa Morelet 1851Succinea recisa Morelet 1851; Test. Noviss. II:14.Fischer & Crosse, Von Martens 1898; Biol. Centr. Amer.:339.Pilsbry 1910; Proc. Acad. Nat. Sci. Phila. 62:503.Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 72:6.Richards 1938; Proc. Amer. Philo. Soc., Type Locality.Under bark of trees, around Lago de Izabal, Dept. Izabal, Guatemala. Distribution.PANAM (Tate 1859). Canal Zone: Las Cascades (Pilsbry 1910). COSTA RICA, Prov. Cartago: (Von Martens). Dept. Rio San Juan: near the Rio San Juan (Von Martens 1898). Region Autonoma Atlantico Sur: Isla del Maz (Richards 1939). HONDURAS, Dept. Islas de la Baha: Isla de Roatn, between Coxen Hole and French Harbor; West End (Richards 1938). GUATEMALA, Dept. Sacatepquez: Antigua (Von Martens 1898). Succinea (Succinea) rusticana Gould 1846Succinea rusticana Gould 1846; Proc. Boston Soc. Nat. Hist. (shell).Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:128.Type Locality.Oregon. to Baja California Norte (Roth & Sadeghian 2006). BAJA CALIFORNIA NORTE: Ojos Negros, in a slow stream through meadows (Smith et al. 1990). BAJA CALIFORNIA SUR: Sierra Laguna (Smith et al. 1990). Succinea (Succinea) socorroensis Dall 1926Succinea socorroensis Dall 1926; Proc. Calif. Acad. Sci. (4) 15:486; Type Locality.On the north slope of Cerro Evermann, Isla Socorro, Nayarit, Mxico; 2800 ft. alt. Holotype CAS 2210. Distribution.Known only from the type locality. Succinea (Succinea) undulate undulata Say 1829Succinea undulata Say 1829; New Harmony Disseminator of Useful Knowledge 2:230.Fischer & Crosse 1878; Miss. Sci. Mex. Centr. Amer.:335.Type Locality.Mxico. Distribution.GUANAJUATO: Irapuato (Von Martens 1898). JALISCO: Sayula (Von Martens 1898). MXICO. Succinea (Succinea) undulata morchi Dunker 1889Succinea undulata morchi Dunker 1889, in Paetel, Catalog Conchyl. Sammlung., 2:366.Von Martens 1898; Biol. Centr. Amer.:336; Type Locality.Zimapan, Hidalgo, Mxico. Distribution.Known only from the type locality. Succinea (Succinea) Von Martens 1865 6, 7 (shell).Von Martens 1898; Biol. Centr. Amer.:334; pl. 19, Type Locality.Veracruz, Mxico. Distribution.OAXACA. PUEBLA: Paso de Macho; Tehuacan (Von Martens 1898). VERACRUZ: Sierra de Atoyac (Von Martens 1898); Texolo (Pilsbry 1904). Succinea (Succinea) Von Martens 1898 Von Martens 1898; Biol. Centr. Type Locality.Oaxaca, Mxico. Distribution.Known only from the type locality. Succinea (Succinea) Von Martens 1898 Fischer & Crosse, Von Martens 1898; Biol. Centr. Amer.:334.H. B. Baker 1930; Occ. Pap. Mus. Zool. Univ. Mich. (220):5.Type Locality.Puebla, Mxico. Distribution.PUEBLA: Necaxa (H. B. Baker 1930a).

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 87 Superfamily ACHATINELLOIDEA Gulick 1873 Family TORNATELLINIDAE Cooke & Kondo 1961 Genus Tornatellinides Pilsbry 1910Tornatellinides Pilsbry 1910; Nautilus 23:123.Type Species.Tornatellina simplex Pease. Distribution.Japan, Formosa, Polynesian Islands, Australia, Hawaii, Galapagos island, Islas Revillagigedo. Taxonomy.Two subgenera are recognized. The typical Coast of Nayarit. The genus contains numerous species. Two species occur in the study area. Tornatellides mexicana Dall 1926Tornatellides mexicana Dall 1926; Proc. Calif. Acad. Sci. (4) Type Locality.On the south slope of Mt. Evermann, Isla Socorro, Islas Revillagigedo, Nayarit, Mxico; 2000 2800 ft. alt. Holotype CAS 2207. Distribution.NAYARIT: known only from the type locality on Isla Socorro. Tornatellides clarionensis Dall 1926Tornatellides clarionensis Dall 1926; Proc. Calif. Acad. Sci. (4) Nayarit, Mxico; 1040 ft. alt. Holotype CAS 2209. Distribution.NAYARIT: known only from the type Superfamily COCHLICOPOIDEA Pilsbry 1900 Family COCHLICOPIDAE Pilbry 1900 Genus Cochlicopa Frussac 1821Cochlicopa Frussac 1821; Tableaux Systematiques des Animaux Mollusques:24. Cionella Jeffreys 1829; Trans. Linn. Soc. London 16:347. Pilsbry 1948; Land Moll. N. Amer. 2:1045.Type Species.Helix lubrica Mller 1774. Distribution.Holarctic in occurrence. Taxonomy.Four species are recognized in North America (Hubricht 1985:6). One occurs in Mxico. Cochlicopa lubrica (Mller 1774)Helix lubrica Mller 1774; Verm. Hist.:104. Cochlicopa lubrica (Mller). Pilsbry & Johnson 1898; Nautilus 11:127.Pilsbry 1908; Man. Conch. 19:312.Hubricht 1985:6; map 24. Cionella lubrica (Mller). Pilsbry 1948; Land Moll. N. Amer. Acad. Nat. Sci. Phila. 105:165.Type Locality.Denmark. Distribution.Holarctic realm; widespread in North America (Hubricht 1985). CHIHUAHUA: about 8 km W of Colonia Juarez; talus of cliff on the Rio Piedras Verdes, about 3.6 km below Pacheco, 5900 ft. alt.; Sierra de la Brea, 17.5 km from Pearson, on road to Pacheco, 7000 ft. alt. (Pilsbry 1953). NUEVO LEN: near the Hacienda Pablillo (Pilsbry 1953) Superfamily PUPILLOIDEA Turton 1831 Family STROBILOPSIDAE Wenz 1915 Genus Strobilops Pilsbry 1893Strobila Morse 1856; Jour. Portland Soc. Nat. Hist. 1:26 (not Strobila Sars 1835; not Strobila Sodoffsky 1837). Strobilops Pilsbry 1893; Proc. Acad. Nat. Sci. Phila. 44:403 (substitute name for Strobila Morse).Pilsbry 1927; Man. Conch. 28:12, 56.Zilch 1959:177.Type Species.Helix labyrinthica Say 1817. Distribution.North America south to Brazil, Islas Galpagos, Korea, China and Japan. Taxonomy.Five subgenera are recognized. Two subgenera, Strobilops s. s., and Discostrobilops, occur in the study area. A third subgenus from the Cayman Islands, Coelostrobilops, is included in this list because of its geographic proximity to Central America. Subgenus Strobilops Pilsbry 1893 Distribution.North America, Mxico, Central America, south to Brazil. Taxonomy.Twelve species are recognized. Seven species occur in the study area. Strobilops (Strobilops) aenea mexicana Pilsbry 1927Strobilops aenea mexicana Pilsbry 1927; Man. Conch. 28; 32; Phila. 105:165.Correa-Sandoval, Garca-Cubas & Reguero 1998; Acta Zool. Mex. (73):14. Mxico. Holotype ANSP 77194a. Distribution.NUEVO LEN: Diente; Monterey; along the Rio Maurisco, ca. 25 km S of Monterey (Pilsbry 1953). PUEBLA: Necaxa (Pilsbry 1927). SAN LUS POTOS: 6 km Sandoval et al. 1998). Strobilops ( Strobilops) californica Miller & Christensen 1980Strobilops californica Miller & Christensen 1980; Proc. Biol. 15 (map).Type Locality.La Laguna, Sierra de la Victoria, along slope above creek immediately behind La Laguna shelter, Baja California Sur, Mxico; 6500 ft. alt. Holotype USNM 799595. Distribution.Known only from the type locality. Strobilops (Strobilops) hannai Pilsbry 1931Strobilops hannai 3, 3a, 5 (shell).Type Locality.Isla Socorro, Islas Revillagigedo, Nayarit, Mxico. Holotype ANSP 256587a. Distribution.Known only from the type locality.

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88 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Strobilops (Strobilops) piratica Pilsbry 1930Strobilops piratica Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. 82; Type Locality.On the summit of the ridge north of High Peak, Isla de Providencia, Colombia. Holotype ANSP 150860. Distribution.Known only from the type locality. Strobilops (Strobilops) (Tristram 1863) Thristram 1863; Proc. Zool. Soc. Lond. 31:411. (Tristram). Von Martens 1892; Biol. Cent. (Tristram). Pilsbry 1927; Man. Conch. 28:37; Type Locality.Mountain forests of northern Guatemala, [Dept. Alta Verapaz]. Distribution.Known only from the type locality. Strobilops (Strobilops) strebeli strebeli (Pfeiffer 1862)Helix strebeli (shell). Strobilops strebeli (Pfeiffer). Pilsbry 1927; Man. Conch. 28:33; Strobila labyrinthica (Say). Strebel 1880; Beitrag. Mex. Landund Biol. Cent. Amer.:173.Type Locality.Mirador, Veracruz, Mxico. Distribution.VERACRUZ: Mirador; environs of Veracruz (Strebel 1880). Strobilops ( Strobilops) strebeli guatemalensis Hinkley 1920Strobilops strebeli guatemalensis Hinkley 1920; Nautilus 34:52.8 (shell).Type Locality.In beach debris of Lago de Izabal, Jocolo, Dept. Izabal, Guatemala. Holotype ANSP 9272a. Distribution.Known only from the type locality. Strobilops (Strobilops) Pilsbry 1927Strobila labyrinthica Strebel 1880; Beitrag. Mex. Landund Sssw.Helix labyrinthica Say 1817). Pilsbry 1927; Man. Conch. 28:35; pl. Helix labyrinthica Strebel).Type Locality.Neighborhood of Veracruz, Mxico. Distribution.Known only from the type locality. Strobilops (Strobilops) Pilsbry 1927Helix strebeli 7b (shell) (not Helix strebeli Pfeiffer 1862). Pilsbry 1927; Man. Conch. 28:36 Helix strebeli Fischer & Crosse).Type Locality.Mirador, Veracruz, Mxico. Distribution.Known only from the type locality. Subgenus Coelostrobilops Pilsbry 1931Coelostrobilops Pilsbry 1931; Man. Conch. 28; 60.Type Species.Strobilops wenziana Pilsbry 1930. Distribution.Grand Cayman Island. Taxonomy.A single species is recognized. Strobilops (Coelostrobilops) wenziana Pilsbry 1930Strobilops wenziana Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. Type Locality.About midway between North Sound and Red Bay, Grand Cayman Island. Holotype ANSP 150861. Distribution.Known only from the type locality. Subgenus Discostrobilops Pilsbry 1927Discostrobilops Pilsbry 1927; Man. Conch. 28:18, 46.Pilsbry 1948; Land Moll. N. Amer. 2:865.Morrison 1953; Nautilus 65:55.Type Species.Helix hubbardi A. D. Brown 1861. Distribution.Gulf Coastal Plain from Mxico northeast to Georgia and Florida; also Bermuda, Cuba, Jamaica, and the Bahamas Islands; Northwest Mxico, Sonora (?). Taxonomy.Three extant species are recognized. Two occur in Mxico. Strobilops (Discostrobilops) hubbardi (A. D. Brown 1861)Helix hubbardi A. D. Brown 1861; Proc. Acad. Nat. Sci. Phila.:333; Strobilops (Discostrobilops) hubbardi (Brown). Pilsbry 1927; Man. Garca-Cubas & Reguero 1998:14.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):236. Helix vendryesiana Gloyne 1872; Jour. de Conchyl. 19:333. Strobilops hubbardi (Gloyne). Gulick 1904; Proc. Acad. Nat. Sci. Phila. 56:413.Hinkley 1907; Nautilus 21:76.Pilsbry 1948; (shell). (Gloyne). Pilsbry 1931; Man. Pilsbry 1899; Proc. Acad. Nat. Sci. Phila. 51:404.Type Localities. Helix hubbardi: Indianola, Calhoun Co., Texas; holotype ANSP 124. : Jamaica. : mangrove swamp, Biscayne Bay near State Agricultural Station, Miami, Florida. Distribution.The Gulf Coastal Plain from northeastern Mxico to Florida and Georgia; Jamaica, Cuba, Bimini Island and Bermuda. SAN LUS POTOS: Valles (Pilsbry 1931); km 48 on road from Cd. Valles to Agua Buena; 900 m alt. Sandoval et al. 1998). TAMAULIPAS: river drift, Tampico

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 89 W) (Correa-Sandoval & Rodriguez 2002). Strobilops (Discostrobilops) sinaloa Morrison 1953Strobilops (Discostrobilops) sinaloa, Morrison 1953; Nautilus 67; Proc. Biol. Soc. Wash. 93:595.Type Locality.Sinaloa (?). Holotype USNM 592719. Distribution.Known only from the type locality. It is questionable about whether this species actually came from Sinaloa (Morrison 1953). Family VALLONIIDAE Morse 1864 Genus Vallonia Risso 1820Vallonia Risso 1820; Hist. Nat. Eur. Merid., 4:101.Pilsbry 1948; Land Moll. N. Amer. 2:1019.Type Species.Vallonia rosalia Risso 1826 (= Helix costata Mller 1774). Distribution.North America, Europe and North Africa, northern and central Asia, Japan. (Pilsbry 1948). One occurs naturally in Mxico. Vallonia gracilicostata Reinhardt 1883Vallonia gracllicostata Reinhard 1883; Sitzung-Ber. Ges. Naturforsch. Freude Berlin:42.Pilsbry 1948; Land Moll.N. Type Locality.North Dakota, Little Missouri River. Distribution.Rocky Mountains, northwest, south and northeastern United States, northeastern Mxico. NUEVO LEN: Iturbide (Correa-Sandoval 1999). Sterki 1892 Sterki 1892; Nautilus 6:77.Sterki 1893; in Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. 105:165.Type Locality.Woodville, Jackson Co., Alabama. Distribution.Widespread across temperate North America, barely entering northern Mxico. CHIHUAHUA: Sierra de la Brea, 11 mi. from Pearson, on road to Pacheco, 7000 ft. alt.; in cliff talus along the Rio Piedras Verdes, 8 km W of Colonia Juarez; on the Rio Piedras Verdes, 3 km E of Pacheco (Pilsbry 1953). Family VERTIGINIDAE Stimpson 1851 Taxonomy.Fifty-one species and nine subspecies occur in the study area. Subfamily TRUNCATELLININAE Steenberg 1925 Genus Columella Westerlund 1878Edentulina Clessin 1876 (not Edentulina Pfeiffer 1855). Columella Westerlund 1878; Fauna Europaea 2:193.Pilsbry 1926; Man. Conch. 27:232. Sphaeradium Westerlund 1887; Fauna Palearctischen Region Binnenconchylien 3:125 (not Sphaeradium as limited by Von Martens 1860).Type Species.Pupa edentula Draparnaud 1805. Distribution.The Palearctic region; North America as far south as Nicaragua, Hawaii. Taxonomy.About ten species are recognized. Four occur in North America and Central America. One is known from the study area. (Pilsbry 1894) Pilsbry 1894; Proc. Acad. Nat. Sci. Phila. 46:31; Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:769. (Pilsbry). Pilsbry 1926; Man. Conch. 27:246; Holotype ANSP 5096. Distribution.Known only from the type locality. Subfamily PUPILLINAE Turton 1831 Genus Chaenaxis Pilsbry & Ferriss 1906Chaenaxis Pilsbry & Ferriss 1906; Proc. Acad. Nat. Sci. Phila. 58:145.Pilsbry 1916; Man. Conch. 24:1.Pilsbry 1948; Land Moll. N. Amer. 2:916.Bequaert & Miller 1973:172.Type Species. Pilsbry & Ferriss 1906. Distribution.Arizona, Sonora. Taxonomy.A single species is recognized. Chaenaxis tuba (Pilsbry & Ferriss 1906) Pilsbry & Ferriss 1906; Proc. Acad. Nat. Sci. Phila. Chaenaxis tuba (Pilsbry & Ferriss). Pilsbry 1916; Man. Conch. McCoy & Sisk 1964; Southwestern Natur. 9:104.Bequaert & Miller 1973:172.Naranjo-Garcia 1991:168. intuscostata Clapp 1908; Nautilus 22:76, 96; pl. 7, Chaenaxis intuscostata (Clapp). Pilsbry 1916; Man. Conch. 24:3; Chaenaxis intuscostata form Pilsbry 1916; Man. Chaenaxis sonorensis Pilsbry 1953; Proc. Acad, Nat. Sci. Phila. Type Localities.: drift debris of the San Pedro River, one mile east of Benson, Arizona; holotype ANSP 87062. intuscostata : foothills of the Plomos Range, about 8 miles east of Quartzite, Yuma County, Arizona; holotype Carnegie Museum 5769. Chaenaxis intuscostata form Tempe, Arizona; holotype ANSP 48527. Chaenaxis sonorensis: lower slopes of hills on south side of the Rio Sonora, about 0.75 km south of Hermosillo, Sonora, Mxico; 900 ft. alt. Holotype ANSP 167627. Distribution.SONORA: drift of the Rio Magdalena, Magdalena; northern foothills of Cerro Zapoxa, E of Cd. Obregon (Pilsbry 1953); drift of Rio de Bavispe, 21 mi. E

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90 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) 111 W); Sierra del Santo Nio, N of Mina del Milagro, on W); Cerro Prieto, ca. 1.5 km W of Ejido 18 de Agosto (Naranjo-Garcia 1991). Genus Pupilla Leach 1828Pupilla Leach, in Fleming 1828; British Animals:268.Pilsbry 1921; Man. Conch. 26:152.Type Species.Pupa marginata Draparnaud 1801 (= Pupa muscorum Linnaeus 1758). Distribution.Temperate North America, Europe, Africa, Asia and Australia. Taxonomy.Five subgenera are recognized. Two occur in the study area. Subgenus Pupilla Leach 1828 Distribution.Temperate North America, Europe, Africa, Asia and Australia. Taxonomy.About thirty species are recognized. One species occurs in the study area. Pupilla (Pupilla ) hebes (Ancey 1881)Pupa hebes Ancey 1881; Le naturaliste 3:389. Pupilla hebes (Ancey). Pilsbry & Ferriss 1911; Proc. Acad. Nat. Sci. Phila. 63:197. Pupilla (Pupilla ) hebes (Ancey). Pilsbry 1921; Man. Conch. 1953; Proc. Acad. Nat. Sci. Phila. 105:164.Bequaert & Miller 1973:180.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:107. Pupa arizonensis Gabb, in Binney 1885; Man. Amer. land shells:173. Pupilla muscorum idahoensis Henderson & Daniels 1917; Proc. Acad. Nat. Sci. Phila. 69:57. Pipilla hebes nefas Pilsbry & Ferriss 1910; Proc. Acad. Nat. Sci. Phila. 62:135.Type Locality.Pupa hebes: White Pine, White Pine Co., Nevada. Distribution.Washington, Idaho and Wyoming south to New Mexico and Arizona, and extreme northwestern Mxico. BAJA CALIFORNIA NORTE: Sierra San Pedro Martir 2.2 mi. below astronomical observatory, 2700 m alt.; rockslide in a ravine 1.3 km below astronomical observatory, 2700 m alt. (Smith et al. 1990). CHIHUAHUA: Sierra de la Brea, 17.5 km from Peason on road to Pacheco, 7000 ft. alt; along the Rio Piedras Verdes, 3.5 km below Pacheco (Pilsbry 1953). Subgenus Striopupilla Pilsbry 1921Striopupilla Pilsbry 1921; Man. Conch. 26:153.Type Species.Pupa sterkiana Pilsbry 1890. Distribution.Baja California Norte, Mxico. Taxonomy.Three species are recognized, all in the study area. Pupilla (Stiopupilla ) goniodon Pilsbry 1927Pupilla goniodon Pilsbry 1927; Proc. Calif. Acad. Sci. (4) 16:172; Proc. Calif. Acad. Sci. 47:107.Type Locality.Northwest Anchorage, Isla Guadalupe, Baja California Norte, Mxico. Holotype CAS 2574. Distribution.BAJA CALIFORNIA NORTE: Isla Guadalupe; Isla San Benito del Oeste (Smith et al. 1990). Pupilla (Stiopupilla ) guadalupensis Pilsbry 1927Pupilla guadalupensis Pilsbry 1927; Proc. Calif. Cad. Sci, (4) Pupilla (Striopupilla ) guadalupensis Pilsbry.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:108.Type Locality. feet above Northeast Anchoraga, Isla Guadalupe, Baja California Norte, Mxico. Holotype CAS 2575. Distribution.BAJA CALIFORNIA NORTE: known only from Isla Guadalupe (Smith et al. 1990). Pupilla (Striopupilla) sterkiana (Pilsbry 1890)Pupa sterkiana Pilsbry 1890; Proc. Acad. Nat. Sci. Phila. 41:411; pl. Pupilla (Striopupilla ) sterkiana (Pilsbry). Pilsbry 1921; Man. Nat. Sci. Phila. 115:202.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:107.Type Locality.San Ramon, Baja California Norte, Mxico. Lectotype ANSP 60466a (Baker 1963:202). Distribution.BAJA CALIFORNIA NORTE: San Ramon; along ocean beach N and S of Bahia San Quintin; W coast of San Quintin Peninsula, W of Winton Hill; Ensenada (Smith et al. 1990). Subfamily NEASOPUPINAE Steenberg 1925 Distribution.Madeira, South Africa, Australia, Oceania, southern North America south to Brazil. Taxonomy.Seven genera are recognized. Four genera occur in the study area. Genus Bothriopupa Pilsbry 1898Bothriopupa Pilsbry 1898; Nautilus 11:119.Pilsbry 1917; Man. Conch. 24:235.Pilsbry 1948; Land Moll. N. Amer. 2:1010 1011.Type Species. Gould 1848. Distribution.Northern South America, Florida, the

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 91 Greater Antilles and Guatemala. Taxonomy.About eight species are recognized. Five are recorded from the study area. Pilsbry 1917 Pilsbry 1917; Man. Conch. 24:230; pl. 28, Type Locality.Mountains west of Livingston, Dept. Izabal, Guatemala. Holotype ANSP 107534. Distribution.GUATEMALA, Dept. Izabal: mountains of Rio Cavech, back of Cavech Village (Hinkley 1920). Bothriopupa conoidea (Pfeiffer 1853)Pupa conoidea Pfeiffer 1853; Monographia heliciorum Viventium, 3:533. Bothriopupa conoidea (Pfeiffer). Pilsbry 1917; Man. Conch. Type Locality.Demerara. Distribution.Northern South America and Nicaragua. Bothriopupa leucodon (Morelet 1851)Pupa leucodon Morelet 1851; Test. Noviss. 2:13.Fischer & Crosse 2b (shell). Bothriopupa leucodon (Morelet). Pilsbry 1917; Man. Conch. 1937:30.Type Locality.Salama, Dept. Baja Verapaz, Guatemala. Distribution.GUATEMALA: known only from the type locality; no precise locality (Goodrich & van der Schalie 1937). Bothriopupa tenuidens (C. B. Adams 1841)Pupa tenuidens C. B. Adams 1841; Proc. Bost. Soc. Nat. Hist. 1845:15.Jacobson & Boss 1973; Occ. Pap. on Moll., 9:500; Bothriopupa tenuidens (C. B. Adams). Pilsbry 1917; Man. Conch. (distribution map).Type Locality.Jamaica. Lectotype MCZ 270000 (Jacobson & Boss 1973:500). Distribution.Jamaica, Puerto Rico, Cuba, Venezuela and Nicaragua. NICARAGUA, Dept. Managua: Lago de (Gould 1848) Gould 1848; Proc. Bos. Soc. Nat. Hist. 3:40. (Gould). Pilsbry 1917; Man. Conch. 24:230; Canegie Inst. Wash. (457):65.Type Locality.Florida. Distribution.Recorded from Florida and Yucatn. YUCATN: Chotch Cenote, 2 mi. NE of Piste (Bequaert & Clench 1936). Genus Pupisoma Stoliszka 1873Pupisoma Stoliszka 1873; Jour. Asiatic Soc. Bengal, 42:32.Pilsbry 1920; Man. Conch. 26:19.Baker 1927; Proc. Acad. Nat. Sci. Phila. 79:223 (anatomy).Pilsbry 1948; Land Moll. N. Amer. 2:1006.Type Species.Pupisoma lignicola Stoliszka 1873. Distribution.Burma, India, South Africa, Japan, Sri Lanka, Penang, India, Indonesia, Australia, New Caledonia, Abyssinia, Gulf coastal area of the United States south to southern Brazil. Taxonomy.Five subgenera are recognized (Zilch 1959). One occurs in the study area. Subgenus Ptychopatula Pilsbry 1889Ptychopatula Pilsbry 1889; Proc. Acad. Nat. Sci. Phila. 41:191.Pilsbry 1889; Nautilus 3:62.Zilch, 159:173.Type Species.Helix caeca Guppy 1868 (= Pupisoma dioscoricola (C. B. Adams 1845). Distribution.Tropical and subtropical Africa, Asia, Australia and the Americas. Taxonomy.Numerous species. Six species and two subspecies occur in the study area. Pupisoma (Ptychopatula ) bailyi Pilsbry 1934Pupisoma (Ptychopatula ) bailyi Pilsbry 1934; Man. Conch. 28:116; Type Locality.Cuernovaca, Moreles [Morelos]. Holotype ANSP 156099. Distribution.Known only from the type locality. Pupisoma (Ptychopatula ) comicolense H. B. Baker 1927Pupisoma comicolense Baker 1927a; Proc. Acad. Nat. Sci. Phila. Baker 1930a; Occ. Pap. Mus. Zool. Univ. Mich. (220):3; pl. Type Locality.Near Necaxa, Puebla, Mxico. Holotype UMMZ 159752. Distribution.PUEBLA: known only from the immediate vicinity of the type locality (H. B. Baker 1930a). Pupisoma (Ptychopatula ) dioscoricola dioscoricola (C. B. Adams 1845)Helix dioscoricola C. B. Adams 1845; Proc. Boston Soc. Nat. Hist. 2:16.Jacobson & Boss 1963; Occ Baker 1963; Proc. Acad. 11 (lectotype). Helix punctum Morelet 1851; Test. Noviss. II:9. Helix caeca Guppy 1868; Proc. Sci. Assos. Trinidad:241.Fischer &

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92 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Pupisoma dioscoricola (Adams). Pilsbry 1920; Man. Conch. 26:36 Phila. 78:133.Pilsbry 1948; Land Moll. N. Amer. 2:1007 map). Type Localities. Helix dioscoricola: Dublin Castle, Graywick, Jamaica; lectotype MCZ 275929 (Jacobson & Boss 1973:516). Helix punctum: Merida, Yucatn, Mxico. Helix caeca : Trinidad. Distribution.Florida and southern Texas to Brazil, Islas Galpagos. PANAM, Prov. Panam: Juan Mina (Pilsbry E of Cayal; Cd. de Carmen (Thompson 1967). QUINTANA ROO: 7.1 mi. NNW of Xiatil (Thompson 1967). TABASCO: Cardenas (Thompson 1967). YUCATN: Tikul; 0.8 mi. NE of Becanchen (Thompson 1967). Pupisoma ( Ptychopatula) dioscoricola insigne Pilsbry 1920Pupisoma dioscoricola insigne Pilsbry 1920; Man. Conch. 26:39 Baker 1930a; Occ. Pap. Mus. Zool. Univ. Mich. (220):2.Bequaert & Clench 1936; Pub. Canegie Inst. Wash. (457):65.8 (shell).Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):12.Correa-Sandoval 1997; Rev. Biol. Trop. 44/45:140.Correa-Sandoval et al. 1998:14.Correa-Sandoval 2000; Acta Zool. Mex. (79) 8.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (80:235.Correa-Sandoval & Rodriguez 2005; Acta Zool. Mex. 21:59.Type Locality.Brownsville, Texas. Holotype ANSP 109013. Distribution.Eastern Mxico south to Venezuela. GUATEMALA, Dept. Petn: Tikal (Basch 1956). NUEVO LEN: numerous localities in eastern region of the state (Correa-Sandoval et al. 1998). TAMAULIPAS: numerous localities in the southern part of the state (Correa-Sandoval & Rodriguez 2002); El Cielo Biosphere Reserve, (CorreaSandoval & Rodriguez 2005); numerous localities in southern part of state (Correa-Sandoval & Rodriguez 2005). PUEBLA: Necaxa (Baker 1930a). SAN LUIS POTOS: Rio Valles falls; Rio Ganina; Rio Choy Cave (Pilsbry 1920). VERACRUZ: Coatotolapan (Baker 1923a); Hwy. Tuxpan-Poza Rica km. YUCATN: Izamal (Pilsbry 1920); Santa Ana Cenote, Villadolid (Bequaert & Clench 1936). Pupisoma (Ptychopatula ) mediamericanum Pilsbry 1920Pupisoma mediamericanum Pilsbry 1920; Man. Conch. 26:42; Zool. Univ. Mich. (220):4.Pilsbry 1948; Land Moll. N. Amer. Type Locality.Chama, Dept. Alta Verapaz, Guatemala. Holotype and paratypes ANSP 28270; syntypes ANSP 45727. Distribution.GUATEMALA; only from the type locality. PUEBLA: Necaxa (Baker 1927a). VERACRUZ:500 ft. above Orizaba (Pilsbry 1920); Peuela; Sumidero, 2625 3400 ft. alt. (Baker 1930a). Pupisoma (Ptychopatula ) michoacanensis Pilsbry 1920Pupisoma michoacanensis Pilsbry 1920; Man. Conch. 26:40; pl. 4, (shell).Type Locality.Morelia, Michoacn, Mxico. Holotype ANSP 77119. Distribution.Known only from the type locality. Pupisoma (Ptychopatula ) minus Pilsbry 1920Pupisoma minus 11 (shell).Pilsbry 1948; Land Moll. N. Amer. 2:1009; 8:168.Correa-Sandoval et al.. 198; Acta Zool. Mex. 73:14.Correa-Sandoval & Rodriguez 2005; Acta Zool. Mex. 21:59.Type Locality.Snapper Creek, south of Coconut Grove, Dade County, Florida. Holotype ANSP 113390. Distribution.NICARAGUA, Dept. Boaco; Dept. EMALA, Dept. Alta Verapaz: Chama (Pilsbry 1920). NUEVO LUS POTOS: numerous localities in the eastern region of the state (Correa-Sandoval et al. 1998). SONORA: Cerro de Oro, 0.25 mi. from Cerro de Oro, on road to Rayon, 600 m Genus Nesopupa Pilsbry 1900Nesopupa Pilsbry 1900; Proc. Acad. Nat. Sci. Phila. 52:432.Pilsbry & Cooke, in Pilsbry 1920; Man. Conch. 25:274.Type Species.Pupa tantilla Gould 1847. regions, St. Helena Island. Taxonomy.Eleven subgenera are recognized. One subgenus occurs in the study area. Subgenus Cocopupa Pilsbry & Cooke 1920Cocopupa Pilsbry & Cooke 1920; Man. Conch. 25:323.Type Species.Vertigo cocosensis Dall 1900. Distribution.Isla del Coco, Costa Rica. Taxonomy.A single species is known. Nesopupa (Cocopupa) cocosensis (Dall 1900)Vertigo cocosensis Dall 1900; Proc. Acad. Nat. Sci. Phila. 52:98, pl. Nesopupa (Cocopupa) cocosensis (Dall). Pilsbry & Cooke, in (shell).Type Locality.Isla del Coco, Costa Rica. Distribution.Known only from the type locality.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 93 Genus Sterkia Pilsbry 1898Sterkia Pilsbry 1898; Nautilus 11:119.Pilsbry 1920; Man. Conch. 26:49.Pilsbry 1948; Land Moll. N. Amer. 2:1012.Type Species.Pupa calamitosa Pilsbry 1889. Distribution.California south to Baja California Sur; northern South America, Guatemala, the West Indies and south Florida. Taxonomy.Two subgenera are recognized. The genus includes six species, all of which occur in the study area. Subgenus Sterkia Pilsbry 1898 Distribution.Southern California and Baja California Norte. Taxonomy.Two species are recognized. Both occur in Baja California Norte. Sterkia (Sterkia) calamitosa calamitosa (Pilsbry 1889)Pupa calamitosa 16 (shell). Sterkia calamitosa (Pilsbry). Pilsbry 1920; Man. Conch. 26:57; pl. Type Locality.Near the mouth of the Rio San Tomas. Holotype ANSP 11602. Distribution.BAJA CALIFORNIA NORTE: Ensenada de Todos Santos (Pilsbry 1948); hills W of Bahia San Quintin (Smith et al. 1990). Sterkia (Sterkia) calamitosa martiniana Pilsbry 1927Sterkia (Sterkia ) calamitosa martiniana Pilsbry 1927; Proc. Calif. Amer. 2:1014.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Type Locality.Isla San Martin, Baja California Norte, Mxico. Distribution.BAJA CALIFORNIA NORTE: Isla San Martin. Sterkia (Sterkia) hemphilli (Sterki 1890)Pupa hemphilli Sterkia (Sterkia ) hemphilli (Sterki). Pilsbry 1920; Man. Conch. Acad. Sci. (4):16:187.Pilsbry 1948; Land Moll. N. Amer. Type Locality.Banks of the Rio San Tomas, Baja California Norte, Mxico. Holotype Carnegie Museum 62.20384. Distribution.Southern California south to Baja California Norte. BAJA CALIFORNIA NORTE: Ensenada de Todos Santos; W shore of Bahia san Quintin; Punta Abreojos (Smith et al. 1990). Subgenus Metasterkia Pilsbry 1920Metasterkia Pilsbry 1920; Man. Conch. 26:50.Pilsbry 1948; Land Moll. N. Amer. 2:1014.Type Species.Sterkia antilliensis Pilsbry 1920. Distribution.Northern South America, the West Indies, Florida, Guatemala and Baja California Norte. Taxonomy.Four species are recognized. All occur in the study area. Sterkia (Metasterkia) antillensis Pilsbry 1920Sterkia (Metasterkia ) antillensis Pilsbry 1920; Man. Conch. 26:53 Type Locality.El Abra, Viales Prov., Cuba. Distribution.Cuba, Jamaica, Venezuela and Nicaragua. Sterkia (Metasterkia) bakeri Pilsbry 1921Sterkia bakeri 3 (shell).Baker 1923a:3.Baker 1963; Proc. Acad. Nat. Sci. Phila. 115:200.Ty pe Locality.Hacienda de Coatotolapan, between the Rio San Juan and its tributary the Arroyo Hueyapan, Canton de Acayucan, Veracruz, Mxico. Lectotype ANSP 141375 (H. B. Baker 1963:200). Distribution.Known only from the type locality. Sterkia (Metasterkia) eyriesii eyreisii (Drouet 1859)Pupa eyriesii Drouet 1859; Essai sur les mollusques terrestres et Sterkia (Metasterkia) eyriesii eyreisii (Druuet). Pilsbry 1920; Man. der Schalie 1937; Misc. Pub. Mus. Zool. Univ. Mich. (34):30.(shell).Type Locality.Ilet-la-Mre, Guyana. Distribution.French Guyana, Surinam, Trinidad and Guatemala. GUATEMALA, no precise locality (Goodrich & van der Schalie 1937). Dept. Alta Verapaz: Cham (Pilsbry 1920). Taxonomy.Two subspecies are recognized. Sterkia eyriesii rhoadsi Pilsbry 1920, occurs in Florida. Sterkia (Metasterkia) clementina (Sterki 1890)Pupa clementina Vanatta 1916; Nautilus 30:48. Sterkia clementina (Sterki). Pilsbry 1920; Man. Conch. 26:54; pl. 7, 540:9, 10, 13 (shell).Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:110.Type Locality.Northeast Anchorage, Isla Guadalupe, Baja California Norte, Mxico. Holotype Carnegie Museum 62.20392. Distribution.BAJA CALIFORNIA NORTE: Northeast Anchorage, Isla Guadalupe. Subfamily PUPOIDINAE Iredale 1940 Distribution.Europe, Africa, Asia, Australia, Hawaii

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94 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) and North America Taxonomy.Three extant genera are recognized. Genus Pupoides Pfeiffer 1854Pupoides Pfeiffer 1854; Malacologische Bltter, 1:102.Pilsbry 1921; Man. Conch. 26:108, 138.Pilsbry 1948; Land Moll. N. Amer. 2:921. Leucochila Von Martens 1860; Die Heliceen:205. Leucochiloides Pfeiffer 1878; Nomenel. Helic. Viv.:292.Type Species.Pupoides: Bulimus nitidulus Pfeiffer 1839. Leucochila: Cyclostoma marginata Say 1821. Leucochiloides : Bulimus lardeus Pfeiffer 1852. Distribution.All continents. Taxonomy.Four subgenera are recognized. Two subgenera occur in the study area. Three species occur in the study area. Subgenus Pupoides Pfeiffer 1854 Distribution.North America, northern Asia, Europe, and North Africa. Taxonomy.A large number of species have been named. Seven occur in North America, two of which are found in the study area. Pupoides (Pupoides) albilabris (C. B. Adams 1841)Cyclostoma marginata Say 1821; Jour. Acad. Nat. Sci. Phila. 2:172 (not Cyclostoma marginatus G. Fischer 1807). Pupoides maerginatus Pilsbry & Vanatta 1900; Proc. Acad. Nat. Sci. Phila. 52:586.Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. (shell). Pupa fallax Say Gould 1843; Boston Jour. Nat. Hist. 4:357. Pupa albilabris C. B. Adams 1841; Amer. Jour. Sci., 40:271. (new name for Cyclostoma marginata Say). Pupoides albilabris (C. B. Adams 1841). Pilsbry 1948; Land Moll. Proc. Acad. Nat. Sci. Phila. 105:164.Bequaert & Miller 1973:78, 176.Smith, Miller, Christensen & Roth Garcia 1991:168.Type Locality.Upper Missouri. Distribution.Widely distributed in North America. BAJA CALIFORNIA NORTE: Isla Angel de la Guardia; Isla San Lorenzo (Smith et al. 1990). BAJA CALIFORNIA SUR: numerous localities (Smith et al. 1990). SINALOA: Rio Fuerte, San Blas (Pilsbry 1953). SONORA: Rio Mayo, at Navajo; Rio Yaqui, 11 mi. N of Cajeme; Cerro Zaporxa, E of Cameje; Arroyo San Rafael, San Bernardo (Pilsbry 1953); N W); Arroyo Cerro de Oro, ca. 0.25 mi. from Cerro de Oro, on Garcia 1991). CHIHUAHUA: Presa Chihuahua, 7 mi. from Cd. Chihuahua (Bequaert & Miller 1973). NUEVO LEN: Topo Chico, near Monterey (Pilsbry 1904). TAMAULIPAS: of Padilla (Bequaert & Miller 1973). Pupoides (Pupoides) catalinensis Hanna 1923Pupoides catalinensis Hanna 1923; Proc. Calif. Acad. Sci. (4) Type Locality.Isla Santa Catalina, Baja California Sur, Mxico. Distribution.BAJA CALIFORNIA SUR: known only from Isla Santa Catalina (Smith et al. 1990). Subgenus Ischnopupoides Pilsbry 1926Ischnopupoides Pilsbry 1926; Man. Conch. 27:250.Pilsbry 1948; Land Moll. N. Amer. 2:924.Type Species.Bulimus cordatus Pfeiffer 1856. Distribution.SW United States, western Mxico, and Bolivia. Taxonomy.Three species are recognized. One occurs in the study area. Pupoides (Ischnopupoides) cordatus (Pfeiffer 1856)Bulimus cordatus Pfeiffer 1856; Malak. Bltter 3:46. Pupa cordata (Pfeiffer). Binney & Bland, Land and Freshwater Pupoides (Pupoides) cordatus (Pfeiffer). Von Martens; 1893:330.(shell).Type Locality.Mazatln, Sinaloa, Mxico. Distribution.Known only from the type locality. Subfamily GASTROCOPTINAE Pilsbry 1918 Distribution.Australia, Africa, North and South America, Asia, Sri Lanka, the Philippines, and Oceania. Taxonomy.A single extant genus, Gastrocopta, is recognized. Two South American genera are referred questionably to the subfamily. Genus Gastrocopta Wollaston 1878Gastrocopta Wollaston 1878; Testacea Atlantica :515.Pilsbry 1916; Man. Conch. 24:6. Leucochilus Boettger 1881; Conchologische Mittheilungen, 1:64 (not Leucochila Von Martens 1860). Sterki, in Pilsbry 1891; Proc. Acad. Nat. Sci. Phila. 43:315. Sterki 1893; Nautilus 6:101.Type Species.Gastrocopta: Pupa acarus Benson 1856. Leucochilus: Pupa armifera Say 1821. : Pupa Gould 1843. : Pupa hordeacea Gabb of Sterki, = Pilsbry & Vanatta 1900. Distribution.Worldwide. Taxonomy.Nine subgenera are recognized (Zilch 1959:160). Subgenus Gastrocopta Wollaston 1878 Distribution.Tropical and warm portions of America, Africa, Mascarene Island, Sri Lanka and the Philippines. Taxonomy.Numerous species. Four occur in the study area.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 95 Gastrocopta ( Gastrocopta) cristata (Pilsbry & Vanatta 1900) Pilsbry & Vanatta 1900; Proc. Acad. Nat. Gastrocopta (Gastrocopta) cristata (Pilsbry & Vanatta). Pilsbry 8.Branson, McCoy & Sisk 1964; S.W. Nat., 9:104.Bequaert & Miller 1973:169.Type Locality.Camp Verde, Yavapai County, Arizona; 3200 ft. Alt. (Bequaert & Miller 1973). Distribution.Nebraska, Kansas, Oklahoma, west Texas, New Mexico and Arizona and adjacent NW Mxico. BAJA CALIFORNIA NORTE. SONORA: drift of Rio Bevispe, 21 mi. S of Agua Prieta; drift of Rio Yaqui, 4 mi. N of Cd. Obregon (Branson et al. 1964); drift of Rio Sonoyta, Sonoyta (Bequaert & Miller 1973). Gastrocopta (Gastrocopta) pellucida hordeacella (Pilsbry 1890)Pupa hordeacella Pilsbry 1890; Proc. Acad. Nat. Sci. Phila. 44; pl. (Pilsbry). Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:766. Gastrocopta pellucida hordeacella (Pilsbry).Pilsbry 1916; Man. 495 (Shell).Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):12. Sterki 1899; Nautilus 12:128. (Sterki). Pilsbry 1948; Land Moll. Sci. (4) 13:515. Pupa pellucida Pfeiffer 1841. Fischer & Crosse 1873; Miss. Sci. Mex. I:311.Strebel 1880:91.Von Martens 1898:328 (in part). Gastrocopta pellucida (Pfeiffer). Hinkley 1920; Nautilus 34:52.Rehder 1966; Proc. Biol. Soc. Wash. 79:383.Thompson 1967; Bull. Fla. St. Mus. 11:231.Bequaert & Miller 1973:79.Naranjo-Garcia 1991; Amer. Malac. Bull., 8:167.CorreaSandoval 1993; Rev. Biol. Trop. 41:675.Correa-Sandoval 1997; Rev. Biol. Trop. 44/45:140.Correa-Sandoval, GarcaCubas & Reguero 1998; Acta Zooll. Mex., (73):134.CorreaSandoval 2000; Acta Zool. Mex. (79):8.Correa-Sandoval, & (distribution map in Nicaragua).Correa-Sandoval & Salazar 2005; Acta Zool. Mex. 21:59.Type Localities. Pupa pellucida : Cuba (Pilsbry 1916). Pupa hordeacella : New Braunfels, Texas; holotype and paratypes ANSP 60460. : around Jerome, Yavapai County, Arizona; syntypes in the Carnegie Museum. Distribution.Widely distributed in the West Indies, Mxico, and Guatemala. Bequaert and Miller (1973) stated that the species apparently has a continuous distribution over Mxico. NICARAGUA, Dept. Managua: Lago de Xiloa GUATEMALA, Dept. Alta Verapaz: Chama (Hinkley 1920). Dept. Baja Verapaz: Salama. Dept. Guatemala: Barranca de Villalobos, near Cd. Guatemala. Dept. Petn: Tikal (Basch 1959). Dept. Sacatepquez: Antigua, 4500 ft. alt. (Von Martens 1898). BAJA CALIFORNIA SUR: numerous localities (Smith et al. 1990). CAMPECHE: 7.1 mi. SW of Cd. Camopeche; 8.1 mi. SW of Champoton (Thompson 1967). NUEVO LEN: numerous records for southern part of state (Correa-Sandoval & Salazar 2005). QUINTANA ROO: Tulm (Rehder 1966). SAN LUIS POTOS: Cd. Valles (Pilsbry 1916); numerous localities (Correa-Sandoval et al. 1998). SONORA: numerous localities (Naranjo-Garcia 1991). TABASCO: San Juan Bautista (Von Martens 1898). TAMAULIPAS: Cd. Victoria (Pilsbry 1916); Tampico (Hinkley 1907); El Cielo Biosphere Reserve (CorreaSandoval & Rodriguez 2005). YUCATN: Sisal; Merida (Fischer & Crosse 1873); 1.0 mi. SSE of Telchac; 0.8 mi. NE of Becanchen (Thompson 1963). VERACRUZ: Antigua (Pilsbry 1916); Veracruz (Strebel 1880); Rancho El Sol, Sandoval 1999). Taxonomy.Recognition of Gastrocopta pellucida hordeacella (Pilsbry 1890) as a valid subspecies is controversial. Bequaert and Miller (1973) synonymized both hordeacella and with Gastrocopta pellucida pellucida (Pfeiffer 1841). Later, Smith et al. (1990) resurrected hordeacella as a distinct subspecies. (Clapp 1899) is a synonym of hordeacella I follow these authors by assigning all mainland populations to hordeacella Gastrocopta (Gastrocopta) riograndensis (Pilsbry & Vanatta 1900) Pilsbry & Vanatta 1900; Proc. Acad. Nat. Sci. Phila. 52:596. Gastrocopta (Gastrocopta) riograndensis (Pilsbry & Vanatta). 10 (shell).Thompson 1967; Bull. Fla. St. Mus. 11:231.Neck 1980; Veliger 23:180.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:106. (Gould). Hinkley 1907; Nautilus 21:77.Type Locality.Lower Rio Grande valley, river drift at Hidalgo, Hidalgo Co., Texas. Holotype ANSP 60137. Distribution.Southwest Texas, northern Mxico. BAJA CALIFORNIA SUR: Juncalito (Smith et al. 1990). CAMPECHE: 8.1 mi. SW of Chamopoton (Thompson 1967). SAN LUIS POTOS: falls below Cd. Valles; canyon below Los Canoas (Pilsbry 1916). TAMAULIPAS: Rio Panuco Valley, Tampico (Hinkley 1907; Pilsbry 1916). YUCATN:1.0 mi. SSE of Puerto Techac (Thompson 1967). Taxonomy.The identity of specimens from Baja California Sur is uncertain (Smith et al. 1990). Gastrocopta (Gastrocopta) (Gould 1843)

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96 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)14 (shell). Pupa pellucida Pfeiffer (in part Gastrocopta (Gastrocopta) (Gould). Pilsbry 1916; Man. Proc. Acad. Nat. Sci. Phila. 78:133.Rehder 1966; Proc. Biol. Soc. Wash. 79:283.Thompson 1967; Bull. Fla. St. Mus. Type Locality.Matanzas, Cuba. Distribution.Generally distributed throughout the West Indies, including Bermuda; scattered records in Central America and Mxico. PANAM, Canal Zone: Isla Barro Colorado. Prov. Panam: Juan Mina (Pilsbry 1926). Antigua. Dept. Baja Verapaz: Salama. Dept. Guatemala: Barranca de Villalobos, near Cd. Guatemala. CAMPECHE: Isla de Carmen (Pilsbry 1916). MORELOS: Yautepec. QUINTANA ROO: Tulm (Rehder 1966). VERACRUZ: Veracruz, on the beach (Strebel 1880). YUCATN: Merida (Pilsbry 1916). Subgenus Albinula Sterki 1892Albinula Sterki 1892; Nautilus 6:101.Pilsbry 1916; Man. Conch. 24:13.Pilsbry 1948; Land Moll. N. Amer. 2:874.Type Species.Pupa contracta Say 1822. Distribution.Eastern North America. Taxonomy.Six species are recognized. One occurs in the study area. Gastrocopta (Albinula) contracta contracta (Say 1822)Pupa contracta Say 1822; Jour. Acad. Nat. Sci. Phila, 2:374.Pilsbry 1891; Proc. Acad. Nat. Sci. Phila. 43:315. (Say). Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:766.Hinkley 1907; Nautilus 21:78. Gastrocopta (Albinula ) contracta contracta (Say). Pilsbry 1916; 1953; Proc. Acad. Nat. Sci. Phila. 105:161.Correa-Sandoval, Garca-Cubas & Reguero 1998:14.Type Locality.Occoquan, Virginia. Distribution.Eastern North America from Ontario west to Manitoba and south to Florida and Veracruz; Cuba and Jamaica. MORELOS: Yautepec (Pilsbry 1891, 1904). NAYARIT: drift debris of Rio Fuerte, San Blas (Pilsbry 1953). NUEVO LEN: on Rio Mauricio, ca. 25 km S of Monterey (Pilsbry 1953). SAN LUS POTOS: Cascadas de Tamasopo, et al. 1998). SONORA: Arroyo San Rafael, San Bernardo (Pilsbry 1953). TAMAULIPAS: Tampico (Hinkley 1907). VERACRUZ: Texolo; near Orizaba (Pilsbry 1904). Subgenus Geminidens Pilsbry 1930Geminidens Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. 82:351.Zilch 1959:160.Type Species.Bothriopupa geminidens Pilsbry 1917. Distribution.Venezuela, Trinidad, and Panam. Taxonomy.The subgenus includes a single species. Gastrocopta (Geminidens) geminidens (Pilsbry 1917)Bothriopupa geminidens Pilsbry 1917; Man. Conch. 24:228; Sci. Phila. 115:201. Gastrocopta geminidens (Pilsbry). Pilsbry 1926; Man. Conch. 27:228. Gastrocopta (Geminidens) geminidens (Pilsbry). Pilsbry 1930; Proc. Acad. Nat. Sci., Phila., 80:351.Haas 1960:13.Thompson & Type Locality.Cariaquita, [Estado Sucre], Venezuela. Holotype ANSP 105200a (Baker 1963:201). Distribution.Known from Panam, Venezuela, Margarita Island and Trinidad. PANAM, Prov. Panam: Old Panam City (Pilsbry 1930). COSTA RICA: Dept. del Rio Subgenus Immersidens Pilsbry & Vanatta 1900Immersidens Pilsbry & Vanatta 1900; Proc. Acad. Nat. Sci. Phila. 52:606.Type Species. 1898. Distribution.Central Plateau of North America south to Brazil and Argentina. Taxonomy.The subgenus contains numerous species. Seven species and one subspecies occur in the study area. Gastrocopta ( Immersidens) allyni Roth & Christensen 1984Gastrocopta (Immersidens) allyni Roth & Christensen 1984; (distribution map).Smith, Miller, Christensen & Roth 1990; Type Locality.In drift, arroyo 24 km south of La Paz, Baja California Sur, Mxico. Holotype CAS 32993. Distribution.BAJA CALIFORNIA SUR: southern tip of peninsula, numerous locality records (Smith et al. 1990). Gastrocopta (Immersidens) ashmuni (Sterki 1898) Sterki 1898; Nautilus 12:90.Pilsbry & Ferriss 1906; Proc. Acad. Nat. Sci. Phila. 58:144. Gastrocopta (Immersidens) ashmuni (Sterki). Pilsbry 1916; Man. Bequaert & Miller 1973:159.Naranjo-Garcia 1991:166 167.Correa-Sandoval & Rodriguez 2005:59.Type Locality.Santa Rita Mountains, on east slope of range, 20 miles west of Crittenden, Santa Cruz County, Arizona (Bequaert & Miller 1973:159).

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 97 Distribution.CHIHUAHUA: Rio Piedras Verdes, ca. 1953); Presa Chihuahua, 7 mi. S of Cd. Chihuahua, 4800 W) (Correa-Sandoval & Rodriguez 2005). SONORA: Rio Nacozari, 7 mi. S of Nacozari (Branson et al 1964); near Nacozari, 3550 ft. alt.; Nogales; Sierra de Santo Nio, N of Mina El Milagro, on road to Sahuaipa, 4000 t. alt. (ca. N, 110 W); Rancho Pinos Altos, Sierra Nacori, 5800 ft. alt. arroyo, along Rayon-Cerro de Oro road, ca. 4.2 mi. from (Naranjo-Garcia 1991). Gastrocopta (Immersidens) cochisensis (Pilsbry & Ferriss 1910) Pilsbry & Ferriss 1910; Proc. Acad. Nat. Sci. Gastrocopta (Immersidens) cochisensis (Pilsbry & Ferriss). Pilsbry 486 (shell).Bequaert & Miller 1973:163.Type Locality.Tanner Canyon [now Garden Canyon], Huachuca Mountains, Arizona. Holotype and paratypes ANSP 97444. Distribution.CHIHUAHUA: Rio Piedras Verdes, below Pacheco, 5900 ft. alt.; Sierra de la Brea, ca. 17.5 mi. from Mata Ortiz, on road to Pacheco, 7000 ft. alt. (Pilsbry 1953). SINALOA: Rio Fuerte, San Blas (Pilsbry 1953). SONORA: arroyo, 8 km S of Guaymas (Pilsbry 1953). Gastrocopta (Immersidens) dalliana dalliana (Sterki 1898) Sterki 1898; Nautilus 12:91. Gastrocopta dalliana (Sterki). Pilsbry 1916; Man. Conch. 24:49; Gastrocopta (Immersidens) dalliana dalliana (Sterki). Branson, McCoy & Sisk 1964:104.Bequaert & Miller 1973:165.Naranjo-Garcia 191:167. Gastrocopta dalliana media Pilsbry 1916; Man. Conch. 24:50; pl. Type Localities. dalliana : Ephraim Canyon, Nogales, Santa Cruz County, Arizona; 3800 ft. alt. (Bequaert & Miller 1973). Gastrocopta dalliaa media: Montezuma Wells, near Rimrock, Yavapai County, Arizona; holotype ANSP 82918. Distribution.CHIHUAHUA: cliff talus on N side of Rio Piedras Verdes, 8 km above Colonia Juarez, 5900 ft. spring-fed march 60 mi. S of Nogales; San Carlos bay, 10 mi. (Bequaert & Miller 1973); S edge of Sierra El Pinito, ca. 10.1 (Naranjo-Garcia 1991). Gastrocopta (Immersidens) dalliana bilamellata (Sterki & Clapp 1909) Sterki & Clapp 1909; Nautilus 22 126; pl. 8, Gastrocopta bilamellata (Sterki & Clapp). Pilsbry 1916; Man. (shell). Gastrocopta (Immersidens) dalliana bilamellata (Sterki & Clapp). Bequaert & Miller 1973:168.Naranjo-Garcia 1991:167.Type Locality.Foothills of Plomosa Range, about 8 miles east of Quartzite, Yuma County, Arizona. Holotype in the Clapp Collection, Carnegie Museum. Distribution.Southwest Arizona and northwest Sonora. SONORA: S side of Rio Sonora, near Hermosillo, 900 ft. alt. (ca. 29 N, 111 W); Arroya San Rafael, San Bernardo, 900 Zaporxa, E of Obregon (ca. 28 N, 109 W); drift in wash W of Pinacate Peak, ca. 45 mi. NW of Sonoyta, 1200 ft. alt. (ca. Gastrocopta ( Immersidens) gularis Thompson & Lpez 1996Gastrocopta (Immersidens) gularis map). Type Locality.Lago de Xiloa, Dept. Managua, Distribution.COSTA RICA, Prov. Guanacaste: Parque Juan del Rio Gastrocopta (Immersidens) prototypus (Pilsbry 1899) Pilsbry 1899; Proc. Acad. Nat. Sci. Phila.

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98 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)51:400.Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:766; 115:202. Gastrocopta (Immersidens) prototypus (Pilsbry). Pilsbry 1916; Gastrocopta (Immersidens) prototypus prototypus (Pilsbry). Bequaert & Miller 1973:164. Pilsbry & Ferriss 1910; Proc. Gastrocopta (Immersidens) prototypus basidentata Pilsbry 1916; Type Localities. : Huingo, near Lago de Cuitzco, 40 km NW of Morelia, Michoacan (Bequaert & Miller 1973); Lectotpe ANSP 77201a (Baker 1963:202). ; Ash canyon, SE slope of Huachuca Mountains, Cochis County, Arizona, 5000 ft. alt. (Bequaert & Miller 1973:165). Gastrocopta prototypus basidentata : Guatemala City, Guatemala. Distribution.Arizona and widely scattered localities south to Guatemala. GUATEMALA, Dept. Guatemala: Cd. Guatemala. JALISCO: Guadalajara (Pilsbry 1916). MICHOACAN: Huingo. Gastrocopta (Immersidens) rixfordi Hanna 1923Gastrocopta rixfordi Hanna 1923; Proc. Calif. Acad. Sci. (4) Gastrocopta (Immersidens) rixfordi Hanna. Pilsbry 1926; Man. Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:106; Gastrocopta dalliana dalliana (Sterki). Bequaert & Miller 1973:167 (in part; Baja California Sur records).Type Locality.Isla Monserrat, Gulf of California, Baja California Sur, Mxico. Holotype CAS 1094. Distribution.BAJA CALIFORNIA SUR: numerous locality records (Smith et al. 1990). Subgenus Sterki 1893 Sterki 193; Nautilus 6:101.Pilsbry 1948; Land Moll. N. Amer. 2:893.Type Species.Odostomia corticaria Say 1816. Distribution.Eastern North America from Maine and Ontario west to Minnesota and south to Florida and Louisiana; northeastern Mxico. Taxonomy.A single species is recognized. Gastrocopta ( ) corticaria (Say 1816)Odostomia corticaria Say 1816; Nicholsons American Edition Gastrocopta ( ) corticaria (Say). Pilsbry 1916; Man. Garca-Cubas & Reguero 1998:14.Type Locality.Philadelphia, Pennsylvania. Neotype ANSP 64524a (Pilsbry 1948). Distribution.SAN LUS POTOS: km 82, on road from W); km 140 on road from Cd. del Maz-Antigua Morelos, (Correa-Sandoval et al. 1998). Subgenus Vertigopsis Sterki 1893Vertigopsis Sterki 1893; Nautilus 6:101.Pilsbry 1948; Land Moll. N. Amer. 2:886.Type Species. Gould (= Vertigo pentodon Say 1821). Distribution.Eastern North America west to Alberta, Canada and Arizona, and south to Nicaragua. Taxonomy.The subgenus includes four species. Two occur in the study area. Gastrocopta (Vertigopsis) pentodon (Say 1821)Vertigo pentodon Say 1821; Jour. Acad. Nat. Sci. Phila. 2:376. Gastrocopta (Vertigopsis) pentodon (Say). Pilsbry 1916; Man. 41 (shell).Hinkley 1920; Nautilus 34:39, 48, 52.Baker 1930a:5.Goodrich & van der Schalie 1937:30.Pilsbry 1948; Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. 105:161.Bequaert Correa-Sandoval & Rodriguez 2005:59. (Gould). Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:766.Type Locality.Pennsylvania. Distribution.NICARAGUA, Dept. Jinotega; Dept. no precise locality (Goodrich & van der Schalie 1937). Dept. Alta Verapaz: Chama. Dept. Izabal: Jocolo; Quirigua (Hinkley 1920). CHIHUAHUA: talus of cliffs on Rio Piedras Verdes, ca. 4.5 km below Pacheco (Pilsbry 1953). NUEVO LEN: on Rio Maurisco; near the Hacienda Pablillo, above Galeana, 24 mi. N of Padillo (Bequaert & Miller 1973); 0.5 km. W & Salazar 2005). PUEBLA: Necaxa (Baker 1930a). SAN LUIS POTOS: bluffs N of San Dieguito (Pilsbry 1916). TAMAULIPAS: canyon 4 mi. W of Cd. Victoria (Pilsbry 1904 1916). Taxonomy.Synonyms that apply to populations outside of the study area are: Gould 1841, Pupa cincinnatiensis Judge 1878, Dall 1885, Pupa montanella Cocherell 1889 and var. gracilis Sterki 1890 (Pilsbry 1948); and Pupa tappaniana C. B. Adams 1842 (Bequaert & Mlller 1973). Gastrocopta (Vertigopsis) pilsbryana (Sterki 1890)Pupa pilsbryana Sterki 1890; Nautilus 3:123. Gastrocopta pilsbryana (Sterki). Pilsbry 1916; Man. Conch. 24:36 1953; Proc. Acad. Nat. Sci. Phila. 105:161.Branson, McCoy & Sisk 1964; Southwestern Natur. 9:104. Bequaert & Miller 1973:158.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 99Gastrocopta soneri Chamberlin & Jones 1929; Bull. Univ. Utah Gastrocopta pilsbryana amissidens Pilsbry 1934; Man. Conch. Type Localities. Pupa pilsbryana: from the Colorado River, Arizona; holotype Carnegie Museum 62.20382. Gastrocopta stoneri: Cedar City, Utah. Gastrocopta pilsbryana amissidens: San Francisco Mountains, Arizona; holotype ANSP 161437. Distribution.Utah, Arizona, New Mexico and transPecos Texas south to northern Mxico. CHIHUAHUA: Sierra la Brea, ca. 17.7 km by road from Pearson, on road to Pacheco, ca. 7000 ft. alt.; along Rio Piedras Verdes, 4.5 km below Pacheco, 5900 ft. alt. (Pilsbry 1953). SONORA: (Branson et al. 1964). TAMAULIPAS: (Bequaert & Miller 1973). Subfamily VERTIGININAE Stimpson 1851 Type Genus.Vertigo Mller 1774. Taxonomy.A single extant genus is recognized (Zilch 1959:148). Genus Vertigo Mller 1774 Type Species.Vertigo pusilla Mller 1774. Distribution.Practically the entire Holarctic Realm from sea level to 10,000 feet altitude (Pilsbry 1948). Taxonomy.Five subgenera are recognized. Three occur in the study area. Subgenus Vertigo Mller 1774 Distribution.As for the genus. Taxonomy.About 80 species are recognized. Three species and three subspecies occur in the study area. Vertigo (Vertigo) berryi Pilsbry 1919Vertigo berryi (shell). Vertigo (Vertigo) berryi Pilsbry. Pilsbry 1948; Land Moll. N. Amer. Type Locality.Mill Creek canyon, San Bernardino Mountains, California; 4600 ft. alt. Holotype ANSP 105166. Distribution.California and Nevada south to Baja California Norte. BAJA CALIFORNIA NORTE: Valle Trinidad (Pilsbry 1948). Vertigo ( Vertigo) gouldi arizonensis Pilsbry & Vanatta 1900Vertigo coloradoensis arizonensis Pilsbry & Vanatta 1900; Proc. Vertigo (Vertigo) coloradoensis arizonensis Pilsbry & Vanatta. Vertigo (Vertigo) gouldi arizonensis Pilsbry & Vanatta. Pilsbry Bequaert & Miller 1973:186.Type Locality.Top of Mingus Mountain, Yavapai Co., Arizona, 7600 ft. alt. Holotype ANSP 119010. Distribution.CHIHUAHUA: Sierra de la Brea, ca. 16 mi. SW of Colonia Juarez (Bequaert & Miller 1973). Vertigo (Vertigo) gouldi coloradoensis (Cockerell 1891)Pupa coloradoensis Cockerell 1891; British Naturalist:100. Virtigo coloradoensis (Cockerell). Sterki 1892; Nautilus 6:5.Pilsbry 1919; Man. Conch. 25:115. Vertigo (Vertigo) gouldi coloradoensis (Cockerell). Pilsbry 1948: Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. 105:164.Bequaert & Miller 1973:184.Type Locality.Near Swift Creek, Custer Co., Colorado. Distribution.Colorado, Utah and Arizona south to northwestern Mxico. CHIHUAHUA: along the Rio Piedras Verdes, 4.6 km from Pearson, 5900 ft. alt. (Pilsbry 1953). Vertigo (Vertigo) gouldi inserta Pilsbry 1919Vertigo coloradoensis inserta Pilsbry 1919; Man. Conch. 25; 118; Vertigo (Vertigo) gouldi inserta Pilsbry. Pilsbry 1948; Land Moll. N. Sci. Phila. 105:164.Bequaert & Miller 1973:187.Type Locality.Bear Wallow, Santa Catalina Mountains, Pima Co., Arizona. Holotype ANSP 109559. Distribution.CHIHUAHUA: Sierra de la Brea, ca. 16 mi. SW of Colonia Juarez (Bequaert & Miller 1973). Vertigo (Vertigo) Say 1822Vertigo Say 1822 Jour. Acad. Nat. Sci. Phila. 2:375. Vertigo (Vertigo) (Say). Pilsbry 1919; Man. Conch. 25:83 4, 7 (shell).Bequaert & Miller 1973:183.Smith, Miller, 13 (map).Correa-Sandoval et al. 1998:14. (Say). Von Martens 1898; Biol. Cent. Amer.:327.Type Locality.Philadelphia, Pennsylvania. Distribution.Widespread in North America and the West Indies. BAJA CALIFORNIA NORTE: On road to La Playa, 0.3 km SSE of San Jos del Cabo; Sierra Laguna (Smith et al. 1990). SONORA: drift of Rio Sonoyta, at Sonoyta, 5 mi. S of the Arizona border (Bequaert & Miller 1973). VERACRUZ: Veracruz (Von Martens 1898; Pilsbry 1919, questions this record). SAN LUS POTOS: Hwy. San Subgenus Alloptyx Pilsbry 1953Alloptyx Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. 105:164.Type Species.Vertigo hinkleyi Pilsbry 1921. Distribution.Arizona and Chihuahua. Taxonomy.A single species is recognized. Vertigo (Alloptyx ) hinkleyi Pilsbry 1921Vertigo hinkleyi 542:12 (shell). Vertigo (Alloptyx ) hinkleyi Pilsbry. Pilsbry 1953; Proc. Acad. Nat.

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100 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Sci. Phila. 105:164.Type Locality.Cave Canyon near the reservoir, Huachuca Mountains, Arizona. Holotype ANSP 46263. Distribution.CHIHUAHUA; Sierra de la Brea on the road to Pacheco, 17.5 km from Pearson, 7000 ft. alt. (Pilsbry 1953) Subgenus Angustula Sterki 1888Angustula Sterki 1888; Proc. U. S. National Museum, 11:378.Pilsbry 1919; Man. Conch. 25:145.Pilsbry 1948; Land Moll. N. Amer. 2:944.Type Species.Pupa milium Gould 1840. Distribution.Eastern North America, west to South Dakota, Colorado and Arizona, Jamaica, Hispaniola and Bermuda. Taxonomy.Two species are recognized. One occurs in the study area. Vertigo (Angustula) milium (A. A. Gould 1840)Pupa milium Gould 1840; Boston Journal of Natural History, 3:402. Vertigo milium (Gould). Binney 1878; Terr. Moll., 5:215; pl. 71, Type Locality.Oak Island, Chelsea, near Boston, Massachusetts. Distribution.Eastern North America, west to South Dakota, Colorado and Arizona, Jamaica and Hispaniola (Pilsbry 1948). NICARAGUA: common in the hilly TAMAULIPAS: Tampico (Hinkley 1907). Genus Nearctula Sterki 1892Nearctula Sterki 1892; Nautilus 8:5.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:109.Type Species.Pupa californica Rowell,1861 = Pupa rowelli Newcomb 1860 (Smith et al. 1990). Distribution.San Francisco, California south to Bahia San Quintin, Baja California Norte. Taxonomy.The genus includes two species and several subspecies. Two species and three subspecies occur in the study area. Nearctula rowelli diegoensis (Sterki 1890)Pupa californica var. diegoensis Sterki 1890; Nautilus 4:18. Vertigo californica diegoensis (Serki). Pilsbry 1918; Man. Conch. Nearctula rowelli diegoensis (Sterki). Smith, Miller, Christensen & Type Locality.False Bay near Asher Station, San Diego, California. Distribution.BAJA CALIFORNIA NORTE: San Ramon; Isla San Martin (Pilsbry 1948); Bahia Todos Santos; near Bahia San Quintin; W shore of Bahia San Quintin; hills W of Bahia San Quintin (Smith et al. 1990). Nearctula rowelli catalinaria (Sterki, 1890)Vertigo californica catalinaria Sterki 1890; Nautilus 4:8.Pilsbry Nearctula rowelli catalinaria (Sterki). Smith, Miller, Christensen & Type Locality.Santa Catalina Island, Chanel Islands, California. Distribution.California Chanel Islands south along the coast to northern Baja California Norte. BAJA CALIFORNIA NORTE: Isla Guadalupe, about 1000 ft above the landing at Northeast Anchorage; ca. 2 mi. N of S end of island on E side; Sierra San Pedro Martir, along road to astronomical observatory area, 2700 m alt. (Smith et al. 1990). Nearctula rowelli guadalupensis (Pilsbry 1927)Vertigo californica guadalupensis Pilsbry 1927; Proc. Calif. Acad. Nearctula rowelli guadalupensis (Pilsbry). Smith, Miller, Christensen & Roth 1990:109.Type Locality.Isla Guadalupe, Baja California Norte, Mxico. Holotype CAS 2582. Distribution.Known only from the type locality. Nearctula degeneris (Pilsbry 1927)Vertigo degeneris Pilsbry 1927; Proc. Calif. Acad. Sci. (4) 16:175; Nearctula degerneris (Pilsbry). Smith, Miller, Christensen & Roth 1990:110.Type Locality.About 1000 feet above the landing at Northeast Anchorage, Isla Guadalupe, Baja California Norte, Mxico. Holotype CAS 2583. Distribution.Known only from the type locality. Superfamily ORTHALICOIDEA Albers 1860 Family ORTHALICIDAE Albers 1860 Subfamily ORTHALICINAE Albers 1960 Distribution.Tropical South America, the West Indies, south Florida, Central America, and Mxico. Taxonomy.The nomenclature employed in this study follows Breure and Schouten (1985). Six genera are recognized in the subfamily Orthalicinae. One occurs in Central America and Mxico. Genus Orthalicus Beck 1837Orthalicus Beck 1837. Von Martens 1893; Biol. Cent. Amer.:179 182.Rehder 1945; Nautilus 59:29.Richardson 1993:97 117. Oxystyla Schluter 1838.Pilsbry 1899; Man. Conch. 12:101. Zebra Shuttleworth 1858.Type Species.Buccinum zebra Mller 1774 (Rehder 1945). Distribution.From Bolivia and Brazil north to Mxico, the West Indies, and south Florida. Within the Central American-Mexican region the genus generally occurs below

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 101 1500 m altitude. Orthalicus occurs in Mxico at lower elevations along the east coast from southeastern Tamaulipas south to Isthmus of Tehuantepec, northern Chiapas, the Yucatn Peninsula and the Petn region of Guatemala. On the west coast, the genus occurs from central Sinaloa southeast to Chiapas and Guatemala, but it occurs at higher elevations than on the east coast. Orthalicus continues south through Central America, generally at less than 1000 m altitude. Taxonomy.Strebel (1909) monographed the Orthalicinae. He described numerous species and subspecies, many of which were based on specimens from undocumented localities. In several instances he ignored nomeclatural within different species, and employed unconventional taxonomic nomenclature. In many instances he divided single specimen lots into varieties, subspecies and even species. In other instances he placed under a given species name separated localities. These actions caused many taxonomic problems, and a major review of the genus is needed. Tentatively, twenty-seven species plus twelve subspecies of Orthalicus are recognized in this study. Richardson (1993) provided a list of references for the species. Orthalicus boucardi Pfeiffer 1860Orthalicus boucardi Pfeiffer 1860; Proc. Zool. Soc. Lond. 28:138; 1993:98. Orthalicus boucardi-ponderosus (shell). (TL: Tehuantepec). Orthalicus ponderosus-boucardi (TL: Tehuantepec). Oxystyla longa var. boucardi (Pfeiffer). Pilsbry 1899; Man. Conch. Oxystyla longa var. strebeli Pilsbry 1899; Man. Conch. 12:128; pl. Zebra boucardi (Pfeiffer). Strebel 1909; Rev. Unterfam. Zebra boucardi heteromorph melanocheila Strebel 1909; Rev. Zebra boucardi heterom. xanthus Strebel 1909; Rev. Unterfam. Orthalicinen:74. Zebra boucardi heterom albinus Strebel 1909; Rev. Unterfam. Tehuantepec, Oaxaca). Zebra boucardi form adustus Strebel 1909; Rev. Unterfam. Zebra boucardi form zebriola Strebel 1909; Rev. Unterfam. Zebra boucardi form zoniferus Strebel 1909; Rev. Unterfam. Type Localities.As indicated above with TL. (Strebel 1909). OAXACA: Villa Alta; Tehuantepec; Cerro Negro, Tehuantepec; Sierra de Betaza (17.25 N, 96.15 W) (Von Martens 1893). Orthalicus decolor Strebel 1882Orthalicus decolor Strebel 1882; Beitrag. Mex. Landund Sssw.Oxystyla decolor (Strebel). Pilsbry 1899; Man. Conch. 12:131; Zebra mars form decolor (Strebel). Strebel 1909; Rev. Unterfam. Type Locality.Not stated. Distribution.Unknown. Remarks.It is doubtful if this taxon is part of the MxicoCentral American fauna. Strebel (1909) noted its similarity to Orthalicus mars (Pfeiffer 1861), an Ecuadorian species Orthalicus delphinus (Strebel 1909)Zebra delphinus Strebel 1909; Rev. Unterfam. Orthalicinen:30. Orthalicus delphinus (Strebel). Jacobson 1958; Amer. Mus. Nov. (1899):11.Richardson 1993:100. Zebra delphinus form nebulosus Strebel 1909; Rev. Unterfam. Oxystyla delphinus form nebulosa (Strebel). Dall 1926; Proc. Calif. Acad. Sci. (4) 15:473. Zebra delphinus form pumilio Strebel 1909; Rev. Unterfam. Orthalicinen:33. Oxystyla delphinus form nesiotica Dall 1926; Proc. Calif. Acad. Sci. Type Localities. Zebra delphinus: not given. Zebra delphinus form nebulosus: Mazatln, Sinaloa, Mxico. Zebra dephinus form pumilio: Mazatln, Sinaloa, Mxico. Oxystyla delphinus form nesiotica : Isla Mara Madre, Islas Maras, Nayarit, Mxico. Holotype CAS 2194. Distribution.NAYARIT: Isla Maria Magdelena (Dall 1926; Jacobson 1958). SINALOA: Mazatln (Strebel 1909). Orthalicus elegans Rolle 1895Orthalicus elegans Rolle 1895; Nachr. Deutsch. Malak. Gesell. 32:131.Neubert & Jannsen 2004; Arch. fr Moll. 133:236; pl. Orthalicus princeps elegans Rolle. Von Martens 1901; Biol., Cent. Oxystyla princeps var. elegans (Rolle). Pilsbry 1899; Man. Conch. (shell). Zebra elegans (Rolle). Strebel 1909; Rev. Unterfam. Orth.[alicus] colimensis Rolle 1895; Nachr. Deutsch. Malak. Gesell. 32:131.Type Locality.Colima, Mxico. Holotype in the ZMB. Orthalicus ferussaci ferussaci Von Martens 1863Orthalicus ferussaci Von Martens 1863; Monatsber. Akad. Wissen. Berlin 1863:542.Fischer & Crosse 1875; Miss. Sci. Mex.:447.(shell).Breure & Schouten 1985; Zool. Meded. Uitg. d. anatomy).Richardson 1993:101.Prez & Lopz 2002:223 224.

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102 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Oxystyla ferussaci (Von Martens). Pilsbry 1899; Man. Conch. Type Locality.Tehuantepec, Oaxaca, Mxico. Distribution.COSTA RICA, Prov. Guanacaste: Guanacaste; Salinas Bay. GUATEMALA, Dept. Alta Verapaz: Panzos; San Geronimo. Dept. Zacapa: Zacapa. NICARAGUA, Dept. Chontales: Acoyapa (Pilsbry 1899). MICHOACN: El Espinal (Breure & Schouten 1985). OAXACA: Tehuantepec. YUCATN: Shkolak (Pilsbry 1899). Orthalicus ferussaci tricinctus Von Martens 1893Orthalicus melanocheilus (Valenciennes). Fischer & Crosse 1875; Orthalicus ferussaci var. tricinctus Von Martens 1893; Biol. Cent. Oxystyla ferussaci var. tricincta (Von Martens). Pilsbry 1899; Man. Orthalicus maracaibensis tricinctus Von Martens. Richardson 1993:106. Distribution.COSTA RICA, Prov. Guanacaste: Quebrada Vijigual, Rio Saverey Valley, 150 m alt.; El Pozo, 50 m alt. Prov. Puntarenas: Boruco, 450 m alt.; Terraba, 250 270 m alt.; Alto de Mano Tigre, 690 m alt. NICARAGUA: no Orthalicus hackeri (Strebel 1909)Zebra hackeri Strebel 1909; Rev. Unterfam. Orthalicinen:50; pl. 7, Orthalicus hackeri (Strebel). Richardson 1993:103.Type Locality.Tepc, Nayarit, Mxico. Distribution.Known only from the type locality. Orthalicus leucocheilus Crosse & Fischer 1869Orthalicus leucocheilus Crosse & Fischer 1869; Journ. De Conchyl. 17:423.Fischer & Crosse 1875; Miss. Sci. Mex.:459; pl. 18, Oxystyla leucocheila (Fischer & Crosse). Pilsbry 1899; Man. Conch. Type Locality.Orizaba, Veracruz, Mxico. Distribution.Known only from the type locality. Shuttleworth 1856 Shuttleworth 1856; Notitiae Malacologicae 1:64; (shell).Von Martens 1893; Biol. Cent. Amer.:189.Richardson 1993:103.Neubert & Gosteli 2005; Contr. Nat. Hist. 5:23; pl. (Shuttleworth). Pilsbry 1899; Man. Conch. 12:118 (Shuttleworth). Strebel 1909; Rev. Unterfam. Type Locality.Mxico, probably near Veracruz. Distribution.GUATEMALA, Dept. Alta Verapaz: Cobn (Strebel 1909). VERACRUZ: exact locality unknown. Remarks.Beck (1837) is often credited as the author of the name but he only listed the name. 2005). Von Martens 1863 Von Martens 1863; Monatsber. Akad. Wiss. Berlin 1863:542.Fischer & Crosse 1875; Miss. Sci. Mex.:448.Strebel 1882; Beitrag. Mex. Landund Sssw.-Conch. V:29 1959; Occ. Pap. Mus. Zool. Univ. Mich. (611):8. (Von Martens). Pilsbry 1899; Man. Conch. 12:124 106 (shell).Type Locality.Volcn Jorullo, Michoacn, Mxico. Distribution.MICHOACN: N slope of Volcn Jorullo, 2700 ft. alt. (Solem 1959). SINALOA: Mazatln (Pilsbry 1899). Orthalicus longus Pfeiffer 1856Orthalicus longus Pfeiffer 1856; Malak. Bltt. 12:39.Fischer & Strebel 1882; Beitrag. Mex. Landund Sssw.-Conch. V:43; Amer.:189.Solem 1959; Occ. Pap. Mus. Zool. Univ. Mich. (611):6. Oxystyla longa (Pfeiffer). Pilsbry 1899; Man. Conch. (2) 12:126 Zebra longa (Pfeiffer). Strebel 1909; Rev. Unterfam. OrthType Locality.Unknown. Distribution.MICHOACN: Ario. MORELOS: Cuernavaca. OAXACA: Santa [Salina] Cruz Bay; San Geronimo; Tehuantepec; 10 mi. W of Tehuantepec; 20 mi. W of Tehuantepec. Solem (1959) listed San Geronimo, of which there are two such places in Oaxaca. Orthalicus maclurae Von Martens 1893Orthalicus maclurae Von Martens 1893; Biol. Cent. Amer.:188; pl. Oxystyla maclurae (Von Martens). Pilsbry 1899; Man. Conch. Zebra maclurae (Von Martens). Strebel 1909; Rev. Unterfam. Zebra maclurae form turrita Strebel 1909; Rev. Unterfam. Type Localities. Ortalicus maclurae: Cacao, Golfo de Fonseca, Nicaragua [Cacao, Dept. Choluteca, Honduras ?]. Zebra maclurae form turrita: unknown. Distribution.Known only from the type locality. COLIMA (Strebel 1909). Nayarit: Islas Maras (Strebel 1909). OAXACA: Tehuantepec (Strebel 1909). TRINIDAD (Strebel 1909). Remarks.See remarks under Orthalicus quagus. Orthalicus maculiferus (Strebel 1909)Zebra maculiferus Strebel 1909; Rev. Unterfam. Orthalicinen:30;

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 103 Orthalicus maculiferus ( Strebel). Richardson 193:105.Type Locality.Not stated. Distribution.Unknown. Orthalicus melanocheilus melanocheilus (Valenciennes 1833)Bulimus melanocheilus Orthalicus melanocheilus (Valenciennes). Von Martens 1865; Malak. Bltt. 12:45, 70.Von Martens 1893; Biol. Cent. & Schouten 1985; Zool. Meded. Uitg. d. Rijksmus. Nat. Hist. Oxystyla melanocheilus (Valenciennes). Pilsbry 1899; Man. Conch. Orthalicus zebra (Mller). Binney & Bland 1869; Land & Freshwater Type Locality.Arbitrarily restricted to northwestern Mxico by Von Martens (1865). 1895). GUERRERO:17 mi. SW of Acapulco (McGinty 1939); 19.3 mi. W of San Marcos, 300 ft. alt. (Breure & Schouten 1985). NAYARIT: Islas Maras (Pilsbry 1899). SINALOA: Mazatln (Pilsbry 1899). Orthalicus melanocheilus mariae (McGinty 1939)Oxystla melanocheilus mariae McGinty 1939; Nautilus 53:6; pl. 2, Type Locality.Puerto Marquez, near Acapulco, Guerrero, Mxico. Holotype ANSP 174048. Distribution.Known only from the type locality. Orthalicus muelleri ( Strebel 1909)Orthalicus zebra (Mller). Strebel 1882; Beitrag. Mex. Landund Zebra muelleri Strebel 1909; Rev. Unterfam. Orthalicinen:46. Orthalicus muelleri (Strebel). Richardson 1993:107.Type Locality.Mazatln, Sinaloa, Mxico. [Mazatln, Guerrero?]. Distribution.Known only from the type locality. Orthalicus nobilis Rolle 1895Orth.[alicus] nobilis Rolle 1895; Nachr. Deut. malak. Gesell. 27:131.Richardson 1993:107. Orthalicus zoniferus var. nobilis Rolle. Von Martens 1901; Biol. Oxystyla nobilis (Rolle). Pilsbry 1899; Man. Conch. 12:124. Oxystyla zoniferus var. nobilis (Rolle). Pilsbry 1902; Man. Conch. Zebra nobilis (Role). Strebel 1909; Rev. Unterfam. Type Locality.Colima, Mxico. locality. Orthalicus pallidus (Srebel 1909)Zebra pallida Strebel 1909; Rev. Unterfam. Orthalicinen:40; pl. 5, Orthalicus nobilis pallidus (Strebel). Richardson 1993:107.Type Locality.Colima, Mxico. Holotype SMF 26849. Distribution.Known only from the type locality. Orthalicus ponderosus ponderosus Strebel 1882Orthalicus ponderosus Strebel 1882; Beitrag. Mex. Landund Deut. malak. Gesell. 32:130.Solem 1959; Occ. Pap. Mus. Zool. Univ. Mich. (611):7.Richardson 1993:108. Oxystyla ponderosa (Strebel). Pilsbry 1899; Man. Conch. 12:130 Orthalicus boucardi-ponderosus Orthalicus ponderosus-boucardi Strebel 1882:132. Zebra mars (Pfeiffer). Strebel 1909; Rev. Unterfam. Zebra miles Strebel 1909; Rev. Unterfam. Orthalicinen:64; pl. Type Locality.Not stated. Distribution.COLIMA: Culata, near Manzanillo; Colima. GUERRERO: Dos Arroyos, 25 mi. NE of Acapulco, 1000 ft. alt. (Pilsbry 1899); vicinity of Acapulco (McGinty 1939). MICHOACN: Sierra de Coalcoman, trail from Pomare to Maruata, 700 ft. alt.; Sierra de Coalcoman, trail from Los Pozos to Ranch Quemado, 4500 ft. alt. (Solem 1959). NAYARIT: San Blas; Tepc. OAXACA: Tehuantepec (Pilsbry 1899). Remarks.Apparently Strebel took the species name Zebra miles from the type locality of miles NE of Acapulco, Guerrero. Orthalicus ponderosus albatus (McGinty 1939)Oxystyla ponderosa albata McGinty 1939; Nautilus 53:5; pl. 2, Type Locality.Vicinity of Acapulco, Guerrero, Mxico. Holotype ANSP 174049. Distribution.Known only from the type locality. Ortalicus ponderosus balesi (McGinty 1939)Oxystyla ponderosa balesi 2 (shell).Richardson 1993:97.Type Locality.Puerto Marquez, near Acapulco, Mxico. Holotype ANSP 174046. Distribution.Known only from the type locality. Orthalicus princeps princeps (Broderip 1833)Bulimus princeps Broderip 1833. Oxystyla princeps (Broderip). Pilsbry 1899; Man. Conch. 12:113 Richards 1939; Proc. Amer. Philos. Soc. 81:33.Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):10. Orthalicus princeps (Broderip). Von Martens 1893; Biol. Cent. Schalie 1940:5.Solem 1959; Occ. Pap. Mus. Zool. Univ.

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104 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Mich. (611):7.Rehder 1966; Proc. Biol. Soc. Wash. 79:287 288.Thompson 1967; Bull. Fla. St. Mus. 11:244.Breure & Schouten 1985; Zool. Meded. Uitg. d. Rijksmus. Nat. Hist. 1993:10911.Prez & Lopz 2002:225.O.[rthalicus] princeps var. crassiuscula Strebel 1909; Rev. Unterfam. Orthalicinen:23.Type Locality.Bulimus princeps: Conchagua, Dept. La Distribution.Widely distributed from Sinaloa along the west coast and from southeastern Tamaulipas along the east coast of Mxico south to Panam. COSTA RICA, Prov. Guanacaste: Guanacaste (Pilsbry 1899). GUATEMALA, Dept. Alta Verapaz: Cahabon; Cobn; Senahu; Panzos (Pilsbry 1899); Cham (van der Schalie 1940). Dept. Petn: Tikal National Park (Basch 1959). Dept. El Progreso: San Agustin (Pilsbry 1899). Dept. Quetzaltenango: Cerro Zunil (Pilsbry 1899). Dept. Retalhuleu: Retalhuleu; San Francisco Miramar; Capetillo; El Reposo (Pilsbry 1899). Dept. Suchitepequez: Mazatenango (Pilsbry 1899). NICARAGUA: Isla del Maz Versant (Prez & Lopz 2002). PANAM, Canal Zone: Isla Barro Colorado; between Tabernillo and San Pablo. Prov. Chiriqu: Bugaba; Quipo (Pilsbry 1926). EL SALVADOR, several localities (Thompson 1967). CHIAPAS: Monte Libano, 600 m alt.; Monte Libano to El Censo, 600 m alt.; San Lorenzo, midway between Ocosingo and El Real (Bequaert 1957); Escuintla (Solem 1959). COLIMA: Colima. NAYARIT: Tepc (Pilsbry 1899); Islas Maras of Orthalicus delphinus (Jacobson 1958:12). OAXACA: Panistlahuaca; Ixstapa; Cerro de Acatepec, near Tuletepec (Pilsbry 1899); 3.4 mi. WNW Zanatepec, ca. 275 ft. alt. (Breure & Schouten 1985). QUINTANA ROO: Isla Cozumel (Richards 1937; Rehder 1966). SAN LUIS POTOS: SINALOA: Mazatln; Presidio de Mazatln (Pilsbry 1899). TAMAULIPAS: localities in the southeastern part of the state. (Correa & Catstro 2002). VERACRUZ: Papantla; Misantla; Playa Vicente; Coatepec, above Jalapa; Almolonga, below YUCATN: Merida; Tabi; Shkolak; Tekanto; Ticul; Silam (Pilsbry 1899); Chichen Itza (Bequaert & Clench 1936). Orthalicus princeps crossei Von Martens 1893Orthalicus zoniferus crossei Von Martens 1893; Biol. Cent. Amer.:186.Haas & Solem 1960; Nautilus 73:131. Oxystyla princeps crossei (Von Martens). Pilsbry 1899; Man. Conch. Type Locality.Belize. Distribution.BELIZE: Rio Frio Cave, Double Fall (Haas & Solem 1960). Orthalicus princeps deceptor (Pilsbry 1899)Orthalicus obductus Shuttleworth. Fischer & Crosse; 1875:456 (exclusive of South American references and localities).Von Martens 1893; Biol. Cent. Amer.:187 (exclusive of South American references and localities). Zebra obductus (Shuttleworth). Strebel 1909; Rev. Unterfam. South American references and localities). Oxystyla princeps var. deceptor Pilsbry 1899; Man. Conch. 12:116 Sci. Phila. 78:341. Orthalicus princeps deceptor (Pilsbry). Richardson 1993:110. Lctotypa ANSP 5099a (H. B. Baker 1963:227). Distribution.BELIZE: Belize. GUATEMALA, Dept. Alta Verapaz: Cobn; San Geronimo. NICARAGUA, Dept. Von Martens 1893Orthalicus princeps var. Von Martens 1893; Biol. Cent. (Von Martens). Pilsbry 1899; Man. Nat. Sci. Phila. 82:342. (Von Martens). Richardson 1993:110.Type Locality.El Reposo and San Isidro, Dept. Suchitepequez, Guatemala. Distribution.Known only from the type locality. Orthalicus princeps perlongus (Pilsbry 1899)Oxystyla princeps var. perlonga Pilsbry 1899; Man. Conch. 12:117; Orthalicus princeps perlongus (Pilsbry). Richardson 1993:110.Type Locality.Panam. Distribution.PANAM: exact locality unknown. Orthalicus princeps trifractus (Pilsbry 1899)Oxystyla princeps var. trifracta Pilsbry 1899; Man. Conch. 12:115; Type Locality.El Salvador. Lectotype ANSP 28080 (H. Baker 1963:231). Distribution.Known only from the type locality. Orthalicus pseudolongus (Strebel 1909)Zebra pseudo-longus Strebel 1909; Rev. Unterfam. Type Locality.Unknown. Distribution.Unknown Orthalicus quagus (Strebel 1909)Zebra quaga Strebel 1909; Rev. Unterfam. Orthalicinen:41; pl. 6,

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 105Zebra quaga form turrita Strebel 1909; Rev. Unterfam. form aberrans Strebel 1909; Rev. Unterfam. (shell).Neubert & Jannsen 2004; Arch. fr Moll. 133:236; pl. Orthalicus quaga (Strebel). Solem 1959; Occ. Pap. Mus. Zool. Univ. Type Localities.Zebra quaga: not given. Zebra quaga form turrita: Colima, Mxico. form aberrans: Colima; syntype SMF 25454 (Neubert & Jannsen 2004). Distribution.COLIMA: (Strebel). MICHOACN: Cerro de Guzman, near Coalcoman (Solem 1959). Remarks.The name Zebra quaga form turrits (Strebel 1909) is a different taxon than Zebra maclurae form turrita (Strebel 1909) or Zebra sphinx form turrita (Strebel 1909:68). Orthalicus richardsoni (Strebel 1909)Zebra richardsoni Strebel 1909; Rev. Unterfam. Orthalicinen:36; Orthalicus richardsoni (Strebel). Richardson 1993:113.Type Locality.Tepc, Nayarit, Mxico. Distribution.Known only from the type locality. Orthalicus selectus (Strebel 1909)Zebra selectus Strebel 1909; Rev. Unterfam. Orthalicinen:37; pl. Orthalicus selectus (Strebel). Richardson 1993:113.Type Locality.Trinidad. Distribution.GUATEMALA, Dept. Alta Verapaz: Cobn (Strebel 1909). Remarks.The inclusion of this taxon in the Central American fauna is questionable. Orthalicus sphinx sphinx (Strebel 1909)Zebra sphinx Strebel 1909; Rev. Unterfam. Orthalicinen:66; pl. Orthalicus sphinx (Strebel 1909).Richardson 1993:113.Type Locality.Tepc, Nayarit, Mxico. Distribution.Known only from the type locality. Orthalicus sphinx latistriatus (Strebel 1909)Zebra sphinx form latistriata Strebel 1909; Rev. Unterfam. Orthalicus sphinx latistriatus (Strebel). Richardson 1993:113.Type Locality.Not stated. Distribution.Unknown. Orthalicus sphinx turrita (Strebel 1909)Zebra sphinx form turrita Strebel 1909; Rev. Unterfam. Orthalicinen:67. Orthalicus sphinx turrita (Strebel). Richardson 1993:113.Type Locality.An island above San Blas, Nayarit, Mxico. Distribution.Known only from the type locality. Remarks.See remarks under Orthalicus quagus. Orthalicus tepicensis tepicensis (Strebel 1909)Zebra tepicensis Strebel 1909; Rev. Unterfam. Orthalicinen:45; pl. Orthalicus tepicensis (Strebel). Richardson 1993:113.Type Locality.Tepc, Nayarit, Mxico. Distribution.Known only from the type locality. Orthalicus tepicensis rollei (Strebel 1909)Zebra tepicensis form rollei Strebel 1909; Rev. Unterfam. Orthalicus tepicensis rollei (Strebel). Richardson 1993:113.Type Locality.Colima, Mxico. Distribution.Known only from the type locality. Orthalicus torrei (McGinty 1939)Oxystyla torrei McGinty 1939; Nautilus 52:93. McGinty 1939; Type Locality.Vicinity of Acapulco, Guerrero, Mxico. Holotype ANSP 173342. Distribution.Known only from the type locality. Orthalicus uhdeanus Von Martens 1893 var. uhdeanus Von Martens 1893; Biol. Cent. Amer.:189. var. uhdeanus (Von Martens). Pilsbry 1899; Man. Zebra uhdeanus (Von Martens). Strebel 1909; Rev. Unterfam. Type Locality.Michoacn, Mxico. Distribution.GUERRERO: Acapulco (Strebel 1909). Martens 1893). Orthalicus zoniferus Strebel 1882Orthalicus zoniferus Strebel 1882; Beitrag. Mex. Landund Sssw.Pap. Mus. Zool. Univ. Mich. (611):7.Breure & Schouten 1985; Zool. Meded. Uitg. d. Rijksmus. Nat. Hist. Leiden 51:35; Oxystyla zonifera (Strebel). Pilsbry 1899; Man. Conch. 12:124; Zebra zoniferus (Strebel). Strebel 1909; Rev. Unterfam. Type Locality.Iguala, Guerrero, Mxico. 1895) GUERRERO: Iguala; Rancho del Platanillo, near Iguala; Chilpancingo; Venta de Zopilote; Monte Pelegrina, N of Acapulco (Pilsbry 1899); Laguna Coyuca (Solem 1959). JALISCO: hill 1.0 mi. NW of Zapotiltic (Breure & Schouten between Tzitzio and Heutamo; 1.2 mi. S of Charapendaro Subfamily BULIMULINAE Tryon 1867 Distribution.Tropical and subtropical Americas, South

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106 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Taxonomy.Numerous genera and species. The taxonomy used here follows Breure (1978, 1979). Richardson (1995) provided a bibliography of the species. The study area is inhabited by 132 species and 53 subspecies. Genus Plekocheilus Guilding 1828 Type Species.Voluta aurisseleni Born 1780. Distribution.Widely distributed in South America. Taxonomy.Five subgenera are recognized (Breure 1979). One enters the southeastern most region of Panam. Subgenus Eudolichotis Pilsbry 1896 Type Species.Bulimus distortus Bruguire 1789. Distribution.Northern areas of Colombia, Venezuela, Guayana, Surinam, French Guayana, Trinidad, Grenada, and Par, Brazil. Southeastern Panam. Taxonomy.About a dozen species are recognized. A single species occurs in the study area. Plekocheilus (Eudolichotis) distorta panamensis (Pilsbry 1910)Auris distorta panamensis Pilsbry 1910; Proc. Acad. Nat. Sci. Phila. Type Locality.Between Tabernillo and San Pablo, Panam. Holotype ANSP 101314. Distribution.Known only from the type locality. Genus Berendtia Crosse & Fischer 1869Berendtia Crosse & Fischer 1869; Jour. de Conchyl. 17:191.Christensen & Miller 1975:44.Breure 1978; Uitg. Rijkmuseum Nat. Hist. Leiden (164):77.Type Species.Clausilia (Balea ?) taylori Pfeiffer 1861. Distribution.Central Baja California Sur, Mxico. Taxonomy.The genus is monotypic. Berendtia taylori (Pfeiffer 1861)Clausilia (Balea ?) taylori Pfeiffer 1861; Proc. Zool. Soc. Lond. Berendtia taylori (Pfeiffer). Crosse & Fischer 1870; Jour. de Conchyl. Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. Cylindrella (Urocoptis) newcombiana Gabb 1868; Amer. Jour. Type Locality.Clausilia (Balea ?) taylori : unknown. Cylindrella newcombiana: Lower California, especially above Moleje [Moleg]. Distribution.BAJA CALIFORNIA SUR: restricted to the Sierra de la Giganta. Smith et al. (1990) listed numerous localities within this range. Genus Bulimulus Leach 1814 Type Species.Helix exilis Gmelin 1791 Distribution.South America from Argentina and Paraguay north to the Greater Antilles and southeastern Mxico. Taxonomy.Breure (1979) listed about 100 species and subspecies names that have been proposed in the genus. Seven species and three subspecies are recognized in Mxico and Central America. Bulimulus coriaceus (Pfeiffer 1856)Bulimus coriaceus Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:318. Bulimulus coriaceus (Pfeiffer). Fischer & Crosse 1875; Miss. Sci. Cent. Amer.:248.Pilsbry 189; Man. Conch. 11:51; pl. 10, Bulimus behrendti Pfeiffer 1861; Malak. Bltt. 8:168. Bulimulus coriaceaus var. berendti (Pfeiffer). Von Martens 1893; Biol. Cent. Amer.:249.Type Locality.State of Veracruz, Mxico. Distribution.OAXACA: Cacaprieto, near Santa Teapa; San Juan Bautista (Von Martens 1893). VERACRUZ: Atoyac (Von Martens 1893). Bulimulus corneus corneus (Sowerby 1833)Bulimus corneus Sowerby 1833; Proc. Zool. Soc. Lond. 1:37.107 (syntype). Bulimulus (Leptomerus) corneus (Sowerby). Fischer & Crosse 1875; Miss. Sci. Mex.:548. Bulimulus corneus (Sowerby).Pilsbry 1897; Man. Conch. 1996; Estud. Mus. Cienc. Naturales de Avala 101:259 Distribution.COSTA RICA, Prov. Alajuela: Las Palmares. Prov. San Jos: San Jos. Prov. Guanacaste: Sarapiqu (Pilsbry 1897). GUATEMALA, Dept. Retalhuleu: Retalhuleu. Dept. Escuintla: Escuintla. Dept. Zacapa: Zacapa. Bulimulus corneus nubeculatus (Pfeiffer 1853)Bulimus nubeculatus Pfeiffer 1853; Proc. Zool. Soc. Lond. 19:257. Bulimulus corneus var. nubeculatus (Pfeiffer). Von Martens 1893; Biol. Cent. Amer.:247.Pilsbry 1897; Man. Conch. 11:56. Type Locality.Central America. Lectotype BMNH 1975407 (Breure 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168):63). Distribution.Exact locality unknown. Taxonomy.Regarded as a questionable variety of

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 107 Bulimulus corneus by Von Martens (1893) and by Pilsbry, (1897). Bulimulus corneus minor (Von Martens 1893)Bulimulus (Leptomerus) nubeculatus (Pfeiffer). Fischer & Crosse Bulimulus corneus var. minor Von Martens 1893; Biol. Cent. 72 (shell).Type Locality.San Andres Tuxtla, Veracruz, Mxico. Distribution.Known only from the type locality. Taxonomy.Regarded as a questionable variety of Bulimulus corneus by Von Martens (1893) and by Pilsbry (1897). Bulimulus dysoni (Pfeiffer 1846)Bulimus dysoni Pfeiffer 1846; Proc. Zool. Soc. Lond. 14:39. Bulimulus dysoni (Pfeiffer). Fischer & Crosse 1875; Miss. Sci. Mex.:551.Von Martens 1893; Biol. Cent. Amer.:241.Pilsbry 1891; Proc. Acad. Nat. Sci. Phila:315.Pilsbry 1897; Man. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168):62.Type Locality.Honduras. Syntypes BMNH 197453 (Breure 1979:62. Distribution.HONDURAS: exact locality unknown. NICARAGUA, Dept. Chontales: Acoyapa (Martens 1893). GUATEMALA, Dept. Sacatepeques: Duenas (Pilsbry 1897). YUCATN: Tabi (Pilsbry 1891). Bulimulus inermis inermis (Morelet 1851)Bulimus inermis Morelet 1851; Test. Noviss. II:10. Bulimulus (Leptomerus) inermis (Morelet). Fischer & Crosse 1875; Bulimulus inermis (Morelet). Pilsbry 1897; Man. Conch. 11:51; pl. Inst. Wash. (431):534.Type Locality.Palizada, state of Campeche, Mxico. Distribution.Known only from the type locality. Bulimulus inermis major Von Martens 1897Bulimulus inermis var. major Von Martens 1897; Biol. Cent. Amer.:250.Khler 2007; Mitt. Mus. fr Nat. Berlin, Zool. Type Locality.Northern Guatemala, valley of the Rio Cisoy at Rancho Chisoy. Syntype ZMB 112736. Distribution.Known only from the type locality. Bulimulus inutilis (Reeve 1850)Bulimus inutilis Bulimulus inutilis (Reeve). Pilsbry 1897; Man. Conch. 11:73; Hist. Leiden (164):144.Breure 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168):63.Type Locality.Central America. Lectotype BMNH 1975162 (Breure 1978:144). Distribution.Unknown. The taxonomic validity of this form is questionable, and its inclusion in the Central American fauna is doubtful. Bulimulus sarcodes (Pfeiffer 1846)Bulimus sarcodes Pfeiffer 1846; Proc. Zool. Soc. Lond. 14:30. Bulimus (Leptomerus) sardodes Pfeiffer. Albers 1850; Die Heliceen:166. Bulimulus (Leptonerus) sarcodes (Pfeiffer). Von Martens 1893; Biol. Cent. Amer.:248. Bulimulus sarcodes (Pfeiffer). Pilsbry 1897; Man. Conch. 11:56; pl. Type Locality.Honduras. Syntype ZMB 112737 (Kohler 2007:135). Distribution.HONDURAS: not recorded from a Bulimulus unicolor (Sowerby 1833)Bulimus unicolor Sowerby 1833; Proc. Zool. Soc. Lond. 1:72.109 (syntype). Bulimulus unicolor (Sowerby). Pilsbry 1897: Man. Conch. 11:53; Inst. Wash. (431):534.Branson & McCoy 1963; Nautilus 76:105.Thompson 1967; Bull. Fla. St. Mus. 11:244. Bulimulus unicolor unicolor (Reeve). Breure 1978; Uitg. Rijkmuseum Nat. Hist. Leiden (164):147. Bulimus umbraticus Bulimus Petnensis Morelet 1852:10.Neubert & Jannsen (paralectotype). Bulimulus unicolor petenensis (Morelet). Breure 1978; Uitg. Rijkmuseum Nat. Hist. Leiden (164):149. Bulimulus (Leptomeris) istapensis Fischer & Crosse 1875:549; pl. Bulimulus sanmiguelensis Richards 1937; Proc. Amer. Philos. Soc. Bulimulus ochraspiris Branson & McCoy 1965; Neotropica 11:97 1969:106.Type Localities. Bulimus unicolor: Panam, Isla Perico Lectotype BMNH 1975411 (Breure 1978; Uitg. Rijkmuseum Nat. Hist. Leiden (164):148). Bulimus umbraticus: Central America. : Central America. Lectotype BMNH 1975411 (Breure 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168):63). Bulimus petenensis: Petn, Guatemala. Lectotype BMNH 1893.2.4.1176 (Breure 1979:64). Bulimulus istapensis: Istapa, Dept. Escuintla, Guatemala. Bulimulus sanmiguelensis: near the town of San Miguel, Isla Cozumel, Quintana Roo. Holotype ANSP 167746. Bulimulus ochraspiris:17.2 miles of Colorado Museum 26373. Distribution.BELIZE: W of Gales Point; Benque Viejo; Rio Benque Viejo, 1 mi. from cayo; Rio Belize (Thompson 1967). GUATEMALA, Dept. Petn: Paso Caballo; Flores; Puebla Nueva; Remate; island in Lago Eckixil; Uaxactun (Thompson 1967); Tikal National Park (Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):10). Dept. Zacapa: around

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108 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Zacapa (Thompson 1967). Dept. Chiquimula: between Jocotan and Jumusna (Thompson 1967). NICARAGUA: Greytown (Pilsbry 1897). CAMPECHE: numerous localities (Thompson 1967). CHIAPAS. QUINTANA ROO: Isla Cozumel, 1.5 mi. NNE of San Miguel; 7.1 mi. NNW of Xiatil; 2.3 mi. SSE of Xiatil, 4.0 mi. E of Xpujil (Thompson 1967). YUCATN: numerous localities (Pilsbry 1897; Bequaert & Clench 1933); Chichen Itza; San Ignacio; Merida; Uxmal; 0.8 mi. NE of Becanchn; 10.0 mi. NE of Becanchn; 7.0 mi. SSE of Uman; 1.0 mi. SSE of Puerto Telchac (Thompson 1967). COLIMA: 8 mi. SW of Colima (Branson & McCoy 1963). Genus Drymaeus Albers 1850Drymaeus Albers 1950:155. Type species by subsequent designation (Pilsbry 1898): Helix hygrohylaeus Orbigny 1835 (Bolivia). Goniognathus Fischer & Crosse 1875. Type species by original designation: Bulimus lattrei Pfeiffer 1847. Metadrymaeus Pilsbry 1926. Type species by original designation: Bulimus josephus Angus 1878.Distribution.Throughout tropical and subtropical America from Argentina north to Mxico, and throughout the West Indies to Florida. Taxonomy.Drymaeus is a very large genus consisting of nearly 600 named species. The genus includes 65 species and 50 subspecies in Mxico and Central America. Since the units has eluded authors. Recognition of subgenera has proven to be unsustainable because of the numerous intermediate species. The arrangement of species groups presented here (1893) and Pilsbry (1899). Nothing more satisfactory has been proposed during the past century. Many of the species have colorful shells, and local varieties frequently have been named. This has added to confusion when working with such a large group of species. Many of the varieties have little basis for taxonomic recognition. They were proposed as descriptors and not as taxonomic units, although subsequent authors have given them formal recognition. They are included in this account while recognizing that their taxonomic status requires further study. Breure & Eskens (1981) recognized two subgenera, Drymaeus and Mesembrinus, within the study area. They are distinguished by the development of the peristome and by some anatomical differences (Breure 1979:103). The shell in the subgenus Drymaeus outer lip, whereas the outer lip is simple to narrowly expanded in the subgenus Mesembrinus. The distinction is arbitrary except when dealing with extreme cases. Breure (1979) listed nine subgeneric names that have been applied to the subgenus Drymaeus. Three subgenera have been proposed for species of the Central American fauna. Drymaeus is divided into species groups on the basis of similar shell traits. The distinction between species variations. Solem (1955) divided eastern Mexican species into four groups based on penis sheath morphology. Breure and Eskens (1981) examined the anatomy of 49 species from throughout the range of the genus. Their data do not support Solems arrangement. Subgenus Drymaeus Albers 1850 Taxonomy.Thirteen species and twenty subspecies occur in the study area. The following two species groups are referred to the subgenus Drymaeus. Group of Drymaeus josephus Drymaeus (Drymaeus) expansus balboa Pilsbry 1926Drymaeus expansus balboa Pilsbry 1926; Proc. Acad. Nat. Sci. Type Locality.Rio Puerco, Prov. Panam, Panam. Holotype ANSP 140691. Distribution.Known only from the type locality. Drymaeus (Drymaeus) josephus josephus (Angas 1878)Bulimus josephus Angas 1878; Proc. Zool. Soc. Lond. 46:73; pl. 5, Otostomus josephus (Angas). Von Martens 1893; Biol. Cent. Amer.:202; pl. 12. Drymaeus josephus (Angas). Pilsbry 1899; Man. Conch. (2) 12:32; Phila. 72:6.Type Locality.San Jos, Costa Rica. (Von Martens 1893); Guapiles (Pilsbry 1920). Drymaeus josephus concolor (Von Martens 1893)Otostomus josephus var. concolor Von Martens 1893; Biol. Cent. Cent. Amer.:630.Khler 2007; Mitt. Mus. fr Nat. Berlin, Drymaeus josephus var. concolor (Von Martens). Pilsbry 1899; Bulimulus (Leptobrysus) zeledoni Dall 1893; Proc. U. S. Nat. Type Localities. Otostomus josephus var. concolor: Costa Rica; exact locality not stated. Bulimulus zeledoni : Costa Rica; holotype USNM 98231. Distribution.COSTA RICA, Prov. Alajuela: Santa Viejo. Prov. Heredia: Rio Sarapiqu; upper Savegal Valley, 1000 m alt. Prov. Puntarenas: near Terraba; Quebrada de Tocori, Rio Paqueta Valley; El Pital, Rio Naranjo Valley (all records from Von Martens 1893 1901). Drymaeus (Drymaeus) josephus errans Pilsbry 1926Drymaeus (Drymaeus) josephus errans Pilsbry 1926; Proc. Acad. Type Locality.Mono Creek, Prov. Bocas del Toro, Panam. Holotype in the ANSP. Distribution.PANAM, Prov. Bocas del Toro: Senori Creek. Canal Zone: between Tabernilla and San Pablo.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 109 Drymaeus (Drymaeus) josephus maculosus (Von Martens 1893)Otostomus josephus var. maculosus Von Martens 1893; Biol. Cent. Drymaeus josephus var. maculosus (Von Martens). Pilsbry 1899; Type Locality.On the banks of the Rio Pacuare del Sur; about 500 m alt. Distribution.Known only from the type locality. Drymaeus (Drymaeus) megastomus Parodiz 1962Drymaeus megastomus Parodiz 1962; Proc. U. S. Nat. Mus. Type Locality.Costa Rica. Holotype USNM 98230. PANAM: Cerro Campaa (Parodiz 1962). Drymaeus (Drymaeus) zhorquinensis (Angas 1879)Bulimus zhorquinensis Angas 1879; Proc. Zool. Soc. Lond. 47:478; Otostomas zhorquinensis (Angas). Von Martens 1893; Biol. Cent. Amer.:202. Drymaeus zhorquiensis (Angas). Pilsbry 1899; Man. Conch. (2) Type Locality.SE Costa Rica, middle Zhorquin to Distribution.Known only from the type locality. Group of Drymaeus serperastrum Drymaeus (Drymaeus) castus castus (Pfeiffer 1846)Bulimus castus Pfeiffer 1846; Proc. Zool. Soc. Lond. 14:112. Bulimulus (Drymaeus) castus (Pfeiffer). Fischer & Crosse 1875; Otostomus castus (Pfeiffer). Von Martens 1893; Biol. Cent. Drymaeus castus (Pfeiffer). Pilsbry 1899; Man. Conch. (2) 12:43 Distribution.GUATEMALA, Dept. Alta Verapaz: Cobn; Tamah (Von Martens 1893); Chejel (Hinkley 1920) Drymaeus (Drymaeus) castus xantholeucus (Von Martens 1893)Otostomus castus var. xantholeucus Von Martens 1893; Biol. Cent. Drymaeus castus var. xantholeucus (Von Martens). Pilsbry 1899; Type Locality.Sabo [Sabob], Polochic Valley, Dept. Alta Verapaz, Guatemala. Lectotype ZMB 109.941 (Khler 2007). Distribution.Known only from the type locality. Drymaeus (Drymaeus) castus porrectus (Von Martens 1893)Otostomus castus var. porrectus Von Martens 1893; Biol. Cent. Drymaeus castus var. porrectus (Von Martens). Pilsbry 1899; Man. Conch. (2) 12:44.Type Locality.Vera Paz, Guatemala. Lectotype ZMB 109.942 (Khler 2007). Distribution.GUATEMALA, Dept. Alta Verapaz: Cobn or Tamahu (Von Martens 1893). Drymaeus ( Drymaeus) chiapensis chiapensis (Pfeiffer 1866)Bulimus chiapensis Pfeiffer 1866; Malak. Bltt. 13:81. Bulimulus (Drymaeus) chiapensis (Pfeiffer). Fischer & Crosse 1875; Miss. Sci. Mex.:483. Bulimulus chiapensis (Pfeiffer). Strebel 1882; Beitrag. Mex. LandOtostomus chiapensis (Pfeiffer). Von Martens 1893; Biol. Cent. Drymaeus chiapensis (Pfeiffer). Pilsbry 1899; Man. Conch. (2) Type Locality.Cumbre de Manzanilla, Chiapas, Mxico. Distribution.MXICO: Cuautitlan (Strebel 1882). Orizaba; Coatepec (Von Martens 1893). Drymaeus (Drymaeus) chiapensis quadrifasciatus (Von Martens 1893)Otostomus chiapensis var. quatrifasciatus Von Martens 1893; Biol. Drymaeus chiapensis var. quadrifasciatus (Von Martens). Pilsbry Type Locality.Matlaquihahuitl, Veracruz, Mxico. Distribution.VERACRUZ: Cerro de Plumas, near Drymaeus (Drymaeus) chiapensis nebulosus (Von Martens 1893)Otostomus chiapensis var. nebulosus Von Martens 1893; Biol. Cent. Amer.:205. Drymaeus chiapensis var. nebulosus (Von Martens). Pilsbry 1899; Type Locality.Cuautitlan, Mxico (Breure 1979). Distribution.MXICO: Cuautitlan. VERACRUZ: Coatepec; Quimistlan (Von Martens 1893). Drymaeus (Drymaeus) colimensis (Rolle 1895)Otostomus colimensis Rolle 1895; Nach. Deut. Malak. Ges. 32:130.(lectotype). Drymaeus colimensis (Rolle). Pilsbry 1899; Man. Conch. (2) 12:47.Solem 1959; Occ. Pap. Mus. Zool. Univ. Mich. (611):9; Type Locality.Colima, Mxico. Lectotype ZMB 47.660a (Khler 2007).

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110 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Drymaeus (Drymaeus) dombeyanus (Ferussc 1842)Bulimus dombeyanus Ferussc 1842, in Pfeiffer, Symb. Hist. Helic. II:76. Bulimulus (Scutalus) dombyanus (Ferussac). Fischer & Crosse 1873; Miss. Sci. Mex. I:515.Strebel 1882; Beitrag. Mex. Landund Drymaeus dombyanus (Ferussc). Pilsbry 1899; Man. Conch. Sci. Phila. 71:218.Solem 1957 Notulae Naturae (298):11. Bulimus alcantarae Bernardi 1853; Jour. de Conchyl. 4:35; pl. 3, Bulimulus chaperi Crosse & Fischer 1893; Jour. de Conchyl. 41:31; Type Localities. Bulinus dombeyanus: not stated. Bulimus alcantarae: not stated. Bulimulus chaperi: Mescala Island, Lake Chapala, Jalisco, Mxico. Distribution.GUERRERO: Laguna Coyuca, near Acapulco. JALISCO: Presidea Island, Lago de Chapala; Guadalajara. MICHOACN: E side of Cerro de Guzman, Coalcomn, 3200 ft. alt.; W side of Cerro Guzman, Coalcomn, 3500 ft. alt.; Sierra de Coalcomn, 3500 ft. alt.; San Jos de Purua (Solem 1957); Venta de Zopilote (Pilsbry 1899). Drymaeus (Drymaeus) dunkeri dunkeri (Pfeiffer 1846)Bulimus dunkeri Pfeiffer 1846, in Philippi; Abbild. neuer Conch. Bulimulus (Scutalus) dunkeri (Pfeiffer). Fischer & Crosse 1875; Miss. Sci. Mex.:511.Strebel 1882; Beitrag. Mex. Landund Sssw.-Conch. V:76. Otostomus dunkeri (Pfeiffer). Von Martens 1893; Biol. Cent. Drymaeus dunkeri (Pfeiffer). Pilsbry 1899; Man. Conch. (2) 12:45 Notulae Naturae (298):13. Drymaeus (Drymaeus) dunkeri (Pfeiffer). Breure & Eskens 1981; 2 (lectotype).Type Locality.Michoacn, Mxico. Lectotype BMNH 1975512 (Breure & Eskens 1981). Distribution.MICHOACN: Patzcuaro; Tzintzuntzan; Morelia; Uruapan; 2 mi. S of Los Reyes (Solem 1957). NAYARIT: Tepc (Pilsbry 1899). Drymnaeus (Drymaeus) dunkeri forreri (Mousson 1883)Bulimulus forreri Mousson 1883; Jour. de Conchyl. 31:217; pl. 9, Otostomus dunkeri var. forreri (Mousson). Von Martens 1893; 207; Drymaeus dunkeri var. forreri (Mousson). Pilsbry 1899; Man. Type Locality.Ventanas, Durango, Mxico. Martens 1893). DURANGO: type locality. Drymaeus (Drymaeus) fenestratus (Pfeiffer 1846)Bulimus fenestratus Pfeiffer 1846; Proc. Zool. Soc. Lond. 14:26. Bulimulus (Scutalus) fenestratus (Pfeiffer). Fischer & Crosse 1875; Miss. Sci. Mex.:528. Otostomus fenestratus (Pfeiffer). Von Martens 1893; Biol. Cent. Drymaeus fenestratus (Pfeiffer). Pilsbry 1899; Man. Conch. (2) (298):12. Drymaeus (Drymaeus) fenestratus (Pfeiffer). Breure & Eskens 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden (216):22; Bulimulus (Otostomus) piescheli Von Martens 1863; in Monatsber. Akad. Wiss. Berl., 26:541.Khler 2007; Mitt. Mus. fr Nat. Type Localities. Bulimus fenestratus: Mxico. Lectotype BMNH 1975525 (Breure & Eskens 1981). Bulimulus piescheli : Manzanilla, Colima, Mxico Bulimulus piescheli : Manzanillo, Colima; lectotype ZMB 4.581 (Khler 2007). Distribution.COLIMA: Manzanillo (Solem 1957). JALISCO: NW slope of Nevada de Colima, 7500 ft. alt. (Solem 1957); 9.1 mi. SE of San Vicente (Breure & Eskens 1981). MICHOACN: Sierra de Coalcomn, between Pomare and Maruato; Rancho Quemado, 5600 ft. alt.; between Tecatas and Las Higueritas, 5300 ft.; Barranca de Herradero, S of San Pedro Damian, 1500 ft. alt.; above San Jos de los Montaas, 4700 ft. alt.; Cerro Barolosa, N of Coalcomn, 7600 ft. alt. (Solem 1959); 4 mi. S of Capacuano (Breure & Eskens 1981). Drymaeus (Drymaeus) lattrei lattrei (Pfeiffer 1846)Bulimus lattrei Bulimulus delattrei (Pfeiffer). Fischer & Crosse 1785:481; pl. 20, (Name amended). Otostomus delattrei (Pfeiffer). Von Martens 1893; Biol. Cent. Drymaeus lattrei (Pfeiffer). Pilsbry 1899; Man. Conch. (2) 12:41 42 (shell).van der Schalie 1940:4. Drymaeus (Drymaeus) lattrei (Pfeiffer). Breure & Eskens 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden (216):28. Bulimus facillatus Type Locality.Vera Cruz, Mxico. Lectotype BMNH 1975555 (Breure & Eskens 1981). The type locality was corrected to Vera Paz, Guatemala, by Pilsbry (1899:42). Distribution.GUATEMALA, Dept. Alta Verapaz: Cobn; trail to Cobn ca. 3 km E of Samac (van der Schalie 1940). Dept. Baja Verapaz: San Geronimo (Von Martens 1893). Drymaeus ( Drymaeus) lattrei hiabundus (Von Martens 1893)Otostomus delattrei var. hiabundus Von Martens 1893; 205; pl. 12; Drymaeus lattrei var hiabundus (Von Martens). Pilsbry 1899; Man. Type Locality.Cerro Zunil, [Dept. Quetzaltenango], Guatemala. Lectotype ZMB 101.812a (Khler 2007). Distribution.Known only from the type locality.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 111 Drymaeus (Drymaeus) lilacinus lilacinus (Reeve 1949)Bulimus lilacinus 532. Bulimulus lilacinus (Reeve). Fischer & Crosse 1875; Miss. Sci. Otostomus lilacinus (Reeve). Von Martens 1893; Biol. Cent. Drymaeus lilacinus (Reeve). Pilsbry 1899; Man. Conch. (2) 12:35 Drymaeus (Drymaeus) lilacinus (Reeve). Breure & Eskens 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden (216):28. Bulimus patricius Type Localities. Bulimus lilacinus : not given. Bulimus patricius: not given; lectotype BMNH 1874.12.11.220 (Breure & Eskens 1982). Distribution.GUATEMALA, Dept. Solol: San Agustin (Von Martens 1893). Dept. Escuintla: Zapote (Haas 1949). Drymaeus ( Drymaeus) lilacinus crossei (Von Martens 1893)Otostomus lilacinus var. crossei Von Martens 1893; Biol. Cent. Amer.:201. Drymaeus lilacinus var. crossei (Von Martens). Pilsbry 1899; Man. Type Locality.Alta Vera Paz, Guatemala. Distribution.Known only from the type locality. Drymaeus ( Drymaeus) lilacinus ictericus (Von Martens 1893)ar. ictericus Von Martens 1893; Biol. Cent. Nat. Berlin, Zool. 82:146 (syntype). (Not Ototostomus depictus var. ictericus Martens 1873, a Venezuela shell.) Drymaeus lilacinus var. ictericus (Von Martens). Pilsbry 1899; Man. Type Locality.Cerro Zunil, [Dept. Quetzaltenango], Guatemala. Syntype ZMB 109.937 (146). Distribution.Known only from the type locality. Drymaeus ( Drymaeus) lilacinus jansoni (Von Martens 1893)Otostomus lilacinus var. jansoni Von Martens 1893; Biol. Cent. Drymaeus lilacinus var. jansoni (Von Martens). Pilsbry 1899; Man. Type Locality.Nicaragua; lectotype BMNH 1901.6.22.951 (Breure & Eskens 1981). Distribution.NICARAGUA: not known from a Drymaeus (Drymaeus) lilacinus undulosus (Von Martens 1893)Otostomus lilacinus var. undulosus Von Martens 1893; Biol. Cent. Drymaeus lilacinus var. undulosus (Von Martens). Pilsbry 1899; Type Locality.Hacienda de Las Nubes, Cerro Zunil, [Dept. Quetzaltenango], Guatemala; 4000 ft. alt. Lectotype ZMB 109.940a (Khler 2007). Distribution.Known only from the type locality. Drymaeus ( Drymaeus) lilacinus unicolor (Von Martens 1893)Otostomus lilacinus var. unicolor Von Martens 1893; Biol. Cent. Drymaeus lilacinus var. unicolor (Von Martens). Pilsbry 1899; Man. Conch. (2) 12:37.Type Locality.Cerro Zunil, Dept. Quetzaltemnango, Guatemala. Lectotype ZMB 109.919 (Khler 2007). Distribution.GUATEMALA, Dept. Huehuetenango: Cholhuitz; Dept. Quetzaltenango: Cerro Zunil, 4000 ft. alt.; El Esposo, 1000 ft. alt. Dept. Sacatepeqeuez: mountain crest between Antigua and Escuintla; Hacienda Buenavista and Hacienda Helvitia (Von Martens 1893). Drymaeus (Drymaeus) serperastrum (Say 1829)Bulimus serperastrus Say 1829; New Harmony Disseminator, Jan. 1:25. Bulimulus serperastrus (Say). Fischer & Crosse 1875; Miss. Sci. Otostomus serperastrus (Say). Von Martens 1893; Biol. Cent. Amer.:202. Drymaeus serperastrum (Say). Pilsbry 1899; Man. Conch. (2) Pub. Carnegie Inst. Wash. (431):532.Bequaert & Clench 1938; Pub. Canegie Inst. Wash. (491):253.Harry 1950; Occ. Pap. Mus. Zool. Univ. Mich. (524):12.Solem 1955; Occ. Pap. Mus. Branson & McCoy 1963; Nautilus 76:106.Thompson 1967; Bull. Fla. St. Mus. 11:249.Breure & Eskens 1981; Zool. Verh. (reproductive anatomy). Bulimus liebmanni Pfeiffer 1846; Zeitshrift fr Malak.:159. Pfeiffer 1866; Malak. Bltt. 13; 81.Type Localities. Bulimus serperastrum: On the road between Vera Cruz and Mxico City; holotype ANSP 25909 (H. B. Baker 1963:230). Bulimus liebmanni : Mxico. Bulimus : Veracruz, Mxico. Distribution.CAMPECHE: 7.1 mi. SW of Cd. Campeche (Thompson 1967). QUINTANA ROO: Isla Cozumel, 3 km S of San Miguel (Branson & McCoy 1963). SINALOA: 22 mi. NNW of Mazatln (Breure & Eskens 1981). YUCATN: Chichn Itza; Progreso (Bequaert & Clench 1933); Ebitz Cave, Oxkutzcab (Bequaert & Clench 1938); San Ignacio (Solem 1955a); 1.5 mi. S of Libre Union; Piste (Branson & McCoy 1963). VERACRUZ: Jalapa; Colorado, between Jalapa and Veracruz; Paso de San Juan and Loma de Piedra on the Rio Jamapa; between Paso de Ovejas and Mirador; Bobo (Pilsbry 1899). Reported from localities (Pilsbry 1899). Drymaeus (Drymaeus) ziegleri (Pfeiffer 1846)Bulimulus zieglei Pfeiffer 1846; Proc. Zool. Soc. Lond. 14:113.

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112 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Drymaeus ziegleri (Pfeiffer). Pilsbry 1899; Man. Conch. (2) 12:39 Bulimus californicus Type Locality.Not stated. Distribution.SINALOA: Mazatln; Altata (Pilsbry 1899). Subgenus Mesembrinus Albers that have been applied to Mesembrinus. Three names pertain to the Central American fauna: Mesembrinus Albers 1850:157 (type species by subsequent designation, Albers 1860: Helix Ferussac 1821); Antidrymaeus Germain 1907 (type species by subsequent designation, Pilsbry 1926: Bulimulus inusitatus Fulton 1900); and Leptodrymaeus Pilsbry 1946 (type species by original designation: Bulimus dominicus Reeve 1850). Fifty-two species and thirty subspecies of Mesembrinus occur in the study area. The following species groups are referred to Mesembrinus. Group of Drymaeus sulcosus Drymaeus (Mesembrinus) sulcosus (Pfeiffer 1841)Bulimus sulcosus Pfeiffer 1841, Symbolae Hist. Heliciorum 1:43. Bulimulus sulcosus (Pfeiffer). Fischer & Crosse 1875; Miss. Sci. Strebel 1882; Beitrag. Mex. Landund Sssw.-Conch. V:63; pl. Otostomus sulcosus 4 (shell). Drymaeus (Drymaeus) sulcosus (Pfeiffer). Pilsbry 1899; Man. & Eskens 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden Drymaeus (Mesembrinus) sulcosus (Pfeiffer). Thompson 2008:376.Type Locality.Tacubaya, Mxico. There are six states that have a place named Tacubaya: Distrito Federal, Oaxaca, Jalisco, Guanajuato, Chihuahua, and Yucatn. Distribution.GUERRERO: Soledad; mountains near Chilpancingo, 9500 ft. alt. (Von Martens 1893); Cerro Muerto, 2600 ft. alt. (Breure & Eskens 1981). MXICO: near Cd. Mxico; summit de los Aguas Escondidas (Von Martens 1893); Santa Rosa to El Desierto de los Leones (Solem 1955a). VERACRUZ: Tuxpn (Strebel 1882). Drymaeus (Mesembrinus) (Pfeiffer 1856) (Pfeiffer). Fischer & Crosse 1875; Miss. Sci. Mex. Landund Sssw.-Conch. V:79. (Pfeiffer). Von Martens 1893; Biol. Cent. Amer.:213. Drymaeus (Mesembrinus) (Pfeiffer). Pilsbry 1899; 187 (reproductive anatomy). Distribution.OAXACA: Juquila; Yalalag. VERA1893); Sumidero (Solem 1955a); 4.7 mi. S of Huatusco (Breure & Eskens 1982). Drymaeus ( Mesembrinus) botterii (Crosse & Fischer 1875)Bulimulus (Drymaeus) botterii Crosse & Fischer 1875; Jour. de Conchyl. 23:52.Fischer & Crosse 1875; Miss. Sci. Mex.:487; und Sssw.-Conch. V:63. Otostomus sulcosus var. botterii (Crosse & Fischer). Von Martens 1893; Biol. Cent. Amer.:208. Drymaeus botterii (Crosse & Fischer). Pilsbry 1899; Man. Conch. Type Locality.Near Orizaba, Veracruz, Mxico. Distribution.Known only from the type locality. Drymaeus (Mesembrinus) cucullus (Morelet 1851)Bulimus cucullus Morelet 1849; Test. Noviss. I:9. Bulimulus (Scutalus) cucullus (Morelet). Fischer & Crosse 1875; Otostomus cucullus (Morelet). Von Martens 1893; Biol. Cent. Amer.:214. Drymaeus cucullus (Morelet). Pilsbry 1899; Man. Conch. (2) 12:58; Bulimulus (Scutalus) cucullus var. gracilior Fischer & Crosse 1875; Type Locality.Sisal, Yucatn, Mxico. Distribution.Known only from the type locality. Drymaeus (Mesembrinus) ghiesbreghti ghiesbreghti (Pfeiffer 1866)Bulimus ghiesbreghti Pfeiffer 1866; Malak. Bltt. 82. Bulimulus (Scutalus) ghiesbreghti (Pfeiffer). Fischer & Crosse (shell). Otostomus ghiesbreghti (Pfeiffer). Von Martens 1893; Biol. Cent. Drymaeus ghiesbreghti (Pfeiffer). Pilsbry 1899; Man. Conch. (2) Type Locality.Chiapas, Mxico. Distribution.OAXACA: Tlacolula (Von Martens 1893). Drymaeus (Mesembrinus) ghiesbreghti interstitialis (Von Martens 1893)Otostomus ghiesbreghti var. interstitialis Von Martens 1893; Biol. Drymaeus ghiesbreghti var. interstitialis (Von Martens). Pilsbry Type Locality.Cumbres de San Martin, on the NW slope of the Cordillera, Guatemala; lectotype ZMB 112.876 (Khler 2007). Distribution.Known only from the type locality.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 113 Drymaeus (Mesembrinus) ghiesbreghti iodostylus (Pfeiffer 1861)Bulimulus iodostylus Pfeiffer 1861; Proc. Zool. Soc. Lond. 29:23 Bulimulus (Scutalus) iodostylus (Pfeiffer). Fischer & Crosse 1875; Miss. Sci. Mex.:539. Bulimulus iodostylus (Pfeiffer). Strebel 1882; Beitrag. Mex. LandOtostomus ghiesbreghti var. iodostylus (Pfeiffer). Von Martens 1893; Biol. Cent. Amer.:210.Rolle 1895:130. Drymaeus ghiesbreghti var. iodostylus (Pfeiffer). Pilsbry 1899; Oaxaca, Mxico. Distribution.COLIMA: Colima (Rolle 1895). OAXACA: only from the type locality. Drymaeus (Mesembrinus) ghiesbreghti stolli (Von Martens 1887)Bulimulus jonasi var. stolli Von Martens 1887; Sitz.-Ber Ges. Naturf. Freude Berlin 1886:161. Otostomus ghiesbreghti var. stolli Von Martens 1893; Biol. Cent. Drymaeus ghiesbreghti var. stolli (Von Martens). Pilsbry 1899; Type Locality.Llanos of Quetzaltenango, Dept. Quetzaltenango, Guatemala; 6000 ft. alt. Distribution.GUATEMALA, Dept. Chilpancingo: Tecpam, 7000 ft. alt. Dept. Huehuetenango: Hacienda Nubes, slopes of Cerro Zunil. Dept. Sapatapequez: Antigua. Dept. Solol: Argueta; Los Encuentros, 8000 ft. alt. (Von Martens 1893). Drymaeus (Mesembrinus) hegewischi (Pfeiffer 1842)Bulimus hegewischi Pfeiffer 1842; Symbolae ad historiam heliceorum 2:46. Bulimulus (Scutalus) hegewischi (Pfeiffer). Fischer & Crosse 1875; Bulimulus hegewischi (Pfeiffer). Strebel 1882; Beitrag. Mex. LandOtostomus hegewischi (Pfeiffer). Von Martens 1893; Biol. Cent. Drymaeus hegewischi (Pfeiffer). Pilsbry 1899; Man. Conch. (2) Drymaeus (Mesembrinus) hegewischi (Pfeiffer). Breure & Eskens Type Locality.Not stated. Distribution.GUERRERO: Omilteme. JALISCO: Lake Chapala (Pilsbry 1920). MXICO: environs of Cd. Mxico; Toluca. MORELOS: Cuernavaca. OAXACA: Tenango. PUEBLA: Puebla; Tehuacn (all above records from Von Martens 1893); NE of Chietla (Breure & Eskens 1982). QUERTARO: Quertaro (Pilsbry 1926). VERACRUZ: Orizaba. Drymaeus (Mesembrinus) jonasi (Pfeiffer 1846)Bulimus jonasi Pfeiffer 1846, in Philippi, Abbild. neuer Conchyl. Bulimulus (Scutalus) jonasi (Pfeiffer). Fischer & Crosse 1875; Miss. Otostomus jonasi (Pfeiffer). Von Martens 1893; Biol. Cent. Drymaeus jonasi (Pfeiffer). Pilsbry 1899; Man. Conch. (2) 12:54 34:38.Haas 1949; Nautilus 62:136, 13. Type Locality.America Centralis, Vera Cruz (Breure 1979). This is in error for Vera Paz, Guatemala (see Von Martens 1893; Biol. Cent. Amer.:212). Three syntype BMNH 1975557. Distribution.GUATEMALA, Dept. Alta Verapaz: Cobn. Dept. Chimaltenango: Yepocapa (Haas 1949). Dept. Guatemala: Amatitlan (Von Martens 1893); Guatemala (Hinkley 1920; Haas 1949). Dept. Sacatepquez: Antigua; Dueas (Von Martens 1893); Finca San Rafael (Haas 1949). Drymaeus (Mesembrinus) lineolatus (Conrad 1855)Bulimulus lineolatus Conrad 1855; Proc. Acad. Nat. Sci. Phila. 7:32. Otosomus recluzianus var. lineolatus (Conrad). Von Martens 1893; Biol. Cent. Amer.:214. Drymaeus lineolatus (shell).Type Locality.Volcn de Cartago, Prov. Cartago, Costa Rica. Distribution.COSTA RICA, Prov. Cartago: Azahar de Cartago, 1600 m alt. Prov. San Jos: San Jos, 1161 m alt. Drymaeus (Mesembrinus) lirinus (Morelet 1851)Bulimus lirinus Morelet 1851; Test. Noviss. II:11. Bulimulus (Drymaeus) lirinus (Morelet). Fischer & Crosse 1875; Otostomus lirinus (Morelet). Von Martens 1893; Biol. Cent. Amer.:214. Drymaeus lirinus (Morelet). Pilsbry 1899; Man. Conch. (2) 12:57 Type Locality.San Luis de Petn, Dept. Petn, Guatemala. Syntypes BMNH 1893.2.4.1954. (Breure 1979). Distribution.Known only from the type locality. Drymaeus (Mesembrinus) recluzianus recluzianus (Pfeiffer 1847)Bulimus recluzianus Pfeiffer 1847; Zeit. Malak.:82. Bulimulus (Scutalus) recluzianus (Pfeiffer). Fischer & Crosse 1875; Miss. Sci. Mex.:510. Bulimulus recluzianus (Pfeiffer). Strebel 1882; Beitrag. Mex. LandOtostomus recluzianus (Pfeiffer). Von Martens 1893; Biol. Cent. Amer.:213.Type Locality.Chiapas, Mxico. locality. Drymaeus (Mesembrinus) recluzianus martensianus Pilsbry 1899 Conrad. Von Martens 1893; Biol. Cent. Amer.:214.

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114 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Drymaeus recluzianus var. martensianus Pilsbry 1899; Man. Conch. 12:56. Drymaeus (Mesembrinus) recluzianus martensianus Pilsbry. Breure & Eskens 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden Type Locality.Not stated. Distribution.COSTA RICA, Prov. San Jos: San Jos; San Francisco de Dos Rios; La Union, 1100 m alt. (Von Martens 1893). Drymaeus (Mesembrinus) rudis (Anton 1839)Bulimus rudis Anton 1839; Verz. Der. Conch. Sammlungen:43. Bulimulus (Scutalus) rudis (Anton). Fischer & Crosse 1875; Miss. (shell). Bulimulus rudis (Anton). Strebel 1882; Beitrag. Mex. Landund Otostomus rudis (Anton). Von Martens 1893; Biol. Cent. Amer.:209. Drymaeus rudis (Anton). Pilsbry 1899; Man. Conch. (2) 12:49; Drymaeus (Drymaeus) rudis (Anton). Breure & Eskens 1981; Zool. 11213 (reproductive anatomy).Type Locality.Not stated. Distribution.MXICO: environs of Cd. Mxico; Chapultepec (Von Martens 1893). MICHOACN: Angangeo (Von Martens 1893). OAXACA: Oaxaca (Breure & Eskens 1981). MXICO, D. F.: Chapultepec Park (Solem 1955a). Group of Drymaeus attenuatus Drymaeus (Mesembrinus) attenuatus attenuatus (Pfeiffer 1851)Bulimus attenuatus Pfeiffer 1851; Proc. Zool. Soc. Lond. 19:256. Bulimulus (Drymaeus) attenuatus (Pfeiffer). Fischer & Crosse 1875; Bulimulus attenuatus (Pfeiffer). Strebel 1882; Beitrag. Mex. LandOtostomus attenuatus (Pfeiffer). Von Martens 1893; Biol. Cent. Drymaeus attenuatus (Pfeiffer). Pilsbry 1899; Man. Conch. (2) 12: (reproductive anatomy). Bulimus kefersteini Pfeiffer 1866:82. Otostomus attenuatus var. concolor Von Martens 1893; Biol. Cent. Type Localities. Bulimus attenuatus : Vera Cruz, Mxico. Lectotype BMNH 1975458 (Breure & Eskens 1981). Otostomus attenuatus var. concolor: Veracruz; syntypes ZMB. Distribution.COSTA RICA, Prov. Cartago: Cartago Atoyac; Mirador (Von Martens 1893); Sumidero (Solem 1955a). Drymaeus (Mesembrinus) attenuatus pittieri (Von Martens 1893)Otostomus attenuatus var. pittieri Von Martens 1893; Biol. Cent. Amer.:216.Von Martens 1901; Biol. Cent. Amer.:631.(lectotype). Drymaeus attenuatus var. pittieri (Von Martens). Pilsbry 1899; Man. Drymaeus attenuatus pittieri (Von Martens). Pilsbry 1920c; Proc. Acad. Nat. Sci. Phila. 72:5.Type Locality.Alto de Mano Tigre, near Terraba, Puntarenus Prov., Costa Rica; 600 m alt. Lectotype ZMB 48.234 (Khler 2007). Distribution.COSTA RICA, Prov. Cartago: road from Prov. San Jos: Dota (Von Martens 1893). Drymaeus (Mesembrinus) (Pfeiffer 1851)Pfeiffer 1851; Proc. Zool. Soc. Lond. 19:256. Otostomus attenuatus var. (Pfeiffer). Von Martens 1893; Drymaeus attenuatus var. (Pfeiffer). Pilsbry 1899; Man. Type Locality.in republic Mexican. Distribution.COSTA RICA, Prov. San Jos: San Atoyac; Mirador (Von Martens 1893). Drymaeus (Mesembrinus) bugabensis (Von Martens 1893)Otostomus bugabensis Von Martens 1893; Biol. Cent. Amer.:218; Drymaeus bugabensis (Von Martens). Pilsbry 1899; Man. Conch. Drymaeus (Mesembrinus) bugabensis (Von Martens). Breure & Type Locality.Bugaba, Prov. Chiriqu, Panam. Lectotype BMNH 1901.6.22.958 (Breure & Eskens 1982). Distribution.Known only from the type locality. Drymaeus (Mesembrinus) chiriquiensis DaCosta 1901Drymaeus chiriquiensis DaCosta 1901; Proc. Malac. Soc. London Type Locality.Boquete, Prov. Chiriqu, Panam. Distribution.Known only from the type locality. Drymaeus (Mesembrinus) costaricensis (Pfeiffer 1862)Bulimus costaricensis Pfeiffer 1862; Malak. Bltt. 9:152. Otostomus costarricensis (Pfeiffer). Von Martens 1893; Biol. Cent. Amer.:217. Drymaeus costaricensis (Pfeiffer). Pilsbry 1899; Man. Conch. (2) Nat. Sci. Phila. 72:5.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 115Drymaeus (Drymaeus) costaricensis (Pfeiffer). Breure & Eskens 34 (reproductive anatomy). Angas 1875; Proc. Zool. Soc. Lond. 43:73; pl. Type Locality.Costa Rica. Distribution.COSTA RICA, Prov. Alajuela: Alahuela, 900 m alt. Prov. Cartago: Cartago to Navarro (Von 1920). Prov. San Jos: San Jos; San Francisco de Dos Rios NICARAGUA. Dept. Matagalpa:1.5 mi. E of Matagalpa, 2700 ft. alt. (Breure & Eskens 1981). Drymaeus (Mesembrinus) droueti droueti (Pfeiffer 1856)Bulimus drouti 12. Bulimulus droueti (Pfeiffer). Fischer & Crosse 1875; Miss. Sci. 7b (shell). Otostomus droueti (Pfeiffer). Von Martens 1893; Biol. Cent. Amer.:218. Drymaeus droueti (Pfeiffer). Pilsbry 1899; Man. Conch. (2) 12:65 anatomy). Drymaeus bourgeoisaeType Localities. Bulimus droueti Mxico. Drymaeus bourgeoisae: near Paraje Nuevo, near Coatepec; Chirimoyo; San Jos Miahuatlan; Jalapa; Mirador; Orizaba; Tuxpan (Pilsbry 1899); Potrero to Sumidero (Solem 1955a). Drymaeus (Mesembrinus) droueti deletus Solem 1955Drymaeus droueti deletus Solem 1955; Occ. Pap. Mus. Zool. Univ. Type Locality.Sumidero, Veracruz, Mxico. Holotype UMMZ 181389. Distribution.Known only from the type locality. Drymaeus ( Mesembrinus) droueti sporlederi (Pfeiffer 1866)Bulimus sporlederi Pfeiffer 1866; Malak. Bltt. 13:83. Bulimulus sporlederi (Pfeiffer). Fischer & Crosse 1875; Miss. Sci. Drymaeus droueti sporlederi (Pfeiffer). Pilsbry 1899; Man. Conch. Type Locality.Mirador, near Veracruz, Veracruz, Mxico. Distribution.VERACRUZ: Penuela to Sumidero (Solem 1955a). Drymaeus ( Mesembrinus) fenestrellus (Von Martens 1863)Bulimulus (Scutalus) fenestrellus Von Martens 1863; 541.Fischer & Crosse 1875; Miss. Sci. Mex.:529.Khler 2007; Mitt. Mus. Bulimulus fenestrellus anatomy). Otostomus fenestrellus (Von Martens). Von Martens 1893; Biol. Cent. Amer.:214. Drymaeus fenestrellus (Von Martens). Pilsbry 1899; Man. Conch. Drymaeus (Drymaeus) fenestrellus (Von Martens). Breure & Eskens 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden (216):26; Bulimus gealei H. Adams 1867; Proc. Zool. Soc. Lond., 309; pl. 19, Otostomus fenestrellus var. subunicolor Von Martens 1893; Biol. Cent. Amer.:215.Type Locality.Bulimulus (Scutalus) fenestrellus: Patzcuaro, Michoacn, Mxico; lectotype ZMB 4.457a (Khler 2007). Ototstomus fenestrellus var. subunicolor: Puebla, Mxico; lectotype ZMB 18.767a (Khler 2007). Distribution.MICHOACN: near El Tigre (Breure & Eskens 1981). PUEBLA: Matamoros Izucar; Puebla (Von Martens 1893). Drymaeus (Mesembrinus) hepatostomus (Pfeiffer 1861)Bulimus hepatostomus Pfeiffer 1861; Proc. Zool. Soc. Lond. 29:23; Bulimulus (Drymaeus) hepatostomus (Pfeiffer). Fischer & Crosse Otostomus hepatostomus (Pfeiffer). Von Martens 1893; Biol. Cent. Drymaeus hepatostomus (Pfeiffer). Pilsbry 1899; Man. Conch. (2) Drymaeus (Drymaeus) hepatostomus (Pfeiffer). Breure & Eskens 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden (216):28; pl. Type Locality.Tepenistlahuaca, Oaxaca, Mxico. Lectotype BMNH 1975571 (Breure & Eskens 1981). Distribution.OAXACA: Juquila (Von Martens 1893). Drymaeus ( Mesembrinus) inglorius inglorius (Reeve 1848)Bulimus inglorius 368 (shell). Otostomus inglorius (Reeve). Von Martens 1893; Biol. Cent. Amer.:219. Drymaeus inglorius (Reeve). Pilsbry 1899; Man. Conch. (2) 12:67 68; pl. 3, 43 (shell). Drymaeus (Mesembrinus) inglorius (Reeve). Breure & Eskens 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden (216):73; Type Locality.Juquila, Oaxaca, Mxico. Lectotype BMNH 1975536 (Breure & Eskens 1981). Distribution.OAXACA: Oaxaca (Breure & Eskens 1981) Drymaeus (Mesembrinus) inglorius heynemanni (Pfeiffer 1866)Bulinus heynemanni Pfeiffer 1866; Malak. Bltt. 13:89. Bulimulus (Scutalis ) heynemanni (Reeve). Fischer & Crosse 1875;

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116 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Miss. Sci. Mex.:527. Bulimulus heynemanni (Reeve). Strebel 1882; Beitrag. Mex. LandOtostomus inglorius var. heynemanni (Reeve). Von Martens 1893; Biol. Cent. Amer.:220. Drymaeus inglorius heynemanni (Reeve). Pilsbry 1899; Man. Type Locality.Orizaba, Veracruz, Mxico. Distribution.OAXACA: Cerro San Antonio de la Cal; Tlacolula. PUEBLA: Tecomavaca, SE of Tehuacn (Von Martens 1893). VERACRUZ: type locality. Drymaeus (Mesembrinus) necaxanus Solem 1955Drymaeus necaxanus Solem 1955; Occ. Pap. Mus. Zool. Univ. Type Locality.Pine slope near Salto Grande, Necaxa, Puebla, Mxico. Holotype UMMZ 181377. Distribution.Known only from the type locality. Drymaeus (Mesembrinus) (Pfeiffer 1862) Peiffer 1862; Malak. Bltt. 9:153. (Pfeiffer). Von Martens 1893; Biol. Cent. Amer.:218. (Pfeiffer). Pilsbry 1899; Man. Conch. 12:64; pl. Type Locality.Costa Rica. locality. Drymaeus (Mesembrinus) rufescens pinchoti Pilsbry 1930Drymaeus rufescens pinchoti Pilsbry 1930; Proc. Acad. Nat. Sci. Drymaeus (Mesembrinus) rufescens pinchoti Pilsbry. Breure & Eskens 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden anatomy).Type Locality.COLOMBIA, Isla de Providencia: about half a mile inland from South Bay. Holotype ANSP 150852. Distribution.COLOMBIA: known only from Isla de Providencia. Drymaeus (Mesembrinus) sargi sargi (Crosse & Fischer 1875)Bulimulus sargi Crosse & Fischer 1875; Jour. de Conchyl. 23:52. Bulimulus (Scutalus) sargi Crosse & Fischer. Fischer & Crosse Otostomus sargi (Crosse & Fischer). Von Martens 1893; Biol. Cent. Amer.:218. Drymaeus sargi (Crosse & Fischer). Pilsbry 1899; Man. Conch. (2) Type Locality.Tamah, Dept. Alta Verapaz, Guatemala. Distribution.Known only from the type locality. Drymaeus (Mesembrinus) sargi motaguae (Von Martens 1893)Otostomus sargi var. motaguae Von Martens 1893; Biol. Cent. Otostomus sargi ar. montagua Breure 1979.Khler 2007; Mitt. Drymaeus sargi var. motaguae (Von Martens). Pilsbry 1899; Man. Type Locality.Rio Motagua Valley, Dept. Baja Verapaz, Guatemala. Lectotype ZMB 112.877a (Khler 2007). Distribution.Known only from the type locality. Drymaeus ( Mesembrinus) trimarianus (Von Martens 1893)Otostomus trimarianus Von Martens 1893; Biol. Cent. Amer.:216; Drymaeus trimarianus (Von Martens). Pilsbry 1899; Man. Conch. Drymaeus (Mesembrinus) trimarianus (Von Martens). Breure 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168):24.(syntype).Type Locality.Islas Maras, Nayarit, Mxico. Lectotype BMNH 1901.6.22.950 (Breure 1979). Distribution.NAYARIT, Islas Maras: Isla Mara Madre (Jacobson 1958). Group of Drymaeus tripictus Drymaeus (Mesembrinus) irazuensis (Angas 1878)Bulimus irazuensis Angas 1878; Proc. Zool. Soc. Lond. 46:73; pl. 5, Otostomus irazuensis (Angas). Von Martens 1893; Biol. Cent. 1901; Biol. Cent. Amer.:631. Drymaeus irazuensis (Angas). Pilsbry 1899; Man. Conch. (2) Type Locality.Volcn de Iraz, Prov. Cartago, Costa Rica. Distribution.COSTA RICA, Prov. Cartag o: Tierra Blanca, on the south slope of Volcn Irazu 1800 m alt. (Von Martens 1893); Estrella de Cartago (Von Martens 1901). Drymaeus (Mesembrinus) tripictus tripictus (Albers 1857)Bulimus tripictus Albers 1857; Malak. Bltt. 3:97. Otostomus tripictus (Albers). Von Martens 1893; Biol. Cent. Amer.:225. Drymaeus tripictus (Albers). Pilsbry 1899; Man. Conch. (2) 12:69 Bulimulus rhodotrema Von Martens 1868; Malak. Bltt. 15:156.(holotype).Type Locality.Bulimus tripictus : unknown. Bulimulus rhodotrema: Costa Rica; holotype ZMB 14.410 (Khler 2007). locality. Drymaeus (Mesembrinus) tripictus hoffmanni (Von Martens 1893)Otostomus tripictus var. hoffmanni Von Martens 1893; Biol. Cent.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 117Biol. Cent. Amer.:631. Drymaeus tripictus var. hoffmanni (Von Martens). Pilsbry 1899; Drymaeus prestoni DaCosta 1906; Proc. Malac. Soc. London 7:9; Drymaeus prestoni var. cancellata DaCosta 1906; Proc. Malac. Soc. Type Localities. Drymaeus tripictus var. hoffmanni: Woods of San Lorenzo de Dota, Prov. San Jos, Costa Rica; 1300 m alt. Drymaeus prestoni: Chiriqu, Panam. Holotype BMNH 1975478 (Breure 1979). Drymaeus prestoni cancellata: Chiriqu, Panam; holotype BMNH 1907.11.21.13 (Breure 1979). Distribution.COSTA RICA, Prov. Heredia: Heredia (Von Martens 1893); slope of Volcn de Barba, 1600 m alt. Prov. San Jos: Tarbaca, 1700 m alt. PANAM, Prov. Chiriqu: Chiriqu (DaCosta 1906). Drymaeus (Mesembrinus) gabbi (Angas 1879)Bulimus gabbi Angas 1879; Proc. Zool. Soc. Lond. 47:477; pl. 40, Otostomus gabbi (Angas). Von Martens 1893; Biol. Cent. Amer.:207. Drymaeus gabbi (Angas). Pilsbry 1899; Man. Conch. (2) 12:70; Pico Blanco, at an altitude of about 3000 to 6000 ft. Lectotype BMNH 1879.7.22.23 (Breure 1979). Distribution.COSTA RICA, Prov. Heredia: La Paz, on the road to the Rio Sarapiqu (Von Martens 1893). Drymaeus (Mesembrinus) pilsbryi Zetek 1933Drymaeus pilsbryiType Locality.Isla Barro Colorado, Canal Zone, Panam. Holotype ANSP 162124. Distribution.PANAM, Canal Zone: Pedro Miguel. San Blas Com.: San Blas; Ustupu [Ustupo] (Zetek 1933). Group of Drymaeus totonacus Drymaeus (Mesembrinus) albostriatus (Strebel 1882)Bulimulus albostriatus Strebel 1882; Beitrag. Mex. Landund Drymaeus alsostriatus (Strebel). Pilsbry 1899; Man. Conch. (2) 12: Type Locality.Tehuantepec, Oaxaca, Mxico. Distribution.Known only from the type locality. Drymaeus (Mesembrinus) championi (Von Martens 1893)Otostomus championi Von Martens 1893; Biol. Cent. Amer.:222; pl. Drymaeus championi (Von Martens). Pilsbry 1899; Man. Conch. (2) Drymaeus (Drymaeus) championi (Von Martens). Breure & Eskens 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden (216):14.Type Locality.Hacienda de Las Nubes, Cerro Zunil, [Dept. Quetzaltenango], Guatemala. Lectotype BMNH 1901.6.22.451 (Breure & Eskens 1981). Distribution.Known only from the type locality. Drymaeus (Mesembrinus) dominicus (Reeve 1850)Bulimus dominicus Bulimulus dominicus (Reeve). Fischer & Crosse 1875; Miss. Sci. Mex.:540.Strebel 1882; Beitrag. Mex. Landund Sssw.Conch. V:94. Drymaeus dominicus (Reeve). Pilsbry 1899; Man. Conch. (2) 12:3 Clench 1933; Pub. Carnegie Inst. Wash. (431):532.Branson & McCoy 1965:8.Thompson 1967; Bull. Fla. St. Mus. 11:249.Drymaeus (Leptodrymaeus) dominicus (Reeve). Pilsbry 1946; Land Drymaeus (Mesembrinus) dominicus (Reeve). Breure & Eskens Type Locality.Florida, United States of America. Lectotype BMNH 1975199 (Breure & Eskens 1981. Distribution.Widely distributed from Florida to Cuba and Hispaniola, and from eastern Mxico south to Nicaragua. NICARAGUA: San Nicolas (Pilsbry 1899). Depts. Leon, and Chontales 11 mi. E of Cd. Campeche (Branson & McCoy 1965); 19.2 mi. E of Silvituc (Thompson 1967). QUINTANA ROO: 4.0 mi. E of Xpujil (Thompson 1967). TABASCO. VERACRUZ: Mirador; Callejon de Zamarana, near Veracruz (Pilsbry 1899). YUCATN: Chichen Itza (Bequaert & Clench 1933); Labna (Pilsbry 1899). Drymaeus (Mesembrinus) emeus (Say 1829)Bulimus emeus Say 1829a; New Harmony Disseminator of Useful Knowledge:25. Otostomus emeus (Say). Von Martens 1893; Biol. Cent. Amer.:222,; Drymaeus emeus (Say). Pilsbry 1899; Man. Conch. 12:73; pl. 41:675.Correa-Sandoval 1997; Rev. Biol. Trop. 44/45:140.Correa-Sandoval 1999; Rev. Biol. Trop. 47:Correa-Sandoval, Gutirrez & Reza 1998; Acta Zool. Mex. (73):17.CorreaSandoval 2000; Acta Zool. Mex. (79):9.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):238. Drymaeus (Mesembrinus) emeus (Say). Breure & Eskens 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden (216):66; textBulimulus palpaloensis Strebel 1882; Beitrag. Mex. Landund & Jannsen 2004; Archiv fr Molluskenkunde 123:221; pl. 16, Otostomus emeus var. hypozonus Von Martens 1893; Biol. Cent. Amer.:223.Khler 2007; Mitt. Mus. fr Nat. Berlin, Zool. Otostomus emeus var. Von Martens 1893; Biol. (letotype).Type Localities. Bulimus emeus: road from Veracruz to Mxico City. Bulimulus palpaloensis three localities near Misantly, Veracruz, Mxico. Otostomus emeus var. hypozonus: Jalapa, Veracruz; lectotpe ZMB 109.947a (Khler 2007). Otostomus emeus var. :

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118 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Playa Vicente, Veracruz; lectotype ZMB 109946a (Khler 2007). Distribution.Reported from northeastern Mxico south to Espirito Santo, Brazil (Breure & Eskens 1982). NUEVO LEN: Santiago (Correa-Sandoval 1993); Iturbde (Correa-Sandoval 1997). PUEBLA: near Necaxa (Solem 1955a). SAN LUIS POTOS: numerous localities (CorreaSandoval et al.. 1998). TAMAULIPAS: numerous localities (Correa-Sandoval & Rodriguez 2002); El Cielo Biosphere Reserve (Correa-Sandoval &Rodriguez 2005). VERACRUZ: base of Volcn San Martin (Brere & Eskens 1982); San Juan Drymaeus (Mesembrinus) intrapictus Pilsbry 1930Drymaeus intrapictus Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. Type Locality.Tonosi, Prov. Los Santos, Panam. Holotype ANSP 140834. Distribution.Known only from the type locality. Drymaeus (Mesembrinus) perductorum Rehder 1943Drymaeus perductorum (shell).Breure & Eskens 1981; Zool. Verh. Rijkmuseum Nat. Type Locality.Near Las Grutas de Cacahuamilpa, Guerrero, Mxico. Holotype USNM 517552. Distribution.GUERRERO: know only from the immediate vicinity of the type locality. Drymaeus (Mesembrinus) semimaculatus Pilsbry 1898Bulimus maculatus Lea 1839; Trans. Amer. Philos. Soc. 6:86; pl. 23, Bulimus maculatus Bruguiere 1798). Otostomus maculatus (Lea). Von Martens 1893; Biol. Cent. Drymaeus semimaculatus Pilsbry 1898; Man. Conch. (2) 11:297 Sci. Phila. 78:84.Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. 82:340.Morrison 1946; Smiths. Misc. Coll. 106:43. Drymaeus (Mesemnbrinus) semimaculatus Pilsbry. Breure & Eskens 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden (216):84; Type Locality.Cartagena, Colombia. Distribution.Colombia north to Guatemala. PANAM, Archipilago de las Perlas: Isla San Jos (Morrison 1946). Canal Zone: Isla Barro Colorado (Pilsbry 1930d). Prov. Chiriqu: Chiriqu. Prov. Panam: Garachino. COSTA RICA, Prov. Cartago: Tapant (Breure & Eskens 1982). NICARAGUA: San Nicolas (Martens 1893). GUATEMALA, Dept. Petn: Dolores; San Luis. Dept. Retalhuleu: Champerico (Von Martens 1893). Drymaeus (Mesembrinus) totonacus (Strebel 1882)Bulimulus totnacus Strebel 1882; Beitrag. Mex. Landund Sssw.Otostomus totonacus (Strebel). Von Martens 1893; Biol. Cent. Amer.:221. Drymaeus totonacus 13 (shell).Type Locality.Not stated. Distribution.VERACRUZ: Rancho Quilate, near Misantla; Agua Caliente, near Misantla (Strebel 1892). Drymaeus (Mesembrinus) tryoni tryoni (Fischer & Crosse 1875)Bulimus mexicanus (Lamarck). Reeve 1850; Conch. Icon. Bulimus: Bulimulus (Thaumastus) tryoni Fischer & Crosse 1875; Miss. Sci. Mex.:565. Otostomus tryoni (Fischer & Crosse). Von Martens 1893; Biol. Cent. Amer.:232. Drymaeus tryoni (Fischer & Crosse). Pilsbry 1899; Man. Conch. (2) Type Locality.Sinaloa, Mxico. locality. Drymaeus (Mesembrinus) tryoni pochutlensis (Crosse & Fischer 1875)Bulimulus tryoni pochutlensis Crosse & Fischer 1875, in Fischer & Otostomus tryoni var. pochutlensis (Crossse & Fischer). Von Martens 1893; Biol. Cent. Amer.:233. Drymaeus tryoni var pochutlensis (Crosse & Fischer). Pilsbry 1899; Man. Conch. (2) 12:76; pl. 3, 54 (shell).Type Locality.Pochutla, near Chilapa, Guerrero, Mxico. Distribution.Known only from the type locality. Group of Drymaeus sulfureus Drymaeus (Mesembrinus) moricandi moricandi (Pfeiffer 1846)Bulimus moricandi Pfeiffer 1846; Proc. Zool. Soc. Lond. 14:113. Ulimulus (Drymaeus) moricandi (Pfeifffer). Fischer & Crosse 1875; Drymaeus moricandi (Pfeiffer). Pilsbry 1899; Man. Conch. 12:78 Drymaeus (Mesembrinus) moricandi (Pfeiffer). Breure & Eskens 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden (216):78; Type Locality.Mount Cobn, Central America. Lectotype BMNH 1975212 (Breure 1979). Distribution.VERACRUZ: above Rio Metlac 19 mi. SW and 0.8 mi. N of Fortn (Breure & Eskens 1981). Drymaeus moricandi hyalinoalbidus (Fischer & Crosse 1875)Bulimulus (Drymaeus) moricandi var. hyalino-albida Fischer & Crosse 1875; Miss. Sci. Mex.:498. Otostomus moricandi var. hyalino-albida (Fischer & Crosse). Von

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 119Martens 1893; Biol. Cent. Amer.:227. Drymaeus moricandi var. hyalino-albidus (Fischer & Crosse). Pilsbry 1899; Man. Conch. 12:79.Type Locality.Chiapas, Mxico. Distribution.GUATEMALA, Dept. Izabal: Izabal Drymaeus (Mesembrinus) sulfureus (Pfeiffer 1856)Bulimus sulfureus Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:318; pl. Bulimulus sulfureus (Pfeiffer). Fischer & Crosse 1875; Miss. Sci. Otostomus sulfureus (Pfeiffer). Von Martens 1893; Biol. Cent. Cent. Amer.:631. Drymaeus sulfureus (Pfeiffer). Pilsbry 1899; Man. Conch. (2) 12:76 51.Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 72:5.Van der Schalie 1940; Occ. Pap. Mus. Zol. Univ. Mich. (413):4.Haas 1949; Nautilus 62:138.Solem 1955; Occ. Pap. Mus. Zool. Branson & McCoy 1963; Nautilus 76:106.Correa-Sandoval et al. 1998:17.Correa-Sandoval 2000; Acta Zool. Mex. (79):9.Correa-Sandoval 2002; Acta Zool. Mex. (86):238. Otostomus sulfurous var. albidus Von Martens 1893; Biol. Cent. Amer.:226.Khler 2007; Mitt. Mus. fr Nat. Berlin, Zool. Otostomus sulfurous var. gracilior Von Martens 1893; Biol. Cent. Bulimus citronellus Angas 1879; Proc. Zool. Soc. Lond. 47:479; pl. Otostomus sulfureus var. obesus Von Martens 1893; Biol. Cent. Type Localities. Bulimus sulfureus Mxico. Bulimus citronellus: Uren to Lipurio, Costa Rica; lectotype BMNH 1879.7.22.19 (Breure 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168):117). Otostomus sulfurous var. albidus: Teapa, Tabasco; lectotype ZNB 109.947a (Khler 2007). Otostomus sulfurous var. gracilior: Senahu, Dept. Alta Verapaz, Guatemala; lectotype ZMB 109.887a (Khler 2007). Otostomus sulfureus var. obesus: Huatusco, Veracruz, Mxico. Distribution.COSTA RICA, Prov. Alajuela: San near Juan Vias, 2500 ft. alt.; Turrialba; Tuis; Santa Clara. Prov. Heredia: San Juan, Sarapiqu, 100 m alt. Prov. Farms, Rio Surubres, 700 ft. alt. (Pilsbry 1920c). Prov. San Jos: San Jos, 1161 m alt. NICARAGUA, Dept. Chontales: La Libertad. GUATEMALA, Dept. Alta Verapaz: north of Cobn; Chiacam; San Joaquin; Chacoj; Senahu; Panzos (Pilsbry 1899); Cham (Hinkley 1920); Samac; Panzamala (van der Schalie 1940). Dept. Izabal: Jocolo (Hinkley 1920). Dept. Zacapa: Santa Clara (Haas 1949). CAMPECHE: Cd. de Carmen; 16 mi. E of Cd. Campeche (Branson & McCoy 1963). SAN LUIS POTOS: entrance to Las Pozas (Xilitla), W); Carr. TamazunchaleChapulhuacn, 15.5 km from Teapa. NUEVO LEN: Hwy Santiago-Montemorelos, km. 2005). TAMAULIPAS: numerous localities in southern part of state (Correa-Sandoval & Rodriguez 2002); El Cielo Biosphere Reserve (Correa-Sandoval & Rodriguez 2005). Soncoantla; Jalapa (Pilsbry 1899); Sumidero (Solem 1955a); YUCATN: 1.5 mi. S of Libre Union (Branson & McCoy 1963). Group of Drymaeus multilineatus Drymaeus (Mesembrinus) discrepans (Sowerby 1833)Bulimus discrepans Sowerby 1833; Proc. Zool. Soc. Lond. 1:72. Bulimulus (Liostracus) discrepans (Sowerby). Fischer & Crosse 1875; Miss. Sci. Mex.:503. Otostomus discrepans (Sowerby). Von Martens 1893; Biol. Cent. Amer.:230. Drymaeus discrepans (Sowerby). Pilsbry 1899; Man. Conch. (2) 215. El Salvador Distribution.COSTA RICA, Prov. Guanacaste: Salinas Bay. NICARAGUA, Dept. Boaco: Masapa. Dept. Grenada: Salama; San Geronimo, near Salama. Dept. Zacapa: Zacapa. (All records from Von Martens 1893). Drymaeus (Mesembrinus) heterogeneus (Pfeiffer 1866)Bulimus heterogeneus Pfeiffer 1866; Malak. Bltt. 8:118. Bulimulus (Liostracus) heterogeneus (Pfeiffer). Fischer & Crosse 1875; Miss. Sci. Mex.:506. Bulimulus heterogeneus (Pfeiffer). Strebel 1882; Beitrag. Mex. Landund Sssw.-Conch. V:92. Otostomus heterogeneus (Pfeiffer). Von Martens 1893; Biol. Cent. Amer.:235. Drymaeus heterogeneus 23 (shell).Type Locality.Vera Cruz.

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120 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Distribution.NAYARIT: Tepc (Von Martens 1893). VERACRUZ: Mirador; Palo Gacho, Rinconada; Jalapa (Von Martens 1893). Drymaeus (Mesembrinus) inusitatus (Fulton 1900)Bulimulus (Drymaeus) inusitatus Fulton 1900.87.Khler 2007; Drymaeus inusitatus (Fulton). Pilsbry 1902; Man. Conch. (2) Drymaeus (Mesembrinus) inusitatus (Fultn). Breure & Eskens 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden (216):75; Type Locality.Costa Rica; holotype BMNH 1901.4.25.28. near Chiriqu Lagoon (Pilsbry 1926). Drymaeus (Mesembrinus) (Pfeiffer 1842) Pfeiffer 1842; Symbolae ad historiam heliceorum II:48. Bulimulus (Mesembrinus) (Pfeiffer). Fischer & Crosse 1875; Miss. Sci. Mex.:543. (Pfeiffer,). Strebel 1882; Beitrag. Mex. Land(Pfeiffer). Von Martens 1893; Biol. Cent. (Pfeiffer). Pilsbry 1899; Man. Conch. (2) Type Locality.Tehuacn, Puebla, Mxico. Distribution.GUERRERO: Chilpancingo. PUEBLA: SE of Tehuacn; Tecomavaca (Von Martens 1893). Drymaeus ( Mesembrinus) moritinctus (Von Martens 1893)Otostomus moritinctus Von Martens 1893; Biol. Cent. Amer.:228; pl. Drymaeus moritinctus (Von Martens). Pilsbry 1899; Man. Conch. Drymaeus (Mesembrinus) moritinctus (Von Martens). Breure & Eskens 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden (216):78.Khler 2007; Mitt. Mus. fr Nat. Berlin, Zool. Type Locality.Chilpancingo, Guerrero, Mxico; 4600 ft. alt. Lectotype BMNH 1901.6.22.841 (Breure & Eskens 1981). Distribution.GUERRERO: Known only from the vicinity of the type locality. Drymaeus (Mesembrinus) multilineatus (Say 1825)Bulimus multilineatus Say 1825; Journ. Journ. Acad. Nat. Sci. Phila. 5:120. Drymaeus multilineatus (Say). Pilsbry 1899; Man. Conch. (2) Branson & McCoy 1965; Univ. Colorado Stud, Biol., (13):8.Correa-Sandoval, Gutirrez & Reza 1998; Acta Zool. Mex. (73):17.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. Drymaeus (Mesembrinus) multilineatus (Say). Breure & Eskens 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden (216):78; Bulimus sisalensis Morelet 1849; Test. Noviss. I:9.Neubert & 193 (paralectotype).Type Locality.Bulimus multilineatus : Miami, Dade Co., Florida. Neotype ANSP 12309a (Pilsbry 1946). Bulimus sisalensis: Sisal, Yucatn, Mxico. Distribution.South Florida, Colombia, Venezuela, Cd. Campeche (Branson &McCoy 1965:8). OAXACA: 4.0 mi. WNW of Tehantepec (Breure & Eskens 1981). SAN LUIS POTOS: Valles (Hinkley 1907); Rancho El Rodeo; Ejido SandovalSandoval et al. 1998). TAMAULIPAS: numerous localities (Correa-Sandoval & Costro 2002). YUCATN Sisal. Drymaeus (Mesembrinus) semipellucidus (Tristram 1861)Bulimus semipellucidus Tristram 1861, Proc. Zool. Soc. Lond. Bulimulus (Liostracus) semipellucidus (Tristram). Fischer & Crosse 1875; Miss. Sci. Mex.:507. Otostomus semipellucidus (Tristram). Von Martens 1893; Biol. Drymaeus semipellucidus (Tristram). Pilsbry 1899; Man. Conch. (2) Drymaeus (Mesembrinus) semipellucidus (Tristram). Breure & Eskens 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden Type Locality.Guatemala. Distribution.COSTA RICA, Prov. Alajuela: Alajuela. Prov. Guanacaste: Salinas Bay. Prov. Heredia: Puerto Viejo, Rio Sarapiqu (Von Martens 1893). NICARAGUA, Dept. Masaya:2.4 mi. N of Masatepe (Breure & Eskens 1981). Drymaeus ( Mesembrinus) shattucki Bequaert & Clench 1931Drymaeus shattucki Bequaert & Clench 1931:424.Bequaert & Clench 1933; Pub. Carnegie Inst. Wash. (431):533; pl. 68, (457):64.Bequaert & Clench 1938; Pub. Canegie Inst. Wash. (491):258.Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):10.Branson & McCoy 1963; Nautilus 76:106.Type Locality.Chichen Itza, Yucatn, Mxico. Holotype MCZ 79396. Distribution.GUATEMALA, Dept. Petn: Tikal National Park (Basch 1959). QUINTANA ROO:3 km S of San Miguel, Isla Cozumel (Branson & McCoy 1963). YUCATN: Tabi; Progreso (Bequaert & Clench 1933); Ebitz Cave, Oxkutzcab (Bequaert & Clench 1938).

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 121 Drymaeus (Mesembrinus) tropicalis (Morelet 1849)Bulimus tropicalis Morelet 1849; Test. Noviss. I:9.Neubert & 194 (paralectoype). Bulimulus (Liostracus) tropicalis (Morelet). Fischer & Crosse 1875; Bulimulus tropicalis (Morelet). Strebel 1882; Beitrag. Mex. Landund Sssw.-Conch. V:95.Pilsbry 1891; Proc. Acad. Nat. Sci. Phila.:315. Otostomus tropicalis (Morelet). Von Martens 1893; Biol. Cent. Amer.:233. Drymaeus tropicalis (Morelet). Pilsbry 1899; Man. Conch. (2) Pub. Carnegie Inst. Wash. (431):533.Bequaert & Clench 1938; Pub. Canegie Inst. Wash. (491):258.Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):10.Branson & McCoy 1963; Nautilus 76:106.Thompson 1967; Bull. Fla. St. Mus. 11:248. Drymaeus (Mesembrinus) tropicalis (Morelet). Breure & Eskens 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden (216):89. Type Locality.Mxico, Campeche, civic plaza of Cd. Campeche. Lectotype BMNH 1893.2.4.210 (Breure & Eskens 1981). Distribution.GUATEMALA, Dept. Petn: Tikal National Park (Basch 1959). CAMPECHE: 51 mi. E of Cd. Campeche; 32 mi. S of Cd. Campeche (Branson & McCoy 1963); 5.1 mi. NNW of Dzibalchn; 3.6 mi. S of Hopelchn; 5.7 mi. E of Cd. Campeche (Thompson 1967). QUINTANA ROO: 4.0 mi. E of Xpujil (Thompson 1967). YUCATN: Labna; Tabi, S of Merida (Pilsbry 1891); Ebitz Cave, Oxkutzcab (Bequaert & Clench 1938); 0.8 mi. NE of Becanchn; 19.1 mi. SSE of Uman (Thompson 1967). Drymaeus (Mesembrinus) uhdeanus uhdeanus (Von Martens 1893)Bulimulus (Mesembrinus) uhdeanus Von Martens 1863; in Monatsber. Akad. Wiss. Berl. 26:541.Fischer & Crosse 1875; 11b (shell).Khler 2007; Mitt. Mus. fr Nat. Berlin, Zool. Otostomus uhdeanus (Von Martens). Von Martens 1893; Biol. Cent. Drymaeus uhdeanus (Von Martens). Pilsbry 1899; Man. Conch. (2) Pap. Mus. Zool. Univ. Mich. (566):11. Drymaeus (Mesembrinus) uhdeanus (Von Martens). Breure & Eskens 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden Drymaeus herrerae Bartsch 1907; Proc. U. S. Nat. Mus. 32:119, Type Locality.Bulimulus uhdeanus: Mxico; lectotype ZMB 4.575b (Khler 2007). Drymaeus herrerae: Bonanza Zimapan, Hidalgo, Mxico; holotype USNM 192997. Distribution.HIDALGO: Zimapan. JALISCO: Sayula. VERACRUZ: Aculcingo, south of Orizaba (Pilsbry 1899); 8.5 mi. SW of Cd. Mendoza, 5200 ft. alt. (Breure & Eskens 1981). Drymaeus (Mesembrinus) uhdeanus borealis (Von Martens 1893)Otostomus uhdeanus var. borealis Von Martens 1893; Biol. Cent. Drymaeus uhdeanus borealis (Von Martens). Pilsbry 1899; Man. Type Locality.Ventanas, Durango, Mxico; 2000 ft. alt. Lectontype ZMB 101.817a (Khler 2007). Distribution.Known only from the type locality. Drymaeus (Mesembrinus) (Crosse & Fischer 1874)Bulimulus (Scutalus) Crosse & Fischer 1874; Jour. de Conchyl. 22:283.Fischer & Crosse 1875; Miss. Sci. Otostomus uhdeanus var. (Crosse & Fischer). Von (Crosse & Fischer). Pilsbry 1955; Occ. Pap. Mus. Zool. Univ. Mich. (566):11. Type Locality.: Cuernovaca, Morelos. Veracruz, Mxico. Distribution.MORELOS: Cuernovaca. VERACRUZ: Orizaba; Maltrata (Pilsbry 1899). Drymaeus (Mesembrinus) uhdeanus tepicensis (Von Martens 1893)Otostomus uhdeanus var. tepicensis Von Martens 1893; Biol. Cent. Drymaeus uhdeanus tepicensis (Von Martens). Pilsbry 1899; Man. Type Locality.Tepc, Jalisco [Nayarit] Mxico. Distribution.NAYARIT: Isla Mara Madre, Islas Maras (Dall 1926). Group of Drymaeus translucens Drymaeus (Mesembrinus) cozumelensis Richards 1937Drymaeus shattucki cozumelensis Richards 1937; Proc. Amer. Type Locality.On trail from San Miguel to Santa Rita, Isla Cozumel, Quintana Roo, Mxico. Holotype ANSP 167744. Distribution.QUINTANA ROO: known only from Isla Cozumel. Drymaeus (Mesembrinus) hondurasanus (Pfeiffer 1846)Bulimus hondurasanus Pfeiffer 1846; Proc. Zool. Soc. Lond. 14:29. Bulimulus (Liostracus) hondurasanus (Pfeiffer): Fischer & Crosse 1875; Miss. Sci. Mex.:503. Otostomus hondurasanus (Pfeiffer). Von Martens 1893; Biol. Cent. Amer.:232. Drymaeus hondurasanus (Pfeiffer). Pilsbry 1899; Man. Conch. (2)

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122 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Drymaeus (D.) alternans honduranus (Pfeiffer). Haas & Solem 1960; Nautilus 73:131.Type Locality.Honduras. Lectotype in the BMNH 1975265 (Rehder 1966). Distribution.BELIZE: Rio Frio Cave; Kates Lagoon (Haas & Solem 1960). HONDURAS: not known from a Drymaeus (Mesembrinus) mayaorum Rehder 1966Drymaeus mayaorum Rehder 1966; Proc. Biol. Soc. Wash. 79:287; Type Locality.Isla Mujeres, Quintana Roo, Mxico. Holotype USNM 251656. Distribution.QUINTANA ROO: Isla Mujeres; near San Miguel, Isla Cozumel (Rehder 1966). Drymaeus (Mesembrinus) translucens translucens (Broderip 1842)Bulimus translucens Broderip 1842; Proc. Zool. Soc. Lond. 10:31. Bulimulus translucens (Broderip). Von Martens 1893; Biol. Cent. Amer.:250. Drymaeus translucens (Broderip). Pilsbry 1899; Man. Conch. (2) Type Locality.Islas del Rey and Sabogas, Panam. Distribution.PANAM, Archipilago de las Perlas: Isla del Rey; Isla Saboga (Pilsbry 1926b); Isla San Jos (Morrison 1946). NICARAGUA, Dept. Boaco (Prez & Drymaeus ( Mesembrinus) translucens alternans (Beck 1837)Bulimus (Bulimulus) alternans Beck 1837; Index molluscorum.:65. Bulimulus(Liostracus) alternans (Beck). Fischer & Crosse 1875; Otostomus alternans (Beck). Von Martens 1893; Biol. Cent. Amer.:230.Von Martens 1901; Biol. Cent. Amer.:632. Drymaeus alternans (Beck). Pilsbry 1899; Man. Conch. (2) 12:86 Drymaeus translucens alternans (Beck). Pilsbry 1930; Proc. Acad. Drymaeus (Mesembrinus) alternans (Beck). Breure & Eskens 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden (216):47; textType Locality.Isla Saboga, Panam. Distribution.COSTA RICA, Prov. Alajuela: Alajuela (Von Martens 1893). Prov. Cartago: Cartago, 1142 m alt. Prov. Puntarenas: San Diquis Valley, near Terraba, 100 m alt. Prov. San Jos: San Jos; La Uruca, 100 m alt. (Von Martens 1893 1901). GUATEMALA, Dept. Baja Verapaz: San Gronimo, near Salama (Von Martens 1893). Dept. Guatemala: Cd. Guatemala (Hinkley 1920). Dept. Sacatepquez: Capetillo, near Antigua (Von Martens 1901). NICARAGUA, Dept. Matagalpa: 1 mi. E of Matagalpa (Breure & Eskens 1981). de las Perlas: Isla del Rey; Isla Saboga (Pilsbry 1926b). Drymaeus (Mesembrinus) translucens juquilensis (Von Martens 1893)Otostomus alternans var. juquilensis Von Martens 1893; Biol. Cent. Amer.:231. Drymaeus alternans var. juquilensis (Von Martens). Pilsbry 1899; Type Locality.Juquila, Oaxaca, Mxico. Distribution.OAXACA: Juquila. GUATEMALA: Vera Paz (Von Martens 1893). Drymaeus (Mesembrinus) translucens misellus Pilsbry 1926Drymaeus translucens form misellus Pilsbry 1926; Proc. Acad. Nat. Type Locality.Tonosi, Prov. Los Santos, Panam. Holotype ANSP 140281. Distribution.Known only from the type locality. Drymaeus (Mesembrinus) translucens pachecensis Pilsbry 1930Drymaeus translucens form pachenensis Pilsbry 1930d; Proc. Acad. Type Locality.Isla Pacheco, Archipilago de las Perlas, Panam. Holotype ANSP 151302. Distribution.Known only from the type locality. Drymaeus (Mesembrinus) translucens panamensis (Broderip 1833)Bulimus panamensis Broderip 1833; Proc. Zool. Soc. Lond. 1:105. Otostomus panamensis (Broderip). Von Martens 1893; Biol. Cent. Amer.:236. Drymaeus panamensis (Broderip). Pilsbry 1899; Man. Conch. (2) Drymaeus translucens form panamensis (Broderip). Pilsbry 1930; Type Locality.San Miguel, Isla del Rey, Panam. Distribution.PANAM: Isla del Rey, Isla Saboga (Pilsbry 1926). Drymaeus ( Mesembrinus) translucens sororcula Pilsbry 1926Drymaeus translucens sororcula Pilsbry 1926; Proc. Acad. Nat. Sci. Type Locality.Isla Taboga, Panam. Holotype ANSP 45238. Distribution.Known only from the type locality. Drymaeus (Mesembrinus) Pilsbry 1899Drymaeus translucens var. Pilsbry 1899; Man. Conch. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168):124.Type Locality.Nicaragua. Lectotype ANSP 25040a (Breure 1979). known.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 123 Drymaeus ( Mesembrinus) translucens tonosiesis Pilsbry 1930Drymaeus alternans (Beck). Pilsbry 1926; Proc. Acad. Nat. Sci. Drymaeus translucens tonosiesis Pilsbry 1930:340. Type Locality.Tonosi, Prov. Los Santos, Panam. Holotype ANSP 151288. Distribution.Known only from the type locality. Genus Naesiotus Albers 1850Naesiotus Albers 1850:162.Breure 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168):64.Hoffman 1988; Veliger 30:117.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47: 11219. Sonorina Pilsbry 1896: Nautilus 9: 114. Puritanina Jacobson 1958; Amer. Mus. Nov. (1899):7. Hannarabdotus Emerson & Jacobson 1965; Trans. San Diego Soc. Nat. Hist. 13:325.Type Species.Naesiotus: Bulimus nux Broderip 1832. Sonorina: Bulimulus spirifer Gabb 1868. Puritanina : Bulimulus montezuma Dall 1893. Hannarabdotus: Bulimulus Hanna 1923. Distribution.South America, Islas Galpagos, the Windward Islands of the West Indies, northwestern Mxico, and Arizona (Breure 1978; Hoffman 1988). Taxonomy.The generic synonymy given above applies only to the Mexican fauna. Breure (1979:67) listed over 160 names that have been proposed for species of Naesiotus. Twenty species are recognized in the study area. An Arizona species, Naesiotus christenseni, is included because of its Naesiotus altus (Dall 1893)Naesiotus altus (Dall). Hoffman 1988; Veliger 30:420.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. Bulimulus (Leptobrysus) subspirifer Mabille 1895; Bulletin de la Socit Philomathique de Paris (8) 7:67.Christensen & Miller 1976; Western Soc. Malac. Ann. Rep., 9:51.Type Localities. Bulimulus altus: Sierra Laguna, Baja California Sur, Mxico; 3000 ft. alt. ( Christensen 1978). Bulimulus subspirifer: Lower California. Distribution.BAJA CALIFORNIA SUR: 2.3 km from main hwy. on road to microwave station San Bartolo, 430 m alt.; ca. 1 km SE of San Bartolo, 240 m alt.; 2.1 km SE of San Bartolo (Smith et al. 1990). Naesiotus beldingi (Cooper 1892)Bulimulus inscendens var. beldingi Cooper 1892; Proc. Calif. Acad. Sci. (2) 3:209. Bulimulus (Orthotomium) beldingi Cooper. Dall 1895; Proc. U. S. Nat. Mus.19:357.Pilsbry 1898; Man. Conch. 11:149; pl. 25, Naesiotus beldingi (Cooper). Hoffman 1988; Veliger 30:420.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. Bulimulus (Leptobrysus) dismenicus Mabille 1895; Bulletin de la Socit Philomathique de Paris (8) 7:67.Emerson & Jacobson 1964; Trans. San Diego Soc. Nat. Hist. 16:51. Type Localities. Bulimulus beldingi : Sierra Laguna, Punta Arena, near San Jos del Cabo, Baja California Sur, Mxico. Bulimulus dismenicus: Sierra de Puna, Baja California Sur 1800 m alt. Distribution.BAJA CALIFORNIA SUR: several localities in the Sierra Laguna at 610 m alt. (Smith et al. 1990). Naesiotus christenseni (Miller & Reeder 1984)Rabdotus christenseni Miller & Reeder 1984; Bull.Southern Naesiotus christenseni (Miller & Reeder). Hoffman 1988; Veliger 30:417.Type Locality.Arizona, Pima Co.; north end of the Santa Rita Mountains, in a large rockslide on the left bank of south fork of Sycamore Creek, ca. 100 m upstream from Holotype CAS 37525. Distribution.Known only from the type locality. Naesiotus cosmicus (Mabille 1895)Bulimulus (Scutalus) cosmicus Mabille 1895; Bulletin de la Socit Philomathique de Paris (8) 7:68. Bulimulus (Orthotomium) cosmicus Mabille. Pilsbry 1897; Man. Conch. 11:144. Naesiotus cosmicus (Mabille). Hoffman 1988; Veliger 30:420.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. Type Locality.La Sierra du Sud dela Presquile, Baja California Sur, Mxico. Distribution.BAJA CALIFORNIA SUR: stations on the NE side of the Sierra Laguna (Smith et al. 1990). Naesiotus dentifer dentifer (Mabille 1895)Bulimulus (Leptobrysus) dentifer Mabille 1895; Bulletin de la Socit Philomathique de Paris (8) 7:67. Bulimulus (Sonoribna) dentifer Mabille. Pilsbry 1898; Man. Conch. 11:161.Hanna 1923; Proc. Calf. Acad. Sci. (4), anatomy, jaw). Rabdotus (Leptobrysus) dentifer (Mabille) Emerson & Jacobson 1964; Trans. San Diego Soc. Nat. Hist. 16:319. Naesiotus dentifer dentifer (Mabille). Hoffman 1988; Veliger 30:420.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Type Locality.Isla Tortuga, Baja California Sur, Mxico. Distribution.BAJA CALIFORNIA SUR: known only from Isla Tortuga (Smith et al. 1990). Naesioyus dentifer johnstoni (Hanna 1923)Bulimulus johnstoni Hanna 1923; Proc. Calif. Acad. Sci. 12:491; pl. Rabdotus (Hannarabdotus) johnstoni (Hanna). Emerson & Jacobson 1964; Trans. San Diego Soc. Nat. Hist. 16:326. Naesiotus dentifer johnstoni (Hanna). Smith, Miller, Christensen &

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124 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Type Locality.Isla Santa Catalina, Baja California Sur, Mxico. Holotype CAS 1024. Distribution.BAJA CALIFORNIA SUR: known only from Isla Santa Catalina (Smith et al. 1990). Naesiotus dentifer lamellifer (Pilsbry 1897)Bulimulus lamellifer Pilsbry 1897; Nautilus 10:103. Bulimulus (Sororina) lamellifer Pilsbry 1898; Man. Conch. 11:160; (4), 12:495. Rabdotus (Leptobrysus) lamellifer lamellifer (Pilsbry). Emerson & Jacobson 1964; Trans. San Diego Soc. Nat. Hist. 16:321; Naesiotus dentifer lamellifer (Pilsbry). Smith, Miller, Christensen (map). Bulimulus spirifer var. orthelasmus Pilsbry 1898; Man. Conch. 11:159.Christensen & Miller 1976; Western Soc. Malac. Ann. Rep., 9:51. Bulimulus ximenez Hanna 1923; Proceedings of the California Rabdotus (Leptobrysus) ximenez (Pilsbry). Emerson & Jacobson (shell). Bulimulus bakeri Hanna 1923; Proc. Calf. Acad. Sci. (4), 12:500 Bulimulus sanmarcosensis Pilsbry & Lowe 1932; Nautilus 46:49 Bulimulus carmen Pilsbry & Lowe 1932; Nautilus 46:50; Type Localities. Bulimulus lamellifer : Lower California. Bulimulus spirifer var. orthelasmus: not given. Bulimulus ximinez: Marquer Bay, Isla Carmen. Bulimulus bakeri: Punta San Antonio, south of Bahia San Nicolas, Baja California Sur, Mxico; holotype CAS 1017. Bulimulus sanmarcosensis: Isla San Marcos. Bulimulus carmen: Salinas Bay, Isla Carmen. Distribution.BAJA CALIFORNIA SUR: northern peninsular Baja California Sur and Isla San Marcos, Isla Carmen, Isla Danzante, Isla Coronados, and Isla San Jos (Smith et al. 1990). Naesiotus dentifer santacruzensis (Hanna 1923)Bulimulus santacruzensis Hanna 1923; Proc. Calif. Acad. Sci. Hanna 1923; Proc. Calif. Acad. Sci. 12:488; Rabdotus (Hannarebdotus) (Hanna). Emerson & Jacobson (shell). (Hanna). Smith, Miller, Christensen & Type Localities. Bulimulus santacruzensis: Isla Santa Cruz, Baja California Sur, Mxico; holotype CAS 1030. : Isla Monserrate, Baja California Sur, Mxico; holotype CAS 1034. Distribution.BAJA CALIFORNIA SUR: known only from Isla Monserrate and Isla Santa Cruz (Smith et al. 1990). Taxonomy.Smith et al. (1990) used the name for this subspecies, but the name santacruzensis has page priority. Naesiotus durangoanus (Von Martens 1893)Bulimulus durangoanus Von Martens 1893; Biol. Cent. Amer.:246; Type Locality.Villa Lerdo, Durango, Mxico. Distribution.DURANGO: reported only from the type locality. Remarks.There are many samples of this and/or closely related species in the Florida Museum of Natural History from the states of Chihuahua, Durango, Coahuila and that they are properly placed in Naesiotus. Further study is Naesiotus excelsus (Gould 1853)Bulimus excelsus Gould 1853; Jour. Boston Soc. Nat. Hist. 6:376; Bulimulus (Orthotomium) excelsus (Gould). Pilsbry 1897; Man. Naesiotus excelsus (Gould). Hoffman 1988; Veliger 30:420.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. Bulimulus elatus Gould 1853; Jour. Boston Soc. Nat. Hist. 6:406. Bulimulus (Scutatus) cacotycus Mabille 1895; Bulletin de la Socit Philomathique de Paris (8) 7:69.Emerson & Jacobson 1964; Trans. San Diego Soc. Nat. Hist. 16:51. Bulimulus excelsus var. sinaloae Pilsbry 1897; Man. Conch. 11:142.Type Localities. Bulimus excelsus : California. Bulimus elatus: not given. Bulimulus cacotycus : Les Sierras du Sud de la Presquile [Baja California Sur]. Bulimulus excelsus var. sinaloae: Sinaloa (doubtful Hanna 1923; Proc. Calf. Acad. Sci. (4), 12:487; Smith et al. 1990:115). Distribution.BAJA CALIFORNIA SUR: La Paz; N slope of La Calavera Mt. Above Playa Coromuel; Playa Coromuel; 20.6 km E of La Paz on road to Las Cruces; granite cliff near Rancho Viniamos, 2.6 km E of La Paz (Smith et al. 1990). Naesiotus gabbi (Crosse & Fischer 1872)Bulimulus (Scutalus) gabbi Crosse & Fischer 1872; Jour. de Conchyl. 20:223.Fischer & Crosse 1873; Miss. Sci. Mex. Bulimulus (Orthotomium) gabbi Crosse & Fischer. Pilsbry 1898; 35 (shell). Naesiotus gabbi (Crosse & Fischer). Hoffman 1988; Veliger 30:420.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Type Locality.California Mexicana. Distribution.BAJA CALIFORNIA SUR: rock outcrop E of road to La Paz, 4.2 km NE of junct. with road between El Gonzaga and El Obispo 180 m alt.; along road between El

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 125 Obispo and Rancho Tinajitas; arroyo near Microwave Station hwy. at km 77, N of La Paz (Smith et al. 1990). Naesiotus gigantensis (Christensen & Miller 1977)Rabdotus gigantensis Christensen & Miller 1977; Nautilus 91:131 Naesiotus gigantensis (Christensen & Miller). Hoffman 1988; Veliger 30:420.Smith, Miller, Christensen & Roth 1990; Proc. Type Locality.In a large lava rockslide immediately south of the mission at San Javier, Baja California Sur, Mxico; 350 m alt. Holotype CAS 57937. Distribution.BAJA CALIFORNIA SUR: nearly the entire length of the Sierra de la Giganta, numerous localities (Smith et al. 1990). Naesiotus hannai (Pilsbry 1927)Bulimulus hannai Pilsbry 1927; Proc. Calif. Acad. Sci. (4) 17:183 Naesiotus hannai (Pilsbry). Hoffman 1988; Veliger 30:420.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. Type Locality.Isla Margarita, under stones within 1 mi. west and south of the village near the center of the east side of the island, Baja California Sur, Mxico. Holotype CAS 2621. Distribution.BAJA CALIFORNIA SUR: known only from Isla Margarita. Ravine below Mt. Izabal; in rocky outcrops in hills just W of road from Puerto Alcatraz to Puerto Cortez (Smith et al. 1990). Naesiotus harribaueri (Jacobson 1958)Bulimulus (Puritanina ) harribraueri Jacobson 1958; Amer. Mus. Naesiotus haribaueri (Jacobson). Hoffman 1988; Veliger 30:420.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. Type Locality.Sand dunes along Bahia de los Frailes, Baja California Sur, Mxico. Holotype AMNH 74003. Distribution.BAJA CALIFORNIA SUR: several localities along the east coast of Cabo San Lucas (Smith et al. 1990). (Christensen & Miller 1977) Christensen & Miller 1977; Nautilus 91:132 (Christensen & Miller). Hoffman 1988; Veliger 30:420.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Type Locality.In a narrow arroyo approximately 0.5 km inland of the beach at El Limoa, on the west side of Isla Cerrralvo, Baja California Sur, Mxico; 50 m alt. Holotype CAS 57942. Distribution.Known only from the type locality. Naesiotus milleri (Hoffman 1987)Rabdotus milleri Naesiotus milleri (Hoffman). Hoffman 1988; Veliger 30:420.Type Locality.Sonora, Mxico; 5.4 km west of the Rio Yaqui bridge at La Estrella on road to Sahuaripa, in a canyon alt. Holotype SBMNH 3490. Distribution.SONORA: known only from the vicinity of the type locality (Hoffman 1987). Naesiotus montezuma (Dall 1893)Bulimus proteus Binney 1869; Smithsonian Miscellaneous Bulinus proteus Broderip 1832.) Bulimulus (Scutalus) montezuma Dall 1893; Proc. U. S. Nat. Bulimulus (Orthotomium) montezuma Dall. Pilsbry 1898; Man. Bulimulus (Puritanina ) montezuna Ball. Jacobson 1958:101. Naesiotus montezuma (Dall). Hoffman 1988; Veliger 30:420.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. Bulimulus (Scutalus) acholus Mabille 1895; Bulletin de la Socit Philomathique de Paris (8) 7:68.Emerson & Jacobson 1964; Trans. San Diego Soc. Nat. Hist. 16:51.Type Localities.Bulimulus montezuma: Lower California, mostly from the mountainous region. Lectotype USNM 8564 (Jacobson 1958:11). Bulimulus acholus: montagnes de la Base Californie. Distribution.BAJA CALIFORNIA SUR: Cabo San Lucas, numerous localities from 60 m alt. (Smith et al. 1990). Naesiotus nigromontanus (Dall 1897)Bulimulus (alternatus var. ?) nigromontanus Dall 1897; Proc. U. S. Nat. Mus. 19:357. Rabdotus nigromontanus Naesiotus nigromontanus (Dall 1897). Hoffman 1988; Veliger 30:420.Type Locality.Summit of Black Mountain, 12 mi. S of Monument 77 of International Boundary, right bank of Distribution.SONORA: La Loma Colorado; Angostura; Magdalena; Alamos (Hoffman 1987). ARIZONA, Santa Cruz Co.: Pena Blanca Canyon, Pajaritos Mts, ca. 4000 ft. alt. (Bequaert & Miller 1973:137). Naesiotus pallidior (Sowerby 1833)Bulimus pallidior Sowerby 1833; Proc. Zool. Soc. Lond. 1:72. Bulimulus (Orthotomium) pallidior (Sowerby). Pilsbry 1897; Man. Naesiotus pallidior (Sowerby). Hoffman 1988; Veliger 30:420.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. Gould 1853: Jour. Boston Soc. Nat. Hist. 6:375; Bulimulus pallidior var. striatulus Dall 1893:640.Pilsbry 1927;

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126 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Cooper 1894; Proc. Calif. Acad. Sci. (2) 4:134.Type Locality.Not stated. Distribution.BAJA CALIFORNIA SUR: Cabo San Lucas, numerous localities from 60 m alt. (Smith et al. 1990). Naesiotus rimatus (Pfeiffer 1846)Bulimus rimatus Pfeiffer 1846; Proc. Zool. Soc. Lond. 14:112. Bulimulus (Sonorina) rimatus (Pfeiffer). Pilsbry 1898; Man. Conch. Sci. (4), 12:495. Naesiotus rimatus (Pfeiffer). Hoffman 1988; Veliger 30:420.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. Bulimulus (Mesembrinus) inscendens var. bryanti Cooper 1891; Bulimulus (Leptobrysus) bryanti (Cooper). Dall 1893; Proc. U. S. Bulimulus (Leptobrysus) inscendens var. monticola Dall 1893; Proc. U. S. Nat. Mus. 16:643Type Localities. Bulimus rimatus: not given. Bulimulus inscendens var. bryanti: San Jos del Cabo, Baja California Sur, Mxico (restricted by Christensen 1978). Bulimulus inscendens monticolus : Sierra Laguna, Baja California Sur; 900 m alt. (restricted by Christensen 1978). Distribution.BAJA CALIFORNIA SUR: numerous localities on the southern end of Cabo San Lucas below 150 m alt. (Smith et al. 1990). Naesiotus spirifer (Gabb 1868)Bulimus spirifer Bulimulus (Sonorina) spirifer (Gabb). Pilsbry 1898; Man. Conch. Naesiotus spirifer (Gabb). Hoffman 1988; Veliger 30:420.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. Mabille 1895; Bulletin de la Socit Philomathique de Paris (8) 7:66.Emerson & Jacobson 1964; Trans. San Diego Soc. Nat. Hist. 16:51.Type Localities. Bulimus spirifer: in the mountains, among rocks from San Antonio, below La Paz, to near San Borja, and in the mountains perhaps even farther north. : Sierra de la Cachila au Sud de la Paz. Distribution.BAJA CALIFORNIA SUR: Cabo San Lucas, from various localities in the Sierra Cachila and near by areas; 430 m alt. (Smith et al. 1990). (Dall 1893)Bulimulus (Leptobrysus) Dall 1893; Proc. U. S. Nat. Bulimulus (Sonorina) Dall. Pilsbry 1898; Man. Conch. Rabdotus (Leptobrysus) (Dall). Emerson & Jacobson 1964; Trans. San Diego Soc. Nat. Hist. 16:324. (Dall). Hoffman 1988; Veliger 30:420.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. Type Locality.Isla Espiritu Santo, Baja California Sur, Mxico. Holotype USNM 34122. Distribution.BAJA CALIFORNIA SUR: known only from Isla Espiritu Santo (Smith et al.. 1990). Naesiotus xantusi (Binney 1861)Bulimus xanttusi W. G. Binney 1861; Proc. Acad. Nat. Sci. Bulimulus (Orthotomium) xantusi (Binney). Pilsbry 1898; Man. Naesiotus xantusi (Binney). Hoffman 1988; Veliger 30:420.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. Type Locality.Cabo San Lucas, Baja California Sur, Mxico. Distribution.BAJA CALIFORNIA SUR: various localities on the Cabo San Lucas, from 90 m alt. (Smith et al. 1990). Genus Rabdotus Albers 1850Rabdotus Albers 1850; Die Heliceen:164.Pilsbry 1946; Land Moll. N. Amer. 2:4.Distribution.Central Mxico from Puebla north to Texas, the central-southern United States, and Baja California Sur. Taxonomy.Two subgenera and sixteen species are recognized. Subgenus Rabdotus Albers 1850Rabdotus Albers 1850; Die Heliceen:164. Odontostomus Fischer & Crosse 1874; Miss. Sci. Mex. I:473. Globulinus Fischer & Crosse 1874; Miss. Sci. Mex. I:475.Type Species.Rabdotus: Helix dealbatus Say 1821. Odontostomus: Gould 1859. Globulinus: Gould 1859. Distribution.The same as for the genus. Taxonomy.Eleven species and one subspecies occur in the study area. Group of Rabdotus alternatus Rabdotus (Rabdotus) alternatus alternatus (Say 1830)Bulimus alternatus Say 1830; New Harmony Disseminator:25. Bulimulus alternatus alternatus (Say 1830). Pilsbry 1928:117.& McCoy 1963; Nautilus 76:105. Rabdotus alternatus (Say). Correa-Sandoval 1993; Rev. Biol. Trop. 41:675.Correa-Sandoval 1997; Rev. Biol. Trop. 44/45:140.Correa-Sandoval, Garca-Cubas & Reguero 1998:17.CorreaSandoval 2000; Acta Zool. Mex. (79):9.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):238. Bulimus lactarius Menke 1846; in Pfeiffer, Symbolae ad Histori Heliceorum 3:85. stricted by Pilsbry 1946). Holotype ANSP 25694. Distribution.NUEVO LEN: Santa Caterina Valley

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 127 opposite and above the Obispada (Pilsbry 1946); Santiago (Correa-Sandoval-Sadoval 1993); Iturbide (Correa-Sandoval 1997). SAN LUIS POTOS: 0.1 km Ejido Buenavista km 18 W, 1260 m alt.); Ejido Buena Vista, 2 km S of Carr. Antiguo Charro Blanco Branson & McCoy 1963). TAMAULIPAS: Mesa de Solis, near La Lajilla, between Padilla and Jimenez (Pilsbry 1928); numerous records (Correa-Sandoval & Rodriguez 2002). VERACRUZ: La Ordea, Papantla (20 Rabdotus (Rabdotus) alternatus mariae (Albers 1850)Bulimus mariae Albers 1850; Die Heliceen:162. Bulimulus alternatus mariae (Albers). Pilsbry & Ferriss 1906; Proc. Bulimulus (Rabdotus) alternatus mariae (Albers). Pilsbry 1946; Type Locality.Brownsville, Texas (restricted by Pilsbry 1946). Syntype ZMB 112669 (Khler 2007). and Tamaulipas (Pilsbry 1946). Rabdotus (Rabdotus) dealbatus dealbatus (Say 1821)Helix dealbaus ay 1821; Journ. Acad. Nat. Sci. Phila. 2:150. Bulimulus dealbatus (Say). W. G. Binney 1878; Terrestrial Mollusks radula). Bulimulus (Rabdotus) dealbatus (Say). Pilsbry 1946; Land Moll. N. Rabdotus dealbatus dealbatus (Say). Correa-Sandoval 1993; Rev. Biol. Trop. 41:675.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):238.Correa-Sandoval & Salazar 2005; Acta Zool. Mex. (21):61. Bulimus liquabilis Type Localities. Helix dealbatus: Missouri and Alabama. Holotype in the ANSP (Pilsbry 1946). Bulimus liquabilis : Texas; lectotype BMNH 1975422 (Breure 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168):76). : Texas. Distribution.Widely distributed in the eastern United States from Illinois south to Alabama and west to eastern Texas and Oklahoma. Mexican records are as follow. NUEVO LEN: Santiago (Correa-Sandoval 1993; Rev. Biol. Trop. POTOS: Valles (Hinkley 1907). TAMAULIPAS: numerous localities (Correa-Sandoval & Rodriguez 2002). Taxonomy.Six subspecies are recognized, only one of which occurs in Mxico, Rhabdotus d. dealbatus Rabdotus (Rabdotus) pilsbryi (Ferriss 1925)Bulimulus pilsbryi Ferriss 1925; Nautilus 30:25. Bulimulus (Rabdotus) pilsbryi (Ferriss). Pilsbry 1946; Land Moll. Type Locality.Sanderson, Terrell Co., Texas. Holotype ANSP 175851. Distribution.Southwestern Texas, Terrell County. In Mxico this species has been collected at the following Sierra Matasagnas, ca. 50 km W of Ojinaga, 9.6 km E of 1000 m alt. (UF 317157). COAHUILA: 5 KM NW of Quatro Cienegas, 800 m slt. (UF 267670); limestone hill 18.7 km E of Quatro Cienegas, 800 m alt. (UF 178584); 3 km N of Quatro Cienegas, 870 m alt. (UF 39968); NW foothills of the Sierra la Madera, 9.3 km SE, 14.7 km SW of Ocampo, 267692); Cerro Sesteadero 11 km S of San Miguel, 1000 m Rabdotus ( Rabdotus) schiedeanus schiedeanus (Pfeiffer 1841)Bulimus schiedeanus Pfeiffer 1841; Symbolae ad Histori Heliceorum 1:43. Bulimulus schiedeans (Pfeiffer). Von Martens 1893; Biol. Cent. Conch. 11:131.Hinkley 1907; Nautilus 21:72. Bulimulus (Rabdotus) schiedeanus (Pfeiffer). Pilsbry 1946; Land Branson & McCoy 1963; Nautilus 76:105. Bulimus patriarcha W. G. Binney 1858; Manual of North American Bulimulus alternatus hesperius Pilsbry & Ferriss 1924; Nautilus 38:40.Type Localities. Bulimus schiedeanus: Mxico. Bulimus patriarcha: Buena Vista, [south of Saltillo, Coahuila]. Bulimulus alternatus hesperius: east of the Pecos at the High Bridge, Val Verde Co., Texas; holotype in the ANSP. Distribution.Texas, counties along the Rio Grande from Val Verde County to Presidio County; widely distributed but sparse in Mxico from the state of Puebla north to Chihuahua (Branson & McCoy 1963). COAHUILA: around Saltillo (Pilsbry 1946). DURANGO: Villa Lerdo (Pilsbry 1946). Sierra de Tlahualilo, 25 mi. N of Tlahualilo (Solem 1954:4). JALISCO: Laguna de Chapala (Pilsbry 1946). PUEBLA: Tahuacan (Pislbry 1946). TAMAULIPAS: Tampico, in river drift (Hinkley 1907) Taxonomy.Two subspecies are recognized. Rabdotus schiedeanus pecosensis to southwest Texas. Rabdotus (Rabdotus) fonsecanus (Haas 1961)Bulimulus (Rabdotus) fonsecanus Haas 1961; Fieldiana, Zoology Type Locality.Gulf of Fonseca, El Salvador or

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128 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Nicaragua. Holotype: FMNH 106702. Distribution.Known only from the type locality. The Group of Rabdotus (Rabdotus) baileyi (Dall 1893)Bulimulus (Scutalus) baileyi Dall 1893; Proc. U. S. Nat. Mus.16:640 Bulimulus (Orthotomium) baileyi Dall. Pilsbry 1898; Man. Conch. Bulimulus baileyi Dall. Hanna 1923; Proc. Calf. Acad. Sci. (4), 12:487.Type Locality.Cabo San Lucas, Baja California Sur, Mxico. Holotype in the USNM. (Doubtful locality, Hanna 1923; Proc. Calf. Acad. Sci. (4), 12:487.) Distribution.SONORA: Guaymas; Hermosillo; Cerrro Tordilla, between Guaymas and San Marciel; Ortiz (Pilsbry 1898). Rabdotus (Rabdotus) ceralboensis (Hanna 1923)Bulimulus ceralboensis Hanna 1923; Proc. Calif. Acad. Sci. 12:490; Rabdotus (Rabdotus) ceralboensis (Hanna). Emerson & Jacobson 1964; Trans. San Diego Soc. Nat. Hist. 16:327.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:120; textType Locality.West of Ranch house, Cerralvo Island, Baja California Sur, Mxico. Holotype CAS 1021. Distribution.BAJA CALIFORNIA SUR: known only from Cerralvo Island (Smith et al. 1990). Rabdotus (Rabdotus) chamberlini (Hanna 1923)Bulimulus chamberlini Hanna 1923; Proc. Calif. Acad. Sci. 12:494; Rabdotus (Leptobrysus) chamberini (Hanna). Emerson & Jacobson (shell). Rabdotus (Rabdotus) chamberlini (Hanna 1923). Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:120; textType Locality.San Diego Island, Baja California Sur, Mxico. Holotype CAS 1023. Distribution.BAJA CALIFORNIA SUR: known only from Isla San Diego (Smith et al. 1990). Rabdotus (Rabdotus) (Dall 1893)Bulimulus xantusi var. Dall 1893; Proc. U. S. Nat. Mus. 16:642. Dall. Dall 1895; Proc. U. S. Nat. Mus. 18:5. Bulimulus (Orthotomium) Dall. Pilsbry 1897: Man. Conch. Rabdotus (Rabdotus) (Dall). Smith, Miller, Christensen & Roth Bulimulus (Thaumastus) digueti Mabille 1895.Type Localities. Bulimulus digueti: Sierra de La Victoria, Baja California Sur, Mxico. Distribution.BAJA CALIFORNIA SUR: various localities near southern tip of peninsula (Smith et al. 1990). Rabdotus (Rabdotus) pilula (Binney 1861)Bulimulus pilula W. G. Binney 1881; Proc. Acad. Nat. Sci. Phila. Bulimulus (Orthotomium) pilula Binney. Pilsbry 1897; Man. Conch. Rabdotus (Rabdotus) pilula (Binney 1861). Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:121; textBulimulus cooperi Dall 1896; Proc. U. S. Nat. Mus.18:5. Bulimulus (Orthotomium) decipiens Cooper 1895; Proc. Calif. Acad. Sci. (2) 5:164.Dall 1896; Proc. U. S. Nat. Mus.19:358.Type Localities. Bulimulus pilula : Todos Santos Mission, Baja California Sur (Binney & Bland 1869). Bulimulus decipiens : Sierra San Lazaro, Cabo San Lucas. Bulimulus cooperi : San Jos del Cabo. Distribution.BAJA CALIFORNIA SUR: along the southern tip of the peninsula from Cabo San Lucas to San Jos del Cabo (Smith et al. 1990). Rabdotus (Rabdotus) (Gould 1859) Gould 1859; Journal of the Boston Society of Bulimulus (Orthotomium) (Gould). Pilsbry 1897; Man. Rabdotus(Rabdotus) (Gould 1859). Smith, Miller & Gould 1853 (not Bulimus vesicalis Pfeiffer 1859. Bulimulus juarezi Pfeiffer 1866; Proc. Zool. Soc. London 34:832. var. insularis Cooper 1892; Proc. Calif. Acad. Sci. (2) 3:208, 212. Bulimulus insularis var. chinchesis Cooper 1894; Proc. Calif. Acad. Sci. (2) 4:140. Bulimulus (Globulus) recognitus Mabille 1895; Bulletin de la Socit Philomathique de Paris (8) 7:69.Emerson & Jacobson 1964; Trans. San Diego Soc. Nat. Hist. 16:51.Breure 1976:1148 (measurements of holotype).Type Localities. : not stated. Bulimulus juarezi: Mxico; syntype in the BMNH (Breure 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168):76). var. insularis: Espiritu Santo Island, Baja California Sur, Mxico. Bulimulus insularis chinchensis : El Chinche Mts., Baja California Sur. Bulimulus (Globulus) recognitus: Mxico, Baja California; lectotype in the MNHN (Breure 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168):76). Distribution.BAJA CALIFORNIA SUR: numerous localities on the south end of the peninsula (Smith et al. 1990). Subgenus Plicolumna Cooper 1895Plicolumna Cooper 1895; Proc. Calif. Acad. Nat. Sci. (2), 5:164.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:101, 122.Type Species.Rhodea californica ramentosa Cooper 1895. Distribution.Endemic to the Cape region of Baja California Sur. Taxonomy.Five species are recognized (Smith et al. 1990).

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 129 Rabdotus (Plicolumna ) (Cooper 1892)Columna ramentosa var. Cooper 1892; Proc. Calif. Acad. Sci. (2) 3:215. Bulimulus (Plicolumna ) (Cooper). Pilsbry 1898; Man. Rabdotus (Plicolumna ) (Cooper). Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:122; textType Locality.Sierra Laguna, Baja California Sur, Mxico. Distribution.BAJA CALIFORNIA SUR: known from three localities in the Cape region (Smith et al. 1990). Rabdotus (Plicolumna ) artemisia (Binney 1861)Bulimus artemisia W. G. Binney 1861; Proc. Acad. Nat. Sci. Bulimulus artemesia (Binney). Dall 1893; Proceedings of the U.S. Bulimulus (Plicolumna ) artemisia (Binney). Pilsbry 1898; Man. Rabdotus (Plicolumna ) artemisia (Binney 1861). Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:122; Type Locality.Cabo San Lucas, Baja California Sur, Mxico. Distribution.BAJA CALIFORNIA SUR: Cape region of the peninsula (Smith et al. 1990). Rabdotus (Plicolumna ) inscendens (Binney 1861)Bulimus inscendens W. G. Binney 1861; Proc. Acad. Nat. Sci. Bulimulus (Orthotomium) inscendens (Binney). Pilsbry 1898; Man. Rabdotus (Plicolumna ) inscendens (Binney). Emerson & Jacobson (reproductive anatomy).Smith, Miller, Christensen & Roth Type Locality.Not stated. Distribution.BAJA CALIFORNIA SUR: numerous localities in the Cape region (Smith et al. 1990). Rabdotus (Plicolumna ) perhirsutus Miller, Christensen, & Roth 1990Rabdotus (Plicolumna ) perhirsutus Miller, Christensen & Roth 1990, in Smith, Miller, Christensen & Roth. 1990:101, 25 (map).Type Locality.Along Todos Santos-Cabo San Lucas road 2.2 mi. (2.2 km) N of Saucito, Mxico; ca. 1500 ft. (460 m) alt. Holotype SBMNH 35110. Distribution.BAJA CALIFORNIA SUR: known only from the vicinity of the type locality (Smith et al. 1990). Rabdotus (Plicolumna ) ramentosus (Cooper 1991)Rhodea californica subsp. ? ramentosa Cooper 1891; Proc. Calif. Acad. Sci. (2) 3:102. Bulimulus (Plicolumna ) ramentosus (Cooper). Pilsbry 1898; Man. Rabdotus (Plicolumna ) ramentosus (Cooper). Christensen & Miller Type Locality.Mountains north of San Jos del Cabo, Baja California Sur, Mxico. Distribution.BAJA CALIFORNIA SUR: known from Genus Spartocentrum Dall 1895Spartocentrum Dall 1895; Nautilus 9:51.Pilsbry 1902; Man. Conch. 15:15.Christensen & Miller 1975; Nautilus 89: 44. Teneritia Mabille 1897; Bulletin de la Socit Philomathique de Paris (8) 9:78.Type Species.Spartocentrum: Cylindrella irregulare Gabb 1868. Teneritia : Berendtia digueti Mabille 1895. Distribution.Endemic to the mainland of Baja California Sur, Mxico and adjacent islands in the Gulf of California. Taxonomy.Five species are recognized. Spartocentrum digueti (Mabille 1895)Bulimulus (Thaumastus) digueti Mabille 1895; Bulletin de la Socit Philomathique de Paris (8) 7:70. Bulimulus (Orthotomium) digueti Mabille. Pilsbry 1898; Man. Conch. 11:148. Spartocentrum digueti (Mabille). Hanna & Smith 1968:396.Breure 1978; Uitg. Rijkmuseum Nat. Hist. Leiden (164):155; Proc. Calif. Acad. Sci. 47:125.Type Locality.Plateau de San Javier, [Baja California the MNHN (Hanna & Smith 1968; Breure 1978; Uitg. Rijkmuseum Nat. Hist. Leiden (164):155). Distribution.BAJA CALIFORNIA SUR: exact locality unknown (Smith et al. 1990). Spartocentrum eisenianum (Pilsbry 1900)Coelocentrum eisenianum Pilsbry 1900; Proc. Acad. Nat. Sci. Phila. Spartocentrum eisenianum (Pilsbry). Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:125. Coelocentrum eiseni Bartsch 1907:119.Type Locality.Lower California. Holotype ANSP 77888. Distribution.BAJA CALIFORNIA SUR: exact locality unknown (Smith et al. 1990). Spartocentrum insulare (Hanna 1923)Coelocentrum insulare Hanna 1923; Proc. Calif. Acad. Sci. 12:509 Spartocentrum insulare (Hanna). Smith, Miller, Christensen & Roth Coelocentrum oweni Trans. San Diego Soc. Nat. Hist. 16:328.Type Localities. Coelocentrum insulare: Puerto

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130 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Ballandra, Carmen Island, Baja California Sur, Mxico. Holotype CAS 1061. Coelocentrum oweni: Agua Grande, Isla Carmen, Baja California Sur, Mxico. Holotype CAS 1055. Distribution.BAJA CALIFORNIA SUR: Isla Carmen; Isla Montserrate; Isla Danzante; Isla Santa Catalina (Smith et al. 1990). Spartocentrum irregularis (Gabb 1868)Cylindrella (Urocoptis) irregularis (shell). Coelocentrum irregularis (Gabb). Pilsbry 1903; Man. Conch. 15:51; Spartocentrum irregulare (Gabb). Breure 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168):79.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:125; Coelocentrum minorinum gabbi Pilsbry 1900; Proc. Acad. Nat. 17 (shell). Berendtia minorina Mabille 1895; Bulletin de la Socit Philomathique de Paris (8) 7:70. Spartocentrum minorina (Mabille). Hanna & Smith 1968; Proc. Calif. Acad. Sci. (4) 30.Breure 1978; Uitg. Rijkmuseum Nat. Hanna 1923; Proc. Calf. Acad. Sci. (4) shell).Type Localities. Cylindrella irregularis: table lands in the interior of Lower California, especially above Moleje [Mulege]; lectotype ANSP 25075 (H. B. Baker 1963:223). Berendtia minorina: Au dessus de larroyo de Purisima, MNHN (Hanna & Smith 1968). Coelocentrum minorinum gabbi: table lands in the interior of Lower California, especially above Moleje [Mulege]; holotype ANSP 25077a (Baker 1963:222). : Agua Verde Bay, Baja California Sur, Mxico; holotype CAS 1082. Distribution.BAJA CALIFORNIA SUR: numerous localities in the central third of the peninsula (Smith et al. 1990). (Hanna 1923) Hanna 1923; Proc. Calf. Acad. Sci. (4) shell). (Hanna). Christensen & Miller 1975; (shell).Smith, Miller, Christensen & Roth 1990; Proc. Calif. Type Locality.Puerto Escondido, Baja California Sur, Mxico. Holotype CAS 1085. Distribution.BAJA CALIFORNIA SUR: Puerto Escondido; Juncalito; 1.6 km S of Chunque (Smith et al. 1990. Genus Simpulopsis Beck 1838Simpulopsis Beck 1838; Index Molluscorum.:100.Von Martens 1897; Biol. Cent. Amer.:252.Pilsbry 1899; Man. Conch. 12:212.Type Species.Helix (Cochlohydra) sulculosa Ferussac. Distribution.South America, Trinidad and eastern Mxico. Taxonomy.Three species occur in the study area. The uncertain. Simpulopsis aenea Pfeiffer 1861Simpulopsis aenea Pfeiffer 1861; Proc. Zool. Soc. Lond. 29:27.12a, 12b (shell).Von Martens 1897; 253. Simpulopsis (Platysuccinea ) aenea Pilsbry 1899; Man. Conch. Type Locality.Parada, Oaxaca, Mxico. Distribution.Known only from the type locality. Simpulopsis cumingi Pfeiffer 1861Simpulopsis cumingi Pfeiffer 1861; Proc. Zool. Soc. Lond. 29:27; Type Locality.Mxico. Distribution.MXICO (country or state?). Simpulopsis simula (Morelet 1851)Bulimus simula Morelet 1851; Test. Noviss.:11. Simpulopsis simula (Morelet). Fischer & Crosse 1877; Miss. Sci. Biol. Cent. Amer.:252.Pilsbry 1899; Man. Conch. 12:219 Breure 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. 14, 15.Type Locality.Petn, Guatemala. Distribution.HONDURAS, Dept. Olancho: vicinity of Magua Cave, ca. 15 km. km SSW of Gualaco, 940 m alt. (Breure 2008). GUATEMALA, Dept. Petn: only from the type locality. Dept. Alta Verapaz: 15 km by road north of Cobn, 1050 m alt.; 3 km W of Pajl, 1130 m alt.; 11 km W of Lanqun, 1000 m alt. (Breure 2008). CHIAPAS: Ocotal Guatemala), 700 m alt. (Bequaert 1957). Superfamily UROCOPTOIDEA Uit de Weerd 2008 Family EUCALODIIDAE Crosse & Fischer 1868 Distribution.Mxico, Guatemala, Honduras, and Belize. Taxonomy.Five genera are included in the Eucalodiidae. Four are restricted to Mxico and northern Central America: Eucalodium Anisospira, Coelocentrum and Dissotropis. Archegocoptis from Hispaniola is also a member of this group. Four other genera are associated provisionally with the Eucalodiidae: Epirobia, Propilsbrya, Pectinistemma and Prionoloplax. A recent study by Uit de Weerd (2008) necessitated the recognition of the Eucalodiidae as a family distinct from the Urocoptidae and the Holospiridae.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 131 Genus Eucalodium Crosse & Fischer 1868Eucalodium Crosse & Fischer 1868; Jour. de Conchyl. 16:88.Type Species.Pupa decollata Nyst 1841. Distribution.El Salvador and Guatemala north and west in Mxico to Jalisco along the west coast and San Lus Potos along the east coast. Taxonomy.Four subgenera, 24 species and 9 subspecies are recognized. Subgenus Eucalodium Crosse & Fischer 1868 Distribution.Tabasco, Chiapas, and Oaxaca, Mxico; Guatemala. the study area. Eucalodium (Eucalodium) compactum Pilsbry 1893Eucalodium compactum Pilsbry 1893. Proc. Acad. Nat. Sci. Type Locality.Tabasco, Mxico. Holotype ANSP 63388a (H. B. Baker 1963). Distribution.Known only from the type locality. Eucalodium ( Eucalodium) decollatum decollatum (Nyst 1841)Pupa decollata Nyst 1841; Bulletin de lAcadmie Royal de Bruxelles 8:345. Cylindrella decollata (Nyst). Pfeiffer 1856; Mon. helic. viv. VIII:421. Eucalodiun decollatum (Nyst). Fischer & Crosse 1872; Miss. Sci. Cylindrella gheisbreghti Pfeiffer 1856; Proc. Zool. Soc. Lond. Eucalodium ghiesbreghti (Pfeiffer). Crosse & Fischer 1868; Jour. de Conchyl. 14:88.Fischer & Crosse 1872; Miss. Sci. Mex. 111b (shell). Eucalodium decollatum var. gheisbreghti (Pfeiffer). Pilsbry 1902; Type Localities. Pupa decollata : State of Tabasco, Mxico. Cylindrella gheisbreghti: Chiapas, in SE Mxico. The type locality for both nominate taxa is here restricted to limestone hills 5 km E of Teapa, Tabasco, Mxico. Distribution.TABASCO: known for certain only from limestone hills immediately east of Teapa, Tabasco, Mxico. Records for Cobn and Vera Paz, Guatemala require Eucalodium (Eucalodium) decollatum guatemalensis Bartsch 1906Eucalodium decollatum guatemalensis Bartsch 1906. Bull. U. S. Type Locality.Guatemala. Holotype USNM 162307. Distribution.Known only from the type locality. Eucalodium (Eucalodium) mexicanum mexicanum (Pfeiffer 1860)Cylindrella mexicana Pfeiffer 1860. Proc. Zool. Soc. Lond. 28:139. Eucalodium mexicanum mexicanum (Pfeiffer). Fischer & Crosse 1873; Miss. Sci. Mex. I:369.Strebel 1880; Beitrag, Type Locality.Republic of Mxico. Distribution.OAXACA: Juquila, Oaxaca (Fischer & Crosse 1873). Earlier records list Tabasco and Chiapas. Taxonomy.Three subspecies are listed. Two were originally described from unlocalized sources, but they were later recorded from the same locality in Oaxaca. A third subspecies was described from Guatemala. It is unlikely that Eucalodium (Eucalodium) mexicanum major Fischer & Crosse 1873 Eucalodium mexicanum major Fischer & Crosse 1873; Miss. Sci. Mex. I:. Miss. Sci. Mex. I:369.Pilsbry 1902; Man. Conch. 15:8. Type Locality.Here restricted to woods between Tactc and Tamahu, Dept. Alta Verapaz, Guatemala. Distribution.GUATEMALA, Dept. Alta Verapaz: woods between Tactc and Tamahu. Eucalodium ( Eucalodium) mexicanum minor (Pfeiffer 1860)Cylindrella mexicana minor Pfeiffer 1860. Novitat. Conch. III:435; Eucalodium mexicanum minor (Pfeiffer). Fischer & Crosse 1873; 11 (shell).Type Locality.Not given. Distribution.OAXACA: Juquila (Fischer & Crosse 1873). Eucalodium (Eucalodium) otoides Thompson 1968Eucalodium otoides Thompson 1968; Bull. Fla. St. Mus. 12:180 Type Locality.Forested ravine 3.5 mi by road S of Distribution.CHIAPAS: known only from the Subgenus Oligostylus Pilsbry 1895Oligostylus Pilsbry 1895; Nautilus 9:51.Type Species.Eucalodium blandianum Crosse & Fischer 1868. Distribution.El Salvador, Guatemala, and the states of Chiapas, Oaxaca, Veracruz, Guerrero, Michoacn, and Jalisco in Mxico. Taxonomy.Thirteen species are recognized. Eucalodium (Oligostylus) australis Thompson 1963Eucalodium australis Thompson 1963; Proc. Biol. Soc. Wash.

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132 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Type Locality.Hacienda Monte Cristo, Metapan, Dept. Santa Ana, El Salvador. Holotype SMF 161266. Distribution.Known only from the type locality. Eucalodium ( Oligostylus) blandianum Crosse & Fischer 1868Eucalodium blandianum Crosse & Fischer 1868. Jour. de Conchyl. 16:276.Fischer & Crosse 1873; Miss. Sci. Mex. I:; Miss. Sci. Type Locality.Mountains near Orizaba, Veracruz, Mxico. Distribution.VERACRUZ: near Orizaba. Eucalodium (Oligostylus) decurtatum (H. Adams 1872)Cylindrella decurtata H. Adams 1872; Proc. Zool. Soc. Lond. 41:13; Eucalodium decurtatum (H. Adams). Fischer & Crosse 1873; Miss. Type Locality.Putla, Oaxaca, Mxico. Distribution.OAXACA: Putla. Eucalodium (Oligostylus) grande (Pfeiffer 1860)Cylindrella grandis Pfeiffer 1860; Proc. Zool. Soc. Lond. 28:139; Eucalodium grande (Pfeiffer). Fischer & Crosse 1873; Miss. Sci. Type Locality.Juquila, Oaxaca, Mxico. Distribution.Known only from the type locality. Eucalodium (Oligostylus) hegewischi (Bartsch 1947)Bulinus truncatus Pfeiffer 1841; Symb. Hist. Helc. I:43. (Not Bulimus truncatus Brugnoire 1792). Cylindrella truncata Von Martens 1865; Malak. Bltt. 12:13. Eucalodium truncatum (Pfeiffer). Fischer & Crosse 1873; Miss. Sci. Oligostylus hegewischi Bartsch 1947; Jour. Wash. Acad. Sci. 37:286.Bartsch 1948; Jour. Wash. Acad. Sci. 38:350.Type Locality.Angangueo, Michoacn, Mxico; type lost. Distribution.MICHOACN: known from the area near Angangueo, 9000 ft. alt. (Thompson 1968). Records of this species from Omilteme, Guerrero (Von Martens 1897:264) almost certainly pertain to Eucalodium mariae (Bartsch 1947). Eucalodium (Oligostylus) hippocastaneum Dall 1897Eucalodium hippocastaneum Dall 1897; Nautilus 11:61.Pilsbry 1902; Man. Conch. 15:15. Type Locality.San Sabastian, Jalisco, Mxico. Holotype USNM 186137. Distribution.Known only from the type locality. Eucalodium (Oligostylus) insigne Crosse & Fischer 1872Eucalodium insigne Crosse & Fischer 1872; Jour. de Conchyl. Ty pe Locality.Southern Mxico. Distribution.Unknown. Eucalodium (Oligostylus) mariae (Bartsch 1947)Oligostylus mariae Bartsch 1947; Jour. Wash. Acad. Sci. 37:285 Type Locality.Omiltemi [Omilteme], Guerrero, Mxico. Holotype USNM 543582. Distribution.GUERRERO: Omilteme. Eucalodium (Oligostylus) moussonianum Crosse & Fischer 1872Eucalodium moussonianum Crosse & Fischer 1872; Jour. de Conchyl. 20:225.Fischer & Crosse 1873; Miss. Sci. Mex. Type Locality.Not stated. Distribution.Unknown. Eucalodium ( Oligostylus) neglectum Crosse & Fischer 1872Eucalodium neglectum Crosse & Fischer 1872; Jour. de Conchyl. 20:302.Fischer & Crosse 1873; Miss. Sci. Mex. I:373; pl. 14, T ype Locality.Oaxaca, Mxico. Distribution.Known only from the type locality. Eucalodium (Oligostylus) splendidum (Pfeiffer 1860)Cylindrella splendida Pfeiffer 1860; Proc. Zool. Soc. Lond. 28:139; Eucalodium splendidum (Pfeiffer). Fischer & Crosse 1873; Miss. Type Locality.Zacatepec, Oaxaca, Mxico. Distribution.Known only from the type locality. Eucalodium ( Oligostylus) sumichrasti Crosse & Fischer 1878Eucalodium sumichrasti Crosse & Fischer 1878; Jour. de Conchyl. Type Locality.Chiapas, Mxico. Distribution.Known only from the type locality. Eucalodium (Oligostylus) walpoleanum Crosse & Fischer 1872Eucalodium walpoleanum Crosse & Fischer 1872; Jour. de Conchyl. 20:75.Fischer & Crosse 1873; Miss. Sci. Mex. I:377; pl. 14,

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 133 22.Bartsch 1906; Proc. U. S. Nat. Mus. 31:111. Type Locality.Woods of Palenque, Chiapas, Mxico. Distribution.CHIAPAS: Palenque. GUATEMALA, Dept. Alta Verapaz Subgenus Ptychocentrum Bartsch 1943Coelocentrum (Ptychocentrum ) Bartsch 1943; Nautilus 56:91; Type Species.Coelocentrum bourgeoisae Bartch 1943 [=Coelocentrum (Ptychocentrum ) marianum Bartsch 1943]. Distribution.Central Chiapas, Mxico. Taxonomy.This subgenus is monotypic. Eucalodium (Ptychocentrum ) marianum (Bartsch 1943)Coelocentrum (Ptychocentrum) bourgeoisae Bartsch 1943; Nautilus Coelocentrum bourgeoisae Pilsbry 1939). Coelocentrum (Ptychocentrum ) marianum Bartsch 1943; Nautilus 56:144.Solem 1957; Notulae Naturae (298):8.Type Locality.Tecpatn, Chiapas, Mxico. Holotype USNM 536900. Distribution.Known only from the type locality. Subgenus Resupinata Von Martens 1897Resupinata Von Martens 1897; Biol. Cent. Amer.:255.Type Species.Cylindrella speciosa Dunker 1844 (Pilsbry 1953:134). Distribution.Central Veracruz and adjacent northern Oaxaca and San Lus Potos, Mxico. Eucalodium (Resupinata) cereum Strebel 1880Eucalodium cereum Type Locality.San Antonio del Norte, 3 leagues east from Neolingo, on road to Misantla, Veracruz, Mxico. Distribution.Known only from the type locality. Eucalodium (Resupinata) densecostatum Strebel 1880Eucalodium densecostatum Type Locality.Orizaba, Veracruz, Mxico. Distribution.VERACRUZ: Orizaba; Misantla (Bartsch 1906:111). Eucalodium (Resupinata) deshayesianum Crosse & Fischer 1873Eucalodium deshayesianum Crosse & Fischer 1872; Jour. de Conchyl. 20:223.Fischer & Crosse 1873; Miss. Sci. Mex. Type Locality.Southern Mxico. Distribution.Unknown. Eucalodium (Resupinata) edwardsianum Crosse & Fischer 1872Eucalodium edwardsianum Crosse & Fischer 1872. Jour. de Conchyl. 20:224.Fischer & Crosse 1873; Miss. Sci. Mex. Type Locality.Cordova, Veracruz, Mxico. Eucalodium (Resupinata) ischnostele (Pilsbry 1909)Coelocentrum ischnostele Pilsbry 1909; Nautilus 22:139. Eucalodium (Resupinata ) ischnostele (Pilsbry). Pilsbry 1953; Proc. Type Locality.Mecos Falls, San Lus Potos, Mxico. Lectotype ANSP 99487A (H. B. Baker 1963:223). Distribution.SAN LUS POTOS: Mecos Falls; near Tomasopo. Eucalodium (Resupinata) speciosum speciosum (Dunker 1844)Cylindrella speciosum Dunker in Phillipi 1844; Abbild. Beschreib., I:86. Eucalodium speciosum speciosum (Dunker). Fischer & Crosse. Eucalodium boucardi var. minor Fischer & Crosse 1873; Miss. Sci. Type Locality.Unknown. de Misantla, Jalapa. Eucalodium ( Resupinata) speciosum boucardi (Pfeiffer 1856)Cylindrella boucardi Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:321; Eucalodium boucardi (Peiffer). Fischer & Crosse 1873; Miss. Sci. Eucalodium speciosum boucardi (Pfeiffer). Von Martens 1897; Biol. Cent. Amer.:262.Pilsbry 1902; Man. Conch. 15:21; pl. 2, Type Locality.Cordova, Veracruz, Mxico. Eucalodium (Resupinata) Von Martens 1897Eucalodium speciosum (Dunker). Fischer & Crosse 1873; Miss. Sci. Von Martens 1897. Biol. Cent. 101. Type Locality.Chiquihuitl, Veracruz, Mxico. Distribution.VERACRUZ: Chiquihuitl. Eucalodium (Resupinata) speciosum minimum Von Martens 1897Eucalodium speciosum Fischer & Crosse 1873; Miss. Sci. Mex. I:381. Eucalodium speciosum minimum Von Martens 1897; Biol. Cent. Amer.:262.Pilsbry 1902; Man. Conch. 15:22. Type Locality.Cordova, Veracruz, Mxico.

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134 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Eucalodium (Resupinata) speciosum strebeli Von Martens 1897Eucalodium boucardi form B, Strebel 1860; Beitrag, IV:71; pl. 5, Eucalodium speciosum strebeli Von Martens 1897. Biol. Cent. Type Locality.Environs of Jalapa, especially about the brooks on the road to Neolingo, and Chirimoya, Veracruz, Mxico. Distribution.VERACRUZ: environs of Jalapa. Genus Anisospira Strebel 1880Anisospira Strebel 1880; Beitrag. Mex. Landund Sssw.-Conch. IV:77.Pilsbry 1902; Man. Conch. 15:24.Thompson 1968; Bull. Fla. St. Mus. 12:142.Type Species.Cylindrella liebmanni Pfeiffer 1846. Distribution.Known from lowland regions of Colima, Michoacn, and Oaxaca, Mxico. Taxonomy.Anisospira includes two subgenera, seven species, and two subspecies. Subgenus Anisospira Strebel 1880 Distribution.Southern Oaxaca, Mxico. Taxonomy.Three species, one of which has two subspecies Anisospira (Anisospira) dalli dalli (Von Martens 1901)Anisospira strebeli Dall 1897; Proc. U. S. Nat. Mus. 19:353; pl. 33, ot Anisospira strebeli Pfeffer 1887). Eucalodium dalli Von Martens 1901; Biol. Cent. Amer. :633 (new name for Anisospira strebeli Dall 1897). Anisospira dalli (Von Martens). Pilsbry 1902; Man. Conch. 15:26 Anisospira dalli dalli (Von Martens). Thompson 1968; Bull. Fla. St. Type Locality.Huilotepc, Oaxaca, Mxico. Holotype USNM 107366. Distribution.OAXACA: limestone hills west and north of Tehuantepec (Thompson 1968). Anisospira (Anisospira) dalli stringens Thompson 1968Anisospira dalli stringens Thompson 1968; Bull. Fla. St. Mus. Type Locality.Hill about 0.4 mi. S of the Presa Benito Juarez on the Rio Tehuantepec, about 18 mi. NW of Tehuantepec, Oaxaca, Mxico. Holotype UF 19079. Distribution.Known only from the type locality. Anisospira (Anisospira) liebmanni (Pfeiffer 1846)Cylindrella liebmanni Pfeiffer 1846; Zeitschrift fr Malak. 1:59. Anisospira liebmani (Pfeiffer). Strebel 1880; Beitrag, IV: 79; pl. 5, Cylindrella trochaeformis Sowerby, in Reeve 1843; Conch. Icon., Cylindrella hyalina Pfeiffer in Eucalodium (Anisospira) orcutti Dall 1910; Nautilus 24:34. Anisospira orcutti (Dall). Solem 1957; Notulae Naturae (298):6, pl. 1. Liocentrum wilmotiType Localities. Cylindrella liebmanni : unknown. Cylindrella trochaeformis: unknown. Cylindrella hyaline : unknown. Eucalodium (Anisospira) orcutti : Rio Verde, Oaxaca, Mxico; holotype USNM 212319. Liocentrum wilmoti : along the Pan American Highway between Oaxaca and Tehuantepec, Oaxaca, Mxico; holotype USNM 589051. Distribution.Southeastern OAXACA. Anisospira (Anisospira) (Strebel 1880) Strebel 1880; Beitrag, IV:73; pl. 11, (Strebel). Bartsch 1947; Jour. Wash. Acad. Sci. 37:284.Type Locality.Mxico. Distribution.Unknown. Subgenus Trachycion Thompson 1968Trachycion Thompson 1968; Bull. Fla. St. Mus. 12:148.Type Species.Anisospira recticosta townsendi Pilsbry & Cocherell 1903. Distribution.Coastal areas of Colima, Michoacn, and Oaxaca, Mxico. Taxonomy.Four species, one of which has two subspecies. Anisospira (Trachycion) hadromylla Thompson 1968Anisospira hadromylla Thompson 1968; Bull. Fla. St. Mus. 12:150 Type Locality.A small range of limestone hills along Mxico, 200 ft. alt. Holotype UF 19055. Distribution.Known only from the type locality. Anisospira ( Trachycion) recticosta recticosta (Pfeiffer 1847)Cylindrella recticosta Pfeiffer, in Philippi 1847; Abbild. 2:48; pl. Eucalodium recticosta (Pfeiffer). Fischer & Crosse 1873; Miss. Sci. Anisospira recticosta (Pfeiffer). Pilsbry 1902; Man. Conch. 15:299.Anisospira (Trachycion ) recticosta recticosta (Pfeiffer). Thompson 1968; Bull. Fla. St. Mus. 12:147.Type Locality.Unknown. Distribution.Unknown. Anisospira (Trachycion) recticosta townsendi Pilsbry & Cockerell 1903Anisospira recticosta townsendi Pilsbry & Cockerell, in H. A. Pilsbry

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 135 Anisospira (Trachycion ) recticosta townsendi Pilsbry & Cockerell. Thompson 1968; Bull. Fla. St. Mus. 12:150. Type Locality.Cualata, Colima, Mxico. Holotype ANSP 84385. Distribution.Known only from the type locality. Anisospira (Trachycion) strebeli Pfeffer 1887Anisospira strebeli Pfeffer 1887. Verh. Ver. Naturw. Hamburg:21.3. Anisospira (Trachycion ) strebeli Pfeffer. Thomopson 1968; Bull. Fla. St. Mus. 12:150.Breure 1977; Zoolog. Med. 51:305; pl. Eucalodium strebeli (Pfeffer). Von Martens 1897; Biol. Cent. Type Locality.Cerro de Las Plumas, near Puerto Angel, Oaxaca, Mxico. Distribution.OAXACA: known only from the area near Puerto Angel and Pochutla. Anisospira (Trachycion) Breure 1977 Breure 1977. Zoolog. Med. 51:301; Type Locality.Km 151 on the road from Pochutla to Puerto Escondido (ca. 10 km E of Puerto Escondido), Oaxaca, Mxico. Distribution.Known only from the type locality. Genus Coelocentrum Crosse & Fischer 1870Coelocentrum Crosse & Fischer 1870; Jour. de Conchyl. 20:302.Type Species.Cylindrella turris Pfeiffer 1856. Distribution.From Tamaulipas and Guerrero, Mxico SE to Guatemala. Taxonomy.The genus includes six subgenera, 33 species and six subspecies. Subgenus Coelocentrum Crosse & Fischer 1870 Distribution.From Tamaulipas and Guerrero SE to Guatemala. Taxonomy.Nineteen species and six subspecies are recognized. Coelocentrum (Coelocentrum ) anconai Bartsch 1948Coelocentrum anconai Bartsch 1948. Jour. Wash. Acad. Sci. Type Locality.Woods of Ocote, Ocozocoautla, Chiapas, Mxico. Holotype USNM 589052. Distribution.Known only from the type locality. Coelocentrum (Coelocentrum ) anomalum Strebel 1880Coelocentrum anomalum 6. Type Locality.Cobn, Departamento Alta Verapaz, Guatemala. Distribution.Known only from the type locality. Coelocentrum (Coelocentrum ) arctispira arctispira (Pfeiffer 1860)Cylindrella arctispira Pfeiffer 1860; Proc. Zool. Soc. Lond. 28:139; Coelocentrum arctispira (Pfeiffer). Fischer & Crosse 1873; Miss. Coieocentrum arctispira arctispira (Pfeiffer). Pilsbry 1902; Man. Type Locality.Juquila, Oaxaca, Mxico. Distribution.OAXACA: Juquila. TABASCO: Istapa, (Fischer & Crosse 1873). Remarks.The status of the species is uncertain, and The identity of specimens from Istapa, Tabasco also needs Coelocentrum (Coelocentrum ) arctispira estefaniae Pilsbry 1902Coelocentrum arctispira estefaniae Pilsbry 1902; Man. Conch. Coelocentrum actispira (Pfeiffer). Strebel 1880; Beitrag, IV: pl. 14, Type Locality.Here restricted to Quilate, Veracruz, Mxico. Distribution.VERACRUZ: Agua Caliente; Arroyo del Banco (Strebel 1880). Coelocentrum (Coelocentrum ) cataclines Thompson 1968Coelocentrum cataclines Thompson 1968; Bull. Fla. St. Mus. Type Locality.Limestone knoll 15.8 mi. NW by road from Ocozocoautla, Chiapas, Mxico, 2700 ft. alt.; holotype UF 19031. Distribution.Known only from the type locality. Coelocentrum ( Coelocentrum) championi Von Martens 1897Coelocentrum championi Von Martens 1897; Biol. Cent. Amer.:269; 12. Type Locality.Cerro Zunl, [Dept. Altenango], Guatemala; syntypes in the BMNH. Distribution.Known only from the type locality. Remarks.The subgeneric status is uncertain. Pilsbry (1903) related Coelocentrum championi and C. clathratum to C. (Gymnocentrum) on the basis of spiral striae between the costate sculpture of the shell. This similarity spiral threads that occur in There is little other similarity between the two species and Coelocentrum ( Coelocentrum) clathratum Von Martens 1897Coelocentrum clathratum Von Martens 1897; Biol. Cent. Amer.:269:

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136 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Type Locality.Hacienda Buena Vista, in upper Chulhuitz, Costa Cuca, western Guatemala; 3500 ft. altitude. Syntypes in the BMNH. Distribution.Known only from the type locality. Remarks.As with Coelocentrum championi, the subgeneric status of this species is uncertain. Coelocentrum (Coelocentrum ) dispar Pilsbry 1902Coelocentrum dispar Pilsbry 1902; Man. Conch. 15:44; pl. 17, Type Locality.Guatemala. Holotype ANSP 25079. unknown. Coelocentrum (Coelocentrum ) (Morelet 1849) Morelet 1849; Test. Noviss. 1:10. (Morelet). Fischer & Crosse 1873; Miss. Sci. Mus. Zool. Univ. Mich. (413):4.Haas & Solem 1960; Nautilus Type Locality.Forests of Petn, Guatemala. Distribution.GUATEMALA: widely distributed in the Depts. Petn and Alta Verapaz, and adjacent BELIZE. Coelocentrum (Coelocentrum ) gigas Von Martens 1897Coelocentrum gigas Von Martens 1897. Biol. Cent. Amer.:267; pl. Type Locality.Livingston, Departamento Izabal, Guatemala. Synotypes in the BMNH. Distribution.Known only from the type locality. Coelocentrum (Coelocentrum ) huertai Bartsch 1947Coelocentrum huertai Bartsch 1947; Jour. Wash. Acad. Sci. 37:286 Type Locality.Omilteme, Guerrero, Mxico; holotype USNM 543585. Distribution.Known only from the type locality. Coelocentrum (Coelocentrum ) nelsoni Dall 1896Coelocentrum nelsoni 26a-e, 22g. Type Locality.Tuxtla Gutierrez, Chiapas, Mxico. Holotype USNM 107368. Distribution.CHIAPAS: known only from the vicinity of Tuxtla Gutierrez, Chiapas (Thompson 1968). Coelocentrum (Coelocentrum ) penion Thompson & Correa-Sandoval 1994Coelocentrum penion Thompson & Correa-Sandoval 1994; Bull. Type Locality.Sierra de Cucharas, 7 km by road Distribution.Known only from the type locality. Coelocentrum (Coelocentrum ) pfefferi Dall, 1896Coelocentrum pfefferi Dall 1896; Proc. U. S. Nat. Mus. 19:352; pl. Type Locality.Ocozocoautla, Chiapas, Mxico. Holotype USNM 107367. Distribution.CHIAPAS: known only from the vicinity of the type locality. Coelocentrum (Coelocentrum ) pittieri pittieri Bartsch 1906Coelocentrum pittieri pittieri Bartsch 1906; Proc. U. S. Nat. Mus. Type Locality.Cave of Sakalkunta, near Senahu, Departamento Alta Verapaz, Guatemala. Holotype USNM 185492. Distribution.Known only from the type locality. Coelocentrum (Coelocentrum ) pittieri guatemalensis Bartsch 1906Coelocentrum pittieri guatemalensis Bartsch 1906; Proc. U. S. Nat. Type Locality.Secanquim, Departamento Alta Verapaz, Guatemala. Holotype USNM 187469. Distribution.Known only from the type locality. Coelocentrum (Coelocentrum ) stenocion Thompson 1968Coelocentrum stenocion Thompson 1968; Bull. Fla. St. Mus. Type Locality.Limestone cliff 5.6 miles NNE El Ocotito, Guerrero, 2900 ft. alt. Holotype UF 19061. Distribution.Known only from the type locality. Coelocentrum (Coelocentrum ) tanydeira Thompson 1968Coelocentrum tanydeira Thompson 1968; Bull. Fla. St. Mus. Type Locality.Hillside 11.4 miles east of Jilitla [Xilitla], San Lus Potos, 1100 ft. alt. Holotype UF 19040. Distribution.Known only from the type locality. Coelocentrum (Coelocentrum ) tomacella tomacella (Morelet 1849)Cylindrella tomacella Morelet 1849; Test. Noviss. I:10. Coelocentrum tomacella (Morelet). Fischer & Crosse 1873; Miss. Coelocentrum tomacella tomacella (Morelet). Thompson 1968; Bull. Fla. St. Mus. 12:171. Cylindrella moreleti Deshayes 1851; in Ferrusac 1851; Historie Naturel II:227. Type Localities. Cylindrella tomacella : woods of Tabasco and in the ruins of Palenque, State of Chiapas, Mxico. Cylindrella moreleti : woods of Tabasco and in the ruins of Palenque, State of Chiapas. Distribution.Northern CHIAPAS and adjacent TABASCO (Thompson 1968:171).

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 137 Coelocentrum (Coelocentrum ) tomacella adelphion Thompson 1968Coelocentrum tomacella adelphion Thompson 1968; Bull. Fla. St. Type Locality.Ravine 5.4 mi. N of Jitotol, Chiapas, Mxico; 5400 ft. alt. Holotype UF 19052. Distribution.Known only from the type locality. Coelocentrum (Coelocentrum ) tomacella attenuatum (Pfeiffer 1856)Cylindrella attenuatum Pfeiffer 1856; Malak. Blatt., 3:258. Coelocentrum attenuatum (Pfeiffer). Fischer & Crosse 1873; Miss. Sci. Mex. I:104. Coelocentrum attenuatum attenuatum (Pfeiffer). Pilsbry 1902; Man. Type Locality.Chiapas, Mxico. Coelocentrum ( Coelocentrum) (Pfeiffer 1856) Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:380. (Pfeiffer). Fischer & Crosse 1873; Miss. Sci. Amer.:270 (Pfeiffer). Pilsbry 1902; Man. Conch. 1968; Bull. Fla. St. Mus. 12:171.Type Locality.Chiapas; here restricted to a limestone hill 2 km NE of Teapa, Tabasco, Mxico. Distribution.TABASCO: known from the immediate vicinity of Teapa (Thompson 1968). Coelocentrum (Coelocentrum ) tomacella rufescens Von Martens 1897 Von Martens 1897; Biol. Cent. Coelocentrum tomacella rufescens Von Martens. Pilsbry 1902; Type Locality.Mxico. Distribution.Unknown. Coelocentrum (Coelocentrum ) turris (Pfeiffer 1856)Cylindrella turris Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:380; pl. Coelocentrum turris (Pfeiffer). Fischer & Crosse 1873; Miss. Sci. Type Locality.Chiapas; here restricted to limestone hills 5 km E of Teapa, Tabasco, Mxico. Distribution.CHIAPAS: vicinity of Solusuchiapa. TABASCO: vicinity of Teapa (Thompson 1968). Coelocentum (Coelocentrum ) tyla Thompson 1968Coelocentum tyla Thompson 1968; Bull. Fla. St. Mus. 12:176; Type Locality.Ravine 8.6 mi. by road E of Chiapa de Corzo, Chiapas, Mxico; 3100 ft. alt. Holotype UF 19044. Distribution.Known only from the type locality. Subgenus Crossostephanus Dall 1908Crossostephanus Dall 1908; Proc. U. S. Nat. Mus. 35:177.Type Species.Coelocentrum palmeri Dall & Bartsch 1908. Distribution.Central Tamaulipas, Mxico. Taxonomy.Six species. Coelocentrum (Crossostephanus ) Thompson & Correa-Sandoval 1994 Thompson & Correa-Sandoval 1994; Bull. Type Locality.Sierra Tamalave, 7 km west-southwest of Adolfo Lopz Mateos, Tamaulipas, Mxico; 350 m alt. Holotype UF 159634. Distribution.TAMAULIPAS: known from the Sierra Tamalave in the vicinity of the type locality. Coelocentrum (Crossostephanus ) hinkleyi Pilsbry 1909Coelocentrum hinkleyi Pilsbry 1909; Nautilus 22:138.Thompson & Correa-Sandoval 1994; Bull. Fla. Mus. Nat. Hist. Type Locality.Highest mountain on south side of river at Mecos Falls, San Lus Potos, Mxico. Lectotype ANSP 9884a (H. B. Baker 1963:223). Distribution.SAN LUS POTOS: recorded only from the type locality and San Diegito. Coelocentrum (Crossostephanus ) palmeri Dall & Bartsch 1908Coelocentrum palmeri Dall & Bartsch 1908; in Dall 1908; Proc. 38. 39, 44, 45, 49. Type Locality.Tamaulipas, Mxico; restricted to the Sierra de Cucharas, west of Gomez Faras, Tamaulipas (Thompson & Correa-Sandoval 1994). Holotype USNM 198083. Distribution.TAMAULIPAS: known only from the Sierra de Cucharas, W of Gomez Faras. Coelocentrum (Crossostephanus ) paucinoda Thompson & Correa-Sandoval 1994Coelocentrum paucinoda Thompson & Correa-Sandoval 1994; Type Locality.Limestone escarpment 3 km east of and Distribution.TAMAULIPAS: known only from the vicinity of the type locality. Coelocentrum (Crossostephanus ) priosculpta Thompson & Correa-Sandoval 1994Coelocentrum priosculpta Thompson & Correa-Sandoval 1994.

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138 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Type Locality.Sierra Grande, 2 km west of Santa Maria de Guadalupe, 21 km west of Ocampo, Tamaulipa, Mxico s; 1100 m alt. Holotype UF 159672. Distribution.TAMAULIPAS: known from the type locality and in immediately adjacent SAN LUS POTOS Coelocentrum (Crossostephanus ) torosum Thompson & Correa-Sandoval 1994Coelocentrum torosum Thompson & Correa 1994; Bull. Fla. Mus. Type Locality.Sierra de Tamaulipas, 7 km northwest Tamaulipas from 50 m alt. Subgenus Elasmocentrum Pilsbry 1902Elasmocentrum Pilsbry 1902; Man. Conch. 15:49.Type Species.Coelocentrum exlex Pilsbry 1902. Distribution.Unknown; attributed to Mxico. Taxonomy.This subgenus is monotypic. Coelocentrum (Elasmocentrum) exlex Pilsbry 1902Coelocentrum exlex 20. Type Locality.Mxico. Holotype ANSP 3938. Distribution.Unknown. Subgenus Gymnocentrum Pilsbry 1942.Liocentrum Pilsbry 1902; Man. Conch. 15:46 (non Liocentrum Karsch 1890). Gymnocentrum Pilsbry 1942; Nautilus 55:105.Type Species. Shuttleworth 1852. Distribution.Central Veracruz, Mxico Taxonomy.This subgenus contains two species. Coelocentrum ( Gymnocentrum) crosseanum (Pfeiffer 1867)Cylindrella crosseana Pfeiffer 1867; Jour. de Conchyl. 15:437. Eucalodium crosseanum (Pfeiffer). Crosse & Fischer 1870; Jour. de Conchyl. 18:22. Coelocentrum crosseanum (Pfeiffer). Fischer & Crosse. 1873; Miss. 25.Type Locality.Orizaba, Veracruz, Mxico. Distribution.VERACRUZ: known from the type locality and Cuautlatitlan (Von Martens 1897:272). Coelocentrum ( Gymnocentrum) (Shuttleworth 1852)Cylindrella Shuttleworth 1852; Mittheil. der Natur. Ges. Bern:296. (Shuttleworth). Crosse & Fischer 1870; Jour. de Conchyl. 18:22. (Shuttleworth). Fischer & Crosse 1873; Type Locality.Cordova, Veracruz, Mxico. Syntypes Naturhistorisches Museum Bern 18863/2 (Neubert & Gosteli 2003). Subgenus Ptychodonta Bartsch 1906Ptychodonta Bartsch 1906; Proc. U. S. Nat. Mus. 31:118.Type Species.Coelocentrum astrophorea Dall 1897. Distribution.Central Hidalgo and eastern Quertaro, Mxico. Taxonomy.Four species are recognized. Coelocentrum (Ptychodonta ) astrophorea Dall 1897Coelocentrum astrophorea Dall 1897; Nautilus 11:62.Pilsbry 1902; Man. Conch. 15:45.Bartsch 1906; Proc. U. S. Nat. Mus. Fla. Mus. Nat. Hist. 36:148. Coelocentrum acanthophorea Von Martens 1901; Biol. Cent. Amer.:634 (lapsus for Coelocentrum astrophorea Dall). Holotype USNM 134696. Distribution.Known only from the type locality. Coelocentrum (Ptychodonta ) brachyacron Thompson & Correa-Sandoval 1994Coelocentrum brachyacron Thompson & Correa-Sandoval 1994; Type Locality. km southeast of Tres Lagunas, Quertaro, Mxico; 2600 m alt. Holotype UF 159627. Distribution.QUERTARO: known only from the vicinity of the type locality. Coelocentrum (Ptychodonta ) endolophus Pilsbry 1953Coelocentrum endolophus Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. 1994; Bull. Fla. Mus. Nat. Hist. 36:152. Type Locality.Unknown; incorrectly given as between Chilpancingo and a small town nearby called Mazatln, Guerrero, Mxico (see Thompson & CorreaSandoval 1994:152). Holotype ANSP 190964. Distribution.HIDALGO: about 10 km SSW of Jacala. Coelocentrum (Ptychodonta ) telescopium Thompson & Correa-Sandoval 1994Coelocentrum telescopium Thompson & Correa-Sandoval 1994; Type Locality. km southwest of El Lobo, Quertaro, Mxico; 2100 m alt. Holotype UF 34493. Distribution.Known only from the type locality. Subgenus Schizopile Pilsbry 1939Schizopile Pilsbry 1939; Nautilus 53:27.Type Species.Coelocentrum bourgeoisae Pilsbry 1939. Distribution.Northwest Guerrero, Mxico.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 139 Taxonomy.This subgenus is monotypic. Coelocentrum (Schizopile) bourgeoisae Pilsbry 1939Coelocentrum bourgeoisae 5.Type Locality.Zihuatanejo, Guerrero, Mxico. Lectotype ANSP 174058a (H. B. Baker 1963:221). Distribution.Known only from the type locality. Nomen dubium associated with Coelocentrum Cylindrella ludersi Pfeiffer 1859.Monographia Helicorum Viventium, IV:721.Remarks.Pfeiffer related this species to other Coelocentrum known at the time. Pilsbry (1903, 15:46) doubted that C. ludersi is a species of Coelocentrum Genus Dissotropis Bartsch 1906Dissotropis Bartsch 1906; Bull. U. S. Nat. Mus. 113.Type Species. Anisospira (Dissotropis) stearnsi Bartsch 1906. Distribution.Southwestern Mxico in the states of Jalisco, Colima, Michoacn, and Guerrero. Two species are reported from Mazatln, Sinaloa, but these records need Taxonomy.Five species are recognized. Dissotropis amplaxis Thompson 1968Dissotropis amplaxis Thompson 1968; Bull. Fla. St. Mus. 12:161 Type Locality.A xeric hillside along the NW side of the highway from Colima to Tecoman, 1.9 mi. SW of Tecolapa, Colima, Mxico. Holotype UF 19764. Distribution.Known only from the type locality. Dissotropis blandi (Bartsch 1906)Anisospira (Dissotropis) blandi Bartsch 1906; Proc. U. S. Nat. (298):7. Dissotropis blandi (Bartsch). Thompson 1968; Bull. Fla. St. Mus. 12:154.Type Locality.Near Mazatln, Sinaloa, Mxico. This surely is an error. Here-in restricted to near Mazatln, Guerrero, Mxico. Holotype USNM 58055. Distribution.GUERRERO: known only from near Mazatln. Dissotropis castaneum Thompson 1968Dissotropis castaneum Thompson 1968; Bull. Fla. St. Mus. 12:155 Type Locality.Limesone sinkhole 6.0 mi. SW and 6.5 mi. E of Pihaumo, Jalisco, Mxico. Holotype UF 19758. Distribution.Known only from the type locality. Dissotropis henryi (Solem 1957)Anisospira (Dissotropis) henryi Solem 1957; Notulae Naturae Dissotropis henryi (Solem). Thompson 1968; Bull. Fla. St. Mus. 12:154.Type Locality.La Placita (= Sulatillo), Sierra de Coalcomn, Michoacn, Mxico. Holotype UMMZ 185499. Distribution.Known only from the type locality. Dissotropis stearnsi (Bartsch 1906)Anisospira (Dissotropis) stearnsi Bartsch 1906; Proc. U. S. Nat. (298):7. Dissotropis stearnsi (Bartsch). Thompson 1968; Bull. Fla. St. Mus. 12:154.Type Locality.Near Mazatln, Sinaloa, Mxico. This surely is an error. Here-in restricted to near Mazatln, Guerrero, Mxico. Holotype USNM 59055b. Distribution.Known only from the type locality. Genus Epirobia Strebel 1880Epirobia Strebel 1880; Beitrag, IV: 77, 85.Von Martens 1897; Biol. Cent.-Amer.:283.Pilsbry & Vanatta 1898; Proc. Acad. Nat. Sci. Phila.:281.Pilsbry 1902; Man. Conch. 15:59. Type Species.Cylindrella polygyra Pfeiffer 1856 (Pilsbry & Vanatta 1898). Distribution.Northern Guatemala; northern Chiapas, Taxonomy.Two subgenera, eight species and two subspecies are recognized. Subgenus Epirobia Strebel & Pfeffer 1880 Distribution.Northern Guatemala; northern Chiapas and central Veracruz, Mxico. Taxonomy.Seven species and two subspecies are recognized. Epirobia apiostoma (Pfeiffer 1856)Cylindrella apiostoma Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:322.Holospira (Epirobia) apiostoma (Pfeiffer). Von Martens 1897; Biol. Cent. Amer:283. Epirobia apiostoma (Pfeiffer). Strebel 1880; Beitrag, IV: 88; pl. 5, Type Locality.Cordova, Veracruz, Mxico. cave at Cacahuatla. Epirobia berendti berendti (Pfeiffer 1866)Cylindrella berendti Pfeiffer 1866; Malak. Bltt. 8:87.Fischer & Crosse 1873; Miss. Sci. Mex. I:409. Epirobia berendti Holospira berendti (Pfeiffer). Von Martens 1897; Biol. Cent. Amer.: 281.Type Locality.Toxpan [Tuxpan], on the slope to Cerro Matlaguiahuitl, near Cordova, Veracruz, Mxico. Distribution.Known only from the type locality.

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140 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Epirobia berendti albida (Fischer & Crosse 1873)Cylindrella berendtl albida Fischer & Crosse 1873; Miss. Sci. Mex. I:409. Holospira berendti var. albida (Fischer & Crosse). Von Martens 1897; Biol. Cent. Amer.:281. Epirobia berendti albida (Fischer & Crosse). Strebel 1880; Beitrag, Type Locality.Chiapas, Mxico. Distribution.CHIAPAS: unknown. Epirobia gassiesi (Pfeiffer 1867)Cylindrella gassiesi Pfeiffer 1867; Jour. de Conchyl. 15:438.Holospira (Epirobia) gassiesi (Pfeiffer). Von Martens 1897; Biol. Cent. Amer.:283. Epirobia gassiesi (Pfeiffer). Pilsbry 1902; Man. Conch. 15: 63; pl. Type Locality.Chiapas, Mxico. locality. Epirobia lurida Thompson 1976Epirobia lurida (shell), 3 (anatomy). Type Locality.Low rolling limestone hills 15.8 mi. [25.5 km] NW of Ocozocoautla on road to Mal Paso, Chiapas, Mxico; 2700 ft. alt. [830 m]. Holotype UF 22449. Distribution.Known only from the type locality. Epirobia polygyra (Pfeiffer 1856)Cylindrella polygyra Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:322; Holospira (Epirobia) polygyra (Pfeiffer). Von Martens 1897; Biol. Cent. Amer.:284. Epirobia polygyra pl. 13.Pilsbry & Vanatta 1898; Proc. Acad. Nat. Sci. Phila. 31:120.Type Locality.Cordova, Veracruz, Mxico. Distribution.Known only from the type locality. Epirobia polygyrella (Von Martens 1863)Cylindrella polygyrella Von Martens 1863; Proc. Zool. Soc. Lond. 31:411.Von Martens 1876; Jahrb. Deut. Malak. Ges. 3:261, Epirobia morini (Von Martens). Strebel 1880; Beitrag, IV:87, pl. 5, Nat. Mus. 31:120. Holospira (Epirobia) polygyrella (Von Martens). Von Martens 1897; Type Locality.Cobn, Dept. Alta Verapz, Guatemala. Distribution.Known only from the type locality. Epirobia swiftiana swiftiana (Crosse 1863)Cylindrella swiftiana Crosse 1863; Jour. de Conchyl. 11:388.Holospira (Epirobia) swiftiana (Crosse). Von Martens 1897; Biol. Cent. Amer.:284. Epirobia swiftiana swiftiana (Crosse). Pilsbry 1902; Man. Conch. Type Locality.Unknown. Distribution.Unknown. Epirobia swiftiana alternans Thompson 1976Epirobia swiftiana alternans Thompson 1976; Nautilus 90:41; Type Locality.Limestone ledge along a ravine 4.5 mi. [10.2 km] N of Jitotol, Chiapas, Mxico; 5400 ft. alt. [1662 m]. Holotype UF 22451. Distribution.Known only from the type locality. Subgenus Gyrocion Pilsbry 1904Gyrocion Pilsbry 1904a; Proc. Acad. Nat. Sci. Phila. 55:765. T ype Species. Epirobia (Gyrocion) mirabilis Pilsbry 1902. Taxonomy.Taxonomic status uncertain, see remarks below. Epirobia (Gyrocion) mirabilis Pilsbry 1902Epirobia mirabilis Pilsbry 1904a; Proc. Acad. Nat. Sci. Phila. Mxico. Holotype ANSP 85914. Distribution.Known only from the type locality. Remarks.The holotype and paratype, which are the only known specimens, are immature and consist only of the upper whorls of the spire. They are similar in all respects to juvenile Coelocentrum and not Epirobia (Thompson 1968). Genus Propilsbrya Bartsch 1906Propilsbrya Bartsch 1906; Proc. U. S. Nat. Mus. 31:121.Type Species.Epirobia (Propilsbrya) nelsoni Bartsch 1906. Distribution.Higher altitudes of mountains in southern Taxonomy.Two species are recognized. Propilsbrya nelsoni (Bartsch 1906)Epirobia (Propilsbrya) nelsoni Bartsch 1906; Bull. U. S. Nat. Mus. Propilsbrya nelsoni (Bartsch). Rehder 1940; Jour. Wash. Acad. Sci. 30:316.Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. 105:136; Type Locality.Sierra Guadalupe, Coahuila, Mxico. Holotype USNM 187504. Distribution.COAHUILA: known only from the Sierra Guadalupe. Pilsbry (1953) recorded the species from 13 km back of the Hacienda Guadalupe, 7500 ft. alt. Propilsbrya potosiana (Pilsbry 1953)Propilsbrya (Stalactella ) potosiana Pilsbry 1953; Proc. Acad. Nat.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 141 Salazar 2005; Acta Zool. Mex. (21):61.Type Locality.Cerro Potos, near Galeana, Nuevo Holotype ANSP 164586. Distribution.Known only from the type locality. Genus Pectinistemma Rehder 1940.Pectinistemma Rehder 1940; Jour. Wash. Acad. Sci. 30:315.Type Species.Propilsbrya koesteri Rehder 1940. Distribution.High altitudes of mountains in southern Taxonomy.Two species are recognized. Pectinistemma infernilla Pilsbry 1953Propilsbrya (Pectinistemma ) infernilla Pilsbry 1953; Proc. Acad. 164148. Distribution.NUEVO LEN: known only from the vicinity of the type locality. Pectinistemma koestneri Rehder 1940Propilsbrya (Pectinistemma ) koestneri Rehder 1940; Jour. Wash. Salazar 2005; Acta Zool. Mex. (21):61. Mxico. Holotype USNM 535762. Distribution.NUEVO LEN: known only from Cerro Potos. Family HOLOSPIRIDAE Pilsbry 1946 Distribution.Mxico from Oaxaca north to Texas, New Mexico, and Arizona. Taxonomy.Six genera are recognized. Anatomical information and rRNA sequence data necessitated the recognition of Holospiridae as a distinct family (Uit de Weerd 2008). Genus Haplocion Pilsbry 1902Haplocion Pilsbry 1902; Man. Conch. 15:89. Liostemma Bartsch 1906; Proc. U. S. Nat. Mus. 19:144. Type Species.Haplocion: Holospira pasonis Dall 1895. Liostemma: Holospira hamiltoni Dall 1897. Distribution.The typical group of species is found in Chihuahua, Coahuila, and adjacent Texas. Haplocion plumbea (Roth & Megaw 1989) is known from the early Tertiary of Chihuahua. A small group of species from Guerrero and the State of Mxico appear not to be closely related to the typical group. Taxonomy.Fourteen species are recognized, 11 in Mxico. Haplocion campoi Bartsch 1943Haplocion campoi Bartsch 1943; Jour. Wash. Acad. Sci. 33:56; Type Locality.Las Grutas, Cacahuamilpa, Guerrero, Mxico. Holotype USNM 536880. Distribution.Known only from the type locality. Haplocion coahuilensis (W. G. Binney 1865)Cylindrella coahuilensis W. G. Binney 1865; Amer. Jour. Conch.., Gongylostoma coahuilensis (Binney). Tryon 1867; Amer. Jour. Holospira coahuilensis (Binney). Fischer & Crosse 1873; Miss. Sci. Mex. I:334.Pilsbry 1902; Man. Conch. 15:91; pl. 23, Bartsch 1943; Jour. Wash. Acad. Sci. 33:56.Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. 105:151. Metastoma coahuilensis (Binney). Dall 1897; Proc. U. S. Nat. Mus. 19:351.Type Locality.Cienega Grande, Coahuila, Mxico. Holotype USNM 9150. Distribution.Known only from the type locality. Haplocion durangoensis Bartsch 1906Haplocion (Liostemma) durangoensis Bartsch 1906:142; pl. 3, Type Locality.Durango, Durango, Mxico. Holotype USNM 187981. Distribution.DURANGO: Durango; Sierra de Tlahualilo, 25 miles N of Tlahualilo, Durango. Haplocion greggi (Drake 1951)Coelostemma greggi Drake 1951; Revista Soc. Malac., 8:40; pl. Type Locality.Foothills of the Sierra de Almoloya, near Cuevo Diablo, near Salaices, Chihuahua, Mxico. Holotype USNM 601628. Distribution.Known only from the type locality. Haplocion hamiltoni (Dall 1897)Holospira (Haplostemma) hamiltoni Dall 1897; Nautilus 11:38.Holospira (Liostemma) hamiltoni Dall. Bartsch 1906; Proc. U. S. Nat. Mus. 31:141. Holospira (Haplocion) hamiltoni Dall. Pilsbry 1946; Land Moll. N. Type Locality.Rio Grande Mountains, Brewster Co., Texas; 3500 feet alt. Holotype USNM 107759. Distribution.Known from Brewster Co., Texas and adjacent Chihuahua. CHIHUAHUA: Rio Conchas, just Haplocion mariae Bartsch 1942Haplocion mariae Bartsch 1942; Jour. Wash. Acad. Sci. 32:187 Type Locality.Ixtapan de La Sal, Mxico. Holotype USNM 536037. Distribution.Known only from the type locality.

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142 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Haplocion mathewsoni Bartsch 1942Haplocion mathewsoni Bartsch 1942. Jour. Wash. Acad. Sci. 32:188; Type Locality.State of Mxico. Holotype USNM 526036. Distribution.MXICO: exact locality unknown. Haplocion pasonis (Dall 1896)Holospira pasonis Dall 1895; Nautilus 8:112. Holospira (Metastoma) pasonis Dall. Dall 1896; Proc. U. S. Nat. Holospira (Haplocion) pasonis Dall. Bartsch 1906; Proc. U. S. 5.Bartsch 1943; Jour. Wash. Acad. Sci. 33:56.Pilsbry 1946; Type Locality.Mule Canyon, El Paso Co., Texas; 4000 feet alt. Holotype USNM 129032. Distribution.TEXAS: Red Mule Canyon, El Paso; Davis Mountains. CHIHUAHUA: Rio Conchos, not far Haplocion semisculpta (Stearns 1890)Holospira semisculpta Stearns 1890; Proc. U. S. Nat. Mus. 13:208; Holospira (Haplocion) semisculpta Stearns. Pilsbry 1902; Man. Mus. 19:144. Haplocion semisculpta (Stearns). Bartsch 1943; Jour. Wash. Acad. Sci. 33:56.Type Locality.Limestone cliffs in a canyon above San Carlos, Chihuahua, Mxico. Holotype USNM 102310. Distribution.Known only from the type locality. Haplocion wilmoti Bartsch 1947Haplocion wilmoti 3. Type Locality.Mountains near Chihuahua, Chihuahua, Mxico. Holotype USNM 543595. Distribution.Known only from the type locality. Haplocion yucatanenisis (Bartsch 1906).Holospira (Liostemma) yucatanensis Bartsch 1906; Proc. U. S. Nat. Holospira (Haplocion) yucatanensis Bartsch. Pilsbry 1952; Naurtilus, 66:69.Pilsbry 1953: Proc. Acad. Nat. Sci. Phila. 105:151. Haplocion yucatanensis (Bartsch). Drake 1952; Rev. Soc. Malac. Type Locality.Incorrectly given as Mujeres Island [Isla Mujeres], Yucatn, Mxico. Holotype USNM 187980. Distribution.TEXAS: Brewster Co., Hot Springs International Park (Pilsbry 1952). COAHUILA: Boquillas (Drake 1952). Genus Coelostemma Dall 1895Coelostemma Dall 1895; Proc. U. S. Nat. Mus. 19:347. Megaxis Pilsbry 1946; Land Mollusca of North America 2:123. Type Species.Coelostemma: Holospira elizabethae Pilsbry 1889. Megaxis: Holospira fusca Von Martens 1897. Distribution.MXICO: widespread from Veracruz, Puebla, and Oaxaca north to Colima, San Luis Potosi, Coahuila, Durango, Chihuahua and New Mexico. four species and two subspecies. Subgenus Coelostemma Dall 1895 Distribution.Oaxaca north to Colima and Coahuila. are recognized. Coelostemma (Coelostemma ) adana (Bartsch 1926)Holospira (Coelostemma) adana Bartsch 1926; Proc. U. S. Nat. Coelostemma adana (Bartsch). Bartsch 1943; Jour. Wash. Acad. Sci. 33:58.Type Locality.River drift from along the Rio Balsas, Rio Balsas Station, Guerrero, Mxico. Holotype USNM 363133. Distribution.Unknown. Coelostemma (Coelostemma ) adria (Bartsch 1926)Holospira (Coelostemma) adria Bartsch 1926; Proc. U. S. Nat. Mus. Coelostemma adria (Bartsch). Bartsch 1943; Jour. Wash Acad. Sci. 33:58.Type Locality.River drift from along the Rio Balsas, Rio Balsas Station, Guerrero, Mxico. Holotype USNM 363131. Distribution.Unknown. Coelostemma (Coelostemma ) anaclasta Thompson 1971Coelostemma anaclasta Thompson 1971; Bull. Fla. Mus. Nat. Hist. 15:280. Type Locality.A small limestone ledge at the head of a small ravine about 300 yards west of the Colima-Manzanillo highway, 1.9 mi. NE of Tecolapa, Colima, Mxico; 700 ft. alt. Holotype UF 20927. Distribution.COLIMA: known from the area south and east of Cd. Colima. Coelostemma (Coelostemma ) anconai Bartsch 1951Coelostemma anconai Bartsch 1951; Jour. Wash. Acad. Sci. 41:146, Type Locality.Ixcatiopn, Guerrero, Mxico. Holotype USNM 595019. Distribution.Known only from the type locality. Coelostemma (Coelostemma ) balesi Pilsbry 1954Coelostemma balesi Type Locality.Km post 175, near Chilpancingo, on road to Acapulco, Guerrero, Mxico. Holotype ANSP 191110. Distribution.Known only from the type locality.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 143 Coelostemma (Coelostemma ) balsasensis (Bartsch 1926)Holospira (Coelostemma) balsasensis Bartsch 1926; Proc. U. S. Coelostemma balsasensis (Bartsch). Bartsch 1943; Jour. Wash. Acad. Sci. 33:58.Type Locality.Balsas Station, Guerrero, Mxico. Holotype USNM 363129. Distribution.GUERRERO. Uncertain. Bartsch (1926) Coelostemma and Holospira that were collected by C. R. Orcutt from river drift along the Rio Balsas at Balsas Station, Guerrero. It is not clear whether the type specimens of C. balsasensis also were collected from river drift or from a natural habitat at Balsas Station. Coelostemma ( Coelostemma) bartschi Pilsbry & Clapp 1909Holospira bartschi Pilsbry and Clapp 1909; Nautilus 22:11415; Haplocion bartschi (Pilsbry & Clapp). Bartsch 1943; Jour. Wash. Acad. Sci. 33:56.Type Locality.In crevices in the limestone rock 1000 ft. above the Rio Balsas, Balsas, Guerrero, Mxico. Lectotype ANSP 117441 (H. B. Baker 1963:221). Distribution.Known only from the type locality. Coelostemma (Coelostemma ) bembix Thompson 1971Coelostemma bembix Thompson 1971; Bull. Fla. Mus. Nat. Hist. Type Locality.Limestone ledge above an old road cut along Federal Highway 95 where it crosses a tributary to the Rio Balsas, 12.6 mi. N of Zumpango del Rio, Guerrero, Mxico; 2300 ft. alt. Holotype UF 20901. Distribution.Known only from the type locality. Coelostemma (Coelostemma ) bourgeoisana bourgeoisana Bartsch 1942Coelostemma bourgeoisana Bartsch 1942; Jour. Wash. Acad. Sci.s, Coelostemma bourgeoisana bourgeoisana Bartsch 1942. Thompson 1971; Bull. Fla. Mus. Nat. Hist. 15:271. Type Locality.Ixtapn de la Sal, Mxico. Holotype USNM 536885. Distribution.MXICO: common on limestone terrain about the type locality; 1875 m alt. Coelostemma (Coelostemma ) bourgeoisana antricola Bartsch 1943Coelostemma antricola Bartsch 1943; Jour. Wash. Acad. Sci. 33:58; Coelostemma bourgeoisana antricola Bartsch. Thompson 1971; Bull. Fla. Mus. Nat. Hist. 15:272.Type Locality.Ravine near Las Grutas, Cacahuamilpa, Guerrero, Mxico. Holotype USNM 536885. Distribution.GUERRERO: near Taxco, and southern MXICO; 1275 m alt. Coelostemma ( Coelostemma) chilpancingoensis Pilsbry 1953Coelostemma chilpancingoensis Pilsbry 1953; Proc. Acad. Nat. Sci. Type Locality.Between Chilpancingo and a small town nearby called Mazatln, Guerrero, Mxico. Holotype ANSP 190967. Distribution.Known only from the type locality. Coelostemma (Coelostemma ) dalli (Pilsbry 1902)Holospira (Coelostemma) dalli Pilsbry 1902; Man. Conch. 15:100 31:149. Coelostemma dalli (Pilsbry). Bartsch 1943; Jour. Wash. Acad. Sci. 33:58.Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. 105:157; pl. Holospira strebeliana Pilsbry 1902; Man Conch. 15:101; pl. Type Localities. Holospira dalli : Sierra de Guadalupe, Coahuila, Mxico. Holotype ANSP 58095. Holospira strebeliana : Sierra de Guadalupe, Coahuila, Mxico. Lectotype ANSP 58096a (Baker 1963:225). Distribution.COAHUILA: Sierra de Guadalupe; 2275 m alt. Coelostemma (Coelostemma ) eclipes Thompson 1971Coelostemma eclipes Thompson 1971; Bull. Fla. Mus. Nat. Hist. Type Locality.Limestone ledge 16.4 mi. N of Zumpango del Rio, Guerrero, Mxico; 2300 ft. alt. Holotype UF 21210. Distribution.Known only from the type locality. Coelostemma (Coelostemma ) elizabethae (Pilsbry 1898)Holospira (Coelostemma) elizabethae Pilsbry 1889; Proc. Acad. Coelostemma elizabethae (Pilsbry). Bartsch 1943; Jour. Wash. Acad. 52. Von Martens 1897; Biol. Cent. Amer.:277; pl. Type Localities.Holospira elizabethae : Amula, Guerrero, Mxico. Lectotype ANSP 25049a (H. B. Baker 1963:222). : Amula, Guerrero, Mxico; syntypes in the BMNH. Distribution.GUERRERO: area east of Chilpancingo, between Tixtla and Chilapa, 1600 m alt. Coelostemma (Coelostemma ) fusca (Von Martens 1897)Holospira fusca Von Martens 1897; Biol. Cent. Amer.:281; pl. 16, Holospira (Haplocion) fusca (Von Martens). Pilsbry 1902; Man. Proc. U. S. Nat. Mus. 31:146. Haplocion fusca (Von Martens). Bartsch 1943; Jour. Wash. Acad. Sci. 33:56. Coelostemma fusca (Von Martens). Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. 105:156.Thompson 1971; Bull. Fla. Sta. Mus.,

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144 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)15:283.Thompson 1988; Bull. Fla. State Mus. 30:50. Holospira (Megaxis) fusca Von Martens. Pilsbry 1946; Land Moll. N. Amer. II:123.Type Locality.Omilteme, Guerrero, Mxico. Syntypes in the BMNH. Distribution.GUERRERO: commonly distributed in the mountains east and west of Chilpancingo; 1975 m alt. Coelostemma (Coelostemma ) hazelae Pilsbry 1953Coelostemma hazelae Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. Type Locality.Between Chilpancingo and a small town nearby called Mazatln, Guerrero, Mxico. Holotype ANSP 190966. Distribution.GUERRERO: known from a small area near Mazatln; 1425 m alt. Coelostemma (Coelostemma ) herrerae (Bartsch 1906).Holospira (Coelostemma herrerae Bartsch 1906; Proc. U. S. Nat. Coelostemma herrerae (Bartsch). Bartsch 1943; Jour. Wash. Acad. Sci. 58.Type Locality.Silaca Yuapn (= Silacayoapn), Oaxaca, Mxico. Holotype USNM 188180. Distribution.Known only from the type locality. Coelostemma (Coelostemma ) iqualaensis (Bartsch 1926)Holospira (Coelostemma) iqualaensis Bartsch 1926; Proc. U. S. Coelostemma igualaensis (Bartsch). Bartsch 1943; Jour. Wash. Acad. Sci. 33:58.Thompson 1971; Bull. Fla. State Mus. Type Locality.Iquala, Guerrero, Mxico. Holotype USNM 363135. Distribution.GUERRERO: Iquala, and the area immediately north and east; 1000 m alt. Coelostemma (Coelostemma ) leucostoma Thompson 1971Coelostemma leucostoma Thompson 1971; Bull. Fla. State Mus. Type Locality.Ruins of Xochicalco, Morelos; 4900 ft. alt. Holotype UF 21145. Distribution.Known only from the type locality. Coelostemma (Coelostemma ) lichenophora (Bartsch 1906)Holospira (Coelostemma) lichenophora Bartsch 1906; Proc. U. S. Coelostemma lichenophora (Bartsch). Bartsch 1943; Jour. Wash. Acad. Sci. 33:58. type USNM 134699. Distribution.Known only from the type locality. Coelostemma (Coelostemma ) lissocentrum Pilsbry 1953Coelostemma lissocentrum Pilsbry 1953; Proc. Acad. Nat. Sci. 10,000 ft. alt. Holotype ANSP 190968. Distribution.Known only from the type locality. Coelostemma (Coelostemma ) microstoma (Pfeiffer 1861)Cylindrella microstoma Pfeiffer 1861; Proc. Zool. Soc. Lond. 29:27.Pfeiffer 1862a:81.Pfeiffer 1868:390. Holospira microstoma (Pfeiffer). Fischer & Crosse; 1873:337; pl. Coelostemma (?) microstoma (Pfeiffer). Bartsch 1943:58. Coelostemma microstoma (Pfeiffer). Thompson & Mihalcik 2005; Type Locality.Unknown. Lectotype BMNH 1996161 (Thompson & Mihalcik 2005). Distribution.Unknown. Coelostemma (Coelostemma ) notogastor Thompson 1971Coelostemma notogastor Thompson 1971; Bull. Fla. State Mus. Type Locality.Limestone hill 14.2 mi. SW of Sola de Vega, Oaxaca, Mxico; 6500 ft. alt. Holotype UF 20910. Distribution.OAXACA: various localities near Sola de Vega and Tlapacoyan. Coelostemma (Coelostemma ) presidioensis Bartsch 1943Coelostemma presidioensis Bartsch 1943; Jour. Wash. Acad. Sci. Type Locality.Presidio, Veracruz, Mxico. Holotype USNM 536886. Distribution.Known only from the type locality. Coelostemma (Coelostemma ) richardi Thompson 1971Coelostemma richardi Thompson 1971; Bull. Fla. Sta. Mus., Type Locality.Limestone hillside 1.3 mi. NE of Tonal, Oaxaca, Mxico. Holotype UF 20905. Distribution.Known only from the type locality. Coelostemma (Coelostemma ) saltillensis Pilsbry 1953Coelostemma saltillensis Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. Type Locality.Mountain on the right of the highest pass on the road from Saltillo to Diamante, Coahuila, Mxico; 7900 ft. alt. Holotype ANSP 164078. Distribution.COAHUILA: known from the type locality and San Lorenzo Canyon SE of Saltillo on the Coahuila-Zacatecas Railroad. Coelostemma ( Coelostemma) scaphopleuron Thompson 1988Coelostemma scaphopleuron Thompson 1988; Bull. Fla. Mus. Nat. Type Locality.Limestone ridge 1.5 km. west-northwest W). Holotype UF 93145. Distribution.Known only from the type locality.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 145 Subgenus Apertaxis Pilsbry 1953Apertaxis Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. 105:159.Type Species.Coelostemma amplaxis Pilsbry 1953 (= Epirobia coahuilensis Bartsch 1906). Distribution.Sierra Guadalupe, Coahuila, Mxico. Taxonomy.One species. The generic relationships of Apertaxis remain uncertain. Coelostemma (Apertaxis) coahuilensis (Bartsch 1906)Epirobia coahuilensis Bartsch 1906; Proc. U. S. Nat. Mus. 31:121; Coelostemma (Apertaxis ) amplaxis Pilsbry 1953; Proc. Acad. Nat. Type Localities.Epirobia coahuilensis : Sierra Guadalupe, Coahuila, Mxico. Holotype USNM 187505. Coelostemma amplaxis: Sierra Guadalupe back of the Hacienda Guadalupe, Coahuila, Mxico; 7500 ft. alt. Holotype ANSP 191189. Distribution.COAHUILA: known only from the Sierra Guadalupe; 2250 m alt. Subgenus Crycoryne Thompson 1971Crycoryne Thompson 1971; Bull. Fla. St. Mus. 15:289.Type Species.Coelocentrum astraxis Thompson 1971. Distribution.Eastern Durango, Mxico. Taxonomy.This subgenus is monotypic. Coelostemma (Crycoryne) astraxis Thompson 1971Coelostemma (Crycoryne) astraxis Thompson 1971; Bull. Fla. State Type Locality.Limestone bluff 9.0 mi. south-southwest of Picardias, Durango, Mxico; 4400 ft. alt. Holotype UF 20903. Distribution.DURANGO: known from a small area along the east and south edges of the Sierra El Rosario. Subgenus Goniapex Thompson 1988Goniapex Thompson 1988; Bull. Fla. State Mus. 33:91.Type Species.Coelostemma pyrgonasta Thompson 1988. Distribution.Southern New Mexico; Chihuahua, Mxico. Taxonomy.This subgenus contains six species. Coelostemma (Goniapex) attenuapex Thompson 1988Coelostemma (Goniapex) attenuapex Thompson 1988; Bull. Fla. Type Locality.Limestone ridge 12.7 km southeast of Ciudad Camargo, Chihuahua, Mxico. Holotype UF 93144. Distribution.Known only from the type locality. Coelostemma (Goniapex) bryantwalkeri (Pilsbry 1917)Holospira bryantwalkeri Pilsbry 1917; Nautilus 30:125; pl. 4, Coelostemma (Goniapex ) bryantwalkeri (Pilsbry). Thompson 1988; Type Locality.River drift along the Rio Conchas, not Mxico. Holotype UMMZ 140145. Distribution.CHIHUAHUA: exact locality unknown. Coelostemma (Goniapex) freytagi Bartsch 1950Coelostemma freytagi Bartsch 1950; Jour. Wash. Acad. Sci. 40:265; Coelostemma (Goniapex) freytagi Bartsch. Thompson 1988; Bull. Coelostemma marrsi 1. Type Localities. Coelostemma freytagi : 29 km west of Ciudad Jimenez, Chihuahua, Mxico. Holotype USNM 601851. Coelostemma marrsi: Cueva Diablo, near Salaices, Chihuahua, Mxico. Holotype USNM 601645. Distribution.CHIHUAHUA: Sierra Almoloya, near Salaices, Chihuahua. Coelostemma (Goniapex) pyrgonasta Thompson 1988Coelostemma (Goniapex) pyrgonasta Thompson 1988; Bull. Fla. Type Locality.West-northwest side of Bishops Cap Mountain, Dona Ana County, New Mexico, 1600 m alt. Distribution.Known only from the type locality. Coelostemma (Goniapex) reiteri Drake 1951Coelostemma reiteri 5. Coelostemma (Goniapex) reiteri Drake. Thompson 1988; Bull. Fla. southeast end of the Sierra Remedios, Chihuahua, Mxico. Holotype USNM 601627. Distribution.CHIHUAHUA: low limestone mountain ranges extending NW from Los Remedios to about halfway between Ciudad Jmenez and Ciudad Camargo. Coelostemma (Goniapex) townsendi (Bartsch 1906) Holospira (Haplocion) townsendi Bartsch 1906; Proc. U. S. Nat. Haplocion townsendi Bartsch. Bartsch 1943: Jour. Wash. Acad. Sci. 33:56. Coelostemma (Goniapex ) townsendi (Bartsch).Thompson 1988; Bull. Fla. State Mus. 33:100.Type Locality.Cerro Chilote, 20 km north of Ciudad Chihuahua and due east of Aldama, Chihuahua, Mxico. Holotype USNM 10925. Distribution.Known only from the type locality. Subgenus Styloptyx Thompson 1971Styloptyx Thompson 1971; Bull. Fla. State Mus. 15:292.Type Species.Coelostemma fornax fornax Thompson 1971. Distribution.A small area in eastern Durango near

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146 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Taxonomy.One species and two subspecies are recognized. Coelostemma (Styloptyx ) fornax fornax Thompson 1971Coelostemma (Styloptyx ) fornax fornax Thompson 1971; Bull. Fla. Type Locality.Limestone mountain side 2.2 mi. [3.65 km]. S of the Presa Francisco Zarca [Zarco], Durango, Mxico; 4100 ft. [1262 m] alt. The Presa Francisco Zarco is about 19 km NE of Pedricea, and about 8 km N of the Torreon-Durango highway. Holotype UF 20895. Distribution.DURANGO: known only from a small area south of the Rio Nazas and east of the Presa Francisco Zarco. Coelostemma (Styloptyx ) fornax ix Thompson 1971Coelostemma (Styloptyx ) fornax ix Thompson 1971; Bull. Fla. Sta. Type Locality.Limestone hillside 1.0 mi. east of Dinamita, Durango, Mxico; 4200 ft. alt. Dinamita is a small Holotype UF 20907. Distribution.Known only from the type locality. Genus Hendersoniella Dall 1905Hendersonia Dall 1905; Smithson. Misc. Coll. 45:187 (not Hendersonia Wagner 1905; Gastropoda, Prosobranchia). Hendersoniella Dall 1905; Proc. Biol. Soc. Wash. 18:189.Type Species.Hendersonia palmeri Dall 1905. Potos, Mxico. Taxonomy.Three species and two subspecies are recognized. Hendersoniella christmani Thompson & Correa-Sandoval 1994Hendersoniella christmani Thompson & Correa-Sandoval 1994; Type Locality.Limestone hillside on south side of Arroyo San Juan, 2.5 km WNW of La Cienega, Municipio Distribution.NUEVO LEN: known only from the vicinity of the type locality. Hendersoniella lux lux Thompson & Correa-Sandoval 1994Hendersoniella lux lux Thompson & Correa-Sandoval 1994; Bull. Type Locality. km west of Santa Catarina, Valle de Las Fantasmas, San Lus Potos, San Lus Potos, Mxico, Distribution.SAN LUS POTOS: known only from the vicinity of the type locality. Hendersoniella lux chonomphix Thompson & Correa-Sandoval 1994Hendersoniella lux chonomphix Thompson & Correa-Sandoval Type Locality.Valle de las Fantasmas, hillside 15.5 km W of Santa Catarina, San Lus Potos, Mxico, 1770 m alt. Distribution.Known only from the type locality. Hendersoniella palmeri (Dall 1905)Hendersonia palmeri Dall 1905; Smiths. Misc. Coll. 48:187; Hendersoniella palmeri (Dall 1905). Dall 1905; Proc. Biol. Soc. Wash. 18:189.Thompson & Correa-Sandoval 1994; Bull. Fla. Hendersoniella palmeri simplex Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. 105:160. Type Localities. Hendersonia palmeri: Sierra Alvarz, San Lus Potos, Mxico. Holotype USNM 110385. Hendersoniella pameri simplex: in crevices and under stones north of the railroad station, Alvarz, San Lus Potos, Mxico. Holotype ANSP 164312. Distribution.SAN LUS POTOS: known only from the immediate vicinity of Alvarz, San Lus Potos. Genus Holospira Von Martens 1860Acera Albers 1850. (Not Acera Cuvier 1810). Holospira Von Martens; in Albers 1860:209.Thompson 1998:87 89.Thompson & Mihalcik 2005; Bull. Fla. Mus. Nat. Hist. 43:66.Type Species.Cylindrella golfussi Menke 1847, not Cylindrella piloceri Pfeiffer 1841 (OPINION 1932; International Commission of Zoological Nomenclature; Opinion 1932:206). Distribution.Oaxaca north and west to Arizona, New Mexico, and Texas. Taxonomy.Seven subgenera, 74 species, and 11 subspecies occur in Mxico. Subgenus Holospira Von Martens 1860 Distribution.From Oaxaca, Puebla, adjacent Veracruz and Guerrero north to Arizona, New Mexico and Texas. Taxonomy.Forty-eight species and two subspecies currently are recognized in Mxico. Additional species occur in Texas, New Mexico, and Arizona. The following are considered synonyms of the subgenus Holospira: Eudistemma Dall 1895 (type species: Holospira arizonensis Stearns 1890); Distomospira Dall 1895 (type species Holospira bilamellata Dall 1895); Tristemma Bartsch 1906 (type species: Holospira ferrissi Pilsbry 1905 [non Tristemma Brandt 1835, Coleoptera]); and Malinchia Bartsch 1945 (type species: Holospira ferrissi Pilsbry 1905). Holospira (Holospira) acanthidia Thompson & Mihalcik 2005Holospira acanthidia Thompson & Mihalcik 2005; Bull. Fla. Mus.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 147Type Locality.Limestone bluff 6.3 km northwest of Tehuacn, Puebla, Mxico; 1750 m altitude. Holotype: UF 34377. Distribution.Known only from the type locality. Holospira ( Holospira) Hanna & Hertlein 1929 Hanna and Hertlein 1929; Proc. Calif. Acad. Hanna & Hertline. Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. 105:151. Type Locality.On the road to Paredon, 16 km north of Ramos Arizpe. Holotype California Academy Science. Distribution.Known only from the type locality. Holospira (Holospira) albertoi Bartsch 1947Holospira albertoi Bartsch 1947; Jour. Wash. Acad. Sci. 37:141 Type Locality.Steep west bank of gully on SE lower very close to the Oaxaca border along the Pan American Highway; 1400 m alt. Holotype USNM 543495. Distribution.PUEBLA: known only from the immediate vicinity of the type locality. Holospira (Holospira) amalthea Bartsch 1926Holospira amalthea Bartsch 1926; Proc. U. S. Nat. Mus. 70:9; Holotype USNM 363140. Distribution.Known only from the type locality. Holospira (Holospira) andromeda Bartsch 1926Holospira andromeda Bartsch 1926; Proc. U. S. Nat. Mus. 70:7; Type Locality.Not stated. Holotype USNM 363138. Distribution.Unknown. Holospira (Holospira) arellanoi (Bartsch 1945)Malinchia arellanoi 6 (shell). Holospira arellanoi (Bartsch). Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. 105:141.Type Locality.Limestone hill 10 km northnortheast of Cadereyta, Quertaro, Mxico. Holotype USNM 431959. Distribution.Known only from the type locality. Holospira (Holospira) aurantiaca Thompson & Mihalcik 2005Holospira aurantiaca Thompson & Mihalcik 2005; Bull. Fla. Mus. Nat. Hist. 43:83; Figs. 130.Type Locality.Limestone canyon 2 km southeast of Distribution.Known only from the immediate vicinity of the type locality. Holospira (Holospira) bachia Bartsch 1926.Holospira bachia Bartsch 1926; Proc. U. S. Nat. Mus. 70: pl. 1, Type Locality.River drift along the Rio Balsas, Rio Balsas Station, Guerrero, Mxico. Holotype USNM 363152. Distribution.Unknown. Holospira (Holospira) catorceana Pilsbry 1953Holospira catorceana Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. Type Locality.Sierra Catorce, near the top of the northward slope facing Santana [Potero], San Lus Potos, Mxico. Holotype ANSP 164197. Distribution.Known only from the type locality. Holospira (Holospira) colymis Thompson & Mihalcik 2005Holospira colymis Thompson & Mihalcik 2005; Bull. Fla. Mus. Type Locality. Limestone hillside 13.5 km west southwest of San Bartola Teontepec, Puebla, Mxico; 2120 Distribution.PUEBLA: known only from the vicinity of the type locality; the north facing slope of a limestone hill about 7 km by road N of Los Reyes Metzontla 1800 m alt. Holospira (Holospira) cyclostoma Pilsbry 1953Holospira cyclostoma Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. Type Locality.Drift in the Rio Gochico, about 7 km from San Bernardo, Sonora, Mxico; 900 ft. alt. Holotype ANSP 166411. Distribution.SONORA; not known for certain. This species has been collected from stream drift at the type locality and at San Bls, Sinaloa, Mxico. Holospira (Holospira) denserpens Thompson & Mihalcik 2005Holospira denserpens Thompson & Mihalcik 2005; Bull. Fla. Mus. Type Locality.Limestone hill 1 km east of Azumbilla, 23 km north of Tehuacn, Puebla, Mxico; 2150 m alt. Distribution.Known only from the type locality. Holospira ( Holospira) eburnea Thompson & Mihalcik 2005Holospira eburnea Thompson & Mihalcik 2005; Bull. Fla. Mus. Type Locality.Limestone hill-top 7.6 km southDistribution.Known only from the type locality.

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148 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Holospira (Holospira) fortisculpta Thompson & Mihalcik 2005Holospira fortisculpta Thompson & Mihalcik 2005; Bull. Fla. Mus. Type Locality. km northwest of Atenco, Puebla, Mxico; 2650 m alt. Holotype: UF 34293. Distribution.Known from the vicinity of the type locality. PUEBLA: limestone ridge 1 km SW of Coyotepec, Holospira (Holospira) fergusoni Gilbertson & Naranjo-Garcia 2010Holospira (Holospira) fergusoni Gilbertson & Naranjo-Garcia Type Locality.Municipio Arteaga, Sierra La Viga, 0.5 W). 2650 m alt. Holotype: LACM 3112. Distribution.Known only from the type locality. Holospira (Holospira) goniostoma (Pfeiffer 1856)Cylindrella goniostoma Pfeiffer 1856; Malak. Bltt. 3:47. Holosira goniostoma (Pfeiffer).Fischer & Crosse 1873; Miss. Sci. Type Locality.Mxico. Lectotype BMNH 1996153 (Thompson & Mihalcik 2005:68). Distribution.Unknown. Holospira ( Holospira) haploplax Thompson & Mihalcik 2005Holospira haploplax Thompson & Mihalcik 2005; Bull. Fla. Mus. Type Locality..3 km east of Cd. Puebla, Puebla, Mxico; 2525 m alt. Holotype UF 34381. Distribution.Known only from the type locality. Holospira (Holospira) hinkleyi Pilsbry 1907Holospira hinkleyi Pilsbry 1907; Nautilus 21:27.Pilsbry 1953; Correa-Sandoval, Gutirrez & Reza 1998; Acta Zool. Mex. (73):16.Type Locality.El Abra, San Lus Potos, Mxico. Holotype ANSP 96575. Distribution.East-central SAN LUS POTOS: Holospira (Holospira) hogeana Von Martens 1897Holospira teres hogeana Von Martens 1897. Biol. Cent. Amer.:280; 71. Holospira hogeana Von Martens. Thompson & Mihalcik 2005; Type Locality.Maltrate, Veracruz, Mxico, 2299 m alt. and the City of Mxico, a little west of Orizaba, eastern slope of the plateau. Lectotype BMNH 1901.6.22.1903 (Thompson & Mihalcik 2005:77). Distribution.PUEBLA: 1 km NE of Chapulco 1940 m alt. 1.5 km NNE Chapulco 1900 m alt.; 3 km NNE Chapulco 1940 m alt.; 4 km NNE Chapulco 1940 m alt.; 5 km WNW Azumbilla, 2400 m alt. (UF 34288); 3 km NW of Tecamachalco, 2140 m alt.; 3.7 km SE of Tecamachalco, 2300 m alt.; limestone escarpment 23 km W of the PueblaVeracruz state line, 10 km E of Esperanza, 2350 m alt. (ca. Mendoza, 1750 m alt. (Thompson & Mihalcik 2005). Holospira (Holospira) hyperia Bartsch 1926Holospira hyperia Bartsch 1926; Proc. U. S. Nat. Mus. 70:6; pl. 105:141.Thompson & Mihalcik 2005; Bull. Fla. Mus. Nat. Type Locality.Esperanza, Puebla, Mxico. Holotype USNM 363146. Distribution.Known only from the type locality. Holospira (Holospira) infanta Bartsch 1906Holospira infanta Bartsch 1906; Proc. U. S. Nat. Mus. 31:129; Type Locality.Sierra Guadalupe, Coahuila, Mxico. Holotype USNM 187650. Distribution.Known only from the type locality. Holospira (Holospira) kriegeri Drake 1950Holospira kriegeriType Locality.Caon del Diablo, Sierra de Tamaulipas, Tamaulipas, Mxico. Holotype USNM 601629). Distribution.Known only from the type locality. The type specimens were collected from an archeological cave deposit, the exact locality of which is not given in Drake (1950). MacNeish (1947:2) vaguely indicated by map that the locality is along the Arroyo Las Palmas, due south of Soto La Marina. Holospira (Holospira) maxwelli Pilsbry 1953Holospira maxwelli Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. Veracruz, Mxico.. Holotype ANSP 191105. Distribution.Known only from the type locality.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 149 Holospira (Holospira) melea Bartsch 1926Holospira melea 11.Pilsbry 1953:141.Thompson & Mihalcik 2005; Bull. Fla. Type Locality.Ixcaquixtla, Puebla, Mxico. Holotype ANSP 191105. Distribution.Known only from the type locality. Holospira (Holospira) mexicana Bartsch 1906Holospira mexicana Bartsch 1906; Proc. U. S. Nat. Mus. 31:127 Type Locality.Southwestern Mxico. Holotype USNM 73987. Distribution.Unknown. Holospira (Holospira) mitraensis Bartsch 1926Holospira mitraensis Bartsch 1926; Proc. U. S. Nat. Mus. 70:101; Type Locality.La Mitra Mountain, Monterrey, Nuevo Distribution.Known only from the type locality. Holospira (Holospira) Bartsch 1925 Bartsch 1925; Proc. U. S. Nat. Mus. 67:2; Type Locality.East of Monclova, Coahuila, Mxico. Holotype USNM 361962. Distribution.Known only from the type locality. Holospira (Holospira) nelsoni Pilsbry 1902Holospira nelsoni Pilsbry 1902; Man. Conch. 15:79; pl. 22, (reproductive anatomy).Bartsch 1906; Proc. U. S. Nat. Mus. 31:132.H. B. Baker 1965; Proc. Acad. Nat. Sci. Phila. 115:224. Type Locality.Sierra Guadalupe, Coahuila, Mxico; 6500 ft. alt. Lectotype ANSP 58094a (H. B. Baker 1963). Distribution.COAHUILA: known only from the type locality and at 9500 ft alt. in the Sierra Guadalupe (Bartsch 1906). Holospira (Holospira) oaxacana Bartsch 1906Holospira oaxacana Bartsch 1906; Proc. U. S. Nat. Mus. 31:132 Type Locality.Tomelln, Oaxaca, Mxico. Lectotype USNM 175085 (Thompson & Mihalcik 2005:80). Distribution.Known only from the type locality. Holospira (Holospira) orcutti Bartsch 1925Holospira orcutti Bartsch 1925; Proc. U. S. Nat. Mus. 67:1; pl. Type Locality.A limestone paredon at Coahuila, Mxico. This is amended here-in to Paredon, Coahuila. Holotype USNM 361961. Distribution.Known only from the type locality. Holospira (Holospira) painteri Bartsch 1906Holospira painteri Bartsch 1906; Proc. U. S. Nat. Mus. 31:130; Type Locality.Tehuacn, Puebla, Mxico. Holotype USNM 187675. Distribution.Known only from the type locality. Holosira (Holospira) palmeri Bartsch 1906Holosira palmeri Bartsch 1906; Proc. U. S. Nat. Mus. 31:128 Bartsch 1926; Proc. U. S. Nat. Mus. 70:6; Type Localities. Holospira palmeri: Sierra Alvarz, San Lus Potos, Mxico; holotype USNM 100388. Holospira : Alvarez, San Lus Potos; holotype USNM 36316. Distribution.SAN LUS POTOS: known only from the immediate vicinity of the type localities. Holospira (Holospira) pedroana pedroana Bartsch 1926Holospira pedroana pedroana Bartsch 1926; Proc. U. S. Nat. Mus. Holospira creightoni Bartsch 1926; Proc. U. S. Nat. Mus. 70:8, Type Localities. Holospira pedroana pedroana: hills north of San Pedron, San Lus Potos, Mxico; holotype USNM 362176. Holospira creightoni: hill east of San Pedro, San Lus Potos; holotype USNM 326178. Distribution.SAN LUS POTOS: known from the vicinity of the type locality. Holospira (Holospira) Pilsbry 1953 Pilsbry 1953. Proc. Acad. Nat. Sci. Phila. 105:145; pl. 7, 4d (shell). T ype Locality.Hill southeast of the reservoir east of San Pedro, up to about 7100 feet on slope facing east and north, San Lus Potos, Mxico. Holotype ANSP 164377. Distribution.Known only from the type locality. Holospira (Holospira) pfeifferi (Menke 1847)Cylindrella pfeifferi Menke 1847; Zeitsch. fr Malak., 4:1. Holospira pfeifferi (Menke). Strebel 1880; Bietrag, IV:84; pl. 13, 51.Thompson & Mihalcik 2005; Bull. Fla. Mus. Nat. Hist. Type Locality.Tehuacn, Puebla, Mxico. Lectotype BMNH 1996134 (Thompson & Mihalcik 2005:80). Distribution.Known only from the type locality. Holospira (Holospira) picta Bartsch 1925Holospira picta Bartsch 1925; Proc. U. S. Nat. Mus. 67:3, pl. 1, Type Locality.East of Monclova, Coahuila, Mxico; holotype USNM 361964. Distribution.Known only from the type locality.

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150 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Holospira (Holospira) politecnica (Bartsch 1945)Malinchia politecnica Bartsch 1945; Jour. Wash. Acad. Sci. 35:94; Holospira politecnica (Bartsch). Pilsbry 1953: Proc. Acad. Nat. Sci. Phila. 105:141.Type Locality. km north-northwest of Cadereyte, Quertaro, Mxico. Holotype USNM 341961. Distribution.Known only from the type locality. Holospira (Holospira) queretaroensis (Bartsch 1945)Malinchia queretaroensis Bartsch 1945; Jour. Wash. Acad. Sci. Type Locality.Limestone hill 10 km northnorthwest of Cadereyte, Quertaro, Mxico. Holotype USNM 431963. Distribution.Known only from the type locality. Holospira (Holospira) rehderi Bartsch 1947Holospira rehderi Bartsch 1947; Jour. Wash. Acad. Sci. 37:287 Holospira morelosensis Bartsch 1947; Jour. Wash. Acad. Sci. Type Localities. Holospira rehderi: Chietla, Puebla, Mxico; holotype USNM 543589. Holospira morelosensis: Tlaquiltenango, Morelos; holotype USNM 543591. Distribution.Known from a broad area in eastern Morelos and adjacent Puebla and Oaxaca, from 1000 m alt. MORELOS:7.2 km SSW of Ticumn, 1033 m alt.; 12.6 km SSE of Izucar de Matamoros, 1700 m alt.; 13 km SE of Izucar de Matamoros, 1475 m alt. OAXACA: 2.7 km limestone hill 7 km E of Teposcolula, km Post 7, 2160 m alt.; 2 km NE of Tonal, 1540 m alt.; 13.4 km NE of Tonal, 2060 m alt.; 13.5 km N of Tonal, 2090 m alt. Holospira ( Holospira) rhinion Thompson & Mihalcik 2005Holospira rhinion Thompson & Mihalcik 2005; Bull. Fla. Mus. Nat. Type Locality.Limestone hillside on the west side of Federal Highway 131, 15 km southeast of Tehuacn, Puebla, Mxico; 1540 m altitude. Holotype UF 34375. Distribution.PUEBLA: known only from the immediate vicinity of the type locality. Holospira (Holospira) scololaema Thompson & Mihalcik 2005Holospira scololaema Thompson & Mihalcik 2005; Bull. Fla. Mus. Type Locality.Limestone mountain side 1 km northeast of Yehualtepec, Puebla, Mxico; 2340 m alt. Holotype UF 34289. Distribution.Known only from the type locality. Holospira (Holospira) stenopylis Pilsbry 1953Holospira stenopylis Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. Type Locality.Slopes on the north side of the valley northeast of San Pedro, San Lus Potos, Mxico. Holotype ANSP 164376. Distribution.Known only from the type locality. Holospira (Holospira) temeroso Pilsbry 1953Holospira temeroso Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. Mountain, on the trail up from Aranzaza, Zacatecas, Mxico; 9250 feet alt. Holotype ANSP 164131. Distribution.Known only from the type locality. Holospira (Holospira) teotitlana Bartsch 1945Holospira teotitlana teotitlana Bartsch 1945; Jour. Wash. Acad. Sci. Bartsch 1945; Jour. Wash. Acad. Sci. Holospira teotitlana Bartsch. Thompson & Mihalcik; Bull. Fla. Type Localities. Holospira teotitlana teotitlana : foothills of Cerro Blanco (Cerro de Tizatepec) at Teotitln del Camino, Oaxaca, Mxico; holotype USNM 431954. : Cerro Tizatepec, near Ignacio Mejia, Oaxaca; holotype USNM 431957. Distribution.Eastern Puebla and adjacent Oaxaca in the Tehuacn Tomelln Valley, at 790 meters altitude. PUEBLA: 10.7 km N of Teotitln del Camino, 950 m alt. ; 2.5 km N of Teotitln del Camino; 10 km NW of Coxcatln, 880 m alt; 11.5 km SSE of Coxcatln. OAXACA: 2.7 km Holospira (Holospira) teres (Menke 1847)Cylindrella teres Menke 1847; Zeitschrift fr Malakologische 4:1.29. Holospira teres (Menke). Fischer & Crosse 1873; Miss. Sci. Mex. I:327.Von Martens 1897, Biol. Cent. Amer:279; pl. 17, 20.Haas 1933; Archiv f. Molluskenkunde 65:271.Bartsch 1943; Jour. Wash. Acad. Sci. 33:58.Thompson & Mihalcik Holospira teres minor Von Martens 1897; Biol. Cent. Amer.:278.Type Locality.State of Puebla, Mxico. Lectotype SMF 7023 (Thompson & Mihalcik 2005:75). Distribution.Unknown.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 151 Holospira (Holospira) tetralasma Pilsbry 1902Holospira tetralasm Pilsbry 1902. Man. Conch. 15:73; pl. 21, Mus. Nat. Hist. 43:79.Type Locality.Mxico. Holotype in the Dohrn Collection at the Stettin Museum, which is now Muzeum Narodowe, Szczecin, Poland, The Dohrn mollusk collection was totally destroyed in the 1939 War (Dance 1986:219). Distribution.Unknown. Holospira (Holospira) topochicoana Bartsch 1926Holospira topochicoana Bartsch 1926; Proc. U. S. Nat. Mus. 70:12 Type Locality.Topochico Mountain, Monterey, Nuevo Distribution.Known only from the type locality. Holospira (Holospira) torrei Pilsbry 1935Holospira torrei Pilsbry 1935. Proc. Acad. Nat. Sci. Phila. 87:2; pl. Type Locality.Mxico. Holotype USNM 363143. Distribution.Unknown. Hiolospira (Holospira) wilmoti Bartsch 1951Holospira wilmoti 2. Coelostemma wilmoti (Bartsch). Thompson 2008:356.Type Locality.Cerro del Fraile, near Villa Garcia, Distribution.Known only from the type locality. Holospira ( Holospira) zygoptyx Thompson & Mihalcik 2005Holospira zygoptyx Thompson & Mihalcik 2005; Bull. Fla. Mus. Type Locality..0 km by road south of San Antonia W). Holotype: UF 190771. Distribution.Known only from the type locality. Subgenus Allocoryphe Pilsbry 1946Allocoryphe Pilsbry 1946; Land Moll. N. Amer. 2:123.Type Species.Holospira minima Von Martens 1897. Distribution.Central Sonora, Mxico. Taxonomy.Two species are recognized. Holospira (Allocoryphe ) guaymasensis (Bartsch 1943)Haplocion guaymasensis Bartsch 1943; Jour. Wash. Acad. Sci. Holospira (Allocoryphe) guaymasensis (Bartsch). Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. 105:152.Type Locality.Guaymas, Sonora, Mxico; holotype USNM 536883. Distribution.Known only from the type locality. Holospira (Allocoryphe ) minima Von Martens 1897Holospira pfeifferi var. minima Von Martens 1897; Biol. Cent. Holospira pfeifferi var. percostata Pilsbry 1902; Man. Conch. 15:95; Haplocion minima Bartsch 1943; Jour. Wash. Acad. Sci. 33:56. Holospira (Allocoryphe ) minima Von Martens. Pilsbry 1953; Proc. Land Moll. N. Amer. II:123.Thompson 1988; Bull. Fla. Mus. (shell), 3a (reproductive anatomy).Type Localities. var. minima: Mxico. Holospira minima var. percostata: N.W. Mxico. Holotype in the ANSP. Distribution.SONORA: immediate vicinity of Hermosilla. Subgenus Bostrichocentrum Strebel 1880Bostrichocentrum Strebel 1880; Beitrag. Mex. Landund Sssw.Conch. IV:80.Type Species.Cylindrella tryoni Pfeiffer 1867. Distribution.The subgenus is concentrated in southcentral Mxico in northeastern Oaxaca, Puebla, and adjacent Veracruz. Two species also occur in northeastern Mxico. Taxonomy.Fifteen species are recognized. Holospira (Bostrichocentrum) anomala Pilsbry 1953Holospira anomala Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. Type Locality.Tepeaca, about 22 km east of Cd. Puebla, Puebla, Mxico. Holotype ANSP 191108. Distribution.Known only from the type locality. Holospira ( Bostrichocentrum) centicostata Thompson 1964Holospira (Bostrichocentrum ) centicostata Thompson 1964; Type Locality.A caliche hillside 14.4 miles northwest of Acatln, Puebla, Mxico; 4300 feet alt. Holotype UMMZ 213218. Distribution.Known only from the type locality. Holospira (Bostrichocentrum) eurybia Bartsch 1926Holospira (Bostrichocentrum ) eurybia Bartsch 1926; Proc. U. S. Bostrichocentrum eurybia (Bartsch). Bartsch 1943; Jour. Wash. Acad. Sci. 33:55.Type Locality.River drift, Rio Balsas Station, Guerrero, Mxico. Holotype USNM 363147. Distribution.Unknown. Holospira (Bostrichocentrum) galathea Bartsch 1926Holospira (Bostrichocentrum) galathea Bartsch 1926; Proc. U. S. Bostrichocentrym galathea (Bartsch). Bartsch 1943; Jour. Wash. Acad. Sci. 33:55.

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152 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Type Locality.River drift, Rio Balsas Station, Guerrero, Mxico. Holotype USNM 363148. Distribution.Unknown. Holospira (Bostrichocentrum) gealei (H. Adams 1872)Cylindrella (Holospira) gealei H. Adams 1872; Proc. Zool. Soc. Holospira tryoni gealei (H. Adams). Pilsbry 1902; Man. Conch. Bostrichocentrym gealei (H. Adams). Bartsch 1943; Jour. Wash. Acad. Sci. 33:55.Type Locality.Putla, Puebla, Mxico; syntype in the BM(NH). Distribution.Known only from the type locality. Holospira (Bostrichocentrum) goldmani Bartsch 1906Holospira (Bostrichocentrum) goldmani Bartsch 1906; Proc. Bostrichocentrum goldmani (Bartsch). Bartsch 1943; Jour. Wash. Acad. Sci. 33:55.Bartsch 1947; Jour. Wash. Acad. Sci. 37:142.Type Locality.Tamazulapn, Oaxaca, Mxico. Holotype USNM 187793. Distribution.OAXACA: Tamazulapan; 5.7 miles NW American Highway km. post 304; 2.2 miles SE of Chila. Holospira (Bostrichocentrum) hidalgoensis Bartsch 1906Holospira (Bostrichocentrum ) hidalgoensis Bartsch 1906; Proc. U. Bostrichocentrym hidalgoensis (Bartsch). Bartsch 1943; Jour. Wash. Acad. Sci. 33:55.Type Locality.Zimapn, Hidalgo, Mxico. Holotype USNM 187982. Distribution.Known only from the type locality. Holospira (Bostrichocentrum) perplexa Thompson 1964Holospira (Bostrichocentrum ) perplexa Thompson 1964; Type Locality.Hill 10.3 mi. northwest of Huahapn de Distribution.Known only from the type locality. Holospira (Bostrichocentrum) pilsbryi Dall 1895Holospira pilsbryi Dall 1895; Proc. U. S. Nat. Mus. 18:4. Holospira (Bostrichocentrum ) pilsbryi Dall. Pilsbry; 1902; Man. Bostrichocentrum pilsbryi (Dall). Bartsch 1943; Jour. Wash. Acad. Sci. 33:55.Type Locality.Around sulfer springs near the City of Puebla, Puebla, Mxico. Holotype USNM 56932. Distribution.PUEBLA: vicinity of the type locality and Tepeaca. Holospira (Bostrichocentrum) pupa Thompson 1964Holospir (Bostrichocentrum) pupa Thompson 1964. Malacologia Type Locality. miles southeast of Acatln, Puebla, Mxico; 4900 feet alt. Holotype UMMZ 213217. Distribution.Known only from the type locality. Holospira (Bostrichocentrum) ronzoni Bartsch 1943Bostrichocentrum ronzoni Bartsch 1943; Jour. Wash. Acad. Sci. Type Locality.Pajaro Verde, Puebla, Mxico. Holotype USNM 536874. Distribution.PUEBLA: known for certain only from Orizaba, Veracruz (Bartsch 1943:56). Holospira (Bostrichocentrum) tamaulipensis Bartsch 1906Holospira (Bostrichocentrum ) tamaulipensis Bartsch 1906; Proc. U. Bostrichocentrum tamaulipensis (Bartsch). Bartsch 1943; Jour. Wash. Acad. Sci. 33:55.Type Locality.[Ciudad] Camargo, Tamaulipas, Mxico. Holotype 187979. Distribution.Known only from the type locality. The type locality is so far removed from the rest of the range of Bostrichocentrum as to raise doubt about its authenticity. Holospira (Bostrichocentrum) tryoni (Pfeiffer 1867)Cylindrella tryoni Pfeiffer 1867; Journal de Conchyl., 15:438. Holospira tryoni (Pfeiffer). Crosse & Fischer 1870; Jour. de Conchyl. Cent. Amer.:276.Cross 1892; Jour. de Conchyl. 40:267; Holospira (Bostrichocentrum ) tryoni (Pfeiffer). Pilsbry 1902; Man. Bostrichocentrum tryoni (Pfeiffer). Strebel 1880; Bietrag., IV:81; pl. Acad. Sci. 33:55. Holospira tryoni appressa Fischer & Crosse 1873; Miss. Sci. Mex. I:331. Type Localities. Cylindrella tryoni: Izucr, Puebla, Mxico. Holospira tryoni appressa: Izucr, Puebla, Mxico. Distribution.Known only from the type locality. Holospira (Bostrichocentrum) Dall 1895 Dall 1895; Proc. U. S. Nat. Mus. 18:4. Holospira (Bostrichocentrum ) Dall. Pilsbry 1902; Man. Conch. 15:85. (Dall). Bartsch 1943; Jour. Wash. Acad. Sci. 33:55.Type Locality.Misantla, Veracruz, Mxico. Holotype in the USNM. Distribution.Known only from the type locality. Holospira ( Bostrichocentrum) (Bartsch 1943) Bartsch 1943; Jour. Wash. Acad. Type Locality.In the neighborhood of Orizaba or

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 153 536877. Distribution.VERACRUZ: questionably known from the state. Subgenus Millerspira Gilbertson & Naranjo-Garcia 2004Millerella Gilbertson & Naranjo-Garcia 1998; Veliger 41:91 (non Millerella Thompson 1942; Protista). Millerspira Gilbertson & Naranjo-Garcia 2004; Veliger 47:157. Type Species.Holospira milleri Gilbertson 1989. Distribution.East-central Sonora, Mxico. Taxonomy.Two species. Holospira (Millerspira) milleri Gilbertson 1989Holospira (Millerella ) milleri Gilbertson 1998; Veliger 32:91; Holospira (Millerspira) milleri Gilbertson & Naranjo-Garcia 1998; Veliger 41:314. Type Locality.Sonora, on the east side of the Rio Yaqui, under conglomerate rocks (containing calcite) in a ravine near the mouth of the Rio Alamo, ca. 1.5 km south of Distribution.Known only from the type locality. Holospira (Millerspira) hoffmani Gilbertson & Naranjo-Garcia 1998Holospira (Millerella) hoffmani Gilbertson & Naranjo-Garcia 1998; Type Locality.Sierra Batamonte (near El Milagro Mine, 20.5 km E of Rio Yaqui Bridge along Hwy. 15 from Estrella to Bacanora, Sonora, Mxico (2830.5 W). Holotype SBMNH 143186. Distribution.SONORA: known only from the vicinity of the type locality. Subgenus Prionoloplax Pilsbry 1953Prionoloplax Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. 105:140.Type Species.Holospira odontoplax Pilsbry 1953. Distribution.Southern Coahuila, Mxico. Taxonomy.This subgenus is monotypic. Holospira (Prionoloplax) odontoplax (Pilsbry 1953)Holospira (Prionoloplax ) odontoplax Pilsbry 1953; Proc. Acad. Type Locality.First mountain of the Sierra de Penitente on the right side of the road from Saltillo to Diamonte, at the highest pass, Coahuila, Mxico; 7800 feet alt. Holotype ANSP 164077. Distribution.Known only from the type locality. Subgenus Sonoraloa Gilbertson 1993Sonoraloa Gilbertson 1993; Amer. Malac. Bull., 10:72.Type Species.Cylindrella remondi Gabb 1865. Distribution.Sonora and Sinaloa, Mxico. Taxonomy.The subgenus includes four species and seven subspecies. Holospira (Sonoraloa) dentaxis dentaxis Pilsbry 1953Holospira dentaxis dentaxis Pilsbry 1953; Proc. Acad. Nat. Sci. Holospira (Sonoraloa) dentaxis dentaxis Pilsbry. Gilbertson 1993; Amer. Malac. Bull., 10:72. Type Locality.River drift at ford of Rio Yaqui about ANSP 166484. Holospira ( Sonoraloa) dentaxis alamellata Gilbertson 1993Holospira (Sonoraloa) dentaxis alamellata Gilbertson 1993; Amer. Malac. Bull., 10:73. Type Locality.Riperian gorge, south side of Hwy 15, 7.5 km by road west of Sahuaripe, Sonora, Mxico; 700 m Distribution.Known only from the type locality. Holospira (Sonoraloa) dentaxis lamellata Pilsbry 1953Holospira dentaxis lamellata Pilsbry 1953; Proc. Acad. Nat. Sci. Bull., 10:75. Type Locality.River drift at ford of Rio Yaqui about ANSP 190760. Distribution.SONORA: exact locality unknown. Holospira (Sonoraloa) dentaxis potamia Pilsbry 1953Holospira dentaxis potamia Pilsbry 1953; Proc. Acad. Nat. Sci. Bull., 10:75. Type Locality.River drift at ford of Rio Yaqui about ANSP 191107. Distribution.SONORA: exact locality unknown. Holospira (Sonoraloa) dentaxis striatella Pilsbry 1953Holospira dentaxis striatella Pilsbry 1953; Proc. Acad. Nat. Sci. Bull., 10:75. Type Locality.River drift at ford of Rio Yaqui about ANSP 191106. Distribution.SONORA: exact locality unknown. Holospira (Sonoraloa) kinonis J. Baily & R. Baily 1940Holospira kinonis Baily & Baily 1940; Nautilus 53:94; pl. 12, Type Locality.Tidal drift 2 miles south of Cochare, near Guaymas, Sonora, Mxico. Holotype ANSP 174953. Distribution.SONORA: exact locality unknown. Holospira (Sonoraloa) mazatlanica (Bartsch 1943)Haplocion mazatlanica Bartsch 1943; Jour. Wash. Acad. Sci. 33:56;

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154 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Holospira (Sonoraloa) mazatlanica (Bartsch). Gilbertson 1993; Amer. Malac. Bull., 10:73. Type Locality.Drift at Mazatln, Sinaloa, Mxico. Holotype USNM 536884). Distribution.Unknown. Holospira (Sonoraloa) remondi remondi (Gabb 1865)Cylindrella remondi 10. Holospira remondi (Gabb). Tryon 1867; Amer. Jour. Conch. 3:313.Holospira remondi remondi (Gabb). Pilsbry 1953; Proc. Acad. Nat. Malac. Bull., 10:72. Haplocion remondi (Gabb). Bartsch 1943; Jour. Wash. Acad. Sci. 33:56.Type Locality..5 leagues (ca. 6 km) from Arivechi, Sahuaripe Valley, Sonora, Mxico. Lectotype ANSP 166484. Distribution.SONORA: known only from the type locality and from river drift in the Rio Yaqui north of Ciudad Holospira (Sonoraloa) remondi forticostata Pilsbry 1953Holospira remondi forticostata Pilsbry 1953; Proc. Acad. Nat. Sci. Type Locality.Drift debris of the Rio Yaqui at the ford Holotype ANSP 166406. Distribution.SONORA: exact locality unknown. Holospira (Sonoraloa) Pilsbry 1953Holospira Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. (reproductive anatomy). Type Locality.Drift of Laguna Presa Rodriguez, Hermosillo, Sonora, Mxico. Holotype ANSP 188316. Distribution.SONORA: exact locality unknown. Holospira (Sonoraloa) remondi yaquensis Pilsbry 1953Holospira remondi yaquensis Pilsbry 1953; Proc. Acad. Nat. Sci. Malac. Bull., 10:73. Type Locality.Drift debris of the Rio Yaqui at the ford Holotype ANSP 166405. Distribution.SONORA: exact locality unknown. Subgenus Stalactella Bartsch 1906Stalactella Bartsch 1906; Bull. U. S. Nat. Mus. 31:151.Rehder 1940:315, Pilsbry 1953:135.Thompson & Mihalcik 2005; Bull. Fla. Mus. Nat. Hist. 43:89.Type Species.Holospira (Stalactella) rosei Bartsch 1906. Distribution.Southern Puebla and adjacent Oaxaca, Mxico. Taxonomy.Five species. Previous authors (e.g., Pilsbry 1953) treated Stalactella as a subgenus of Propilsbrya. Internal and external aspects of the shell indicate that Stalactella is derived from a species-group of Holospira s. s. that is localized in southern Puebla, and no close relationship exists with Propilsbrya. Holospira (Stalactella ) chazumbae Thompson & Mihalcik 2005Holospira (Stalactella) chazumbae Thompson & Mihalcik 2005:94 Type Locality.Hillside 2 km northeast of Santiago Chazumba, Oaxaca, Mxico; 1970 m alt. Holotype UF 34299. Distribution.Known only from the type locality. Holospira (Stalactella ) cremnobates Thompson & Mihalcik 2005Holospira (Stalactella ) cremnobates Thompson & Mihalcik 2005; Type Locality. km north of Tehuacn, Puebla, Mxico; 1990 m alt. (18 UF 190789. Distribution.Known only from the type locality. Holospira (Stalactella ) marmorata Thompson & Mihalcik 2005Holospira (Stalactella ) marmorata Thompson & Mihalcik 2005; Type Locality..5 km WSW of San Bartolo north side of the highway. Snails were found under caliche slabs and limestone boulders. Holotype UF 233188. Distribution.Six km N of Santa Cruz Nueva, Puebla. 1760 m alt. (18 km S of San Vicente Coyotepec, 2080 m alt. (18 N, 97 Coyotepec 1880 m alt. (UF 233190). Holospira ( Stalactella) psectra Thompson & Mihalcik 2005Holospira (Stalactella ) psectra Thompson & Mihalcik 2005; Bull. Type Locality.A limestone cliff on the south slope of Cerro Caolalote, 2 km east of El Carmen, Puebla, Mxico; is at the base of a south-facing bare limestone cliff. Holotype UF 233197. Distribution.Known only from the type locality. Holospira (Stalactella ) rosei Bartsch 1906Holospira (Stalactella ) rosei Bartsch 1906; Proc. U. S. Nat. Mus. 171. Type Locality.Tehuacn, Puebla, Mxico. Holotype

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 155 USNM 188181. Distribution.Known only from the type locality. Names of uncertain status associated with Holospira Cylindrella cretacea Pfeiffer 1860.Cylindrella cretacea Pfeiffer 1860; Proc. Zool. Soc. Lond. 28:140.Type Locality.Mxico. Syntype in the Cuming Collection, BMNH. Distribution.Unknown. Remarks.The interior structure of the shell is unknown thereby preventing subgeneric assignment. Pilsbry (1902) commented on its external similarity to Haplocion coahuilensis (Binney 1865). Cylindrella (Holospira) imbricata Von Martens 1863Cylindrella (Holospira) imbricata Von Martens 1863; Monatber. Berlin Akademie der Wissenschafte:540.Fischer & Crosse 1873; Miss. Sci. Mex. I:336.Von Martens 1897, Biol. Cent. Type Locality.Mxico. Distribution.Unknown. Remarks.The interior of the shell is unknown. Externally it resembles some species of Coelostemma and Bostrichocentrum Cylindrella pilocerei Pfeiffer 1841Cylindrella pilocerei Pfeiffer 1841; Symbolae ad Hist. Helic. 1:47.Type Locality.Cuautla de Las Amilpas, Puebla [Morelos]; syntype apparently lost. A nomum dubium (OPINION 1932; International Commission of Zoological Nomenclature; Bulletin of Zoological Nomenclature, 56:206; September 1999). Distribution.Known only from the type locality; presumed to be extinct. Genus Metastoma Strebel 1880Metastoma Strebel 1880; Beitrag, IV:80.Pilsbry 1902; Man. Conch. 15:96.Pilsbry 1946; Land Moll. N. Amer. II:114.Thompson 1971; Bull. Fla. St. Mus. 15:299. Type Species.Cylindrella roemeri Pfeiffer 1848, by original designation. Distribution.Southwestern Texas, southeastern New Mexico, northeastern Chihuahua, and northern Coahuila. Taxonomy.A single species is recognized. Metastoma roemeri (Pfeiffer 1848)Cylindrella roemeri Pfeiffer 1848; Mono. Helic. Viv., 2:382. Holospira roemeri (Pfeiffer).Binney 1878; Terrest. Moll. N. Amer., U. S. Nat. Mus. 31:148.Pilsbry 1946; Land Moll. N. Amer. Metastoma roemeri (Pfeiffer).Thompson 1971; Bull. Fla. St. Mus. 15:300.Type Locality.Around New Braunfels, Comal County, Texas. Distribution.Southwestern Texas and southeastern New Mexico. Previously unrecorded from Mxico. CHIHUAHUA: limestone knoll, 7.5 km NE of Coyame, La Encantada, Caon La Ventana de Boquilla, 27 km WNW Ranco La Babia, 91 km NW Melechor Mosquiz, 1150 m alt. limestone knoll, 2.5 km W of La Cuesta, 1500 m alt. (UF mountain 3 km E La Cuesta, 1380 m alt. (UF 244929); Sierra La Encantada, canyon 3 km ESE La Cuesta, 1420 m alt. (UF 244930). Family UROCOPTIDAE Pilsbry 1898 Distribution.The West Indies in general, and adjacent areas of the mainland, including Florida, Mxico. Central America, Colombia, and Venezuela. Taxonomy.There are numerous genera in the West Indies. Two genera representing two subfamilies have colonized Mxico and Central America. The results from a recent study by Uit de Weerd (2008) necessitated the recognition of the Eucalodiidae, Epirobiidae, and Holospiridae as families distinct from the Urocoptidae. Subfamily BRACHYPODELLINAE H. B. Baker 1956 Genus Brachypodella Beck 1837Brachypus Guilding 1828 (not Brachypus Swainson 1824). Brachypodella Beck 1837 Index Moll. :89.Pilsbry & Vanatta 1898: 277, 278.Pilsbry 1904:40. Brachypodisca Agassiz 1847 (emendation for Brachypodella ). (type species: Helix (Cochlodina) antiperersa Ferussc 1821).Type Species.Brachypodella : Helix (Cochlodina ) Ferussc 1821 (Pilsbry & Vanatta 1898). Distribution.Antilles except for western Cuba and the Bahama Islands; mainland of Trinidad and Venezuela; southeastern Mxico and northern Guatemala. Taxonomy.About six subgenera and 50 species. Only one subgenus occurs in Mxico and Central America. Subgenus Brachypodella Beck 1837 Distribution.Hispaniola southeast through the Lesser Antilles, Trinidad and Venezuela, Guatemala, and southern Mxico. Taxonomy.Numerous species. Five species and three subspecies occur in Mxico and Central America. Brachypodella bourguignatiana (Ancey 1886)Cylindrella bourguignatiana Ancey 1886; Bull. Soc. Malac. France 2:243.Pilsbry 1891; Proc. Acad. Nat. Sci. Phila. 43:316; pl. Brachypodella bourguignatiana (Ancey). Pilsbry 1904; Man.

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156 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Philos. Soc. 79:176.Type Locality.Isla de tila, Honduras. Distribution.HONDURAS: tila and Roatn, Dept. Islas de la Baha. Brachypodella dubia (Pilsbry 1891)Cylindrella spelucae var. dubia Pilsbry 1891; Proc. Phila. Acad. Brachypodella speluncae (Morelet) (in part). Pilsbry 1904; Man. 1933; Pub. Carnegie Inst. Wash. (431):535.Berquaert & Clench 1936; Pub. Carnegie Inst. Wash. (457): 65.Bequaert & Clench 1938; Pub. Carnegie Inst. Wash. (491):258.Richards 1937; Proc. Amer. Philos. Soc. 77:254.Harry 1950; Occ. Pap. Mus. Zool. Univ. Mich.; (524):16.Branson and McCoy 1965; Univ. Col. Stud. Biol., (13):9. Brachypodella speluncae var. dubia (Pilsbry). Pilsbry 1904; Man. Conch. 16:71. Brachypodella dubia (Pilsbry). Thompson 1967; Bull. Fla. Sta. Type Locality.Labn, Yucatn, Mxico. Lectotype ANSP 62027a (H. B. Baker 1963:222). Distribution.The Yucanan Peninsula of Mxico. CAMPECHE: 19.2 mi. S of Pixtn; 6.1 mi. SW of Seybaplaya; 7.1 mi. SW of Campeche; 5.1 mi. NNW of Dzilbalchn; 5.1 mi. W of Tikinml. QUINTANA ROO: 7.1 mi. NW of Xiatil; Isla Cozumel, 1.5 km NE of San Miquel. YUCATN: Labn; 10.0 mi. NE of Becanchn; cave at Tabi; Ticul; between Sitilpech and Tunkas. Brachypodella morini morini (Morelet 1849)Cylindrella morini Morelet 1849; Test. Noviss. I:11.Pfeiffer 1849; Monogr. Helic. Viv., 3:578.Fischer & Crosse 1873; Miss. Sci. in Conch. Icon., 43:338. Holospira morini(Morelet). Von Martens 1897; Biol. Cent. Amer.: 285. Brachypodella morini.(Morelet). Pilsbry 1904; Man. Conch. 16:67 Type Locality.Vera Pz, Guatemala. Distribution.GUATEMALA, Dept. Alta Verapz, Brachypodella morini salpinx (Tristram 1861)Cylindrella salpinx Tristram 1861; Proc. Zool. Soc. Lond. 29:231.Fischer & Crosse 1873; Miss. Sci. Mex. I:415. Holospira morini var. salpinx (Tristam). Von Martens 1897; Biol. Brachypodella morini var. salpinx (Tristam). Pilsbry 1904; Man. Type Locality.Probably near Cobn or Lanqun, Dept. Alta Verapz, Guatemala. Distribution.Known only from the type locality. Brachypodella morini sargi (Von Martens 1897)Holospira morini var. sargi Von Martens 1897; Biol. Cent. Amer.:285. Brachypodella morini form sargi (Von Martens). Pilsbry 1904; Man. Conch. 16:68. Guatemala. Distribution.Known only from the type locality. Brachypodella speluncae (Morelet 1852)Cylindrella speluncae Morelet 1852; Test. Nov., 2:12.Fischer & Holospira speluncae (Morelet). Von Martens 1897; Biol. Cent. Amer.:284. Brachypodella speluncae (Morelet). Pilsbry 1904; Man. Conch. 16: 1937; Misc. Pub. Mus. Zool. Univ. Mich. (34):28.Thompson Cylindrella costulata Morelet 1851; Test. Noviss. II:12, (not Cylindrella costulata C. B. Adams 1849).Type Locality.Walls of the Cave Jobitsinal, Dept. Petn, Guatemala. Distribution.GUATEMALA, Dept. Petn: knoll 1.2 mi. S of Flores; Cubixinal Cave, S of Flores; 7 mi. SW of San Benito; 2 km S of Puebla Nueva; knoll E of La Libertad; west shore Lago de Petnxl; San Andres; Tabi; Ticl; between Sitilpech and Tuncas, Labn. Brachypodella subtilis subtilis (Morelet 1849)Cylindrella subtilis Morelet 1849; Test. Noviss. I:11.Fischer & Holospira subtilis (Morelet). Von Martens 1897; Biol. Cent. Amer.:284. Brachypodella subtilis (Morelet). Pilsbry 1904; Man. Conch. 16:69: 73:131.Type Locality.Forests of Petn, Guatemala. Distribution.GUATEMALA: known from the type locality. BELIZE: Cayo Dist.: Rio Frio Cave East, about 2 mi. from Augustine on the Mountain Pine Ridge. Brachypodella subtilis pulchella (Von Martens 1886)Cylindrella (Gongylostoma) pulchella Von Martens 1886; Sitz. Ges. Nat. Freude Berlin:161 Holospira morini var. pulchella (Von Martens). Von Martens 1897; Brachypodella morini var. pulchella (Von Martens). Pilsbry 1904; Brachypodella subtilis pulchella (Von Martens). Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 71:218.Type Locality.Livingston, Dept. Izabl, Guatemala. Distribution.Known only from the type locality. Doubtful name associated with Mexican and Guatemalan Brachypodella Brachypodella subula Frussac 1821Clausilia subula Frussac 1821:61, (nomen nudum).Deshayas, in Lamarck 1838:216. Brachypodella subula (Frussac). Beck 1837:89.Pilsbry 1904;

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 157 Type Locality.Unknown. Remarks.The type is lost, and its internal shell structure is unknown. Pilsbry (1904) suggests that this taxon is most similar to species of Brachypodella described from Guatemala. Subfamily MICROCERAMINAE Pilsbry 1904 Genus Microceramus Pilsbry & Vanatta 1898Microceramus Pilsbry & Vanatta 1904; Nautilus 11:107.Type Species. Pilsbry 1898. Distribution.Greater Antilles, Curacao, and Central America north to Texas and Florida. Taxonomy.Only a single subgenus is recognized. Floreziella Weyrauch 1967, from Peru is treated as a synonym by Richardson (1991:69), but its anatomy remains unknown. About 35 species are recognized. Three species and two subspecies occur in the study area. Microceramus concisus concisus (Morelet 1849)Cylindrella concisus Morelet 1849; Test. Noviss. I:12. Macroceramus concisus (Morelet). Petit 1850; Jour. de Conchyl. 1:379.Fischer & Crosse 1873; Miss. Sci. Mex. I:421; pl. 18, Martens 1897; Biol. Cent. Amer.:287. Microceramus concisus (Morelet). Pilsbry 1904; Man. Conch. Phila. 82:221.Haas and Solem 1960; Nautilus 73:131. Macroceramus polystreptus Tristram 1861; Proc. Zool. Soc. Lond. Type Localities.Cylindrella concisa: Yucatan, Mxico. Macrocermus polystreptus: Cobn, Dept. Alta Verapaz, Guatemala. Distribution.Nicaragua, Honduras, Isla de San Andrs, Isla del Cisne Grande, Isla del Cisne Pequeo, northern Guatemala, Belize, Yucatn, and Campeche. Microceramus concisus arctispirus (Ancey 1886)Macroceramus gossei var. arctispirus Ancey 1886; Ann. de Malacologie II:242. Microceramus consisus var. arctispirus (Ancey). Pilsbry 1904; Amer. Philos. Soc. 79:173.Type Locality.Isla de tila, Honduras. Distribution.HONDURAS, Dept. Islas de la Baha: Roatn and Isla de tila. Microceramus kieneri (Pfeiffer 1846)Bulimus kieneri Pfeiffer 1846; Proc. Zool. Soc. Lond. 14:40. Macroceramus kieneri (Pfeiffer). Pfeiffer 1859. Monographia heliceorum viventium 4:689.Bland 1883; Ann. New York Microcerumus kieneri (Pfeiffer). Pilsbry 1904; Man. Conch. 16:154; Type Locality.Honduras. Distribution.Unknown. Microceramus mexicanus (Von Martens 1897)Macroceramus concisus var. mexicanus Von Martens 1897; Biol. Microcermus mexicanus (Von Martens). Pilsbry 1904; Man. Conch. 31:157.Pilsbry 1953; Proc. Acad. Nat. Sci. Phila. 105:161.Correa-Sandoval 1993; Rev. Biol. Trop. 41:685.-CorreaSandoval 1998; Acta Zool. Mex. (73):16.Correa-Sandoval 2002; Acta Zool. Mex. 86:239.Correa-Sandoval 2000; Acta Zool. Mex. 79:9. (Gould). Strebel 1880; Beitrag. Mex. Type Locality.Here restricted to Orizaba, Veracruz, Mxico. Distribution.Widely distributed and common. Reported from the Mexican states of: MXICO, MORELOS, JALISCO, NUEVO LEN, SAN LUS POTOS, TAMAULIPAS and VERACRUZ. Superfamily ACHATINOIDEA Swainson 1840 Family FERUSSACIDAE Bourguignat 1883 Genus Cecilioides Frussac 1814Cecilioides Frussac 1814; Mem. Geologiques:48. Pilsbry 1908; Man. Conch. 20:1.Hugh Watson 1828; Jour. of Conch. 18:217 (anatomy).Pilsbry 1946; Land Moll. N. Amer. 2:184.Zilch 1959:338. Caecilioides Hermannsen 1846; Ind. Gen. Malac., 1:150 (emendation for Cecilioides ).Type Species.Bulimus acicula Bruguire (= Buccinum aciculums Mller 1774). Distribution.Europe, Africa, southern Asia, Philippines, Oceania, and the American tropics. Three occur naturally in the neotropical region. Subgenus Karolus De Folin 1870Karolus De Folin 1870; Les Fonds des Mer tude internationale sur les particularits nouvelles des regions sous-marines 1:182 189.Pilsbry 1916; Nautilus 19:119. Caecilianopsis Pilsbry 1907; Nautilus 21:28.Pilsbry 1909; Man. Conch. 20:5, 38.Type Species. Caecilioides (Caecilianopsis ) jod Pilsbry 1907 (= Karolus primus De Folin 1870). Distribution.Tropical America. Introduced elsewhere. Taxonomy.The subgenus includes three species. One species, Cecilioides consobrinus, is widespread in the American tropics, and consists of several subspecies. The typical subspecies occurs in Cuba. Another subspecies, C. c. primus, is found in Mxico and Central America. Cecilioides (Karolus) consobrinus primus (De Folin 1870)Karolus primus De Folin 1870; Les Fonds des Mer :189; pl. 28, Caecilioides consobrinus primus (De Folin). Pilsbry 1916; Nautilus 29:119.Harry 1950; Occ. Pap. Mus. Zool. Univ. Mich. (524):19. Cecilioides consobrinus primus (De Folin). Dall 1926; Proc. Calif. Acad. Sci. (4) 15:471472.Baker 1930:14.Thompson 1967; Bull. Fla. St. Mus. 11:240. Crosse & Fischer 1877; Jour. de

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158 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) (shell).Martens 1898:324. Caecilioides consobrinus var. (Crosse & Fischer). 79 (shell).Goodrich & van der Schalie 1937:29.Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):9. (Crosse & Fischer). CorreaSandoval, Guerrero & Reza 1998:15.Correa-Sandoval 2000; Acta Zool. Mex. (79):9.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):238. Caecilioides (Caecilianopsis ) jod Pilsbry 1907; Nautilus 21:28. Cecilioides consobrinus Type Localities. Karolus primus: Veracruz, Mxico. : among debris drifted out of the Rio Antigua, Veracruz, Mxico. Caecilioides jod: drift debris along the Rio Panuco, Tamaulipas, Mxico. Verapaz: Chama (Hinkley 1920b). Dept. Izabal: Quiragua; Sierra Cavech; Jocolo (Hinkley 1920b). Dept. Petn: Tikal (Thompson 1967). NAYARIT: Isla Mara Madre, Islas Maras; Isla Socorro (Dall 1926). PUEBLA: Necaxa, 3000 ft. alt. (Baker 1930a). NUEVO LEN: Iturbide (Correa-Sandoval 1997). SAN LUS POTOS: numerous localities (CorreaSandoval et al. 1998). TAMAULIPAS: numerous localities (Correa-Sandoval & Rodriguez 2002). VERACRUZ: Rio YUCATN: near Chichen Itza (Harry 1950); 0.8 mi. NE of Becanchn (Thompson 1967). Taxonomy.There is a tendency in current literature to disregard subspecies, and cite only species names, which automatically take on the name of the typical subspecies. It is presumptive of later workers to assume which subspecies was intended. Such is the case with the work by Prez & Cecilioides consobrinus consobrinus in the study area. Subgenus Caecilianopsis Pilsbry 1907Caecilianopsis Pilsbry 1907; Nautilus 21:28.Type Species.Cecilioides (Caecilianopsis ) jod Pilsbry 1907. Distribution.Northeastern Mxico. Taxonomy.The subgenus contains a single species. Cecilioides (Caecilianopis ) jod Pilsbry 1907Cecilioides (Caecilianopis ) jod Pilsbry 1907; Nautilus 21:28.Hinkley 1907; Nautilus 21:77.Type Locality.In river drift, Tampico, Tamaulipas, Mxico. Distribution.TAMAULIPAS: known only from the tpe locality. Subgenus Geostilba Crosse 1867Geostilba Crosse 1867; Jour. de Conchyl. 15:184.Pilsbry 1909; Man. Conch. 20:43.Type Species.Geostilba caledonica Crosse 1867. Distribution.Tropics of both hemispheres. Taxonomy.Many species. One occurs in the study area. Cecilioides (Geostilba) aperta (Swainson 1840)Macrospira aperta Swainson 1840; Treatise on Malacology:335; Geostilbs aperta (Swainson). E. A. Smith 1895; Proc. Zool. Soc. Lond. 63:307. Cecilioides aperta (Swainson). Pilsbry 1946; Land Moll. N. Amer. Achatina gundlachi Pfeiffer 1850; Zeit. fr Malak. 7:80. Cecilioides gundlachi (Pfeiffer). Pilsbry 1909; Man. Conch. 20:43.Type Localities. Macrospira aperta: West Indies. Achatina gundlachi : Cuba. Distribution.West Indies in general; introduced Family SUBULINIDAE Fischer & Crosse 1877 Distribution.Tropical America east through Africa and southeast Asia. Taxonomy.Numerous genera. Subfamily SUBULININAE Fischer & Crosse 1877 Taxonomy.Most Middle American Subulinidae had been grouped in the genus Leptinaria. A species critical to this Allopeas gracilis (Hutton 1834), which was placed variously within Opeas and then Lamellaxis, which in turn was considered a subgenus of Leptinaria. H. B. Baker (1927 1935 1945) pointed out the anatomical differences between Opeas, Beckianum Allopeas Leptopeas, Lamellaxis and Leptinaria. The anatomies of only a few species among these genera are known. Authors continued to unite most of the Middle American species within Leptinaria because of the lack of discrete shell differences between it and Lamellaxis, Leptopeas, and Allopeas Opeas, Allopeas Leptopeas, Lamellaxis, and Beckianum as separate genera because of their known anatomies. Genus Allopeas H. B. Baker 1935Allopeas H. B. Baker 1935; Nautilus 48:84.H. B. Baker 1945; Nautilus 58:88 (anatomy).Type Species.Bulimus gracilis Hutton 1834. Distribution.Tropical, subtropical, and many temperate regions of the paleotropical regions of the world. Allopeas gracilis and a few closely related species are readily dispersed elsewhere through human activities. Taxonomy.Several species. Conventionally, Allopeas was treated as a subgenus of Lamellaxis. More recently it is recognized as a separate genus because of its anatomical dissimilarity with Lamellaxis and other genera.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 159 Allopeas gracilis (Hutton 1934)Bulimus gracilis Hutton 1834; Jour. As. Soc. Bengal 3:93. Opeas gracilis (Hutton). Pilsbry 1906; Manual of Conchology (2) British India, II (Trochomorphidae Janellidae):355.Morrison 1946; Smiths. Misc. Coll. 106:41. Lamellaxis gracilis (Hutton). Thompson 1967; 241. Lamellaxis (Allopeas) gracilis (Hutton). H. B. Baker 1934; Nautilus 48:84.H. B. Baker 1945; Nautilus 58:88 (anatomy). 8 (pallial organs), 9 (reproductive anatomy), 10 f, g (shell).Correa-Sandoval et al. 1998:15.Correa-Sandoval 2002:126; map. Achatina subula Pfeiffer 1839; Wiegmanns Archive f. Naturg., 1:352. Opeas subula (Pfeiffer). Von Martens 1898; Biol. Cent. Amer.:291 29. Bulimus octonoides 22.Type Locality.Mirzapur, Uttar Pradesh State, India. Distribution.This is the worlds most widely distributed land snail (Pilsbry 1946). It is known from the following localities in the study area. PANAM, Archipilago de las Perlas: Isla San Jos (Morrison 1946). Prov. Panam: Panam; Quipo (Pilsbry 1926a); Mt. Hope (Pilsbry 1930). 2002). HONDURAS, Dept. Islas de la Baha: Isla de tila. GUATEMALA, Dept. Alta Verapaz: Cobn. Dept. Petn: Tikal (Basch 1959). Dept. Sacatepquez: Antigua. CAMPECHE: various localities (Thompson 1967). SAN LUS POTOS: numerous localities (Correa-Sandoval 1998). TABASCO: San Juan Bautista (Pils-bry 1906). TAMAULIPAS: Valle de Aguayo, Cd. Victoria (Correa-Sandoval & Rodriguez 2002). YUCATN: Merida. Genus Beckianum H. B. Baker 1961Beckianum 2002:120. Synopeas Jousseaume 1889 (not Synopeas Forster 1856, Hymenoptera). Diaopeas Haas 1962:55.Naranjo-Garcia & Fahy 2010; Amer. Maloc. Bull., 28:66.Type Species.Beckianum : Bulimus beckianus Pfeiffer 1846. Diaopeas: Bulimus beckianus Pfeiffer 1846. Distribution.Tropical America in general from Brazil and Bolivia north to Mxico, and the West Indies. Taxonomy.Diaopeas is an objective junior synonym of Beckianum because it has the same type species. Two species of Beckianum are recognized. Beckianum beckianum beckianum (Pfeiffer 1846)Bulimus beckianus Pfeiffer 1846; Symbolae ad Historiam Heliciorum 3:82. Opeas beckianum (Pfeiffer). Pilsbry 1906; Man. Conch. 18:189 48, 32. Synopeas beckianum (Pfeiffer). H. B. Baker 1940:93.H. B. Baker 1945; Nautilus 58:91 (anatomy).H. B. Baker 1947; Nautilus 61: pl,1 (reproductive anatomy).Thompson 1957:101.Bequaert 1957:216. Beckianum beckianum (Pfeiffer). H. B. Baker 1961; Nautilus 75:64.Correa-Sandoval et al. 1998:15.Correa-Sandoval 2000; Acta Zool. Mex. (79):9.Correa-Sandoval & Rodriguez 2002; Acta Diaopeas beckianum (shell).Naranjo-Garcia & Fahy 2010; Amer. Malc. Bull., 28:66. Bulimus curaccasensis Opeas curaccasensis (Reeve). Fischer & Crosse 1877:599; pl. 26, Opeas micra var. caraccasensis (Reeve). Von Martens 1898; Biol. Distribution.The same as for the genus. The species determine its original distribution. PANAM, Prov. Bocas del Toro: San San Creek; Mono Creek. Prov. Los Santos: Tonosi. Isla Taboga (Pilsbry 1926a). Prov. Panam: ruins Cahuita (Pilsbry 1926b). Prov. Guanacaste: Bahia de Salinas. Prov. Puntarenas: Savana de Guacimo, 200 m alt.; Turubanes, Dept. Alta Verapaz: Chama (Hinkley 1920b). Dept. Izabal: Quirigua, Sierra Cavech; Jocolo (Hinkley 1920b). Dept. Petn: Tikal (Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):9). BELIZE: Cayo Dist.: Rio Frio Cave (Haas & Solem 1960). CAMPECHE. CHIAPAS: Ocosingo, 850 m alt.; Laguna Ocotal, 950 m alt.; Monte Libano to El Censo, 600 700 m alt. (Bequaert 1957). PUEBLA: Necaxa (H. B. Baker 1940). SAN LUS POTOS: numerous localities (CorreaSandoval et al. 1998). TABASCO: 0.5,0 mi. E of Teapa (Thompson 1957). TAMAULIPAS: Gomez Farias (CorreaSandoval & Rodriguez 2002); El Cielo Biosphere Reserve (Correa-Sandoval & Rodriguez 2005). VERACRUZ: Atoyac Orizaba; Dos Arroyos; Barranca de Mahuistlan, near Jalapa; Pacho; Mirador, 2630 ft. alt. (Von Martens 1898); Rancho Beckianum beckianum gabbianum (Angas 1879)Stenogyrus gabbiana Angas 1879; Proc. Zool. Soc. Lond. 47:485; Opeas beckianum var gabbianum (Angus). Pilsbry 1906; Man. Type Locality.Costa Rica. Distribution.Known only from the type locality.

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160 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Beckianum sinistrum (Von Martens 1898)Leptinaria sinistra Von Martens 1898; Biologia Centrali-Americana; Beckianum sinistrum (Von Martens).Prez and Lopez 1995; Type Locality.Central Nicaragua: Acoyapa on the north shore of Lake Nicaragua. Distribution.COSTA RICA, Prov. Guanacaste: Palo Verde. NICARAGUA, region of the two great lakes, Dept. Boaco: Rio Santa Rita, Tustepec; Tecolostote; El Boqueron, Teustepe. Dept. Carazo: Chacocente Biological Station. Dept. Lon: Asososca; Laguna Asososca; Laguna Monte Galan. Dept. Managua: Laguna Xiloa. Dept. Masaya: Laguna Genus Lamellaxis Strebel & Pfeffer 1882Lamellaxis Strebel & Pfeffer 1882; Beitrag. Mex. Landund Sssw.Conch.:109.Pilsbry 1907; Man. Conch.; 18:287.H. B. Baker 1927c:17.H. B. Baker 1945:85, 88.Pilsbry 1946; Land Moll. N. Amer. 2:175.Type Species.Spiraxis mexicanus Pfeiffer 1866. Distribution.This is a tropical American genus which is distributed from Mxico south to Bolivia. occur in the study area. Other species occur in South America. The arrangement of species given below follows Von Martens (1898) and Pilsbry (1907). Formerly most of the species were placed in Leptinaria, subgenus Lamellaxis. Data provided by H. B. Baker (1927c. 1945) necessitated the recognition of Lamellaxis as a separate genus. These nonlamellate Mexican and Central American species form a cohesive group of similar appearing species. Lamellaxis lacks a parietal lamella, and the outer lateral teeth of the radula are tri-cuspid, where-as Leptinaria has a parietal lamella at some stage of development, and the outer laterals are bi-cuspid. Lamellaxis tamaulipensis (Pilsbry 1903)Leptinaria tamaulipensis Pilsbry 1903; Proc. Acad. Nat. Sci. Phila. Garca-Cubas & Reguero 1998:15.Correa-Sandoval & 2002:139.Naranjo-Garcia & Fahy 2010; Amer. Malaco. Bull., 28:66.Type Locality.A caon about 4 miles west of [Ciudad] Victoria, Tamaulipas, Mxico; about 3000 ft. alt. Syntypes ANSP 85909. Distribution.NICARAGUA, Dept. Chinandega (Pm alt. (Correa-Sandoval, Garca-Cubas & Reguero 1998). TAMAULIPAS: between km 652, between Santa InezSanta Llera (Jacobson 1952); numerous localities (CorreaSandoval & Rodriguez 202); El Cielo Biosphere Reserve (Corra-Sandoval & Rodriguez 2005). Lamellaxis martensi martensi (Pfeiffer 1856)Bulimus martensi Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:318. Spiraxis martensi 9 (shell). Leptinaria martensi (Pfeiffer). Von Martens 1898; Biol. Cent. (shell).Dall 1926; Proc. Calif. Acad. Sci. (4) 15:472; pl. 35, Wash. (431):536.Bequaert & Clench 1936; Pub. Canegie Inst. Wash. (457):65.Goodrich & van der Schalie 1937:29.Richards 1937; Proc. Amer. Philos. Soc. 77:254.Rehder 1966; Proc. Biol. Soc. Wash. 79:283). Lamellaxis (Lamellaxis ) martensi (Pfeiffer). H. B. Baker 1940; Nautilus 53:93.H. B. Baker 1945:91.Harry 1950; Occ. Pap. Mus. Zool. Univ. Mich. (524):18.Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):9. Distribution.GUATEMALA, Dept. Petn: Tikal (Basch 1959). NAYARIT: Islas Maras, Isla Mara Madre; Isla Maria Magdalena, (Dall 1926). VERACRUZ: Antigua 1300 ft. alt. (H. B. Baker 1940). QUINTANA ROO: San Miguel; San Gerbacio, Isla Cozumel (Richards 1937; Rehder 1966). YUCATN: Chichen Itza; Balam-Canche Cave, 4.8 km E of Chichen Itza (Bequaert &Clench 1933 1936); 8 mi. NW of Chichen Itza, near road to Merida (Harry 1950). Lamellaxis martensi modestus Strebel 1882Lamellaxis modestus Strebel 1882; Beitrag. Mex. Landund 31 (shell). Leptinaria martensi var. modesta (Strebel). Pilsbry 1907:308; pl. Lamellaxis (Lamellaxis) martensi form modestus (Strebel). H. B. Baker 1940:93.H. B. Baker 1945; Nautilus 58:91.Type Locality.Misantla, Veracruz, Mxico. Distribution.VERACRUZ: Misantla; Atoyac to Lamellaxis mexicanus mexicanus (Pfeiffer 1866)Spiraxis mexicanus Pfeiffer 1866; Malak. Bltt. 13:84.Fischer & Crosse 1877:617. Lamellaxis mexicanus (Pfeiffer). Strebel 1882; Beitrag. Mex. Land7a, 38 (shell). Leptinaria (Lamellaxis) mexicana (Pfeiffer). Von Martens 1898; Biol. Cent. Amer.:316.Pilsbry 1907; Man. Conch. 18:306 & Reguero 1998:15.Correa-Sandoval 2000; Acta Zool. Mex. (79):9.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):237. Lamellaxis (Lamellaxis) mexicanus (Pfeiffer). H. B. Baker 1940; Nautilus 53:93.Richards 1938:173.Richards 1939; Proc. Amer. Philos. Soc. 81:34.H. B. Baker 1946; Nautilus 59: 4 (reproductive system).Correa-Sandoval et al. 1998:15.Correa-Sandoval 1999:9.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):236.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 161 Von Martens 1898; Biol. Cent. Amer.:316.Type Localities. Spiraxis mexicanus: Mirador, Veracruz, Mxico. Leptinaria mexicana var. turreta: Agua Caliente [near Misatla], Veracruz, Mxico. Distribution.NICARAGUA: Quin Bluff, S end of Isla Grande del Maz; Isla de Ometepe (Richards 1939). HONDURAS, Dept. Islas de la Baha: Between Coxen Hole and French Harbor; West End, Isla de Roatn (Richards 1938). PUEBLA: Necaxa, 2215 ft. alt. (H. B. Baker 1940). SAN LUIS POTOS: numerous localities in eastern part of state (Correa-Sandoval, Garca-Cubas & Reguero 1998). TAMAULIPAS: Gomez Farias; Valle de los Ovnis, San Jos; Rio Frio (Correa-Sandoval & Rodriguez 2002). VERACRUZ: Misantla; Caada de Coatepec, near Jalapa (Pilsbry 1907); northern part of state, numerous localities (Correa-Sandoval 1999). (Von Martens 1898)Leptinaria mexicanus var. Von Martens 1898; Biol. Cent. Amer.:316.Pilsbry 1907; Man. Conch. 18:307; pl. 41, Lamellaxis (Lamellaxis ) mexicanus form (Von Martens). H. B. Baker 1940:93.Type Locality.Mirador, Veracruz, Mxico. Distribution.PUEBLA: Necaxa, 2215 ft. alt. (H. B. Baker 1940). VERACRUZ: Mirador; Agua Caliente, near Misantla (Von Martens 1898); Texola (Pilsbry 1907). Lamellaxis mexicanus utilensis (Pilsbry 1907)Leptinaria mexicanus utilensis Pilsbry 1907; Man. Conch. 18:307 Type Locality.Isla de tila, Dept. Islas de la Baha, Honduras. Distribution.Known only from the type locality. Lamellaxis pittieri pittieri (Von Martens 1898)Leptinaria pittieri Von Martens 1898; Biol. Cent. Amer.:317; pl. 18, (shell). Distribution.COSTA RICA, Prov. Alajuela: Alajuela; Bruschik, Alta Tararia. Prov. Puntarenas: middle course of the Rio Pacuare del Sur. Prov. San Jos: San Francisco de Guadalupe; La Ueruca, 1100 m alt.; La Palma (Von Martens 1898). Lamellaxis pittieri obliquatus (Von Martens 1898)Leptinaria pittieri var. oblquatus Von Martens 1898; Biol. Cent. Type Locality.Tarbaca, [Prov. San Jos], Costa Rica; 1700 m alt. Distribution.Known only from the type locality. Lamellaxis hyalinus (Tate 1870)Tornatellina hyalina Tate 1870; American Journal of Conchology, 5:157. Leptinaria hyalina (Tate). Von Martens 1898; Biol. Cent. Type Locality.Toro Rapids on the Rio San Juan; Dept. Rio San Juan, Nicaragua. Distribution.Known only from the type locality. Taxonomy.Von Martens (1898) stated that Lamellaxis pittieri may be a synonym of L. hyalinus. Lamellaxis exiguus (Von Martens 1898)Leptinaria exigua Von Martens 1898; Biol. Cent. Amer.:318; pl. 18, (shell). Lamellaxis exiguus (Von Martens). Bequaert 1957:216.Type Locality.Teapa, Tabasco, Mxico. Distribution.CHIAPAS: Laguna Ocotal, 950 m alt.; Ocosingo, 1000 m alt. (Bequaert 1957). TABASCO: Teapa. Lamellaxis interstriatus (Tate 1870)Tornatellina interstriata Tate 1870; American Journal of Leptinaria interstriata (Tate). Von Martens 1898; Biol. Cent. Amer.:318.Pilsbry 1907; Man. Conch. 18:3101; pl. 41, Lamellaxis interstriatus (Tate). Thompon 2008:522.NaranjoGarcia & Fahy 2010; Amer. Malaco. Bull., 28:66.Type Locality.On an island in the lagoon of Boca del Toro, Panam. Distribution.Known only from the type locality. COSTA RICA, Prov. Puntarenas: plains of the Rio Grande de Lamellaxis crenulatus (Von Martens 1898)Leptinaria crenulata Von Martens 1898; Biol. Cent. Amer.:318; pl. Costa Rica. Distribution.Known only from the type locality. Lamellaxis guatemalensis guatemalensis (Crosse & Fischer 1877)Spiraxis guatemalensis Crosse & Fischer 1877; Jour. de Conchyl. Leptinaria guatemalensis (Crosse & Fischer). Von Martens 1898; Biol. Cent. Amer.:319.Pilsbry 1907; Man. Conch. 18:312; pl. 2002:131.Type Locality.Cobn, Dept. Alta Verapaz, Guatemala. Distribution.GUATEMALA, Dept. Izabal: Quiri-gua; Sierra Cavech; Jocolo (Hinkley 1920b). Dept. Retalhueleu: Retalhueleu (Von Martens 1898). NICARAGUA: various Lamellaxis guatemalensis majusculus (Von Martens 1898)Leptinaria guatemalensis var. majuscule Von Martens 1892; Biol.

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162 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Type Locality.None selected. Distribution.COSTA RICA, Prov. Guanacaste: Guanacaste. Prov. Puntarenas: valley of the Rio Grande de Terraba, Palmar, 30 m alt.; Golfo Dulce. Prov. San Jos: San Jos (Von Martens 1898). Strebel 1882 Strebel 1882; Beitrag. Mex. Landund (Strebel). Von Martens 1898; Biol. Cent. 17 (shell).Pilsbry 1926a; Proc. Acad. Nat. Sci. Phila. 78:90; Phila. 82:343.Goodrich & van der Schalie 1937:29. Type Locality.San Miguel Jucume, Guatemala [probably San Miguel Turucu in the Polochic Valley, Von Martens 1898; Biol. Cent. Amer.:319]. Tabernilla and San Pedro; Los Cascades; Isla Barro Colorado; road SE of Empire. (Pilsbry 1926a). Prov. Panam: ruins of Old Panam City (Pilsbry 1930). Isla Taboga (Pilsbry 1926a); Isla San Jos, Archipilago de las Perlas (Morrison 1946). HONDURAS, Dept. Copn: Copn (Von Martens 1898). GUATEMALA: only from the type locality. Lamellaxis strebelianus (Pilsbry 1907)Leptinaria strebeliana Pilsbry 1907; Man. Conch. 18:313; pl. Distribution.Known only from the type locality. Lamellaxis simpsoni (Ancey 1886)Nothus simpsoni Ancey 1886; Annales de Malacologie, 2:245. Leptinaria simpsoni (Ancey). Pilsbry 1907; Man. Conch. 18:314; pl. Type Locality.Isla de tila, Dept. Islas de la Baha, Honduras. Distribution.Known only from the type locality. Lamellaxis fordianus (Ancey 1886)Nothus fordianus Ancey 1886; Annales de Malacologie, 2:248. Leptinaria fordiana (Ancey). Pilsbry 1907; Man. Conch. 18:34; Type Locality.Isla de tila, Dept. Islas de la Baha, Honduras. Distribution.BELIZE: Cayo Dist.: Rio Frio Cave (Haas & Solem 1960). HONDURAS: only from the type locality. Lamellaxis yucatanensis (Pilsbry 1907)Leptinaria yucatanensis Pilsbry 1907; Man. Conch. 18:315; pl. Type Locality.Labna, Yucatn, Mxico. Syntypes in the ANSP. Distribution.Known only from the type locality. Lamellaxis biolleyi (Von Martens 1898)Leptinaria biolleyi Von Martens 1898; Biol. Cent. Amer.:319; pl. Type Locality.Near San Jos, Costa Rica. Lamellaxis costaricanus (Von Martens 1898)Leptinaria costaricana Von Martens 1898; Biol. Cent. Amer.:320, Distribution.COSTA RICA, Prov. Puntarenas: Terraba; El Pital; Rio Naranjo valley; plains of the Rio Corredor; Golfo Dulce region; Quebrada de Java; Djiri; Durania; San Blas valley; Rio Diquis valley. Prov. San Jos: San Jos (Von Martens 1898). PANAM, Prov. Los Santos: Tonosi (Pilsbry 1926a). Lamellaxis panamensis panamensis (Pilsbry 1910)Leptinaria panamensis Pilsbry 1910; Proc. Acad. Nat. Sci. Phila. Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. 82:343.Morrison 1946; Smiths. Misc. Coll. 106:41.Type Locality.between San Pedro and Tabernilla, Distribution.PANAM, Isla San Jos, Archipilago Road E of Empire, Canal Zone. Prov. Panam: ruins of old Panam City (Pilsbry 1930). Lamellaxis panamensis tabogensis ( Pilsbry 1930)Leptinaria pananensis Pilsbry. Pilsbry 1926; Proc. Acad. Nat. Sci. Leptinaria panamensis tabogensis Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. 82:343.Type Locality.Isla Taboga, Panam. Distribution.Known only from the type locality. Lamellaxis imperforatus Strebel 1882Lamellaxis imperforatus Strebel 1882; Beitrag. Mex. Landund Naranjo-Garcia & Fahy 2010; Amer. Malaco. Bull., 28:66. Leptinaria imperforata (Strebel). Pilsbry 1907; Man. Conch. Type Locality.Jalapa, Veracruz, Mxico. Distribution.Known only from the type locality. Lamellaxis hapaloides (Von Martens 1898)Leptinaria hapaloides Von Martens 1898; Biol. Cent. Amer.:321; pl. Type Locality.Plains of the Rio Grande de Terraba, Prov. Puntarenas, Costa Rica; 20 m alt. Distribution.Known only from the type locality.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 163 Lamellaxis ambiguus (Von Martens 1898)Leptinaria ambigua Von Martens 1898; Biol. Cent. Amer.:321; pl. Viejo. Prov. San Jos: La Palma (Von Martens 1898). Genus Leptopeas H. B. Baker 1927Leptopeas H. B. Baker 1927c:17.H. B. Baker 1945; Nautilus 58:85.Type Species.Leptinaria bequaerti Pilsbry 1926a. Distribution.Brazil, Venezuela, and British Guayana, north to Mxico; Known from Cuba and Hispaniola in the West Indies. Introduced elsewhere. Taxonomy.Eight species occur in the study area. Leptopeas argutus (Pilsbry 1906)Opeas argutum Lamellaxis (Leptopeas) argutus (Pilsbry). H. B. Baker 1940; Nautilus 53:92.H. B. Baker 1945; Nautilus 58:90.Lamellaxis argutus (Pilsbry). Naranjo-Garcia & Fahy 2010; Amer. Malaco. Bull., 28:66.Type Locality.Orizaba, Veracruz, Mxico; 500 ft. alt. above the city. Distribution.PUEBLA: Necaxa (H. B. Baker 1940). 1940); Texolo (Pilsbry 1906). Leptopeas bocourtianum bocourtianum (Crosse & Fischer 1869)Stenogyra bocourtiana Crosse & Fischer 1869; Journ. De Conchyl. 17:424. Opeas bocourtianum (Crosse & Fischer). Fischer & Crosse 87, 88 (shell).Type Locality.Dept. Alta Verapaz, Guatemala; in the stomach of Euglandina plicatula. Distribution.Known only from the type locality. Leptopeas bocourtianum pittieri (Von Martens 1898)Opeas bocourtianum var. pittieri Von Martens 1898; Biol. Cent. Type Locality.La Palma, Costa Rica, 1500 m alt. There are at least three places named La Palma in Costa Rica. Distribution.Known only from the type locality. Leptopeas gladiolus (Crosse & Fischer 1877)Opeas gladiolus Crosse & Fischer 1877; Jour. de Conchyl. 25:272.Type Locality.Guatemala. Distribution.Known only from the type locality. YUCATN: Merida (Martine 1898). Leptopeas colimense (Crosse & Fischer 1869)Stenogyra colimensis Crosse & Fischer 1869; Jour. de Conchyl. 17:424. Opeas colimense (Crosse & Fischer). Fischer & Crosse 1877:603; Proc. Acad. Nat. Sci. Phila.:343.Naranjo-Garcia & Fahy 2010; Amer. Malaco. Bull., 28:66.Type Locality.Colima, Mxico (Pilsbry 1930). COLIMA: type locality. VERACRUZ: Leptopeas guatemalense guatemalense (Strebel 1882)Opeas guatemalensis Strebel 1882; Beitrag. Mex. Landund Opeas guatemalense Strebel. Von Martens 1898; Biol. Cent. Amer.:293.Pilsbry 1906; Man. Conch. 18:213; pl. 29, Type Locality.Cobn, Dept. Alta Verapaz, Guatemala. Distribution.COSTA RICA, Prov. Puntarenas: along the Rio Coto, Golfo Dulce. GUATEMALA, Dept. Alta Verapaz: Cobn. Dept. Sacatepquez: Antigua (Von Martens 1898). Leptopeas guatemalense majus Von Martens 1898Opeas guatemalense var. majus Von Martens 1898; Biol. Cent. Type Locality.Miramar, near San Francisco, Dept. Escuintla, Guatemala. Distribution.Known only from the type locality. Leptopeas micra micra (Orbigny 1835)Helix micra Orbigny 1835; Magazin de Zool.:9. Opeas micra (Orbigny). Von Martens 1898; Biol. Cent. Amer.:294 296.Pilsbry 1906; Man. Conch. 18:193.H. B. Baker 1927c:10 (radula).Pilsbry 1926a; Proc. Acad. Nat. Sci. Phila. Lamellaxis micra (Orbigny). Richards 1937; Proc. Amer. Philos. Soc. 77:254.Richards 1938:173.Richards 1939; Proc. Amer. Philos. Soc. 81:34.Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):9.Correa-Sandoval, Garca-Cubas & Reguero 1998:15.Correa-Sandoval 1999:9.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):237. Lamellaxis (Leptopeas) micra (Orbigny). H. B. Baker 1945; Nautilus 58:90.Thompson 1957:101. Lamellaxis (Allopeas) micra (Orbigny).Pilsbry 1946; Land Moll. Bulimus octonoides C. B. Adams 1845; Proceedings of the Boston Society of Natural History:12. Opeas octonoides of authory. Bulimus tryonianus Tate 1870; American Journal of Conchology, Opeas micra var. tryonianum (Tate). Pilsbry 1906; Man. Conch. Sci. Phila.. 78:93. Opeas tryonianum var. Von Martens 1898; Biol. Cent.

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164 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Opeas octonoides var. Von Martens. Pilsbry 1906; Man. Allopeas micra micra (Orbigny). Naranjo-Garcia & Fahy 2010; Amer. Malaco. Bull., 28:66. Leptopeas micra (Orbigny). Thompson 2008:541.Type Localities. Helix micra: Eastern-most foothills of the Andes, not far from Santa Cruz de la Sierra, Bolivia. Bulimus octonoides: Jamaica. Bulimus tryonianus: island in the lagoon of Bocas del Toro, Panam. Opeas micra var. tryonianum: Turubanes [Turrucares], Prov. Alajuela, Costa Rica. Distribution.Bolivia to Mxico and the West Indies (Pilsbry 1946). PANAM, Prov. Bocas del Toro: island in he Darin (Pilsbry 1926a); Puerto Bello; Isla Barro Colorado (Pilsbry 1930). Prov. Panam: ruins of Old Panam City. Cahuita (Pilsbry 1926b). NICARAGUA, numerous localities Isla Grande del Maz (Richards 1939). Dept. Managua: Managua (Jacobson 1968). HONDURAS, Dept. Islas de la Baha: Isla de tila; Isla de Roatn (Richards 1938). GUATEMALA, Dept. Izabal: Quirigua; Esmeralda; Jocolo (Hinkley 1920b). Dept. Petn: Tikl (Basch 1959). SAN LUS POTOS: numerous localities (Correa-Sandoval et al.. 1998). TABASCO: San Juan Bautista (Pilsbry 1906); 0.5.0 mi. E of Teapa (Thompson 1957). TAMAULIPAS: Tampico (Hinkley 1907); El Cielo Biosphere Reserve (CorreaSandoval & Rodriguez 2005). VERACRUZ: San Rafael Jicaltepec; Antigua; Texolo; Veracruz (Pilsbry 1906); Rancho YUCATN: Izamal; Progreso; Tekanto; Ticul; Tunkas (Pilsbry 1906). Leptopeas micra mazatlanica Pilsbry 1931Opeas micra mazatlanica 12.Naranjo-Garcia & Fahy 2010; Amer. Malaco. Bull., 28:66.Type Locality.Under stones on the trail to the summit, Isla del Faru, Mazatlkan, Sinaloa, Mxico. Holotype ANSP 151314. Distribution.Known only from the type locality. Leptopeas semistriatum (Morelet 1851)Bulimus semistriatus Morelet 1851; Test. Noviss. II:10. Bulimulus semistriatus (Morelet). Fischer & Crosse 1877:555; pl. Opeas semistriatum (Morelet). Von Martens 1898; Biol. Cent. Amer.:296,Pilsbry 1906; Man. Conch. 18:212; pl. 29, Lamellaxis semistriatus (Morelet). Naranjo-Garcia & Fahy 2010; Amer. Malaco. Bull., 28:66.Type Locality.Forests of Palenque, Chiapas, Mxico. Distribution.Known only from the type locality. Leptopeas yucatanense (Pilsbry 1906)Opeas yucatanense 69 (shell).Bequaert & Clench 1933; Pub. Carnegie Inst. Wash. (431):536.Bequaert & Clench 1938; Pub. Canegie Inst. Wash. (491):258.Type Locality.Ticul, Yucatn, Mxico. Syntypes in the ANSP. Distribution.YUCATN: Balaam Canche Cave, Chichen Itza; Chac Mol Cave, Tohil (Bequaert & Clench 1938). Genus Leptinaria Beck 1839Leptinaria Beck 1837; Index Molluscorum:79.Fischer & Crosse 1877:622.Pilsbry 1907; Man. Conch. 18:284.H. B. Baker 1927c:22.H. B. Baker 1945:88. Type Species.Helix unilamellata Orbigny 1835. Distribution.South America, north to Mxico and the West Indies. Taxonomy.About a dozen species are included in Leptinaria. Nine occur in the study area. Leptinaria lamellata lamellata (Potiez & Michaud 1838)Achatina lamellata Portiez & Michaud 1838; Galerie des Leptinaria lamellata (Potiez & Michaud). Pilsbry 1907; Man. Solem 1960; Nautilus 73:130.Haas 1962:52.Prez & Type Localities. Achatina lamellata : not given. Bulimus unilamellata : last foothills of the Andes in Bolivia, at Petaca, near the Rio Piray, 20 leagues from Santa Cruz de la Sierra. Distribution.The species is widely distributed in South America north through the West Indies and Central America. 2002). BELIZE: Stan Creek Dist.: Stan Creek (Haas & Solem 1960). Leptinaria lamellata concentrica (Reeve 1849)Achatina concentrica Leptinaria lamellata var. concentrica (Reeve). Pilsbry 1907; Man. Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. 82:343.H. B. Baker Achatina lamellosa Moricand, in Reeve 1849; Conch. Con., 5: Jousseaume 1887; Bull. Soc. Zool. De rance, Type Localities. Achatina concentrica : Bolivia.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 165 Achatina lamellose: unknown. : Coca, Ecuador. Distribution.Widely distributed in South America, the West Indies and Central America. PANAM, Taboga island; Isla Pacheco (Pilsbry 1930); Isla San Jos, Archipilago de las Perlas (Morrison 1946; Smiths. Misc. Coll. 106:40). Prov. Pedro; Isla Barro Colorado. Prov. Los Santos: Tonosi. Prov. Panam: Panam; ruins of Old Panam City; near Darin (Pilsbry 1926). Leptinaria unilamellata (Orbigny 1842)Bulimus unilamellata Orbigny 1842; Voyage dans lAmrique Mridionale:257. Leptinaria lamellate (Pfeiffer), Pilsbry 1907; Man. Conch. 18:290. Leptinaria unilamellata (Orbigny). H. B. Baker 1945; Nautilus 58:91.H. B. Baker 1947; Nautilus 61: pl. 1 (reproductive system).Type Locality.Last foothills of the Andes, at Petaca, near the Rio Piray, 20 leagues from Santa Cruz de la Sierra. Distribution.Widely distributed in South America. Leptinaria ambigua Von Martens 1898Leptinaria ambigua Von Martens 1898; Biol. Cent. Amer.:321; pl. Type Locality.Not selected. Viejo; La Palma (Von Martens 1898). Von Martens 1898 Von Martens 1898; Biol. Cent. Amer.:322; pl. Distribution.COSTA RICA, Prov. Puntarenas: Santa Clara; Golfo Dulce; Turubares (Von Martens 1898). Leptinaria elisae Tristram 1861Leptinaria elisae Tristram 1861; Proc. Zool. Soc. Lond. 29:231.Fischer & Crosse 1877:625.Von Martens 1898; Biol. Cent. 34:52.Goodrich & van der Schalie 1937:29.Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):9.Type Locality.Cobn, Dept. Alta Verapaz, Guatemala. Distribution.GUATEMALA, Dept. Alta Verapaz: Cobn; Chama, 950 ft. alt. Dept. Izabal: Maya Farms, Quirigua, Sierra Cavech (Hinkley 1920). Dept. Petn: Tikal (Basch 1959). Leptinaria emmelinae Tristram 1861Leptinaria emmelinae Tristram 1861; Proc. Zool. Soc. Lond. 35, 36 (shell).Type Locality.Cobn, Dept. Alta Verapaz, Guatemala. Distribution.Known only from the type locality. Leptinaria solida Von Martens 1898Leptinaria solida Von Martens 1898; Biol. Cent. Amer.:321; pl. 18, Type Locality.Not selected. Viejo. Prov. Puntarenas: Alta de Mano Tigre, near Terraba (Von Martens 1898). Leptinaria stolli Von Martens 1898Leptinaria stolli Von Martens 1898; Biol. Cent. Amer.:316; pl. 19, 31 (shell).Type Locality.Retalhuleu, Dept. Retalhuleu, Guatemala. Distribution.Known only from the type locality. Hinkley 1920 Hinkley 1920; Nautilus 33:80.Hinkley Type Locality.Livingston, Dept. Izabal, Guatemala. Lectotype ANSP 107608a (H. B. Baker 1963:217). Distribution.GUATEMALA, Dept. Izabal: Sierra Cavech; Maya Farms, Quirigua; Jocolo (Hinkley 1920b). Genus Ochrodermella Pilsbry 1907Ochrodermella Pilsbry 1907; Man. Conch. (2):18:327.Zilch 1959:351.H. B. Baker 1945:87.Type Species.Leptinaria (Neosubulina) Von Martensi Dall 1900. Distribution.Central America. Taxonomy.Three species are recognized. Ochrodermella cumingiana (Pfeiffer 1849)Tornatellina cumingiana Pfeiffer 1849; Proc. Zool. Soc. Lond. 17:134.Von Martens 1898; Biol. Cent. Amer.:324. Ochroderma cumingiana (Pfeiffer). Ancey 1903; Jour. de Conchyl. 51:102. Ochroderma (Ochrodermella) cumingiana (Pfeiffer). Pilsbry 1907; Type Locality.Real Llejos, Nicaragua. Distribution.COSTA RICA: Isla del Coco. Ochrodermella martensi (Dall 1900)Leptinaria (Neosubulina) martensi Dall 1900; Proc. Acad. Nat. Sci. Ochroderma (Ochrodermella) martensi (Dall 1900). Pilsbry 1290.Type Locality.Isla de Coco, Costa Rica. Distribution.Known only from the type locality. Ochrodermella pittieri (Von Martens 1898)Tornatellina pittieri Von Martens 1898; Gesel. Nat. Freunds zu Berlin 198:157.Von Martens 1901; Biol. Cent. Amer.:640; pl.

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166 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Ochroderma (Ochrodermella) pittieri (Von Martens). Pilsbry Ochrodermella pittieri (Von Martens). H. B. Baker 1945:87. Type Locality.Isla de Coco, Costa Rica. Distribution.Known only from the type locality. Genus Opeas Albers 1850Opeas Albers 1850; Die Heliceen:175.Pilsbry 1906; Man. Conch. 18:123.H. B. Baker 1927c:8. H. B. Baker 1945:85.Type Species.Helix goodalli Mller 1822 (= Bulimus pumilus Pfeiffer 1840). Distribution.Worldwide in tropical, subtropical and many temperate regions of the world. Some species are readily transported through human activities, which account for much of the distribution of the genus. Taxonomy.Numerous species. The exact number is uncertain because many included species are known only from shells, and no doubt anatomical investigations will study area. Opeas adamsi Pilsbry 1906Opeas panamensise (Pilsbry). Pilsbry 1910; Proc. Acad. Nat. Sci. Phila. 62:503. Opeas adamsi (shell).Pilsbry 1926a; Proc. Acad. Nat. Sci. Phila. 78:93.Thompson 1968; Nautilus 81:105.Type Locality.Panam. Distribution.PANAM, Prov. Darin: 2.5 mi.SW of El Real (Thompson 1968). Opeas pumilum (Pfeiffer 1840)Helix goodalli Mller 1822, Ann. Philos., n. s. 3:381. (Not Helix goodalli Frussac 1821). Opeas goodalli (Mller). Pilsbry 1906; Manual of Conchology (2) Turton 1831; Manual of he land and freshwater Bulimus Lamarck 1822). Bulimus pumilus Pfeiffer 1840; Archiv. Naturg.:252. Opeas pumilum (Pfeiffer). Pilsbry 1910; Nautilus 24:31.Hinkley 1920b:43, 45, 48.Pilsbry 1926a; Proc. Acad. Nat. Sci. Phila. (radula).H. B. Baker 1940: Nautilus 58:86.Pilsbry 1946; (shell).Morrison 1946; Smiths. Misc. Coll. 106:41.Prez & Type Locality.Bristol, England (introduced). Distribution.Worldwide in tropical, subtropical and many temperate regions, as for the genus. PANAM, Prov. Los Cascades. Prov. Panam: Panam (Pilsbry 1926a); ruins of Old Panam City (Pilsbry 1930). Isla San Jos, Archipilago Izabal: Sierra Cavech; Esmeralda; Jocolo (Hinkley 1920b). TABASCO: San Juan Bautista (Pilsbry 1906). Opeas rarum Miller 1879Opeas rarum 15:471Type Locality.Guayaquil, Ecuador. Distribution.Guatemala and eastern Mxico (Strebel 1882). NAYARIT: Isla Maria Magdalena, Islas Maras (Dall 1926). Genus Subulina Beck 1837Subulina Beck 1837; Index Molluscorum :76.Pilsbry 1906; Man. Conch. 18:71, 220.H. B. Baker 1927c:2 (anatomy).Zilch 1959:342.Type Species.Helix octona Bruguire 1789. Distribution.Tropical South America. One species, Subulina octona, has been introduced throughout much of the world through human agency. Taxonomy.Only two or three species are recognized. One occurs in the study area. Subulina octona (Bruguire 1792)Bulimus octona Bruguire 1792; Encyclopdie Methodique:325. Subulina octona (Bruguire). Von Martens 1898; Biol. Cent. Amer.:298.Pilsbry 1906; Man. Conch. 18:222.Pilsbry 1926a: Proc. Acad. Nat. Sci. Phila. 78:89.Goodrich & van der Schalie 1937:29.Richards 1938:173.Pilsbry 1946; (jaw).Branson & McCoy 1963; Nautilus 76:107.Thompson 144; map. Achatina trochlea Pfeifer 1842; Symbolae ad historiam heliceorum 2:59. Subulia octona var. strebeli Von Martens 1898; Biol. Cent. Amer.:299.Type Localities. Helix octona: probably South America. The species was already widely disseminated by humans long before it was studied by naturalists. Achatina trochlea : Mxico. Subulina octona var. strebeli : Cd. Campeche, Campeche. Distribution.The species is nearly ubiquitous in the study area at lower elevations. PANAM, Isla San Jos, Archipilago de las Perlas (Morrison 1946; Smiths. Misc. Coll. 106:40). Prov. Bocas de Toro: Bocas del Toro; San San Creek. Prov. Los Santos: Tonosi; Anon; Cd. Panam (Pilsbry 1926a). COSTA RICA, Prov. Cartago: Turrialba, 1926b) Talamanca. Prov. Puntarenas: Puerto Lagarto, 70 m. alt.; Palmar, Rio Grande de Terraba valley; Golfo Dulce; Santo Domingo; Coto (Von Martens 1898). NICARAGUA: HONDURAS, Dept. Islas de la Baha: Isla de Roatn (Richards 1938). El SALVADOR. GUATEMALA, Dept. Alta Verapaz: Cobn. Dept. Huehuetenango: Hacienda

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 167 Helvetia, in upper Cholhuitz; Costa Cuco. Dept. Izabal: Sierra Cavech, 1 mi. back of Cavech Village (Hinkley 1920b); Puerto Barrios (Goodrich & van der Schalie 1937). BELIZE: Belize City. CAMPECHE: Cd. Campeche; Cd. Carmen (Branson & McCoy 1963). CHIAPAS. TABASCO: Teapa; San Juan Bautista; Villahermosa (Branson & McCoy 1963). SAN LUS POTOS: Ro Moctezuma, Tamazunchale Cd. Victoria (Correa-Sandoval & Rodriguez 2002); El Cielo Biosphere Reserve (Correa-Sandoval & Rodriguez 2005). Merida. Subfamily OBELISCINAE Thiele 1931 Distribution.Tropical America. Taxonomy.Five genera are recognized (Zilch 1959). One genus occurs in the study area. Genus Obeliscus Beck 1837Obeliscus Beck 1837; Index Molluscorum. :61.Pilsbry 1907; Man. Conch. 18:240.Zilch 1959:357.Type Species.Helix (Cochlicella ) obeliscus Moricand 1833. Distribution.South America from Brazil and Bolivia north to Panam; Cuba, Hispaniola, and Puerto Rico. Taxonomy.Eight subgenera are recognized, one in the study area. Subgenus Ischnocion Pilsbry 1908Ischnocion Pilsbry 1908; Man. Conch. 18:324.H. B. Baker 1927; Occ. Pap. Mus. Zool. Univ. Mich. (182):6.Zilch 1959:358 359.Type Species.Neosubulina (Ischnocion) triptyx Pilsbry 1908. Distribution.Colombia and southern Panam. Taxonomy.A single species is recognized. Obeliscus (Ischnocion) triptyx (Pilsbry 1908)Neosubulina (Ischnocion) triptyx Pilsbry 1908; Man. Conch. 1323.Thompson 1968; Nautilus 81:105.Type Locality.Colombia. Distribution.PANAM, Prov. Darin: 2.5 mi. SW of El Real (Thompson 1968). Subfamily RUMININAE Wenz 1923 Distribution.Africa and Mediterranean Europe, east to India. Taxonomy.The subfamily includes only a few genera. Genus Rumina Risso 1828Rumina Risso 1826; Histoire naturele LEurope meridionale, 4:79.Pilsbry 1905; Man. Conch. 17:211.Type Species.Helix decollata Linnaeus 1758. Distribution.Native to the Mediterranean region of Europe, Africa and Asia, and introduced elsewhere. Taxonomy.A single species is recognized in Mxico. Rumina decollata (Linnaeus 1758)Helix decollata Linnaeus 1758; Systema Naturae (10):773. Rumina decollata (Linnaeus). Pilsbry 1905; Man. Conch. 17:212 82a, 82b, 82c (shell).Correa-Sandoval et al. 1998:15.CorreaSandoval 1999:9.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):236.Correa-Sandoval & Rodriguez 2005:60.Type Locality.Southern Europe. Distribution.This species is common throughout the arid regions of northern Mxico, although it is often ignored by collectors because of its introduced status. COAHUILA: Parras de La Fuente (UMMZ). NUEVO LEN: Laguna Rodriguez 2005); Iturbide (Correa-Sandoval 1997). SAN numerous localities (Correa-Sandoval & Rodriguez 2002). Superfamily TESTACELLOIDEA Gray 1840 Family SPIRAXIDAE H. B. Baker 1939 B. Baker (1939b, 1941a, 1941b, 1943a). Three subfamilies, representing 263 species and an additional 40 subspecies, are recognized in Mxico and Central America. Most species are local endemism. The majority of the species are known from early colonial history with limited accessibility into many of the regions, and the very small number of malacologists who have worked there. Large areas of the study area remain very poorly surveyed, and undoubtedly the number of spiraxid species will be vastly increased with further biodiversity surveys of the many biotic provinces that comprise Mxico and Central America. Even areas that are relatively well known, such as central Veracruz, yield new taxa with each additional investigation. Subfamily EUGLANDININAE H. B. Baker 1941Euglandinarum H. B. Baker 1941; Nautilus 55:54.Thompson 2010; Revista de Biologia Tropical 58:196.Type Genus.Euglandina Crosse & Fischer 1870. Distribution.Southeastern United States south through Mxico and Central America to Peru, Bolivia, and French Guyana. The subfamily is absent from the West Indies except for coastal islands such as Cozumel, tila, Isla del Maz, etc. genera. Currently 112 species are recognized in Mxico and Central America.

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168 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Genus Euglandina Crosse & Fischer 1870Euglandina Crosse & Fischer, in Fischer & Crosse 1870; Miss. Sci. Mex. I:97.Pilsbry 1907; Man. Conch. 19:175.H. B. Baker 1941; Nautilus 55:54.H. B. Baker 1943; Proc. Acad. Nat. Sci. Phila. 95:71. Glandina Strebel 1875:1.Von Martens 1891; Biol. Cent. Amer.:49.Type Species.Achatina lignaria Reeve 1849 (= Glandina aurata Morelet 1849). Distribution.Southeastern United States and Texas south to Brazil, Peru, and Bolivia. The central radiation of the genus is in Mxico and northern Central America. Taxonomy.Euglandina includes three subgenera. Forty-four species are recognized in Mxico and Central America. Subgenus Euglandina Crosse & Fischer 1870 Distribution.Southeastern United States south to Bolivia, Brazil and Peru. Taxonomy.Twenty-two species and six subspecies occur in the study area. Euglandina (Euglandina) aurata (Morelet 1849)Glandina aurata Morelet 1849; Test. Noviss. I:12.Fischer & Martens 1890; Biol. Cent. Amer.:57. Oleacina aurata (Morelet). Gray 1855:33.Pfeiffer 1859:642.Euglandina aurata (Morelet). Pilsbry 1907; Man. Conch. 19:188.Thompson 1987; Bull. Fla. Stat. Mus. 30: 40. Achatina lignaria Reeve 1849; Conch. Icon., 6, Achatina 27 (shell). Oleacina lignaria (Reeve). Gray 1855:34.Type Locality.Glandina aurata: Woods of Vera Paz, Guatemala. Achatina lignaria : not given. Distribution.GUATEMALA, Dept. Alta Verapaz. Recorded from Cobn, and from mountain forests between Tepn (Tecpn?) and Totonicapan, 8,000,000 ft. alt. Dept. Chimaltenango: Santa Elena, near Tecpn (Thompson 1987). Euglandina (Euglandina) bailyi M. Smith 1950Euglandina bailyi Type Locality.Close to highway, three miles east of Chilpancingo, Guerrero, Mxico. Holotype UF 105927. Distribution.Known only from the type locality. Euglandina (Euglandina) binneyana (Pfeiffer 1845)Achatina (Oleacina) fusiformis Pfeiffer 1845; Proc. Zool. Soc. Lond. 13:75. (not Achatina lubrica var. fusiforme Picard. 1840). Glandina fusiformis (Pfeiffer). Morelet 1852; Jour. de Conchyl. Achatina (Oleacina ) binneyana Pfeiffer 1855; Proc. Zool. Soc. Lond. 23:117.Pfeiffer 1859; Monog. Hel. Viv., 4:638. Glandina fusiformis var.? binneana (Pfeiffer). Von Martens 1891:58. Euglandina binneyana (Pfeiffer). Pilsbry 1907; Man. Conch. 19:188, 320.Type Localities.Not known. Distribution.GUATEMALA, Dept. Alta Verapaz: Cobn. Baja Verapaz: Purula, upper part of the Polochc Valley (Von Martens 1891). Euglandina (Euglandina) cognata (Strebel 1875)Glandina cognata Strebel 1875; Beitrag. Mex. Landund Sssw.Euglandina cognata (Strebel). Pilsbry 1908; Man. Conch. 19:195.Type Locality.Tehuantepec, Oaxaca, Mxico. Distribution.Known only from the type locality. Euglandina (Euglandina) cuneus (Von Martens 1891)Glandina cuneus Von Martens 1891; Biol. Cent. Amer.:56; pl. 3, Euglandina cuneus 18 (shell).Type Locality.Omilteme, Guerrero, Mxico. Distribution.Known only from the type locality. Euglandina (Euglandina) dactylus (Broderip 1832)Achatina dactylus Broderip 1832; Proc. Zool. Soc. Lond. :32. Euglandina dactylus 1 (shell). Glandina striata of Authors 1832.Type Locality.Isla Tumaco, near the Ecuador Colombia border. Distribution.PANAM, Prov. Panam: Rio Puerco. COLOMBIA: near Cartago (all Pilsbry 1926). Euglandina (Euglandina) daudebarti dardebarti (Deshayes 1850)Achatina dardebarti Deshayes 1850; in Ferussacs Hist. Nat. Moll. Terr., ii:183. Glandina audebarti (Deshayes). Fischer & Crosse 1870; Miss. Sci. Oleacina audebarti (Deshayes). Tryon 1885; Man. Conch. 1:41; pl. Euglandina daudebarti (Deshayes). Pilsbry 1908; Man. Conch. 19:195.Type Locality.Not stated. Distribution.COLIMA: Colima (Von Martens 1901). GUERRERO: Venta de Zopilote. OAXACA: Tehuantepec. PUEBLA: Chietla. VERACRUZ: environs of Veracruz; Chiquihuitl (all records from Von Martens 1891). Euglandina (Euglandina) daudebarti amoena (Von Martens 1865)Glandina amoena 8 (shell).Fischer & Crosse 1870; Miss. Sci. Mex. I:114.Glandina audebarti e-h, m-n (shell). Glandina audebarti var. amoena Von Martens 1891; Biol. Cent. Euglandina daudebarti var. amoena (Von Martens). Pilsbry 1908;

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 169Man. Conch. 19:195.Type Locality.Mxico. Distribution.VERACRUZ: environs of Veracruz (Strebel 1875). Euglandina (Euglandina) daudebarti jalapana (Von Martens 1891)Glandina longula var. jalapana Von Martens 1891; Biol. Cent. Euglandina daudebarti var. jalapana (Von Martens). Pilsbry 1908; Type Locality.Jalapa, Veracruz, Mxico. Distribution.VERACRUZ: Misantla (Von Martens 1891). Euglandina (Euglandina) daudebarti miradorensis (Strebel 1878)Glandina audebarti form B. Strebel 1875; Beitrag. Mex. Landund Glandina miradorensis Strebel 1878; Beitrag. Mex. Landund Glandina audebaerti var. miradorensis (Strebel). Von Martens 1891; Biol. Cent. Amer.:63.Type Locality.Mirador, Veracruz, Mxico; 2630 ft. alt. Distribution.Known only from the type locality. Euglandina (Euglandina) gigantea Pilsbry 1926Euglandina sowerbyana form B, Strebel 1875; Beitrag. Mex. LandEuglandina gigantea Pilsbry 1926; Proc. Acad. Nat. Sci., 78: 128; Euglandina gigantea gabbi Pilsbry 1926; Proc. Acad. Nat. Sci., 78: Type Localities. Euglandina gigantea : Salinas Bay, in southwestern Costa Rica. Amended to Bahia de Salinas, near La Cruz, Guancaste Prov., Costa Rica (Thompson 1987). Euglandina gigantea gabbi: no locality given. Distribution.Widely distributed in Costa Rica from near sea-level to 2600 m alt. Also known from Chiriqu Province, Panam. COSTA RICA, Prov. Alajuela: Rio Frio; San Carlos; Turubares, 250 m alt. (Turrucares?). Prov. Cartago: Estrella, near Cartago, 1500 m alt.; Tapant, 1300 m alt.; Nicoya; Tilaran; Cache (Cache). Guanacasta Prov.: Rio Jesus Maria. Prov. Heredia: Sarapiqui (on Sarapiqui); Reventazon). Prov. Puntarenas: Coto, 10 m. alt.; Rincon; 2.5 km NE of Monte Verde; Terraba, 700 m alt. Prov. San Jos: San Jos, 116 m alt. PANAM, Prov. Chiriqu: Boquete; Chiriqu; Volcan de Chiriqu, 3000 ft. alt. Euglandina (Euglandina) huingensis (Pilsbry 1903)Glandina huingensis Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. Euglandina huingensis Pilsbry 1907; Man. Conch. 19:185; pl. 23 Type Locality.Huingo, Michoacn, Mxico. Syntypes ANSP 77179. Distribution.Known only from the type locality. Euglandina (Euglandina) immemorata Pilsbry 1907Euglandina immemorata Pilsbry 1907; Man. Conch. 19:192; pl. 24, 41:685.Type Locality.Texas. Holotype: ANSP 11777. Distribution.Northeastern Mxico. Reported from Texas, but without localized information. NUEVO LEN: Santiago (Correa-Sandoval 1993). Euglandina (Euglandina) indusiata (Pfeiffer 1860)Oleacina indusiata Pfeiffer 1860; Proc. Zool. Soc. Lond. 28:138 139. Glandina indusiata Euglandina indusiata (Pfeiffer). Pilsbry 1907, Man. Conch. 19:185.Type Locality.La Parada, in Oaxaca, Mxico. Distribution.MICHOACN: Angangeo. OAXACA: Oaxaca (Von Martens 1891). Euglandina ( Euglandina) lamyi (Fischer & Chatelet 1903)Glandina lamyi Fischer & Chatelet 1903; Jour. de Conchyl. 51:321; Euglandina lamyi (Fischer & Chatelet). Pilsbry 1908; Man. Conch. & Reguero 1998; Acta Zool. Mex. (73):16.Correa-Sandoval & Rodriguez 2005; Acta Zool. Mex. (86):237.Type Locality.Cardenas, San Luis Potos, Mxico. Distribution.SAN LUIS POTOS: 2 km above Presa (Correa-Sandoval et al. 1998). TAMAULIPAS: La Tapona W) (Correa-Sandoval & Rodriguez 2005). Euglandina (Euglandina) liebmanni (Pfeiffer 1846)Achatina liebmanni Pfeiffer 1846; Zeit. fr Malak.:159. Glandina liebmanni (Pfeiffer). Fischer & Crosse 1870; Miss. Sci. Strebel 1878; Beitrag. Mex. Landund Sssw.-Conch. III:7, Euglandina liebmanni (Pfeiffer).Pilsbry 1908; Man. Conch. 19:195.Pilsbry 1909; Nautilus 22:114. Glandina nympha Fischer & Crosse 1870; Miss. Sci. Mex. I:115; pl. Type Localities. Achatina liebmanni : Mxico. Glandina nympha: Mxico. Distribution.COLIMA (Rolle 1895). GUERRERO: Chilpancingo (Von Martens 1892); Balsas, in limestone crevices above the river, 2000 ft. alt. (Pilsbry 1909). Euglandina (Euglandina) Dall 1908 Dall 1908; Proc. U. S. Nat. Mus. 35:180; pl. 29, Type Locality.Between Bolanos and Guadalajara,

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170 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Jalisco, Mxico. Distribution.Known only from the type locality. Euglandina (Euglandina) michoacanensis (Pilsbry 1899)Glandina michoacanensis Pilsbry 1899:397.Pilsbry 1904; Proc. Martens 1901; Biol. Cent. Amer.:610. Euglandina michoacanensis (Pilsbry). Pilsbry 1907, Man. Conch. Type Locality.Uruapan, Michoacn, Mxico. Holotype ANSP 77181a. Distribution.Known only from the type locality. Euglandina (Euglandina) pan Thompson 1987Euglandina pan Thompson 1987; Bull. Fla. Stat. Mus. 30:38; Type Locality.Finca Las Delicia, near Barillas, Dept. Guatemala, Guatemala. Holotype USNM 487385. Distribution.GUATEMALA: known only from moderate elevations in central and southern Dept. Guatemala. Euglandina (Euglendina) pilsbryi Bartsch 1909Euglandina pilsbryi Bartsch 1909; Proc. U. S. Nat. Mus. 37:322; pl. Sci. Phila. 77:307.Type Locality.Bolanos, Jalisco, Mxico. Holotype USNM 207776. Distribution.Known only from the type locality. Euglandina ( Euglandina) pinicola (Fischer & Crosse 1870)Glandina plicatula var. pinicola Fischer & Crosse 1870; Miss. Sci. Glandina pinicola (Fischer & Crosse). Von Martens 1891:64. Euglandina pinicola Pilsbry 1908:196.Type Locality.Totonicapam [Totonicapn], Dept. Totonicapn, Guatemala. Distribution.Known only from the type locality. Euglandina (Euglandina) radula (Strebel 1875)Glandina radula Strebel 1875; Beitrag. Mex. Landund Sssw.Type Locality.Tehuantepec, Oaxaca, Mxico. Distribution.Known only from the type locality. Euglandina (Euglandina) sowerbyana sowerbyana (Pfeiffer 1846)Achatina (Glandina) sowerbyana Pfeiffer 1846; Proc. Zool. Soc. Lond. 14:32. Glandina sowerbyana (Pfeiffer). Albers 1850:198.Fischer & 1878:34; pls. 15 (anatomy).Von Martens 1890:55. Oleacina sowerbyana (Pfeiffer). Gray 1855:33.Tryon 1885; Man. Euglandina sowerbyana Pilsbry 1907, Man. Conch. 19:186. Glandina lignaria Euglandina sowerbyana sowerbyana (Pfeiffer). Thompson 1987; Type Locality.Totontepc, Oaxaca, Mxico. Syntype in BMNH. Distribution.Known from eastern Mxico in the states of Veracruz and immediately adjacent areas of Oaxaca and Puebla. OAXACA: Tontontepc. PUEBLA: Puerto Morales, near Acultzingo, 2770 m alt. VERACRUZ: Catamaco; Cerro Mano Blanco, near Catamaco; Cerro Chicahuaxtla near Cuantlapan; Jalpan; pine forest near Veracruz; Volcan San Martn; Volcan Tuxla (Volcan San Martn), 1475 m alt.; Misantla; San Jos; San Juan Miachutln; Pacho; Jalapa; Mirador; Cerro Necoxtla between Jalapa and Orizaba, 3000 5000 ft.; Orizaba. (Thompson 1987). Euglandina (Euglandina) sowerbyana estephaniae (Strebel 1875)Glandina sowerbyana form D, Strebel 1875; Beitrag. Mex. LandGlandina estephaniae Strebel 1878; Beitrag. Mex. Landund Euglandina sowerbyana estephaniae (Strebel). Pilsbry 1907; Man. Conch. 19:186.H. B. Baker 1941; Nautilus 55:51.Thompson Type Locality.Miahuatln, between Jalapa and Misantla, Veracruz, Mxico. Distribution.VERACRUZ: known only from the 3400 ft. alt. (H. B. Baker 1941). Euglandina (Euglandina) striata (Mller 1774)Buccinum striatum O. F. Mller 1774; Hist. Verm. 2:149. Glandina striata (Mller). Von Martens 1891; Biol. Cent. Amer.:79 80. Euglandina (Euglandina ) striata (Mller). Pilsbry 1907; Man. Conch. 19:176.Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. Type Locality.Unknown. Distribution.Widely distributed in South America in Colombia, Peru, Bolivia, and Brazil (Von Martens 1891). PANAM: Isla Barro Colorado (Pilsbry 1930). Euglandina ( Euglandina) texasiana texasiana (Pfeiffer 1856)Achatina texasiana Pfeiffer 1856; Novitates Conchologicae 1:82; Glandina texasiana .W. G. Binney 1859; Boston Jour. Nat. Hist. Oleacina texasiana. (shell). Euglandina texasiana .Pilsbry 1907; Man. Conch. 19:190; Trop. 41:685.Correa-Sandoval, Garca-Cubas & Reguero 1998:16.Correa-Sandoval 2000; Acta Zool. Mex. (79):9.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):237.Type Locality.Texas. Distribution.Northeastern Mxico and extreme southern Texas. NUEVO LEN: Santiago (Correa-Sandoval 1993). SAN LUS POTOS: El Abra, Valles (Pilsbry

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 171 1907); numerous localities (Correa-Sandoval et al. 1998). TAMAULIPAS: Tampico; numerous localities (CorreaSandoval & Rodriguez 2002). Euglandina (Euglandina) texasiana angustior Pilsbry & Vanatta 1936Euglandina texasiana angustior Pilsbry & Vanatta 1936:97; pl. 7, Type Locality.Gonzalez, Tamaulipas, Mxico. Holotype ANSP 162640. Distribution.Southeastern TAMAULIPAS and adjacent SAN LUS POTOS. Euglandina (Euglandina) titan Thompson 1987Euglandina titan Thompson 1987; Bull. Fla. St. Mus. 30:42; Type Locality.Montaas del Mico, 4 km WSW of Puerto Santo Toms at 800 m elevation in a tropical rainforest, Dept. Izabel, Guatemala. Holotype UF 35307. Distribution.Known only from the type locality. Euglandina (Euglandina) (Lea 1834) Lea 1832; Trans. Amer. Philos. Soc. 5:84. Achatina coronata Pfeiffer 1846; Zeit. fr Malak. 158:158. Glandina coulteri Gray, in Pfeiffer, Monogr. Helic. Viv. 1:642. Glandina uhdeana Gray. Von Martens 1863; in Monatsber. Akad. Wiss. Berl. 26:540. Glandina guttata Crosse & Fischer 1869; Jour. de Conchyl. 17:250. Tryon 1866; Amer. Jour. Conch. 2:226, pl. 16, ). (Lea). Pilsbry 1907, Man. Conch. 19:185.H. B. Baker 1943; Proc. Acad. Nat. Sci. Phila. 95:10 Type Localities.: Mxico. Glandina coulteri: no locality given. Achatina coronata: Mxico. Glandina uhdeana: Veracruz, Mxico. Glandina guttata : Mxico, near Puebla, Mxico. GUERRERO: Omilteme. HIDALGO: Zimapn. MXICO: Volcn de Mexicalcingo; Pirmides; Teotihuacn; Guajimalpa. OAXACA: El Punto; Cuicatln; Huajuapn turnoff to Nacaltepc; 0.7 mi. W of Tlapacoyn, 5000 ft. alt.; Oaxaca; Juquila. PUEBLA:3.6 mi. SW of Chapulco; 8.1 mi. SW Izucar de Matamoros,4800 ft. alt; Hwy. 150, 1.5 mi. SW of Veracruz-Puebla border; Cd. Puebla, Mxico; Tecamachalco, 7500 m alt.; 1.1 mi. N of Tecamachalco; Tehuacn; Tepeaca. Adjacent area at altitudes of 1285 m. VERACRUZ: Guajimalpa; Teotihuacan; Jalapa (all records from Thompson 1987). Subgenus Singleya H. B. Baker 1941Singleya H. B. Baker 1941:54.Type Species.By original designation: Glandina singleyana (W. G. Binney 1892). Distribution.Southern Texas south to Panam. Taxonomy.Twenty-one species and four subspecies are recognized. Cosmomenus, which is treated as a separate subgenus, was described as a section of Singleya on the basis of the origin of the penis retractor muscle. The penis retractor in Singleya originates on the inner wall of the lung, while in Cosmomenus it originates on the left side of the columellar retractor muscle. The assignment of species to either subgenus on the basis of shell characters is arbitrary. Most of the following species are known only on the basis of the shell. They are place in Singleya because it is the senior name. Euglandina (Singleya) anomala anomala (Angas 1879)Glandina (Oleacina) anomala Angas 1879; Proc. Zool. Soc. Lond. Euglandina anomala anomala (Angas). Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. 82:344.Type Locality.Hilly region of Costa Rica. record. Euglandina (Singleya) anomala barrocoloradensis Pilsbry 1930Euglandina anomala barrocoloradensis Pilsbry 1930; Proc. Acad. Type Locality.Isla Barro Colorado, Panam. Holotype ANSP 151391. Distribution.Known only from the type locality. Euglandina (Singleya) balesi Pilsbry 1938Euglandina balesi Pilsbry 1938; Nautilus 52:16; Nautilus 53: pl. 2, Type Locality.Puerto Marques, south of Acapulco, [Sinaloa], Mxico. Holotype ANSP 170440. Distribution.Known only from the type locality. Euglandina ( Singleya) candida candida (Shuttleworth 1852)Achatina (Polyphema ) candida Shuttleworth 1852; Mitt. Naturf. Euglandina candida (Shuttleworth). Pilsbry 1908; Man. Conch. Euglandina (Singleya ) candida (Shuttleworth). H. B. Baker 1941; Nautilus 55:60. Glandina simplex Strebel 1875; Beitrag. Mex. Landund Sssw.Type Localities. Achatina candida: Mxico. Syntype

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172 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Naturhistorische Museum Bern 18951/1 (Neubert & Gosteli 2003). Glandina simplex : Oaxaca, Mxico. Distribution.OAXACA. VERACRUZ: Atoyac, 1300 1450 ft. alt. (H. B. Baker 1941). Euglandina (Singleya) candida conularis (Pfeiffer 1855)Achatina (Glandina) conularis Pfeiffer 1855; Proc. Zool. Soc. Lond. 23:100. Glandina conularis (Pfeiffer). Von Martens 1891; Biol. Cent. Euglandina conularis (Pfeiffer). Pilsbry 1908; Man. Conch. 19:197 Euglandina (Singleya) condida var. conularia (Pfeiffer). H. B. Baker 1941; Nautilus 55:60.Type Locality.Mxico. 3000 ft. alt. (H. B. Baker 1941). Euglandina (Singleya) carminensis (Morelet 1849)Glandina carminensis Morelet 1849; Test. Noviss. I:14.Morelet Martens 1891:61.Fiscer & Crosse 1870:109.Strebel 1875:37; Euglandina carmenensis (Morelet). Pilsbry 1908; Man. Conch. 19:195.Bequaert & Clench 1933; Pub. Carnegie Inst. Wash. (431):528.Thompson 1967; Bull. Fla. St. Mus. 11:232.Type Locality.Isla de Carmen, Campeche, Mxico. Distribution.CAMPECHE: 7.1 mi. SW of Campeche; 2.2 mi. S of Pixtn (Thompson 1967). YUCATN: Chankom (Bequaert & Clench 1933). GUATEMALA, Dept. Quetzaltenango: Cerro Zunil, near Quetzaltenango (Von Martens 1891). HONDURAS, Dept. Islas de la Baha: Isla de tila (Von Martens 1901). Euglandina (Singleya) corneola (W. G. Binney 1857)Glandina corneola W. G. Binney 1857, Proc. Acad. Nat. Sci. Phila.:189. Euglandina corneolaa (Binney). Pilsbry 1908; Man. Conch. Garca-Cubas & Reguero 1998:16.Correa-Sandoval 2000; Acta Zool. Mex. (79):9.Type Locality.Not stated. Distribution.SAN LUIS POTOS: Valles (Pilsbry 1908); numerous localities (Correa-Sandoval et al. 1998). Euglandina (Singleya) decussata (Deshayes 1840)Achatina decussata Deshayes 1840, in Ferussacs His. Nat. Moll. Glandina decussata (Deshayes). Fischer & Crosse 1870; Miss. Sci. Mex. I:112.Strebel 1875; Beitrag. Mex. Landund Sssw.Biol. Cent. Amer.:58. Euglandina decussata (Deshayes). Pilsbry 1908; Man. Conch. 19:188.Hinkley 1920; Nautilus 34:39, 43, 48, 51.H. B. Baker 1923; Occ. Pap. Mus. Zool. Univ. Mich. (135):8. Euglandina (Singleya) decussata (Deshayes). H. B. Baker 1943; Type Locality.Not stated. Distribution.GUATEMALA, Dept. Alta Verapaz: Cobn; Chama (Hinkley 1920). Dept. Baja Verapaz: Senahu; Panzos (Von Martens 1891). Dept. Izabal: mountains of Cavech, back of Cavech Village; Jocolo (Hinkley 1920). Dept. Quirigua: Maya Farms (Hinkley 1920). VERACRUZ: Cuatotolapam (H. B. Baker 1923). Euglandina (Singleya) (Von Martens 1891) Von Martens 1891; Biol. Cent. Amer.:67; pl. 4, (Von Martens).Pilsbry 1908; Man. Conch. Acad. Nat. Sci. Phila. 77:307.Type Locality.Mazatln, Sinaloa (?). Distribution.SINALOA (?). Euglandina (Singleya) ghiesbreghti (Pfeiffer 1856)Oleacina ghiesbreghti Pfeiffer 1856; Malak. Bltt, 3:235. Glandina ghiesbreghti (Pfeiffer). Strebel 1875; Beitrag. Mex. LandMartens 1891; Biol. Cent. Amer.:58. Euglandina ghiesbreghti (Pfeiffer). Pilsbry 1907; Man. Conch. 19:188.Bequaert 1957:218.Type Locality.Chiapas, Mxico. Distribution.CHIAPAS: Laguna Ocotal, 950 m alt.; El Censo to Laguna Ocotal, 700 m alt. (Bequaert 1957). TABASCO: San Juan Bautista (Von Martens 1901). Euglandina (Singleya) hererrae (Contreras 1923)Glandina hererrae Contreras 1923; Boletin Soc. Estud. Biol. 1:13. Euglandina hererrae (Contreras).Pilsbry and Cockerell 1925; Proc. Acad. Nat. Sci. Phila. 77:307.Type Locality.Chapala, Jalisco, Mxico. Distribution.Known only from the type locality. Euglandina (Singleya) insignis (Pfeiffer 1855)Achatina (Glandina) insignis Pfeiffer 1855; Proc. Zool. Soc. Lond. Glandina insignis (Pfeiffer). Fischer & Crosse 1870; Miss. Sci. Oleacina insignis (Pfeiffer). Tryon 1885, Man. Conch. 1:35; pl. 7, Glandina liebamnni var. insignis (Pfeiffer). Von Martens 1891:62. Euglandina insignis (Pfeiffer). Pilsbry 1908; Man. Conch. 19:195.Pilsbry and Cockerell 1925; Proc. Acad. Nat. Sci. Phila. 77:307.Type Locality.Unknown. Distribution.NAYARIT: San Blas; Tepc (Von Martens 1891); Islas Maras (Pilsbry & Cockerell 1925). Euglandina (Singleya) jacksoni Pilsbry & Vanatta 1936Euglandina jacksoni Pilsbry & Vanatta 1936, Nautilus 49:97; pl. 7, (86:237.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 173 Type Locality.Gonzalez, Tamaulipas, Mxico. Holotype: ANSP 162639. Distribution.TAMAULIPAS: Gonzalez (Pilsbry & Vanatta 1937);El Moro, Hyw Cd. Victoria-Soto la Marina Rodriguez 2002). Euglandina (Singleya) longula (Fischer & Crosse 1870)Glandina longula Fischer & Crosse 1870; Miss. Sci. Mex. I:111; pl. Euglandina longula (Fischer & Crosse). Pilsbry 1908; Man. Conch. 19:196.Type Locality.Rio San Juan, northeast Mxico. Distribution.Unknown. The precise type locality has not been determined. Euglandina (Singleya) lowei Pilsbry 1931Euglandina lowei (shell).Type Locality.Isla Mara Madre, Islas Maras, Nayarit, Mxico. Holotype ANSP 151320. Distribution.Known only from the type locality. Euglandina (Singleya) mazatlanica mazatlanica (Von Martens 1891)Glandina mazatlnica Von Martens 1891; Biol. Cent. Amer.:65; pl. Euglandina mazatlnica (Von Martens). Pilsbry 1908; Man. Conch. 1925; Proc. Acad. Nat. Sci. Phila. 77:307.Dall 1926; Proc. Calif Acad. Sci. ser. 4, 15:470.Type Locality.Mazatln, Sinaloa, Mxico. Syntype Berlin Museum 35386. Distribution.NAYARIT: Islas Maras (Dall 1926). SINALOA: Mazatln. Euglandina (Singleya) (Von Martens 1891)Glandina mazatlanica var. Von Martens 1891; Biol. (Von Martens). Pilsbry 1908: Type Locality.Islas Maras, Nayarit, Mxico. Distribution.Known only from the type locality. Euglandina (Singleya) pseudoturris (Strebel 1875)Glandina pseudoturris Strebel 1875; Beitrag. Mex. Landund Euglandina pseudoturris (Strebel). Pilsbry 1908; Man. Conch. 19:197.Type Locality.Juquila, Oaxaca, Mxico. Distribution.GUERRERO: Tiera Colorada. OAXACA: Juquila. Euglandina (Singleya) singleyana (W. G. Binney 1892)Glandina decussata Deshayes. W. G. Binney 1878; Terr. Moll. N. Amer. 5:86. Glandina singleyana W. G. Binney 1892; 4th Suppl. Bull. Mus. Euglandina singleyana. Pilsbry and Ferriss 1906; Proc. Acad. Nat. Sci. Phila. 58:143.Pilsbry 1907; Man. Conch. 19:189; pl. 24, (18):12 (eggs).H. B. Baker 1943; Proc. Acad. Nat. Sci. Phila. Type Locality.San Antonio, Bexar County, Texas. Distribution.Southern Texas and adjacent Tamaulipas, Mxico. TAMAULIPAS. Euglandina (Singleya) tenella (Strebel 1875)Glandina tenella Strebel 1875; Beitrag. Mex. Landund Sssw.Biol. Cent. Amer.:59. Euglandina tenella (Strebel). Pilsbry 1907; Man. Conch. 19:188.Type Locality.First woods 0.5 hour from Veracruz [City], Veracruz, Mxico. Distribution.VERACRUZ: known only from the vicinity of Cd. Veracruz. Euglandina (Singleya) turris turris (Pfeiffer 1846)Achatina (Glandina) turris Pfeiffer 1846; Symb. Hist. Helic. 3:91. Glandina turris (Pfeiffer). W. G. Binney 1869; Land and Freshw. Euglandina turris (Pfeiffer).Pilsbry and Cockerell 1926; Proc. Acad. Nat. Sci. Phila. 77:308.Type Locality.Mxico. Distribution.SINALOA: Mazatln (Von Martens 1891). Euglandina (Singleya) turris longurio Pilsbry & Cockerell 1926Euglandina turris longurio Pilsbry and Cockerell 1926; Proc. Acad. Type Locality.Mazatln, Sinaloa, Mxico. Holotype ANSP 24797. Distribution.JALISCO: Guadalajara. NAYARIT: Islas Maras. SINALOA: Mazatln. Euglandina (Singleya) wani (Jacobson 1968)Streptostyla (Chersomitra) wani Jacobson 1968; Nautilus 81:118; Type Locality.Near Wani (Huani), Nicaragua. Holotype USNM 426028. Distribution.Known only from the type locality. Subgenus Cosmomenus H. B. Baker 1941Cosmomenus H. B. Baker 1941; Nautilus 55:54.Type Species.Glandina cumingi Beck 1837. Distribution.Yucatn to Venezuela. Taxonomy.Two species are recognized as belonging to Cosmomenus on the basis of the origin of the penis retractor muscle.

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174 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Euglandina (Cosmomenus) cumingi (Beck 1837)Glandina cumingi Beck 1837; Index Moll.:78.Von Martens 1891; Euglandina cumingi (Beck). Pilsbry 1908; Man. Conch. 19:195.Richards 1938:172.H. B. Baker 1943; Proc. Acad. Nat. (radula).Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):8.Jacobson 1968; Amer. Mus. Novit. (1899):117.Prez & Lopz 2002:151. Glandina petiti (Deshayes). Strebel 1875:40. Glandina rosea (Frussac). Fischer & Crosse 1870; Miss. Sci. Mex. I:107 (in part Type Locality.Not stated. Distribution.TABASCO: Teapa (Von Martens 1891). COSTA RICA: San Jos (Von Martens 1891); Alto de Mano Tigre, near Terraba, 600 m alt. (Von Martens 1901). GUATEMALA, Dept. Retalhuleu: Retahluleu; El Reposo, 800 ft. alt.; Costa Cuca, 2500 ft. alt.; Cholluitz, on the slope of Volcn de Santa Maria (Von Martens 1891). Dept. Petn: Tikal National Park (Basch 1959). HONDURAS, Dept. Islas de la Baha: between Coxen Hole and French Harbout, Isla de Roatn (Richards 1938). NICARAGUA: numerous localities Bonanza, Moravian Mission (Jacobson 1968). PANAM, Prov. Chiriqu: Chiriqu. Prov. Panam: Cd. Panam; between Tabernilla and San Pablo. Prov. Los Santos: Tonos (Pilsbry 1926). Canal Zone: Isla Barro Colorado (Pilsbry 1930). VENEZUELA: La Fria (H. B. Baker 1943). Euglandina (Cosmomenus) cylindracea (Phillips 1846)Glandina cylindracea Phillips 1846; Proc. Acad. Nat. Sci. Phila. Euglandina cylindracea (Phillips). Pilsbry 1908; Man. Conch 19:198.Bequaert & Clench 1933; Pub. Carnegie Inst. Wash. (431):528.Richards 1937; Proc. Amer. Philos. Soc. 77:251.Thompson 1967; Bull. Fla. St. Mus. 11:232. Euglandina (Cosmomenus) cylindracea (Phillips). H. B. Baker 1941; Nautilus 55:60.H. B. Baker 1943; Proc. Acad. Nat. Sci. Achatina largillierti Pfeiffer 1846; Symbolae Hist. Helic., 3:90. Glandina largillierti (Pfeiffer). Fischer & Crosse 1870:116.Von Achatina yucatanensis Pfeiffer 1846; Symbolae Hist. Helic. 3:92. Glandina yucatanensis Achatina carnea Pfeiffer 1854; Proc. Zool. Soc. Lond. (1852):157.Type Localities. Glandina cylindracea : Yucatn, Mxico. Achatina largillierti : Yucatn, Mxico. Achatina yucatanensis : Yucatn, Mxico. Achatina carnea: Central America. Distribution.CAMPECHE: 7.2 mi. S of Pixtn; 2.2 mi. S of Pixtn; Cd. d Carmin; 6.1 mi. SW of Sebaplaya; 6.2 mi. SE of Sebaplaya; 7.1 mi. SW of Campeche; 5.7 mi. E of Campeche; 3.6 mi. S of Hopelchen; 5.1 mi. NNW of Dzilbachn; 5.1 mi. W of Tikinml; 10.2 mi. E of Escarcega (Thompson 1967). QUINTANA ROO: San Miguel, Isla Cozumel (Richards 1937); Chetumal (Haas & Solem 1960); 7.1 mi. NNW of Xiatil (Thompson 1967). YUCATN: Chichen Itza; Chankom; Dzitas; Temax; Progreso; cave of Actun Lara, near Ticul; Actun Has, Hacienda Yocat (Bequaert & Clench 1933); 0.8 mi. NE of Becanchn; Uxmal; 19.1 mi. SSE of Uxmal; 7.0 mi. SSE of Uman; Progreso (Thompson 1967). Genus Guillarmodia H. B. Baker 1941Guillarmodia H. B. Baker 1941; Nautilus 55:54, 57.Thompson 1995; Bull. Fla. Mus. Nat. Hist. 39:54.Type Species.Euglandina pupa H. B. Baker 1941). Distribution.Mxico in general with one species from Costa Rica. Taxonomy.Guillarmodia was previously treated as a subgenus of Euglandina. Its distinct shell characters and reproductive anatomy justify recognition as a separate genus, with two subgenera. Forty-one species and seven subspecies are recognized within the genus. Subgenus Guillarmodia H. B. Baker 1941 Distribution.Mxico, generally from states along the Taxonomy.Fourteen species are recognized. Guillarmodia (Guillarmodia) arthritica (Thompson 1995)Euglandina arthritica Thompson 1995; Bull. Fla. Mus. Nat. Hist. Type Locality.An oak-forested limestone hillside 7.5 km ENE of Ixcateopn, Guerrero, Mxico; 2650 m alt. Holotype UF 34666. Distribution.Known only from the type locality. Guillarmodia ( Guillarmodia) brachystyla (Thompson 1995)Euglandina (Guillarmodia) brachystyla Thompson 1995; Bull. Fla. Type Locality.A small karst limestone knoll 2 km east of Punta Troncones, Guerrero, Mxico; 60 m alt. Holotype UF 193587. Distribution.Known only from the type locality. Guillarmodia (Guillarmodia) comma (Thompson 1995)Euglandina comma Thompson 1995; Bull. Fla. Mus. Nat. Hist. Type Locality.Southeast side of the Rio Ixtapa, 6 km W); 20 m alt. Pantla is a small village on the coastal highway 23 km NW of Zihuatanejo. The type locality is in a lowland mesic forest overlying a substrata consisting of lateritic soils and limestone. Holotype UF 34663. Distribution.Found along coast areas of northwestern Guerrero, Mxico. GUERRERO: 2 km SE of Pantla, 30 m alt.; 3.5 km NW of Zihuatanejo, 130 m alt.; 4.5 km NW of Zihatanejo, 100 m alt.; limestone hill 3 km E of Naranjillo; 2

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 175 W), 130 m alt. Guillarmodia (Guillarmodia) cymatophora (Pilsbry 1910)Euglandina cymatophora Pilsbry 1910; Proc. Acad. Nat. Sci. Phila. Type Locality.Side of the canyon below Los Canoas, San Luis Potos, Mxico. Holotype ANSP 98612. Distribution.Known only from the type locality. Guillarmodia (Guillarmodia) dorsalis (Thompson 1963)Euglandina (Guillarmodia) dorsalis 3, 4.Type Locality.One mile north of Pomero, Michoacn, Mxico, 700 ft alt. Holotype UMMZ 213222. Distribution.Known only from the type locality. Guillarmodia (Guillarmodia) elegans (Von Martens 1895)Salasiella elegans Von Martens in Rolle 1895; Von Martens 1895; Euglandina (Guillarmodia) elegans (Von Martens). Thompson Type Locality.Colima, Mxico. Lectotype Berlin Museum 47661 (Thompson 1963). Distribution.Known only from the type locality. Guillarmodia (Guillarmodia) gracilior (Thompson 1995)Euglandina gracilior Thompson 1995; Bull. Fla. Mus. Nat. Hist. Type Locality.An open oak forest 12 km by road substrata with clusters of palms, shrubs and sp. growing among the oaks. Holotype UF 194115. Distribution.GUERRERO: known only from intermediate altitudes along a limestone ridge southwest of 1285 m altitude that is dominated by sparse growths of oaks (Quercus sp.); 10 km SW of Xochilapa, 1700 m alt.; 11.5 km SW of Xochilapa, 1750 m alt.; 12 km by road SW of Xochilapa, 1700 m alt.; 15 km SW of Xochilapa, 2200 m alt.; 1485 m alt. Guillarmodia (Guillarmodia) kingi (Thompson 1995)Euglandina kingi Thompson 1995; Bull. Fla. Mus. Nat. Hist. 39:62 Type Locality.Among boulders in an open semi-xeric scrub forest on a limestone hillside, on the west side of the Rio Tehuntepec, 25 km NW of the Presa Benito Juarz (dam), Oaxaca, Mxico; 300 m alt. Holotype UF 34691. OAXACA:30 km NW, 3 km NE of Tehuntepec, 200 m alt.; 13.5 km NW of Tehuntepec, 150 m alt.; 24 km NW, 3 km NE of Tehuntepec, 210 m alt.; 3.5 km NW of Mixtequilla, N of La Ventosa, 210 m alt.; limestone hill 10 km E of La Ventosa, 50 m alt.; 7.5 km N of La Ventosa, 100 m alt.; 15 km W); limestone mtn. 3 km W of Santiago Astata, 100 m alt. Guillarmodia (Guillarmodia) mariana (Dall 1926)Euglandina mariana Dall 1926; Proc. Calif. Acad. Sci., ser. 4 Type Locality.Isla Maria Magdelena, Islas Maras, Nayarit, Mxico. Holotype CAS 2190. Distribution.Known only from the type locality. Guillarmodia (Guillarmodia) minuta (Pilsbry 1910)Streptostyla minuta Pilsbry 1910; Proc. Acad. Nat. Sci. Phila. Type Locality.Mountain sides of canyon below Los Canoas, San Luis Potos, Mxico. Holotype ANSP 98595. Distribution.Known only from the type locality. Taxonomy.See Streptostyla minuta. The general structure of the shell indicates a closer relationship with Guillarmodia than with Streptostyla Guillarmodia (Guillarmodia) multispira (Pfeiffer 1861)Oleacina multispira Pfeiffer 1861; Novitat. Conch. 2:163; pl. 44, Glandina multiapira (Pfeiffer). Tryon 1885; Man. Conch. 1:30; pl. Euglandina multispira (Pfeiffer). Pilsbry 1908; Man. Conch. 19:207.Type Locality.Juquila, Oaxaca, Mxico. Distribution.Known only from the type locality. Guillarmodia (Guillarmodia) nelsoni (Bartsch 1909)Euglandina nelsoni Bartsch 1909; Proc, U. S. Nat. Mus. 37:321; pl. Nat. Sci. Phila. 77:307.Type Locality.Acaponeta, Tepc [Nayarit], Mxico. Syntypes USNM 207784. Distribution.Known only from the type locality. Guillarmodia (Guillarmodia) pupa (H. B. Baker 1941)Euglandina (Guillarmodia) pupa H. B. Baker 1941; Nautilus 55:57; Type Locality.Atoyc, Veracruz, Mxico; 1300 ft. alt. Holotype in the UMMZ. Distribution.Eastern Mxico in central Veracruz and immediately adjacent Oaxaca; 100 m alt. OAXACA: limestone range 1 km W of Cedral, 100 m alt.; 4 km NW of Temascal, 100 m alt.; 3 km S of Acatln, 100 m alt.; 4 km SW

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176 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) of Acatln, 100 m alt. VERACRUZ:3 km NE of Atoyc, 640 m alt.; 1 km NW of Atoyaquillas, ca. 7 km NNW of Paraje alt.; Cerro de Las Palmas, 1 km N of San Mateo, ca. 3 km NE of Comalapa, 400 m alt.; 6 km NE of Comalapa, 250 m alt. Guillarmodia (Guillarmodia) pygmaea (Pilsbry & Vanatta 1936)Euglandina pygmaea Pilsbry & Vanatta 1936. Nautilus 49:98; pl. 7, Type Locality.Gonzalez, Tamaulipas, Mxico. Holotype ANSP 162641. Distribution.Known only from the type locality. Guillarmodia ( Guillarmodia) stenotrema (Thompson 1995)Euglandina stenotrema Thompson 1995; Bull. Fla. Mus. Nat. Hist. Type Locality.A limestone hillside 4.5 km ENE of Ixcateopn, Guererro; 2500 m alt. The area is forested with a dense growth of junipers among limestone boulders. km WSW of Taxco. Holotype UF 193042. terrain in northern Guerrero, Mxico, north of the Rio Balsas from 1100 m alt GUERRERO: ca 10 km NE of Chapa, Ixcateopn, 2250 m alt.; 2 km ENE of Ixcateopn, 2100 m 1650 m alt.; 5 km E of Teloloapn 1850 m alt.; 25 km E of Teloloapn 1850 m alt.; 18 km N of Tonalapa; 1100 m alt.; 9 km S of Buenavista de Cuellar, 1350 m alt.; ca. 20 km NNW Subgenus Proameria H. B. Baker 1941Subgenus Proameria H. B. Baker 1941:54, 57.Type Species.Euglandina saxitilis saxitilis H. B. Baker 1941. Distribution.Mxico in general south to Oaxaca and Veracruz. One species is recorded from Costa Rica. A species from Peru, Euglandina haasi Thompson 1982, also appears to belong here. Taxonomy.Twenty-seven species and seven subspecies are recognized. Guillarmodia (Proameria) albersi albersi (Pfeiffer 1854)Achatina (Glandina) albersi Pfeiffer 1854; Proc. Zool. Soc. Lond. 22:295. Glandina albersi (Pfeiffer). Fischer & Crosse 1870; Miss. Sci. Mex. Euglandina albersi (Pfeiffer). Pilsbry 1908; Man. Conch. 19:201.Pilsbry and Cockerell 1926; Proc. Acad. Nat. Sci. Phila. 77:307.Dall 1926; Proc. Calif. Acad. Sci., ser. 4 15:470.Type Locality.Gulf of California. Distribution.COLIMA: Sierra Madre de Colima (Von Martens 1891). NAYARIT: Tepc; Islas Maras (Dall 1926). SINALOA: Mazatln (Von Martens 1891). Guillarmodia (Proameria) albersi infanta (Von Martens 1891)Glandina albersi var. infanta Von Martens 1891; Biol. Cent. Euglandina albersi var. infanta (Von Martens). Pilsbry 1908; Man. Conch. 19:201.Type Locality.Tepc, Nayarit, Mxico. Distribution.Known only from the type locality. Guillarmodia (Proameria) alticola (Pilsbry 1903) Pilsbry 1903; Proc. Acad. Nat. Sci. Euglandina alticola (Pilsbry). Pilsbry 1908; Man. Conch. 19:194 Correa-Sandoval 1993; Rv. Biol. Trop., 41:685. Mxico. Syntypes ANSP 77174. Distribution.NUEVO LEN: Santiago (CorreaSandoval 1993). Guillarmodia (Proameria) attenuata (Pfeiffer 1851)Achatina (Glandina) attenuata Pfeiffer 1851; Proc. Zool. Soc. Lond. 19:259.Pfeiffer, in Martini & Chemnitz, Syst. Conch. Cab., ed. 2: Achatina Glandina attenuata (Pfeiffer). Tryon 1885; Man. Conch. 1:22; pl. Euglandina attenuata (Pfeiffer). Pilsbry 1908; Man. Conch. 19:208.Type Locality.Central America. Distribution.Unknown. This species has not been found since its original discovery. Guillarmodia ( Proameria) bellula (Crosse & Fischer 1869)Glandina bellula Crosse & Fischer 1869; Jour. de Conchyl. 17:425.Euglandina bellula (Crosse & Fischer). Pilsbry 1908; Man. Conch. 19:207.Type Locality.Mxico. Distribution.Unknown. Guillarmodia (Proameria) chasonae Pilsbry & Cockerell 1926Euglandina chasonae Pislbry and Cockerell 1926; Proc. Acad. Nat. Type Locality.Tepc, Nayarit, Mxico. Holotype ANSP 139942. Distribution.Known only from the type locality. Guillarmodia ( Proameria) conferta conferta (Pfeiffer 1861)Oleacina conferta Pfeiffer 1861; Proc. Zool. Soc. Lond. 29:26. Glandina conferta (Pfeiffer). Fischer & Crosse 1870:131 (in part).

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 177 Euglandina conferta (Pfeiffer). Pilsbry 1908; Man. Conch. 19:210.Type Locality.Juquila, Oaxaca, Mxico. Distribution.Known only from the type locality. VERACRUZ: Huatusco; Jalapa; Mirador (Von Martens 1891). Guillarmodia (Proameria) conferta crossei (Von Martens 1891)Glandina conferta (Pfeiffer). Fischer & Crosse 1870; Miss. Sci. Glandina conferta var. crossei Von Martens 1891; Biol. Cent. Amer.:71.Type Locality.Orizaba(?), Veracruz, Mxico. Distribution.VERACRUZ(?). Guillarmodia (Proameria)(Pfeiffer 1856)Achatina (Varicella ) Lond. 24:321. (Pfeiffer). Fischer & Crosse 1870; Miss. Sci. (Pfeiffer). Pilsbry 1908; Man. Conch. 19:209. Euglandina (Proameria) (Pfeiffer). H. B. Baker 1941; Nautilus 55:60.H. B. Baker 1943; Proc. Acad. Nat. Sci. Phila. 95:8. 1875); Sumidero, 3400 ft. alt. (H. B. Baker 1941). Guillarmodia (Proameria) dalli (Pilsbry 1899)Glandina dalli Pilsbry 1899; Proc. Acad. Nat. Sci. Phila. 51:396.5a (shell). Euglandina dalli (Pilsbry). Pilsbry 1908; Man. Conch. 19:207; Trop. 41:685.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (73):237. Mxico. Syntypes ANSP 77173. Distribution.NUEVO LEN: Diente; Santiago (Correa-Sandoval 1993). TAMAULIPAS: numerous localities in southern part of state (Correa-Sandoal & Castro 2002). Guillarmodia (Proameria) delicata Pilsbry 1903Glandina delicata Pilsbry 1903; Proc. Acad. Nat. Sci. Phila. 55:771 Euglandina delicata (Pilsbry). Pilsbry 1908; Man. Conch. 19:194; Type Locality.Canyon 4 miles west of [Ciudad] Victoria, Tamaulipas, Mxico. Syntypes ANSP 85917. Distribution.Known only from the type locality. Guillarmodia (Proameria) delicatula delicatula (Shuttleworth 1852)Achatina (Polyphemus) delicatula Shuttleworth 1852; Mitt. Naturf. Ges. Bern 1852:202. Gandina delicatula (Shuttleworth). Fischer & Crosse 1870:92.Von Euglandina delicatula (Shuttleworth). Pilsbry 1908; Man. Conch. Syntype: Naturhistorische Museum Bern 18844/1 (Neubert and Gosteli 2003). (Von Martens 1891). Guillarmodia (Proameria) delicatula major (Von Martens 1891)Glandina delicatula var. major Von Martens 1891; Biol. Cent. Euglandina delicatula var. major (Von Martens). Pilsbry 1908; Type Locality.Coatepec, Veracruz, Mxico. Distribution.Known only from the type locality. Guillarmodia (Proameria) H. B. Baker 1941Euglandina (Proameria) H. B. Baker 1941; Type Locality.Above Necaxa, Puebla, Mxico; 4925 ft. alt. Holotype in the UMMZ. Distribution.Known only from the type locality. Guillarmodia (Proameria) (Pfeiffer 1855)Achatina (Glandina) Pfeiffer 1855; Proc. Zool. Soc. Lond. 23:100. (Pfeiffer). Pilsbry 1908; Man. Conch. 19:200; pl. Type Locality.Orizaba, Veracruz, Mxico. Distribution.Known only from the type locality. Guillarmodia (Proameria) (Von Martens 1891) Von Martens 1891; Biol. Cent. Amer.:74; pl. 5, (Von Martens). Pilsbry 1908; Man. Conch. Type Locality.Toluca, State of Mxico. Distribution.Known only from the type locality. Guillarmodia (Proameria) mitriformis (Angas 1879)Glandina (Oleacina ) mitriformis Angas 1879; Proc. Zool. Soc. Oleacina mitriformis (Angas). Tryon 1885; Man. Conch. 1:35; pl. Glandina mitriformis Angas. Von Martens 1891; Biol. Cent. Amer.:75. Euglandina mitriformis (Angas). Pilsbry 1908:201. Zhorquin to Cuabre, Costa Rica. Distribution.Known only from the type locality.

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178 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Guillarmodia (Proameria) oblonga (Pfeiffer 1866)Oleacina oblonga Pfeiffer 1866; Malak. Bltt. 13:86. Glandina oblonga (Pfeiffer). Von Martens 1891; Biol. Cent. Euglandina oblonga (Pfeiffer 1866). Pilsbry 1908; Man. Conch. Type Locality.Mirador, Veracruz, Mxico. Distribution.PUEBLA: Tehuacn (Von Martens 1891). VERACRUZ: Mirador. Guillarmodia (Proameria) orizabae (Pfeiffer 1856)Achatina (Varicella) orizabae Pfeiffer 1856; Proc. Zool. Soc. Lond. Glandina orizabae 8a (shell).Strebel 1875:52.Von Martens 1891:73.Type Locality.Volcn de Orizaba, at a height of 3000 meters, Veracruz, Mxico. Distribution.VERACRUZ: Las Vigas, on the edge of the plateau NW of Jalapa, 7900 ft. alt. (Von Martens 1891). Guillarmodia (Proameria) polita (Strebel 1875)Glandina polita Strebel 1875; Beitrag. Mex. Landund Sssw.Euglandina (Proameria) polita (Strebel). H. B. Baker 1941; Nautilus 55:57.H. B. Baker 1943; Proc. Acad. Nat. Sci. Phila. 95:8.Type Locality.A short distance in the Barranca Mihuistlan, near Coatepec, Veracruz, Mxico. Distribution.Known only from the type locality. Guillarmodia ( Proameria) potosiana potosiana Pilsbry 1908Euglandina oblonga var. potosiana Pilsbry 1908; Man. Conch. & Reguero 1996; Acta Zool. Mex. (73):16.Type Locality.Valles, San Luis Potos, Mxico. Syntypes in the ANSP. Distribution.SAN LUIS POTOS: Cascadas Micos Guillarmodia (Proameria) potosana tamaulipensis (Pilsbry 1908)Glandina oblonga tamaulipensis Pilsbry 1903; Proc. Acad. Nat. Sci. Euglandina oblonga var. tamaulipensis Pilsbry. Pilsbry 1908; Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):237.Type Locality.A canyon about 4 miles west of [Ciudad] Victoria, Tamaulipas, Mxico. Syntypes ANSP 85901. Distribution.NUEVO LEN: Iturbide (Correa-Sandoval 1997); numerous localities in southern part of state (Correa-Sandoval & Salazar 2005). TAMAULIPAS: numerous localities in southern part of state (Corrrea-Sandoval & Catsro 2002). Guillarmodia (Proameria) pulcherrima (Strebel 1883)Glandina monilifera form B, Strebel 1875; Beitrag. Mex. Landund Glandina pulcherrima Strebel 1883:104. Euglandina pulcherrima (Strebel).Pilsbry 1908; Man. Conch. Type Locality.Quautlatitlan, Veracruz, Mxico. Syntype Berlin Museum 30738. Distribution.VERACRUZ: Cuatepec, above Jalapa (Strebel 1883). Guillarmodia (Proameria) rhoadsi (Pilsbry 1899)Glandina rhoadsi Pilsbry 1899; Proc. Acad. Nat. Sci. Phila. 51:395.3b (shell). Euglandina rhoadsi (Pilsbry). Pilsbry 1908; Man. Conch. 19:192 Biol. Tropic., 41:685. Mxico. Syntypes ANSP 77173. Distribution.NUEVO LEN: Diente; Santiago (Correa Sandoval 1993). Guillarmodia ( Proameria) saxitilit saxitilis H. B. Baker 1941Euglandina (Proameria) saxitilit saxitilis H. B. Baker 1941; Nautilus anatomy).Type Locality.Below Necaxa, Puebla, Mxico; 3000 ft. alt.. Holotype in the UMMZ. Distribution.Known only from the type locality. Guillarmodia ( Proameria) H. B. Baker 1941Euglandina (Proameria) H. B. Baker 1941; Type Locality.Tepexic, below Necaxa, Puebla, Mxico; 2215 ft. alt.. Holotype in the UMMZ. Distribution.Known only from the type locality. Guillarmodia (Proameria) H. B. Baker 1941Euglandina (Proameria) H. B. Baker 1941; Type Locality.Above Necaxa, Puebla, Mxico. Holotype in the UMMZ. Distribution.Known only from the type locality. Guillarmodia (Proameria) sayula (Von Martens 1891)Glandina turgida var. sayula Von Martens 1891; Biol. Cent. Euglandina turgida var. sayula (Von Martens). Pilsbry 1908; Man. Type Locality.Sayula, Jalisco, Mxico. Distribution.JALISCO: Sayula. Guillarmodia (Proameria) speciosa (Pfeiffer 1856)Achatina (Varicella) speciosa Pfeiffer 1856; Proc. Zool. Soc. Lond.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 179Glandina speciosa (Pfeiffer). Fischer & Crosse 1870; Miss. Sci. Euglandina speciosa (Pfeiffer). Pilsbry 1908; Man. Conch. 19:210. Distribution.VERACRUZ: Huatusca; Orizaba (Von Martens 1891). Guillarmodia (Proameria) sulcifera (Von Martens 1891)Glandina sulcifera Von Martens 1891; Biol. Cent. Amer.:74; pl. 5, Euglandina sulcifera (Von Martens). Pilsbry 1908; Man. Conch. Type Locality.Jalisco, Mxico. Distribution.Known only from the type locality. Guillarmodia (Proameria) tepicensis Pilsbry & Cockerell 1926Euglandina tepicensis Pilsbry and Cockerell 1926; Proc. Acad. Nat. Type Locality.Tepc, Nayarit, Mxico. Holotype ANSP 139943. Distribution.Known only from the type locality. Guillarmodia (Proameria) tortillana (Pfeiffer 1846)Achatina (Glandina) tortillana Pfeiffer 1846; Proc. Zool. Soc. Lond. 14:32. Achatina tortillana Glandina tortillana (Pfeiffer). Strebel 1875; Beitrag. Mex. Landund Sssw.-Conch. II:50.Von Martens 1891:72. Oleacina tortillana (Pfeiffer). Tryon 1885; Man. Conch. 1:35; pl. Euglandina tortillana (Pfeiffer). Pilsbry 1908; Man. Conch. 19:201.Type Locality.Tortilla, Central America. Distribution.Unknown. Guillarmodia (Proameria) turgida (Pfeiffer 1861)Oleacina turgida Pfeiffer 1861; Proc. Zool. Soc. Lond. 29:26. Glandina turgida (Pfeiffer). Fischer & Crosse 1870; Miss. Sci. Mex. Euglandina turgida (Pfeiffer). Pilsbry 1908; Man. Conch. 19:199; Type Locality.Juquila, Oaxaca, Mxico. Distribution.Known only from the type locality. Guillarmodia (Proameria) (Pilsbry 1903) Pilsbry 1903; Proc. Acad. Nat. Sci. Phila. 55: (Pilsbry). Pilsbry 1908; Man. Conch. Rodriguez 2002; Acta Zool. Mex. (86):237.Type Locality.Canyon 4 miles west of [Ciudad] Victoria, Tamaulipas, Mxico. Lectotype ANSP 85918a (H. B. Baker 1963:219). Distribution.TAMAULIPAS: numerous localities in southern part of state (Correa-Sandoval & Rodriguez 2002). Genus Varicoglandina Pilsbry 1908 Varicoglandina Pilsbry 1908:201.Type Species.Glandina monilifera Pfeiffer 1845. Distribution.Southern Veracruz and adjacent Oaxaca, Chiapas, and Guatemala. Taxonomy.Five species and one subspecies are recognized. Varicoglandina constricta (Thompson 1995)Euglandina constricta Thompson 1995; Bull. Fla. Mus. Nat. Hist. Type Locality.A limestone ridge 3 km WNW of Mazn W). Holotype UF 190175. Distribution.Known only from the type locality. Varicoglandina monilifera monilifera (Pfeiffer 1845)Glandina monilifera Pfeiffer 1845; Proc. Zool. Soc. Lond. 13:75.Von Martens 1891; Biol. Cent. Amer.:75. Glandina monilifera Euglandina monilifera (Pfeiffer). Pilsbry 1908; Man. Conch. 34:43, 51.Bequaert 1957:217.Type Locality.Cobn, [Dept. Alta Verapaz], Guatemala. Distribution.CHIAPAS: Monte Alban, 950 m. alt.; El Censo to Laguma Ocotal, 1000 m alt. (Bequaert 1937). GUERRERO: Omilteme (Von Martens 1891). GUATEMALA, Dept. Alta Verapaz: mountains of Cobn; north of Chama (Hinkley 1920). Dept. Izabal: mountains of Cavech, above Cavech Village; Jocalo (Hinkley 1920). Varicoglandina monilifera rubella (Morelet 1849)Glandina rubella Morelet 1849; Test. Noviss. I:14. Euglandina monilifera var. rubella (Morelet 1849). Pilsbry 1908; Glandina iheringi Pilsbry 1900; Nautilus 14:4.Von Martens 1901; Biol. Cent. Amer.:612. Euglandina iheringi (Pilsbry). Pilsbry 1908; Man. Conch. 19:211; Type Localities. Glandina rubella: Province of Vera Paz, Guatemala. Glandina iheringi : Alta Vera Paz, Guatemala. Distribution.GUATEMALA: known only from the type localities. Varicoglandina nana (Shuttleworth 1852)Achatina (Polyphemus) nana Shuttleworth 1852; Mitt. Naturf. Ges. Bern 1852:202. Glandina nana (Shuttleworth). Fischer & Crosse 1870; Miss. Sci. Mex. Landund Sssw.-Conch. II:39.Von Martens 1891; Euglandina nana (Shuttleworth). Pilsbry 1908; Man. Conch. 19:201. Vericoglandina nana (Shuttleworth). Neubert & Gosteli 2003; Type Locality.Veracruz, Mxico. Syntype Naturhistorische Museum Bern 18845/1 (Neubert & Gosteli 2003). Martens 1891).

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180 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Varicoglandina rubiginosa (Thompson 1995)Euglandina (Varicoglandina ) rubiginosa Thompson 1995; Bull. Type Locality.A limestone knoll 11 km S of Cobn, Distribution.Known only from a small area in the Dept. Alta Verapaz, Guatemala. GUATEMALA; Dept. Alta Varicoglandina stigmatica (Shuttleworth 1852)Achatina (Polyphema) stigmatica Shuttleworth 1852; Mitt. Naturf. Ges. Bern 1852:2002. Glandina stigmatica (Shuttleworth). Fischer & Crosse 1870; Miss. Euglandina stigmatica (Shuttleworth). Pilsbry 1908; Man. Conch. 19:210.Neubert & Gosteli 2003; Contributions to Natural Euglandina (Varicoglandina ) stigmatica (Shuttleworth). H. B. Baker 1941; Nautilus 55:60.H. B. Baker 1943; Proc. Acad. 13 (radula). Syntype: Naturhistorische Museum Bern 18846/1 (Neubert & Gosteli 2003). (Von Martens 1891); Peuela to Sumidero, 2625 ft. alt. (H. B. Baker 1941). Genus Streptostylella Pilsbry 1907Streptostylella Pilsbry 1907; Man. Conch. 19:161.H. B. Baker Type Species.Streptostyla botteriana Crosse & Fischer 1869. Distribution.Central Veracruz, Mxico. Taxonomy.A single species is recognized. Streptostylella botteriana (Crosse & Fischer 1869)Streptostyla botteriana Crosse & Fischer 1869; Jour. de Conchyl. 18:190.Fischer & Crosse 1870; Miss. Sci. Mex. I:25; pl. 2, 95 (shell).Von Martens 1891; Biol. Cent. Amer.:85. Streptostyla (Streptostylella ) botteriana Crosse & Fischer. Pilsbry 1907; Man. Conch. 19:161.Thompson 2009; Arch. Moll. Type Locality.Orizaba, Veracruz, Mxico. Distribution.VERACRUZ: W slpe of Cerro de Las Palmas, San Bartolo, 851 m alt. (18.9433 N, 96.9283 W); Cerro de Las Palmas, 1 km N of San Mateo, ca. 3 km N of 2009). Genus Varicoturris Pilsbry 1907Vericoturris Pilsbry 1907; Man. Conch. 19:161.H. B. Baker 1941; Nautilus 55:56.Thompson 1995; Bull. Fla. Mus. Nat. Hist. 39:47.Thompson 2009; Arch. Molluskenkunde 138:64 65. Ghiesbreghtia H. B. Baker 1941; Nautilus 55:52, 54.Type Species.Varicoturris: Spiraxis dubia Pfeiffer 1856. Ghiesbreghtia : Euglandina (Ghiesbreghtia ) H. B. Baker. Distribution.Southeastern Mxico and northern Guatemala. Taxonomy.The genus contains six species. Varicoturris dubia (Pfeiffer 1856)Spiraxis dubia Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:378.Pfeiffer 1859. Monographia heliceorum viventium, IV:580. Streptostyla dubia (Pfeiffer). Fischer & Crosse 1875; Miss. Sci. Mex. Amr. Cent.:65.Von Martens 1891; Biol. Cent. Amer.:90; pl. 65. Euglandina dubia (Pfeiffer), H. B. Baker 1941; Nautilus 55:54.(shell).Type Locality.Chiapas, Mxico. Syntypes BMNH 1991144. Distribution.Mxico in extreme northern Chiapas and Adjacent Tabasco. CHIAPAS:3 km N of Ixhuatn, 550 m alt. TABASCO: 4 km E of Teapa (Thompson 1995). Varicoturris elegans Thompson 2009Varicoturris elegans Thompson 2009; Arch. Molluskenkunde Type Locality.Limestone range on the northwest shore of Presa Miguel Aleman 11 km southwest, 3.5 m. northeast of Tierra Blanca, Oaxaca, Mxico; 100 m alt. (18.405 N, 96.460 W). Distribution.Known only from the type locality. (H. B. Baker 1941) H. B. Baker 1941; Nautilus 6 (shell). Mxico; 2700 ft. alt. Holotype in the UMMZ. Distribution.VERACRUZ: Cerro de las Palmas, 1 km 1995). Varicoturris hadra Thompson 2009Varicoturris hadra Thompson 2009: Arch. Molluskenkunde Type Locality.Limestone ridge 12 km northwest of Bethania, Oaxaca, Mxico; 100 m alt. (17.968 N, 96.115 W). Distribution.Known only from the type locality. Varicoturris huehuetenangoensis (Thompson 1995)Euglandina huehuetenangoensis Thompson 1995; Bull. Fla. Mus.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 181 Dept. Huehuetenango, Guatemala. Holotype UF 190305. Distribution.GUATEMALA: known only from the immediate vicinity of the type locality. Varicoturris pycnoptyx (Thompson 1995)Euglandina pycnoptyx Thompson 1995; Bull. Fla. Mus. Nat. Hist. Type Locality.Limestone bluff 6 km NE of Comalapa, Veracruz, Mxico. Holotype UF 81946. Distribution.VERACRUZ: known only from the immediate vicinity of the type locality. Genus Pittieria Von Martens 1901Pittieria Von Martens 1901; Biol. Cent. Amer.:617.Pilsbry 1907; Man. Conch 19:162.H. B. Baker 1941; Nautilus 55:55.Type Species.Streptostyla (Pittieria ) bicolor Von Martens 1901. Distribution.Central Mxico south to northern Panam. Taxonomy.Three subgenera and fourteen species are recognized. Subgenus Pittieria Von Martens 1901 Distribution.Costa Rica. Taxonomy.Two species are recognized. Pittieria (Pittieria ) bicolor (Von Martens 1901)Streptostyla (Pittieria) bicolor Von Martens 1901; Biol. Cent. Type Locality.Uiskar, in Alta Talamanca, Costa Rica. Distribution.Known only from the type locality. Pittieria (Pittieria ) pittieri (Von Martens 1901)Glandina pittieri Von Martens 1901; Biol. Cent. Amer.:611; pl. 28, Euglandina pittieri (Von Martens). Pilsbry 1908; Man. Conch. Type Locality.Between Mokri and Ukutschka, near Talamanca, Costa Rica. Distribution.Known only from the type locality. Subgenus Pilsbry 1908 Pilsbry 1908; Man. Conch. 19:201.H. B. Baker 1941; Nautilus 55:55.H. B. Baker 1943; Proc. Acad. Nat. Sci. Phila. 95; Thompson 1995; Bull. Fla. Mus. Nat. Hist. 39:12.Type Species.Oleacina underwoodi Fulton 1897 Distribution.Central Veracruz and Guerrero south to northern Panam. Taxonomy.Nine species are recognized. Pittieria ( ) aurantiaca (Angas 1879)Glandina aurantiaca Angas 1979; Proc. Zool. Soc. Lond. 47:480; Euglandina () aurantiaca (Angas). Pilsbry 1908; Man. Conch. 19:204.Pilsbry 1926; Proc. Acad. Nat. Sci. Type Locality.From the hill country, Costa Rica, Distribution.COSTA RICA: Puerto Viejo; Rio Sarapiqu; San Miguel, on the road to Sarapiqu; Turrialba; Tuis, 600 m alt.; Santa Clara, 200 m alt.; Talamanca; Valley of Baca; Terraba (all Von Martens 1901). PANAM: Bocas del Toro (Pilsbry 1926). Pittieria ( ) broctontomlini ( Pilsbry 1926)Euglandina broctontomlini Pilsbry 1926; Proc. Acad. Nat. Sci. Type Locality.Vesta Farms, Estrella Vally, Costa Rica. Holotype ACSP 132557a. Distribution.COSTA RICA: Cahuita; Rio Jiminey, 50 ft. alt. PANAM, Prov. Bocas del Toro: Mono Creek. Pittierria ( ) chiriquiensis (Da Costa 1900)Glandina chiriquiensis Martens 1901; Biol. Cent. Amer.:612. Euglandina chiriquiensis (Da Costa). Pilsbry 1908; Man. Conch. Pittierria chiriquiensis(Da Costa). H. B. Baker 1943; Proc. Acad. Type Locality.Chiriqu, Panam. Distribution.PANAM: Near Santa Clara, Volcan de Chiriqu, 4100 ft. alt. (Pilsbry 1908). Pittieria ( ) decidua (Pfeiffer 1861)Oleacina decidua Pfeiffer 1861; Proc. Zool. Soc. Lond. 29:26. Glandina decidua (Pfeiffer). Fischer & Crosse 1970:122. Von Euglandina decidua (Pfeiffer). Pilsbry 1908; Man. Conch. Type Locality.Juquila, Oaxaca, Mxico. Distribution.Known only from the type locality. Pittieria ( ) izabellina (Pfeiffer 1846)Achatina (Glandina) izabellina Pfeiffer 1846; Proc. Zool. Soc. Euglandina isabelina (Pfeiffer). Pilsbry 1908; Man. Conch. 19:203 204.Pilsbry 1926:97.Type Locality.Mxico. Distribution.Unknown. Pittieria ( ) lanceolata (Von Martens 1891)Glandina lanceolata Von Martens 1891; Biol. Cent. Amer.:69; pl. 4, Euglandina lanceolata (Von Martens). Pilsbry 1908; Man. Conch. Type Locality.Omilteme, Guerrero, Mxico. Distribution.Known only from the type locality. Pittieria ( ) obtusa (Pfeiffer 1844)Glandina obtusa Pfeiffer 1844; in

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182 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Euglandina obtusa Pilsbry 1908; Man. Conch. 19:204.Type Locality.Ralejo, Nicaragua. (Pilsbry 1908). Pittieria ( ) tryoniana Pilsbry 1908Euglandina tryoniana Pilsbry 1908; Man. Conch. 19:203; pl. 27, Type Locality.Central America. Holotype ANSP 24814. Distribution.Unknown. Pittieria ( ) underwoodi (Fulton 1897)Oleacina underwoodi Fulton 1897; Ann. & Mag. Nat. Hist. 6th ser. Glandina underwoodi (Fulton). Von Martens 1901; Biol. Cent. Amer.:612. Euglandina uderwoodi (Fulton). Pilsbry 1908; Man. Conch. Type Locality.Asaha Centago [Azahar de Cartago], Costa Rica. Distribution.Known only from the type locality. Subgenus Shuttleworthia H. B. Baker 1941Shuttleworthia H. B. Baker 1941; Nautilus 55:55.H. B. Baker 1943; Proc. Acad. Nat. Sci. Phila. 95:11.Type Species.Pittieria arborea H. B. Baker 1941. Distribution.Eastern Mxico in the states of Puebla and Veracruz. Taxonomy.Three species are recognized. Pittierria (Shuttleworthia) ambigua (Pfeiffer 1856)Achatina ambigua Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:321. Glandina ambigua (Pfeiffer). Fischer & Crosse 1870; Miss. Sci. Oleacina ambigua (Pfeiffer). Tryon 1885; Man. Conch. 1:23; pl. 4, Euglandina ambigua (Pfeiffer). Pilsbry 1908; Man. Conch. 19:201 Pittieria (Shuttleworthia) ambigua (Pfeiffer). H. B. Baker 1941; Nautilus 55:59. Distribution.VERACRUZ: Bajada del Ojo de Agua, Pittierria (Shuttleworthia) arborea H. B. Baker 1941Pittierria arborea 13.H. B. Baker 1943; Proc. Acad. Nat. Sci. Phila. 95:11 (radula), 28 (pallial organs).Type Locality.Below Necaxa, Puebla, Mxico; 3120 ft. alt. Holotype in the UMMZ. Distribution.Known only from the type locality. Pittierria ( Shuttleworthia) (Crosse & Fischer 1869) Crosse & Fischer 1869; Jour. de Conchyl. 17:426.Fischer & Crosse 1870; Miss. Sci. Mex. I:132; pl. 6, (Crosse & Fischer). Pilsbry 1908; Man. Conch. 19:201. Pittieria (Shuttlewortia) (Crosse & Fischer). H. B. Baker 1941; Nautilus 55:59.Type Locality.Probably in the environs of Cordova, Veracruz, Mxico. Distribution.Known only from the type locality. Subfamily STREPTOSTYLINAE H. B. Baker 1941Streptostylarum H. B. Baker 1941.Thompson 2010; Revista de Biologia Tropical 58:196.Type Genus.Streptostyla Shuttleworth 1852. Distribution.Mxico and Central America. A single species is reported from Venezuela. Taxonomy.Seventy-four species are recognized in the study area. Genus Myxastyla Thompson 1995Myxastyla Thompson 1995; Bull. Fla. Mus. Nat. Hist. 39:73.Type Species.Streptostyla coxeni Richards 1938. Distribution.Northern Central America. Taxonomy.Three species are recognized. Myxastyla coxeni (Richards 1938)Streptostyla coxeni Richards 1938; Proc. Amer. Philos. Soc. 79:172; Myxastyla coxeni (Richards). Thompson 1995; Bull. Fla. Mus. Nat. Type Locality.Limestone outcrops between Coxen Hole and West End, Isla de Roatn, Dept. Islas de la Baha, Honduras. Holotype ANSP 170020. Distribution.HONDURAS: apparently endemic to Isla de Roatn. Other species of Myxastyla currently undescribed, occur elsewhere in Honduras. Myxastyla hyalina Thompson 1995Myxastyla hyalina Thompson 1995; Bull. Fla. Mus. Nat. Hist. Type Locality.Limestone knoll 17.5 km by road NW Distribution.GUATEMALA, Dept. Alta Verapaz: 11 km W of San Cristobal Verapaz, 1120 m alt.; 1.5 km SE of San Juan Chamelco; 1300 m alt.; 2 km ESE of Cajj; 1250 m alt.; 11 km S of Cobn; 1350 m alt.; 14 km N of Cobn; 990 m alt. (Thompson 1995). Myxastyla pycnota Thompson 1995Myxastyla pycnota Thompson 1995; Bull. Fla. Mus. Nat. Hist. Type Locality.Gorge along the Rio Selequa 12 km SSE of La Democracia, Dept. Huehuetenango, Guatemala; 950 m alt. HOLOTYPE UF 190415. Distribution.Known only from the type locality.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 183 Genus Oryzosoma Pilsbry 1891Oryzosoma Pilsbry 1891; Nautilus 5:9.Pilsbry 1891; Proc. Acad. Nat. Sci. Phila. 43:311.Pilsbry 1907; Man. Conch 19:163.Type Species. Streptostyla (Orizosoma ) tabiensis Pilsbry 1891. Distribution.Yucatn, Mxico. Taxonomy.The single known species is known only from a single bleached shell. Oryzosoma tabiensis (Pilsbry 1891)Streptostyla (Orizosoma ) tabiensis Pilsbry 1891; Nautilus 5:9.6 (shell). Oryzosoma tabiensis (Pilsbry). Pilsbry, Man. Conch 19:163; pl. 30, Type Locality.Cave in the mountains near the Hacienda Tabi, Yucatn, Mxico. Holotype ANSP 61630. Distribution.Known only from the type locality. Genus Salasiella Strebel 1878Salasiella Strebel 1878; Beitrag. Mex. Landund Sssw.-Conch. III:29.H. B. Baker 1941; Nautilus 54:80.Type Species.Salasiella joaquinae Strebel 1877. Distribution.Mxico south to Panam. Taxonomy.Two subgenera are recognized. The genus includes twelve species. Subgenus Salasiella Strebel 1878 Distribution.Mxico south to Panam, Isla de Providencia, Colombia. Taxonomy.Nine species are included in the subgenus. Salasiella (Salasiella ) browni Pilsbry 1910Salasiella browni Pilsbry 1910; Proc. Acad. Nat. Sci. Phila. 62:508; 82:345.H. B. Baker 1923; Occ. Pap. Mus. Zool. Univ. Mich. (135):7.Type Locality.Las Cascades, Canal Zone, Panam. Holotype ANSP 101325. Distribution.PANAM: Las Cascades; between Tabernillo and San Pablo (Pilsbry 1926); Isla Barro Colorado (Pilsbry 1930) Salasiella (Salasiella ) corni Richards 1939Salasiella corni Richards 1939; Proc. Amer. Philos. Soc. 81:32; Type Locality.Near Quin Bluff, Isla del Maz, Nicaragua. Holotype ANSP 173926. Distribution.Known only from the type locality. Salasiella (Salasiella ) guadalajarensis Pilsbry & Cockerell 1926Salasiella guadalajarensis Pilsbry and Cockerell 1926; Proc. Acad. Type Locality.Guadalajara, Jalisco, Mxico. Holotype: ANSP 107474. Distribution.Known only from the type locality. Salasiella (Salasiella ) guatemalensis Pilsbry 1920Salasiella guatemalensis Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. Baker 1923; Occ. Pap. Mus. Zool. Univ. Mich. (135):7.Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):8.Type Locality.Gualan, [Dept. Zacapa], Guatemala. Holotype ANSP 114838. Distribution.GUATEMALA: Jocola (Pilsbry 1920). Dept. Petn: Tikal National Park (Basch 1959). Salasiella (Salasiella ) hinkleyi Pilsbry 1920Salasiella hinkleyi Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 71:212; Sandoval & Rodriguez 2002; Acta Zool. Mex. (79):237.Type Locality.Mountain side near the Tamosopo Sugar Companys place, San Luis Potos, Mxico. Holotype ANSP 45198a. Distribution.NUEVO LEN: Dientew, near Monterrey (Pilsbry 1920). SAN LUIS POTOS: El Abra (Pilsbry 1920); numerous localities (Correa-Sandoval et al. 1998). NUEVO LEN: Diente, near Monterrey (Pilsbry 1920). SAN LUIS POTOS: El Abra (Pilsbry 1920). TAMAULIPAS: Los Ejido Cuauhtemoc (Correa-Sandoval & Rodriguez 2002). Salasiella (Salasiella ) joaquinae Strebel 1878Salasiella (Salasiella ) joaquinae Strebel 1878; Beitrag. Mex. LandPilsbry 1907; Man. Conch. 19:172.Type Locality.Jalapa, Veracruz, Mxico. Distribution.NUEVO LEN: Diente, near Monterrey (Pilsbry 1907). VERACRUZ: only from the type locality. Salasiella (Salasiella ) margaritacea (Pfeiffer 1856)Achatina margaritacea Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:321. Salasiella margaritacea (Pfeiffer). Von Martens 1891; Biol. Cent. Zool. Univ. Mich. (135):6. Coatotolopam (H. B. Baker 1923). Salasiella (Salasiella ) pulchella (Pfeiffer 1856)Achatina (Oleacina) pulchella Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:379 (not Achatina pulchella Spix 1827). Glandina pulchella (Pfeiffer). Fischer & Crosse 1872:136. Salasiella pulchella (Pfeiffer). Von Martens 1891; Biol. Cent. Univ. Mich. (135):7.Jacobson 1968; Nautilus 81:118. Salasiella pfeifferi Pilsbry 1899; Proc. Acad. Nat. Sci. Phila. 51:398 (new name for Achatina pulchella Pfeiffer 1856).

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184 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Type Locality.Chiapas, Mxico. Distribution.CHIAPAS: type locality. NICARAGUA, Dept. Zelaya: 78 mi. W of San Carlos, along the Rio Cocos (Jacobson 1968). Salasiella (Salasiella ) subcylindrica Pilsbry 1903Salasiella subcylindrica Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. Mxico. Holotype ANSP 77169a. Distribution.Known only from the type locality. TABASCO: 0.5.0 mi. E of Teapa (Thompson 1957). Salasiella (?) camerata H. B. Baker 1941Oleacina ? (Salasiella?) camerata H. B. Baker 1941; Nautilus 55:55; Type Locality.Tepexic, below Necaxa, Puebla, Mxico; 2200 ft. alt. Holotype in the UMMZ. Distribution.PUEBLA: known only from the type locality and a near-by locality above Necaxa, 4925 ft. alt. (H. B. Baker 1941). Subgenus Perpusilla H. B. Baker 1941Perpusilla H. B. Baker 1941; Nautilus 54:81.Type Species.Salasiella perpusilla (Pfeiffer 1866). Distribution.Eastern Mxico in the states of Nuevo Taxonomy.Three species are recognized. Salasiella (Perpusilla) minima Pilsbry 1907Salasiella minima Salasiella (Perpusilla ) minima Pilsbry. H. B. Baker 1941; Nautilus Correa-Sadoval & Salazar 2005; Acta Zool. Mex. 21:60.Type Locality.Hills around Orizaba, about 500 ft. above the town, Veracruz, Mxico. Holotype ANSP 58070. Distribution.VERACRUZ: type locality. PUEBLA: near Necaxa (H. B. Baker 1941). NUEVO LEN: ROAD Salasiella (Perpusilla) modesta (Pfeiffer 1862)Oleacina modesta Pfeiffer 1862; Malak. Bltt. 9:98. Glandina modesta (Pfeiffer). Fischer & Crosse 1872:135.Strebel Salasiella modesta (Pfeiffer). Strebel 1878; Beitrag. Mex. LandBasch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):8. Salasiella (Perpusilla) modesta (Pfeiffer). H. B. Baker 1941; Type Locality.Veracruz, Mxico. Distribution.PUEBLA: near Necaxa (H. B. Baker GUATEMALA, Dept. Petn: Tikal National Park (Basch 1959). Salasiella (Perpusilla) perpusilla (Pfeiffer 1866)Oleacina perpusilla Pfeiffer 1866; Malak. Bltt. 13:86. Glandina perpusilla (Pfeiffer). Fischer & Crosse 1872:134; pl. 3, (shell). Salasiella (Perpusilla) perpusilla (Pfeiffer). H. B. Baker 1941; Type Locality.Mirador, Veracruz, Mxico. Distribution.NUEVO LEN: Diente, near Monterrey (Pilsbry 1907). PUEBLA: Necaxa (H. B. Baker 1941). VERACRUZ: woods of Pacho, near Jalapa (Strebel 1877). Genus Strebelia Crosse & Fischer 1868Physella Pfeiffer 1861; Malak. Bltt. 8:71 (non Physella Haldeman 1842). Strebelia Crosse & Fischer 1868; Jour. de Conchyl. 16:90. Pilsbry 1907; Man. Conch. 19: xxvii. Spirobulla Ancey 1881; Le Naturaliste, 1:484.Type Species.Physella berendti Pfeiffer 1861. Distribution.Eastern Mxico in the states of Veracruz and Hidalgo. Taxonomy.The genus is monotypic. Strebelia berendti (Pfeiffer 1861)Physella berendti Strebelia berendti (Pfeiffer). Crosse & Fisher 1868; Jour. de Conchyl. 1904; Proc. Academy of Natural Sciences of Philadelphia:774 .Type Locality.Here restricted to Mirador, Veracruz, Mxico. Distribution.VERACRUZ: Mirado; Coiscomatepec (Pfeiffer 1861); Pacho, near Jalapa (Strebel 1878); Texolo (Pilsbry 1903). Genus Streptostyla Shuttleworth 1852Streptostyla Shuttleworth 1852; Mittheilungen fr Naturforschenden Gesellschaft in Bern:203.Pilsbry 1907; Man. Conch. 19:144 145.H. B. Baker 1927; Nautilus 41:21.H. B. Baker 1941; Nautilus 55:53.Thompson 1967; Bull. Fla. St. Mus. 11:233 234.Type Species.By tautonymy, Achatina streptostyla Pfeiffer 1846. Distribution.Mxico, Central America, and Cuba. A single species is reported from Venezuela. Taxonomy.Fifty-six species and fourteen subspecies are recognized in Mxico and Central America. Four subgenera, Streptostyla Chersomitra, Peteniella and Eustreptostyla occur in the study area. Another subgenus, Rectoleacina occurs in Cuba. The distinctions between the subgenera Streptostyla and Chersomitra are primarily anatomical. The numerous species that remain unknown

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 185 anatomically are retained in the subgenus Streptostyla with the exceptions of those placed in Eustreptostyla and Peteniella Within Streptostyla species are grouped according to shell shape only as a matter of convenience following Von Martens (1892) and Pilsbry (1897). The arrangement does to achieve without further anatomical or genetic information. Subgenus Streptostyla Shuttleworth 1852 Distribution.Mxico and Central America. Taxonomy.Forty-two species and twelve subspecies are recognized. Taxa with Oliviform Shell Streptostyla (Streptostyla ) Ancey 1903 Ancey 1903; Nautilus 17:56. (Not Ancey. Pilsbry 1907; Man. Conch. 19:160 161.Type Locality.Central America. Distribution.None recorded. Taxonomy.Apparently Anceys name was a typographic error, which was corrected to by Pilsbry (1907). Streptostyla (Streptostyla ) cylindracea (Pfeiffer 1846)Achatina cylindracea Pfeiffer 1846; Proc. Zool. Soc. Lond. 14:31. Streptostyla cylindracea (Pfeiffer). Fischer & Crosse 1870:30.Angas 1879; Proc. Zool. Soc. Lond. 47:482.Tryon 1885; Man. Type Locality.Tortilla, Central America. (Von Martens 1891). Streptostyla (Streptostyla ) gabbi Pilsbry 1907Streptostyla boucardi (Pfeiffer) var. (?). Angas 1879; Proc. Zool. Soc. Lond. 47:482. Streptostyla boucardi (Pfeiffer). Tryon 1885; Man. Conch. 1:46; pl. Streptostyla gabbi 2, 3 (shell).Type Locality.On the ridge between Tiloria and Zorquin, Costa Rica. Distribution.COSTA RICA: Cabeceras de Tarazu, 2000 m alt.; Alahuela; Tarbaca; Diquis, below Terraba; Caas Gordas (Von Martens 1901). Streptostyla (Streoptostyla ) irrigua irrigua (Shuttleworth 1852)Spiraxis (Streptostyla) irrigua Shuttleworth 1852; Mitt. Natrf. Ges. Bern 1852:205. Streptostyla irrigua (Shuttleworth). Fischer & Crosse 1870; Miss. 58.Von Martens 1891; Biol. Cent. Amer.:92.H. B. Baker syntype: Naturhistorische Museum Bern 18837/1 (Neubert & Gosteli 2003). alt. PUEBLA: Necaxa (H. B. Baker 1943) Streptostyla (Streptostyla ) irrigua cingulata Crosse & Fisher 1869Streptostyla cingulata Crosse & Fischer 1869; Jour. de Conchyl. Streptostyla irrigua cingulatum Crosse & Fischer. Pilsbry 1907, Man. Conch. 19:146.Type Locality.[San Andres] Tuxtla, Veracruz, Mxico. Distribution.VERACRUZ: Orizaba (Von Martens 1891). Streptostyla (Streptostyla ) irrigua quirozi Strebel 1878Streptostyla quirozi Strebel 1878; Beitrag. Mex. Landund Sssw.Streptostyla shuttleworthi var. quirozi Strebel. Von Martens 1891; Biol. Cent. Amer.:93. Streptostyla irrigua quirozi Strebel. Pilsbry 1907; Man. Conch. 19:146.H. B. Baker 1941; Nautilus 55:60; H. B. Baker 1943; Proc. Acad. Nat. Sci. Phila. 95:7.Type Locality.Not stated. Distribution.VERACRUZ: Coatepec; Miahuatlan; San Antonio de Monte (Strebel 1877); Tezuitlan; Misantla (Von Martens 1891). PUEBLA: Necaxa, 2120 ft. alt. (H. B. Baker 1943). Streptostyla (Streptostyla ) irrigua similis Strebel 1878Streptostyla similis Strebel 1878; Beitrag. Mex. Landund Sssw.Streptostyla shuttleworthi var. similis Strebel. Von Martens 1891; Biol. Cent. Amer.:93.Type Locality.Misantla, Veracruz, Mxico. Distribution.VERACRUZ: Orizaba; Veracruz OAXACA: Juquila (Von Martens 1891). Streptostyla (Streptostyla ) Von Martens 1891Streptostyla shuttleworthi var. Fischer & Crosse 1870; Miss. Sci. Streptostyla shuttleworthi Streptostyla shutttleworthi var. Von Martens 1891; Biol. Cent. Amer.:93. Streptostyla irrigua var. Von Martens. Pilsbry 1907; Man. Conch. 19:146. Distribution.VERACRUZ: Orizaba; Nautla (Von Martens 1891). Streptostyla (Streptostyla ) lattrei lattrei ( Pfeiffer 1845)Achatina lattrei Pfeiffer 1845; Proc. Zool. Soc. Lond. 13:138.

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186 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Reeve 1849; Conch. Icon, 6, Achatina Streptostyla delattrei (Pfeiffer). Fischer & Crosse 1870; Miss. Sci. Streptostylla lattrei (Pfeiffer). Pilsbry 1907; Man. Conch. 19:145.Hinkley 1920; Nautilus 34:43, 51. Morelet 1849; Test. Noviss. I:13.Type Locality.Achatina lattrei: Not given. Glandina : Vera Paz, Guatemala. Distribution.GUATEMALA. Alta Verapaz: Cobn; Chama (Hinkley 1920). Dept. Baja Verapaz: Panzos; Senahu, 2800 ft. alt. Dept. Izabal: Izabal (Von Martens 1891); mountains of Rio Cavech, above Cavech Village (Hinkley 1920). Streptostyla (Streptostyla ) lattrei edwardsiana Crosse & Fischer 1869Streptostyla delattrei var. edwardsiana Crosse & Fischer 1869; Jour. de Conchyl. 17:29.Fischer & Crosse 1870; Miss. Sci. Mex. Streptostyla lattrei var. edwardsiana Crosse & Fischer. Pilsbry 1907; Man. Conch. 19:145.Type Locality.Orizaba, Veracruz, Mxico. Distribution.Known only from the type locality. Streptostyla ( Streptostyla) lattrei sallei Crosse & Fischer 1869Streptostyla delattrei var. sallei Crosse & Fischer 1869; Jour. de Streptostyla sallei Crosse & Fischer. Tryon 1885; Man. Conch. 1:44; Streptostyla lattrei var. sallei Crosse & Fischer. Pilsbry 1907; Man. Conch. 19:145.Type Locality.Orizaba, Veracruz, Mxico. Distribution.Known only from the type locality. Streptostyla (Streptostyla ) Pilsbry 1899Streptostyla Pilsbry 1899; Proc. Acad. Nat. Sci. Phila. 51:397.Pilsbry 1903, Proc. Acad. Nat. Sci. Phila. 55:774; pl. 41:605.Correa-Sandoval & Salazar 2005; Acta Zool. Mex. (21):60. Mxico. Lectotype ANSP 77153a (H. B. Baker 1963:219). Distribution.NUEVO LEN: Santiago (CorreaSandoval 1993); numerous localities (Correa-Sandoval & Salazar 2005). Streptostyla (Streptostyla ) palmeri Dall 1905Streptostyla palmeri Dall 1905; Smiths. Misc. Coll. 48:191.Pilsbry Sandoval, Garca-Cubas & Reguero 1998; Acta Zool. Mex. (73):16.Type Locality.Sierra Alvarz, San Luis Potos, Mxico; 7200 ft. alt. Holotype USNM 110394. Distribution.SAN LUIS POTOS: 2 km NW Poza de l Sandoval & Rodrigez 2002) Streptostyla (Streptostyla ) potosiana Dall 1905Streptostyla potosiana Dall 1905; Smiths. Misc. Coll. 48:190; pl. (shell).Correa-Sandoval, Garca-Cubas & Reguero 1998; Acta Zool. Mex. (73):16.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):237.Type Locality.Sierra Alvarz, San Luis Potos, Mxico; at a height of 7200 ft. Holotype USNM 110395. Distribution.SAN LUIS POTOS: Hwy. Ro VerdeW) (Correa-Sandoval et al. 1998). TAMAULIPAS: La W) (Correa-Sandoval & Rodriguez 2002) Streptostyla (Streptostyla ) shuttleworthi (Pfeiffer 1856)Spiraxis shuttleworthi Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:320; Streptostyla shuttleworthi Martens 1891:92.Pilsbry 1907; Man. Conch. 19:146. Spiraxis bullacea Pfeiffer 1866; Malak. Bltt. 13:84.Von Martens 1891; Biol. Cent. Amer.:93. Distribution.CHIAPAS. TABASCO: Teapa. VERATaxa with Coniform Shell Streptostyla (Streptostyla ) conulus Von Martens 1891Streptostyla conulus Von Martens 1891; Biol. Cent. Amer.:94; pl. (shell).Type Locality.Sayula, Jalisco, Mxico. Distribution.Known only from the type locality. Streptostyla (Streptostyla ) gracilis Pilsbry 1907Streptostyla gracilis Pilsbry 1907; Man. Conch. 19:148; pl. 28, Garca-Cubas & Reguero 1998; Acta Zool. Mex. (73):16.Correa-Sandoval 2000; Acta Zool. Mex. (79):9.CorreaSandoval & Rodriguez 2002; Acta Zool. Mex. 86:237.Type Locality.Valles, San Luis Potos, Mxico. Holotype ANSP 92802a (Baker 1963:219). Distribution.SAN LUIS POTOS: numerous localities (Correa-Sandoval et al. 1998). TAMAULIPAS: Km 636, on the road between Santa Inez and Llera (Jacobson 1956); numerous localities (Correa-Sandoval & Rodriguez 2002). W) (Correa-Sandoval 2000).

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 187 Streptostyla (Streptostyla ) streptostyla streptostyla (Pfeiffer 1846)Achatina streptostyla Pfeiffer 1846; Zeitschrift fr Malakozoologie:159. Streptostyla streptostyla .H. B. Baker 1927; Nautilus 41:21.H. B. Baker 1941; Nautilus 55:60.H. B. Baker 1943; Proc. Acad. Nat. Sci. Phila. 95:6. Streptostyla coniformis Streptostyla pfeifferi Fischer & Crosse 1870; Miss. Sci. Mex. I:27.Tryon 1885; Man. Conch. 1:47.Type Locality.Mxico. Distribution.VERACRUZ: Orizaba; Sumidero 2625 3400 ft. alt. (H. B. Baker 1941). Streptostyla (Streptostyla ) streptostyla coniformis (Shuttleworth 1852)Spiraxis (Streptostyla ) coniformis Shuttleworth 1852; Mitt. Naturf. Ges. Bern 1852:206. Stretostyla coniformis (Shuttleworth). Fischer & Crosse 1870:43.Streptostyla blandiana Crosse & Fischer 1869; Jour. de Conchyl. Streptostyla pfeifferi Crosse & Fischer, in Fischer & Crosse 1870:27. Streptostyla streptostyla form coniformis (Shuttleworth). H. B. Baker 1941; Nautilus. 55:60.Type Locality.Spiraxis coniformis: Cordova, Veracruz, Mxico.; Naturhistorische Museum Bern 18835/1 (Neubert & Gosteli 2003). Streptostyla blandiana: Cordova, Veracruz, Mxico. Streptostyla pfeifferi : Mxico. Coatepec; Orizaba (Von Martens 1891). Taxa with Turdiga Shell Streptostyla (Streptostyla ) chiriquiana Von Martens 1901 Da Costa 1900; Proc. Malac. Soc. London Spiraxis (Streptostyla ) Shuttleworth 1852). Streptostyla chiriquiana Von Martens 1901; Biol. Cent. Amer.:615.Pilsbry 1907 (new name for Da Costa 1900); Man. Conch. 19:151.Tomlin 1927; Nautilus 41:130. Streptostyla pallidus Da Costa 1901; Proc. Malac. Soc. London 9:185 (new name for Da Costa 1900).Type Locality.Chiriqu, Panam. Distribution.Known only from the type locality. Streptostyla (Streptostyla ) costaricensis Da Costa 1904Streptostyla costaricensis Da Costa 1904; Proc. Malac. Soc. London Type Locality.Azajar de Cartago, Costa Rica. Distribution.Known only from the type locality. Streptostyla (Streptostyla ) labida (Morelet 1851)Glandina labida Morelet 1851; Test. Noviss. II:13. Streptostyla labida (Morelet). Fischer & Crosse 1870; Miss. Sci. Type Locality.Vera Paz [Alta Verapaz], Guatemala. Distribution.Known only from the type locality. Von Martens (1901) reported this species from Costa Rica. Streptostyla (Streptostyla ) obesa Von Martens 1891Streptostyla obesa Von Martens 1891; Biol. Cent. Amer.:95; pl. 5, 71 (shell).Type Locality.Honduras. Distribution.Known only from the type locality. Streptostyla ( Streptostyla) turgidula turgidula (Pfeiffer 1856)Spiraxis turgidula Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:320; pl. Streptostyla turgidula (Pfeiffer). Fischer & Crosse 1870; Miss. Sci. 1891; Biol. Cent. Amer.:95.H. B. Baker 1941:60. Distribution.VERACRUZ: Jalapa (Strebel 1877); (H. B. Baker 1941:60). Von Martens (1901) reports this species from COSTA RICA. Streptostyla (Streptostyla ) turgidula guatemalensis Fischer & Crosse 1870Streptostyla turgidula guatemalensis Fischer & Crosse 1870; Miss. Cent. Amer.:96.Pilsbry 1907; Man. Conch. 19:150. Streptostyla schneideri Streptostyla turgidula producta Pilsbry. Hinkley 1920; Nautilus 34:40, 43, 51 (a nomen nudum).Type Localities.San Agustin, [Dept. Solol], Guatemala (Von Martens 1891). Distribution.GUATEMALA. Dept. Alta Verapaz: Chama (Hinkley 1920). Dept. Izabal: Maya Farms, Quirigua; mountains of Rio Cavech, above Cavech Village (Hinkley 1920). Dept. Retalhuleu: Hacienda Helvetia, 3000 ft. alt.; Hacienda Buenavista, 3500 ft. alt.; Hacienda Germania (Von Martens 1891). Streptostyla (Streptostyla ) nebulosa Dall 1896Streptostyla nebulosa Dall 1896; Proc. U. S. Nat. Mus. 19:364; Type Locality.San Cristobal, Chiapas, Mxico. Holotype USNM 107369. Distribution.Known only from the type locality. Streptostyla (Streptostyla ) sumichrasti Ancey 1903Streptostyla sumichrasti Ancey 1903; Nautilus 17:56.Pilsbry 1907; Type Locality.Isthmus of Tehuantepec. Distribution.OAXACA (?).

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188 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Streptostyla (Streptostyla ) Jacobson 1966 Jacobson 1966; Nautilus 79:102; Type Locality.Quemigas [Quimiquas] or Rio Negro, approximately 20 miles northeast of Bonanza, Zelaya estimated elevation 1300 ft. Distribution.Known only from the type locality. Streptostyla (Streptostyla ) Angas 1879 Angas 1879; Proc. Zool. Soc. Lond. 47:482; Martens 1901; Biol. Cent. Amer.:616.Pilsbry 1907; Man. Nautilus 41:131.Type Locality.Hills of Uren, Costa Rica. Holotype in the ASNSP. Tierra, on road to Sarapiqu, 600 m alt.; Tuis, 600 m alt.; Schukuluk, in Alto Uren; between Mokri and Ukatschka, Alta Talamanca; La Hondura, 1000 m alt.; slopes of Volcn Barba (Von Martens 1901). PANAM, Prov. Bocas del Toro, Mono Creek (Pilsbry 1926). Taxa with Physaeform Shell Streptostyla (Streptostyla ) biconica Pfeiffer 1856Spiraxis biconica Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:378. Streptostyla biconica (Pfeiffer). Fischer & Crosse 1870; Miss. Sci. 79 (shell).Type Locality.Chiapas, Mxico. Distribution.Known only from the type locality. Streptostyla ( Streptostyla) binneyana Crosse & Fischer 1869Streptostyla (Streptostyla) binneyana Crosse & Fischer 1869; Jour. var. binneyana Crosse & Fischer. Von Type Locality.Totonicapam [Totonicapan], [Dept. Totonicapan], Guatemala. Distribution.Known only from the type locality. Streptostyla (Streptostyla ) crassa Strebel 1878Streptostyla crassa Strebel 1878; Beitrag. Mex. Landund Sssw.Streptostyla delibuta var. crassa Strebel. Von Martens 1892; Biol. Cent. Amer.:97.Type Locality.Cobn, [Dept. Alta Verapaz], Guatemala. Distribution.Known only from the type locality. Streptostyla (Streptostyla ) delibuta (Morelet 1851)Glandina delibuta Morelet 1851; Test. Noviss. II:13. Streptostyla delibuta (Morelet). Fischer & Crosse 1870; Miss. Sci. 77, 78, 80 (shell).Hinkley 1920; Nautilus 34:43, 51.Type Locality.Vera Paz, Guatemala. Distribution.GUATEMALA, Dept. Alta Verapaz: Chama (Hinkley 1920). Dept. Izabl: environs of Izabl (Von Martens 1892); mountains of Cavech, above Cavech Village (Hinkely 1920). Von Martens (1901) reports this species from Costa Rica. Streptostyla (Streptostyla ) (Shuttleworth 1852)Spiraxis (Streptostyla ) Shuttleworth 1852; Mitt. Naturf. Ges. Bern 1852:206. (Shuttleworth). Fischer & Crosse 1870; Miss. Sci. Mex. I:39.Tryon 1885; Man. Conch. 1:48; pl. 11, Syntypes: Naturhistorische Museum Bern 18836/2 (Neubert & Gosteli 2003). Streptostyla ( Streptostyla) (Pfeiffer 1861)Spiraxis boucardi Pfeiffer 1861; Proc. Zool. Soc. Lond. 29:24. Streptostyla boucardi (Pfeiffer). Fischer & Crosse 1870; Miss. Sci. 19. (Pfeiffer). Pilsbry 1907; Man. Type Locality.Juquila, Oaxaca, Mxico. Distribution.Known only from the type locality. Streptostyla (Streptostyla ) Crosse & Fischer 1869 Crosse & Fischer 1869; Jour. de Conchyl. Biol. Cent. Amer.:99.Type Locality.Orizaba, Veracruz, Mxico. Distribution.Known only from the type locality. Streptostyla (Streptostyla ) glandiformis Crosse & Fischer 1869Streptostyla glandiformis Crosse & Fischer 1869; Jour. de Conchyl. Von Martens 1892; Biol. Cent. Amer.:99.Pilsbry 1907; Man. Conch. 19:158.H. B. Baker 1941:60.Type Locality.Orizaba, Veracruz, Mxico. 3125 ft. alt. (H. B. Baker 1941). Streptostyla (Streptostyla ) oblonga (Pfeiffer 1856)Spiraxis oblonga Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:378. Streptostyla oblonga (Pfeiffer). Fischer & Crosse 1870; Miss. Sci. Mex. I:57.Von Martens 1892; Biol. Cent. Amer.:100.Pilsbry 1907; Man. Conch. 19:159.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 189 Type Locality.Chiapas, Mxico. Distribution.Known only from the type locality. Streptostyla (Streptostyla ) plicatula Strebel 1878Streptostyla plicatula Strebel 1878; Beitrag. Mex. Landund Sssw.1907; Man. Conch. 19:157.H. B. Baker 1941; Nautilus 55:60.Type Locality.Orizaba, Veracruz, Mxico. Distribution.VERACRUZ: Atoyac, 1300 ft. alt. (H. B. Baker 1941). Streptostyla (Streptostyla ) thomsoni Ancey 1888Streptostyla thomsoni Ancey 1886; Ann. de Malac.:257.Von Martens 1892; Biol. Cent. Amer.:98.Pilsbry 1907; Man. 73:130.Type Locality.Isla de tila, Dept. Islas de la Baha, Honduras. Distribution.BELIZE: Cayo Dist.: Rio Frio Cave, 2 mi. from Augustine (Haas & Solem 1960; Nautilus 73:130). HONDURAS: type locality. Streptostyla (Streptostyla ) Rehder 1942 Rehder 1942; Jour. Wash. Acad. Sci. 32:351 Type Locality.Cervantes, Prov. Cartago, Costa Rica; 1480 m alt. Holotype USNM 536020. Distribution.COSTA RICA, Prov. Cartago: Navarro. Prov. San Jos: Tablazo 1800 m alt.; La Verbina, 1000 m alt. (Rehder 1942). Streptostyla (Streptostyla ) (Morelet 1849) Morelet 1849; Test. Noviss. I:15. (Morelet). Von Martens 1892; Biol. Cent. Pub. Carnegie Inst. Wash. (431):529.Branson & McCoy 1963; Nautilus 76:103).Thompson 1967; Bull. Fla. St. Mus. Streptostyla yucatanensis Pilsbry 1907; Man. Conch. 19:153; pl. 30, Inst. Wash. (431):529.H. B. Baker 1941; Nautilus 55:60. Streptostyla yucatanensis var. distorta Pilsbry 1907; Man. Conch. Type Localities. : environs of Merida, Yucatn, Mxico. Streptostyla yucatanensis : Tekanto, Yucatn, Mxico. Streptostyla yucatanensis var. distorta: Tekanto, Yucatn, Mxico. Distribution.CAMPECHE: many localities (Thompson 1967). QUINTANA ROO. YUCATN: 19.1 mi. SSE of Uman; 0.8 mi. NE of Becanchn (Thompson 1967). Von Martens (1901:615) reported this species from COSTA RICA. Taxa with Aplectiform Shell Streptostyla ( Streptostyla) boyeriana Crosse & Fischer 1869Streptostyla boyeriana Crosse & Fischer 1869, Jour. de Conchyl. Von Martens 1892; Biol. Cent. Amer.:102.Type Locality.Orizaba, Veracruz, Mxico. Distribution.Known only from the type locality. Streptostyla (Streptostyla ) dysoni (Pfeiffer 1846)Achatina dysoni Pfeiffer 1846; Proc. Zool. Soc. Lond. 14:32. Streptostyla dysoni (Pfeiffer). Tryon 1885; Man. Conch. 1:48; pl. Pilsbry 1907, Man. Conch. 19:159.Haas & Solem 1960; Nautilus 73:130.Type Locality.Honduras. Distribution.BELIZE: Cayo Dist.: Rio Frio Cave, 2 mi. from Augustine (Haas & Solem 1960). HONDURAS. NICARAGUA, Dept. Chontales: Castillo Fort (Von Martens 1892). Streptostyla (Streptostyla ) jacobsoni Pilsbry 1951Streptostyla jacobsoni (shell).Type Locality.Km 636, on the road between Santa 1952:112]. Holotype ANSP 187058. Distribution.Known only from the type locality. Streptostyla ( Streptostyla) meridana meridana (Morelet 1849)Glandina meridana Morelet 1849; Test. Noviss. I:15. Streptostyla meridana (Morelet). Fischer & Crosse 1890:32.Von (shell).Thompson 1967; Bull. Fla. St. Mus. 11:238; textStreptostyla Streptostyla maslini Branson & McCoy 1962; Nautilus Streptostyla toltecorum Branson & McCoy 1962; Nautilus 76:8; Type Localities. Glandina meridana: Merida, Yucatn, Mxico. Streptostyla maslini: 19 miles east of Merida, Yucatn; holotype UMMZ 210557. Streptostyla toltecorum: 3 miles east of Campeche, Campeche; holotype UMMZ 210558. Distribution.CAMPECHE: numerous localities (Thompson 1967). QUINTANA ROO: 7.1 mi. NNW of Xiatil. YUCATN: 7.0 mi. SSE of Uan; 1.0 mi. SSE of Puerto Telchac; 0.8 mi. NE of Becanchn; 30 mi. S of Uxmal; 19 mi. E of Merida (Branson & McCoy 1962). GUATEMALA, Dept. Petn: Tikal National Park (Basch 1959). Streptostyla (Streptostyla ) meridana cobanensis (Tristram 1861)Spiraxis cobanensis Tristram 1861; Proc. Zool. Soc. Lond. 29:231. Streptostyla cobanensis (Tristram). Fischer & Crosse 1870; Miss. Sci. Mex. I:52. Streptostyla meridana cobanensis (Tristram). Von Martens 1892; Biol. Cent. Amer.:101.Pilsbry 1907; Man. Conch. 19:155; pl. Streptostyla cornea Crosse & Fischer 1869; Jour. de Conchyl.

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190 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Type Locality.Spiraxis cobanensis: Cobn, [Dept. Alta Verapaz], Guatemala. Streptostyla cornea : Guatemala. Distribution.GUATEMALA, Dept. Solol: San Agustin; Hacinda Helvetia, upper Rio Cholhuitz, 3000 ft. alt. (Von Martens 1891). TABASCO: 0.5.0 mi. E of Teapa (Thompson 1957). Von Martens (1901:616) reports this species from Costa Rica. Streptostyla (Streptostyla ) propinqua Thompson 1963Streptostyla propinqua Thompson 1963; Proc. Biol. Soc. Wash. Type Locality.Hacienda Monte Cristo, Metapan, Dept. Santa Ana, El Salvador; 2200 m alt. Holotype UMMZ 195760. Distribution.Known only from the type locality. Streptostyla ( Streptostyla) sololensis Crosse & Fischer 1869Streptostyla sololensis Crosse & Fischer 1869; Jour. de Conchyl. 1920; Nautilus. 34:51.Haas 1949; Nautilus 62:137.Type Locality.Mountains of Solol, Dept. Solol, Guatemala. Distribution.GUATEMALA, Dept. Alta Verapaz: Chama (Hinkley 1920). Dept. Chimultenango: Ruins of Iximche (Von Martens 1892); W slope of the Volcano Acatenango (Haas 1949). Dept. Sapatepequez: Finca San Rafael (Haas 1949). Von Martens (1901:616) reports this species from Costa Rica. Streptostyla (Streptostyla ) Strebel 1878 Strebel 1878; Beitrag. Mex. Landund Sssw.Amer.:102.H. B. Baker 1941; Nautilus 55:60.Type Locality.Jalapa, Veracruz, Mxico. 3000 ft. alt. (H. B. Baker 1941). Taxa with Subturrite Shell Streptostyla (Streptostyla ) mohriana (Pfeiffer 1862)Spiraxis mohriana Pfeiffer 1862; Malak. Bltt. 9:97. Streptostyla mohriana (Pfeiffer). Fischer & Crosse 1870; Miss. Sci. Mex. I:65.Von Martens 1892; Biol. Cent. Amer.:102.Pilsbry Type Locality.Cerro Borrego, near Orizaba, Veracruz, Mxico. Distribution.Known only from the type locality. Streptostyla (Streptostyla ) pilsbryi Richards 1937Streptostyla pilsbryi Richards 1937; Proc. Amer. Philos. Soc. Type Locality.Near the ruins of San Gerbaio, Isla Cozumel, Quintana Roo. Holotype ANSP 167748. Distribution.QUINTANA ROO: only from the type locality. Streptostyla ( Streptostyla) sargi sargi Crosse & Fischer 1869Streptostyla sargi Crosse & Fischer 1875; Jour. de Conchyl. 1892; Biol. Cent. Amer.:102.Hinkley 1920; Nautilus 34:51.Type Locality.Cobn, Guatemala. Distribution.GUATEMALA, Dept. Alta Verapaz: Chama (Hinkley 1920). Dept. Baja Verapaz: Senahu, 2800 ft. alt. (Von Martens 1892). Streptostyla ( Streptostyla) sargi championi Von Martens 1901Streptostyla sargi var. championi Von Martens 1901; Biol. Cent. Type Locality.Sabo, Dept. Alta Verapaz, Guatemala. Distribution.Known only from the type locality. Streptostyla (Streptostyla ) sargi pallidior Crosse & Fischer 1876Streptostyla sargi var. pallidior Crosse & Fischer 1876; Jour. de (shell).Type Locality.Cobn, Guatemala. Distribution.Known only from the type locality. Subgenus Chersomitra Von Martens 1860Chersomitra Von Martens 1860; in Albers, Die Heliceen:33.Pilsbry 1907; Man. Conch. 19:145.Type Species.Glandina nigricans Pfeiffer 1845. Distribution.From central Veracruz south to Guatemala. Taxonomy.Six species are assigned to the subgenus. Streptostyla (Chersomitra) chiapensis Pilsbry 1909Streptostyla limnaeiformis var. Von Martens 1892; Biol. Spirixis [Streptostyla ] Pfeiffer 1856.) Streptostyla limnaeiformis chiapensis Pilsbry 1909; Man. Conch. 20:111.Bequaert 1957:217.Type Locality.Chiapas, Mxico. Distribution.CHIAPAS: Laguna Ocotal, 950 m alt. (Bequaert 1957). Streptostyla ( Chersomitra) limneiformis (Shuttleworth 1852)Spiraxis (Streptostyla ) limneiformis Shuttleworth 1852; Mit. Natruf. Ges. Bern 1852:206. Streptostyla limneiformis (Shuttleworth). Fischer & Crosse 1870; Miss. Sci. Mex. I:39.Pilsbry 1907; Man. Conch. 19:159.Von Martens 1892; Biol. Cent. Amer.:100. Streptostyla lymneiformis (Shuttleworth). Tryon 1885; Man. Conch.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 191Thompson 1967; Bull. Fla. St. Mus. 11:234.Neubert & Gosteli Type Locality.Cordova [Cordob], Veracruz, Mxico; syntypes: Naturhistorische Museum Bern 18839 (Neubert & Gosteli 2003). Baker 1941) Streptostyla (Chersomitra) lurida (Shuttleworth 1852)Spiraxis (Streptostyla ) lurida Shuttleworth 1852; Mitt. Naturf. Ges. Bern 1852:205. Streptostyla (Chersomitra) lurida (Shuttleworth). Fischer & Pilsbry 1907; Man. Conch. 19:150.Neubert & Gosteli 2003; Streptostyla bocourti Crosse & Fischer 1869; Jour. de Conchyl. 62:137.Type Locality.Spiraxis lurida Veracruz, Mxico; syntypes: Naturhistorische Museum Bern 18838/2 (Neubert & Gosteli 2003). Streptostyla bocourti: Cobn, Guatemala. TABASCO: San Juan Bautista. GUATEMALA, Dept. Alta Verapaz: Cobn (Von Martens 1891). Dept. Sacatepquez: Finca San Rafael (Haas 1949). Von Martens (1901) reports this species from COSTA RICA. Streptostyla ( Chersomitra) mitraeformis (Shuttleworth 1852)Spiraxis (Streptostyla ) mitraeformis Shuttleworth 1852; Mitt. Naurf. Ges. Bern 1852:205. Streptostyla mitraeformis (Shuttleworth). Fischer & Crosse 1870; 1891:90.Von Martens 1901; Biol. Cent. Amer.:614. Streptostyla mitraeformis var. minor Strebel 1878; Beitrag. Mex. Streptostyla (Chersomitra) mitraeformis (Shuttleworth). Neubert & Gosteli 2003; Contributions to Natural History 1:36; pl. 11, Type Locality.Spiraxis mitraeformis: Cordova Museum Bern 18840/3. Streptostyla mitraeformis minor: Cobn, Guatemala. Distribution.TABASCO (Von Martens 1891). Martens 1891). GUATEMALA, Dept. Petn: Lago de Petn. Dept. Alta Verapaz: Cobn (Von Martens 1891). Von Martens (1901:614) reports this species from Costa Rica. Streptostyla (Chersomitra) nigricans (Pfeiffer 1845)Glandina nigricans Pfeiffer 1845; Proc. Zool. Soc. Lond. 13:75. Streptostyla nigricans (Pfeiffer). Fischer & Crosse 1870; Miss. Sci. Cent. Amer.:90.Hinkley 1920; Nautilus 34:50. Streptostyla (Chersomitra) nigricans (Pfeiffer). Thompson 1967; 1C (reproductive system), 1F (kidney).Type Locality.Vera Cruz. Von Martens (1891) suggested that this was in error for Vera Paz. alt. (Thompson 1967). TABASCO: 0.5 mi. E of Teapa (Thompson 1957). GUATEMALA, Dept. Alta Verapaz: Cobn (Von Martens 1891); Chejel, 2000 ft. alt. (Hinkley 1920). Dept. Petn: Lago de Petn (Von Martens 1891) Streptostyla (Chersomitra) physodes (Shuttleworth 1852)Spiraxis (Streptostyla) physodes Shutttleworth 1852; Mitt. Naurf. Ges. Bern 1852:207. Streptostyla physodes (Shuttleworth). Fischer & Crosse 1870; Miss. 4 (reproductive system).Thompson 1967; Bull. Fla. St. Mus. 11:234.Neubert & Gosteli 2003; Contributions to Natural Spiraxis auriculacea Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:320. Streptostyla auriculacea (Pfeiffer). Fischer & Crosse 1870:55Type Localities. Spiraxis physodes: Cordova Museum Bern 18841/3 (Neubert & Gosteli 2003). Spiraxis auriculacea (H. B. Baker 1941); Pacho, near Jalapa; Mirador; Huatusco (Von Martens 1892). Subgenus Eustreptostyla H. B. Baker 1927Eustreptostyla H. B. Baker 1927; Nautilus 41:21.H. B. Baker 1941; Nautilus 55:53.Zilch 1959; Handb. Palaoz., 6 (2):456.Type Species.Spiraxis nicoleti Shuttleworth 1852. Distribution.Northeastern Mxico from Tamaulipas and San Luis Potos south to Central Veracruz. Taxonomy.Six species and two subspecies are recognized. The shell of Eustreptostyla is characterized by having a strongly twisted columella and having strong growth threads below the suture. The type species is large with a thick, solid shell about 30 mm high. Streptostyla (Eustreptostyla) nicoleti nicoleti (Shuttleworth 1852)Spiraxis (Streptostyla) nicoleti Shuttleworth 1852, Mittheilungen der naturforschenden Gessellschaft Bern, 249:204. Spiraxis nicoleti Pfeiffer 1853; Monogr. Helic. Viv., 3:477. Achatina nicoleti Pfeiffer, in Martini & Chemnitz, Syst. Conch.Cab., ed. 2, Achatina Streptostyla nicoleti Shuttleworth. Fischer & Crosse 1870; Miss. Sci. Mex. I:23.Strebel 1878:6, 12, 49; pl. 3 (anatomy); 1891:85.H. B. Baker 1943; Proc. Acad. Nat. Sci. Phila. 95:5;

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192 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Streptosstyla (Eustreptostyla ) nicoleti nicoleti Shuttleworth. syntypes: Naturhistorische Museum Bern 18842/2 (Neubert & Gosteli 2003). Distribution.OAXACA: limestone hill 0.5 km W of El Cedral (18.311 N, 96.303 W); Bosque Natural de Temascal, 0.5 km W of El Cedral,100 m alt.; Bosque Natural de Temascal, 1 km SW of El Cedral, 100 m alt. (Thompson 2009). VERACRUZ: Coatepc; Jalapa (Von Martens 1891); Misantla; San Isidro (Strebel 1878); Sumidero (H. B. Baker 1943). Streptostyla ( Eustreptostyla) nicoleti atypica H. B. Baker 1941Streptostyla nicoleti Streptostyla nicoleti atypica H. B. Baker 1941; Nautilus 55:55.H. B. Baker 1943; Proc. Acad. Nat. Sci. Phila. 95:6.Type Locality.Rio Necaxa Gorge, Puebla, Mxico; 2625 ft. alt. Holotype in the UMMZ. Distribution.PUEBLA: type locality. VERACRUZ: San Juan Miahuatln (Strebel 1877). Streptostyla (Eustreptostyla) Von Martens 1891Streptostyla nicoleti var. Von Martens 1891; Biol. Cent. Amer.:85. Streptostyla nicoleti (Shuttleworth). Fischer & Crosse 1870; Miss. Streptostyla nicoleti (Shuttleworth) form B. Strebel 1875; Beitrag. Veracruz, Mxico. Distribution.VERACRUZ: Orizaba (Strebel 1875). Streptostyla (Eustreptostyla) bartschi Dall 1908Streptostyla bartschi Dall 1908; Proc. U. S. Nat. Mus. 35:178 Reguero 1998; Acta Zool. Mex. (73):16.Correa-Sandoval 2000; Acta Zool. Mex. 79:9.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (79):237.Type Locality.Tamaulipas, Mxico. Holotype USNM 19800. Distribution.SAN LUS POTOS: 6 km. SW of numerous localities in te southern region of the state (CorreaSandoval & Rodriguez 2002); El Cielo Biosphere Reserve (Corra-Sandoval & Rodriguez 2005). VERACRUZ: Rancho Streptostyla (Eustreptostyla) jilitlana Dall 1908Streptostyla jilitlana Dall 1908; Proc. U. S. Nat. Mus. 35:179 Reguero 1998; Acta Zol. Mex. (73):16.Type Locality.Jilitla [Xilitla], San Luis Potos, Mxico. Holotype USNM 107821. Distribution.SAN LUS POTOS: km 140, E of W) (Correa-Sandoval et al. 1998). Streptostyla (Eustreptostyla) rupecula Thompson 2009Streptostyla (Eustreptostyla ) rupecula Thompson 2009; Arch. Type Locality. km northwest of Comalapa, Veracruz, Mxico; 499 m alt. (18.7 N, 96.8667 W). Holotype UF 89102. Distribution.VERACRUZ: known only from the immediate vicinity of the type locality. Streptostyla (Eustreptostyla) toyuca Dall 1908Streptostyla toyuca Dall 1908; Proc. U. S. Nat. Mus. 35:179; pl. 29, Type Locality.Metlal Toyuca [Metlaltoyuca], Puebla, Mxico (20.666 N, 97.816 W). Holotype USNM 107822. Distribution.Known only from the type locality. Streptostyla (Eustreptostyla) supracostata Pilsbry 1910Streptostyla supracostata Pilsbry 1910; Proc. Acad. Nat. Sci. Phila. Cubas & Reguero 1998; Acta Zool. Mex. (730:16.Type Locality.At a cave near San Dieguito, San Luis Potos, Mxico. Lectotype ANSP 99031a (H. B. Baker 1963:219). Distribution.SAN LUIS POTOS: a canyon below Las Canoas; El Ambra (Pilsbry 1909); 6 km SW of Xilitla, 830 m Subgenus Peteniella Pilsbry 1907Petenia Crosse & Fischer 1869; Jour. de Conchyl. 17:35.Fischer & Crosse 1870:66. (Not Petnia Gnther 1862, Pisces). Peteniella Pilsbry 1907; Man. Conch. 19:161.Type Species.Glandina ligulata Morelet 1849. Distribution.Eastern Guatemala, Chiapas and Veracruz, Mxico. Taxonomy.Two species are recognized. Streptostyla (Peteniella ) catenata (Pfeiffer 1856)Spiraxis catenata Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:378. Streptostyla catenata (Pfeiffer). Fischer & Crosse 1870; Miss. Sci.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 193 Type Locality.Chiapas, Mxico. Distribution.CHIAPAS: exact locality unknown. Streptostyla (Peteniella ) ligulata (Morelet 1849)Glandina ligulata Morelet 1849; Test. Noviss. I:12.Morelet Petnia ligulata (Morelet). Crosse & Fischer 1869; Jour. de Conchyl. Streptostyla ligulata (Morelet). Tryon 1885; Man. Conch. 1:50; pl. Streptostyla (Peteniella) ligulata (Morelet). Pilsbry 1907; Man. Conch. 19:161.Hinkley 1920; Nautilus 34:40.Type Locality.Woods of Petn and Palenque. Distribution.GUATEMALA, Dept. Izabal: Livingston (Hinkley 1920). Dept. Petn. CHIAPAS: Palenque. 1892). Subfamily SPIRAXINAE H. B. Baker 1939Spiraxinae H. B. Baker 1939; Nautilus 53:9.Thompson 2010; Rev. Biol. Trop. 58:195.Type Genus.Spiraxis C. B. Adams 1850. Distribution. Spiraxis The subgenera Spiraxis s. s., Repressaxis H. B. Baker 1939, Dignaxis H. B. Baker 1939 and Euspiraxis Pfeiffer 1854 are known from Jamaica and Hispaniola. The subgenus are referred to different genera. Taxonomy.Five genera and seventy-seven species of Spiraxinae are recognized in the study area. Traditionally Spiraxis has been used as an all-inclusive genus for species of to different genera on the basis of shell morphology. H. B. Baker (1939b) discussed the anatomy of the subfamily. He emphasized that the subgenera and sections he recognized should more properly be ranked as genera and subgenera on the basis of anatomical differentiation. This ranking was followed by Zilch (1960) and is adapted here. The large number of species assigned to Volutaxis and Pseudosubulina remains arbitrary in many cases because of the lack of anatomical information. See Thompson (2010) for a more detailed discussion of the taxonomy of this subfamily. Genus Mayaxis Thompson 1995Mayaxis Thompson 1995; Bull. Fla. Mus. Nat. Hist. 39:79.Thompson 2010; Rev. Biol. Trop. 58:197.Type Species.Myaxis leei Thompson 1995. Distribution.Chiapas and Tabasco, Guatemala and northern Honduras. Taxonomy.Nine species are placed in the genus. Previously, three species, Myaxis cilindrella Myaxis porrecta and Myaxis stolli Subulina. They are now placed in Myaxis on the basis of shell morphology and for biogeographic reasons Mayaxis chiapensis (Pfeiffer 1856)Achatina chiapensis Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:379. Subulina chiapensis (Pfeiffer). Fischer & Crosse 1877:637; pl. 26, Pseudosubulina chiapensis (Pfeiffer). Strebel 1882; Beitrag. Mex. Biol. Cent. Amer.:303.Pilsbry 1907; Man. Conch. 19:3. Mayaxis chiapensis (Pfeiffer). Thompson 1995; Bull. Fla. Mus. Nat. Hist. 39:82.Type Locality.Chiapas, Mxico. Distribution.CHIAPAS: known from the type locality. A record of this species from Guatemala is questioned by Von Martens (1898:303). Mayaxis cylindrella (Morelet 1851)Achatina cylindrella Morelet 1851; Test. Noviss. II:12. Subulina cylindrella (Morelet). Fischer & Crosse 1878:634; pl. 25, Nautilus 62:136, 138.Type Locality.Woods of Petn, near San Luis, Guatemala. Distribution.GUATEMALA, Dept. Chimaltenango: Finca Panajabal; Yabocapa. Dept. Escuintla: Zapote (Haas 1949). Mayaxis fortis (Von Martens 1898)Pseudosubulina fortis Von Martens 1898; Biol. Cent. Amer.:304; pl. Myaxis fortis (Von Martens). Thompson 1995; Bull. Fla. Mus. Nat. Hist. 39:82.Type Locality.Restricted to El Reposo, western Guatemala, 800 ft. alt. (Thompson 1995). Distribution.GUATEMALA, Dept. Escuintla: Miramar. Dept. Jutiapa: Zapote. Dept. Quetzaltenango: Mercedes. Dept. Retalhuleu: near San Francisco (Von Martens 1898). Mayaxis leei Thompson 1995Myaxis leei 51 (reproductive system), 52 (penis), 53 (radula), 54 (shell).Type Locality.Cerro Santa Barbara, Finca Las Quebradas, 3 km W of Nueva Esperanza, Dept. Santa Brbara, Honduras; 1280 m alt. Holotype UF 193387. Distribution.HONDURAS: known only from the immediate vicinity of the type locality. Mayaxis lirifera (Morelet 1851)Achatina lirifera Morelet 1851; Test. Noviss. II:12. Subulina lirifera 12. Pseudosubulina lirifera (Morelet). Von Martens 1898; Biol. Cent. Myaxis lirifera (Morelet). Thompson 1995; Bull. Fla. Mus. Nat. Hist. 39:83.Type Locality.Woods of Petn, near San Lus, Guatemala. Distribution.GUATEMALA, Dept. Petn: type

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194 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) locality. Dept. Izabal: Livingston (Von Martens 1898). Mayaxis martensiana (Pilsbry 1920)Pseudosubulina martensiana Pilsbry 1920; Proc. Acad. Nat. Sci. Myaxis martensiana (Pilsbry). Thompson 1995; Bull. Fla. Mus. Nat. Hist. 39:82.Type Locality.Mountains west of Livingston, Dept. Izabal, Guatemala. Distribution.GUATEMALA, Dept. Izabal: Las Escobas; mountains of Rio Cavech, above Cavech Village (Hinkley 1920). Mayaxis mitescens (Von Martens 1898)Pseudosubulina mitescens Von Martens 1898; Biol. Cent. Amer.:304; 10.Hinkley 1920; Nautilus 34:52. Myaxis mitescens (Von Martens). Thompson 1995; Bull. Fla. Mus. Nat. Hist. 39:82.Type Locality.Dueas, near Antigua, Dept. Sacatepeque, Guatemala; 5000 ft. alt. Distribution.GUATEMALA, Dept. Alta Verapaz: Chama (Hinkley 1920). Mayaxis porrecta (Von Martens 1898)Subulina porrecta Von Martens 1898; Biol. Cent. Amer.:300; pl. 17, 23 (shell).Type Locality.Teapa, Tabasco, Mxico. Distribution.Known only from the type locality. Mayaxis stolli (Von Martens 1898)Subulina stolli Von Martens 1898; Biol. Cent. Amer.:300; pl. 17, (shell).Type Locality.Vera Paz, Guatemala. Distribution.Known only from the type locality. Genus Miraradula H. B. Baker 1939Miraradula H. B. Baker 1939; Nautilus 53:10.Thompson 2010; Rev. Biol. Trop. 56:196.Type Species.Volutaxis similaris Strebel 1882. Distribution.Southeastern Mxico in the states of Puebla, Veracruz and Chiapas. Taxonomy.Two species are recognized. Miraradula miradornsis (Strebel 1882)Volutaxis miradorensis Strebel 1882; Beitrag. Mex. Landund Spiraxis miradorensis (Strebel). Von Martens 1898; Biol. Cent. (shell). Spiraxis (Miraradula ?) miradorensis (Strebel). H. B. Baker 1940; Nautilus 53:93.Type Locality.Mirador, Veracruz, Mxico. Distribution.VERACRUZ: Peuela to Sumidero [14 Miraradula similaris (Strebel 1882)Volutaxis similaris Strebel 1882; Beitrag. Mex. Landund Sssw.Spiraxis similaris (Strebel). Von Martens 1898; Biol. Cent. Amer.:310. Spiraxis (Miraradula) similaris (Strebel). H. B. Baker 1939; Spiraxis similaris (Strebel). Bequaert 1957:217.Type Locality.Pacho, near Jalapa, Veracruz, Mxico. Distribution.CHIAPAS: Laguna Ocotal to El Censo, 600 m alt. (Bequaert 1957). PUEBLA: Necaxa (H. B. Baker 1939b). VERACRUZ: Peuela to Sumidero, 2625 3400 ft. alt. (H. B. Baker 1940). Genus Pseudosubulina Strebel 1882Pseudosubulina Strebel 1882; Beitrag. Mex. Landund Sssw.Conch. V:117.Pilsbry 1907; Man. Conch. 19:1.H. B. Baker 1939; Nautilus 53:11.Thompson 2010; Rev. Biol. Trop. 58:197.Type Species.Pseudosubulina berendti Strebel 1882. Distribution.Mxico and Guatemala; a single species, Pseudosubulina exilis (Pfeiffer 1839), occurs in Cuba. Taxonomy.Twenty-two species and four subspecies are recognized in the study area. Pseudosubulina arcuata (H. B. Baker 1939)Spiraxis (Pseudosubulina) arcuatus H. B. Baker 1939; Nautilus Veracruz, Mxico. Holotype in the UMMZ. Distribution.Known only from the type locality. Pseudosubulina berendti berendti (Pfeiffer 1862)Achatina berendti Pfeiffer 1862; Malak. Bltt. 9:98. Subulina berendti (Pfeiffer). Fischer & Crosse 1877:635; pl. 26, Pseudosubulia berendti (Pfeiffer). Strebel 1882; Beitrag. Mex. (reproductive system).Tryon 1885; Man. Conch. 1:50; pl. 9, 1907; Man. Conch. 19:3.H. B. Baker 1926b:9.H. B. Baker Nautilus 53:94.Type Locality.Orizaba, Veracruz, Mxico. Distribution.PUEBLA: Necaxa, 3000 ft. alt. (H. B. Baker 1940:94). VERACRUZ: Pacho; Soncoautla; Consolapa; Dos Arroyos; Las Vigas (Von Martens 1898). Pseudosubulina berendti gracilior von Martens 1898Pseudosubulina berendti form B. Strebel 1882; Beitrag. Mex. LandPseudosubulina berendti var. gracilior von Martens 1898; Biol. Cent. Amer.:303.Type Locality.Veracruz, Veracruz, Mxico.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 195 Martens 1898). Pseudosubulina berendti ocidentalis Pilsbry 1899Pseudosubulina berendti occidentalis Pilsbry 1899; Proc. Acad. Nat. Sci. Phila. 51:398.Pilsbry 1903; Proc. Acad. Nat. Sci. Phila. Type Locality.Uruapam [Uruapan], Michoacn, Mxico. Syntypes ANSP 77161. Distribution.Known only from the type locality. Pseudosubulina borealis (Pilsbry 1903)Spiraxis (?) borealis Pilsbry 1903; Proc. Acad. Nat. Sci. Phila. Mxico. Syntypes ANSP 77166. Distribution.Known only from the type locality. PUEBLA: Necaxa, 4600 ft. alt (H. B. Baker 1940). Pseudosubulina caduca (H. B. Baker 1939)Spiraxis (Pseudosubulina) caducus H. B. Baker 1939a:133; pl. 4, Type Locality.Las Tortolas [2 km. N of east end of] the UMMZ. Distribution.Known only from the type locality. Pseudosubulina cheatumi Pilsbry 1950Pseudosubulina cheatumi Pilsbry 1950; Nautilus 64:55 (not Type Locality.Casa Grande Peak, Chisos Mountains, Texas. Holotype ANSP 186649. Distribution.Known only from the type locality. This species is included because of the proximity of the type locality to Mxico. Pseudosubulina costata (H. B. Baker 1939)Spiraxis (Pseudosubulina) costatus H. B. Baker 1939a:133; pl. Veracruz, Mxico; 1300 ft. alt. Holotype in the UMMZ. 3000 ft. alt. (H. B. Baker 1940). Pseudosubulina eiseniana (Cooper 1893)Melaniella eiseniana Cooper 1893; Proc. Cal. Acad. Sci. 3:339; pl. Pseudosubulina eiseniana (Cooper). Dall 1896; Proc. U. S. Nat. (map).Type Locality.Under stones, Cape region, Baja California Sur, Mxico. Distribution.BAJA CALIFORNIA SUR: several localities on southern tip of peninsula (Smith et al. 1990). Dall 1926 Dall 1926; Proc. Calif. Acad. Sci., ser. 4 Type Locality.Cerro Evermann, Isla Socorro, Nayarit, Mxico; 2000 ft. alt. Holotype CAS 2191. Distribution.NAYARIT: known only from the type locality of Isla Socorro. Pseudosubulina insularis Pilsbry 1930Pseudosubulina insularis Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. Type Locality.Isla de Providencia, western slope below the cleft of Split Hill, Colombia. Holotype ANSP 150172. Distribution.COLOMBIA: known only from the immediate vicinity of the type locality, Isla de Providencia. Pseudosubulina irregularis irregularis Pilsbry 1907Pseudosubulina irregularis Pilsbry 1907; Man. Conch. 19:7; pl. 6, Type Locality.Texola, Veracruz, Mxico. Holotype ANSP 77147a. Distribution.Known only from the type locality. Pseudosubulina irregularis negligens (H. B. Baker 1939)Spiraxis (Pseudosubulina) irregularis negligens H. B. Baker (shell).H. B. Baker 1940; Nautilus 53:94.Type Locality.Necaxa, Puebla, Mxico; 5000 ft. alt. Holotype in the UMMZ. Distribution.PUEBLA: known only from the vicinity of the type locality (H. B. Baker 1940). Pseudosubulina martensiana Pilsbry 1920Pseudosubulina martensiana Pilsbry 1920; Proc. Acad. Nat. Sci. Type Locality.Mountains west of Livingston, [Dept. Izabal], Guatemala. Holotype ANSP 107535. Distribution.Known only from the type locality. Pseudosubulina orizabensis Pilsbry 1907Pseudosubulina orizabensis Pilsbry 1907; Man. Conch. 19:7; pl. 2, Type Locality.Orizaba, Veracruz, Mxico. Holotype ANSP 61545. alt. (H. B. Baker 1940). (H. B. Baker 1939)Spiraxis (Pseudosubulina) H. B. Baker 1939; Nautilus B. Baker 1940; Nautilus 53:94.Type Locality.Necaxa, Puebla, Mxico; 5000 ft. alt. Holotype in the UMMZ.

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196 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Distribution.VERACRUZ: known from the vicinity of the type locality (H. B. Baker 1940). Pseudosubulina robusta von Martens 1898Pseudosubulina robusta von Martens 1898; Biol. Cent. Amer.:304; Type Locality.Omilteme, Guerrero, Mxico; 8000 ft. alt. Distribution.Known only from the type locality. Pseudosubulina ruthae Pilsbry 1954Pseudosubulina ruthae Type Locality.On the beach at False Bay above high tide level, California. Holotype ANSP 190114. Distribution.BAJA CALIFORNIA NORTE(?): unknown. Pseudosubulina von Martens 1898Pseudosubulina von Martens 1898; Biol. Cent. Amer.:305; 36 (shell).Pilsbry 1907; Man. Conch. 19:6.Hinkley 1920; Nautilus 34:52.Type Locality.Vera Paz, Guatemala. Distribution.GUATEMALA, Dept. Alta Verapaz: Chama (Hinkley 1920). Pseudosubulina sargi Crosse & Fischer 1877Subulina sargi Crosse & Fischer 1877; Jour. de Conchyl. 25:272.Pseudosubulina sargi (Crosse & Fischer). Von Martens 1898; Biol. Cent. Amer.:303.Type Locality.Cobn, Dept. Alta Verapaz, Guatemala. Distribution.GUATEMALA: type locality. TABASCO: Teapa (Von Martens 1898). Pseudosubulina splendens Thompson 1959Pseudosubulina splendens Thompson 1959; Nautilus 72:115; pl. 12, Type Locality.Cobn-Sebol Road, 55 miles northeast of Cobn, Guatemala. Holotype UMMZ 195985. Distribution.Known only from the type locality. Pseudosubulina tastensis (Cooper 1894)Melaniella tastensis Cooper 1894; Proc. Calif. Acad. Sci. 4:141; pl. Pseudosubulina tastensis (Cooper). Pilsbry 1907; Man. Conch. Proc. Calif. Acad. Sci. 47:132.Type Locality.Saltito Pass, just north of the Sierra El Taste, 3200 ft. alt., Baja California Sur, Mxico. Distribution.BAJA CALIFORNIA SUR: numerous localities on southern tip of peninsula (Smith et al. 1990). Pseudosubulina texoloensis Pilsbry 1899Pseudosubulina texoloensis Pilsbry 1899; Proc. Acad. Nat. Sci. Phila. 51:398.Pilsbry 1903; Proc. Acad. Nat. Sci. Phila. Type Locality.Texolo, Veracruz, Mxico. Holotype ANSP 77104a. Distribution.Known only from the type locality. Pseudosubulina trypanodes (Pfeiffer 1856)Achatina trypanodes Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:379. Pseudosubulina trypanodes (Pfeiffer). Von Martens 1898; Biol. Type Locality.Mxico. Distribution.Unknown. Pseudosubulina (H. B. Baker 1939)Spiraxis (Pseudosubulina) H. B. Baker 1939; Nautilus (shell).H. B. Baker 1940; Nautilus 53:94.Type Locality.Tenango Hills [3 km S of] Necaxa, Puebla, Mxico; 4000 ft. alt. Holotype in the UMMZ. Distribution.Known only from the type locality. Genus Micromena H. B. Baker 1939Micromena H. B. Baker 1939; Nautilus 53:11.Thompson 2010; Rev. Biol. Trop. 58:196.Type Species.Spiraxis (Micromena) minutus H. B. Baker 1939. Distribution.Jamaica and Mxico. Taxonomy.Three species are recognized, two in the study area. Baker (1939) proposed Micromena as a subgenus of Pseudosubulina. Thompson (2010) raised it to full generic rank. Micromena minuscula (H. B. Baker 1940)Spiraxis (Micromena) minusculus H. B. Baker 1940; Nautilus 53:92, Pseudosubulina (Micromena) minuscule (H. B. Baker). Thompson 2008: 651.Type Locality.Below Necaxa, Puebla, Mxico; 2625 ft. alt. Holotype in the UMMZ. doba, 2650 ft. alt. Micromena minuta (H. B. Baker 1939)Spiraxis (Micromena) minutus H. B. Baker 1939; Nautilus 53:14 Pseudosubulina (Micromena) minuta (H. B. Baker). Thompson 2008:651.Type Locality.Below Necaxa, Puebla, Mxico; 3000 ft. alt. Holotype in the UMMZ. Distribution.PUEBLA: known only from the immediate vicinity of the type locality (H. B. Baker 1940). Genus Rectaxis H. B. Baker 1926Spiraxis (Rectaxis ) H. B. Baker 1926; Occasional papers of the Museum of Zoology, University of Michigan, (167):7.H. B. Baker 1939; Nautilus 53:101.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 197Rectaxis H. B. Baker. Zilch 1959; Handbuch der Paleozoologie, 6 (2):448.Thompson 2010; Rev. Biol. Trop. 58:196.Type Species.Pseudosubulina (Rectaxis ) decussatus H. B. Baker 1926; (Venezuela). Distribution.Venezuela north to Veracruz. Taxonomy.Twelve species are recognized (Thompson 2010). (Goodrich & van der Schalie 1937) Goodrich & van der Schalie 1937; Misc. Pub. 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):9. El Paso de Los Caballos, Dept. Petn, Guatemala. Holotype in the UMMZ. Distribution.GUATEMALA, Dept. Petn: Tikal National Park (Basch 1959). Rectaxis canalizonalis (Pilsbry 1930)Pseudosubulina canalizonalis Pilsbry 1930; Proc. Acad. Nat. Sci. Spiraxis canalizonalis (Pilsbry). H. B. Baker 1940; Nautilus 53:92.Baker 1963; Proc. Acad. Nat. Sci. Phila. 115:218.Thompson Type Locality.Roadside bank southeast of Empire, [Isla Barro Colorado], Panam. Holotype ANSP 151350 (lost; see Thompson 2010). Distribution.COSTA RICA, Prov. Cartago InterAmerican Agricultureal Institute, Turrialba (Thompson 2010). PANAM: in addition to the type locality, ruins of Old Panam City (Pilsbry 1930). Prov. Darin 6.5 km W of Boca de Cope (Thompson 2010). Rectaxis confertestriatus (Strebel 1882)Volutaxis confertestriatus Strebel 1882; Beitrag. Mex. Landund Spiraxis confertestriatus (Strebel). Von Martens 1898; Biol. Cent. (shell). Spiraxis (Rectaxis ) confertestriatus (Strebel).H. B. Baker 1939; Nautilus 53:50.Type Locality.Mirador, Veracruz, Mxico. Distribution.Known only from the type locality. Rectaxis funibus (Goodrich & van der Schalie 1937)Spiraxis funibus Goodrich & van der Schalie 1937; Misc. Pub. Mus. Occ. Pap. Mus. Zool. Univ. Mich. (612):9. El Paso de Los Caballos, Dept. Petn, Guatemala. Holotype in the UMMZ. Distribution.GUATEMALA, Dept. Petn: Tikal National Park (Basch 1959). Rectaxis granum (H. B. Baker 1939)Spiraxis (Rectaxis ) granum H. B. Baker 1939; Nautilus 53:49; Type Locality.Below Necaxa, Puebla, Mxico. Holotype in the UMMZ. Distribution.PUEBLA. VERACRUZ: Peuela, 2700 ft. alt. (H. B. Baker 1940). Rectaxis intermedius (Strebel 1882)Volutaxis intermedius Strebel 1882; Beitrag. Mex. Landund Spiraxis intermedius (Strebel). Von Martens 1898; Biol. Cent. Amer.:310. Spiraxis (Rectaxis) intermedius (Strebel). H. B. Baker 1939; Nautilus 53:13.H. B. Baker 1940; Nautilus 53:94.Type Locality.Pacho, near Jalapa, Veracruz, Mxico. Distribution.VERACRUZ: Peuela to Sumidero [14 Rectaxis pagodus Thompson 2010Rectaxis pagodus (shell).Type Locality.La Lola Agriculture Experimental Holotype UF 80842. Distribution.Known only from the type locality. Rectaxis paulisculpta (Rehder 1942)Spiraxis (Rectaxis ) paulisculpta Rehder 1942; Jour. Wash. Acad. Rectaxis paulisculpta (Rehder). Thompson 2010; Rev. Biol. Trop. Type Locality.Santa Maria, Prov. San Jos, Costa Rica; 1550 m alt. Holotype USNM 536016. Distribution.Known only from the type locality. Rectaxis pittieri (Von Martens 1898)Spiraxis nitidus var. pittieri Von Martens 1898; Biol. Cent. Type Locality.El Pital, in the valley of the Rio Naranjo, Costa Rica; 250 m alt. Distribution.COSTA RICA: type locality; Savana de Guacino, valley of the Rio Brus (Von Martens 1898). Rectaxis rhabdus (Pilsbry 1907)Spiraxis rhabdus 26.H. B. Baker 1939; Nautilus 53:50.Type Locality.Texolo, Veracruz, Mxico. Syntypes in the ANSP. Distribution.Known only from the type locality. Rectaxis subnitidus (H. B. Baker 1939)Spiraxis (Rectaxis ) subnitidus H. B. Baker 1939; Nautilus 53:51: 53:93.Type Locality.Above Necaxa, Puebla, Mxico; 5000 ft. alt. Holotype in the UMMZ.

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198 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Distribution.Known only from the type locality. Rectaxis subtilis subtilis (H. B. Baker 1939)Spiraxis (Rectaxis ) subtilis H. B. Baker 1939; Nautilus; 53:50; pl. 9, Type Locality.Las Tortolas [2 km. N of east end of], the UMMZ. Distribution.Known only from the type locality. (H. B. Baker 1939)Spiraxis (Rectaxis) H. B. Baker 1939; Nautilus 53:93.Type Locality.Below Necaxa, Puebla, Mxico; 2625 ft. alt. Holotype in the UMMZ. Distribution.Known only from the type locality. Genus Volutaxis Strebel 1882Volutaxis Strebel 1882; Beitrag. Mex. Landund Sssw.-Conch. V:119.Pilsbry 1907; Man. Conch. 19:20. H. B. Baker 1939; Nautilus 53:101. Zilch 1960:448.Thompson 2010; Rev. Biol. Trop. 58:196, 200.Type Species.Bulimus sulciferus Morelet 1851. Distribution.Mxico, Central America and the Greater Antilles. Taxonomy.Volutaxis consists of three subgenera: Volutaxis Mirapex, and Versutaxis (Thompson 2010). Subgenus Volutaxis Strebel 1882 Distribution.Mxico and Guatemala south to Costa Rica. Additional species occur in Cuba, Jamaica and Hispaniola. Taxonomy.Twenty species and four subspecies occur in the study area. Volutaxis (Volutaxis) blandiana (Pilsbry 1909)Spiraxis blandi Crosse & Fischer 1877; Jour. de Conchyl. 25; 217.Martens 1898; Biol. Cent. Amer.:311 (not Spiraxis ( ) blandi Crosse 1873). Volutaxis blandi (Crosse & Fischer). Tryon 1885; Man. Conch. 1: Spiraxis blandiana Pilsbry 1909; Man. Conch. 20:111.Type Locality.State of Veracruz, Mxico. Distribution.Known only from the type locality. Volutaxis (Volutaxis) cacahuamilpensis (Herrera 1891)Spiraxis cacahuamilpensis Herrera 1891; Memorias y Revista de (shell).Type Locality.Caves of Cacahuamilpa, Guererro, Mxico. Distribution.Known only from the type locality. Taxonomy.The type specimen is a juvenile consisting only of the initial four whorls of the shell. Volutaxis (Volutaxis) confertecostatus Strebel 1882Volutaxis confertecostatus Strebel 1882; Beitrag. Mex. Landund Spiraxis confertecostatus (Strebel). Von Martens 1898; Biol. Cent. Amer.:310.Type Locality.Pacho and Dos Arroyos, near Jalapa, Veracruz, Mxico. Distribution.Known only from the type locality. Volutaxis (Volutaxis) delicatus (Pilsbry 1907)Spiraxis delicatus 23 (shell).Type Locality.Uruapam [Uruapan], Michoacn, Mxico. Holotype ANSP 77099a. Distribution.MICHOACN: type locality. VERACRUZ: Texolo (Pilsbry 1907). Volutaxis (Volutaxis) eburneus Thompson 2010Volutaxis (Volutaxis ) eburneus Thompson 2010; Rev. Biol. Trop. Type Locality.Hitoy Cerera Biological Preserve, La Distribution.COSTA RICA: known only from the vicinity of the type locality. Volutaxis (Volutaxis) fallax (H. B. Baker 1940)Spiraxis (Volutaxis) fallax H. B. Baker 1940; Nautilus 53:90; pl. 11, Holotype in the UMMZ. 2625 ft. alt. PUEBLA: Necaxa (H. B. Baker 1940). Volutaxis (Volutaxis) linearis (Pfeiffer 1866)Spiraxis linearis Pfeiffer 1866; Malak. Bltt. 13; 84.Fischer & Biol. Cent. Amer.:311. Volutaxis linearis Strebel 1882; Beitrag. Mex. Landund Sssw.Type Locality.Veracruz, Veracruz, Mxico; on the beach under pieces of wood cast ashore. Distribution.Known only from the type locality. Volutaxis (Volutaxis) (Pilsbry 1920) Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 34:44.Type Locality.Mountains west of Livingston [Dept, Izabal], Guatemala. Holotype ANSP 17537a. Distribution.GUATEMALA, Dept. Izabal: mountains of Rio Cavech, above Cavech Village (Hinkley 1920). Volutaxis (Volutaxis) longior (Pilsbry 1920)Spiraxis longior Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 71:214;

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 199 Type Locality.Mountains west of Livingston, [Dept. Izabal], Guatemala. Holotype ANSP 107525. Distribution.GUATEMALA, Dept. Izabal : mountains of Rio Cavech, above Cavech Village (Hinkley 1920). Volutaxis (Volutaxis) maya (Bequaert & Clench 1931)Spiraxis (Volutaxis) maya Bequaert & Clench 1931; Occ. Pap. Boston Soc. Nat. Hist. 5:423.Bequaert & Clench 1933; Pub. Type Locality.Chichen Itza, Yucatn, Mxico. Holotype MCZ 85799. Distribution.Known only from the type locality. Volutaxis (Volutaxis) nitidus nitidus Strebel 1882Volutaxis nitidus Strebel 1882; Beitrag. Mex. Landund Sssw.Spiraxis nitidus (Strebel). Von Martens 1898; Biol. Cent. Amer.:310. Spiraxis nitidus var. major Von Martens 1898; Biol. Cent. Amer.:310; Spiraxis nitidus var. minor Von Martens 1989:310.Type Locality.Pacho and Mirador, Veracruz, Mxico. Distribution.VERACRUZ: Las Vigas; Camino de Obispo (Von Martens 1898). Taxonomy.Von Martens (1898:310) recognized var. minor Von Martens 1898 and var. major Von Martens 1898. The two names have been used numerous times in this and other families as descriptors of variation. Thus, they are not accorded formal taxonomic status. Volutaxis ( Volutaxis) nitidus persulcatus (H. B. Baker 1940)Spiraxis (Volutaxis) nitidus persulcatus H. B. Baker 1940; Nautilus Type Locality.Below Necaxa, Puebla, Mxico; 2625 ft. alt. Holotype in the UMMZ. Distribution.PUEBLA: Necaxa. VERACRUZ: Peuela 2625 ft. alt. (H. B. Baker 1940). Volutaxis (Volutaxis) rhoadsae (Pilsbry 1899)Opeas rhoadsae Pilsbry 1899; Proc. Acad. Nat. Sci. Phila. 51:399.(shell).Naranjo-Garcia & Fahy 2010; Amer. Malaco. Bull., 28:66. Spiraxis rhoadsae (Pilsbry). Pilsbry 1907; Man. Conch. 19:26; pl. Mxico. Lectotype ANSP 77103a (Baker 1963:219). Distribution.Known only from the type locality. Volutaxis (Volutaxis) scalariopsis (Morelet 1851)Bulimus scalariopsis Morelet 1851; Test. Noviss. II:11. Spiraxis scalariopsis (Morelet). Fischer & Crosse, 1877:609; pl. 25, Thompson 1957; Nautilus 70:101.Bequaert 1957:217.Type Locality.Petn, Guatemala. Distribution.Known only from the type locality. CHIAPAS: Monte Lebano to El Censo, 600 m alt. (Bequaert 1957). TABASCO: 0.5.0 mi. E of Teapa (Thompson 1957). Volutaxis (Volutaxis) scalella (Von Martens 1898)Spiraxis scalella Von Martens 1898; Biol. Cent. Amer.:311; pl. 18, 16 (shell).Type Locality.El Pital, in the valley of the Rio Naranjo [Prov. Puntarenas ?], Costa Rica; 200 m alt. Distribution.Known only from the type locality. Volutaxis (Volutaxis) strebeli (Pilsbry 1907)Spiraxis strebeli 28 (shell).Type Locality.Texolo, Veracruz, Mxico. Holotype ANSP 77165a. Distribution.Known only from the type locality. Volutaxis (Volutaxis) subulinus (H. B. Baker 1940)Spiraxis (Volutaxis ) subulinus H. B. Baker 1940; Nautilus 53:90; Type Locality.Above Necaxa, Puebla, Mxico; 5000 ft. alt. Holotype in the UMMZ. Distribution.Known only from the type locality. Volutaxis (Volutaxis) sulciferus sulciferus (Morelet 1851)Bulimus sulciferus Morelet 1851; Test. Noviss. II:12. Spiraxis sulciferus (Morelet). Fischer & Crosse 1877:610; pl. 25, Spiraxis (Volutaxis ) sulciferus sulciferus (Morelet). H. B. Baker Nautilus 53:93. Volutaxis sulciferus (Morelet). Strebel 1882; Beitrag. Mex. Land(shell). Volutaxis (Volutaxis) sulciferus sulciferus (Morelet).Thompson Type Locality.Palenque, Chiapas, Mxico. Distribution.BELIZE: Cayo Dist. Rio Frio Cave, 2 mi. from Augustine (Haas & Solem 1960). GUATEMALA, Dept. Retalhuleu: Retalhuleu (Von Martens 1898). CHIAPAS: Palenque. OAXACA: 9.2 km NE of Valle Nacionl, 250 m alt. (Thompson 2010). VERACRUZ: localities near Misantla; Mirador (Von Martens 1898); Atoyac, [20 km. ESE alt. (H. B. Baker 1940). Volutaxis (Volutaxis) sulciferus atoyacensis (H. B. Baker 1940)Spiraxis (Volutaxis ) sulciferus atoyacensis H. B. Baker 1940; Type Locality.Atoyac, Veracruz, Mxico; 1300 ft. alt. Holotype in the UMMZ. Distribution.VERACRUZ: Atoyac, [20 km. ESE of];

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200 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Volutaxis (Volutaxis) sulciferus cobanensis (Fischer & Crosse 1877)Spiraxis sulciferus var. cobanensis Fischer & Crosse 1877:610; Amer.:308.Type Locality.Cobn, Dept. Alta Verapaz, Guatemala. Distribution.Known only from the type locality. Volutaxis ( Volutaxis) tenuecostatus tenuecostatus Strebel 1882Volutaxis (Volutaxis ) tenuecostatus Strebel 1882; Beitrag. Mex. Spiraxis (Volutaxis ) tenuecostatus tenuecostatus H. B. Baker 1940; Nautilus 53:94.Type Locality.Agua Caliente, near Misantla, Veracruz, Mxico. Distribution.PUEBLA: Necaxa (H. B. Baker 1940). VERACRUZ: type locality. Volutaxis ( Volutaxis) tenuecostatus obesus (H. B. Baker 1940)Spiraxis (Volutaxis) tenuecostatus obesus H. B. Baker 1940; Type Locality.Tenango Hills, near Necaxa, Puebla, Mxico; 4600 ft. alt. Holotype in the UMMZ. Distribution.Known only from the type locality. Volutaxis (Volutaxis) tenuis (Pfeiffer 1868)Spiraxis tenuis Pfeiffer 1868: Malak. Bltt. 15:84.Fischer & Crosse Type Locality.Orizaba, Veracruz, Mxico. Distribution.Known only from the type locality. Volutaxis (Volutaxis) uruapamensis (Pilsbry 1899)Spiraxis uruapamensis Pilsbry 1899; Proc. Acad. Nat. Sci. Phila. 51:398.Pilsbry 1903; Proc. Acad. Nat. Sci. Phila. 55:775; pl. 12 (shell).Type Locality.Uruapam [Uruapan], Michoacn, Mxico. Holotype ANSP 77160a. Distribution.Known only from the type locality. Subgenus Mirapex H. B. Baker 1939Mirapex H. B. Baker 1939; Nautilus 53:13.Thompson 2010; Rev. Biol. Trop. 58:200.Type Species.Spiraxis (Mirapex) acus enigmaticus H. B. Baker 1939. Distribution.Central Veracruz at moderate elevations Taxonomy.Mirapex includes a single species. Volutaxis (Mirapex) enigmaticus (H. B. Baker 1939)Spiraxis (Mirapex) acus enigmaticus H. B. Baker 1939; Nautilus B. Baker 1940; Nautilus 53:93.Type Locality.Las Tortolas [2 km. N of east end of], UMMZ. Distribution.Known only from the type locality. Subgenus Versutaxis H. B. Baker 1939Versutaxis H. B. Baker 1939; Nautilus 53:33.Thompson 2010; Rev. Biol. Trop. 58:200.Type Species. Spiraxis (Versutaxis) opeas H. B. Baker 1939. Distribution.Central Mxico, including Michoacn, Puebla, and Veracruz. Taxonomy.Seven species are recognized (Thompson 2010). Volutaxis (Versutaxis) arctatus (H. B. Baker 1940)Spiraxis (Versutaxis) arctatus H. B. Baker 1940; Nautilus 53:84; pl. Type Locality.Below Necaxa, Puebla, Mxico; 2625 ft. alt. Holotype in the UMMZ. Distribution.Known only from the type locality. Volutaxis (Versutaxis) futilis (H. B. Baker 1939)Spiraxis (Versutaxis) futilis H. B. Baker 1939; Nautilus 53:52; pl. 9, Type Locality.Tenago Hill, Necaxa, Puebla, Mxico; 4600 ft. alt. Holotype in the UMMZ. Distribution.PUEBLA: known only from the immediate vicinity of the type locality. Volutaxis (Versutaxis) opeas (H. B. Baker 1939)Spiraxis (Versutaxis) opeas H. B. Baker 1939; Nautilus 53:13; pl. 3, Type Locality.Above Necaxa, Puebla, Mxico; 5000 ft. alt. Holotype in the UMMZ. Distribution.Known only from the type locality. Volutaxis (Versutaxis) odiosus (Pilsbry 1899)Opeas odiosum Pilsby 1899; Proc. Acad. Nat. Sci. Phila. 51:399.(shell). Spiraxis odiosus (Pilsbry). H. B. Baker 1939; Nautilus 53:13.Type Locality.Patzcuaro, Michoacn, Mxico. Holotype ANSP 77100a. Distribution.Known only from the type locality. Volutaxis (Versutaxis) patzcuarensis (Pilsbry 1899)Opeas patzcuarensis Pilsbry 1899; Proc. Acad. Nat. Sci. Phila. 51:399.Pilsbry 1903; Proc. Acad. Nat. Sci. Phila. 55: pl. 50, 23 (shell).Naranjo-Garcia & Fahy 2010; Amer. Malac. Bull., 28:66. Spiraxis putzcuarensis (Pilsbry). H. B. Baker 1939; Nautilus 53:52.Type Locality.Patzcuaro, Michoacn, Mxico. Syntypes ANSP 77151. Distribution.Known only from the type locality.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 201 Volutaxis (Versutaxis) subgranum (H. B. Baker 1939)Spiraxis (Versutaxis) subgranum H. B. Baker 1939; Nautilus 53:51; Type Locality.Las Tortolas [2 km. N of east end of], Distribution.Known only from the type locality. Volutaxis (Versutaxis) subopeas (H. B. Baker 1939)Spiraxis (Versutaxis) subopeas H. B. Baker 1939; Nautilus 53:52; Type Locality.Below Necaxa, Puebla, Mxico; 2625 ft. alt. Holotype in the UMMZ. Distribution.Known only from the type locality. Volutaxis (Mirapex ?) acus (Shuttleworth 1852)Spiraxis acus Shuttleworth 1852:207.Fischer & Crosse 1877:614.Taxonomy.Spiraxis acus Shuttleworth is a nomen dubium. It has not been recognized since its original description because of its vague description, because it has (Neubert & Gosteli 2003:11). The name had been associated with Mirapex because of H. B. Bakers (1939b) designation of Volutaxis enigmaticus as a subspecies of Spiraxis acus. Volutaxis (Volutaxis ?) costatostriatus (Pfeiffer 1856)Bulimus costatostriatus Pfeiffer 1856; Proc. Zool. Soc. Lond. Volutaxis (?) costatostriatus (Pfeiffer). Pilsbry 1907; Man. Conch. 19:24. Distribution.Known only from the type locality. Taxonomy.This is a nomum dubium. The species was reported since its original description. Superfamily STREPTAXOIDEA Gray 1860 Family STREPTAXIDAE Gray 1860 Subfamily ENNEINAE Bourguignat 1883 Genus Sinoennea Kobelt 1904Sinoennea Kobelt 1904; Nachr. Deut. Malak. Gezel. 36:28.Type Species.Pupa strophiodes Gredler 1881. Distribution.East Asia, including Japan, China, Vietnam, Malaysia, and southern India. Taxonomy.Two subgenera are recognized. One has been introduced into the study area. Subgenus Indoennea Kobelt 1904Indoennea Kobelt 1904; Nachr. Deut. Malak. Ges. 36:28Type Species. Ennea blanfordiana Godwin-Austin 1872. Distribution.Primarily southern India and the Himalayas. One species has been introduced extensively around the tropical and subtropical. Taxonomy.Numerous species in southern Asia. One species has been introduced into the study area. Sinoennea (Indoennea) bicolor (Hutton 1834)Pupa bicolor Hutton 1834; Jour. Asian Soc. Bengal 3:86. Ennea (Huttonella ) bicolor (Hutton). Kobelt 1905; in Martini & Chemnitz System. Conchyl-Cabinet, Rhytididae & Enneidae, Gulella (Indoennea) bicolor (Hutton). Pilsbry 1919; Bull. Amer. Mus. Nat. Hist. 40:223.Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:98.Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. 82:345.Correa-Sandoval 1999; Acta Zool. Mex. (78):179 181. Sinoennea (Indoennea) bicolor (Hutton). Thompson 2008:666. Pupa largilliarti Philippi 1844; Zeitshr. Malac., 165. Pupa mellita Gould 1846; Boston Soc. Nat. Hist. 2:98. Ennea ceylonica Pfeiffer 1855; Proc. Zool. Soc. Lond. 23:9. Pupa caffaeicola Craven 1880; Proc. Zool. Soc. Lond. 48:215; pl. Type Locality.West Indies. Distribution.Widely distributed throughout the tropical and subtropical regions of the world. The origin of the species is unknown, but is presumed to be in southern India. PANAM: Cd. Panam; Ancon; Isla Tobago (Pilsbry 1926). VERACRUZ: El Tajn (Correa-Sandoval 1999). Genus Streptostele Dohrn 1866Streptostela Dohrn 1866; Malak. Bltt. 13:128.Type Species.Bulimus fastigiatus Morelet 1848. Distribution.Africa and adjacent islands, but not Madagascar. Taxonomy.Four subgenera are recognized (Zilch 1960). One has been introduced into the study area. Subgenus Tomostele Ancey 1885Tomostele Ancey 1885; Bull. Soc. Malac. France 2:143.Pilsbry 1919; Bull. Amer. Mus. Nat. Hist. 40:180.Type Species.Achatina musaecola Morelet 1860. Distribution.Tropical west Africa. Taxonomy.A few species, one of which has been widely introduced into tropical regions of the world. Streptostele (Tomostele ) musaecola (Morelet 1860)Achatina musaecola Morelet 1860; Jour. de Conchyl. 8:190. Streptostele (Tomostele) musaecola (Morelet). Pilsbry 1919; Bull. & Bermdez 2003; Malacologia 45:185.Thompson 2008:267. Luntia insignis Smith 1898; Jour. Conch. 9:27. Leptinaria insignis 44:303.Type Locality.Gabon. Syntypes BMNH 1898.12.5.18 (3). Distribution.Native range from Guinea to the Congo (Pilsbry 1919); widely introduced in tropical regions around

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202 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Arenal, El Mango, 455 m alt.; El Arenal, El Pochote, 455 m Superfamily RHYTIDOIDEA Pilsbry 1893 Family SCOLODONTIDAE H. B. Baker 1925Scolotontinae H. B. Baker 1925; Nautilus 38:86.Tillier 1880; Mem. Mus. Nat. dHist. Nat., 118.Hausdorf 2003; Jour. Moll. Stud., 69:183. Systrophiidae Thiele 1926. (Name not available from Thiele 1921, who used the vernacular name Systrophiiden).Type Genus.Scolodonta Dring 1875. Distribution.The Neotropical realm from southern Argentina north to eastern Mxico and the Lesser Antilles. Taxonomy.Eight genera are recognized. Five occur in the study area. Genus Drepanostomella Bourguignat 1889Drepanostomella Bourguignat 1889:42.H. B. Baker 1925a; Occ. Pap. Mus. Zool. Univ. Mich. (156):15, 17, 24.Type Species. Helix (Drepanostomella ) ammoniformis Orbigny 1835. Distribution.Ecuador north to Guatemala. Taxonomy.Six species are recognized. Two occur in the study area. Drepanostomella pinchoti Pilsbry 1930Drepanostomella pinchoti Pilsbry 1930d; Proc. Acad. Nat. Sci. 2002:228.Thompson 2008:668.Type Locality.Foothills near the Rio Madingo, Gulf of San Blas, Republic of Panam. Holotype ANSP 152048. Distribution.Nicaragua: Dept. Boaco 2002). PANAM: only from the type locality. Drepanostomella stolli (Von Martens 1892)Hyalinia stolli 15c (shell).Martens 1901:619. Ammoniceras stolli (Von Martens). Hinkley 1920; Nautilus 34:39, 51. Drepanostomella stolli (Von Martens). Pilsbry 1930; Proc. Acad. 2008:668.Type Locality.Retalhuleu, Dept. Retalhuleu, Guatemala. Distribution.COSTA RICA: San Jos, 1135 m alt; Puerto Viejo (Martens 1892 1901); Cahuita (Pilsbry 1930). GUATEMALA, Dept. Alta Verapaz: Cham (Hinkley 1920). Dept. Izabal: Quirigua (Hinkley 1920). Genus Guestieria Crosse 1872Guestieria Crosse 1872:290.H. B. Baker 1925; Occ. Pap. Mus. Zool. Univ. Mich. (156):14 18.Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. 82:347.Type Species.Helix powesiana Pfeiffer 1848. Distribution.Ecuador north to Panam. Taxonomy.Six species are recognized. One occurs in the study area. Guestieria isthmica Pilsbry 1930Guestieria isthmica Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. 6b (shell).Type Locality.Foothills near Rio Madingo, Gulf of San Blas, Panam. Holotype ANSP 152647. Distribution.Known only from the type locality. Genus Miradiscops H. B. Baker 1925Miradiscops H. B. Baker 1925; Occ. Pap. Mus. Zool. Univ. Mich. (156):17, 34.H. B. Baker 1929; Proc. Acad. Nat. Sci. Phila. 81 Type Species. H. B. Baker 1925. Distribution.Venezuela north to eastern Mxico. Taxonomy.Eight species are recognized. Seven occur in the study area. Miradiscops balboa Pilsbry 1930Miradiscops balboa Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. Type Locality.Ruins of Old Panam City, Panam. Holotype ANSP 151302. Distribution.PANAM: Isla Barro Colorado, Canal Zone (Pilsbry 1930). Miradiscops haplocochlion Thompson 1967Miradiscops haplocochlion Thompson 1967; Bull. Fla. State Campeche, Mxico. Holotype UF 19058. Distribution.Known only from the type locality. Miradiscops maya (Pilsbry 1920)Pseudohyalina maya Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. Mus. Zool. Univ. Mich. (156):12.Thompson 2008:670.Type Locality.Maya farms, Quirigua, Dept. Izabal, Guatemala. Holotype ANSP 107511. Distribution.GUATEMALA: known only from the type locality. VENEZUELA (H. B. Baker 1925). Miradiscops opal (Pilsbry 1920)Pseudohyalina opal Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. Pap. Mus. Zool. Univ. Mich. (156):13. Miradiscops opal (Pilsbry). H. B. Baker 1929; Proc. Acad. Nat. Sci. Holotype ANSP 48523. Distribution.NICARAGUA: various localities along

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 203 (H. B. Baker 1929a). SAN LUS POTOS: numerous localities in the eastern part of the state (Correa-Sandoval et al. 1998). Venezuela (H. B. Baker 1925). Miradiscops panamensis Pilsbry 1930Miradiscops panamensis Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. Thompson 2008:670.Type Locality.Ruins of Old Panam City, Panam. Holotype ANSP 151365. Distribution.NICARAGUA: numerous localities only from the type locality. Miradiscops puncticipitis (Pilsbry 1926)Pseudohyalina puncticipitis Pilsbry 1926; Proc. Acad. Nat. Sci. Miradiscops puncticipitis (Pilsbry). H. B. Baker 1929; Proc. (penis).Correa-Sandoval, Garca-Cubas & Reguero 1998:19.Thompson 2008:671.Type Locality.Cham, Dept. Alta Verapaz, Guatemala. Holotype ANSP 45656. Distribution.GUATEMALA: Chama. PUEBLA: Necaxa (H. B. Baker 1929). SAN LUS POTOS: Vega W) (Correa-Sandoval et al. 1998). Miradiscops ridicula Pilsbry 1930Miradiscops ridicula Pilsbry 1930b; Proc. Acad. Nat. Sci. Phila. Type Locality.Valley opening westward, north of High Peak, Isla de Providencia, Colombia. Holotype ANSP 150163. Distribution.Known only from the type locality. Genus Occultator Pilsbry 1926Occultator Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:129.Type Species.Occultator olssoni Pilsbry 1926. Distribution.Costa Rica. Taxonomy.A single species is recognized. Occultator olssoni Pisbry 1926Occultator olssoni Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. Holotype ANSP 104675. Distribution.Known only from the type locality. Genus Systrophia Pfeiffer 1855Systrophia Pfeiffer 1855; Proc. Zool. Soc. Lond. 23:136.H. B. Baker 1925; Occ. Pap. Mus. Zool. Univ. Mich. (156):19.Thiele 1927:3101.H. B. Baker 1928; Nautilus 41:124 Scolodonta Dring 1875:438.Type Species.Systrophia: Helix systropha Albers 1854. Scolodonta : Scolodonta semperi Dring 1875. Distribution.South America from Argentina north to Costa Rica. Taxonomy.Three subgenera are recognized (Zilch 1960). One occurs in the study area. Subgenus Systrophiella H. B. Baker 1925Systrophiella H. B. Baker 1925; Occ. Pap. Mus. Zool. Univ. Mich. (156):17, 31.H. B. Baker 1925; Nautilus 38:86.H. B. Baker 1926; Occ. Pap. Mus. Zool. Univ. Mich. (167):5.H. B. Baker 1928b; Nautilus 41:125.Type Species.Scolodonta (Systrophiella ) eudiscus H. B. Baker 1925 Distribution.Northern and northwestern South America north to Costa Rica. Taxonomy.Numerous species. Three species occur in the study area. Systrophia (Systrophiella) costaricana Rehder 1942Systrophia (Systrophiella ) costaricana Rehder 1942; Jour. Wash. Type Locality.Coto, on the Golfo Dulce, Prov. Puntarenas, Costa Rica. Holotype USNM 536023. Distribution.Known only from the type locality. Systrophia (Systarophiella) zeteki (Pilsbry 1920)Scolodonta zeteki Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 72:195; Systrophia (Systarophiella) zeteki (Pilsbry). Thompson 2008:673.Type Locality.Gamboa, Canal Zone, Panam. Holotype ANSP 114079. Distribution.PANAM: Gatn; Isla Barro Colorado; Gamboa (Pilsbry 1926). Systrophia (?) antoni (Pfeiffer 1842)Helix antoni Pfeiffer 1842:22. Scolodonta antoni (Pfeiffer). H. B. Baker 1925; Occ. Pap. Mus. Zool. Univ. Mich. (156):30.Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:101. Systrophia (?) antoni (Pfeiffer). Thompson 2008:673.Type Locality.Panam (?). Distribution.Known only from the type locality. Superfamily ACAVOIDEA Pilsbry 1895 Family MEGOMPHICIDAE Baker 1930 Distribution.Western North America from Montana and Idaho south to extreme northwestern Mxico. Taxonomy.The family contains four genera. One genus occurs in Mxico. Genus Glyptostoma Bland & Binney 1873Glyptotoma Bland & Binney 1873. Proc. Acad. Nat. Sci. Phila.:244.Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. 82:309.Berry 1938; Jour. Ento. Zool. Pom. Coll., 30:55.Pilsbry 1939; Land Moll. N. Amer. I:567.Type Species.Helix newberryana W. G. Binney 1858. Distribution.Los Angeles County, California, south to Ensanada, Baja California Norte.

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204 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Taxonomy.The genus contains two species. A single species and subspecies occurs in Mxico. Glyptostoma newberryanum depressum Bryant 1902Glyptostoma newberryanum depressum Bryant 1902; Nautilus 16:70.Berry 1928; Jour. Ent. Zool. Pom. Coll. 20:75.Pilsbry Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. Type Locality.Bluffs north of Ensanada de Todo Santos, Baja California Norte, Mxico. Holotype in the SDMNH. Distribution.BAJA CALIFORNIA NORTE: vicinity of the type locality; Caada Macho Gero (Smith et al. 1990). Superfamily PUNCTOIDEA Morse 1864 Family PUNCTIDAE Morse 1864Punctidae Morse 1864.Solem 1982.Type Genus.Punctum Morse 1964. Distribution.Holarctic, scattered localities in Africa, southern Australia, New Zealand, sub-Antarctic islands. Taxonomy.Numerous genera. One occurs naturally within the study area. A second genus from southern Australia has been introduced. Genus Paralaoma Iredale 1913 Paralaoma Iredale 1913; Proc. Malacol. Soc. London 10:380.Type Species.Paralaoma raoulensis Iredale 1913. Distribution.Kermadec Group of islands. One species has been distributed widely by human agency.. Taxonomy.A few species have been described. (Shuttleworth 1852) Shuttleworth 1852. Zonites diegoensis Hemphill, in Binney 1862. Punctum conspectum var. pasadenae Pilsbry 1896; Nautilus 10:21. Paralaoma caputspinulae (Reeve). Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:127. (Shuttleworth). Roth & Sadeghian 2006:42, 61.Type Localities. : Canary islands. Zonites diegoensis: near Julian City, San Diego County, California. Punctum conspectum pasadenae : Pasadena, California. Distribution.A non-native species that has been introduced widely throughout temperate regions of the world. Reported in Mxico from BAJA CALIFORNIA SUR: San Javier, 1200 ft. alt.; 1 km E of San Javier; 15.7 km W of San Javier (Smith et al. 1990). JALISCO (Roth & Sadeghian 2006). Genus Punctum Morse 1864Punctum Morse 1864:27. H. B. Baker 1930; Proc. Acad. Nat. Sci. Phila. 82:5.Pilsbry 1948; Land Moll. N. Amer. 2:641.Type Species.Helix minutissima Lea 1841. Distribution.Holarctic in distribution south to Panam in the New World; South Africa. Taxonomy.Zilch (1959) listed three subgenera. Subgenus Punctum Morse 1864 Distribution.As for the genus. Taxonomy.Numerous species. One species with three subspecies occurs in Mxico. Punctum ( Punctum) minutissimum minutissimum (Lea 1841)Helix minutissimus Lea 1841:17. Punctum minutissimum Mus. Zool. Univ. Mich. (220):5.Pilsbry 1948; Land Moll. N. Cubas & Reguero 1998:14.Thompson 2008:676. Punctun pygmaeum (Draparnaud). Dall 1926; Proc. Calif. Acad. Sci., ser. 4 15:481.Type Locality.Helix minutissimus: vicinity of Cincinnati, Ohio, USA. Syntypes USNM 105694. Distribution.Widely distributed throughout temperate North America south to central Mxico. NAYARIT: Isla Socorro, 2000 ft. alt. (Dall 1926). PUEBLA: Necaxa, 2025 5000 ft. alt. (H. B. Baker 1930a). SAN LUS POTOS: numerous localities in the southeastern part of the state below 1320 m alt. (Correa-Sandoval et al. 1998). Punctum (Punctum) minutissimum rotundum Dall 1926Punctum pygmaeum var. rotundum Dall 1926; Proc. Calif. Acad. Sci., ser. 4 15:481. Punctum (Punctum) minutissimum rotundum Dall. Thompson 2008:676.Type Locality.Isla Maria Magdelena, Islas Maras, Nayarit, Mxico. Holotype CAS 2203. Distribution.Known only from the type locality. Subgenus Toltecia Pilsbry 1926Toltecia Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:11617.Pilsbry 1948; Land Moll. N. Amer. 2:649.Roth 1985; Malac. Rev. 18:52.Type Species.Thysanophora jaliscoensis Pilsbry 1926 (= Punctum conspectum jaliscoense ). Distribution.Central America and Mxico from Panam north to California. Taxonomy.Seven species are recognized. Six occur in the study area. Punctum (Toltecia ) burringtoni Pilsbry 1930Punctum (Toltecia ) burringtoni Pilsbry 1930d; Proc. Acad. Nat. Thompson 2008:677.Type Locality.Guatemala. Holotype ANSP 132713. Distribution.GAUTEMALA: known only from the type locality. NICARAGUA, Dept. Chinandega (Prez & Managua. Dept. Matagalpa: La Cartuja; Cd. Dario (Prez & Punctum (Toltecia ) coloba (Pilsbry 1894)Thysanophora coloba Pilsbry 1894; Proc. Acad. Nat. Sci. Phila.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 205 Sci. Phila. 55:763.Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:117. Punctum coloba (Pilsbry). Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. Punctum (Toltecia ) coloba (Pilsbry). Thompson 2008:677. Holotype ANSP 12162. Distribution.Known only from the type locality. This may be the species recorded from Nicaragua by Prez and Lopz (2003) as Thysanophora crintita Fulton 1917. Punctum (Toltecia ) conspectum jaliscoensis (Pilsbry 1926)Thysanophora (Toltecia ) jaliscoensis Pilsbry 1926; Proc. Acad. Nat. Punctum (Toltecia ) conspectum jaliscoensis (Pilsbry). H. B. Baker 1930; Occ. Pap. Mus. Zool. Univ. Mich. (220):5.Pilsbry 1948; Land Moll. N. Amer. 2:652.Thompson 2008:678.Type Locality.Guadalajara, Jalisco, Mxico. Holotype ANSP 44927. Distribution.DISTRITO FEDERAL: Chapultepec Park (Pilsbry 1948). JALISCO: Guadalajara. Punctum (Toltecia ) mazatlanica (Pfeiffer 1856)Helix mazatlanica Pfeiffer 1856; Malak. Bltt. 3:43.Binney & Helix (Patula ) mazatlanica (Pfeiffer). Fischer & Crosse 1872:231. Pseudohyalina mazatlanica (Pfeiffer). Tryon 1866; Man. Conch. Patula mazatlanica (Pfeiffer). Von Martens 1892; Biol. Cent. Amer.:127. Punctum (Toltecia ) mazatlnica (Pfeiffer). Thompson 2008:678.Type Locality.Mazatln, Sinaloa, Mxico. Distribution.SINALOA (?). It is not clear from earlier literature whether the type locality is Mazatln, Sinaloa, or Mazatln, Guerrrero. Punctum (Toltecia ) textilis (Pilsbry 1920)Thysanophora textilis Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. Thysanophora (Toltecia ) textilis Pilsbry. Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:117. Punctum (Toltecia ) textilis (Pilsbry). Thompson 2008:679.Type Locality.Cham, Guatemala. Holotype ANSP 45653a (H. B. Baker 1963:240). Distribution.Known only from the type locality. Taxonomy.The shell characters suggest Microconus (Thysanophoridae). It is tentatively left in Toltecia where Pilsbry (1926) placed it. Punctum (Toltecia ) H. B. Baker 1930Punctum (Toltecia ) H. B. Baker 1930; Occ. Pap. Mus. Zool. Sandoval 1997; Rev. Biol. Trop. 44/45:140.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):236.Thompson 2008:679.Type Locality.Pleasure Gardens, New Braunfels, Comal County, Texas. Holotype ANSP 150609. Distribution.Widely distributed in the eastern United States west to Texas, and south to Veracruz. NUEVO LEN: Iturbide (Corrrea-Sandoval 1997). TAMAULIPAS: Caon Rodriguez 2002). VERACRUZ: Sumidero, 3400 ft. alt. (H. B. Baker 1930a). Family CHAROPIDAE Hutton 1884Charopidae Hutton 1884.Solem 1982.Type Genus.Charopa Albers 1860. Distribution.Widely distributed on the southern Taxonomy.Five subfamilies are recognized (Solem 1982). All New World Charopidae belong in the subfamily Rotadiscinae. Subfamily ROTADISCINAE H. B. Baker 1927Rotadiscinae H. B. Baker 1927; Proc. Acad. Nat. Sci. Phila. 79:228 230..Solem 1982:70.Type Genus.Rotadiscus Pilsbry 1926. Distribution.Western North America south through South America. One genus is present on islands in the Western Australia (Solem 1982). Taxonomy.Six neotropical genera belong in this subfamily (Solem 1982). Genus Chanomphalus Strebel 1880Chanomphalus Strebel 1880; Beitrag. Mex. Landund Sssw.Conch. IV:19.H. B. Baker 1928; Nautilus 41:123.Type Species.Helix elegantula Pfeiffer 1867 (= Thysanophora pilsbryi H. B. Baker 1922). Distribution.Widely distributed from Costa Rica Tamaulipas along the east cost. Taxonomy.Three species are placed in the genus. Two are provisionally assigned here, pending further study. Chanomphalus pilsbryi (H. B. Baker 1927)Helix elegantula Pfeiffer 1867; Malak. Bltt. 14:196 (not Helix elegantula Cristophri & Jan 1832). Chanomphalus elegantulus (Pfeiffer). Strebel 1880; Beitrag. Mex. Pseudohyalinea elegantula (Pfeiffer). Von Martens 1892; Biol. Cent. Amer.:125. Zonitoides elegantulus (Pfeiffer). Hinkley 1907; Nautilus 21:77.Hinkley 1920; Nautilus 34:39, 51. Thysanophora pilsbryi H. B. Baker 1922; Occ. Pap. Mus. Zool. Planogyra (Chanomphalus) pilsbryi (H. B. Baker). H. B. Baker Chanomphalus pilsbryi (H. B. Baker). H. B. Baker 1928; Nautilus 41:123.H. B. Baker 1929; Proc. Acad. Nat. Sci. Phila. 81:265.Correa-Sandoval, Garca-Cubas & Reguero 1998:14.Correa

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206 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)2008:680. Punctum planatum Dall 1926; Proc. Calif. Acad. Sci. (4) 15:482; pl. Type Localities. Helix elegantula : near Veracruz, Veracruz, Mxico. Thysanophora pilsbryi: Hacienda Cuatotolapam, Veracruz, Mxico. Holotype in the UMMZ. Punctum planatum: near village on east side of island, Isla Mara Madre, Islas Maras, Nayarit, Mxico. Holotype CAS 2205. Distribution.COSTA RICA: Tierra Blanca, on the southern slope of Volcn Iraz (Von Martens 1892). Chama (Hinkley 1920). Dept. Izabal: Maya Farms, Quirigua (Hinkley 1920). Dept. Petn: Tikal (Basch 1959). CHIAPAS: Laguna Ocotal, 950 m alt. (Bequaert 1957). NAYARIT: Isla Mara Madre, Islas Maras. PUEBLA: Necaxa 3000 ft. alt. (H. B. Baker 1930a). SAN LUS POTOS: various localities (Correa-Sandoval et al. 1998). TAMAULIPAS: river drift from near Tampico (Hinkley 1907); various localities in the southern part of the state (Correa-Sandoval & Rodriguez 2002). VERACRUZ: environs of Veracruz (Von Martens 1892); Sunidero, 3400 ft.a alt. (H. B. Baker 1930a); Carr. (Correa-Sandoval 1999). Chanomphalus cidarisca (Von Martens 1892)Pseudohyalina cidariscus Von Martens 1892; Biol. Cent. Amer.:126; Chanomphalus cidarisca (Von Martens). Thompson 2008:681.Type Locality.Palenque, Chiapas, Mxico. Syntype(s) in the British Museum of Natural History. Distribution.Known only from the type locality. Taxonomy.This species is provisionally referred to the genus until the microsculpture of the syntypes is reexamined. Chanomphalus tatei Pilsbry 1903Helix blakeana (not Helix blakeana Newcomb 1861). Thysanophora tatei Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. Nat. Sci. Phila. 78:116.Thompson 2008:682.Type Locality.Chontales forest, Nicaragua. Holotype ANSP 58065. Distribution.Known only from the type locality. Taxonomy.The shell is similar in size and shape to Chanomphalus pilsbryi. Genus Radiodiscus Pilsbry & Ferriss 1906Radiodiscus Pilsbry & Ferriss 1906; ANSP, 58:154.H. B. Baker 1927; Proc. Acad. Nat. Sci. Phila. 79:230.Type Species.Radiodiscus millicostatus Pilsbry & Ferriss 1906. Distribution.Western North America from Idaho south through Arizona and New Mexico and south to Argentina. Taxonomy.Three subgenera are recognized. One subgenus occurs within the study area. Subgenus Radiodiscus Pilsbry & Ferriss 1906 Distribution.As for the genus. Taxonomy.About a dozen species are recognized, two in the study area. One is represented by two subspecies. Radiodiscus (Radiodiscus) millicostatus millicostatus Pilsbry & Ferriss 1906Radiodiscus millicostatus Pilsbry & Ferriss 1906, Proc. Acad. Nat. 2002:238. Radiodiscus millicostatus millicostatus Pilsbry & Ferriss. Thompson 2008:683.Type Locality.Carr Canyon, Huachuca Mountains, Cochise County, Arizona. Holotype ANSP 89224. Distribution.Widely distributed in Arizona and New mi, from Pearson, 7000 ft. alt. (Pilsbry 1948). MICHOACAN: Patzcuaro; Morelia (Pilsbry 1948). Ragiodiscus (Radiodiscus) millicostatus costaricanus Pilsbry 1926 Ragiodiscus millicostatus costaricanus 2008:683. Holotype ANSP 140676a (H. B. Baker 1963:232). Distribution.Known only from the type locality. PUEBLA: near Necaxa (H. B. Baker 1927). Radiodiscus (Radiodiscus) proameri H. B. Baker 1930Radiodiscus proameri H. B. Baker 1930; Occ. Pap. Mus. Zool. Type Locality.Necaxa, Puebla, Mxico. Holotype in the UMMZ. Distribution.PUEBLA: Necaxa. VERACRUZ: Sumidero (H. B. Baker 1930a). Genus Rotadiscus Pilsbry 1926Rotadiscus Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:132.Type Species.Helix hermanni Pfeiffer 1866. Distribution.Southeastern Mxico south to Costa Rica; possibly Brasil. Taxonomy.Two species are recognized. One is represented by two subspecies. A third species, Entodonta discoidea Thiele 1927, from Brasil, may also be a Rotodiscus (H. B. Baker 1928b:128). Rotadiscus hermanni hermanni (Pfeiffer 1866)Helix hermanni Pfeiffer 1866; Malak. Bltt. 13:79.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 207Helix (Patula ) hermanni Pfeiffer. Fischer & Crosse 1873; Miss. Sci. Patula hermanni (Pfeiffer). Strebel 1880; Beitrag. Mex. LandAcad. Nat. Sci. Phila. 43:314.Von Martens 1892; Biol. Cent. Amer.:128. Rotadiscus hermanni (Pfeiffer). Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:132.H. B. Baker 1927; Proc. Acad. Nat. Sci. Phila. Rotadiscus hermanni hermanni (Pfeiffer). Thompson 2008:694.Type Locality.Mirador, Veracruz, Mxico. Distribution.GUATEMALA, Dept. Huehuetenango: Volcan de Agua (Von Martens 1892). MICHOACN: Morelia; Patzcuaro; Uruapan (Pilsbry 1903). PUEBLA: Necaxa (H. B. Baker 1927). VERACRUZ: Mirador; Orizaba (Von Martens 1892). H. B. Baker 1930 H. B. Baker 1930; Occ. Pap. Mus. Zool. Type Locality.Between La Venta and El Desierto de los Leones, Mxico State; 9700 ft. alt. Holotype in the UMMZ. Distribution.Known only from the type locality. Rotadiscus pilsbryi Rehder 1942Rotadiscus pilsbryi Rehder 1942; Jour. Wash. Acad. Sci. 32:352; Type Locality.Santa Maria, Prov. San Jos, Costa Rica; 1550 m alt. Holotype USNM 536018 (destroyed). Distribution.Known only from the type locality. Species provisionally referred to the CHAROPIDAE Punctum baschi Thompson 1962Punctum baschi Type Locality.Cobn-Sebal road, 55 miles northeast of Cobn, Dept. Alta Verapaz, Guatemala. Holotype UMMZ. Distribution.Known only from the type locality. Family DISCIDAE Thiele 1931Patulidae Tryon 1866. Goniodiscinae Wagner 1927. Discidae Thiele 1931.Zilch 1959:227.Solem 1982.Type Genus.Discus Fitzinger 1833. Europe, Madeira, and the Canary Islands. Taxonomy.Patulidae Tryon 1866 is the oldest name that has been proposed for this family. Solem (1982) rejected the name Patulidae because at the time it was proposed the type genus Patula was associated with Oreohelix (Oreohelicidae). Discidae is conserved under ICZN Art. 40.2 because of prevailing usage (Bouchet & Rocroi 2005:268). Genus Discus Fitzinger 1833Discus Fitzinger 1833:99.Pilsbry, Land Moll. N. Amer. 1948:508.Zilch 1959:227.Type Species.Helix ruderata H. & A. Adams (H. B. Baker 1932). Distribution.Holarctic Realm. Taxonomy.Nine subgenera are recognized (Zilch 1959). The genus contains numerous species. Subgenus Discus Fitzinger 1833 Distribution.Holarctic Realm. Taxonomy.Numerous species. One occurs in the study area. Discus (Discus) whitneyi (Newcomb 1864)Patula whitneyi Newcomb 1864:118. Discus whitneyi (Newcomb). Roth 1987; Malacological Review 20:129.Thompson 2008:686. Patula cronkhitei Newcomb 1865:180. Discus cronkitei (Newcomb). Pilsbry, Land Moll. N. Amer. II Type Localities. Patula whitneyi : Near Lake Tahoe, California, 6100 ft. alt.; lectotype University of California Museum of Paleontology 10619 (Roth 1987). Patula cronkitei : Klamath Valley, Oregon; location of 5 syntypes CU 26391. Distribution.Widely distributed in North America, extending south into northern Mxico. CHIHUAHUA: Sierra de Brea (Bequaert & Miller 1973). Subgenus Goniodiscus Fitzinger 1833Goniodiscus Fitzinger 1833:98. Mexicodiscus Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:133.Type Species.Goniodiscis: Muhfeld 1810 (not Say 1817). Mexicodiscis : Pilsbry 1904. Distribution.Europe and North America. Taxonomy.Several species. One occurs in Mxico Discus (Goniodiscus)(Pilsbry 1904) Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. Goniodiscus (Mexicodiscus ) (Pilsbry). Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:132.Correa-Sandoval 1999:14.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):236. Goniodiscus (Mexicodiscus ) (Pilsbry). H. B. Baker 1930:14. Discus (Goniodiscus) (Pilsbry). Thompson 2008:687.Type Locality.Canyon 4 miles west of [Ciudad] Victoria, Tamaulipas, Mxico. Holotype ANSP 86907. Distribution.PUEBLA: Necaxa, 3000 ft. alt. (H. B. Baker 1930a). SAN LUS POTOS: numerous localities in southeastern part of state below 1320 m alt. (CorreaSandoval et al. 1998). TAMAULIPAS: numerous localities in southern part of state (Correa-Sandoval & Rodriguez Family HELICODISCIDAE H. B. Baker 1927Helicodiscinae H. B. Baker 1927; Proc. Acad. Nat. Sci. Phila.

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208 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)79:226. Helicodiscidae Solem 1982.Type Genus.Helicodiscus Pilsbry & Ferriss 1906. Distribution.North America south to northern Mxico. Taxonomy.A single genus is recognized. Genus Helicodiscus Morse 1864 Type Species.Helix lineata Say 1817 (= Helicodiscus paralellus Say 1821; not Helix lineata Olivi 1792). Distribution.Generally spread over North America from eastern Mxico, Chihuahua and Arizona to Canada, and the West Indies on Cuba and Jamaica. Taxonomy.Three subgenera are recognize (Zilch 1959). Two occur in Mxico. Subgenus Helicodiscus Morse 1864 Distribution.Continental North America. Taxonomy.About twenty species are recognized. One occurs in Mxico. Helicodiscus (Helicodiscus ) eiganmanni Pilsbry 1900Helicodiscus eiganmanni Pilsbry 190:11.H. B. Baker 1930:14.a-c (shell).Bequaert & Miller 1978:151.Thompson 2008:688.Type Locality.Beaver Cave, near San Marcos, Comal County, Texas. Holotype ANSP 78730. Distribution.Western United States from South Dakota, Utah, Colorado, Arizona, New Mexico and Texas south to Mxico. CHIHUAHUA: canyon above Colonia Juarez; Sierra de Brea, 11 mi. from Pearson; on Rio Piedras Verdes, near Pacheco (Pilsbry 1948). PUEBLA: near Necaxa, 4600 ft. alt. (H. B. Baker 1930a). SONORA: San Bernardino (Pilsbry 1948). Subgenus Lucilla Lowe 1852Lucilla Lowe 1852; Ann. Mag. Nat. Hist. ser. 2, 9:275. Hebetodisus H. B. Baker 1929; Nautilus 42:86.Pilsbry 1948:635.Type Species.Luilla: Zonites singeyanus Pilsbry 1890. Hebetodiscus: Helicodiscus singleyanus inermis H. B. Baker 1929. Distribution.Eastern United States south to Puebla and Baja California Sur. Taxonomy.Three species are recognized. One occurs in Mxico. Helicodiscus (Lucilla ) singleyanus (Pilsbry 1889)Zonites singleyana Pilsbry 1889; Proc. Acad. Nat. Sci. Phila. 41:84. Zonitoides singleyana (Pilsbry). Hinkley 1907; Nautilus 21:77. Helicodiscus (Hebetodiscus) singleyanus singleyanus (Pilsbry). H. 13.Pilsbry 1948; Land Moll. N. Amer. 2:636; textCalif. Acad. Sci. 47:127. Lucilla singleyana (Pilsbry). Roth & Sadeghian 2006; Santa Barbara Mus. Nat. Hist. Contrib. Sci., (3):9. Helicodiscus (Lucilla ) singleyanus (Pilsbry). Thompson 2008:689.Type Locality.New Braunfels, Comal County, Texas. Holotype ANSP 160058. Distribution.Widely distributed in North America east of the Rocky Mountains and south into northern Mxico. Widely introduced elsewhere. BAJA CALIFORNIA SUR: 0.3 km SSE of San Jos del Cabo; Isla Santa Catalina (Smith et al. 1990). PUEBLA: near Necaxa (H. B. Baker 1930a). TAMAULIPAS: Tampico (Hinkley 1907). Family OREOHELICIDAE Pilsbry 1939 Distribution.Western North America from Alberta, Canada south to northern Mxico. Taxonomy.Two genera are recognized. Genus Oreohelix Pilsbry 1904Oriohelix Pilsbry 1904; Nautilus 17:131.Pilsbry 1905; Proc. Acad. Nat. Sci. Phila. 57:268.Pilsbry 1916; Proc. Acad. Nat. Sci. Phila. 68:340.Henderson and Daniels 1916; Proc. Acad. Nat. Sci. Phila. 68:315.Pilsbry 1917; Proc. Acad. Nat. Sci. Phila. 69:42.Henderson 1918; Proc. Malac. Soc. London 13:21.Henderson 1924; Univ. Colorado Studies, 13:109.Pilsbry 1934; Proc. Acad. Nat. Sci. Phila. 85:383.Henderson 1936; Univ. Colorado Studies, 23:87.Pilsbry 1939; Land Moll. N. Amer. I:412.Type Species.Helix strigosa Gould 1846. Distribution.Southern Saskatchewan and British Colombia south through the mountain states to northern Sonora and from Saskatchewan eastward to North Dakota. Taxonomy.Twenty-six species and numerous subspecies are recognized. A single species occurs in Mxico. Oreohelix concentrata concentrata (Dall 1895)Patula strigosa var. concentrata Dall 1895; Proc. U. S. Nat. Mus. 18:1.Dall 1896; Proc. U. S. Nat. Mus. 19:336. Oreohelix strigosa concentrata (Dall). Pilsbry 1905; Proc. Acad. (shells). Oreohelix concentrata (Dall). Pilsbry 1916; Proc. Acad. Nat. Sci. 1939; Land Moll. N. Amer. I:501; 326, 327, 329 (shell), 328 (reproductive system). Pyramidula strigosa huachucana Pilsbry 1902. Proc. Acad. Nat. Sci. Phila. 54:511. Oreohelix strigosa huachucana (Pilsbry). Pilsbry 1905; Proc. Acad. (shell). Oreohelix concentrata huachucana (Pilsbry). Pilsbry 1916; Proc. Acad. Nat. Sci. Phila. 68:354.Pilsbry and Ferriss 1923; Proc. Oreohelix concentrata concentrata (Dall). Bequaert and Miller 1973; Moll. Arid SW:130.Thompson 2008:690.Type Locality.Patula strigosa var. concentrata : Summit of Huachuca Mountains, Arizona. Holotype USNM 129999. Pyramidula strigosa huachucana: Holotype ANSP 83370. Distribution.Southeastern Arizona and immediately adjacent Sonora. Another subspecies, Oreohelix concentrata grahamensis Gregg and Miller 1972, occurs in the Pinaleo

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 209 Mountains, Arizona. SONORA: Sierra San Jos 5 mi. S of the Arizona border, E of Rio San Pedro (Bequaert & Miller 1973). Genus Radiocentrum Pilsbry 1905Radiocentrum Pilsbry 1905; Proc. Acad. Nat. Sci. Phil., 57:283.Pilsbry 1939; Land Moll. N. Amer. 1:540.Bubrakzai, Miller and Ward 1975; Bull. Amer. Malac. Union, 40:10.Type Species.Oreohelix chiricahuana Pilsbry 1905. Distribution.Southern New Mexico and Arizona south to Tamaulipas, Coahuila, Chihuahua, and Baja California Sur. Possibly also Santa Catalina Island, California. Taxonomy.Twelve species are recognized Seven occur in Mxico. Radiocentrum almoloya (Drake 1949)Oreohelix (Radiocentrum ) almoloya Drake 1949; Nautilus 62:110 112; pl. 8.Miller 1973; Veliger 15:332. Radiocentrum almoloya (Drake). Thompson 2008:692.Type Locality.Rocky cactae and bush covered hillside mile radius northwest of Cuevo Diablo, a limestone sink two miles northwest of Salaices, Chihuahua, Mxico; ca. 5200 ft. alt. Holotype ANSP 185106. Distribution.Known only from the type locality. Radiocentrum caenosa (Pilsbry 1953)Oreohelix (Radiocentrum) caenosa Pilsbry 1953; Proc. Acad. Nat. Radiocentrum caenosa (Pilsbry). Thompson 208:692.Type Locality.Below a cliff at 6200 ft. alt. on the western slope of a mountain about 4.5 mi. southeast of Pearson (Mata Ortiz), Chihuahua, Mxico. Holotype ANSP 166154. Distribution.Known only from the type locality. Radiocentrum discus Christensen & Miller 1976Radiocentrum discus Christensen & Miller 1976; Veliger. 18:378 34 (map).Thompson 2008:692.Type Locality.Along the road from Valle Perdido to ranch at Bajada del Molino, in rockslide on north-facing slope of large arroyo south of the road, Baja California Sur 55827. Distribution.BAJA CALIFORNIA SUR: known only from the vicinity of the type locality. Radiocentrum exorbitans (Miller 1973)Oreohelix (Radiocentrum) exorbitans Miller 1973; Veliger 15:332 Radiocentrum exorbitans (Miller). Christensen and Miller 1976; Veliger 18:378.Smith, Miller, Christensen & Roth 1990; Proc. Type Locality.Sierra de La Gigante, in rock slides 37 km by road southwest of Loreto, Baja California Sur, CAS (Geology) 532676. Distribution.BAJA CALIFORNIA SUR: known only from the type locality and 1 km SE of San Bartolo. Radiocentrum labrenana (Pilsbry 1948)Oreohelix (Radiocentrum ) labrenana Pilsbry 1948; Proc. Acad. Nat. Radiocentrum labrenana (Pilsbry). Thompson 2008:693.Type Locality.Sierra de La Brea in a rock slide above the road from Pearson [Mata Ortiz] to Pacheco, Chihuahua, Mxico; 7000 ft. alt. Holotype ANSP 166587. Distribution.Known only from the type locality. Radiocentrum orientalis Metcalf 1980Radiocentrum orientalis (shell).Thompson 2008:693.Type Locality.Upper end of Caon El Bonito, Serranias del Burro, Mcpo. de Villa Acuna, Coahuila, Mxico; 1680 m Distribution.COAHUILA: known only from the type locality as late Pleistocene and Recent shells. (Pilsbry 1904) Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:7 (Pilsbry). Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):238. (Pilsbry). Thompson 2008:693.Type Locality.Canyon 4 miles west of Ciudad Victoria, Tamaulipas, Mxico. Lectotype ANSP 85907 (H. B. Baker 1963:233). Distribution.TAMAULIPAS: numerous records in southern part of state (Correa-Sandoval & Rodriguez 2002). Remarks.Assignment of this species to Radiocentrum is arbitrary. It is known only from dead shells. Its embryonic shell sculpture is consistent with Radiocentrum Superfamily SAGDOIDEA Pilsbry 1895 Family SAGDIDAE Pilsbry 1895 Genus Hojeda H. B. Baker 1926Hojeda H. B. Baker 1926; Occ. Pap. Mus. Zool. Univ. Mich. 167:15.Type Species. H. B. Baker 1924. the Bahamas, and Florida The exact distribution of the genus is poorly known. Taxonomy.Two species are recognized. (H. B. Baker 1924) H. B. Baker 1924 Occ. Pap. Mus. Zool. Thysanophora (Hojeda) H. B. Baker 1926; Occ. Pap. Mus.

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210 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) (H. B. Baker). Thompson 2008:694.Type Locality.Aruba, Dutch West Indies. Distribution.Known only from the type locality. Genus Hyalosagda Albers 1860Hyalosagda Albers 1860; in Von Martens, Die Heliceen, 2:77.Type Species.Helix similis C. B. Adams 1849. Distribution.Jamaica, Hispaniola, Cuba, and southern Guatemala. Taxonomy.The genus includes a few species. Subgenus Aerotrochus Pilsbry 1926Aerotrochus Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:120.Type Species.Helix subpyramidalis C. B. Adams 1845. Distribution.Hispaniola, Cuba, Jamaica, and the Dept. Petn, Guatemala. Taxonomy.A single species occurs in the study area. Hyalosagda (Aerotrochus) turbonella (Morelet 1851)Helix turbonella Morelet 1851; Test. Noviss. II:9. Helix (Patula ) turbonella Morelet. Fischer Crosse 1872:229; pl. 12, Patula turbonella (Morelet). Von Martens 1892; Biol. Cent. Hyalosagda turbonella (Morelet). Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):11. Hyalosagda (Aerotrochus) turbonella (Morelet). Thompson 2008:695.Type Locality.Forests of Petn, Guatemala. Distribution.BELIZE: Cayo Dist.: Mountain Pine J. Grego). GUATEMALA, Dept. Petn: Tikal (Basch 1959). Taxonomy.The generic assignment of this species is problematic. Genus Xenodiscula Pilsbry 1919Xenodiscula Pilsbry 1919:206.Goodrich & Van der Schalie 1937; Misc. Publ. Mus. Zool. Univ. Mich. (34):27.Type Species. Pilsbry 1919. Distribution.Venezuela north to eastern Mxico. Taxonomy.Two species are recognized. One occurs in the study area. Xenodiscula taintori Goodrich & Van der Schalie 1937Xenodiscula taintori Goodrich & Van der Schalie 1937; Misc. Publ. Type Locality.Woodland just east of El Paso de Los Caballos, Dept. Petn, Guatemala. Holotype in the UMMZ. Distribution.GUATEMALA, Dept. Alta Verapaz; Dept. Petn (Goodrich & van der Schalie 1937). NICARAGUA, Dept. Boaca: El Caracol: Dept. Matagalpa: Cd. Dario; Selva Negra; Fuente Pura. Dept. Rio San Juan: Almendro (Prez & Superfamily HELICARIONOIDEA Baurguignat 1877 Family HELICARIONIDAE Baurguignat 1877 Subfamily EUCONULIDAE H. B. Baker 1928 islands, North America, Central America, and South America. Taxonomy.Numerous genera are recognized (Zilch 1959:271). Thirty species of Helicarionidae, subfamily Euconulinae are recognized in Mxico and Central America. Genus Euconulus Reinhardt 1883Euconulus Reinhardt 1883; Beitrage zur Landeskunde Oesterreichs, 9:94.Pilsbry 1946; Land Moll. N. Amer. 2:234.Type Species. Mller 1774. Distribution.Holarctic realm in general, Hawaii, and Tahiti. Taxonomy.Six subgenera are recognized. Only one occurs in the study area. Subgenus Euconulus Reinhardt 1883 Distribution.Holarctic realm in general. Taxonomy.Numerous species. A single species occurs in the study area. (Mller 1774) Mller 1774; Hist. Vermium, 2:56. (Mller). Pilsbry 1908; Nautilus 22:25.Pilsbry 1926; Nautilus 40:68.Baker 1930; Proc. Acad. Nat. Sci. Phila. anatomy).Pilsbry 1946; Land Moll. N. Amer. 2:235; 1973:143.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:129.Thompson 2008:697.Type Locality.Fredrichsdal, Denmark. Distribution.Almost throughout the Holarctic realm (Pilsbry 1946). The species barely enters Mxico along its northern border with Arizona and California. BAJA CALIFORNIA NORTE: Sierra San Pedro Mrtir, 2700 m alt. (Smith et al. 1990). CHIHUAHUA: NW part of state (Bequaert & Miller 1973). SONORA: ESE part of Sierra (Naranjo-Garcia 1991). Genus Habroconus Fischer & Crosse 1872Habroconus Fischer & Crosse 1872:154.Baker 1928c: Proc. Acad. Nat. Sci. Phila. 80:11.Zilch 1959:279, 280.Type Species.Helix selenkai Pfeiffer 1866. Distribution.South America, the West Indies and Central America north to Mxico. Taxonomy.Five subgenera are recognized. All occur in the study area Subgenus Habroconus Fischer & Crosse 1872. Distribution.Northern South America, Central America, Mxico, and the West Indies. Taxonomy.Five species and one subspecies occur in the study area.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 211 Habroconus ( Habroconus) championi (Von Martens 1892)Guppya championi Von Martens 1892; Biol. Cent. Amer.:119; Amer.:619.van der Schalie 1940; Occasional papers Museum of Zoology, University of Michigan, (413):4. Habroconus championi 174. Habroconus (Habroconus) championi (Von Martens). Thompson 2008:698.Type Locality.Purula [Purulha], towards the head of the Polochc Valley, Dept. Baja Verapaz, Guatemala. Distribution.COSTA RICA, Prov. Cartago: Hacienda El Roble, between Irazu and Turrialba. Prov. San Jos: environs of San Jos (Von Martens 1892); San Cristobal; Tarbaca; 1500 m alt. (Von Martens 1901). NICARAGUA, Alta Verapaz: Panzamala (van der Schalie 1940). Dept. Baja Verapaz: Purulja. Dept. Huehuetenango: Cholhuitz (Von Martens 1892). Habroconus (Habroconus) costaricanus costaricanus (Pilsbry 1920)Guppya costaricana Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 72:4; Habroconus (Habroconus) costaricanus costaricanus (Pilsbry). Thompson 2008:699.Type Locality.Alajuela, Prov. Alajuela, Costa Rica; 3200 ft alt. Holotype ANSP 105285. Distribution.Known only from the type locality. Habroconus ( Habroconus) costaricanus elatior (Pilsbry 1920)Guppya costaricana elatior Pilsbry 1920; Proc. Acad. Nat. Sci. Habroconus (Habroconus) costaricanus elatior (Pilsbry). Thompson 2008:699. Vias, Costa Rica; 2500 ft. alt. Holotype ANSP 105276. Distribution.Known only from the type locality. Habroconus (Habroconus) elegans Strebel 1880Habroconus elegans Strebel 1880; Beitrag. Mex. Landund Sssw.Stenopus elegans (Strebel). Pilsbry 1891; Proc. Acad. Nat. Sci. Phila. 43:312. Guppya elegans (Strebel). Von Martens 1892; Biol. Cent. Amer.:120.Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:768. Habroconus (Habroconus) elegans Strebel. Thompson 2008:699.Type Locality.Woods of Pacho, near Jalapa, Veracruz, Mxico. Distribution.GUATEMALA, Dept. Baja Verapaz: Salama (Von Martens 1892). VERACRUZ: Orizaba (Pilsbry 1891). Habroconus (Habroconus) selenkai (Pfeiffer 1866)Zonites selenkai Pfeiffer 1866; Malak. Bltt. 12:77. Zonites (Habroconus) selenkai (Pfeiffer). Fischer & Crosse Habroconus selenkae (Pfeiffer). Strebel 1880; Beitrag. Mex. Land(radula).Baker 1930a; Occasional papers of the Museum of Habroconus (Habroconus) selenkai (Pfeiffer). Thompson 2008:700.Type Locality.Mirador, Veracruz (Baker 1930a:24). Distribution.NICARAGUA: various localities in the Necaxa Falls, 2215 ft. alt. (Baker 1930a). VERACRUZ: Jalapa; Hacienda Mirador, near Jalapa; Agua Caliente, near Misantla (Strebel 1880); Atoyac, 1300 ft. alt. (Baker 1930a). Habroconus (Habroconus) trochulinus (Morelet 1851)Helix trochulina Morelet 1851; Test. Noviss. II:10. Zonites (Habroconus) trochulinus (Morelet). Fischer & Crosse 1872:172. Guppya trochulina (Morelet). Von Martens 1892; Biol. Cent. Biol. Cent. Amer.:619.Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:768. Guppya (Habroconus) trochulina (Morelet). Baker 1922; Occasional papers of the Museum of Zoology, University of Michigan, Habroconus (Habroconus) trochulinus (Morelet). Baker 1928c; (reproductive anatomy).Baker 1930: Occasional papers of the Museum of Zoology, University of Michigan, (220):22; pl. Zoology, 116:219.Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):12.Thompson 2008:700.Type Locality.Woods of Petn, near San Lus, Dept. Petn, Guatemala. Distribution.Venezuela north to Veracruz and Puebla (Baker 1930a). COSTA RICA, Prov. Alajuela: San Cristobal; between Mokri and Ukatschka, Alta Talamanca (Von Martens 1901). Prov. San Jos: environs of San Jos, near the streamlet Atajueleta, 1135 m alt. (Von Martens 1892); San Francisco de Guadalupe. Prov. Puntarenas: El Pital, valley of the Rio Naranjo, 200 m alt.; Quebrada de Java, in the valley of the Rio Brus, 900 m alt. (Von Martens 1901). GUATEMALA, Dept. Petn: San Luis (Von Martens 1892); Tikal National Park (Basch 1959). CHIAPAS: Selva Lacandona, Laguna Ocotal to El Censo, 700 m alt.; El Real, 600 m. alt. (Bequaert 1957). MICHOACAN: Patzcuaro; Morelia (Pilsbry 1904). PUEBLA: Necaxa, 2625 ft. alt. (Baker 1930a). VERACRUZ: Texolo (Pilsbry 1904); Coatotolapam; Lago de Catemaco (Baker 1922); Peuela to Sumidero, 2625 ft. alt. (Baker 1930a). Subgenus Cocosconus Baker 1941Cocosconus Baker 1941; Bull. Bernice P. Bishop Museum (166):223.Type Species.Guppya hopkinsi Dall 1900. Distribution.Isla del Coco, Costa Rica. Holotype USNM 108524. Taxonomy.A single species is recognized.

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212 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Habroconus (Cocosconus) hopkinsi (Dall 1900)Guppya hopkinsi Dall 1900; Proc. Acad. Nat. Sci. Phila. 52:97; pl. (Pfeiffer). Ancey 1903; Jour. de Conchyl. 51:101. Habroconus (Cocosconus) hoptkinsi (Dall 1900). Baker 1941; Bull. Type Locality.Isla del Coco, Costa Rica. Distribution.Known only from the type locality. Subgenus Cocoslens H. B. Baker 1941Cocoslens Baker 1941; Bull. Bernice P. Bishop Museum (166):223.Type Species.Habroconus (Cocoslens) pallidaus H. B, Baker 1941. Distribution.Isla del Coco, Costa Rica. Taxonomy.A single species is recognized. Habroconus (Cocoslens) pallidaus H. B. Baker 1941Habroconus (Cocoslens) pallidaus H. B. Baker 1941; Bull. Bernice Type Locality.Wafer Bay, Isla del Coco, Costa Rica. Holotype ANSP 170394. Distribution.Known only from the type locality. Subgenus Ernstia Jousseaume 1889Ernstia Jousseaume 1889; Mem. Soc. Zool. France, 2:232.H. B. Baker 1928; Proc. Acad. Nat. Sci. Phila. 80:12.Type Species.Ernstia ernsti Jousseaume 1889. Distribution.Northern South America and Central America north to Mxico. Taxonomy.Two species occur in the study area. Others occur in South America. Habroconus (Ernstia) elegantulus (Pilsbry 1919)Guppya elegantula Pilsbry 1919; Proc. Acad. Nat. Sci. Phila. Euconulus elegantulus (Pilsbry). H. B. Baker 1922; Occ. Pap. Mus. Habroconus (Ernstia) elegantulus (Pilsbry). H. B. Baker 1928c; B. Baker 1930a; Occ. Pap. Mus. Zool. Univ. Mich. (220):22.Haas 1949; Nautilus 62:137, 138.Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):12.Correa-Sandoval 1997; Rev. Biol. Trop. 44/45:140.Correa-Sandoval, Garca-Cubas & Reguero 1998:14.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):236.Thompson 2008:703.Type Locality.Canyon and falls below Valles, San Luis Potosi, Mxico. Holotype in the ANSP. Distribution.GUATEMALA, Dept. Alta Verapaz: Cama (Hinkley 1920). Dept. Chimaltenango: Finca Montserrat; Yepocapa (Haas 1949). Dept. Escuintla: Zapote (Haas 1949). Dept. Izabal: Jocolo; Maya Farms, Quirigua (Hinkley 1920). Dept. Petn: Tikal National Park (Basch 1959). Dept. Sacatepquez: Finca San Rafael (Haas 1949). JALISCO: Guadalajara (Pilsbry 1919). MICHOACAN: Uruapan (Pilsbry 1919). MORELOS: Yautepec Pilsbry 1919). NUEVO LEN: Diente, near Monterrey (Pilsbry 1919). PUEBLA: Necaxa, 2215 ft. alt. (H. B. Baker 1928 1930). NUEVO LEN: Iturbide (Correa-Sandoval 1997); Santiago, Montemorelos, Zaragoza (Correa-Sandoval & Salazar, (2005). SAN LUIS POTOS: Valles; numerous localities (Correa-Sandoval et al. 1998). TAMAULIPAS: canyon 4 mi. W of Ciudad Victoria, 3000 ft. alt. (Pilsbry 1919). VERACRUZ: Orizaba (Pilsbry 1919); Coatotolapam (H. B. Baker 1922). Atoyac to Sumidero, 1300 ft. alt. (H. B. Baker 1930a). Habroconus (Ernstia) zeteki Pilsbry 1930Habroconus (Ernstia) zeteki Pilsbry 1930d; Proc. Acad. Nat. Sci. Smiths. Misc. Coll. 106:42.Thompson 2008:704.Type Locality.Ruins of Old Panam City, Panam. Holotype ANSP 151367. Distribution.PANAM, Archipilago de las Perlas: Isla San Jos (Morrison 1964). Prov. Panam: Old Panam City. Subgenus Pseudoguppya H. B. Baker 1925.Pseudoguppya H.B. Baker 1925; Occ. Pap. Mus. Zool. Univ. Mich. (156):10.Type Species.Helix cassiquiensis Pfeiffer 1853. Distribution.Northern South America and Central America. Taxonomy.Five species occur in the study area. Others occur in northern South America. Habroconus (Pseudoguppya) browni (Pilsbry 1910)Guppya browni Pilsbry 1910; Proc. Acad. Nat. Sci. Phila. 62:509; Euconulus (Pseudoguppya) browni (Pilsbry). H.B. Baker 1925a:11.Pilsbry 1926a; Proc. Acad. Nat. Sci. Phila. 78:100. Habroconus browni (Pilsbry). Pilsbry 1930d; Proc. Acad. Nat. Sci. Phila. 82:351. Habroconus (Pseudoguppya) browni (Pilsbry). Thompson 2008:704.Type Locality.Between Tabernillo and San Pablo, Canal Zone, Panam. Holotype ANSP 101319. Distribution.Known only from the type locality. Habroconus (Pseudoguppya) (Pilsbry 1920) Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 72:3; Habroconus ( Pseudoguppya) (Pilsbry). Thompson 2008:704.Type Locality.Stream near the railroad west of Juan Vias, Costa Rica; 330 ft. alt. Holotype ANSP 105266. Distribution.Known only from the type locality. Habroconus (Pseudoguppya) (Pfeiffer 1846) Pfeiffer 1846; Symbolae, 3:66. Patula (Trochomorpha) (Pfeiffer). Tryon 1887; Man. Guppya hopkinsi conulus Von Martens 1902; Sitz. Ges. Naturf. Berlin:59.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 213Guppya fultoni Gude 1903; Proceedings Malacological Society, Habroconus (Pseudoguppya) (Pfeiffer). H. B. Baker 2008:705.Type Locality.Isla del Coco, Costa Rica. Distribution.Known only from the type locality. Habroconus (Pseudoguppya) pittieri (Von Martens 1892)Guppya pittieri Von Martens 1892; Biol. Cent. Amer.:121; pl. 6, Euconulus (?) pittieri (Von Martens). H. B. Baker 1922; Occ. Pap. Mus. Zool. Univ. Mich. (106):48. Habroconus pittieri (Von Martens). Thompson 1967; Bull. Fla. St. Mus. 11:243. Euconulus pittieri Habroconus (Pseudoguppya) pittieri (Von Martens). Thompson 2008:705.Type Locality.San Francisco de los Rios, near San Jos; 1100 m alt., Costa Rica. Distribution.COSTA RICA, Prov. San Jos: San Francisco de los Rios; Urraca, near San Jos, 1135 m alt. (Von Martens 1892). NICARAGUA: numerous localities along mi. NNW of Dzilbachen; 8.1 mi. SW of Champoton; 7.1 mi. S of Pixtun; 19.2 mi. E of Silvituc (Thompson 1967). QUINTANA ROO:4.0 mi. E of Xpujil, Campeche; 7.1 mi. NNW Xiatil (Thompson 1967). VERACRUZ: Catamaco (H. B. Baker 1922). YUCATN: 0.8 mi. NE of Becanchen (Thompson 1967). Habroconus (Pseudoguppya) utillensis (Ancey 1886)Conulus utillensis Guppya utillensis (Ancey). Von Martens 1892; Biol. Cent. Amer.:122. Habroconus (Pseudoguppya) utillensis (Ancey). Thompson 2008:706.Type Locality.Isla de tila, Dept. Islas de la Baha, Honduras. Distribution.Known only from the type locality. Genus Guppya Mrch 1867 Type Species. Mrch (error for Conulus Guppy 1866, in part) = Helix gundlachi Pfeiffer 1840 (see Baker 1925:7). Distribution.Central America, Mxico, southern United States, and the West Indies. Taxonomy.Numerous species are placed in Guppya, but the generic allocations of many are doubtful because anatomical characteristics of only a few are known. Thirteen species are recognized in the study area. Guppya angasi Von Martens 1892Stenopus guildingi Angas 1879; Proc. Zool. Soc. Lond. 47:484; pl. Stenopus guildingi Bland 1865). Guppya angasi Von Martens 1892; Biol. Cent. Amer.:120 (substitute name for Stenopus guildingi Angas).Von Martens 1901; Biol. Cent. Amer.:619.Thompson 2008:706.Type Locality.Costa Rica. Mokri and Ukatschka, Alta Talamanca (Von Martens 1901). Prov. Puntarenas: Quebrada de Java, 900 m alt.; Caas Gordas (Von Martens 1901). Guppya biolleyi Von Martens 1892Guppya biolleyi Von Martens 1892; Biol. Cent. Amer.:121; pl. Amer.:620.Fluck 1905; Nautilus 19:78.H. B. Baker 1928c; B. Baker 1930a; Occ. Pap. Mus. Zool. Univ. Mich. 220:21.Thompson 2008:707.Type Locality.Hacienda Helvetia, Costa Cuca, Dept. Quetzaltenango, Guatemala (H. B. Baker 1930a). (Von Martens 1901). Prov. San Jos: La Urraca; San Francisco de Los Rios (Von Martens 1892). NICARAGUA: Rama Key, Helvetia, Costa Cuca (Von Martens 1892). PUEBLA: Necaxa, 2625 ft. alt. (H. B. Baker 1930a). Guppya capsula Dall 1926Guppya capsula Dall 1926; Proc. Calif. Acad. Sci. (4) 15:479 Type Locality.North slope of Cerro Evermann, Isla Socorro, Nayarit, Mxico; 2000 ft. alt. Holotype CAS 2202. Distribution.Known only from the type locality. (Morelet 1851) Morelet 1851; Test. Noviss. II:9. Zonites (Hyalinia ) (Morelet). Fischer & Crosse 1872:177. Guppya gundlachi (Pfeiffer). Von Martens 1892; Biol. Cent. Amer.:122 (in part). (Morelet). Thompson 2008:708.Type Locality.Isla de Carmen, at the head of the Laguna de Terminos, Campeche, Mxico. Distribution.Known only from the type locality. Guppya gundlachi gundlachi (Pfeiffer 1840)Helix pusilla Pfeiffer 1839; Archiv fr Naturg. 1:351 (not Helix pusilla Lowe 1833). Helix gundlachi Pfeiffer 1840; Archiv fr Naturg. 1:250 (substitute name for Helix pusilla Pfeiffer). Guppya gundlachi (Pfeiffer). Tate 1870; American Journal of Conchology, 5:155.Von Martens 1892; Biol. Cent. Amer.:122 123.Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:768.Hinkley 1920; Nautilus 34:39, 48, 51.H. B. 1922; Occ. Pap. (radula).Pilsbry 1926a; Proc. Acad. Nat. Sci. Phila. 78:100.Pilsbry 1930d; Proc. Acad. Nat. Sci. Phila. 82:351.Richards 120a (shell).Basch 1958:11.Thompson 1967c:243.CorreaSandoval 1999:8.Correa-Sandoval & Rodriguez 2002; Acta

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214 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Guppya gundlachi gundlachi (Pfeiffer). Thompson 2008:708. Helix simulans C. B. Adams 1849; Contributions to Conchology, 2:35.Type Localities. Helix pusilla: Cuba. Helix simulans: Jamaica. Distribution.Florida and Texas south to Panam, Venezuela and Trinidad, and the West Indies. PANAM: Canal Zone: Isla Barro Colorado; Juan Mina (Pilsbry 1926a); Mt. Hope (Pilsbry 1930d). Prov. Panam; Cd. Panam (Pilsbry 1926a); ruins of Old Panam City (Pilsbry 1930d). Rio San Juan (Tate 1870). HONDURAS: Isla de Roatn, between Coxen Hole and French Harbor (Richards 1938). GUATEMALA, Dept. Alta Verapaz: Chama (Hinkley 1920). Dept. Izabal: Cavech; Esmeralda; Jocolo (Hinkley 1920). Dept. Petn: Tikal National Park (Basch 1958). CAMPECHE: 8.1 mi. SW of Champoton (Thompson 1967). TAMAULIPAS: near Ciudad Victoria; canyon 4 mi. W of Ciudad Victoria (Pilebry 1904). QUINTANA ROO: 4.0 mi. E of Xpujil, Campeche; 1.5 km. NNE of San Miguel, Isla Cozumel (Thompson 1967). TAMAULIPAS: numerous localities (Correa-Sandoval & Rodriguez 2002). VERACRUZ: Coatotolapam (H. B. Baker 1922); Naranjo Guppya gundlachi orosciana Von Martens 1892Guppya orosciana (shell). Guppya gundlachi orosciana Von Martens. H. B. Baker 1928c; anatomy.H. B. Baker 1930a; Occ. Pap. Mus. Zool. Univ. Mich. (220):23.Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):12.Thompson 2008:709. Distribution.COSTA RICA: type locality. GUATEMALA; Dept. Alta Verapaz: Cobn (Von Martens 1892). Dept. Petn: Tikal National Park (Basch 1959). Sumidera, 2625 ft. alt. (H. B. Baker 1930a). Guppya jalisco Pilsbry 1919Guppya jalisco Pilsbry 1919; Proc. Acad. Nat. Sci. Phila. 71:216; pl. Type Locality.Guadalajara, Jalisco, Mxico. Holotype ANSP 44830. Distribution.JALISCO: Guadalajara. Guppya micans (Angas 1879)Stenopus micans Angas 1879; Proc. Zool. Soc. Lond. 47:485; pl. 40, Guppya micans (Angas). Von Martens 1892; Biol. Cent. Amer.:121 (shell).Thompson 2008:710.Type Locality.Costa Rica. Alta Tararia (Von Martens 1901). Guppya micra Pilsbry 1904Guppya micra Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:769; pl. 2008:710.Type Locality.Patzcuaro, Michoacn, Mxico. Lectotype ANSP 85913a (Baker 1963:236). Distribution.MICHOACAN: Patzcuaro (Pilsbry 1904). TAMAULIPAS: canyon 4 mi. W of Cd. Victoria Gyppya montanicola Dall 1926Gyppya montanicola Dall 1926; Proc. Calif. Acad. Sci., ser. 4 Type Locality.North slope of Cerro Evermann, Isla Socorro, Nayarit, Mxico; 2000 ft. alt. Holotype CAS 2201. Distribution.Known only from the type locality. Guppya perforata Dall 1926Guppya perforata Dall 1926; Proc. Calif. Acad. Sci., ser. 4 15:478; Type Locality.Isla Mara Madre, Islas Maras, Nayarit, Mxico. Holotype CAS 2198. Distribution.NAYARIT: known only from Isla Mara Madre and Isla Maria Magdelena (Dall 1926). Guppya socorroana Dall 1926Guppya socorroana Dall 1926; Proc. Calif. Acad. Sci., ser. 4 Type Locality.Isla Socorro, Nayarit, Mxico; 2000 ft alt. Holotype CAS 2199. Distribution.NAYARIT: Isla Socorro; Isla Maria Magdelena (Dall 1926). Guppya spirulata (Pfeiffer 1845)Helix sirulata Pfeiffer 1845; Monogr. Helic. Vivent., 1:37. Guppya (?) spirulata (Pfeiffer). Von Martens 1892; Biol. Cent. Amer.:123. Guppya spirulata (Pfeiffer). Thompson 2008:711.Type Locality.Realejo, Dept. Chinandega, Nicaragua. Distribution.Known only from the type locality. Guppya sterkii punctum H. B. Baker 1930Guppya sterkii punctum Baker 1930a; Occ. Pap. Mus. Zool. Univ. 2008:711.Type Locality.Near Necaxa, Puebla, Mxico; 2625 ft. alt. Holotype in the UMMZ. Distribution.PUEBLA: known only from the immediate vicinity of the type locality.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 215 Genus Habe 1948Habe 1946; Venus, 14:167.Type Species.Macroclamys fulgans Gude 1900. Distribution.Japan. Widely introduced throughout the tropical regions of the world. Taxonomy.Two species are recognized. One is introduced into the study area. (Gude 1900)Macrochlamys fulgens Gude 1900; Proc. Malacol. Soc. London4:70. (Gude 1900).Habe 1946; Venus, 14:167.Azuma 1982; Colored Illustrations Land Snails Japan [in Trop. 46:369 (life history).Barrientos 2000; Rev. Biol. Trop. 48; 71 (ecology, distribution).Thompson 2008:712.Type Locality.Japan. Distribution.Japan. Widely introduced in the tropical Humid tropical zones throughout the country (Barrientos 2000). Genus Velifera W. G. Binney 1879Velifera Binney 1879; Ann. New York Acad. Sci. 1:257.Type Species.Velifera gabbi W. G. Binney 1879. Distribution.Costa Rica. Taxonomy.The genus is monotypic. Velifera gabbi W. G. Binney 1879Velifera gabbi Binney 1879; Ann. New York Acad. Sci. 1:257, ibid.Type Locality.Central Costa Rica, on the banks of the Rio Blanco, 3000 ft. alt. Distribution.Known only from the type locality. Superfamily GASTRODONTOIDAE Tryon 1866 Family GASTRODONTIDAE Tryon 1866 Distribution.This is a Nearctic family consisting of six genera. Two occur in Mxico and Central America. Genus Striatura Morse 1864Striatura Morse 1864; Jour. Portland Soc. Nat. Hist. 1:17.Baker 1928c; Proc. Acad. Nat. Sci. Phila. 80:33.Pilsbry 1946; Land Moll. N. Amer. 2:487.Type Species.Helix milium Morse 1859. Distribution.North America, Hawaiian Islands. Taxonomy.Three subgenera are recognized. One subgenus occurs in Mxico and Central America. Subgenus Striatura Morse 1864. Distribution.North America south to Nicaragua. Taxonomy.Two species occur in Mxico and Central America. Striatura (Striatura) meridionalis (Pilsbry & Ferriss 1906)Vitrea milium meridionalis Pilsbry & Ferriss 1906; Proc. Acad. Nat. Sci. Phila. 58:152. Striatura (Pseudohyalina ) meridionalis (Pilsbry & Ferriss). Baker 1930a; Occ. Pap. Mus. Zool. Univ. Mich. (220):36; pl. 11, Striatura (Striatura) meridionalis (Pilsbry & Ferriss). Pilsbry 1946; 2008:713. Striatura meridionalis (Pilsbry & Ferriss). Bequaert & Miller Type Locality.Along the Guadalupe River above New Braunfels, Texas. Holotype ANSP 90724. Distribution.NICARAGUA, Dept. Leon (Prez & alt. (Bequaert & Miller 1973). NUEVO LEN: Pablillo. (Pilsbry 1946). Striatura (Striatura) pugetensis (Dall 1895)Patulastra? pugetensis Dall 1895; Nautilus 8:130. Pilsbry 1896; Nautilus 9:18. Striatura milium pugetensis (Dall). Pilsbry 1927; Proc. Calif. Acad. Sci. 16:169. Striatura pugetensis (Dall). Baker 1930; Occ. Pap. Mus. Zool. Univ. Striatura (Pseudohyalis) pugetensis (Dall). Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:129. Striatura (Striatura) pugetensis (Dall). Thompson 2008:714.Type Locality.Seattle, Washington. Holotype USNM 107541. Distribution.Found from British Columbia and Montana south to southern California and Baja California. BAJA CALIFORNIA NORTE: Isla Guadalupe, hills above Northeast Anchorage, 300 m alt.; 3.2 km N of S end of island on E side (Smith et al. 1990). Genus Zonitoides Lehmann 1862Zonitoides Lehmann 1862; Malak. Bltter 9:111.Type Species.Helix nitidus Mller 1774. Distribution.Throughout most of the temperate Holarctic realm. Taxonomy.Three subgenera are recognized. One subgenus occurs in the study area. Subgenus Zonitella Baker 1828Zonitella Baker 1828; Proc. Acad. Nat. Sci. Phila. 80:33, 39.Pilsbry 1946; Land Moll. N. Amer. 2:475, 480.Type Species.Helix arboreus Say 1816. Distribution.North America south to Costa Rica. Taxonomy.Seven species occur in the study area. Aside from Zonitoides arboreas and Z. hoffmanni, the taxonomic status of the others remains to be determined. Zonitoides (Zonitella ) arboreus (Say 1816)Helix arboreus Say 1816; [Nicholsons] American Edit. British

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216 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Hyalinia arborea (Say). Von Martens 1892; Biol. Cent. Amer.:116 Zonitoides (Zonitella ) arboreus (Say 1816). Pilsbry 1946; Land Nautilus 62:137, 138.Correa-Sandoval, Gutierrez, & Reza 1998:14.Thompson 2008:715.Type Locality.Probably Philadelphia, Pennsylvania (Baker 1930a). Distribution.Throughout North America south to Costa Rica, Cuba, Hispaniola, Jamaica, and the Lesser Antilles. Introduced widely by its association with live plants. GUATEMALA, Dept. Chimaltenango: Finca Montserrat (Haas 1949). Dept. Zacapa: Santa Clara, N of Cabaas, Sierra de las Minas (Haas 1949). PUEBLA: Necaxa (Baker 1930a). SAN LUS POTOS: km 48 on road from Cd. Valles-Agua Zonitoides (Zonitella ) hoffmanni (Von Martens 1892)Hyalina hoffmanni Von Martens 1892; Biol. Cent. Amer.:115, pl. 6, Zonitoides hoffmanni (Von Martens). Pilsbry 1920:5.Baker 1929; Thompson 2008:716.Type Locality.Quebrada Honda, Costa Rica. Distribution.COSTA RICA, Prov. Cartago: Hacienda El Roble, between Volcan Irazu and Volcan Turrialba (Von Cachi, 3300 ft. alt. (Pilsbry 1920). Prov. San Jos: San Jos (Von Martens 1892). PANAM, Canal Zone. Prov. Chiriqu: Boquete (Pilsbry 1926a). Zonitoides (Zonitella ) glomerulus (Von Martens 1892)Hyalinia glomerula Von Martens 1892; Biol. Cent. Amer.:115; pl. 6, Zonitoides (Zonitella ) glomerulus (Von Martens). Thompson 2008:716.Type Locality.San Luis, Dept. Petn, Guatemala. Distribution.Known only from the type locality. Zonitoides (Zonitella ) Pilsbry 1926Zonitoides Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. Holotype ANSP 140311. Distribution.Known only from the type locality. Zonitoides (Zonitella ) nitidopsis (Morelet 1851)Helix nitidopsis Morelet 1851; Test. Noviss. II:8. Zonites (Hyalinia ) nitidopsis (Morelet). Fischer & Crosse 1872:178. Hyalinia nitidopsis (Morelet). Von Martens 1892; Biol. Cent. Zonitoides (Zonitella ) nitidopsis (Morelet). Thompson 2008:717.Type Locality.Salama, Dept. Baja Verapaz, Guatemala. Distribution.Known only from the type locality. Zonitoides (Zonitella ) ostauri Pilsbry 1926Zonitoides ostauri Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:100 Type Locality.Drift of San Juan Creek, Prov. Bocas del Toro, Panam. Holotype ANSP 140714. Distribution.Known only from the type locality. Zonitoides (Zonitella ) tehuantepecensis (Crosse & Fischer 1870)Zonites tehuantepecensis Crosse & Fischer 1870; Jour. de Conchy. 18:297. Zonites (Hyalinia ) tehuantepecensis Crosse & Fischer. Fischer & Hyalinia tehuantepecesis (Crosse & Fischer). Von Martens 1892; Biol. Cent. Amer.:116. Zonitoides (Zonitella ) tehuantepecensis (Crosse & Fischer). Thompson 2008:717.Type Locality.Tehuantepec, Oaxaca, Mxico. Distribution.GUATEMALA, Dept. Alta Verapaz: Senahu (Von Martens 1892). OAXACA: Tehuantepec. Superfamily ZONITOIDEA Mrch 1864 Family ZONITIDAE Mrch 1864 Subfamily ZONITINAE Mrch 1864 Distribution.North America and Europe. Taxonomy.The subfamily contains many genera (Zilch 1959:242). Twenty-eight (28) species of Zonitidae are recognized in Mxico and Central America. Genus Glyphyalinia Von Martens 1892Glyphyalinia Von Martens 1892; Biol. Cent. Amer.:117.Baker 1928; Proc. Acad. Nat. Sci. Phila. 80:19.Pilsbry 1946; Land Moll. N. Amer. 2:255, 288.Type Species.Helix indentata Say 1822 (Pilsbry 1946). Distribution.North and Middle America. Taxonomy.Seven species are recognized. One species and two subspecies occur in the study area. Glyphyalinia indentata indentata (Say 1822)Helix indentata Say 1822:372. Hyalinia (Glyphyalinia ) indentata (Say). Von Martens 1892; Biol. Cent. Amer.:117. Vitrea indentata (Say). Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:768. Retinella (Glyphyalinia )indentata (Say). Baker 1930b; Proc. Acad. Nat. Sci. Phila. 82:209.Pilsbry 1946; Land Moll. N. Rev. Biol. Trop. 41:675.Correa-Sandoval 1997; Rev. Biol. Glyphyalinia indentata (Say). Naranjo-Garcia 1991:168, 169.Correa-Sandoval, Garca-Cubas & Reguero 1998:41.Thompson 2008:719. northern Philadelphia and New Jersey.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 217 Distribution.Generally distributed in the temperate part (Pilsbry 1904). MORELOS: Yautepec (Pilsbry 1891). LEN: Iturbide (Correa-Sandoval 1997); Montemorelos, Zaragosa (Correa-Sandoval & Salzar (2005); Santiago (Correa-Sandoval 1993). SAN LUS POTOS: Ro Gallinas, Sandoval et al. 1998). SONORA: numerous localities in the eastern part of the state (Naranjo-Garcia 1991). Remarks.Some Central American and Mexican records may pertain to the following subspecies. Glyphyalinia indentata paucilirata (Morelet 1849)Helix paucilirata Morelet 1851; Test. Noviss. II:8. Zonites (Hyalinia ) paucilirata (Morelet). Fischer & Crosse 1872:173.Pilsbry 1891; Proc. Acad. Nat. Sci. Phila. 43:312. Hyalinia pauciliraa (Morelet). Von Martens 1892; Biol. Cent. Retinella (Glyphyalinia ) indentata paucilirata (Morelet). Baker 1930a; Occ. Pap. Mus. Zool. Univ. Mich. (220):24.Baker & Roth 1990; Proc. Calif. Acad. Sci. 47:129. Zonites indentatus var. umbilicatus Cocherell 1893; British Nat., 3:81. Glyphyalinia indentata paucilirata (Morelet). Thompson 2008:219.Type Localities. Helix paucilirata : Salama, Dept. Baja Verapaz, Guatemala. Zonites indentatus var. umbilicatus : Lee Co., Texas. Distribution.Widely distributed in the mid-west and central United states to Arizona, New Mexico and Texas south to Guatemala. GUATEMALA, Dept. Alto Verapaz: Salama. BAJA CALIFORNIA SUR: Sierra Laguna; Sierra de la Giganta, ca. 1.6 km from Pie de la Cuesta, along trail to Guajademi, 640 m alt.; 19 km SW of San Miguel Comond; San Javier; along road from Valle Perdido to Bajada del Molino, 430 m alt.; summit of trail from La Burrera to La Laguna, 2100 m alt.; La Laguna 1980 m alt. (Smith et al. 1990). DIST. FEDERAL: San Juan Teotihuacan, 7510 ft. alt. (Baker 1930a). PUEBLA: Necaxa, 4430 ft. alt. (Baker 1930a). Pilsbry (1946) recorded this snail from DURANGO, JALISCO, MICHOACAN, and MORELOS Genus Cooke 1921 Cooke 1921; Occasional Pap. Bernice Bishop Museum, 7:271.Baker 1941; Bulletin Bernice P. Bishop Museum, 166:328. Perpolita Baker 1928c; Proc. Acad. Nat. Sci. Phila. 80:15.Pilsbry 1946; Land Moll. N. Amer. 2:255, 256.Type Species.Types by original designations. : Vitrea pauxillus Gould 1846. Perpolita : Helix electrina Gould 1841. Distribution.Palearctic realm and Hawaiian islands. Taxonomy.Many Palearctic, Nearctic and Hawaiian species. One species with two subspecies occurs in Mxico. (Pfeiffer 1867)Helix subhyalina Pfeiffer 1867; Malak. Bltt. 14:196. Hyalinia subhyalina Zonites (Hyalinia ) subhyalina (Pfeiffer). Fischer & Crosse 1872; 172.Pilsbry 1891; Proc. Acad. Nat. Sci. Phila. 43:312. Hyalinia subhyalina (Pfeiffer). Von Martens 1892; Biol. Cent. Amer.:117. R[etinella ] (Perpolita ) subhyalina subhyalina (Pfeiffer). Baker 1930a; Occ. Pap. Mus. Zool. Univ. Mich. 220:28.Baker 1930b; Retinella () subhyalina (Pfeiffer). Baker 1941; Bulletin Bernice P. Bishop Museum, 166:329. (Pfeiffer). Thompson 2008:720.Type Locality.Uncertain; Mirador or near Vera Cruz, Veracruz, Mxico (Strebel & Pfeffer 1880). Distribution.PUEBLA: Necaxa, 4265 ft. alt. (Baker 1930a). VERACRUZ: Orizaba (Baker 1930a). (Dall 1926)Zonitoides socorroensis Dall 1926; Proc. Calif. Acad. Sci. (4) Retinella ((Perpolita ) subhyalina socorroensis (Dall). Baker 1930b; Proc. Acad. Nat. Sci. Phila. 82:200. (Dall). Thompson 2008:721.Type Locality.Cerro Evermann, Isla Socorro, Nayarit, Mxico; 2000 ft. alt. Holotype CAS 2206. Distribution.Known only from the type locality. Genus Hawaiia Gude 1911Hawaiia Gude 1911; Proc. Malac. Soc. London 9:272.Baker 1930a; Occasional Papers of the Museum of Zoology, University of Michigan, (220):34.Pilsbry 1930b; Proc. Acad. Nat. Sci. Phila. 82:247.Pilsbry 1946; Land Moll. N. Amer. 2:418. Baker 1928c; Proc. Acad. Nat. Sci. Phila. 80:24.Type Species.Hawaiia: Helix kawaiensis Pfeiffer 1854 (Helix minuscule Binney 1840) (see Pilsbry 1930b:247). : Helix minuscule Binney 1840. Distribution.North America from Alaska and Maine to Florida and south to Costa Rica, Cuba, Hispaniola, Jamaica, Puerto Rico, and the Virgin Islands. Taxonomy.The genus contains two or more species and several subspecies. Two species occur in the study area. Hawaiia minuscula minuscula (Binney 1840)Helix minuscule Binney 1840; Boston Journal of Natural History, 3:435. Chanomphalus minusculus (Binney). Strebel 1880; Beitrag. Mex. Landund Sssw.-Conch. IV:19. Pseudohyalina minuscule (Binney). Von Martens 1892; Biol. Cent. Amer.:124. Zonitoides minusculus (Binney). Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:768.Hinkley 1920; Nautilus 34:39. (Binney 1840). Baker 1928c; Proc. Acad.

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218 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Hawaiia minuscula (Binney 1840). Bequaert & Clench 1938; Pub. Carnegie Inst. Wash. (491):253, 254.Pilsbry 1946; Land Bequaert & Miller 1973:145.Correa-Sandoval 1997; Rev. Biol. Trop. 44/45:140.Correa-Sandoval, Garca-Cubas & Reguero 1998:14.Correa-Sandoval 1999:8.Correa-Sandoval 2002:183. Hawaiia minuscula minuscula (Binney). Thompson 2008:721.Type Locality.Ohio. Lectotype ANSP (Pilsbry 1946:421). Distribution.Widely distributed as for the genus. (Hinkley 1920). Dept. Petn: Tikal National Park (Basch 1959). CAMPECHE: Isla de Carmen; Palizada (Von Martens 1892). CHIAPAS: Palenque. CHIHUAHUA, SONORA and BAJA CALIFORNIA (Bequaert & Miller 1973). NUEVO LEN: Topo Chico, near Monterrey (Pilsbry 1904); Iturbide (Correa-Sandoval 1997). SAN LUS POTOS: numerous localities in eastern part of state (Correa-Sandoval et al. numerous localities (Correa-Sandoval & Rodriguez 2002). YUCATN: Sazich Cave, Calcehtok (Bequaert & Clench Sandoval 1999). Hawaiia minuscula neomexicana (Cockerell & Pilsbry 1900)Zonitoides neomexicans Cocherell & Pilsbry 1900; Nautilus 13:114. (Cockerell & Pilsbry). Baker (anatomy). Hawaiia minuscula neomexicana (Cockerell & Pilsbry). Baker 1930a; Occas. Pap. Mus. Zool. Univ. Michigan, (220):35.(shell).Thompson 2008:722. Punctum pygmaeum var. albeolum Dall 1926; Proceedings California Academy of Sciences, ser. 4, 15:481.Type Localities. Zonitoides neomexicanus: Dripping Spring, Organ Mountains, Texas. Punctum pygmaeum var. albeolum: Isla Maria Magdelena, Islas Maras, Nayarit, Mxico. Holotype CAS 2203. Distribution.Texas and New Mexico south to Nayarit and Puebla, Mxico; Colombia. NAYARIT: Isla Maria Magdelena; Mt. Evermann, Isla Mara Madre, 2000 ft. alt. (Dall 1926). PUEBLA: Necaxa (Baker 1930a). COLOMBIA: Isla de Providencia (Pilsbry 1930). Hawaiia minuscula permodesta (Strebel 1880)Hyalinia permodesta Strebel 1880; Beitrag. Mex. Landund Sssw.(Strebel). Baker 1929; Proc. ?Hawaiia minuscula permodesta (Strebel 1880). Pilsbry 1946; Thompson 2008:723. Hawaiia minuscule Type Locality.Mirador, Veracruz, Mxico. Distribution.CAMPECHE: 8.1 mi. SW of Champoton; 7.2 mi. SW of Pixtun (Thompson 1967). VERACRUZ: Mirador. various regions of the study area is problematic. Thompson (1967) could not assign specimens from Campeche to subspecies; and Smith et al. (1990) hesitated to assign specimens from Baja California Sur to species. Hawaiia pentagyra (Pilsbry 1907)Zonitoides pentagyra 8 (shell).Hinkley 1907; Nautilus 21: Hawaiia pentagyra (Pilsbry). Thompson 2008:724.Type Locality.River drift, Tampico, Tamaulipas, Mxico. Holotype ANSP 93796. Distribution.Known only from the type locality. Genus MesomphixMesomphix 1911; Proc. Acad. Nat. Sci. Phila. 63:478.Baker 1930a; Occas. Pap. Mus. Zool. Univ. Michigan, (220):24.Pilsbry 1946, Land Moll. N. Amer. 2:305, 319. Omphix Pilsbry 1911; Proc. Acad. Nat. Sci. Phila. 63:479. Micromphix Pilsbry 1911; Proc. Acad. Nat. Sci. Phila. 63:479.Type Species.Mesomphix: 1819 (= Baker 1933; not Helix l Linnaeus 1766. Omphix: Helix inornatus Say 1821. Micromphix: Helix subplasus Binney 1842. Distribution.Eastern North America from Ontario and Quebec to Guatemala, in regions of moderate or high humidity (Pilsbry 1946). Taxonomy.Four subgenera are recognized. Three occur in the study area. The subgenus Mesomphix is an Appalachian group primarily, with some species extending outward from the mountains into adjacent regions. It does not occur in the study area. Subgenus OmphalinaOmphalina Pilsbry 1894; Proc. Acad. Nat. Sci. Phila. 46:14.Pilsbry 1911; Proc. Acad. Nat. Sci. Phila. 63:469. Von Martens 1892; Biol. Cent. Amer.:104.Baker 1930a; Occas. Pap. Mus. Zool. Univ. Michigan, (220):26.Type Species.Omphalina cuprea Distribution.Eastern North America from Quebec south to Guatemala. Taxonomy.Omphalina which occur in Mxico and Central America. Mesomphix ( Omphalina) lucubratus lucubratus (Say 1829)Helix lucubrata Say 1829; New Harmony Disseminator, II:229. Hyalina lucubrata (Say). Tryon 1866; Man. Conch. 2:247; pl. 18,

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 219 Omphalina lucubrata (Say). Von Martens 1892; Biol. Cent. Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:767. Mesomphix (Omphalina) lucubratus lucubratus (Say). Baker 1930a; Occas. Pap. Mus. Zool. Univ. Michigan, (220):27.Thompson 2008:725.Type Locality.Mxico. Distribution.OAXACA. VERACRUZ: Las Vigas; Mesomphix (Omphalina) lucubratus caducus (Pfeiffer 1846)Helix caduca Pfeiffer 1846; Malak. Bltt. 3:159. Zonites caducus Binney & Bland; land and freshwater shells of Moreletia caduca (Pfeiffer). Strebel 1880; Beitrag. Mex. Landund Omphalina caduca (Pfeiffer). Von Martens 1892; Biol. Cent. Amer.:108. Mesomphix (Omphalina) lucubratus caducus (Pfeiffer). Baker 1930a; Occas. Pap. Mus. Zool. Univ. Michigan, (220):26.Thompson 2008:725. Distribution.VERACRUZ: Quilate, near Misantla; Soncoautla, near Coatepec; San Antonia Huatusco; Cerro de (Von Martens 1892); Peuela to Sumidero, 2625 ft. alt. (Baker 1930a). Mesomphix (Omphalina) lucubratus deppianus (Von Martens 1892)Omphalina lucubrata deppiana Von Martens 1892; Biol. Cent. Mesomphix (Omphalina) lucubratus deppianus (Von Martens). Thompson 2008:726.Type Locality.Orizaba, Veracruz, Mxico. Distribution.Known only from the type locality. Mesomphix (Omphalina) lucubratus fasciatus (Fischer & Crosse 1872)Zonites (Moreletia) caducus fasciata Fischer & Crosse Moreletia angiomphala Strebel 1880; Beitrag. Mex. Landund Omphalina lucubrata fasciata (Fischer & Crosse). Von Martens 1892; Biol. Cent. Amer.:107. Mesomphix (Omphalina) lucubratus fasciatus (Fischer & Crosse). Thompson 2008:726.Type Localities. Zonites fasciata : Jacale, near the Volcn de Orizaba, Mxico. Moreletia angiomphala: not Distribution.VERACRUZ: Cajetas; San Antonia del Monte, near Neolingo; Cuautlatitlan, between Jico and Perote; Ixhuacan, between Jalapa and Quimistla; Coatepec, near jalapa (Strebel 1880); Jalapa; Jacale; Mirador (Von Martens 1892). Remarks.Baker (1930a:2) stated that angiostomus might be a distinct species. Mesomphix (Omphalina) (Fischer & Crosse 1872)Zonites (Moreletia ) lucubratus Fischer & Crosse 1872:161; Omphalina (Fischer & Crosse). Von Martens 1892; Biol. Cent. Amer.:107. Mesomphix (Omphalina) (Fischer & Crosse). Thompson 2008:727.Type Locality.Tepanasacualco, Oaxaca, Mxico. Distribution.Known only from the type locality. Mesomphix (Omphalina) lucubratus strebelianus (Von Martens 1892)Moreletia fuliginosa form B. Strebel 1880; Beitrag. Mex. Landund Omphalina caduca strebeliana Von Martens 1892; Biol. Cent. Amer.:108. Mesomphix (Omphalina) lucubratus strebelianus (Von Martens). Baker 1930a; Occas. Pap. Mus. Zool. Univ. Michigan, (220):28.Thompson 2008:727.Type Locality.Near Misantla, Veracruz, Mxico (Baker 1930:28). Distribution.PUEBLA: Necaxa (Baker 1930a). VERACRUZ: Quilate; Hirial; Palpoala; Misantla; Cajetes, near San Juan Miahuatlan, between Jalapa and Misantla (Von Martens 1892). Mesomphix (Omphalina) martensianus (Pilsbry 1903)Omphalina martensiana Pilsbry 1903; Proc. Acad. Nat. Sci. Phila. Mesomphix (Omphalina) martensianus (Pilsbry). Baker 1930a; Occas. Pap. Mus. Zool. Univ. Michigan, (220):28.Thompson 2008:727.Type Locality.Huehuetenango, Guatemala. Holotype ANSP 85521. Distribution.Known only from the type locality. Mesomphix (Omphalina) paradensis (Pfeiffer 1860)Helix paradensis Pfeiffer 1860; Malak. Bltt. 7:233. Zonites (Moreletia ) paradensis (Pfeiffer). Fischer & Crosse Moreletia paradensis (Pfeiffer). Strebel 1880; Beitrag. Mex. Land(shell). Moreletia dohrnii Strebel 1880; Beitrag. Mex. Landund Sssw.Mesomphix (Omphalina) paradensis (Pfeiffer). Baker 1930a; Occas. Pap. Mus. Zool. Univ. Michigan, (220):28.Thompson 2008:728.Type Locality.Helix paradensis: Parada, Oaxaca, Mxico. Moreletia dohrnii Distribution.OAXACA: Parada. VERACRUZ:

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220 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Cajetes, near San Juan Miahuatlan; Consolapa, near Coatepec; Coautlatitlan (Strebel 1880). Mesomphix (Omphalina) pittieri (Bartsch 1909)Omphalina pittieri Mesomphix (Omphalina) pittieri (Bartsch). Thompson 2008:728.Type Locality.In the vicinity of Secanquim, Dept. Alta Verapaz, Guatemala. Holotype USNM 207783. Distribution.Known only from the type locality. Mesomphix (Omphalina) zonites (Pfeiffer 1845)Helix zonites Pfeiffer 1845; Proc. Zool. Soc. Lond. 13:127. Zonites (Moreletia ) metonomasticus Fischer & Crosse 1872:157; pl. Moreletia metonomastica (Fischer & Crosse). Strebel 1880:5. Omphalina (Moreletia ) zonites (Pfeiffer). Von Martens 1892; Biol. Cent. Amer.:106. Mesomphix (Omphalina) zonites (Pfeiffer). Baker 1930a; Occas. Pap. Mus. Zool. Univ. Michigan, (220):28.Thompson 2008:729.Type Localities. Helix zonites: not stated. Zonites metonomasticus: Tuxtla [Gutierrez], Chiapas, Mxico. Distribution.CHIAPAS: Tuxtla Gutierrez. Subgenus Moreletia Gray 1855Moreletia Gray 1855; Catalogue of the pulmonata or air breathing Mollusca in the collection of the British Museum:148.Type Species.Helix euryomphala Pfeiffer 1845. Distribution.Guatemala. Taxonomy.The subgenus contains a single species. Mesomphix (Moreletia ) euryomphala (Pfeiffer 1845)Helix euryomphala Pfeiffer 1845; Proc. Zool. Soc. Lond. 13:127. Zonites (Moreletia ) euryomphala (Pfeiffer). Gray 1855; Catalogue of the Pulmonata or air-breathing Mollusca in the collection of the British Museum:148.Fischer & Crosse 1872:155; pl. 7, Moreletia euryomphala (Pfeiffer). Von Martens 1875; Proc. Zool. Soc. Lond. 43:648.Strebel 1880:5. Mesomphix euryomphala (Pfeiffer). van der Schalie 1940:4. Omphalina (Moreletia ) euryomphala (Pfeiffer). Von Martens 1892; Biol. Cent. Amer.:106. Helix cymbalum Morelet 1849; Test. Noviss. I:7. Mesomphix (Moreletia ) euryomphala (Pfeiffer). Thompson 2008:229.Type Localities. Helix euryomphala: Cobn, Dept. Alta Verapaz, Guatemala. Helix cymbalum: mountain forests of Vera Paz, Guatemala. Distribution.GUATEMALA, Dept. Alta Verapaz: mountains of Alta Verapaz; Cobn (Von Martens 1892). Subgenus Zonyalina Von Martens 1865Zonyalina Von Martens 1865; Malak. Bltt. 12:16.Baker 1930a; Occas. Pap. Mus. Zool., Univ. Michigan, (220):29.Zilch 1959:242.Type Species.Helix bilineata Pfeiffer 1845. Distribution.Eastern Mxico Taxonomy.Three species are recognized. Mesomphix (Zonyalina) bilineatus (Pfeiffer 1845)Helix bilineata Pfeiffer, 1845; Proc. Zool. Soc. Lond. 13:128. Hyaline (Zonyalina) bilineata (Pfeiffer). Von Martens, Malak. Bltt. 12:16. Zonites (Moreletia ) bilineatus (Pfeiffer). Fischer & Crosse 1872:167. Zonyalina bilineatus 19 (jaw). Omphalina bilineata (Pfeiffer). Von Martens 1892; Biol. Cent. apicalis ).Pilsbry 1903; Proc. Acad. Nat. Sci. Phila. 55:707.Bequaert 1957; Bull. Mus. Comp. Zool. 116:220. Zonites (Zonyalina) bilineatus (Pfeiffer). Pilsbry 1891; Proc. Acad. Nat. Sci. Phila. 43:312. Mesomphix (Zonalina) bilineatus (Pfeiffer). Baker 1930a; Occ. Pap. Type Locality.Not stated. Distribution.CHIAPAS: Laguna Ocotal, 950 m alt. Jalapa; Pacho, near Jalapa; Cuautlatitlan; Orizaba (Von (Baker 1930a). Mesomphix (Zonyalina) modestus (Von Martens 1892)Zonites (Moreletia) bilineatus Omphalina modesta Von Martens 1892; Biol. Cent. Amer.:11011; Amer.:617. Mesomphix (Zonyalina) modestus (Strebel). Baker 1930a; Occas. Pap. Mus. Zool. Univ. Michigan, (220):29.Thompson 2008:731.Type Locality.Jalapa, Veracruz, Mxico. Distribution.VERACRUZ: Jalapa; Las Vigas, NW of Jalapa; Nacimiento de Quilate, near Misantla (Von Martens 1892). COSTA RICA, Prov. San Jos: La Palma, 1600 m alt.; La Paz, 750 m alt. Remarks.Von Martens (1901:617) recorded this species from Costa Rica, on the basis of a report by Biolley Mesomphix ( Zonyalina) tuxtlensis (Crosse & Fischer 1870)Zonites tuxtlaensis Crosse & Fischer 1870; Jour. de Conchyl. Omphalina tuxtlensis (Crosse & Fischer). Von Martens 1892; Biol. Cent. Amer.:110. Mesomphix (Zonyalina) tuxtlensis (Crosse & Fischer). Baker 1930a; Occas. Pap. Mus. Zool. Univ. Michigan, (220):29.Thompson 2008:731.Type Locality.Tuxtla [San Andres Tuxtla ?], Veracruz, Mxico. Distribution.Known only from the type locality. Genus Patulopsis Strebel 1880Patulopsis Strebel 1880; Beitrag. Mex. Landund Sssw.-Conch. IV:16.Type Species.Patulopsis carinata Strebel 1880.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 221 Distribution.Eastern Mxico in the states of Veracruz and Puebla. Taxonomy.Two subgenera are recognized. Subgenus Patulopsis Strebel 1880 Distribution.Eastern Mxico from the states of Puebla and Veracruz. Taxonomy.A single species is recognized. Patulopsis (Patulopsis) carinatus Strebel 1880Patulopsis carinatus Omphalina (Patulopsis) carinata (Strebel). Von Martens 1892; Biol. Cent. Amer.:113.Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:708. Mesomphix (Patulopsis) carinatus (Strebel). Baker 1930a; Occasional papers of the Museum of Zoology, University anatomy). Patulopsis (Patulopsis) carinatus Strebel. Thompson 2008:732.Type Locality.Woods of Pacho, near Jalapa, Veracruz, Mxico. Distribution.VERACRUZ: Jalapa; Pacho, near Jalapa; Coatepec (Von Martens 1892). PUEBLA: above Necaxa, 4700 ft. alt. (Baker 1930a). Subgenus Omphalinella Baker 1930Omphalinella Baker 1930; Occ. Pap. Mus. Zool. Univ. Mich. (220):26, 29.Type Locality. Pfeiffer 1856. Tamaulipas, Puebla, Veracruz and Guerrero. Taxonomy.Four species and two subspecies are assigned to this subgenus. Patulopsis (Omphalinella ) montereyensis montereyensis (Pilsbry 1899)Omphalina montereyensis Pilsbry 1899; Proc. Acad. Nat. Sci. Phila. 51:395.Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:767; Mesompix (Omphalinella ) montereyensis montereyensis (Pilsbry). Baker 1930a; Occ. Pap. Mus. Zool. Univ. Mich. (220):31. Mesomophix montereyensis (Pilsbry). Correa-Sandoval 1993; Rev. Biol. Trop. 41:675.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):236. Patulopsis (Omphalinella ) montereyensis montereyensis (Pilsbry). Thompson 2008:733. Mxico. Holotype ANSP 77110. Distribution.NUEVO LEN: Diente; Santiago (Correa-Sandoval 1993). TAMAULIPAS: numerous localities in the west part of the state (Correa-Sandoval & Rodriguez 2002). Patulopsis (Omphalinella ) (Pilsbry 1903) Pilsbry 1903; Proc. Acad. Nat. Mesomphix (Omphalinella ) m (Pilsbry). Baker 1930a, (220):31.Solem 1954; Nautilus 68:7.CorreaSandoval, Garca-Cubas & Reguero 1998:14.CorreaSandoval & Rodriguez 2002; Acta Zool. Mex. (86):236. Patulopsis (Omphalinella ) (Pilsbry). Thompson 2008:733).Type Locality.Caon 4 miles west of Ciudad Victoria, Tamaulipas, Mxico. Holotype ANSP 85912a. Distribution.SAN LUIS POTOS; Cd. del MazSandoval et al. 1998). NUEVO LEN: numerous localties in southern part of state (Correa-Sandoval & Salazar 2005). TAMAULIPAS: Ranch del Cielo, 7 km N of Gomez Farias; 20 km N of Chamal (Solem 1954); numerous localities (Correa-Sandoval & Rodriguez 2002). Patulopsis (Omphalinella ) salleana (Von Martens 1892) (shell) (not Pfeiffer 1856). Omphalina salleana Von Martens 1892; Biol. Cent. Amer.:112. (Substitute name for Fischer & Crosse 1872). Mesomphix (Omphilinella ) salleana (Von Martens). Baker 1930a; Occ. pa. Mus. Zool. Univ. Mich. (220):31. Patulopsis (Omphalinella ) salleana (Von Martens). Thompson 2008:734. Distribution.Known only from the type locality. Patulopsis (Omphalinella ) sculptus (Von Martens 1892)Omphalina sculpta Von Martens 1892; Biol. Cent. Amer.:113; pl. 6, Mesomphix (Omphalinella ) sculptus (Von Martens). Baker 1930a; Occ. Pap. Mus. Zool. Univ. Mich. (220):31. Patulopsis (Omphalinella ) sculptus (Von Martens). Thompson 2008:734.Type Locality.Omilteme, Guerrero, Mxico. Distribution.Known only from the type locality. Patulopis (Omphalinella ) (Pfeiffer 1856) Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:318.(shell). Mesomphix (Omphalinella ) (Pfeiffer). Baker 1930a; Occasional papers of the Museum of Zoology, Patulopis (Omphalinella) (Pfeiffer). Thompson 2008:734. Sumidero, 1300 ft. alt. (Baker 1930a) Patulopsis (Omphalinella ) (Strebel 1880)Zonyalina jalapensis Strebel 1880; Beitrag. Mex. Landund Sssw.-

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222 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) (Strebel). Von Martens 1892; Biol. Cent. Amer.:111. Mesomphix (Omphalinella ) (Strebel). Baker 1930a; Occas. Pap. Mus. Zool. Univ. Michigan, (220):31. Patulopsis (Omphalinella ) (Strebel). Thompson 2008:725.Type Locality.Jalapa, Veracruz, Mxico. Distribution.GUATEMALA, Dept. Alta Verapaz: Cobn. VERACRUZ: Jalapa; environs of Jalapa; Pacho; Consolapa; Soneautla; Coatepec; Dos Arroyos, between Jalapa and Naolingo (Von Martens 1892). Genus Pycnogyra Strebel 1880Pycnogyra Strebel 1880; Beitrag. Mex. Landund Sssw.-Conch. IV:20.Baker 1928; Proc. Acad. Nat. Sci. Phila. 80:27.Zilch 1959:240.Type Species.Helix berendti Pfeiffer 1861. Distribution.Eastern Mxico from the states of Veracruz and Puebla. Taxonomy.The genus is monotypic. Pycnogyra berendti (Pfeiffer 1861)Helix berendti Pfeiffer 1861; Malak. Bltt. 8:72. Pycnogyra berendti 55:768.Baker 1928; Proc. Acad. Nat. Sci. Phila. 80:27; pl. Pseudohyalina berendti (Pfeiffer). Von Martens 1892; Biol. Cent. Amer.:126.Type Locality.Hacienda Mirador, near Jalapa, Veracruz, Mxico. Distribution.VERACRUZ: Mirador; Coatepec; Consolapa; Dos Arroyos (Von Martens 1892); Texolo (Pilsbry 1904). PUEBLA: Necaxa (Baker 1928). Superfamily LIMACOIDEA Lamark 1801 Family LIMACIDAE Lamarck 1801 Distribution.Western Palearctic region. Three genera have been introduced widely through human agency. A single genus, Deroceras, occurs naturally in North America. Genus Limax Linnaeus 1758 Type Species.Limax maximus Linnaeus 1758. Distribution.Europe, adjacent Asia, and Africa. Introduced widely in temperate and tropical areas of the world. Taxonomy.Numerous species, two of which have been introduced into the study area. Limax maximus Linnaeus 1758 Type Locality.Sweden. Distribution.Europe, Asia Minor, and Algeria. MXICO: Guajimalpa to El Desierto de Leones, 8525 ft. alt. (H. B. Baker 1930a). Linnaeus 1758 Type Locality.Western Europe. Distribution.PUEBLA: Huachinango, 4300 ft. alt. (H. B, Baker 1930)4580ft. alt. (H. B. Baker 1930a). Genus Deroceras Type Species. Mller 1774. Distribution.Entire Palearctic region and both North America and South America. Widely disseminated by human activities. Taxonomy.Over 70 species have been described, but fewer than half are recognizably described. Three occur in the study area, two of doubtful validity. Deroceras cobanensis (Crosse & Fischer 1872)Limax cobanensis Crosse & Fischer 1872; Jour. de Conchyl. 20:159.Von Martens 1898; Biol. Cent. Amer.:349. Deroceras cobanenesis (Crosse & Fischer). H. B. Baker 1930a; Occas. Pap. Mus. Zool. Univ. Michigan (220):42.Thompson 2008:737.Type Locality.Cobn, Dept. Alta Verapaz, Guatemala. Distribution.Known only from the type locality. Taxonomy.The taxonomic status of this species is questionable (H. B. Baker 1930a). Deroceras costaricensis (Cockerell 1890)Agriolimax californicus subsp. costaricensis Cockerell 1890; Ann. & Mag. Nat. Hist. 6:278. Agriolimax costaricesnsis Cockerell. Von Martens 1898; Biol. Cent. Amer.:345. Deroceras costaricensis (Cockerell). Thompson 2008:737.Type Locality.Costa Rica. Distribution.Known only from the type locality. Taxonomy.The taxonomic status of this species requires further study. (Mller 1774) Mller 1774; Verm. Terr. et Fluv., II:1. Limax (Hyrolimax) Mller. Von Martens 1898; Biol. Cent. Amer.:347. (Mller). H. B. Baker 1930; Occ. Pap. Mus. Zool. 7 (terminal genitalia).Pilsbry 1948; Land Moll. N. Amer. Thompson 2008:739.Type Locality.Denmark Distribution.North America generally, from the Arctic to Central America. The southern limit is not determined. Von Martens (1898) gives the following records. COSTA RICA, Prov. San Jos: Borubeta, on the Rio Uren. NICARAGUA, Dept. Chontales: Javali, in Chontales. Dept. 2002). GUATEMALA, Dept. Alta Verapaz: Cobn. Dept. Guatemala: Guatemala City. Dept. Sacatepquez: Antigua. Dept. Totonicapan: Plateau of Totonicapan. VERACRUZ: Jalapa

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 223 Taxonomy.H. B. Baker (1930a) lists the following names from the study area as probable synonyms of Deroceras : Limax gracilis Limax campestris Binney 1841; Limax semitectus Mch 1857; Krynichia americana Tata 1870; Limax guatemalensis Crosse & Fischer 1870; Limax stenuras Strebel & Pfeffer 1880; Limax jalapensis Strebel & Pfeffer 1880; Limax berendti Strebel & Pfeffer 1880; Adriolimax motaguensis Cockerell 1914; and Agriolimax berendti var. pictus Cockerell 1897. Genus Milax Gray 1855 Type Species.Limax gagatus Draparnaud. Distribution.Europe, particularly in the Mediterranian region. Taxonomy.About 20 species are recognized. A single species has been introduced into the study area. Milax gagatus (Draparnaud 1801) Type Locality.Not stated. Distribution.Reported for Mxico from a limited area, but can be expected to have been introduced widely in Mxico and elsewhere. MXICO: Desierto de Los Leones, 9850 ft. alt. (H. B. Baker 1930a). Superfamily ARIONOIDEA Gray 1840 Family BINNEYIDAE Cockerell 1891 Subfamily BINNEYINAE Cockerell 1891 Taxonomy.Two genera are recognized. One occurs in Baja California Norte. Genus Binneya Cooper 1863Binneya Cooper 1863; Proc. Calif. Acad. Sci. 3:62.Pilsbry & Vanatta 1898; Proc. Acad. Nat. Sci. Phila. 50:229.Pilsbry 1948; Land Moll. N. Amer. 2:732.Type Species.Binneya notabilis Cooper 1863. Distribution.Small islands off the coast of southern California and Baja California Norte. Taxonomy.Two subgenera are recognized. Both occur in Baja California Norte. Subgenus Binneya Cooper 1863 Distribution.CALIFORNIA: Santa Barbara Island; San Nicolas Island (fossil). Baja California Norte (?). Taxonomy.The subgenus is monotypic. Binneya (Binneya) notabilis Cooper 1863Binneya notablis 15.Pilsbry & Vanatta 1908; Proc. Acad. Nat. Sci. Phila. (shell).Smith, Miller, Christensen & Roth 1990; Proc. Calif. Binneya (Binneya ) notabilis Cooper. Thompson 2008:740.Type Locality.Santa Barbara Island, California. Distribution.CALIFORNIA: Santa Barbara Island, San Nicolas Island. BAJA CALIFORNIA NORTE: hills north of Bahia San Quintin. Taxonomy.The identity of the Mexican population is uncertain, and it may involve a distinct species (Pilsbry 1948). Subgenus Alluthyra Pilsbry 1948Alluthyra Pilsbry 1948; Land Moll. N. Amer. 2:737.Type Species.Binneya guadalupensis Pilsbry 1927. Distribution.Endemic to Isla Guadalupe, Baja California Norte. Taxonomy.A single species is recognized. Binney (Alluthyra) guadalupensis Pilsbry 1927Binneya guadalupensis Pilsbry 1927; Proc. Calif. Acad. Sci. 16:169; 2d (reproductive anatomy), 3 (jaw).Pilsbry 1948; Land. Moll. D, E (animal).Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:128.Thompson 2008:741.Type Locality. miles south of Northeast Anchorage, Isla Guadalupe, Baja California Norte, Mxico. Syntypes CAS 2566. Distribution.BAJA CALIFORNIA NORTE: Isla Guadalupe. FAMILY PHILOMYCHIDAE Gray 1847 Distribution.East Asia, North America south to Colombia. Taxonomy.Four genera generally are recognized. Philomycus Pallifera and Megapallifera are New World genera. Pallifera is the only genus that occurs in the study area. Genus Pallifera Morse 1864Pallifera Morse 1864; Jour. Portland Soc. Nat. Hist. 1:8.Pilsbry 1948; Land Moll. N. Amer. 2:759.Type Species.Philomychus dorsalis Binney 1842. Distribution.Eastern North America, eastern and Central Mxico, Costa Rica, Colombia. Taxonomy.Seven species are recognized. One species and three subspecies are recorded from the study area. Two other species are considered nomina dubia. Pallifera costaricensis costaricensis (Mrch 1858)Philomycus costaricensis Mrch 1858, in Vidensk, Meddelelser Kjobenh. 1857:341. Mrch 1860; Malak. Bltt. 6:109.Von Martens 1898; Biol. Cent. Amer.:346.Cockerell 1913; Nautilus 27:3. Tebennophorus costaricensis (Mrch). Binney 1879; Ann. New (radula). Pallifera costaricensis (Mrch). Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:99.Baker 1930a; Occ. Pap. Mus. Zool. Univ. Mich. 9220:15.Thompson 2008:741.Type Locality.Central Costa Rica, 2000 ft. alt.

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224 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Distribution.COSTA RICA: the type locality. PANAM, Prov. Bocas del Toro: Bocas del Toro (Cockerell 1913). Pallifera costaricensis alticola Baker 1930Pallifera costaricensis alticola Baker 1930a; Occ. Pap. Mus. de Los Leones, Mxico; 9850 ft. alt. Holotype in the UMMZ. Distribution.Known only from the type locality. Pallfera costaricensis crosseana (Strebel 1880)Tebennophorus crosseana Philomychus crosseanus (Strebel). Von Martens 1898; Biol. Cent. Amer.:346.Pilsbry 1904; Proc. Acad. Nat. Sci. Phila. 55:770. Pallfera costaricensis crosseana (Strebel & Pfeffer). Baker 1930a; Type Locality.Not given. Distribution.PUEBLA: Necaxa, 4265 ft. alt. 3400 ft. alt. (Baker 1930a); Texolo (Pilsbry 1904). Pallifera sallei (Crosse & Fischer 1869)Philomycus sallei Crosse & Fischer 1869; Jour. de Conchyl. 17:190.Thompson 2008:743.Type Locality.Playa Vicente, Veracruz, Mxico. Distribution.Known only from the type locality. Taxonomy.Referred to Pallifera but considered a nomen dubium by Baker (1930a:18). Pallifera auratus (Tate 1870)Tebennophorus auratus Tate 1870; Amer. Jour. Conch. 5:153. Philomychus auratus (Tate). Von Martens 1898; Biol. Cent. Amer.:340.Thompson 2008:743.Type Locality.Javali, Dept. Chontales, Nicaragua. Distribution.Known only from the type locality. Taxonomy.Referred to Pallifera but considered a nomen dubium by Baker (1930a:18). Family CAMAENIDAE Pilsbry 1895 Subfamily PLEURODONTINAE Ihering 1912 Genus Labyrinthus Beck 1837Labyrinthus Beck 1837.Solem 1966; Fieldiana: Zoology 50:37.Type Species.Helix otis Lightfoot 1786 (= Helix labyrinthus Deshayes 1838) (Herrmannsen 1846). Distribution.Southeastern Nicaragua south to Madre level to 2200 m alt. Taxonomy.About thirty species and ten subspecies are recognized in Labyrinthus (Solem 1966). Five species and seven subspecies occur in the study area. Labyrinthus otis orthorhinus (Pilsbry 1910)Pleurodonte (Labyrinthus) otis orthorhinus Pilsbry 1910; Proc. Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:75.Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. 82:339. Labyrinthus otis orthorhinus (Pilsbry). Solem 1966; Fieldiana: 2008:746. Panam. Holotype ANSP 101308a (Baker 1963:246). mountains near mouth of Rio Atrato (Solem 1966). PANAM, Prov. Darin: Pintupo; Tacarcuna. Comarco de San Blas: forest near Armila; hills near Rio Mandingo, head of Golfo de San Blas. Prov. Panam: mountains near Gaspasalama, Rio Mamon; Rio Indio, near Madden Lake; between Chepo and Rio Plantanal; along ridge of Rio Chico, Tucuman Airport, 9 km from Cerro Azul; slope of Cerro Cabra; hills above Arraijn; Cerro Trinidad; Cerro Campana. Canal Zone: between Tabernillo and San Pablo; near Empire; near Gorgona; Alhajuela; Gatun; Isla Barro Colorado; Rio Juanito, near Pegero, Cocl del Norte. Prov. Cocl: El Valle. Prov. Veraguas: Azuero Peninsula, Tres Puntas; Avacada; Mangillo. COSTA RICA, Prov. San Jos: Santa Maria de Dota, 1500 m alt. Labyrinthus quadridentatus quadridentatus (Broderip 1832)Caracolls quadridentata Broderip 1832; Proc. Zool. Soc. Lond. :30. Labyrinthus quadridentatus (Broderip). Von Martens 1892; Biol. Cent. Amer.:176.Von Martens 1901; Biol. Cent. Amer.:628. Labyrinthus quadridentatus quadridentatus (Broderip). Solem 2008:744.Type Locality.Golfo Dulce, Costa Rica. Distribution.PANAM, Prov. Comarca de Baru: Puerto Armuelles (Solem 1966). COSTA RICA, Prov. Puntarenas: Coto; Quebrada de la Palma, Rio Coto basin (Solem 1966); Trraba, 200 m alt.; Alto de Mano Tigre, near Trraba, 690 m alt.; El Pozo, Rio Grande de Trraba; El Pital, Rio Naranjo valley, 200 m alt.; Quebrada Labyrinthus quadridentatus biollyi Solem 1966Labyrinthus quadridentatus biollyi Solem 1966; Fieldiana: Zoology Type Locality.Guaitil de Pirris, Prov. San Jos, Costa Rica. Holotype Carnegie Museum 62.575. Distribution.COSTA RICA, Prov. Alajuela: Piedras Negras. Prov. Cartago: Azahar de Cartago. Prov. San Jos: Guaitil de Pirris (Solem 1966).

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 225 Labyrinthus triplicatus (Von Martens 1868)Helix triplicata Von Martens 1868; Malak. Bltt. 15:156. Helix (Labyrinthus) triplicata (Von Martens). Pilsbry 1889; Man. Labyrinthus triplicatus (Von Martens). Von Martens 1892; Biol. Biol. Cent. Amer.:628.Solem 1966; Fieldiana: Zoology Helix aesopus Angas 1878; Proc. Zool. Soc. Lond. 46:72; pl. 5, Helix (Labyrinthus) triplicatus var. aesopus Angas.Pilsbry 1889; Pleurodonte (Labyrinthus) aesopus (Angas). Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:127.Type Localities. Helix triplicata : Costa Rica. Helix aesopus: Buena Vista, [Prov. Alajuela?], Costa Rica. Distribution.PANAM. COSTA RICA, Prov. Cartago: Turrialba; Navarro; Platanillo, 600 m alt.; Cervantes, 1480 m alt.; Valle de Tuis, 620 m alt.; Tuis, 650 m alt.; Talamanca valley (Solem 1966). Prov. Heredia: Sarapiqu (Von Martens Labyrinthus subplanatus sipunculatus (Forbes 1850)Helix labyrinthus var. sipunculata Forbes 1850; Proc. Zool. Soc. Pleurodonte (Labyrinthus) otis sipuncuculata (Forbes). Pilsbry 1910; Proc. Acad. Nat. Sci. Phila. 62:504. Pleurodonte (Labyrinthus) sipuncuculata (Forbes). Pilsbry 1926; Labyrinthus subplanatus sipunculatus (Forbes). Solem 1966: 59 (map) .Thompson 2008:745. Helix annulifera (shell). Helix (Labyrinthus) labyrinthus annulifera Pfeiffer. Pilsbry 1889; Pleurodonte (Labyrinthus) sipunculata annulifera (Forbes). Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:74. Pleurodonte (Labyrinthus) goldmani Dall 1912; Smiths. Misc. Coll. Type Localities. Helix labyrinthus var. sipunculatus: Panam. Helix annulifera: Panam. Pleurodonte (Labyrinthus) goldmani: Pirr Range, Darin, Panam. Victoria, Boca del Pepe, Rio Baudo; Nucu; mountains near mouth of Rio Atrato (Solem 1966). PANAM, Prov. Darin: Pirri Range; Paya; Rio Tucut. Prov. Panam: Cerro Campana; Sierra Chucanti 1800 ft. alt. (Solem 1966). Labyrinthus uncigerus uncigerus (Petit 1838)Caracolla uncigera Petit 1838; Guerins Mag. de Zool.:113. Helix (Labyrinthus) uncigera (Petit). Pilsbry 1889; Man. Conch. Pleurodonte (Labyrinthus) uncigera (Petit). Pilsbry 1910; Proc. Labyrinthus uncigerus (Petit). Von Martens 1892; Biol. Cent. Amer.:177. Labyrinthus unciger (Petit). Solem 1966; Fieldiana: Zoology 50:95 Helix uncigera var. conoidea Ancey 1890; Bull. Soc. Malac. France 7:152. Helix uncigera anopla Ancey 1890; Bull. Soc. Malac. France 7:152. Pleurodonte (Labyrinthus) tenaculum Dall 1909; Smiths. Misc. Type Localities. Caracolla uncigera: Panam. Helix uncigera var. conoidea: Colombia. Helix uncigera var. anopla: Colombia. Pleurodone (Labyrinthus) tenaculum : Mountains near mouth of Rio Atrato, Sierra Darin, Panama. Distribution.COLOMBIA, Dept. Choco; Arqua, 1.5 hrs. walk from Unguia; Acandi; Golfo de Urab; mountains near mouth of Rio Atrato (Solem 1966). PANAM, Canal Zone: Rio Pequen; Salamanca Hydrographic Station, Rio Pequen. Prov. Darin: Rio Puerco. Prov. Panam: mountains around Gaspasalama, upper Rio Mamoni; half-way from Cerro Azul to Mandingo; Cerro Campana. Comarco de San Blas: Puerto Olvaldia (Solem 1966). Labyrinthus uncigerus chiriquiensis (Pilsbry 1910)Pleurodonte (Labyrinthus) uncingera chiriquiensis Pilsbry 1910; Pleurodonte (Labyrinthus) chiriquiensis Pilsbry. Pilsbry 1926; Proc. Labyrinthus unciger form chiriquiensis (Pilsbry 1910). Solem 1966; Labyrinthus unciger form chiriquiensis (Pilsbry).Thompson 2008:747.Type Locality.Chiriqu Lagoon, Prov. Chiriqu, Panam. Lectotype ANSP 1612a (Baker 1963:244). Distribution.PANAM, Canal Zone: near Madden Lake. Prov. Chiriqu: Chiriqu Lagoon. Prov. Cocl: El Valle; El Valle, 2400 ft. alt.; trail to Las Minas; Club Campestre, El Valle. Prov. Panam: Cerro Campana (Solem 1966). Labyrinthus uncigerus tau (Pilsbry 1926)Pleurodonte (Labyrinthus) uncigera tau Pilsbry 1926; Proc. Acad. Labyrinthus unciger form tau (Pilsbry). Solem 1966; Fieldiana, Labyrinthus uncigera tau (Pilsbry). Thompson 2008:748.Type Locality.Archipilago de Bocas del Toro, Prov. Bocas del Toro, Panam. Distribution.COLOMBIA, Dept. Choco: Acandi Cocl: El Valle. Prov. Bocas del Torro: Archipilago de Bocas del Toro; Mono Creek, Almirante (Solem 1966). Genus Zachrysia Pilsbry 1894Zachrysia Pilsbry 1894; Man. Conch. Ser. 2, 9:97Type Species.Helix auricoma Frussac 1821. Distribution.Cuba. Taxonomy.Four subgenera are recognized. One has been introduced into the study area. Subgenus Zachrysia Pilsbry 1894 Taxonomy.Several species are native to Cuba. One

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226 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) species, with two subspecies, has been introduced into the study area. Zachrysia auricoma auricoma (Frussac 1821)Helix (Helicogena ) auricoma Frussac 1821. Helix (Thelidomus) auricoma Frussac. Pilsbry 1889; Man. Conch. Pleurodonte (Zachrysia) auricoma (Frussac). Pilsbry 1894; Man. Conch. 9:96. Zachrysia auricoma (Frussac). Bequaert & Clench 1936; Pub. Carnegie Inst. Wash. (457):64.Type Locality.Cuba. Distribution.PANAM, Prov. Panam: Cd. Panam (Bequaert & Clench 1936). (Pilsbry 1894)Pleurodonte (Zachrysia) Pilsbry 1894; Man. Pilsbry. Pilsbry 1926; Proc. Clench 1936; Pub. Carnegie Inst. Wash. (457):64.Thompson 2008:749.Type Locality.Havana, Cuba. Lectotype ANSP 77459a (Pilsbry 1926:76; H. B. Baker 1963:245). Distribution.A Cuban species that has been widely introduced to other tropical regions. PANAM, Prov. Panam: Cathedral Plaza, Cd. Panam (Pilsbry 1926). YUCATN: Quinta; Merida (Bequaert & Clench 1936). Family SOLAROPSIDAE Nordsieck 1986 Genus Psadara K. Miller 1878Psadara Miller 1878; Malac. Bltt. 25:162.Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:112.Type Species.Helix selenostoma Pfeiffer; 1852. Distribution.Southern Brazil, Bolivia, and Peru north to Guayana and Colombia; Costa Rica. Taxonomy.Many species. One occurs in the study area. Psadara tiloriensis (Angas 1879)Helix (Solaropsis) tiloriensis Angas 1879; Proc. Zool. Soc. Amer.:159.Von Martens 1901; Biol. Cent. Amer.:626. Psadara tiloriensis (Angas). Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:11213.Thompson 2008:750.Type Locality.Between the rivers Tilorio and Zhorquin Viejo the Rio Sarapiqu (Von Martens 1901). Family HUMBOLDTIANIDAE Pilsbry 1939 Distribution.Southern Texas south to Puebla and Veracruz. Taxonomy.Two genera are recognized. Genus Bunnya H. B. Baker 1942Bunnya H. B. Baker 1942; Nautilus 56:37.Miller 1987; Veliger 29:308.Type Species.Bunnya bernadinae H. B. Baker 1942. Distribution.Western Mxico: States of Durango, Jalisco, Mxico, Nayarit, and Sinaloa. Taxonomy.Two species are recognized. Bunnya bernadinae H. B. Baker 1942Bunnya bernadinae H. B. Baker 1942; Nautilus 56:37; pl. 5, Thompson 2008:750.Type Locality.Walls of old monastery, Desierto de Distribution.JALISCO: in addition to the type locality it is known from Temexcaltgepec. Bunnya naranjoae Miller 1987Bunnya naranjoae Type Locality.Along road from El Chante to Guizr, 1.5 km south of Rancho Manantln, (or 16 km south of El Distribution.Western slopes of the Sierra Madre Occidental from the vicinity of Mazatln, Sinaloa in the north to the Sierra de Manantln in southwestern Jalisco. DURANGO: ravine along the Mazatln-Durango highway 3 km E of Santa Lucia (15 km west of Loberas summit). JALISCO: type locality. NAYARIT: ravine along the TepcPuerto Vallarta highway at km 42 marker from Tepc; in riparian valley of Rio Manantlan from the type locality to 1.1 km upstream in the vicinity of the abandoned sawmill at Rincon de Manantlan. Genus Humboldtiana Von Ihering 1892Humboldtiana Ihering 1892; Zoologie 54:472.Pilsbry 1927; Proc. Acad. Nat. Sci. Phila. 79:165.Pilsbry 1939; Land Moll. N. Amer. I:395.Burch & Thompson 1957; Occ. Pap. Mus. Zool., 590:1; map 1.Solem 1974; Veliger 16:359.Nordsieck 1987; Arch. fr Mollusk. 118:23.Thompson & Brewer 2000; Bull. Fla. Mus. Nat. Hist. 43:50.Type Species.Helix humboldtiana Pfeiffer 1847. Distribution.Southern Texas south to the states of Mxico and Veracruz. Taxonomy.Six subgenera are recognized. The genus contains 54 species of which 43 have been described from Mxico. Subgenus Aglotrochus Thompson 2006Aglotrochus Thompson 2006; Bull. Fla. Mus. Nat. Hist. 46:92.Type Species.Humboldtiana tanymastix Thompson 2006. Distribution.Chihuahua, Mxico. Taxonomy.This subgenus contains a single known species. Humboldtiana (Aglotrochus) tanymastix Thompson 2006Humboldtiana tanymastix Thompson 2006; Bull. Fla. Mus. Nat.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 227 anatomy). Humboldtiana (Aglotrochus) tanymastix Thompson. Thompson 2008:751.Type Locality.Sierra Santo Domingo, 4 km westDistribution. Known only from the type locality. Subgenus Clydonacme Thompson 2006Clydonacme Thompson 2006; Bull. Fla. Mus. Nat. Hist. 46:75.Type Species.Humboldtiana regula Thompson 2006. Distribution.Chihuahua, Mxico. Taxonomy.This subgenus contains six species. Humboldtiana (Clydonacme ) hogeana (von Martens 1892)Helix humboldtiana hgeana von Martens 1892; Biol. Centr. Humboldtiana hgeana (von Martens).Pilsbry 1927; Proc. Acad. 9 (shell).Pilsbry 1935; Proc. Acad. Nat. Sci. Phila. 87:2; pl. 1, 1948; Proc. Acad. Nat. Sci. Phila. 100:202. Humboldtiana hogeana (von Martens). Solem 1954; Nautilus 68:5.Solem 1974; Veliger 16:363.Thompson 1967; Nautilus 81:25.Thompson 2006; Bull. Fla. Mus. Nat. Hist. 46:90; Humboldtiana (Clydonacme ) hogeana (von Martens). Thompson 2008:752.Type Locality.Chihuahua. Lectotype BMNH 1901 6 (Thompson 2006). Distribution.CHIHUAHUA. Von Martens (1892) described this species from Chihuahua. It is not clear whether the type locality is the city or the state. Whether the species actually occurred in Cd. Chihuahua can no longer be determined. The city has expanded since the Eighteenth Century from being a small pueblo of less than 1,500 people then to a vast metropolis of 840,000 people now, and nothing remains there that resembles a natural habitat that could harbor a Humboldtiana. Humboldtiana (Clydonacme ) oberon Thompson 2006Humboldtiana oberon Thompson 2006; Bull. Fla. Mus. Nat. Hist. anatomy). Humboldtiana (Clydonacme ) oberon Thompson. Thompson 2008:753.Type Locality.Sierra Victorino, 17 km west of Nueva 1900 m alt. Holotype UF 317164. Distribution.Known only from the type locality. Humboldtiana (Clydonacme ) princeps Thompson 2006Humboldtiana princeps Thompson 2006; Bull. Fla. Mus. Nat. anatomy). Humboldtiana (Clydonacme) princeps Thompson. Thompson 2008:753.Type Locality. km southwest of Cd. Chihuahua, Holotype UF 350039. Distribution.Known only from the type locality. Humboldtiana (Clydonacme ) regula Thompson 2006Humboldtiana regula Thompson 2006; Bull. Fla. Mus. Nat. Hist. system). Humboldtiana (Clydonacme ) regula Thompson. Thompson 2008:753.Type Locality.Sierra de la Catarina, highway pass ca. 12 km northeast of Ignacio Zaragoza, Chihuahua, Mxico Distribution.CHIHUAHUA: known only from the immediate vicinity of the type locality:21.6 km NE of Ignacio Humboldtiana (Clydonacme ) spectabile Thompson 2006Humboldtiana spectabile Thompson 2006; Bull. Fla. Mus. Nat. Hist. Humboldtiana (Clydonacme ) spectabile Thompson. Thompson 2008:754.Type Locality.Rocky outcrop in the foothills of the Sierra La Catarina, Buenaventura, Chihuahua, Mxico Distribution.CHIHUAHUA: known from the type locality in the foothills of the Sierra La Catarina, which lies about 20 km southwest of Buenaventura. The range is oriented north-south, and is about 60 km long and about 10 km wide near its middle. Humboldtiana (Clydonacme ) titania Thompson 2006Humboldtiana titania Thompson 2006; Bull. Fla. Mus. Nat. Hist. anatomy). Humboldtiana (Clydonacme ) titania Thompson. Thompson 2008:754.Type Locality. Sierra Victorino, 0.6 km southeast of Cumbres Majalca Visitors Center, Chihuahua, Mxico 2060 m alt. Holotype UF 358679. Distribution.Known only from the type locality. Subgenus Gymnopallax Thompson 2006Gymnopallax Thompson 2006; Bull. Fla. Mus. Nat. Hist. 46:72.Type Species.Humboldtiana cicatricosa Thompson 2006. Distribution.Chihuahua, Mxico. Taxonomy.Three species are recognized. Humboldtiana (Gymnopallax) cicatricosa Thompson 2006Humboldtiana cicatricosa Thompson 2006; Bull. Fla. Mus. Nat.

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228 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Humboldtiana (Gymnopallax) cicatricosa Thompson. Thompson 2008:755.Type Locality. 4 km north-northeast of Rancho Blanco, ca. 31 km NNE of San Juanito, along highway from W); 2180 m alt. Holotype UF 317169; Distribution.Known only from the type locality. Humboldtiana (Gymnopallax) ootamorum Meja, Naranjo-Garcia & Polaco 2009Humboldtiana ootamorum Megia, Naranjo-Garcia & Polaco 2009; anatomy).Type Locality.Mesa el Campanero (= Mesa de Enmedio), Barrana El Salto (West side of Mesa), Sonora, CNMO 1188. Distribution.SONORA: Known only from the immediate vicinity of the type locality. Humboldtiana (Gymnopallax) Thompson & Mejia 2006. Thompson & Mejia 2006; Nautilus 120:28 Humboldtiana (Gymnopallax ) Thompson & Mejia. Thompson 2008:755.Type Locality.Corareachi, 4.4 km north, 0.4 km west W); 2000 m alt. Holotype UF 353714. Distribution.Known only from the type locality. Subgenus Humboldtiana von Ihering 1892 Distribution.Southern Texas south to the states of Mxico and Veracruz. Taxonomy.The subgenus contains 38 species and subspecies, with 31 species in Mxico. Humboldtiana (Humboldtiana ) balanites Thompson 2006Humboldtiana balanites Thompson 2006; Bull. Fla. Mus. Nat. Humboldtiana (Humboldtiana) balanites Thompson. Thompson 2008:756.Type Locality..3 km north of San Ignacia Arareco, ca. 5 km south of Creel, Chihuahua, Mxico; 2280 m alt. Distribution.CHIHUAHUA: known from the type locality and a near-by locality: 6.6 km E of San Ignacio Humboldtiana (Humboldtiana ) bicincta Thompson & Brewer 2000Humboldtiana bicincta Thompson & Brewer 2000; Bull. Fla. Mus. anatomy). Humboldtiana (Humboldtiana) bicincta Thompson & Brewer. Thompson 2008:756. km south by road from Las Crucitas, Zacatecas, Mxico 2500 Distribution.Known only from the type locality. Humboldtiana (Humboldtiana ) buffoniana (Pfeiffer 1845)Helix buffoniana Pfeiffer 1845; Zeit. fr Malak. 2:152; Syst. Conch. in Philippi 1847; Helix humboldtiana var. buffoniana Pfeiffer.Fischer & Crosse Biol. Cent. Amer.:148. Helix (Lysinoe) humboldtiana Pfeiffer.von Martens 1892; Biol. Cent. Amer.:147. Helix matronula Uhde, in von Martens 1892; Biol. Cent. Amer.:148. Humboldtiana buffoniana (Pfeiffer).Pilsbry 1927; Proc. Acad. Humboldtiana (Humboldtiana) buffoniana (Pfeiffer). Thompson 2008:757.Type Locality.Rio Frio, northern slope of Volcn Ixtaccihuatl, on the road to Puebla, Mxico; 2600 m alt. Distribution.MXICO, DISTRITO FEDERAL: at high altitudes in mountains forming the western border of the Distrito Federal and along the southeastern border of the State (near and above La Venta and Guajimalpa or Coaximalpn, about 22 km SW of Mxico City, ca. 3,000 m alt.; Santa Rosa, 2600 m alt. Humboldtiana (Humboldtiana ) chrysogona Pilsbry 1948Humboldtiana chrysogona Pilsbry 1948; Proc. Nat. Sci. Phila. Humboldtiana (Humboldtiana) chrysogona Pilsbry. Thompson 2008:757.Type Locality.Low limestone bluff at lower border Mxico; 8,500 ft. alt. Holotype ANSP 164055. Distribution.ZACATECAS: known only from the immediate vicinity of the type locality. Humboldtiana (Humboldtiana ) corruga Thompson & Mejia 2006Humboldtiana corruga Thompson & Mejia 2006; Nautilus 120:25 Humboldtiana (Humboldtiana) corruga Thompson & Mejia. Thompson 2008:757.Type Locality..8 km south and 0.3 km west of Holotype UF 358872. Distribution.Known only from the type locality. Humboldtiana (Humboldtiana ) durangoensis Solem 1954Humboldtiana durangoensis Solem 1954; Nautilus 68:4, pl. 1,

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 229Humboldtiana (Humboldtiana) durangoensis Solem. Thompson 2008:757.Type Locality.Pine woods at 8000 ft., Laguna del Progresso, 30 miles north northwest of Los Coyotes and 25 miles north northwest of El Salto, Durango, Mxico. Holotype UMMZ 169746. Distribution.DURANGO: Sierra Madre Occidental from the drainage basin of the Rio Mezquital near Durango City, N to the Rio Mayo in SW Chihuahua, and eastern Sinaloa (Solem 1955). CHIHUAHUA: no locality given. DURANGO: 2.5 mi. W of San Lus; 8000 ft. alt.; El Boneto, W). SINALOA:13.4 mi. NE of Santa Lucia, 7100 ft. alt. (UF 130599). Humboldtiana (Humboldtiana ) edesma Thompson & Brewer 2000Humboldtiana edesma Thompson & Brewer 2000; Bull. Fla. Mus. 41 (reproductive anatomy). Humboldtiana (Humboldtiana) edesma Thompson & Brewer. Thompson 2008:758.Type Locality.Sierra San Francisco de Desmontes, east slope of Cerro La Mota, Cueva Ahumada 5 km southwest of Distribution.NUEVO LEN: known from the Sierra San Francisco de Desmontes. The sierra is a low, dry limestone border east of Saltillo. The type locality is at the northeast end of the range. A population from the southwest end of the range is referred to this species. COAHUILA: Cuesta de Humboldtiana (Humboldtiana ) eulaliae Metcalf 1983Humboldtiana eulaliae 1 (shell).Thompson 2006; Bull. Fla. Mus. Nat. Hist. 46:97. Humboldtiana (Humboldtiana) eulaliae Metcalf. Thompson 2008:758.Type Locality.A canyon on the west side of the Sierra Santa Eulalia 1.3 km east of El Pinolero and 7.5 km north and 1.5 km east of Penoles, Chihuahua, Mxico; 1250 m alt. Distribution.Known only from the type locality. Humboldtiana (Humboldtiana ) fasciata Burch & Thompson 1957Humboldtiana fasciata Burch & Thompson 1957; Occ. Pap. Mus. Humboldtiana (Humboldtiana) fasciata Burch & Thompson. Thompson 2008:759.Type Locality.Minerl del Monte, Hidalgo, Mxico; 9,300 ft. alt. Holotype UMMZ 191473. Distribution.Known only from the type locality. Humboldtiana (Humboldtiana ) fortis Pilsbry 1940Humboldtiana fortis Pilsbry 1940; Nautilus 53:140.Pilsbry 1948; Nautilus 54:47. Humboldtiana (Humboldtiana) fortis Pilsbry. Thompson 2008:759.Type Locality.Cerro Potos, northwest of Galeana, ft. alt. Holotype ANSP 164045. Distribution.Known only from the type locality. Humboldtiana (Humboldtiana ) globosa Burch & Thompson 1957Humboltiana globosa Burch & Thompson 1957; Occ. Pap. Mus. Humboldtiana (Humboldtiana) globosa Burch & Thompson. Thompson 2008:759.Type Locality.Three miles east of Perote, La Molina, Veracruz, Mxico; 8,000 ft. alt. Holotype UMMZ 191476. Distribution.Known only from the type locality. Humboldtiana (Humboldtiana ) gradyi Thompson & Brewer 2000Humboldtiana gradyi Thompson & Brewer 2000; Bull. Fla. Mus. anatomy). Humboldtiana (Humboldtiana) gradyi Thompson & Brewer. Thompson 2008:760.Type Locality.Area near the Dos de Diciembre Microwave Tower, 2 km southeast, 5 km east of Dos de Distribution.DURANGO; known only from the vicinity of the type locality. Humboldtiana ( Humboldtiana) humboldtiana (Pfeiffer 1847)Helix humboldtiana Valenciennes Pfeiffer 1847, Symb. Hist. in Helix badiocincta Weigmann, in Pfeiffer 1845; Symbolae Hist. 2:37. (A nomum nudum, but described as equivalent to H. humboldtiana.) Helix (Pomatia) humboldtiana (Pfeiffer).Fischer & Crosse 1872; Helix (Lysinoe) humboldtiana var. hegewishi von Martens 1892; Biol. Cent. Amer.:147. Humbolditana humboldtiana (Pfeiffer). Pilsbry 1927; Proc. Acad. (shell). Humboldtiana (Humboldtiana) humboldtiana (Pfeiffer). Thompson 2008:760. Pilsbry (1927:174) expressed doubt about this locality. Distribution.Unknown.

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230 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Humboldtiana (Humboldtiana ) inferior Pilsbry 1948Humboldtiana montezuma inferior Pilsbry 1948; Proc. Acad. (genitalia). Humboldtiana inferior Pilsbry. Thompson 2006; Nautilus 120:23.Thompson 2008:760.Type Locality.About 3 km from Hacienda Encinal, near Pablillo, on the trail to Alamar, Municipio de Galeana, Distribution.Known only from the type locality. Humboldtiana (Humboldtiana ) Thompson 2006 Humboldtiana (Humboldtiana) Thompson. Thompson 2008:761.Type Locality.Highway 68 at km post 116, Las Noria, Anteojitos; 1800 m alt. Holotype UF 130588. Distribution.Known only from the type locality. Humboldtiana (Humboldtiana ) latizona Thompson & Brewer 2000Humboldtiana latizona Thompson and Brewer 2000; Bull. Fla. Humboldtiana (Humboldtiana) latizona Thompson & Brewer. Thompson 2008:761.Type Locality.Sierra San Lorenzo, Mesitas Coloradas, 7.5 km east of Pedricea 1850 m alt., Durango, Mxico Distribution.Known only from the type locality. Humboldtiana (Humboldtiana ) malenae Metcalf 1983Humboildtiana malenae 1. Humboldtiana (Humboldtiana) malenae Metcalf. Thompson 2008:761.Type Locality.Boundary area between Municipios Ocampo and Muzquiz, on Coahuila State Highway 53, 6.2 km slightly north of east from Tres Caminos (village) above head of north branch of Caada La Virgen, Chihuahua, USNM 784768. Distribution.Known only from the type locality. Humboldtiana (Humboldtiana ) montezuma Pilsbry 1940Humboldtiana montezuma Pilsbry 1940; Nautilus 53:140.Pilsbry Humboldtiana (Humboldtiana) montezuma Pilsbry. Thompson 2008:762. 10,000 ft. alt. Holotype ANSP 164062. Distribution.Known only from the type locality. Humboldtiana (Humboldtiana ) Pilsbry 1927 Pilsbry 1927; Proc. Acad. 2 (shell). .Pilsbry 1948; Proc. Acad. Nat. Sci. (genitalia). Lysinoe humboldtiana buffoniana (Pfeiffer).Pilsbry 1903; Proc. Acad. Nat. Sci. Phila. 55:762. Humboldtiana (Humboldtiana) Pilsbry. Thompson 2008:762. Mxico. Holotype ANSP 85596. Distribution.COAHUILA: San Lorenza Canyon, SE of Saltillo, 7200 ft. alt.; Carneros Pass, 8,300 ft. alt.; Sierra Guadelupe, La Cuchilla. NUEVO LEN: mountains around Monterrey; Alamar Canyon; 2 km from Alamar on trail to Pablillo. SAN LUS POTOS: Catorce; Santana, 9700 ft. alt. Humboldtiana (Humboldtiana ) pergranulosa Solem 1955Humboldtiana pergranulosa Solem 1955; Nautilus 69:42; pl. 3, Humboldtiana (Humboldtiana) pergranulosa Solem. Thmpson 2008:762.Type Locality.San Jos Range of mountains in Restricted to near Jaralito (Solem 1974). Holotype ANSP 194820. Distribution.Known only from the type locality. Humboldtiana (Humboldtiana ) pilsbryi Solem 1954Humboldtiana pilsbryi 4 (shell). Humboldtiana (Humboldtiana) pilsbryi Solem. Thompson 2008:763.Type Locality.A humid pine-oak-madroo-yucca forest, two miles by road west of Ojitos Mine, about 4 miles west of Chihue, a few miles east of China [? Chinas, 23.8667 N, 99.4667 W], northwest of Ciudad Victoria, Tamaulipas, Mxico; 8600 ft. Holotype UMMZ 181280. Distribution.Known only from the type locality. Humboldtiana (Humboldtiana ) pinicola Thompson & Brewer 2000Humboldtiana pinicola Thompson & Brewer 2000; Bull. Fla. (reproductive anatomy). Humboldtiana (Humboldtiana) pinicola Thompson & Brewer. Thompson 2008:763.Type Locality.Southeast side of Cerro Puerto El Pino, limestone outcrop in an oak-pine forest ca. 6 km by road southwest of Pinal de Amoles, Quertaro, Mxico; 2650 m Distribution.Known only from the type locality. Humboldtiana (Humboldtiana ) potosiana Pilsbry 1927Pomatia humboldtiana Valenciennes. W. G. Binney 1879; Bull.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 231(radula). Humboldtiana potosiana Pilsbry 1927; Proc. Acad. Nat. Sci. Phila. (shell). Humboldtiana (Humboldtiana) potosiana Pilsbry. Thompson 2008:673.Type Locality.Sierra de San Miguelito, south of Cd. San Lus Potos, San Lus Potos, Mxico. Holotype ANSP 31836. Distribution.Known only from the type locality. Humboldtiana (Humboldtiana ) queretaroana (Dall 1897)Helix (Lysinoe) queretaroana Dall 1897; Nautilus 11:73. Humboldtiana queretaroana (Dall). Pilsbry 1927; Proc. Acad. Nat. (shell). Humboldtiana (Humboldtiana) queretaroana (Dall). Thompson 2008:764.Type Locality.Pinal de Amoles, Quertaro, Mxico; 8,000,000 ft. Holotype USNM 134691. Distribution.QUERTARO: known only from the immediate vicinity of the type locality. Humboldtiana (Humboldtiana ) riskindi Fullington & Zimmerman 1977Humboldtiana riskindi Fullington & Zimmerman 1977; Veliger Humboldtiana (Humboldtiana) riskindi Fullington & Zimmerman. Thompson 2008:764.Type Locality.Sierra de La Gloria, Caon Obscuro Chiquillo, 17 km east of Castaos, Coahuila, Mxico; 1300 m Natural History 5357. Distribution.Known only from the type locality. Humboldtiana (Humboldtiana ) Meja, Naranjo-Garcia & Polaco 2009Humboldtiana (Humboldtiana) Meja, Naranjo13 (reproductive anatomy).Type Locality..1 km south and 6.8 km wst of W), 2290 m alt. Holotype: Coleccion Malacologica de la Subdireccion de Laboratorios y Apoyo Academico del INAH, Mxico (DP) 691. Distribution.Known only from the type locality. Humboldtiana (Humboldtiana ) striata Burch & Thompson 1957Humboldtiana striata Burch & Thompson 1957; Occ. Pap. Mus. Humboldtiana (Humboldtiana) striata Burch & Thompson. Thompson 2008:764.Type Locality.Paso Cortez, Distrito Federal, Mxico; 10,500 ft. alt. Holotype UMMZ 192247. Distribution.Known only from the type locality. Humboldtiana (Humboldtiana ) taylori Drake 1951Humboldtiana taylori Drake 1951; Rev. Soc. Malacol:93; pl. Humboldtiana (Humboldtiana) taylori Drake. Thompson 2008:765.Type Locality.From the surface of cultural deposits of archeological site C96, Caon Media Luna, Fronteriza Range (Serranias del Burro), 16 miles south of Boquillas, northern Coahuila, Mxico. Corrected by Metcalf and Riskind (1979) Holotype USNM 596939. Distribution.COAHUILA: known only from the vicinity of the type locality: Sierra Madera del Carmen, Municipio de Villa Ocampo, ca. 2 km NNW of Loomis Humboldtiana (Humboldtiana ) tescola Thompson 1967Humboldtiana tescola Thompson 1967; Nautilus 81:22; text Humboldtiana (Humboldtiana) tescola Thompson. Thompson 2008:765.Type Locality.Knoll 15.7 miles southwest of San Tiburcio, Zacatecas, Mxico; 7600 ft. alt. Holotype UF 19752. Distribution.Known only from the type locality. Humboldtiana (Humboldtiana ) thompsoni Meja, Naranjo-Garcia & Polaco 2009Humboldtiana (Humboldtiana) thompsoni Meja, Naranjo-Garcia (reproductive anatomy).Type Locality.Canon de Carretas, 11.1 km north Malacologica de la Subdireccion de Laboratorios y Apoyo Academico del INAH, Mxico (DP) 692. Distribution.Known only from the type locality. Humboldtiana (Humboldtiana ) torrei Pilsbry 1935Humboldtiana torrei Pilsbry 1935; Proc. Acad. Nat. Sci. Phila. 87:1; Humboldtiana (Humboldtiana) torrei Pilsbry. Thompson 2008:765.Type Locality.San Antonio y Santa Rosalia [now Chihuahua, Chihuahua, Mxico. Holotype ANSP 162325. Distribution.Known only from the type locality. Subgenus Oreades Thompson & Brewer 2000Oreades Thompson & Brewer 2000; Bull. Fla. Mus. Nat. Hist. 43:66.Type Species.Humboldtiana porterae Thompson & Brewer 2000.

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232 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Taxonomy.A single species is recognized. Humboldtiana ( Oreades) porterae Thompson & Brewer 2000Humboldtiana porterae Thompson & Brewer 2000; Bull. Fla. Mus. anatomy). Humboldtiana (Oreades) porterae Thompson & Brewer. Thompson 2008:766.Type Locality.Caon de Garcia, 7 km northeast of Distribution.Known only from the type locality. Subgenus Polyomphala Thompson & Brewer 2000Polyomphala Thompson & Brewer 2000; Bull. Fla. Mus. Nat. Hist. 43:71.Type Species.Humboldtiana oreina Thompson & Brewer 2000. Distribution.Sierra La Encantada, northwestern Coahuila, Mxico. Taxonomy.Two species are recognized. Humboldtiana (Polyomphala ) oreina Thompson & Brewer 2000Humboldtiana oreina Thompson & Brewer 2000; Bull. Fla. Mus. (reproductive anatomy). Humboldtiana (Polyomphala) oreina Thompson & Brewer. Thompson 2008:767.Type Locality.Sierra La Encantada, La Ventana, Caon de Boquilla, 27 km WNW of Hacienda La Babia, 91 km NW of Melchor Musquz, Coahuila, Mxico; 1150 m alt. Distribution.Known only from the type locality. Humboldtiana ( Polyomphala) plana Metcalf & Riskind 1976Humboldtiana plana Metcalf & Riskind 1976; Nautilus 90:99; Humboldtiana (Polyomphala ) plana Metcalf & Riskind. Thompson 65 (shell). Thompson 2008:767.Type Locality.Sierra de Santa Rosa near the summit exposure in a massive limestone cliff, Mcpo. de Muzquz, Coahuila, Mxico. Holotype Delaware Museum of Natural History 106681. Distribution.Known only from the type locality. Family HELMINTHOGLYPTIDAE Pilsbry 1939 Subfamily HELMINTHOGLYPTINAE Pilsbry 1939 Genus Cahuillus Roth 1996Cahuillus Roth 1996; Veliger 39:40.Type Species.Sonorella walcottiana Bartsch 1903. Distribution.Southern California and adjacent northwestern Mxico. Taxonomy.Three species are recognized. Two occur in Mxico. Cahuillus greggi (Miller 1967)Sonorella greggi Cahuillus greggi (Miller).Roth 1996; Veliger 39:40.Thompson 2008:768.Type Locality.Sierra Purica, Sonora, Mxico, in igneous rock outcroppings in northeastern-facing ravine, on south bank of large canyon which runs easterly from saddle between the two highest peaks at the south end of the range Distribution.Known only from the type locality. Cahuillus mexicanus (Pilsbry & Lowe 1934)Microarionta rowelli mexicana Pilsbry & Lowe 1934; Nautilus 48:67.Pilsbry 1939; Land Moll. N. Amer. I:230. Cahuillus mexicanus (Pilsbry & Lowe).Roth 1996:40.Thompson 2008:768.Type Locality.Sierra San Francisco, in a range of granite mountains just south of the trail to Punta Peasco, 12 miles south of Sonoyta, Sonora, Mxico. Holotype not cited. Distribution.Known only from the type locality. Genus Eremarionta Pilsbry 1913Eremarionta Pilsbry 1913; Proc. Acad. Nat. Sci. Phila.:382.Pilsbry 1939; Land Moll. N. Amer. I:226.Bequaert & Miller 1973; Moll. Arid SW:106.Roth 1996; Veliger 39:41.Type Species.Micrarionta desertorum Pilsbry & Ferriss 1908. Distribution.California and Arizona in low mountain ranges in the Mohave, Colorado and Yuma Deserts; northwestern Sonora and northeastern Baja California. Taxonomy.Numerous species and subspecies. Four species and one subspecies occur in Mxico. Eremarionta ellipsostoma (Pilsbry 1894)Epiphragmophora ellipsostoma Pilsbry 1894; Nautilus 8:81.Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:142. Eremarionta ellipsostoma (Pilsbry). Thompson:2008:769.Type Locality.San Juan del Norte, probably on the east coast of Lower California. The original label accompanying the holotype bears the inscription ...SE Nicaragua, which was later scratched over in pencil. Holotype ANSP 10745. Distribution.The geographic occurrence and the Eremarionta indioensis (Yates 1890)Helix (Arionta) carpenteri variety indioesis Yates 1890; Nautilus 4:63. Sonorella indioesis (Yates). Bartsch 1904; Smiths. Misc. Coll. Micrarionta indioensis (Yates). Berry 1922; Proc. Acad. Nat. Sci. Phila. 74:93.Willett 1930; Nautilus 43:115.Pilsbry 1939;

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 233Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. Eremarionta indioensis (Yates). Thompson 2008:768.Type Locality.On the south side of the canyon among granite talus, Indio, Riverside County, California. Holotype ANSP 62145. Distribution.Riverside Co., California south to northwestern Mxico. BAJA CALIFORNIA NORTE: Caon de Guadalupe, Sierra Jurez; canyon ca. 16 km S of Caon de Guadalupe; E side of Sierra Jurez ca. 0.25 mi. above fork of Caon Carrizo. Eremarionta rowelli rowelli (Newcomb 1865)Helix rowelli Newcomb 1865; Proc. Calif. Acad. Sci. 3:181. Sonorella rowelli (Newcomb). Bartsch 1904; Smiths. Misc. Coll. 47:119 (in part). Micrarionta rowelli (Newcomb). Pilsbry & Ferriss 1923; Proc. Eremarionta rowelli rowelli (Newcomb). Bequaert & Miller 1973; Moll. Arid SW:106.Thompson 2008:769.Type Locality.Tinajas Atlas, in Tinajas Atlas Mountains, 26 miles south of Wellton and ca. 45 miles Newcomb Collection 27517, Cornell University. Distribution.Tinajas Atlas Mountains and the Sierra Pinta in Yuma County, Arizona and in adjacent Mxico. SONORA: Sierra del Tuseral just south of the International Border, 50 mi. SE of Wellton; Sierra Tinajas Atlas, along Hwy. 2, 8.9 mi. W of El Puerto. Eremarionta rowelli bechteli (Emerson & Jacobson 1964)Micrarionta (Eremarionta) rowelli bechteli Emerson & Jacobson 1964; Trans. San Diego Soc. Nat. Hist. 16: Trans. San Diego Eremarionta rowelli bechteli (Emerson & Jacobson 1964).Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47: 2008:770.Type Locality.In rock slide, Isla San Esteben, Sonora, Mxico. Holotype CAS 36377. Distribution.Known only from the type locality. Eremarionta ultima (Pilsbry 1916)Sonorella ultima (shell). Eremarionta ultima (Pilsbry). Thompson 2008:770.Type Locality.Sinaloa, Mxico. Holotype ANSP 58124. locality. Genus Greggelix Miller 1972Greggelix Miller 1972; Nautilus 85:128.Type Species.Helix indigena Mabille 1895. Distribution.Endemic to Baja California Sur. Taxonomy.Three species are recognized. Greggelix indigena (Mabille 1895)Helix indigena Mabille 1895; Bull. Soc. Philomath. Paris., 8th ser., 7:64. Helix digueti Mabille 1895; Bull. Soc. Philomath. Paris., 8th ser., 7:65. Greggelix indigena (Mabille).Miller 1972; Nautilus 85:128; Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:140; textSonorella loheii lioderma Pilsbry 1904; Nautilus 18:59.Pilsbry Type Localities.Above 800 m on peaks of the Sierra, throughout most of the central part of the Peninsula of California. Helix digueti Sonorella loheii lioderma: near Muleje [Muleg], Baja California Sur, Mxico; holotype ANSP 58107. Distribution.BAJA CALIFORNIA SUR: San Jos Comond; 19 km SW San Miguel Comond; Mesa de San Alejo, NW of San Javier, Sierra de La Gigante, 800 m alt.; Arroyo de La Purisima, 4.8 km E of San Isidro, along road to Caripol; near Muleg. Greggelix loehrii (Gabb 1867)Helix lhrii (shell). Sonorella lohrii (Gabb). Pilsbry 1900; Proc. Acad. Nat. Sci. Phila.:561.Bartsch 1904; Proc. U. S. Nat. Mus. 47:197; pl. 32, Sonorella loehrii (Gabb). Miller 1972; Nautilus 85:128. Greggelix loehrii (Gabb). Miller 1981; Proc. Biol. Soc. Wash. Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:140; Helix steganella Mabille 1895; Bull. Soc. Philomath. Paris., 8th ser., 7:64. Mabille 1895; Bull. Soc. Philomath. Paris., 8th ser., 7:65.Type Locality.Helix lhrii: higher table lands from near Moleje [Muleg], Baja California Sur, Mxico; holotype ANSP 58106. Helix steganella : above 800 m on peaks of the Sierra, throughout most of the central part of the Peninsula of California. : Lower California. Distribution.BAJA CALIFORNIA SUR: known only from the immediate vicinity of the type locality and San Javier. Greggelix punctata Miller 1981Greggelix punctata Miller 1981; Proc. Biol. Soc. Wash. 94:732; Thompson 2008:772.Type Locality.Gulf slope of Sierra de la Gigante, southwest of Muleje [Muleg], 2.9 miles south of El Potrero, along trail from Pie de La Cueta to Guajademi, in rock slide about 1.5 miles from Pie de La Cueta, Baja California Sur,

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234 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Mxico; ca. 2450 m alt. Holotype USNM 792140. Distribution.Known only from the type locality. Genus Helminthoglypta Ancey 1887Helminthoglypta Ancey 1887; Conchologists Exchange, 1:76.Pilsbry 1895; Man. Conch. 9:193 (in part).Pilsbry 1939; Land Moll. N. Amer. I:63.Roth 1996; Veliger 39:32.Type Species.Helix tudiculata Binney 1843. Distribution.Southwest Oregon south to Baja California Norte. Taxonomy.Three subgenera are recognized (Roth 1996). Two occur in Mxico. Subgenus Helminthoglypta Ancey 1887 Distribution.Southwest Oregon south to Baja California Norte. Taxonomy.Numerous species and subspecies. One occurs in Mxico. Helminthoglypta ( Helminthoglypta) tudiculata (Binney 1843)Helix tudiculata Binney 1843; Boston Jour. Nat. Hist. 4:360; pl. 20 (shell). Helminthoglypta tudiculata (Binney). Berry 1928; Jour. Ento. Zool. Pomona Coll. 20:79.Ingles 1935; Proc. Malac. Soc. Lond., & Roth 1990; Proc. Calif. Acad. Sci. 47:133.Thompson 2008:773. Helix tudiculata var. binneyi Hemphill 1890, in Binney; Terr. Moll, N. A., 3rd suppl.:219.Type Localities. Helix tudiculata : San Diego, San Helix tudiculata var. binneyi : mountains of San Diego County, California; holotype CAS 2481. Distribution.Los Angeles and San Bernardino Counties, California, and adjacent Mxico. BAJA CALIFORNIA NORTE: near Bajia de Todos Santos; Caada Macho Gero; Punta Banda; San Antonia Canyon, ca. 8 km N of Johnson Ranch; Johnson Ranch. Subgenus Charodotes Pilsbry 1939Charodotes Pilsbry 1939; Land Moll. N. Amer. I:170.Roth and Hochberg 1992; Veliger 35:339.Roth 1996; Veliger 39:32.Type Species.Helix traskii Newcomb 1861. Distribution.Southern California south to Baja California Norte. Taxonomy.About 16 species are recognized. Five species and three subspecies occur in Mxico. Helminthoglypta (Charodotes) coelata (Bartsch 1916)Epiphragmophora traskii coelata Bartsch 1916; Proc. U. S. Nat. Helminthoglypta traski coelata (Bartsch). Pilsbry 1939; Land Moll. Helminthoglypta (Charodotes) coelata (Bartsch). Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:133.Thompson 2008:774. Diego County, California. Holotype USNM 124747. Distribution.Along the coast of San Diego County California south to extreme northwestern Mxico. BAJA CALIFORNIA NORTE: 11 km S of Tijuana. Helminthoglypta ( Charodotes) hannai hannai Pilsbry 1927Helminthoglypta hannai Pilsbry 1927; Proc. Calif. Acad. Sci. Helminthoglypta (Charodotes) hannai hannai Pilsbry. Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:133.Thompson 2008:774.Type Locality.Pine Ridge, Isla Guadalupe, Baja California Norte, Mxico; 900 m alt. Holotype CAS 2561. Distribution.BAJA CALIFORNIA NORTE: known only from Isla Guadalupe. Helminthoglypte (Charodotes) hannai diodon Pilsbry 1927Helminthoglypte hannai diodon Pilsbry 1927; Proc. Calif. Acad. Helminthoglypte (Charodotes) hannai diodon Pilsbry. Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:133.Thompson 2008:774.Type Locality.Northeast Anchorage, Isla Guadalupe, Baja California Norte, Mxico. Holotype CAS 2562. Distribution.BAJA CALIFORNIA NORTE: known only from the immediate vicinity of the type locality. Helminthoglypta (Charodotes) misiona Chase 1937Helminthoglypta traski misiona Chase 1937; Nautilus 51:60; pl. 4, 47 (shell). Helminthoglypta (Charodotes) misiona Chase. Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:134.Thompson 2008:775.Type Locality.A rockslide near the San DiegoEnsanada highway about 40 miles [64 km] south of Tijuana, Baja California Norte, Mxico. Holotype in LACM. Distribution.Known only from the vicinity of the type locality. BAJA CALIFORNIA NORTE: rockslide on S side Tijuana-Ensanada highway; El Progresso, Sandoval Ranch area, ca. 48 km S of Tijuana. Helminthoglypta (Charodotes) reederi Miller 1981Helminthoglypta reederi Helminthoglypta (Charodotes) reederi Miller. Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:134; textType Locality.Sierra San Pedro Mrtr, in rock outcrops in a small canyon which crosses the road to the astronomical observatory, at a distance of about 2.5 km below the observatory housing area, Baja California Norte, Mxico; 2575 m alt. Holotype CAS 19732. Distribution.Known only from the type locality.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 235 Helminthoglypta (Charodotes) traski coronadoensis (Bartsch 1916)Epiphragmophora traskii coronadoensis Bartsch 1916; Proc. Calif. (shell). Helminthoglypta traski coronadoensis (Bartsch). Pilsbry 1939; Helminthoglypta (Charodotes) traskii coronadoensis (Bartsch). Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. Epiphragmophora traskii chrysoderma Berry 1920; Proc. Calif. Helminthoglypta (Charodotes) traski coronadoensis (Bartsch). Thompson 2008:776.Type Localities.Epiphragmophora traskii coronadoensis: Ilsa Los Coronados, Baja California Norte, Mxico; holotype in the USNM. Epiphragmophora traskii chrysoderma: Isla Coronado Sur, among loose talus on higher portion of south end; holotype in the CAS. Distribution.BAJA CALIFORNIA NORTE: known only from Isla Los Coronados, and Isla Coronado Sur. Genus Herpeteros Berry 1947Herpeteros Nautilus 85:134. Roth 1996; Veliger 39:33.Type Species.Micrarionta (Eremarionta) inglesiana Berry 1928. Distribution.Baja California Norte and Baja California Sur. Taxonomy.Five species are placed in the genus. Only two are known anatomically. The generic assignment of the other three is provisional. Herpeteros chacei (Willett 1940)Micrarionta (Eremarionta) chacei Willett 1940; Bull. Calif. Acad. Sci. 39:81; pl. 12. Sonorelix (Herpeteros) chacei (Willett). Smith, Miller, Christensen Herpeteros chacei (Willett). Thompson 2008:776.Type Locality.Lower end of El Tigre Canyon, about 9 miles [14.4 km] north of Ensanada, Baja California Norte, Mxico. Distribution.Known only from the type locality. (Pilsbry 1927) Pilsbry 1927; Proc. Calif. Acad. Sci. 4th 1973; Moll. Arid SW:110. Sonorelix (Herpeteros) (Pilsbry). Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47: Proc. (Pilsbry). Thompson 2008:777.Type Locality.Turtle Bay [Bajia Tortugas], Baja California Norte, Mxico. Holotype CAS 2618. Distribution.Known only from the type locality. Herpeteros inglesiana (Berry 1928)Micrarionta (Eremarionta) inglesiana Berry 1928; Jour. Ent. Zool. Sonorelix (Herpeteros) inglesiana Malacol., 1:11.Miller 1972; Nautilus 85:134.Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. Herpeteros inglesiana (Berry). Thompson 2008:777.Type Locality.On moist north slope of the Red Rock [Pea Colorado] under rocks, about 3 miles [4.8 km] from the sea and 0.5 miles [0.8 km] from Hamilton Ranch, west of Santo Domingo, Baja California Norte, Mxico. Holotype in the SBMNH. Distribution.BAJA CALIFORNIA NORTE: known only from the vicinity of the type locality. Herpeteros merrilli (Bartsch 1904)Sonorella merrilli Bartsch 1904; Smiths. Misc. Coll. 47:192; Sonorelix (Herpeteros) merrilli (Bartsch). Smith, Miller, Christensen Herpeteros merrilli (Bartsch). Thompson 2008:778.Type Locality.Below San Quintin, Baja California Norte, Mxico. Holotype USNM 125260. Distribution.Known only from the type locality. Herpeteros peninsularis (Pilsbry 1916)Sonorella peninsularis 4 (shell). Micrarionta peninsularis (Pilsbry). Hanna 1927; Proc. Calif. Acad. Sonorelix (Herpeteros) peninsularis (Pilsbry). Miller 1972; Nautilus Herpeteros peninsularis (Pilsbry). Thompson 2008:778.Type Locality.Trinidad, on the west coast near [60 km south of] San Borga [Borja], Baja California Norte, Mxico (Hanna 1927:503). Holotype ANSP 58127. Distribution.Widely distributed in the southern half of BAJA CALIFORNIA NORTE, and extreme northern BAJA CALIFORNIA SUR. Genus Martirelix Miller 1982Martirelix Miller 1982; Veliger 24:345.Roth 1996; Veliger 39:33.Type Species.Gregghelix babrakzaii Miller 1982. Distribution.Known only from Baja California Norte. Taxonomy.Two species are recognized. Martirelix babrakzaii (Miller 1982)Greggelix babrakzaii Thompson 2008:779.Type Locality.Caon Diablito, about 1 km up the canyon from its mouth, east slope of the Sierra San Pedro Martir, Baja California Norte, Mxico; 700 m alt. (31.1 N, Distribution.Known only from the type locality. Martirelix huertai (Miller & Roth 1990)Greggelix huertai Miller & Roth 1990; Proc. Calif. Acad. Sci.

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236 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Thompson 2008:779.Type Locality.Cerro de La Mina de San Jos, in rock piles on north side of entrance to the mine; Baja California Holotype SBMNH 35113. Distribution.Known only from the type locality. Genus Micrarionta Ancey 1880Micrarionta Ancey 1880; Le Naturaliste, 1:334.Pilsbry 1939; Land Moll. N. Amer. I:201.Pearce 1990; Malac. Rev. 23:1.Type Species.Helix facta Newcomb 1864. Distribution.Islands off southern California; Isla Guadalupe, Baja California Norte, Mxico. Taxonomy.Eleven species are recognized. One occurs in Mxico. Micrarionta guadalupiana (Dall 1898)Epiphragmophora (Micrarionata) guadelupiana Dall, in Pilsbry & (genitalia).Dall 1900; Proc. Acad. Nat. Sci. Phila.:101; Micrarionta guadalupiana (Dall). Pilsbry 1927; Proc. Calif. Acad. guadalupiana.) Smith, Miller, Christensen & Roth 1990; Proc. Calif. Acad. Sci. 47:134.Pearce 1990; Malac. Rev. 23:7.Thompson 2008:780.Type Locality.Isla Guadalupe, Baja California Norte, Mxico. Holotype USNM 129999. to Isla Guadalupe. Genus Xerarionta Pilsbry 1913Xerarionta Pilsbry 1913; Proc. Acad. Nat. Sci. Phila.:382.Pilsbry 1939; Land Moll. N. Amer. I:214.Miller 1981; Veliger 24:46.Type Species. Tryon 1866 (= Helix canescens Adams and Reeve 1848). Distribution.Southwestern California south to Baja California Sur, Mxico. Taxonomy.Two subgenera are recognized (Roth 1996:32). Subgenus Xerarionta Pilsbry 1913 Type Species. Newcomb. Distribution.Southwestern California south to Baja California Sur, Mxico. Taxonomy.Seven species and four subspecies. Three species and three subspecies occur in Mxico. Xerarionta (Xerarionta) areolata (Pfeiffer 1845)Helix areolata Sowerby Pfeiffer 1845; Zeitschr. fr Malak., 2:145.Fischer & Crosse 1872; Miss. Sci. Mex. I:262; pl. 11, Epiphragmophora (Micrarionta) areolata (Sowerby). Pilsbry 1898; Epiphragmophora areolata (Sowerby). Dall 1900; Proc. Acad. Nat. Sci. Phila.:100. Micrarionta areolata (Sowerby Pfeiffer). Pilsbry 1913; Proc. th Xerarionta areolata (Pfeiffer 1845). Smith, Miller, Christensen & Helix areolata var. albida Hemphill 1890; Cat. N. A. shells. (Nomum nudum) Helix areolata var. examinata Cooper 1892; Proc. Calif. Acad. Sci. Micrarionta areolata examinata Pilsbry 1913; Proc. Acad. Nat. Sci. Phila.:392. Micrarionta areolata var. arida Pilsbry 1913; Proc. Acad. Nat. Sci. Acad. Nat. Sci. Phila. 115:248. Micarionta areolata var. scammoni Pilsbry 1913; Proc. Acad. Nat. Acad. Nat. Sci. Phila. 114:249. Micrarionta aspersa var. aspersa Pilsbry 1913; Proc. Acad. Nat. Proc. Acad. Nat. Sci. Phila. 115:248. Xerarionta (Xerarionta) areolata (Pfeiffer). Thompson 2008:780.Type Localities. Helix areolata: California; restricted to Magdalena Bay, Baja California Sur (Pilsbry 1913:319). Helix areolata var. examinata : Isla Espritu Santo, Baja California Sur, Mxico. Micrarionta areolata var. arida: Magdalena Bay, Baja California Sur, Mxico; lectotype ANSP 76208a (H. B. Baker 1963:249). Micarionta areolata var. scammoni: Magdalena Bay, Baja California Sur, Mxico; lectotype ANSP 10302a (H. B. Baker 1963:249). Micrarionta aspersa var. aspersa: Magdalena Bay, Baja California Sur, Mxico; lectotype ANSP 10265a (H. B. Baker 1963:249). Distribution.BAJA CALIFORNIA SUR: Smith et al. (1990:136) list numerous localities for extant populations and the eastern regions. Xerarionta (Xerarionta) (Pfeiffer 1845) Pfeiffer 1845; Zeitschr. f. Malak., 2:152. Syst. Conch. Cab., Helix (Pfeiffer). Dall 1900; Proc. Acad. Nat. Sci. Phila.:100. (Pfeiffer). Pilsbry 1913; Proc. Acad. Nat. Sci. 1927; Proc. Calif. Acad. Sci. 4th ser., 16:179. (Pfeiffer). Smith, Miller, Christensen & Roth Xerarionta (Xerarionta) (Pfeiffer). Thompson 2008:782.Type Locality.California; restricted to Puerto San Bartolom, Baja California Norte (Pilsbry 1919:288). Distribution.BAJA CALIFORNIA NORTE; known Xerarionta (Xerarionta) (Adams & Reeve 1848)Helix canescens Adams & Reeve 1848; Voy. Samarang:62; pl. 16,

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 237 (Adams & Reeve). Pilsbry 1913; Micrarionta canescens (Adams & Reeve). Pilsbry 1927; Proc. Calif. (Adams & Reeve). Smith, Miller, 37 (map). Newcomb Tryon 1866; Amer. Jour. Conch. 2:316; Stearns 1867; Proc. Calif. Acad. Sci. 3:328. Epiphragmophora (Miscarionta) (Newcomb). Pilsbry & (genitalia). (Newc. Tryon). Pilsbry 1913; Proc. Acad. (Newc. Tryon). Pilsbry 1927; (shell). Helix areolata var. cedroensis Hemphill 1890; Cat. N. A. shells. (A nomem nudum.) Epiphragmophora leucanthea Dall 1900; Proc. Acad. Nat. Sci. Pilsbry 1913; Proc. Acad. Nat. Sci. Acad. Nat. Sci. Phila. 115:248. Xerarionta (Xerarionta) (Adams & Reeve). Thompson 2008:782.Type Localities. Helix canescens: Africa. Helix : Cerros Island, Baja California Norte, Mxico. Epiphragmophora leucanthea : east side of Cerros Island, Baja California Norte, Mxico; syntypes USNM 107627. : Isla Cedros, Baja California Norte, Mxico; lectotype ANSP 10304 (H. B. Baker 1963:248). Distribution.BAJA CALIFORNIA NORTE: numerous localities on Isla Cedros. BAJA CALIFORNIA SUR: Isla Navidad. Xerarionta (Xerarionta) (Dall 1900)Epiphragmophora crassula Dall 1900; Proc. Acad. Nat. Sci. (Dall). Pilsbry 1913; Proc. Acad. Nat. (Dall). Smith, Miller, Christensen & Roth Xerarionta (Xerarionta) (Dall). Thompson 2008:783.Type Locality.Isla Navidad, Baja California Sur, Mxico. Holotype in the USNM. Distribution.BAJA CALIFORNIA SUR: living colonies are known only from Isla Navidad; sub-fossil on Isla San Geronimo. Xerarionta (Xerarionta) pandorae (Forbes 1850)Helix pandorae Forbes 1850; Proc. Zool. Soc. Lond. 18:55; pl. 9, Epiphragmophora pandorae (Forbes). Dall 1900; Proc. Acad. Nat. Sci. Phila.:101. Micrarionta pandorae (Forbes). Pilsbry 1913; Proc. Acad. Nat. Sci. Xerarionta pandorae (Forbes). Smith, Miller, Christensen & Roth Helix demascensis Gould 1856; Proc. Boston Soc. Nat. Hist. 6:11.(shell). Epiphragmophora (Micrarionta) pandorae bonitosensis Pilsbry Xerarionta (Xerarionta) pandorae (Forbes), Thompson 2008:783.Type Locality.Helix pandorae: unknown. Helix demascensis: desert region east of California; lectotype Cornell Univ. Paleo. Coll. 27244. Epiphragmophora (Micrarionta) pandorae bonitosensis: Las Bonitos Island, off Lower California; holotype ANSP 66092a. Distribution.BAJA CALIFORNIA NORTE: Isla San Benito. Subgenus Plesarionta Pilsbry 1939Plesarionta Pilsbry 1939; Land Moll. N. Amer. I:212.Miller 1981; Veliger 24:46.Roth 1996; Veliger 39:32.Type Species.Helix stearnsiana Gabb 1867. Norte and southern California. Taxonomy.Two species are recognized. Xerarionta (Plesarionta) orcutti (Dall 1900)Epiphragmophora orcutti Dall 1900; Proc. Acad. Nat. Sci. Epiphragmophora traski tularensis Hemphill Pilsbry 1895; Nautilus 9:82. (A nomum nudum, not Arionta tudiculata tularensis Hemphill 1892.) Epiphragmophora traski tularica Bartsch 1916; Proc. U. S. Nat. Plesarionta orcutti (Dall). Roth 1982; Bull. South. Calif. Acad. Sci. Xerarionta (Plesarionta ) orcutti (Dall). Thompson 2008:784.Type Locality.Rosario Mesas, Baja California Norte, Mxico. Holotype in the USNM. Distribution.BAJA CALIFORNIA NORTE: mesa about 2.4 km N of El Rosario; El Rosario, 30 m alt. Xerarionta (Plesarionta) stearnsiana (Gabb 1868)Helix stearnsiana 310 (shell).Fischer & Crosse 1872; Miss. Sci. Mex. 1:248; pl. Epiphragmophora (Micrarionta) stearnsiana (Gabb). Pilsbry 1898; Micrarionta stearnsiana (Gabb). Hanna 1925; Proc. Calif. Acad. Sci. 14:247.Berry 1928; Jour. Entom. Zool. Pomona Coll. Acad. Sci. 16:178.Pilsbry 1939; Land Moll. N. Amer. I:212 Plesarionta stearnsiana (Gabb). Smith, Miller, Christensen & Roth Xerarionta (Plesarionta ) stearnsiana (Gabb). Thompson 2008:785.Type Locality.San Toms to a little beyond [El]

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238 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Rosario, Baja California Norte, Mxico. Lectotype ANSP 10831 (Pilsbry 1939). Distribution.San Diego Co., California south to Baja California Norte. BAJA CALIFORNIA NORTE: Numerous Miller, Christensen & Roth 1990); La Zonja, Sierra San Pedro Martr. Subfamily SONORELLINAE Pilsbry 1939 Genus Sonorella Pilsbry 1900Sonorella Pilsbry 1900; Proc. Acad. Nat. Sci. Phila.:556.Pilsbry 1939; Land Moll. N. Amer. I:267.Bequaert & Miller 1973:11012. Masculus Pilsbry 1939; Land Moll. N. Amer. I:377. Sonoranax Pilsbry 1939; Land Moll. N. Amer. I:391.Type Species. Sonorella : Epiphragmophora hachitana Dall 1895. Masculus: Pilsbry 1905. Sonoranax: Sonorella dalli Bartsch 1904). Distribution.Arid southwestern United States in Arizona, New Mexico and Texas south to Chihuahua and Sonora, Mxico. Taxonomy.Two subgenera are recognized, Sonorella s. s. and Sonoranax (Roth 1996:32). Only the former occurs in Mxico. Twenty species and one subspecies are reported from Mxico. Sonorella (Sonorella) burgessi Naranjo-Garcia 1988Sonorella burgessi Sonorella (Sonorella ) burgessi Naranjo-Garcia. Thompson 2008:786.Type Locality.Sierra el Viejo, at base of northeast facing limestone cliff on ridge southwest of canyon junction where road from west turns north, ca. 7 km east of El Plomito, SBMNH 34933. Distribution.Known only from the type locality. Sonorella (Sonorella) cananea Naranjo-Garcia 1988Sonorella cananea 1 (shell), 2 (genitalia). Sonorella (Sonorella) cananea Naranjo-Garcia. Thompson 2008:786.Type Locality.Sierra Elenita, in igneous rock slide along right bank of creek approximately 0.1 mi. (0.16 km) southeast of Highway 2, along arroyo El Quince, which crosses Highway 2 at 2.1 mi. (approximately 3.38 km) west of Puerto Cananea, a pass approximately 8 mi. (approximately W). Distribution.SONORA: known from the Sierra Elenita and two localities in the Sierra Mariquita. SONORA: Sierra W. Sonorella (Sonorella) goldmani Bartsch 1904Sonorella goldmani Bartsch 1904; Smiths. Misc. Coll. 47:192; pl. Sonorella (Sonorella ) goldmani Bartsch. Thompson 2008:787.Type Locality.Mountains near Lago de Santa Maria, Chihuahua, Mxico. Holotype: USNM 174933. Distribution.Known only from the type locality. Sonorella (Sonorella) hachitana hachitana (Dall 1895)Epiphragmophora hachitana Dall 1895; Proc. U. S. Nat. Mus. 18:2.Dall 1896; Bull. U. S. Nat. Mus. 19:338 (in part). Sonorella hachitana (Dall). Pilsbry 1901; Proc. Acad. Nat. Sci. Phila.:556.Bartsch 1904; Smiths. Misc. Coll. 47:190; pl. 31, Phila.:257 (in part).Pilsbry & Ferriss 1915; Proc. Acad. Nat. Pilsbry & Ferriss 1923; Proc. Acad. Nat. Sci. Phila.:65.Pilsbry (genitalia).Bequaert & Miller 1973; Moll. Arid SW:125. Sonorella (Sonorella) hachitana hachitana (Dall). Thompson 2008:787. Land & Gilbertson 2010; Proc. Biol. Soc. Wash. 123:67.Type Locality.Big Hatchet Peak, Big Hatchet Mountains, Hidalgo Co., New Mexico. Holotype USNM 130004. Distribution.Known only from the Big Hatchet Mountains in New Mexico. Bequaret and Miller (1973:125) locality. Presumably this is also in the Sierra Hacheta Grande. Other subspecies occur in New Mexico and Texas. Sonorella (Sonorella) madreana Naranjo-Garcia 1989Sonorella madreana Sonorella (Sonorella ) madreana Naranjo-Garcia. Thompson 2008:788.Type Locality.Sierra Las Minitas, ca. 2 km southeast of Rancho Jucaral buildings, Sonora, Mxico; 1400 m alt. Distribution.SONORA: known only from the immediate vicinity of the type locality. Sonorella (Sonorella) magdalenensis (Stearns 1890)Helix magdalenensis Stearns 1890; Proc. U. S. Nat. Mus. 13:207 Sonorella magdalenensis (Stearns). Bequaert & Miller 1973; Moll. Arid. SW:121. Sonorella tumamocensis Pilsbry & Ferriss 1915; Proc. Acad. Nat. 214 (shell). Sonorella sitiens arida Pilsbry & Ferriss 1915; Proc. Acad. Nat. Sci. Sonorella arida Pilsbry & Ferriss.Pilsbry 1939; Land Moll. (genitalia). Sonorella hinkleyi Pilsbry & Ferriss 1919; Nautilus 33:19.Pilsbry Sonorella hinkleyi fraterna Ferriss 1919; Nautilus 33:38 (A nomum nudum).

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 239Sonorella hinkleyi tumacacori Pilsbry & Ferriss 1919; Nautilus 33:19. Sonorella cayetanensis Pilsbry & Ferriss 1919; Nautilus 33:19. Sonorella hinkleyi cayetanensis Pilsbry & Ferriss. Pilsbry & Ferriss Sonorella linearis Pilsbry & Ferriss 1923; Proc. Acad. Nat. Sci. Sonorella tumamocensis linearis Pilsbry & Ferriss. Pilsbry 1939; Sonorella (Sonorella) magdalenensis (Stearns). Thompson 2008:788.Type Localities. Helix magdalenensisi: restricted to Sierra Magdalena, ca. 1 mi. north of Magdalena; 3650 ft. alt. (Bequaert & Miller 1973); holotype USNM. Sonorella tumamocensis: Tumamoc Hill at west city limits of Tucson, Arizona; 2750 ft. alt.; holotype ANSP 112245. Sonorella sitiens arida: south end of Cerro Colorado, ca. 2 mi. from Cerro Colorado Mine, Pima County, Arizona; holotype ANSP 112160. Sonorella hinkleyi : San Cayetana Mountains on southern most peak [Mt. Shibell], 2 mi. above Calabasa, Santa Cruz County, Arizona; holotype ANSP 43735. Sonorella cayetanensis: from near the top to rather low on the highest peak of the Cayetano Mountains, Santa Cruz County, Sonorella linearis: North end of Santa Rita Mountains on west side of saddle overlooking Helvetia, Pima County, Arizona; holotype ANSP 130996. SONORA: Sierra Pajaritos ca. 24 mi. by road E of Ures, 3000 ft. alt. Sonorella (Sonorella) mearnsi Bartsch 1904Sonorella mearnsi Bartsch 1904; Smiths. Misc. Coll. 47:194; SW:125. Sonorella (Sonorella ) mearnsi Bartsch. Thompson 2008:789.Type Locality.Sierra San Jos, Sonora, Mxico, about four miles south of the Arizona border and a few miles east of the Rio San Pedro. Holotype USNM 130003. Distribution.Known only from the type locality. Sonorella (Sonorella) mormonorum mormonorum Pilsbry 1948Sonorella mormonorum Pilsbry 1939; Land Moll. N. Amer. I:271 (a nomum nudum).Pilsbry 1948; Proc. Acad. Nat. Sci. Phila. Sonorella (Sonorella ) mormonorum mormonorum Pilsbry. Thompson 2008:789.Type Locality.Canyon on the Rio Piedras Verdes 3.5 miles above Colonia Juaraz, in talus of cliffs on south side of river, Chihuahua, Mxico; 5200 ft. alt. Holotype ANSP 166160. Distribution.Known only from the immediate vicinity of the type locality. Sonorella ( Sonorella) mormonorum huasabensis Miller 1967 Sonorella mormonum huasabensis Miller 1967; Nautilus 81:2; pl. 2008:790.Type Locality.In northwest facing rockslides about to Bavispe, Sonora, Mxico; ca. 3600 ft. alt. Holotype ANSP 312763. Distribution.Known only from the type locality. Sonorella (Sonorella) nelsoni Bartsch 1904Sonorella nelsoni Bartsch 1904; Smiths. Misc. Coll. 47:191; pl. Sonorella (Sonorella ) nelsoni Bartsch. Thompson 2008:790.Type Locality.Mountains near Lago de Santa Maria, Chihuahua, Mxico. Holotype USNM 174934. Distribution.Known only from the type locality. Sonorella (Sonorella) nixoni Miller 1967Sonorella nixoni Sonorella (Sonorella ) nixoni Miller. Thompson 2008:790.Type Locality.La Angustura, Sonora, in rocks just south of the main road at the village, overlooking the west rim of La Angustura Dam on the Rio Bavispe, Sonora, Mxico; 3100 ft. alt. Holotype ANSP 310361. Distribution.SONORA: rockslide on mountain south of road from El Tajo to La Anustura, 8.4 mi. from El Tajo, 4600 ft. alt; Pilares de Nacozari, ca. 1 mi. E of Nacozari; 4200 ft. alt. Sonorella (Sonorella) pennelli Pilsbry 1948Sonorella pennelli Pilsbry 1939; Land Moll. N. Amer. I:271 (A nomum nudum).Pilsbry 1948; Proc. Acad. Nat. Sci. Phila. Sonorella (Sonorella ) pennelli Pilsbry. Thompson 2008:791.Type Locality.Sierra de La Brea, a steep rock slide above the road from Pearson [Mata Ortiz] to Pacheco, Chihuahua, Mxico; 7000 ft. alt. Holotype ANSP 164081. Distribution.Known only from the type locality. Sonorella (Sonorella) perhirsuta Miller 1967Sonorella perhirsuta Sonorella (Sonorella ) perhirsuta Miller. Thompson 2008; 791.Type Locality.In granite rock piles in a ravine on the north-east face of the high peak, in mountains between Moctezuma and Huasabas, about 2 miles north of the Moctezuma-Huasabas road at a point 16.6 road miles east of Moctezuma, Sonora, Mxico; 4650 ft. alt. Holotype ANSP 312765. Distribution.Known only from the type locality. Sonorella (Sonorella) pratti Naranjo-Garcia 1988Sonorella pratti (shell), 4 (genitalia). Sonorella (Sonorella ) pratti Naranjo-Garcia. Thompson 2008:791.Type Locality.Sierra El Viejo, in north-facing limestone rockpile at base of cliffs, at mouth of large central

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240 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) canyon, running westerly, at point 9.5 road miles (15.2 km) north and 4.5 road miles (7.2 km) east of El Plomito, Sonora Distribution.Known only from the type locality. Sonorella (Sonorella) rothi Naranjo-Garcia 1988Sonorella rothi Sonorella (Sonorella ) rothi Naranjo-Garcia. Thompson 2008:792.Type Locality.Northwest end of Sierra Pico, in igneous rocks at base of cliffs, along road from El Plomito to Puerto Libertad, 14.9 miles (23.8 km) from El Plomito, SBMNH 34934. Distribution.Known only from the type locality. Sonorella (Sonorella) sasabe Naranjo-Garcia 1989Sonorella sasabe (shell), 3 (genitalia). Sonorella (Sonorella) sasabe Naranjo-Garcia. Thompson 2008:792. El Sasabe, 4 km south of Sasabe on road to Altar, Sonora, SBMNH 34954. Distribution.Known only from the type locality. Sonorella (Sonorella) seri Naranjo-Garcia 1988Sonorella seri Sonorella (Sonorella ) seri Naranjo-Garcia. Thompson 2008:792.Type Locality.Sierra El Viejo, in north-facing limestone piles at north end of range, Sonora, Mxico; 550 m alt. (30.1 N, 112.5 W). Holotype SBMNH 34935. Distribution.Known only from the type locality. Sonorella (Sonorella) sitiens montezuma Pilsbry & Ferriss 1919Sonorella sitiens montezuma Pilsbry & Ferriss 1919; Nautilus 33:20.Pilsbry & Ferriss 1923; Proc. Acad. Nat. Sci. Phila. Sonorella (Sonorella ) sitiens montezuma Pilsbry & Ferriss. Thompson 2008:793.Type Locality.South end of Huachuca Mountains in a deep double gulch west of last rock fronted mountain on south side of Montezuma Canyon, Cochis Co., Arizona. Distribution.Huachuca Mountains and Patagonia Mountains in southern Arizona, and in Mxico. SONORA: ft. alt. Sonorella (Sonorella) torreonica Naranjo-Garcia 1989Sonorella torreonica Sonorella (Sonorella ) torreonica Naranjo-Garcia. Thompson 2008:793. km) southeast of Magdalena, on road to Cucurpe, Sonora, SBMNH 34951. Distribution.Known only from the type locality. Sonorella ( Sonorella) walkeri walkeri Pilsbry & Ferriss 1915Sonorella walkeri Pilsbry & Ferriss 1915; Proc. Acad Nat. Sci. (genitalia). Sonorella walkeri walkeri Pilsbry & Ferriss. Bequaert & Miller 1973: Moll. Arid SW:115. Sonorella montana Pilsbry & Ferriss 1919; Nautilus 33:19. Sonorella walkeri montana Pilsbry & Ferriss 1923; Proc. Acad. Nat. Sonorella walkeri aguacalientensis Pilsbry & Ferriss 1923; Proc. Acad. Nat. Sci. Phila. 75:68. Sonorella (Sonorella) walkeri walkeri Pilsbry & Ferriss. Thompson 2008:793.Type Locality.Sonorella walkeri: Walnut Branch, Agua Caliente Canyon, Santa Rita Mountains, Santa Cruz County, Arizona; holotype ANSP 112164. Sonorella montana: Montana Mine, Montana Peak, in Bear Canyon, Pajarito Mountains, Santa Cruz Co., Arizona; holotype ANSP 43724. Distribution.SONORA: known from the Santa Rita Mountains and the Pajarito Mountains, which extend across the International Border from Arizona into Sonora. Sonorella (Sonorella) walteri Naranjo-Garcia 1989Sonorella walteri Sonorella (Sonorella) walteri Naranjo-Garcia. Thompson 2008:794.Type Locality.Cerro Gallardo, in north facing rheolite rock piles below cliffs, ca. 1.5 km south of Rancho Gallardo W). Holotype SBMNH 34953. Distribution.Known only from the type locality. Family XANTHONICHIDAE Strebel & Pfeffer 1880 Distribution.The Neotropical realm. Within the study area the family is widely distributed from Panam north to Tamaulipas, Mxico on the east coast, and to Sonora on the west coast. Taxonomy.The Xanthonichidae includes ten Mexican and Central American genera. Nordsieck (1997) and Schileyko (1998) proposed various subfamilies and tribes to accommodate the genera. The phylogeny and suprageneric taxonomy of the Middle American helicids is controversial, because anatomical data are not available for many species, evolution within reproductive anatomies, as surely has Xanthonichidae requires further investigations (Miller & Naranjo-Garcia 1991). Subfamily LEPTARIONTINAE Nordsieck 1987 Genus Leptarionta Fischer & Crosse 1872Leptarionta Fischer & Crosse 1872:253.Pilsbry 1927; Proc. Acad,

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 241Nat. Sci. Phila. 79:136.Zilch 1960:651. Type Species.Helix bicincta Pfeiffer 1841. Distribution.Central Veracruz south to Chiapas, Tabasco, Belize, Guatemala, Honduras, Costa Rica, Panam, and Colombia. Taxonomy.Eight species and eight subspecies are recognized. Leptarionta maxwillsmithi is included because it occurs in an immediate adjacent region in Colombia. A Peruvian species, Leptarionta woytkowskii Weyrauch 1960, appears to be an Helicina Leptarionta adela (Angas 1878)Helix adela 8 (shell). Helix (Oxychona) adela Angas. Von Martens 1892; Biol. Cent. Amer.:158.Von Martens 1901; Biol. Cent. Amer.:625. Oxychona adela (Angas 1878). Pilsbry 1894; Man. Conch. 9:189. Leptarionta adela (Angas). Thompson 2008:795.Type Locality.Navarro, [Prov. Cartago], Costa Rica. Distribution.COSTA RICA, Prov. Cartago: Azahar, 1500 m alt. (Von Martens 1901). Leptarionta altispira (Von Martens 1892)Helix (Oxychona) altispira Von Martens 1892; Biol. Cent. Oxychona altispira (Von Martens 1892). Pilsbry 1894; Man. Conch. 9:191. Leptarionta altispira (Von Martens). Thompson 2008:795.Type Locality.Honduras. Distribution.Known only from the type locality. Leptarionta bicincta (Pfeiffer 1841)Helix bicincta Pfeiffer 1841; Symbolae Hist. Helic., 1:38. Helix (Leptarionata) bicincta Pfeiffer. Fischer & Crosse 1872:253; Corasia bicincta shell). Helix (Oxychona) bicincta (Pfeiffer). Von Martens 1892; Biol. Cent. Amer.:157.Pilsbry 1894; Man. Conch. 9:191. Leptarionta bicincta (shell).Thompson 2008:796.Type Locality.Oaxaca, Mxico. Distribution.OAXACA: Juquila; Panistlahuaca (Von Orizaba (Von Martens 1892). Leptarionta costaricensis (Roth 1856)Helix costaricensis Roth, in Pfeiffer 1856; Novitates Conchologicae Helix boucardi Angas 1878; Proc. Zool. Soc. Lond. 46:72; pl. 5, Helix costaricensis var. steiniana Ancey 1890; Bull. Soc. Malac. France 7:155. Helix costaricensis var. Ancey 1890; Bull. Soc. Malac. France 7:156. Helix (Oxychona) costaricensis (Roth). Pilsbry 1889; Man. Conch. Martens 1892; Biol. Cent. Amer.:158.Von Martens 1901; Biol. Cent. Amer.:626. Leptarionta costaricensis (Roth). Thompson 2008:796.Type Locality.Costa Rica. Distribution.COSTA RICA, Prov. Cartago: Navarro; Cabecar (Von Martens 1892); Puente de Tierra, on the road to Sarapiqui; Santa Clara, 250 m alt.; between Mokri and Ukatschka; between Mokri and Uiskar; Alta Talamanca (Von Martens 1901). Leptarionta guillarmodi (Shuttleworth 1852)Helix guillarmodi Shuttleworth 1852. Mitt. Naturf. Ges. Bern:199. Helix (Corasia) guillarmodi Shuttleworth. Fischer & Crosse Corasia guillarmodi (Shuttleworth). Strebel 1880:50. Helix (Oxychona) guillarmodi Shuttleworth. Pilsbry 1889; Man. 1892; Biol. Cent. Amer.:158. Leptarionta guillarmodi (Shuttleworth). Pilsbry 1927; Proc. Acad. Neubert & Gosteli 2003; Contributions to Natural History Naturhistorisches Museum Bern 19080/3 (Neubert & Gosteli 2003:27). 1892); Las Tortolas, in limestone foothills ca. 2 km N of Leptarionta maxwellsmithi Pilsby 1930Leptarionta maxwellsmithi Pilsbry 1930; Nautilus 43:11617; textHolotype ANSP 150243. Distribution.Known only from the type locality. Leptarionta trigonostoma trigonostoma (Pfeiffer 1844)Helix trigonostoma Pfeiffer 1844; in Phillipi, Abbildungen neuer Helix (Geotrochus) trigonostoma Pfeiffer. Fischer & Crosse Helix (Oxychona) trigonostoma Pfeiffer. Pilsbry 1889; Man. Conch. Von Martens 1892; Biol. Cent. Amer.:154. Leptarionta trigonostoma (Pfeiffer). Bequaert 1957:223. Leptarionta trigonostoma trigonostoma (Pfeiffer). Thompson 2008:798.Type Locality.Dept. Alta Verapaz, Guatemala. Distribution.HONDURAS (von Marten 1892). GUATEMALA, Dept. Alta Verapaz: Senahu (von Marten 1892). Dept. Chimaltenango: Yepocapa (Bequaert 1957). Dept. Petn (Von Martens 1892). Dept. Solola: Finca Montequina, Atitlan (Bequaert 1957). CHIAPAS: Laguna Ocotal, 950 m alt.; El Censo to Laguna Ocotal, 700 m alt.; Monte Libano to El Censo, 600 m alt. (Bequaert 1957). TABASCO.

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242 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) (Fischer & Crosse 1872)Helix trigonostoma var. Fischer & Crosse 1872:291; Helix (Oxychona) trigonostoma var. Fischer & Crosse. Von Martens 1892; Biol. Cent. Amer.:154. (Fischer & Crosse). Pilsbry 1894; Man. Conch. 9:190. (Fischer & Crosse). Thompson 2008:798.Type Locality.Mountains of the Dept. Vera Paz, Guatemala. Distribution.GUATEMALA, Dept. Alta Verapaz: Cobn. Dept. Guatemala: near Cd. Guatemala, 5000 ft. alt. (Von Martens 1892). Leptarionta trigonostoma freytagiana (Von Martens 1892)Helix trigonostoma var. freytagiana Von Martens 1892; Biol. Cent. Oxychona trigonostoma freytagiana (Von Martens). Pilsbry 1894; Man. Conch. 9:191. Leptarionta trigonostoma freytagiana (Von Martens). Thompson 2008:798.Type Locality.Honduras. Distribution.Known only from the type locality. Leptarionta trigonostoma intermedia (Fischer & Crosse 1872)Helix trigonostoma var. intermedia Fischer & Crosse 1872:291; pl. Oxychona trigonostoma intermedia (Fischer & Crosse). Pilsbry 1894; Man. Conch. 9:190. Leptarionta trigonostoma intermedia (Fischer & Crosse). Thompson 2008:799.Type Locality.San Agustn, Dept. San Marcos, Guatemala. Distribution.GUATEMALA, Dept. Alta Verapaz: Senahu (Von Martens 1892). Dept. Quetzaltenango: Cerro Zunil (Von Martens 1892). Dept. San Marcos: San Agustn. Leptarionta trigonostoma salleana (Pfeiffer 1849)Helix salleana Pfeiffer 1849; Mon. Heliceorum Vivntium, 3:173. Helix trigonostoma var. salleana Pfeiffer. Fischer & Crosse 1872:291. Helix trigonostoma var. obscura Fischer & Crosse 1872:291; pl. 11, Helix (Oxychona) trigonostoma var. salleana Pfeiffer. Von Martens 1892; Biol. Cent. Amer.:154. Leptarionta trigonostoma salleana (Pfeiffer). Thompson 1957; Nautilus 70:101.Thompson 2008:799.Type Locality.Along the Rio San Juan, Guatemala. Distribution.GUATEMALA, Dept. Izabal: Sierra del Mico, south of the Golfo Dulce (Von Martens 1892). Dept. San Marcos: San Agustn (Von Martens 1892). TABASCO: 0.5.0 mi. E of Taepa (Thompson 1957). Leptinaria trigonostoma stolliana (Von Martens 1892)Helix trigonostoma var. stolliana Von Martens 1892; Biol. Cent. Oxychona trigonostoma stolliana (Von Martens). Pilsbry 1894; Leptinaria trigonostoma stolliana (Von Martens). Thompson 2008:799.Type Locality.Not designated. Distribution.GUATEMALA, Dept. Escuintla: Buenavista; Miramar; Hacienda Helvetia; Hacienda San Francisco (Von Martens 1892). Dept. Quetzaltenango: Cerro Zunil (Von Martens 1892). Dept. Retalhuleu: El Reposo (Von Martens 1892). Dept. San Marcos: San Agustn (Von Martens 1892). Gude 1903 Gude 1903; Proc. Malac. Soc. London 5:263; Gude. Thompson 2008:800.Type Locality.Chiriqu, Panam. Distribution.Known only from the type locality. Rehder 1942 Thompson 2008:800.Type Locality.Prov. Chiriqu, Panam. Holotype USNM 536030. Distribution.Known only from the type locality. Leptarionta zhorquinensisis (Angas 1879)Helix zhorquinensis Angas 1879; Proc. Zool. Soc. Lond. 47:475; pl. Helix (Oxychona) zhorquinensis Angas. Pilsbry 1889; Man. Con., Oxychona zhorquinensis (Angas). Pilsbry 1894; Man. Conch. 9:190. Leptarionta zhorquinensis (Angas). Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:127. Thompson 2008:800.Type Locality.Along the Rio Zhorquin, Costa Rica. Point (Pilsbry 1926). Genus Tryonigens Pilsbry 1927Tryonigens Pilsbry 1927; Proc. Acad. Nat. Sci. Phila. 79:189.Solem 1959; Occ. Pap. Mus. Zool. Univ. Mich. (611):12.Type Species.Helix rmondi Tryon 1862. Distribution.Coastal western Mxico from Sonora south to Colima. Taxonomy.A single species is recognized. Tryonigens remondi (Tryon 1863)Helix rmondi Tryon 1863; Proc. Acad. Nat. Sci. Phila. 15:281. Arionta rmondi (Tryon). Tryon 1866; Amer. Jour. Conch. 2:318, Ancey 1887; Conchologists Exchange, 2:63. Helix trypanomphala var. remondi Tryon. Von Martens 1892; Biol. Cent. Amer.:143. Tryonigens remondi (Tryon). Pilsbry 1927; Proc. Acad. Nat. Sci. (reproductive anatomy).Solem 1959; Occ. Pap. Mus. Zool. Univ. Mich. (610):12.Thompson 2008:801.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 243 Type Locality.Helix rmondi: near Mazatln, Sinaloa, Mxico; holotype ANSP 58122. : Ventanas, Durango, Mxico. Distribution.CHIHUAHUA: Rio Mayo, Guasaremos (Solem 1959). COLIMA: Manzanillo (Pilsbry 1927). DURANGO: Ventanas. GUERRERO: Agua de Obispo (Solem 1959). JALISCO: Sayul (Von Martens 1892); Puebla de Santa Catarina (Solem 1959). MICHOACN: Cerro Guzman, Coalcoman (Solem 1959). NAYARIT: Tepc; Rio Tepc, 5 mi. below Tepc (Solem 1959). SINALOA: Mazatln; Rosario; Panuco 1800 ft. alt. (Solem 1959). SONORA: Guaymas (Tryon 1866). Subfamily LYSINOINAE Hoffmann 1928 Taxonomy.A single genus is placed in this subfamily. Genus Lysinoe H. Adams & A. Adams 1855Lysinoe H. Adams & A. Adams 1855; Genera of Recent Mollusca (2):203.Pilsbry 1894; Man. Conch. 9:191.Type Species.Helix ghiesbreghti Nyst 1841. Distribution.Chiapas, Guatemala, El Salvador and Honduras. A species from Jalisco is included in the genus, Taxonomy.Four species and six subspecies are recognized. Lysinoe eximia eximia (Pfeiffer 1844)Helix eximia Pfeiffer 1844; in Philippi, Abbildungen neuer Conchyl., Helix (Odontura) eximia Pfeiffer. Fischer & Crosse 1872:242; (nervous system). Odontura eximia (Pfeifer). Strebel 1880:36. Helix (Lysinoe) eximia Pfeiffer. Von Martens 1892; Biol. Cent. Amer.:149. Lysinoe eximia (Pfeiffer). Pilsbry 1894; Man. Conch. 9:192; pl. 60, Lysinoe eximia eximia (Pfeiffer). Thompson 2008:802.Type Locality.Not given. Distribution.GUATEMALA, Dept. Alta Verapaz: Cobn. Dept. Huehuetenango: upper Cholhuitz; near San Martin, ca. 6000 ft. alt. (Von Martens 1892). Dept. Sacatepquez: Dueas (Fischer & Crosse 1872). Lysinoe eximia stolli (Von Martens 1892)Helix (Odonture) eximia Pfeiffer. Fischer & Crosse 1873; Miss. Sci. Helix (Lysinoe) eximia var. stolli Von Martens 1892; Biol. Cent. Amer.:149. Lysinoe eximia stolli (Martens). Pilsbry 1894; Man. Conch. 9:192.Thompson 2008:803.Type Locality.Vera Paz, Guatemala. Distribution.Known only from the type locality. Lysinoe ghiesbreghti ghiesbreghti (Nyst 1841)Helix ghiesbreghti (shell). Helix (Odontura) ghiesbreghti Nyst. Fischer & Crosse 1872:245, pl. Odontura ghiesbreghti (Nyst). Strebel 1872:37. Helix (Lysinoe) ghiesbrghti Nyst. Von Martens 1892; Biol. Cent. Lysinoe ghiesbreghti (Nyst). Pilsbry 1894; Man. Conch. 9:192; pl. Lysinoe ghiesbreghti ghiesbreghti (Nyst). Thompson 2008:803.Type Locality.Chiapas, Mxico. Distribution.GUATEMALA: Dept. Alta Verapaz: Colocality). Lysinoe ghiesbreghti bizonata (Von Martens 1892)Helix (Odontura) ghiesbreght var. Fischer & Crosse 1872:245, 247. Helix (Lysinoe) ghiesbreghti var. bizonata Von Martens 1892; Biol. Cent. Amer.:151. Lysenoe ghiesbreghti bizona (Von Martens). Pilsbry 1894; Man. Conch. 9:192.Thompson 2008:804.Type Locality.Cordierra de San Marcos, Honduras; ca. 2660 m alt. Distribution.Known only from the type locality. Lysinoe ghiesbreghti (Von Martens 1892)Helix (Odontura) ghiesbreghti var. Fischer & Crosse 1872:245; pl. Helix (Lysinoe) ghiesbreghti var. Von Martens 1892; Biol. Cent. Amer.:151. Lysinoe ghiesbreghti (Von Martens). Pilsbry 1894; Man. Conch. 9:192.Thompson 2008:804.Type Locality.Tolimn, in the hills above San Lucas, near Lago de Atitln, Dept. Solola, Guatemala. Distribution.Known only from the type locality. Lysinoe ghiesbreghti rufozonata (Von Martens 1892)Helix (Odontura) ghiesbreghti var. Fischer & Crosse 1872:247. Helix (Lysinoe) ghiesbreghti var. rufo-zonata Von Martens 1892; Biol. Cent. Amer.:151. Lysinoe ghiesbreghti rufozonata (Von Martens). Pilsbry 1894; Man. Conch. 9:192.Thompson 2008:804.Type Locality.Not given. Distribution.GUATEMALA (?). Lysinoe ghiesbreghti subaurantia (Von Martens 1892)Helix (Lysinoe) ghiesbreghti var. subaurantia Von Martens 1892; Lysinoe ghiesbreghti subaurantia (Von Martens). Pilsbry 1894; Man. Conch. 9:192.Thompson 2008:805.Type Locality.Purula [Purulha], Dept. Baja Verapaz, Guatemala. Distribution.GUATEMALA, Dept. Baja Verapaz: Purulha. Dept. Quetzaltenango: Cerro Zunil (Von Martens 1892). Lysinoe ghiesbreghti strubelli (Von Martens 1892)Helix (Lysinoe) ghiesbreghti var. strubelli Von Martens 1892; Biol.

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244 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Lysinoe ghiesbreghti strubelli (Von Martens). Pilsbry 1894; Man. Conch. 9:192.Thompson 2008:805.Type Locality.Cabo Gracias a Dios, Honduras. Distribution.Known only from the type locality. Lysinoe sebastiana (Dall 1897)Helix (Lysinoe) sebastiana Dall 1897:73. Lysinoe sebastiana (Dall). Solem 1955; Occ. Pap. Mus. Zool. Thompson 2008:805.Type Locality.Near San Sebastian, Jalisco, Mxico. Holotype in the USNM. Distribution.JALISCO: Sierra Autln, ca. 20 mi. SSE of Autln, Jalisco (Solem 1955). Lysinoe starretti Thompson 1963Lysinoe staretti Thompson 1963; Proc. Biol. Soc. Wash. 76:27; 1, C-D, a-f (reproductive anatomy).Type Locality.Hacienda Monte Cristo, Metapan, Dept. Santa Ana, El Salvador; 2200 m alt. Holotype UMMZ 195327. Distribution.Known only from the type locality. Subfamily TRICHODISCININAE Schileyko 1998 and from Tamaulipas on the east coast south to Panam. Taxonomy.Three genera are included. The subfamily Miraveralliinae Schileyko 1998 is a synonym. Genus Ancey 1887Coelospira Ancey 1886; Conch. Exchange, 1:20 (not Coelospira Hall 1864: Brachiopoda). Ancey 1887; Conch. Exchange, 1:54. Pilsbry 1894; Man. Conch. 9:197 198.Type Species.Helix mac-neili Crosse 1873. Distribution.Southern Costa Rica and northwestern Panam. Taxonomy.A single species is recognized. (Crosse 1873)Helix mac-neili Crosse 1873; Jour. de Conchyl. 21:67.Crosse Helix ( ) macneili (Crosse). Pilsbry 1889; Man. Conch. Amer.:137.Von Martens 1901; Biol. Cent. Amer.:623. Epiphragmophora ( ) macneili (Crosse). Pilsbry 1895; Man. Conch. 9:197. (Crosse). Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:73.Thompson 2008:806.Type Locality.Chiriqu Lagoon, Costa Rica [Panam]. Distribution.PANAM, Prov. Bocas del Toro: drift of San San Creek. Prov. Chiriqu: Chiriqu Lagoon (Pilsbry 1926). COSTA RICA, Prov. Cartago: Tus, 600 m lower foothills from Parismina to the hills of Zhorquin (Von Martens 1892). Prov. Puntarenas: Alto de Mano Tigre, 690 m alt.; Terraba (Von Martens 1892). Genus Baker 1922 Baker 1922; Occ. Pap. Mus. Zool. Univ. Mich. (106):60.Solem 1959; Occ. Pap. Mus. Zool. Univ. Mich. (611):1.Type Species.Helix sumichrasti Crosse & Fischer 1872. Distribution.Southern Mxico in the states of Chiapas, Oaxaca, Guerrero, Michoacn and Veracruz. Taxonomy.Four species are recognized. (Thompson 1859) ( ) Thompson 1959; Occ. Pap. Mus. (reproductive anatomy). (Thompson). Thompson 2008:807.Type Locality.El Fuente, south slope of Volcn San Martn, Sierra de las Tuxtlas, Veracruz, Mxico; 1170 m alt. Holotype UMMZ 196002. Distribution.Known only from the type locality. (Crosse & Fischer 1872)Helix sargi Crosse & Fischer 1872; Jour. de Conchyl. 20:146.Von Martens 1874; Proc. Zool. Soc. Lond. 42:648. Helix (Aglaia) sargi Crosse & Fischer. Crosse & Fischer 1873; Jour. Helix (Trichodiscina) sargi Crosse & Fischer. Von Martens 1892; Biol. Cent. Amer.:157. Trichodiscina sargi (Crosse & Fischer). Hinkley 1920; Nautilus 34:50. ( ) sargi (Crosse & Fischer 1872). Solem 13 (shell). (Crosse & Fischer). Thompson 2008:809.Type Locality.Tamah, [Dept. Alta Verapaz], Guatemala. Distribution.GUATEMALA, Dept. Alta Verapaz: Tamah; Cobn (Von Martens 1892); Chejel (Hinkley 1920). (Crosse & Fischer 1872)Helix sumichrasti Crosse & Fischer 1872; Jour. de Conchyl. 20:146. Helix (Trichia) sumichrasti Crosse & Fischer 1873; Jour. de Helix (Trichodiscina ) sumichrasti Crosse & Fischer. Von Martens 1892; Biol. Cent. Amer.:137. ( ) sumichrasti (Crosse & Fischer). Baker 1922; (Crosse & Fischer). Thompson. 2008:808.Type Locality.[Santiago] Laollaga, Isthmus of

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 245 Tehuantepec, Oaxaca (16.58 N, 95.2 W) (Solem 1959). Distribution.OAXACA: Laollaga. Remarks.Records by Solem (1959) from Veracruz and Michoacn are doubtful. (Dall 1910) Epiphragmophora (Trichodiscina ) Dall 1910; Nautilus 24:35 (not illustrated). ) sumichrasti (Crosse & Fischer). Solem 1959; Occ. Pap. Mus. Zool. Univ. Mich. (611):3 (in part). (Dall). Thompson 2008:809.Type Locality.Near the hot springs, along the Rio Verde, Oaxaca, Mxico. Further restricted to Rio Verde, Oaxaca (17.057 N, 94.751 W) (Solem 1959). Holotype USNM 212318. Distribution.Known only from the type locality. Genus Trichodiscina Von Martens 1892Trichodiscus Strebel 1880; Beitrag. Mex. Landund Sssw.-Conch. IV:32 (not Trichodiscus Ehrenberg 1830, Protozoa). Trichodiscina Von Martens 1892; Biol. Cent. Amer.:133.Prez & Type Species.Helix coactiliata Frussac 1838. Distribution.Venezuela north through Central America to Tamaulipas and Sinaloa. Taxonomy.Six species and one subspecies are recognized. Trichodiscina crinita Fulton 1917 belongs in Thysanophora (Setidiscus ). Trichodiscina coactiliata (Frussac 1838)Helix coactiliata Deshayes 1839, in Frussac, Hist. Nat. Moll. Helix (patula) coactiliata (Frussac). Fischer & Crosse 1872:234. Trichodiscus coactiliata (Frussac). Strebel 1880:34.Prez & Patula coactiliata (Frussac). Pilsbry 1889; Proc. Acad. Nat. Sci. Phila. 43:314. Helix (Trichodiscina ) coactiliata (Frussac). Von Martens 1892; Biol. Cent. Amer.:133, Epiphragmophora coactiliata (Frussac). Pilsbry 1895; Man. Conch. 9:191.Fluck 1905; Nautilus 19:78. (Trichodiscina) coactiliata (Frussac). Hinkley 1907; Nautilus 21:71.Baker 1922; Occ. Pap. Mus. Zool. Univ. Mich. Occ. Pap. Mus. Zool. Univ. Mich. (167):22.Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:73.Bequaert & Clench 1933; Pub. Carnegie Inst. Wash. (431):531.Bequaert & Clench 1936; Pub. Carnegie Inst. Wash. (457):64.Richards 1937; Proc. Amer. Philos. Soc. 77:252.Richards 1938:172.Morrison 1946; Smiths. Misc. Coll. 106:44.Haas 1949; Nautilus 62:138.Harry 1950; Occ. Pap. Mus. Zool. Univ. Mich. (524):11.Bequaert 1957:224.Basch 1959; Occ. Pap. Mus. Zool. Univ. Mich. (612):11.Haas & Solem 1966; Nautilus 73:131.Thompson 1967; Bull. Fla. Sta. Mus., 11:253. Helix nystiana Pfeiffer 1845; Proc. Zool. Soc. Lond. 13:130. Helix bridgesi 9, 10 (shell). Helix parkeri Tryon 1867; Amer. Jour. Conch. 3:106.Type Localities.Helix coactiliata : not given. Helix nystiana: Realejo, Nicaragua. Helix bridgesi: Nicaragua. Helix parkeri : Nicaragua. Distribution.Eastern Mxico from Tamaulipas south through Central America to Trinidad and Venezuela (Baker 1926). PANAM, Archipilago de las Perlas: Isla San Jos (Morrison 1946). Prov. Los Santos: Tonos (Pilsbry 1926). 2002). Dept. Chinandega: Realejo (Von Martens 1892). Reg. Aut. Atlantico Norte: Weilawas Hill, near Wani (Fluck 1905). HONDURAS, Isla de Roatn: between Coden Hole and French Harbor; North Side; West End (Richards 1938). BELIZE: Rio Frio Cave (Haas & Solem 1966). GUATEMALA, Dept. Escuintla: Zapote (Haas 1949). Dept. Izabal: near Livingston. Dept. Petn: Tikal National Park (Basch 1959). CAMPECHE:5.1 mi. NNE of Dzibalchen; 3.6 mi. S of Hopelchen; 6.1 mi. E of Seyaplaya; 7.2 mi. S of Pixtun; 10.2 mi. E of Escarcega; 19.2 mi. E of Silvituc (Thompson 1967). CHIAPAS: Laguna Ocotal, 950 m alt.; El Real, 600 m alt. (Bequaert 1957). QUINTANA ROO: Isla Cozumel, 3 mi. N of San Miguel (Richards 1937); Chetumal (Haas & Solem 1966); 4.0 mi. E of Xpujil, Campeche (Thompson 1967). TAMAULIPAS: Tampico (Hinkley 1907). VERACRUZ: Coatotolapam (Baker 1922). YUCATN: Tablas; Labna (Pilsbry 1891); Chichen Itza (Bequaert & Clench 1933); Xtolok Cenote, near Chichen Itza (Bequaert & Clench 1936); 8 mi. NW of Chichen Itza (Harry 1950). (Pfeiffer 1858) Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:318. Helix (Patula ) coactiata var. Pfeiffer. Fischer & Crosse 2008:811. (Pfeiffer). Von Martens 1892; Biol. Cent. Amer.:135. (Trichodiscina ) (Pfeiffer). Baker 1927; Proc. (Dall). Correa-Sandoval 1993; Rev. Biol. Trop. 41:675.Correa-Sandoval, Garca-Cubas & Reguero 1998:17.Correa-Sandoval 2000:9. Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):238.Type Locality.Tuspan [Tuxpam], south of Tampico, Veracruz, Mxico. Distribution.GUERRERO: Omilteme (Von Martens 1892). NUEVO LEN: Santiago (Correa-Sandoval 1993). SAN LUS POTOS: numerous localities (Correa-Sandoval et al.. 1998). TAMAULIPAS: numerous localities in southern part of state (Correa-Sandoval & Rodriguez 2002). VERACRUZ: Nacimiento de Quilate, Cerro de Espadilla, Rancho Guerrero, all in the vicinity of Misantla; numerous localities in the vicinity of Jalapa; hacienda Mirador; Soledad;

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246 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Trichodiscina hinkleyi Pilsbry 1919Trichodiscina hinkleyi Thompson 2008:812. (Pilsbry). Hinkley 1920; Nautilus 34:43, 48.Type Locality.Livingston, [Dept. Izabal], Guatemala. Holotype ANSP 107533. Distribution.GUATEMALA, Dept. Izabal: mountains of Rio Cavech, and those back of Cavech Village; across the Rio Dulcey from Livingston, Jocola (Hinkley 1920). Trichodiscina oajacensis (Kock 1842)Helix oajacensis Kock, in Pfeiffer 1842; Symbolae ad Historiam Heliciorum, 2:35. Helix (Discus) oajacensis Kock. Fischer & Crosse 1872:237. Trichodiscus oajacensis (Kock). Strebel 1880:35.Tryon 1887; Epiphragmophora (Trichodiscina ) oajacensis (Kock). Pilsbry 1895; Man. Conch. 9:199. Helix (Trichodiscina) oajacensis (Kock). Von Martens 1892; Biol. Cent. Amer.:136.Type Locality.Oaxaca, Mxico. Distribution.Known only from the type locality. Trichodiscina sinaloa Pilsbry 1954Trichodiscina sinaloa Thompson 2008:812.Type Locality.Drift of the Rio Fuerte, San Blas, Sinaloa, Mxico. Holotype ANSP 166601. Distribution.Known only from the type locality. Trichodiscina suturalis suturalis (Pfeiffer 1846)Helix suturalis Pfeiffer 1846; Proc. Zool. Soc. Lond. 14:37.Reeve 1851; Conch. Icon., Helix 626 (shell). Helix (Patula ) coactiliata suturalis Pfeiffer. Fischer & Crosse 1872:234. Helix (Trichodiscus) suturalis Pfeiffer. Tryon 1887; Man. Conch. Helix (Trichodiscina) suturalis Pfeiffer. Von Martens 1892; Biol. Cent. Amer.:136. (Trichodiscina) suturalis (Pfeiffer). Baker 1922; Occ. Pap. Mus. Zool. Univ. Mich. (106):58. (Pfeiffer). Goodrich & van der Schalie 1937:28.Thompson 1957; Nautilus 70:101.Thompson 1967c:254. Trichodiscina suturalis suturalis (Pfeiffer). Thompson 2008:813.Type Locality.Honduras. Distribution.HONDURAS. GUATEMALA (Goodrich near Salama. Dept. Petn: San Luis (Von Martens 1892). CAMPECHE:5.1 mi. NNW of Dzibalchen; 19.2 mi. E of Silvituc (Thompson 1967c). QUINTANA ROO:4.0 mi. E of Xpujil, Campeche; 2.3 mi. SSE of Xiatil (Thompson 1967c). TABASCO: 0.5.0 mi. E of Teapa (Thompson 1957). VERACRUZ: Cuatotolapam (Baker 1922). Trichodiscina suturalis pressula (Morelet 1851)Helix pressula Morelet 1851; Test. Noviss. II:8. Helix (Patula ) pressula Morelet. Fischer & Crosse 1872:236; pl. 12, Trichodiscus pressula (Morelet). Strebel 1880:34. Helix (Trichodiscus) pressula Morelet. Tryon 1887; Man. Conch. Helix (Trichodiscina) suturalis var. pressula Morelet. Von Martens (Morelet). Haas 1949; Nautilus 62:137. Helix almonte Tristram 1863; Proc. Zool. Soc. Lond. 31:411. Helix almonteana Tristram, Fischer & Crosse 1872:231. Trichodiscina suturalis pressula (Morelet).Thomposn 2008:813.Type Locality.Istapa, near San Jos, Dept. Escuintla, Guatemala. Distribution.COSTA RICA, Prov. San Jos: Uruca. GUATEMALA, Dept. Baja Verapaz: Purulha. Dept. Chimaltenango: Finca Montserrat, W slope of Volcn Acatenango (Haas 1949). Dept. Guatemala: limestone mountain La Pedrera N of Cd. Guatemala. Dept. Quetzaltenango: Las Mercedes (Von Martens 1892). Subfamily XANTHONICHINAE Strebel & Pfeffer 1880Xanthonicinae Strebel & Pfeffer, in Strebel 1880; Beitrag. Mex. Landund Sssw.-Conch. IV:25. Metastracini Nordseick 1978; Arch. fr Molluskenkunde, 118:22 23.Distribution.Mxico and Central America. Taxonomy.Four genera are recognized. Genus Cryptostracon Binney 1879Cryptostracon Binney 1879; Ann. New York Acad. Sci. 1:258.Pilsbry 1900; Proc. Malac. Soc. London 4:27.Cuezzo 1997; Amer. Malac. Bull., 14:7.Type Species.Cryptostracon gabbi Binney 1879. Distribution.Costa Rica. Taxonomy.Two species are recognized. Cuezzo 1997 Cuezzo 1997; Amer. Malac. Bull., (radula).Thompson 2008:814.Type Locality.Costa Rica, Prov. Puntarenas, Corcovado National Park, Estacion Sirena, along Rio Los Patos; 10 m alt. Holotype UCR-INBIO 648087. Distribution.COSTA RICA: known only from the vicinity of the type locality. Cryptostracon gabbi Binney 1879Cryptostracon gabbi Binney 1879; Ann. New York Acad. Sci. Acad. Nat. Sci. Phila. 115:248.Cuezzo 1997; Amer. Malac. Bull., 14:108.Thompson 2008:815.Type Locality.Flanks of Pico Blanco, 5000 ft. elevation. Lectotype ANSP 246310 (Baker 1963:248).

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 247 Distribution.Known only from the type locality. Genus Metastracon Pilsbry 1900Metastracon Pilsbry 1900; Proc. Malac. Soc. London 4:24.Type Species.Metastracon mima Pilsbry 1900. Distribution.Michoacn, Mxico. Taxonomy.A single species is recognized. Metastracon mima Pilsbry 1900Metastracon mima Pilsbry 1900; Proc. Malac. Soc. London 4:27; 2008:815.Type Locality.Morelia, Michoacn, Mxico. Holotype ANSP 77245. Distribution.MICHOACN: Morelia; Uruapan (Pilsbry 1900). Genus Semiconchula Naranjo-Garcia, Polaco & Pearce 2000Semiconchula Naranjo-Garcia, Polaco & Pearce 2000; Archiv fr Molluskenkunde 128:156.Type Species.Semiconchula custepecana NaranjoGarcia, Polaco & Pearce 2000. Distribution.Southern Chiapas, Mxico. Taxonomy.Two species are recognized. Naranjo-Garcia 2003 Naranjo-Garcia 2003; Proc. Calif. Acad. Sci. 54:225.Thompson 2008:816.Type Locality.West of San Cristobal de Las Casas; steep, heavily wooded slope near the summit of Mukta vits (Cerro Huitepec), Chiapas, Mxico. Holotype CAS 77540. Distribution.Known only from the type locality. Semiconchula custepecana Naranjo-Garcia, Polaco & Pearce 2000 Semiconchula custepecana Naranjo-Garcia, Polaco & Pearce 2000. Thompson 2008:816.Type Locality.North side of Sierra Madre de Chiapas, east bank of Rio Custepec, ca. 4 km north by road from Finca Custepec toward Jaltenango, Chiapas, Mxico, 1040 m. Among riparian vegetation. Holotype Coleccion Nacional de Moluscos, Universidad Nacional Autonoma de Mxico 326. Distribution.CHIAPAS: known from four localities within 10 km of the type locality. Genus Xanthonyx Crosse & Fischer 1867Xanthonyx Crosse & Fischer 1867; Jour. de Conchyl. 15:223.Fischer & Crosse 1872:192.Strebel 1880:26.Type Species.Vitrina sumichrasti Brot 1867. Distribution.Eastern Mxico from San Luis Potosi south to Chiapas. Taxonomy.Five species are recognized. Xanthonyx chiapensis (Pfeiffer 1856)Simpulopsis chiapensis Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:377.Reeve, 13, ( Simpulopsis Xanthonyx chiapensis (Pfeiffer). Fischer & Crosse 1872:203.Thompson 2008:217.Type Locality.Chiapas, Mxico. Distribution.Known only from the type locality. (Pfeiffer 1856) Pfeiffer 1856; Proc. Zool. Soc. Lond. (Pfeiffer). Crosse & Fischer 1867: Jour. de 3a (shell).Strebel 1880; Beitrag. Mex. Landund Sssw.7G (reproductive anatomy).Naranjo-Garcia, Polaco & Pearce 2000, Arch. fr Mollusk. 128:160.Thompson 2008:817. BMNH 1992132 (Naranjo-Garcia et al. 2000:160). Monte, between Jalapa and Misantla (Strebel 1880). Xanthonyx potosiana Dall 1905Xanthonyx potosiana Dall 1905; Smithsonian Miscellaneous Sandoval & Rodriguez 2002; Acta Zool. Mex. 2(86):61.Correa-Sandoval & Salazar 2005; Acta Zool. Mex. 21:61.Correa-Sandoval et al. 2007; Acta Zool. Mex. 23:162.Thompson 2008:217.Type Locality.Sierra Alvarz, San Lus Potos, Mxico; 7900 ft. alt. Holotype USNM 110396. Distribution.NUEVO LEN: El Salto, Zaragoza (Correa-Sandoval & Salazar 2005). SAN LUS POTOS: Sierra Alvarz. TAMAULIPAS: Aniceto Medrono (CorrreaSandoval & Rodriguez 2002); El Cielo Biosphere Reserve (Correa-Sandoval & Rodriguez 2005). Xanthonyx salleanus (Pfeifffer 1856)Simpulopsis salleana Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:319; Xanthonyx salleanus (Pfeiffer).Crosse & Fischer 1867; Jour. de 1900; Proc. Malac. Soc. London 4:27; pl. 3, 12 (animal), (retractor muscles).Naranjo-Garcia, Polaco & Pearce 2000, Arch. fr Mollusk. 128:160.Thompson 2008:818. BMNH 1992131 (Naranjo-Garcia et al. 2000:160). Distribution.OAXACA (Von Martens 1897). VERA

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248 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Xanthonyx sumichrasti (Brot 1867)Vitrina sumichrasti 2 (shell). Xanthonyx sumichrasti (Brot). Crosse & Fischer 1867; Jour. de Garcia, Polaco & Pearce 2000; Arch. fr Mollusk. 128:160.Thompson 2008:818.Type Locality.Mxico. Holotype Museum dHistorie Naturelle de Geneve 993/213 (Naranjo-Garcia et al. 2000:160). Distribution.MXICO: without nearer indication of locality. Family POLYGYRIDAE Pilsbry 1895 Subfamily POLYGYRINAE Pilsbry 1895 Distribution.Temperate North America south to northern Central America. One species has been widely introduced to other tropical and subtropical regions. Taxonomy.Emberton (1995) listed twelve genera belonging to this subfamily. Genus Ashmunella Pilsbry & Cockerell 1899Ashmunella Pilsbry & Cockerell 1899; Nautilus 12:107.Pilsbry & Cockerell 1899; Proc. Acad. Nat. Sci. Phila. 51:188.Ancey & Mordoch 1901; Jour. Malac., 8:73.Pilsbry 1905; Proc. Acad. Nat. Sci. Phila:223.Pilsbry & Ferriss 1910; Proc. Acad. Nat. Sci. Phila. 61:503.Pilsbry & Ferriss; 1910; Proc. Acad. Nat. Sci. Phila. 62:95.Pilsbry & Ferriss 1917; Proc. Acad. Nat. Sci. Phila. 69:88.Pilsbry 1940; Land Moll. N. Amer. I:912.Bequaert & Clench 1973: Moll. Arid S.W.:37, 130.Type Species.Polygyra miorhyssa Dall 1898 (Pilsbry 1905:223. Distribution.Western Texas, New Mexico, and southeastern Arizona, south into Chihuahua, Mxico. Taxonomy.Numerous species. Five occur in Mxico. Ashmunella intricata Pilsbry 1948Ashmunella intricata Pilsbry 1948; Proc. Acad. Nat. Sci. Phila. Type Locality.Mountain side about 4.5 miles southeast of Pearson [Mata Ortz], among stones at the base of a cliff, Chihuahua, Mxico; 6200 ft. alt. Holotype ANSP 166324. Distribution.Known only from the type locality. Ashmunella juarazensis Pilsbry 1948Ashmunella juarazensis Pilsbry 1948; Proc. Acad. Nat. Sci. Phila. Type Locality.South side of the Rio Piedras Verdes 3.5 miles above Colonia Juarz, Chihuahua, Mxico; 5200 ft. alt. Holotype ANSP 166161. Distribution.CHIHUAHUA: known only from the immediate vicinity of the type locality. Ashmunella meridionalis Pilsbry 1948Ashmunella meridionalis Pilsbry 1948; Proc. Acad. Nat. Sci. Phila. Type Locality.Barranca Colorado, 21 miles south of Mieca, Chihuahua, Mxico. Holotype ANSP 164084. Distribution.CHIHUAHUA: known only from the vicinity of the type locality. Ashmunella milesi Reeder 1993Ashmunella milesi 5 (genitalia).Type Locality.Northern Sonora, Mxico, west of Cananea; south-facing talus slope along road to microwave W. Holotype SBMNH 35609. Distribution.Known only from the type locality. Pilsbry 1948 Pilsbry 1948; Proc. Acad. Nat. Sci. Phila. Type Locality.Sierra de La Brea, in a rock slide above the road from Colonia Juarz and Pearson [Mata Ortiz] to Pacheco, Chihuahua, Mxico; 700 ft. alt. Holotype: ANSP 166158. Distribution.Known only from the type locality Genus Erectidens Pilsbry 1953Erectidens trichalis Pilsbry 1953; Nautilus 67:47.Bogan 1992; Malac. Rev. 25:131.Type Species.Erectidens trichalis Pilsbry 1953. Distribution.Known only from river drift along the Mxico. Taxonomy.A single species is recognized. It is known only from the holotype. Pilsbry (1953b:46) tentatively placed Erectidens in the Polygyridae. Other authors placed it in other (Bogan 1992:131). Erectidens trichalis Pilsbry 1953Erectidens trichalis Type Locality.Rio Maurisco near its junction with the short stream on the eastern slope of the Sierra Madre Oriental Holotype ANSP 164748. Distribution.Known only from the type locality. Genus Giffordius Pilsbry 1930Giffordius Pilsbry 1930; Nautilus 43:143.Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. 82:249.Type Species.Giffordius pichoti Pilsbry 1930. Distribution.Isla de Providencia, Colombia. Taxonomy.Two species are recognized. Giffordius pinchoti Pilsbry 1930Giffordius pinchoti Pilsbry 1930; Nautilus 43:143.Pilsbry 1930;

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 249 Type Locality.Valley opening west, north of High Peak, Isla de Providencia. Holotype ANSP 150865. Distribution.Known only from Isla de Providencia. Giffordius corneliae Pilsbry 1930Giffordius corneliae Pilsbry 1930; Nautilus 43:143.Pilsbry 1930; (shell).Thompson 2008:822.Type Locality.Valley opening west, north of High Peak, Isla de Providencia. Holotype ANSP 150864. Distribution.Known only from Isla de Providencia. Genus Polygyra Say 1818Polygyra Say 1818: Jour. Acad. Nat. Sci. Phila. 1:276.W. G. Binney 1878:262.Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. 82:311.Pilsbry 1940; Land Moll. N. Amer. I:578.Emberton 1995; Malacologia 37:89. Cyclodoma Swainson 1840; Treatise on Malacology:193. Ulostoma Albers 1856; Die Heliceen:95. Anchistoma H. and A. Adams 1855; Genera of Recent Mollusca, 2:205. (Name ammended to Angistoma; Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. 82:311.)Type Species.Polygyra : Say 1818 (Hermannsen 1847:317). Cyclodoma: Say 1818 (Pilsbry 1930). Ulostoma: Say 1818 (Pilsbry 1930). Anchistoma: Say 1818 (Pilsbry 1930). Distribution.The southeastern United States south throughout most of Mxico to Guatemala and Belize. Taxonomy.A recent analysis of the phylogeny of the Polygyridae recognized within the clade Polygyri seven subclades of unequal value (Emberton 1995). Each of these subclades was elevated to generic rank. Polygyri equals Polygyra plus Praticolella and Giffordius. The remaining Polygyra The phylogeny proposed by Emberton produced a new understanding of group relationships, but the proposed clear advantages over previous nomenclatorial systems. For purposes of this document Giffordius, Praticolella and Polygyra are recognized as separate genera. Six subgenera of Polygyra are recognized. Four occur in Mxico. Two others, Lobosculum, and Daedalochila are extralimital. Subgenus Polygyra Say 1818 Distribution.The Bahama Islands, Bermuda, Cuba, and the coastal zone of the southeastern United States south to Yucatn, Mxico. Taxonomy.Four species are recognized. One occurs in Mxico. Polygyra (Polygyra ) cereolus cereolus (Muhlfeld 1818)Helix cereolus Muhlfeld 1818; Gesellschaft naturforschender Polygyra cereolus cereolus (Muhlfeld).W. G. Binney 1878; Terr. 379 (shell).Thompson 2008:823. Helix cereolus var. laminifera W. G. Binney 1858; Proc. Acad. Nat. Sci. Phila.:200 ( nomum nudum). Helix carpenteriana Bland 1860; Ann. Lyceum Nat. Hist. N. Y., 7:138. Polygyra cereolus carpenteriana (Bland). Pilsbry 1894; Man. Conch. 9:73.Bequaert & Clench 1933; Publ. Carnegie Inst. Wash., (431):531.Harry 1950; Occ. Pap. Mus. Zool. Univ. Mich. 534:7, 8.Rehder 1966; Proc. Biol. Soc. Wash. 79:289.Correa-Sandoval, Garca-Cubas & Reguero 1998; Acta Zool. Mex. (73):17.Type Localities. Helix cereolus: restricted to Key West, Florida (Pilsbry 1940:584). Helix carpenteriana : Key Biscayne, Dade Co., Florida. Lectotype ANSP 10972 (Pilsbry 1940:586). Distribution.Florida and eastern Mxico. QUINTANA ROO: Puerto Morelos; Isla Mujeres. YUCATN: 1.7 km SSE Puerto Telchac (UF 19190); Progreso; 5.3 km S Progreso (UF 19189). SAN LUS POTOS: Laguna Media Luna, 10 km SSW of Rio Verde (UF 151961, UF 189664). Subgenus Acutidens Pilsbry 1956Acutidens Pilsbry 1956; Proc. Acad. Nat. Sci. Phila. 108:26.Type Species.Helix (Polygyra ) acute-dentata W. G. Binney 1858. Distribution.Coastal regions of Sinaloa, Mxico. Taxonomy.The subgenus contains a single species. Polygyra (Acutidens) acutedentata (W. G. Binney 1858)Helix (Polygyra ) acute-dentata W. G. Binney 1858; Proc. Acad. Nat. Sci. Phila. ( for 1857), 9:183. Helix acutedentata Binney. W. G. Binney 1869; Land and Fresh Polygyra acutedentata (Binney). Von Martens 1892; Biol. Cent. Amer.:172.Pilsbry 1956; Proc. Acad. Nat. Sci. Phila. 108:26 Daedalocheila acutidentata (Binney). Emberton 1995; Malacologia 37:91. Helix (Polygyra) loisa W. B. Binney 1858; Proc. Acad. Nat. Sci. Phila. (for 1857), 9:183. Helix acutedentata var. quinquedentata Fischer & Crosse 1872; Miss. Sci. Mex. I:289. Polygyra (Acutidens ) acutedentata (W. G. Binney). Thompson 2008:824. Helix acutedentata var. minor Fischer & Crosse 1872; Miss. Sci. Mex. I:289. Helix unguifera 111b.Martens 1892; Biol. Cent. Amer.:172, 627. (A nomum nudum).Type Locality.Helix acuta-dentata : Mazatln, Sinaloa, Mxico; holotype ANSP 33449. Helix loisa: Mazatln, Sinaloa, Mxico; holotype: ANSP 33450. Helix acutedentata var. quinquedentata : not given. Helix acutedentata var. minor: along streams at Mazatln, Sinaloa, Mxico. Distribution.SINALOA: known only from the vicinity

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250 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) of the type locality; highway ca. 4 km N of Mazatln; ca. 10 km N of Mazatln. Subgenus Linisia Pilsbry 1930Linisia Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. 82:315.Pratt 1981; Unpubl. thesis:57.Emberton 1995; Malacologia 37:90. Erymodon Pilsbry 1956; Proc. Acad. Nat. Sci. Phila. 108:20.Solem 1959; Occ. Pap. Mus. Zool. Univ. Mich. (611): 10. Monophysis Pilsbry 1956; Proc. Acad. Nat. Sci. Phila. 108:25. Solidens Pilsbry 1956; Proc. Acad. Nat. Sci. Phila. 108:27.Type Species.Linisia: Helix (Polygyra) anilis Gabb 1865. Erymodon: Pfeiffer 1845. Monophysis: Helix (Polygyra) behri Gabb 1865. Solidens: Helix (Polygyra) anilis Gabb 1865. Distribution.Texas, Oklahoma and Louisiana south throughout most of Mxico to the Dept. Petn, Guatemala, and Belize. are recognized. All occur or are presumed to occur in Mxico and adjacent Central America. Polygyra (Linisia) albicostulata Pilsbry 1896Polygyra albicostulata Pilsbry 1896; Proc. Acad. Nat. Sci. Linisia albicostulata (Pilsbry). Emberton 1995; Malacologia 37:90. Polygyra (Linisia) albicostulata Pilsbry. Thompson 2008:826.Type Locality.Mxico. Holotype presumed to be in the ANSP. Distribution.NAYARIT: Drift debris of the Rio Grande de Santiago at Yago. Polygyra (Linisia) anilis (Gabb 1865)Helix anilis (shell).Fischer & Crosse 1872; Miss. Sci. Mex. I:269. Polygyra analis (Gabb). Tryon 1867; Amer. Jour. Conch. 3:158, Amer. I:172.Pilsbry 1956; Proc. Acad. Nat. Sci. Phila. 108:27; Linisia anlis (Gabb). Emberton 1995; Malacologia 37:90. Helix solidens Mabille 1895; Bull. Soc. Philomath. Paris, (8), 7:65. Polygyra (Linisia) anilis (Gabb). Thompson 2008:826.Type Localities. Helix anilis: Near Guaymas, Sonora, Mxico; lectotype: ANSP 33447a (H. B. Baker 1963:241). Helix solidens: south of the village of Guaymas, Sonora, Mxico. Distribution.Coastal lowland of Sonora. SONORA: near Ortiz Station; Pitahaya; numerous places along highway from Cameja to Navojoa. Polygyra (Linisia) ariadnae (Pfeiffer 1848)Helix ariadnae Pfeiffer 1848; Zeitschr. f. Malak., 120.Pfeiffer, in Martini & Chemnitz, Syst. Conch. Cab., (2 ed.), Helix I:372; Helix (Polygyra ) texasiana Pfeiffer. Fischer & Crosse 1872; Miss. Polygyra ariadnae (Pfeiffer). Binney 1869; Terr. Air-breath. Moll. 80:117.Pilsbry 1956; Proc. Acad. Nat. Sci. Phila. 108:27. Daidalocheila ariadnae (Binney). Emberton 1995; Malacologia 37:91. Helix couchiana Lea 1857; Proc. Acad. Nat. Sci. Phila. 9:102.Lea 1857; Obs. Genus Unio Polygyra (Linisia) ariadnae (Pfeiffer). Thompson 2008:826.Type Localities. Helix ariadnae: not given. Helix couchiana : Texas. Distribution.TAMAULIPAS: 1 mi. N of Jimenz; Mesa de Solis, near La Lajilla, between Padilla and Jimenz; km 155, 9 km SW of San Fernando; Matamoras (UF). Polygyra (Linisia) aulacomphala Pilsbry & Hinkley 1907Polygyra aulacomphala Pilsbry & Hinkley 1907; Nautilus 21:38 Phila. 108:29. Linisia aulacomphala (Pilsbry & Hinkley). Emberton 1995; Malacologia 37:90. Polygyra (Linisia) aulacomphala Pilsbry & Hinkley. Thompson 2008:927.Type Locality.River drift of the Rio Panuco at Tampico, Tamaulipas, Mxico. Holotype ANSP 96595. Distribution.TAMAULIPAS: known only from river drift shells from the type locality, and from river drift of the Rio Temes, W of Maquiscatzn. Polygyra (Linisia) behri (Gabb 1865)Helix (Polygyra ) behrii Gabb 1865; Amer. Jour. Conch. I:208; pl. I:276. Daedalichila behrii (Gabb). Tryon 1867; Amer. Jour. Conch. 3:64, Polygyra behri (Gabb). Von Martens 1892; Biol. Cent. Amer.:171.(shell). Linisia behri (Gabb). Emberton 1995; Malacologia 37:90. Polygyra (Linisia) behri (Gabb). Thompson 2008:828.Type Locality.Near Guaymas, Sonora, Mxico. Holotype ANSP 33448. Distribution.Coastal regions of northwestern Mxico. SINALOA: Topolobampo; Rio Fuerte, San Blas. SONORA: Alamos; Jori; Pitahaya; northern foothills of Cerro Zaporxa, E of Cajeme; south from Cameje to Navojoa on the Rio Mayo. Polygyra (Linisia) bicruris bicruris (Pfeiffer 1857)Helix bicruris Pfeiffer 1857; Proc. Zool. Soc. Lond. 25:109.Pfeiffer 1859; Monogr. Helic. Viv., 4:315. Helix (Polygyra ) bicruris Pfeiffer. Fischer & Crosse 1872; Miss. Polygyra bicruris (Pfeiffer). Von Martens 1892; Biol. Cent. Linisia bicruris (Pfeiffer). Emberton 1995; Malacologia 37:90. Polygyra (Linisia) bicruris bicruris (Pfeiffer). Thompson 2008:828.Type Locality.Mexico. Distribution.DURANGO: Ventanas. NAYARIT: Islas

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 251 Maras. SONORA: Arroyo San Rafael near San Bernardo; Arroyo Gochico ca. 7 km from San Bernard, 300 m alt. Fischer & Crosse (1872) recorded this species from Chiapas. Both records are unlikely. Polygyra (Linisia) bicruris insculpta Pilsbry 1956Polygyra bicruris insculpta Pilsbry 1956, Proc. Acad. Nat. Sci. Polygyra (Linisia) bicruris insculpta Pilsbry. Thompson. 2008:828.Type Locality.Hill in the coastal low land between Yago and the Rio Santiago, Nayarit, Mxico; 500 ft. alt. Holotype ANSP 166441. Distribution.Known only from the type locality. Polygyra (Linisia) cantralli Solem 1957Polygyra cantralli Solem 1957; Notulae Naturae (298):9; pl. 1, Linisia cantralli (Solem). Emberton 1995; Malacologia 37:90. Polygyra (Linisia) cantralli Solem. Thompson 2008:829.Type Locality.Sierra Autln, about 20 miles southeast of Autln, Jalisco, Mxico; 8200 ft. alt. Holotype UMMZ 185506. Distribution.Known only from the type locality. Polygyra (Linisia) euglypta Pilsbry 1896Polygyra euglypta Pilsbry 1896; Proc. Acad. Nat. Sci. Phila. 48:18; Phila. 108:25. Linisia euglypta (Pilsbry). Emberton 1995; Malacologia 37:90. Polygyra (Linisia) euglypta Pilsbry. Thompson 2008:829.Type Locality.Sinaloa. Lectotype: ANSP 33462 (H. B. Baker 1963:241). Distribution.SINALOA: hills around Pnuco, ca. 50 mi. from Mazatln 1800 ft. alt. (Pilsbry 1956). Polygyra (Linisia) hertleini Haas 1961Polygyra hertleini (shell). Linisia hertleini (Haas). Emberton 1995; Malacologia 37:90. Polygyra (Linisia) hertleini Haas. Thompson 2008:829.Type Locality.Tenecatita Bay, Jalisco, Mxico. Holotype: FMNH 106703). Distribution.Known only from the type locality. Polygyra (Linisia) hindsi hindsi (Pfeiffer 1845)Helix hindsi Pfeiffer 1845; Proc. Zool. Soc. Lond. 13:132.Pfeiffer 1845, in Martini and Chemnitz, Syst. Conch. Cab., Helix Conch. Icon., var. hindsi (Pfeiffer). Von Martens 1892; Biol. Cent. Amer.:169.Pilsbry 1920; Proc. Acad. Nat. Sci. Phila.:192. Polygyra hindsi (Pfeiffer). Pilsbry 1956; Proc. Acad. Nat. Sci. Phila. Naturae (298):9. Linisia hindsi (Pfeiffer). Emberton 1995; Malacologia 37:90. Polygyra (Linisia) hindsi hindsi (Pfeiffer). Thompson 2008:829.Type Locality.Texas. Restricted to Tepc, Nayarit, Mxico (Pilsbry 1956). Distribution.JALISCO: Actopan; Chapala; Guadalajara. NAYARIT: banks of the Rio Tepc, ca. 5 km below Tepc; 20 mi. E of Yago. SINALOA: along Rio Fuerte, at San Bls. Polygyra (Linisia) hindsi guadalajarensis Pilsbry 1910Polygyra guadalajarensis Pilsbry 1910; Proc. Acad. Nat. Sci. Phila. Polygyra hindsi guadalajarensis Pilsbry. Pilsbry 1956; Proc. Acad. Nat. Sci. Phila. 108:24. Polygyra (Linisia) hindsi guadalajarensis Pilsbry. Thompson 2008:830:Type Locality.Guadalajara, Jalisco, Mxico. Holotype: ANSP 99513. Distribution.Known only from the type locality. Polygyra (Linisia) hindsi heteroea Pilsbry 1956Polygyra hindsi heteroea Pilsbry 1956; Proc. Acad. Nat. Sci. Phila. Polygyra (Linisia) hindsi heteroea Pilsbry. Thompson 2008:830.Type Locality.Bank of the Laguna Santa Maria, Nayarit, Mxico; at about 2600 ft. alt. Holotype: ANSP 166188. Distribution.Known only from the type locality. Polygyra (Linisia) implicata (Von Martens 1865)Helix implicata Beck Von Martens 1865; Malak. Bl., 12:20.12e (shell). Polygyra implicata (Von Martens). Strebel 1880; Beitrag, 4:47 Polygyra oppilata implicata (Von Martens). Pilsbry 1910; Proc. Acad. Nat. Sci. Phila. (1909):543. Millerelix ? implicata (Von Martens). Emberton 1995; Malacologia 37:91. Polygyra (Linisia) implicata (Von Martens). Thompson 2008:830.Type Locality. Rio Tecoluta, near the shore, about 21 N Lat., Veracruz, Mxico. Distribution.Eastern Mxico from central Veracruz north to Tamaulipas. NUEVO LEN: 2.8 mi. N of Santiago. SAN LUS POTOS: various places around Valles; Mecos; San Dieguito; Tamazunchale; Canoas; Ignacio Agua Buena. TAMAULIPAS: Tampico; drift of Rio Temes W of Magiscatzin; 4 mi. S of Villagrn. VERACRUZ: Agua Caliente; Rancho Guerrero; Camino de Arroyo Hondo; Camino de Obispo; Misantla; Barranca de Mahuistlan between Jalapa and Mirador; San Juan Miahuatlan; Jalapa; Veracruz. Polygyra ( Linisia) matermontana matermontana Pilsbry 1896Polygyra matermontana matermontana Pilsbry 1896; Proc. Acad. Clapp 1909; Nautilus 22:114.Solem 1954; Nautilus 68:6.

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252 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Linisia matermontana (Pilsbry). Emberton 1995; Malacologia 37:90. Polygyra (Linisia) matermontana matermontana Pilsbry. Thompson 2008:831.Type Locality.Sierra Madre, Colima, Mxico. Holotype: ANSP 834a. Distribution.COLIMA: Sierra Madre. GUERRERO: Balsas, in limestone rocks 1000 ft. above the Rio Balsas; Taxco. Polygrya (Linisia) matermontana jaliscoensis Pilsbry 1910Polygrya matermontana jaliscoensis Pilsbry 1910; Proc. Acad. Nat. Sci. Phila. (1909):541.Pilsbry 1920; Proc. Acad. Nat. Sci. Phila. 72:192. Polygrya (Linisia) matermontana jaliscoensis Pilsbry. Thompson 2008:831: Type Locality.Guadalajara, Jalisco, Mxico. Holotype: ANSP 99512. Distribution.Known only from the type locality. Polygyra (Linisia) nelsoni Dall 1897Polygyra nelsoni Dall 1897; Nautilus 11:74.Solem 1957; Notulae Polygyra nelsoni var. colisella Dall 1897; Nautilus 11:74. Linisia nelsoni (Dall). Emberton 1995; Malacologia 37:90. Polygyra (Linisia) nelsoni Dall. Thompson 2008:832.Type Localities. Polygyra nelsoni: near San Sebastian [between Milpillas and San Sebastian], Jalisco, Mxico; 3900 ft. alt.; holotype USNM 190921. Polygyra nelsoni var. colisella : not stated; syntypes USNM 190922. Distribution.JALISCO: La Laguna; vicinity of San Sabastian. Polygyra (Linisia) oppilata (Morelet 1849)Helix oppilata Morelet 1849; Test. Nov., I:8.Fischer & Crosse Polygyra oppilata (Morelet). Strebel 1880; Beitrag, IV:49.Von Martens 1892; Biol. Cent. Amer.:164.Pilsbry 1956; Proc. Poligyra oppilata complicata Pilsbry 1910; Proc. Acad. Nat. Sci. Phila. 61:543. Daedalocheila oppilata (Morelet). Emberton 1995; Malacologia 37:91. Polygyra (Linisia) oppilata (Morelet). Thompson 2008:832.Type Localities. Helix oppilata : Bequaert and Clench [1933:532] restricted to the type locality of Campeche [Littora Yucatanea]; no Campeche record is otherwise reported. Poligyra oppilata complicata : south of Valles, San Lus Potos; lectotype: ANSP 99003a (H. B. Baker 1963:241). Distribution.The species is known from northern Veracruz and adjacent regions of Tamaulipas and San Lus Potos. SAN LUS POTOS: mountains near Valles. TAMAULIPAS: foothills near Gonzalez; Tampico. VERACRUZ: Veracruz. Polygyra (Linisia) pergrandis Solem 1959Polygyra pergrandis Solem 1959; Occ. Pap. Mus. Zool. Univ. Mich. Linisia pergrandis (Solem). Emberton 1995; Malacologia 37:90. Polygyra (Linisia) pergrandis Solem. Thompson 2008:833.Type Locality.Cerro de Barolosa, north of Coalcomn, Sierra de Coalcomn, Michoacn; 9000 ft. alt. Holotype: UMMZ 191180. Distribution.MICHOACN: known only from Cerro de Barolosa above 7500 ft. alt. Polygyra (Linisia) polita Pilsbry & Hinkley 1907Polygyra polita 11 (shell).Pilsbry 1928; Proc. Acad. Nat. Sci. Phila. 80:117.Pratt 1981; Unpubl. thesis:80. Polygyra scintilla Pilsbry & Hubricht 1956; Nautilus 69:94; pl. Linisia polita (Pilsbry & Hinkley). Emberton 1995; Malacologia 37:90. Polygyra (Linisia) polita Pilsbry & Hinkley. Thompson 2008:833.Type Localities.Polygyra polita : in river drift at Tampico, Tamaulipas, Mxico; holotype: ANSP 96597. Polygyra scintella: along a railroad 1.5 miles north of Raymondville, Willacy Co., Texas; holotype: ANSP 196560. Distribution.Extreme southern Texas and northeastern Mxico. NUEVO LEN: river drift, Sabinas Hidalgo. TAMAULIPAS: loess 1.4 mi. S of Ciudad. Mier; drift, Rio San Fernando, San Fernando; 9 mi. SW of Santa Teresa; Mesa de Sols, near La Lajilla, between Padilla and Jimenz. Polygyra (Linisia) ponsonbyi Pilsbry 1896Polygyra ponsonbyi Pilsbry 1896; Proc. Acad. Nat. Sci. Phila.:17; Phila. 108:22. Linisia ponsonbyi (Pilsbry). Emberton 1995; Malacologia 37:90. Polygyra (Linisia) ponsonbyi Pilsbry. Thompson 2008:833.Type Locality.Mxico; holotype: ANSP 67808. Distribution.Unknown. Polygyra (Linisia) rhoadsi Pilsbry 1899Polygyra rhoadsi Pilsbry 1899; Proc. Acad. Nat. Sci. Phila.:393.(shell).Pilsbry 1956; Proc. Acad. Nat. Sci. Phila. 108:29; 41:685. Millerelix rhoadsi (Pilsbry). Emberton 1995; Malacologia 37:91. Polygyra (Linisia) rhoadsi Pilsbry. Thompson 2008:834.Type Locality.Topo Chico, near Monterey, Nuevo 1963:241). Distribution.NUEVO LEN: Topo Chico, near Monterrey; subfossil near an arroyo 4 mi. NW of Pesqueria Chica; 13 mi. W of Santa Catarina; Santiago. Polygyra (Linisia) richardsoni Von Martens 1892Polygyra richardsoni Von Martens 1892; Biol. Cent. Amer.:168, Sci. Phila. 108:24. Polygyra richardsoni var. lingualis Pilsbry 1899; Nautilus 12:144. Polygyra richardsoni form paucicostata Dall 1926; Proc. Calif.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 253 Linisia richardsoni (Von Martens). Emberton 1995; Malacologia 37:90. Polygyra (Linisia) richardsoni Von Martens. Thompson 2008:834.Type Localities.Poligyra richardsoni: Mazatln, Sinaloa, Mxico. Poligyra richardsoni var. lingualis: Rosaria, near Mazatln, Sinaloa, Mxico; lectotype: ANSP 73999a (H. B. Baker 1963:241). Polygyra richardsoni form paucicostata : Islas Maras, Nayarit, Mxico; lectotype: CAS 2211 (H. B. Baker 1963:241). Distribution.COLIMA: Colima. NAYARIT: Islas Maras. SINALOA: many localities in the vicinity of Mazatln. Polygyra (Linisia) suprazonata Pilsbry 1899Polygyra suprazonata Pilsbry 1899; Proc. Acad. Nat. Sci. Phila.:393.Pilsbry 1903; Proc. Acad. Nat. Sci. Phila.:763; Phila. 108:22.Solem 1959; Occ. Pap. Mus. Zool. Univ. Mich. (611):9. Linisia suprazonata (Pilsbry). Emberton 1995; Malacologia 37:90. Polygyra (Linisia) suprazonata Pilsbry. Thompson 2008:835.Type Locality.Tzintzuntzn, Michoacn, Mxico. Lectotype ANSP 77125 (H. B. Baker 1963:241). Distribution.MICHOACN: 1 mi. N of San Pedro Damian, Sierra de Coalcomn; 7.5 mi. N of Uruapn, 6800 ft. alt.; 9.5 mi. N of Carapn, 6800 ft. alt.; Tzintzuntzn. Polygyra (Linisia) tamaulipasensis (Lea 1857)Helix tamaulipasensis Lea 1857; Proc. Acad. Nat. Sci. Phila. 9:102.Lea 1864; Obs. Gen. Unio Polygyra texasiana tamaulipasensis (Lea). Pilsbry & Hubricht; 1956; Nautilus 69:95. Polygyra tamaulipasensis (Lea). Hubricht 1961; Nautilus 75:27. Linisia tamaulipasensis (Lea). Emberton 1995; Malacologia 37:90. Polygyra (Linisia) texasiana texasiana (Moricand). Thompson 2008:835.Type Localities. Helix tamaulipasensis: Tamaulipas, Mxico; holotype USNM. Distribution.Central Texas south to northern Mxico near the United States border. Polygyra (Linisia) texasiana texasiana (Moricand 1833)Helix (Helicodonta) texasiana Moricand 1833; Mem. Soc. Phys. et Hist. de Gneve 6:538. Polygyra texasiana (Moricand). Pilsbry 1940; Land Moll. N. Amer. (shell). Linisia texasiana (Moricand). Emberton 1995; Malacologia 37:90. Triodopsis tridonta Beck 1837; Index Moll.:22 (new name for texasiana Moricand [Pilsbry 1940:617]). Polygyra texasiana tillandsiae Cockerell 1917; Nautilus 21:36. Polygyra (Linisia) texasiana texasiana (Moricand). Thompson 2008:835.Type Localities. Helix texasiana : Pilsbry (1940) restricted the type locality to Camaron Co., Texas. Pratt (1981) suggested that it should be San Antonio, Texas. Syntypes in the Vienna Nuturforschende Museum. Polygyra texasiana tillandsiae : San Benito, Cameron Co., Texas; holotype ANSP 116250. Distribution.Northern part of Tamaulipas, Nuevo to southwestern Missouri, western and southern Arkansas, and western Mississippi. It has been introduced elsewhere in the United States (Pratt 1981). COAHUILA: 9.5 km 244871). TAMAULIPAS: mesquite forest, 35 km NW Cruillas Polygyra (Linisia) texasiana texasensis Pilsbry 1902Polygyra texasensis Pilsbry 1902; Nautilus 16:31. Polygyra texasiana texasensis Pilsbry. Pilsbry & Ferriss 1906; Proc. Polygyra texasiana hyerolia Pilsbry & Ferriss 1906; Proc. Acad. Polygyra (Linisia) texasiana texasensis Pilsbry. Thompson 2008:836.Type Locality.Polygyra texasensis: Texas, Mitchell Co., Colorado City; holotype ANSP 38258. Polygyra texasiana hyperolia: high mesa west of Devils River, Texas; holotype ANSP 91363. Distribution.Southwestern Texas and immediately adjacent Mxico. COAHUILA: bluff opposite Amistad 244871). Polygyra (Linisia) (Pfeiffer 1845) Pfeiffer 1845; Proc. Zool. Soc. Lond. 13:131.Pfeiffer in Martini & Chemnitz; Conch. Cab., Helix 1:373; pl. (Pfeiffer). Von Martens 1892; Biol. Cent. Nat. Sci. Phila.:192.Dall 1926; Proc. Calif. Acad. Sci. (4) 15:476.Pilsbry 1956; Proc. Acad. Nat. Sci. Phila. 108:21; (Pfeiffer). Emberton 1995; Malacologia 37:90. Polygyra nayarta Pilsbry & Cockerell 1925; Proc. Acad. Nat. Sci. Polygyra (Linisia) (Pfeiffer). Thompson 2008:837.Type Localities. : Mxico. Restricted to Tepc, Nayarit, Mxico (Pilsbry 1956:22). Polygyra nayarita: Nayarit. Holotype ANSP 139841. Distribution.Known for certain only from western Mxico in the States of Jalisco and Nayarit. JALISCO: Chapala. NAYARIT: Islas Maras; Isla Mara Madre; banks of Rio Tepc 5 km below Tepc; Cerro San Juan ca. 3 km SSW of Tepc, 4500 ft. alt.; Yago, on the Rio Grande SE Santiago. Subgenus Upsilodon Pilsbry 1930Upsilodon Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. 82:315.Pilsbry 1940; Land Moll. N. Amer. I:637.Emberton 1995; Malacologia 37:91.Type Species.Helix hippocrepis Pfeiffer 1848.

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254 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1) Distribution.South-central Texas and northern Coahuila, Mxico. Taxonomy.Six species are recognized. Five occur in Mxico. Polygyra (Upsilodon) burlesoni Metcalf & Riskind 1978Polygyra burlesoni Metcalf & Riskind 1978; Proc. Biol. Soc. Wash. Daedalochila (Upsilodon) burlesoni (Metcalf & Riskind). Emberton 1995; Malacologia 37:91. Polygyra (Upsilodon) burlesoni Metcalf & Riskind. Thompson 2008:837.Type Locality.Caon Bonito at extreme western end of Caon Zorra, Serranias del Burro, Coahuila, Mxico 751886. Distribution.COAHUILA: known only from the immediate vicinity of the type locality. Polygyra (Upsilodon) dalli Metcalf & Riskind 1978Polygra dalli Metcalf & Riskind 1978; Proc. Biol. Soc. Wash. Daedalochila (Upsilodon) dalli (Metcalf & Riskind). Emberton 1995; Malacologia 37:91. Polygyra (Upsilodon) dalli Metcalf & Riskind. Thompson 2008:838.Type Locality.Sierra de Guadalupe, Coahuila, Mxico; 9500 ft. alt. Holotype USNM 758554. Distribution.Known only from the type locality. Polygyra (Upsilodon) idiogenes Pilsbry 1956Polygyra idiogenes Pilsbry 1956; Proc. Acad. Nat. Sci. Phila. Linisia indiogenes (Pilsbry). Emberton 1995; Malacologia 37:90. Polygyra (Upsilodon) idiogenes Pilsbry. Thompson 2008:838.Type Locality.San Lorenzo Canyon, southeast of Saltillo, Coahuila, Mxico; 7200 ft. alt. Holotype ANSP 164589. Distribution.Known only from the type locality. Polygyra ( Upsilodon) multiplicata Metcalf & Riskind 1978Polygyra multiplicata Metcalf & Riskind 1978; Proc. Biol. Soc. (genitalia). Daedalochila (Upsilodon) multiplicata (Metcalf & Riskind). Emberton 1995; Malacologia 37:91. Polygyra (Upsilodon) multiplicata Metcalf & Riskind. Thompson 2008:238.Type Locality.Caon Bonito at extreme western end of Caon Zorra, Serranias del Burro, Coahuila, 1675 m alt. Distribution.COAHUILA: known only from the immediate vicinity of the type locality. Polygyra (Upsilodon) sterni Metcalf & Riskind 1978Polygyra sterni Metcalf & Riskind 1978; Proc. Biol. Soc. Wash. Daedalochila (Upsilodon) sterni (Metcalf & Riskind). Emberton 1995; Malacologia 37:91. Polygyra (Upsilodon) sterni Metcalf & Riskind. Thompson 2008:839.Type Locality.Lower north slope of the extreme north end of the Sierra San Vicente, Mcpo. Sacremento, immediately north of Fed. Hwy. 30, 2.5 km east of Boquillos, a village between Nadadores and Cuatro Cienegas, Coahuila 758821. Distribution.Known only from the type locality. Polygyra species of uncertain subgeneric status Polygyra chiapensis (Pfeiffer 1856)Helix chiapensis Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:377.Helix (Polygyra ) chiapensis (Pfeiffer).Fischer & Crosse 1872: Miss. Sci. Mex. I:281. Polygyra chiapensis (Pfeiffer).Von Martens 1892; Biol. Cent. Amer.:165.Thompson 2008:839.Type Locality.Chiapas. Distribution.Unknown. Polygyra couloni (Shuttleworth 1852)Helix couloni Shuttleworth 1852; in Bern, Mittheil. (Diagn. 2, p. 17) 197. Polygyra couloni (Shuttleworth). Strebel 1880; Beitrag, IV:46; pl. 2008:839. Linisia couloni (Shuttleworth). Emberton 1995; Malacologia 37:90.Type Locality.Cordova, Veracruz, Mxico. Syntype: Naturhistorisches Museum Bern 19045/1 h (Neubert & Gosteli 2003). Distribution.Trans-Mxico south of the plateau. COLIMA: Colima. GUERRERO: Omilteme. JALISCO: Jalapa; Maltrata; Orizaba. Polygyra dissecta Von Martens 1892Polygyra dissecta Von Martens 1892; Biol. Cent. Amer.,:167; pl. 7, Linisia dissecta (Von Martens). Emberton 1995; Malacologia 37:90.Type Locality.Toluca, Mxico. Distribution.Known only from the type locality. Polygyra dysoni (Shuttleworth 1852)Helix dorfeuilleana Lea. Pfeiffer 1848; Mon. Helic. Viv., 1:410.Pfeiffer in Martini and Chemnitz, Conch. Cab., Helix, 1:377; Helix dysoni Shuttleworth 1852; in Bern, Mittheil. (Diagn. 2, p. 16) 196. Polygyra dysoni (Shuttleworth). Von Martens 1892; Biol. Cent. Amer.:165.Neubert & Gosteli 2003; Contributions to Natural Linisia dysoni (Shuttleworth). Emberton 1995; Malacologia 37:90.Type Locality.Honduras. Here restricted to Belize. Syntype: Naturhistorisches Museum Bern 15276/1 (Neubert

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 255 & Gosteli 2003). Distribution.Unknown. Polygyra plagioglossa (Pfeiffer 1859)Helix plagioglossa Pfeiffer 1859; Proc. Zool. Soc. Lond. 27:26; pl. Helix (Polygyra ) plagioglossa (Pfeiffer). Fischer & Crosse 1872; Polygyra plagioglossa (Pfeiffer). Strebel 1880; Beitrag, 4:45; pl. Thompson 2008:841. Linisia plagioglossa (Pfeiffer). Emberton 1995; Malacologia 37:90.Type Locality.Oaxaca. Distribution.Puebla and Oaxaca (Von Martens 1892). Polygyra yucatanea (Morelet 1849)Helix yucatanea Morelet 1849; Test. Noviss. I:9.Pfeiffer 1853; Mon. Helic. Viv., 3:263.Folin 1865; Jour. de Conchyl. 13:68 (living animal, eggs). Helix (Polygyra ) yucatanea Morelet. Fischer & Crosse 1872; Miss. Polygyra yucatanea (Morelet). Von Martens 1892; Biol. Cent. Amer.:166.Bequaert & Clench 1933; Publ. Carnegie Inst. Wash., (431):531.Goodrich & van der Schalie 1937; Misc. Publ. Mus. Zool. Univ. Mich. (34):27.Rehder 1966; Proc. Biol. Soc. Wash. 79:280.Thompson 1957; Nautilus 70:100.Thompson 2008:841. Linisia yucatanea (Morelet). Emberton 1995; Malacologia 37:90. Helix helictomphala Pfeiffer 1856; Proc. Zool. Soc. Lond. 24:377.Pfeiffer 1858; Mon. Helic. Viv., 4:314. Helix (Polygyra ) helictomphala Pfeiffer. Fischer & Crosse 1872; Polygyra helicomphala (Pfeiffer). Strebel 1880; Beitrag, 4:46; pl. Polygyra yucatanea var. helictomphala (Pfeiffer). Von Martens 1892; Biol. Cent. Amer.:199.Type Localities. Helix yucatanea : Yucatan. Rehder (1966) incorrectly stated that Bequaert and Clench (1933) restricted the type locality to Isla de Carmen, Campeche, Mxico. Helix helictomphala : Chiapas, Mxico. Distribution.CAMPECHE: Isla de Carmen, at Laguna de Terminos. QUINTANA ROO: Icaich. TABASCO: GUATEMALA, Dept. Petn: El Paso de Los Caballos; Laguna Perdido. Genus Praticolella Von Martens 1892Praticola Strebel 1880; Beitrag, IV:38. (Not Praticola Kaup 1837, Aves; not Swainson 1837, Aves;. not Fatio 1867, Mammalia.) Praticolella Von Martens 1892; Biol. Cent. Amer.:138.Pilsbry 1895; Man. Conch. 9:67.Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. 82:318.Pilsbry 1940; Land Moll. N. Amer. 1:688.Emberton 1995; Malacologia 37:90.Type Species.Praticola ocampi Strebel 1880 (= Helix ampla Pfeiffer 1866). Distribution.Southeastern United States and eastern Mxico south to Nicaragua. Taxonomy.The genus contains four subgenera. Two occur in Mxico and Central America. Subgenus Praticolella Von Martens 1892 Distribution.Texas and Arkansas south to Nicaragua. Taxonomy.Eight species are recognized. Six occur in Mxico and Central America. Praticolella (Praticolella ) ampla (Pfeiffer 1866)Helix ampla Pfeiffer 1866; Malak. Bltt. 13:78. Helix (Patula ) ampla Pfeiffer. Fischer & Crosse 1880; Miss. Sci. Mex. 1:227. Praticola ocampi Helix (Praticolella) ampla Pfeiffer. Von Martens 1892; Biol. Cent. Amer.:138. Praticolella ampla (Pfeiffer). Pilsbry 1896; Nautilus 10:59.Emberton 1995; Malacologia 37:90.Thompson 2008:843.Type Locality.Mirador, Veracruz, Mxico. Distribution.VERACRUZ: Almolonga; Jalapa; La Banderilla; Monte de Tataquicapa; Dos Arroyos; Consolapa, near Coatepec; San Antonia del Monte; Mirador; Misantla; Rancho de Guerrero, near Misantla (Von Martens 1892); San Rafael, Jicaltepec (Pilsbry 1896); Praticolella (Praticolella ) berlandieriana (Moricand 1833)Helix berlandieriana Moricand 1833; Mem. Soc. Physique et Helic. Viv., 3:227.Von Martens 1892; Biol. Cent. Amer.:140. Praticolella berlandieriana (Moricand). Pilsbry & Ferriss 1906; 1928; Proc. Acad. Nat. Sci. Phila. 80:117.Pilsbry 1930; Proc. Pilsbry 1956; Proc. Acad. Nat. Sci. Phila. 108:30.CorreaSandoval 1993; Rev. Biol. Trop. 41:685.Emberton 1995; Malacologia 37:90.Correa-Sandoval 1997; Rev. Biol. Trop. 47:140.Correa-Sandoval 2000; Acta Zool. Mex. (79):9.Correa-Sandoval 2002; Acta Zool. Mex. (86):238.Thompson 2008:843.Type Locality.Texas. Distribution.DURANGO: Ventanas. JALISCO: Sayula. NUEVO LEN: Monterrey; Pablillo; Santa Barbara; Santiago; Topo Cicho; between Pablillo valley and Santa Barbara canyon; Santiago; Iturbide (Pilsbry 1956). SAN LUS POTOS: Cd. Valles. TAMAULIPAS: Cd. Victoria; Mesa de Solis, near La Lajilla, between Pablillo and Jimenz. Recorded from numerous localities in northern VERACRUZ, lipas (Correa-Sandoval 1998). Praticolella (Praticolella ) (Pfeiffer 1848) Pfeiffer 1848; Monogr. Helic. Viv., 1:337.

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256 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Helix (Leptarionta) Pfeiffer. Fischer & Crosse, Miss. Sci. Mex. 1:255; Helix (Arionta) Pfeiffer. Von Martens 1892; Biol. Cent. (Pfeiffer). Strebel 1880; Beitrag, IV:41; pl. 13, (Pfeiffer 1848). Emberton 1995; Malacologia 37:90.Thompson 2008:844.Type Locality.Papantla, Veracruz, Mxico. Distribution.Known only from the type locality. Praticolella (Praticolella ) griseola (Pfeiffer 1841)Helix griseola Pfeiffer 1841; Symb. Hist. Hel.., 1:40. Helix berlandieriana var. griseola Pfeiffer. Von Martens 1892; Biol. Praticolella griseala (Pfeiffer). Pilsbry 1896:59.Pilsbry 1940; & McCoy 1965; Univ. Col. Stud. Biol., (13):1.Neck 1977:1.Emberton 1995; Malacologia 37:90.CorreaSandoval 1999; Acta Zool. Mex. (79):9.Correa-Sandoval & Rodriguez 2002; Acta Zool. Mex. (86):238.Prez & Lopz 2002:191.Correa-Sandoval & Salizar 2005; Acta Zool. Mex. 21:61.Thompson 2008:845.Type Locality.Vera Cruz, Veracruz, Mxico. Distribution.Naturally occurring from Tamaulipas south to Nicaragua. Widely introduced into tropical areas elsewhere. Recorded from numerous localities in southern TAMAULIPAS and northern VERACRUZ (Correa-Sandoval 1999 2002). Other records follow. CAMPECHE: 8.1 mi. SW of (Branson & McCoy 1965). SAN LUS POTOS. Cd. Valles. Flores, Petn. NICARAGUA: Volcn de Masaya; Masapa, Boaco Dept. (Von Martens 1901); Grenada; San Ubaldo; (Pres & Lopz 2002). Praticolella (Praticolella ) strebeliana Pilsbry 1899Praticolella strebeliana Pilsbry 1899; Proc. Acad. Nat. Sci. Phila. 51:394.Pilsbry & Ferriss 1904; Proc. Acad. Nat. Sci. Phila. 37:90.Thompson 2008:845.Type Locality.Diente, near Monterey, Nuevo Lon; holotype: ANSP 77128a. Distribution.Known only from the type locality. Praticolella (Praticolella ) taeniata Pilsbry 1940Praticolella berlandieriana taeniata Pilsbry 1940; Land Moll. N. Praticolella taeniata Pilsbry. Emberton 1995; Malacologia 37:90.Thompson 2008:825.Type Locality.Brownsville, Texas. Distribution.Southeastern Texas; Tamaulipas, Nuevo Subgenus Eduardus Pilsbry 1930Eduardus Pilsbry 1930; Proc. Acad. Nat. Sci. Phila. 82:315.Pilsbry 1936; Nautilus 21:140.Emberton 1995; Malacologia 37:90.Type Species.Polygyra martensiana Pilsbry 1907. Distribution.Northeastern Mxico. Taxonomy.The subgenus contains a single species. Praticolella (Eduardus ) martensiana (Pilsbry 1907)Polygyra (?) martensiana Pilsbry 1907; Nautilus 21:26; pl. 9, Praticolella martensiana (Pilsbry). Pilsbry 1936; Nautilus 49:140.Pilsbry 1956; Proc. Acad. Nat. Sci. Phila. 108:30.Emberton 1995; Malacologia 37:90.Correa-Sandoval, Garca-Cubas & Reguero 1998; Acta Zool. Mex. (73):17.Correa-Sandoval 2000; Acta Zool. Mex. (79):9. Praticolella (Eduardus) martensiana (Pilsbry). Thompson 2008:846.Type Locality.Tampico, Tamaulipas, in river drift. Lectotype ANSP 94798a (H. B. Baker 1963:241). Distribution.SAN LUIS POTOS: Ignacio Agua Buena, near Tomasopo; Rio Gallinas, Ejid. Carpintero 1999). TAMAULIPAS: Tampico. VERACRUZ: El Bajia, carretera Naranjos-Tuxpan; La Ordea, Papantla (CorreaSandoval 2000). Family THYSANOPHORIDAE Pilsbry 1926 Distribution.Middle America from Arizona and New Mexico south to Colombia and Venezuela; the Caribbean region; south Florida. Taxonomy.Five genera are recognized. Three genera are found in the study area. One genus, McCleania Bequaert & Clench 1939, is found on Hispaniola and Puerto Rico. Another genus, Microphysala Pilsbry 1926, is found in western North America. Genus Itzamna Pilsbry 1926Itzamna Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:11213.Type Species.Helix sigmoides Morelet 1851. Distribution.Guatemala. Taxonomy.Itzamna is tentatively placed in the Thysanophoridae. A single species is recognized. Itzamna sigmoidius (Morelet 1851)Helix sigmoides Morelet 1851; Test. Noviss. II:9.Fischer & Crosse 1880:38. Tristam 1863; Proc. Zool. Soc. Lond. 31:411. Itzamna signoidius (Morelet). Pilsbry 1926 Proc. Acad. Nat. Sci. Phila. 78:11213.Thompson 2008:847.Type Locality.Vera Paz, Guatemala. Distribution.Known only from the type locality. Genus Microconus Strebel & Pfeffer 1880Microconus Strebel & Pfeffer 1880:29.H. B. Baker 1927; Proc. Acad. Nat. Sci. Phila. 79:236.Thompson 1958; Nautilus 72:5.Type Species.Helix wilhelmi Pfeiffer 1866.

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THOMPSON: Checklist of land and freshwater snails of Mxico and Central America 257 Distribution.Panam north to Puebla, Mxico. Taxonomy.Two subgenera and four species are recognized. Subgenus Microconus Strebel & Pfeffer 1880 Distribution.Guatemala and southeastern Mxico. Taxonomy.Two species are recognized. Microconus (Microconus) rufus Thompson 1958Thysanophora conspercatella (Morelet). Goodrich & van der Schalie 1937; Misc. Pub. Mus. Zool. Univ. Mich. (34):26. Microconus rufus B (shell). Microconus (Microconus) rufus Thompson. Thompson 2008:847.Type Locality.A knoll along the Santa Ana Road, 2 km south of Puebla Nueva, Dept. Petn, Guatemala. Holotype UMMZ 64416. Distribution.GUATEMALA, Dept, Petn: various localities (Thompson 1958). Microconus (Microconus) wilhelmi (Pfeiffer 1866)Helix wilhelmi Pfeiffer 1866; Malak. Bltt. 13:79. Microconus wilhelmi (Pfeiffer). Strebel & Pfeffer 1880:29; pl. Patula wilhelmi (Pfeiffer). Von Martens 1892; Biol. Cent. Amer.:128. Type Locality.Mirador, Veracruz, Mxico. Distribution.PUEBLA: near Necaxa (H. B. Baker 1927). VERACRUZ: Mirador. Subgenus Pulchriconus Thompson 1958Pulchriconus Thompson 1958; Nautilus 72:9.Type Species. Microconus (Pulchriconus) pilsbryi Thompson 1958. Distribution.Nicaragua, Panam. Taxonomy.Two species are recognized. Microconus (Pulchriconus) pilsbryi Thompson 1958Microconus pilsbryi (reproductive anatomy). Microconus (Pulchriconus ) pilsbryi Thompson. Thompson 2008:248.Type Locality..5 km south of Matagalpa, Dept. Matagalpa, Nicaragua. Holotype UMMZ 193100. Distribution.Known only from the type locality. Microconus (Pulchriconus) termitarum Pilsbry 1926Microconus termitarum Pilsbry 1926, Proc. Acad. Nat. Sci. Phila. 72:9. Microconus (Pulchriconus ) termitarum Pilsbry. Thompson 2008:849.Type Locality.Isla Barro Colorado, Canal Zone, Panam. Holotype ANSP 140824. Distribution.Known only from the type locality. Genus Thysanophora Strebel & Pfeffer 1880Thysanophora Strebel & Pfeffer 1880:30.Tryon 1887, Man. Conch. 3:16.Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:113.H. B. Baker 1927, Proc. Acad. Nat. Sci. Phila. 79:240.Pilsbry 1940, Land Moll. N. Amer. 1:984.Type Species.Helix impura Pfeiffer 1866. Type by subsequent designation; Tryon 1887. Distribution.Continental North America from Arizona south to Venezuela. One species occurs in Cuba. Taxonomy.Four subgenera are recognized. All occur in the study area. Twenty-one species are found in the study area. Subgenus Thysanophora Strebel & Pfeffer 1880 Distribution.Western North America south to Colombia and Venezuela; Cuba. Taxonomy.Ten species are recognized in the study area. As a taxonomic convenience various problematic small species of land snails with a simple aperture have been placed in Thysanophora. The genus as now constituted includes several such species. Thysanophora (Thysanophora) amita Pilsbry 1926Thysanophora amita Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. Thysanophora (Thysanophora) amita Pilsbry. Thompson 2008:849.Type Locality.Juan Minas, Canal Zone, Panam. Holotype ANSP 45259. Distribution.PANAM: Juan Minas; Mount Hope (Pilsbry 1930). Thysanophora (Thysanophora) balboa Pilsbry 1926Thysanophora balboa Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. Thysanophora (Thysanophora) balboa Pilsbry. Thompson 2008:850.Type Locality.Las Cascades, Canal Zone, Panam. Holotype ANSP 101339a (H. B. Baker 1963:240). Distribution.PANAM, Canal Zone: Juan Mina. Las Santos Prov.: Tonosi (Pilsbry 1926); Ruins of Old Panam City (Pilsbry 1930). Thysanophora ( Thysanophora) canalis canalis Pilsbry 1910Thysanophora canalis Pilsbry 1910; Proc. Acad. Nat. Sci. Phila. Univ. Mich. (167):13.Pilsbry 1926; Proc. Acad. Nat. Sci. Phila. 78:70. Thysanophora (Thysanophora) canalis canalis Pilsbry. Thompson 2008:850.Type Locality.Las Cascades, Canal Zone, Panam. Holotype ANSP 101329. Distribution. PANAM: known only from the type locality. VENEZUELA (H. B. Baker 1926). Thysanophora (Thysanophora) canalis cariacoensis Pilsbry 1926Thysanophora canalis cariocoensis Pilsbry 1926; Proc. Acad. Nat.

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258 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 50(1)Sci. Phila. 78:79. Thysanophora (Thysanophora) canalis cariacoensis Pilsbry. Thompson 2008:850.Type Locality.Cariaco, Venezuela. Holotype ANSP 28263a (H. B. Baker 1963:240). Distribution.Known only from the type locality. Thysanophora (Thysanophora) clarionensis Dall,1926Thysanophora clarionensis Dall 1926; Proc. Calif. Acad. Sci. (4) Thysa